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<document id="58D798FFBFAB791961D93A997388CEB5" ID-DOI="10.37828/em.2016.8.5" ID-ISSN="2336-9744" ID-ZooBank="urn:lsid:zoobank.org:pub:88013A3B-3801-45C1-9CEE-3F2487EA8622" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720075178013" checkinUser="felipe" docAuthor="Vas, Zoltán" docDate="2016" docId="038787C2FFA3AF267397FE9F523B4CDF" docLanguage="en" docName="EcolMontenegrina.8.38-44.pdf" docOrigin="Ecologica Montenegrina 8" docSource="https://doi.org/10.37828/em.2016.8.5" docStyle="DocumentStyle:17621BF4B6DAF9FC3B559F0A069E6C5F.2:EcolMontenegrina.2014-2016.journal_article" docStyleId="17621BF4B6DAF9FC3B559F0A069E6C5F" docStyleName="EcolMontenegrina.2014-2016.journal_article" docStyleVersion="2" docTitle="Temelucha flavia Vas, 2016, sp. n." docType="treatment" docUuid="4E1F19CA-028C-4429-B842-EA5FA7072D06" docUuidSource="ZooBank" docVersion="4" lastPageNumber="43" masterDocId="FFBEFFBAFFA2AF237300FFD456044F7C" masterDocTitle="Temelucha flavia sp. n. from the southern Mediterranean region (Hymenoptera: Ichneumonidae: Cremastinae)" masterLastPageNumber="44" masterPageNumber="38" pageNumber="39" updateTime="1723577274176" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<treatment id="038787C2FFA3AF267397FE9F523B4CDF" ID-DOI="http://doi.org/10.5281/zenodo.12652729" ID-Zenodo-Dep="12652729" LSID="urn:lsid:zoobank.org:act:4E1F19CA-028C-4429-B842-EA5FA7072D06" httpUri="http://treatment.plazi.org/id/038787C2FFA3AF267397FE9F523B4CDF" lastPageId="5" lastPageNumber="43" pageId="1" pageNumber="39">
<subSubSection id="C334655FFFA3AF227397FE9F57B04E19" box="[151,436,331,357]" pageId="1" pageNumber="39" type="nomenclature">
@ -183,7 +186,7 @@ has well developed notaulus.
Female (
<typeStatus id="54958876FFA3AF2272A6FC3954144B74" box="[422,528,1005,1032]" pageId="1" pageNumber="39" type="holotype">holotype</typeStatus>
) (
<figureCitation id="13152A51FFA3AF22712CFC3954984B74" box="[556,668,1005,1032]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="2.[432,507,988,1012]" captionStartId-1="2.[432,507,2003,2027]" captionStartId-2="3.[401,476,1007,1031]" captionStartId-3="3.[397,472,1957,1981]" captionTargetBox-0="[449,1234,189,969]" captionTargetBox-1="[361,1225,1046,1985]" captionTargetBox-2="[290,1297,189,988]" captionTargetBox-3="[291,1296,1096,1939]" captionTargetId-0="figure-9@2.[353,1234,189,969]" captionTargetId-1="figure-22@2.[361,1225,1046,1985]" captionTargetId-2="figure-14@3.[290,1297,189,988]" captionTargetId-3="figure-28@3.[291,1296,1096,1939]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionText-0="Figure 1. Holotype of Temelucha flavia sp. n., female: lateral view." captionText-1="Figure 2. Holotype of Temelucha flavia sp. n., female: dorsal view." captionText-2="Figure 3. Holotype of Temelucha flavia sp. n., female: head, dorsal view." captionText-3="Figure 4. Holotype of Temelucha flavia sp. n., female: head, frontal view." pageId="1" pageNumber="39">Figs 14</figureCitation>
<figureCitation id="13152A51FFA3AF22712CFC3954984B74" box="[556,668,1005,1032]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="2.[432,507,988,1012]" captionStartId-1="2.[432,507,2003,2027]" captionStartId-2="3.[401,476,1007,1031]" captionStartId-3="3.[397,472,1957,1981]" captionTargetBox-0="[449,1234,189,969]" captionTargetBox-1="[361,1225,1046,1985]" captionTargetBox-2="[290,1297,189,988]" captionTargetBox-3="[291,1296,1096,1939]" captionTargetId-0="figure-9@2.[353,1234,189,969]" captionTargetId-1="figure-22@2.[361,1225,1046,1985]" captionTargetId-2="figure-14@3.[290,1297,189,988]" captionTargetId-3="figure-28@3.[291,1296,1096,1939]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionText-0="Figure 1. Holotype of Temelucha flavia sp. n., female: lateral view." captionText-1="Figure 2. Holotype of Temelucha flavia sp. n., female: dorsal view." captionText-2="Figure 3. Holotype of Temelucha flavia sp. n., female: head, dorsal view." captionText-3="Figure 4. Holotype of Temelucha flavia sp. n., female: head, frontal view." figureDoi-0="http://doi.org/10.5281/zenodo.13314852" figureDoi-1="http://doi.org/10.5281/zenodo.13314856" figureDoi-2="http://doi.org/10.5281/zenodo.13314860" figureDoi-3="http://doi.org/10.5281/zenodo.13314862" httpUri-0="https://zenodo.org/record/13314852/files/figure.png" httpUri-1="https://zenodo.org/record/13314856/files/figure.png" httpUri-2="https://zenodo.org/record/13314860/files/figure.png" httpUri-3="https://zenodo.org/record/13314862/files/figure.png" pageId="1" pageNumber="39">Figs 14</figureCitation>
). Body length c.
<quantity id="4CD69B31FFA3AF227076FC3955C54B74" box="[886,961,1005,1032]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="39" unit="mm" value="8.0">8 mm</quantity>
, fore wing length c.
@ -222,7 +225,7 @@ almost straight, not intercepted. Distal abscissa of
<emphasis id="B95AEAC6FFA3AF227191F83654B74880" box="[657,691,2018,2044]" italics="true" pageId="1" pageNumber="39">Cu</emphasis>
1 spectral.
</paragraph>
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<paragraph id="8B9136D4FFA0AF2172B0FC0852804C8F" blockId="2.[432,1156,987,1012]" box="[432,1156,987,1012]" pageId="2" pageNumber="40">
<emphasis id="B95AEAC6FFA0AF2172B0FC0854124C88" bold="true" box="[432,534,988,1012]" pageId="2" pageNumber="40">Figure 1.</emphasis>
Holotype of
@ -233,7 +236,7 @@ Holotype of
, female: lateral view.
</paragraph>
</caption>
<caption id="DF51665CFFA0AF2172B0F80752874897" box="[432,1155,2003,2027]" pageId="2" pageNumber="40" startId="2.[432,507,2003,2027]" targetBox="[361,1225,1046,1985]" targetPageId="2" targetType="figure">
<caption id="DF51665CFFA0AF2172B0F80752874897" ID-DOI="http://doi.org/10.5281/zenodo.13314856" ID-Zenodo-Dep="13314856" box="[432,1155,2003,2027]" httpUri="https://zenodo.org/record/13314856/files/figure.png" pageId="2" pageNumber="40" startId="2.[432,507,2003,2027]" targetBox="[361,1225,1046,1985]" targetPageId="2" targetType="figure">
<paragraph id="8B9136D4FFA0AF2172B0F80752874897" blockId="2.[432,1155,2003,2027]" box="[432,1155,2003,2027]" pageId="2" pageNumber="40">
<emphasis id="B95AEAC6FFA0AF2172B0F80754124897" bold="true" box="[432,534,2003,2027]" pageId="2" pageNumber="40">Figure 2.</emphasis>
Holotype of
@ -244,7 +247,7 @@ Holotype of
, female: dorsal view.
</paragraph>
</caption>
<caption id="DF51665CFFA1AF207291FC3B52A74B7A" box="[401,1187,1006,1031]" pageId="3" pageNumber="41" startId="3.[401,476,1007,1031]" targetBox="[290,1297,189,988]" targetPageId="3" targetType="figure">
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<paragraph id="8B9136D4FFA1AF207291FC3B52A74B7A" blockId="3.[401,1187,1006,1031]" box="[401,1187,1006,1031]" pageId="3" pageNumber="41">
<emphasis id="B95AEAC6FFA1AF207291FC3B57F24B7B" bold="true" box="[401,502,1007,1031]" pageId="3" pageNumber="41">Figure 3.</emphasis>
Holotype of
@ -255,7 +258,7 @@ Holotype of
, female: head, dorsal view.
</paragraph>
</caption>
<caption id="DF51665CFFA1AF20728DF87152A248C0" box="[397,1190,1956,1981]" pageId="3" pageNumber="41" startId="3.[397,472,1957,1981]" targetBox="[291,1296,1096,1939]" targetPageId="3" targetType="figure">
<caption id="DF51665CFFA1AF20728DF87152A248C0" ID-DOI="http://doi.org/10.5281/zenodo.13314862" ID-Zenodo-Dep="13314862" box="[397,1190,1956,1981]" httpUri="https://zenodo.org/record/13314862/files/figure.png" pageId="3" pageNumber="41" startId="3.[397,472,1957,1981]" targetBox="[291,1296,1096,1939]" targetPageId="3" targetType="figure">
<paragraph id="8B9136D4FFA1AF20728DF87152A248C0" blockId="3.[397,1190,1956,1981]" box="[397,1190,1956,1981]" pageId="3" pageNumber="41">
<emphasis id="B95AEAC6FFA1AF20728DF87157F748C1" bold="true" box="[397,499,1957,1981]" pageId="3" pageNumber="41">Figure 4.</emphasis>
Holotype of
@ -266,7 +269,7 @@ Holotype of
, female: head, frontal view.
</paragraph>
</caption>
<caption id="DF51665CFFA6AF277397FBF355614B43" box="[151,869,1063,1087]" pageId="4" pageNumber="42" startId="4.[151,226,1063,1087]" targetBox="[173,1434,189,1044]" targetPageId="4" targetType="figure">
<caption id="DF51665CFFA6AF277397FBF355614B43" ID-DOI="http://doi.org/10.5281/zenodo.13314864" ID-Zenodo-Dep="13314864" box="[151,869,1063,1087]" httpUri="https://zenodo.org/record/13314864/files/figure.png" pageId="4" pageNumber="42" startId="4.[151,226,1063,1087]" targetBox="[173,1434,189,1044]" targetPageId="4" targetType="figure">
<paragraph id="8B9136D4FFA6AF277397FBF355614B43" blockId="4.[151,869,1063,1087]" box="[151,869,1063,1087]" pageId="4" pageNumber="42">
<emphasis id="B95AEAC6FFA6AF277397FBF356F94B43" bold="true" box="[151,253,1063,1087]" pageId="4" pageNumber="42">Figure 5.</emphasis>
Paratype of
@ -294,7 +297,7 @@ Head yellow except following parts: occiput, narrow perimeter of ocelli and two
The
<typeStatus id="54958876FFA7AF26725EFEFD57C74E3F" box="[350,451,297,323]" pageId="5" pageNumber="43" type="paratype">paratype</typeStatus>
female (
<figureCitation id="13152A51FFA7AF267134FEFC547B4E3F" box="[564,639,296,323]" captionStart="Figure 5" captionStartId="4.[151,226,1063,1087]" captionTargetBox="[173,1434,189,1044]" captionTargetId="figure-9@4.[151,1434,189,1044]" captionTargetPageId="4" captionText="Figure 5. Paratype of Temelucha flavia sp. n., female: lateral view." pageId="5" pageNumber="43">Fig. 5</figureCitation>
<figureCitation id="13152A51FFA7AF267134FEFC547B4E3F" box="[564,639,296,323]" captionStart="Figure 5" captionStartId="4.[151,226,1063,1087]" captionTargetBox="[173,1434,189,1044]" captionTargetId="figure-9@4.[151,1434,189,1044]" captionTargetPageId="4" captionText="Figure 5. Paratype of Temelucha flavia sp. n., female: lateral view." figureDoi="http://doi.org/10.5281/zenodo.13314864" httpUri="https://zenodo.org/record/13314864/files/figure.png" pageId="5" pageNumber="43">Fig. 5</figureCitation>
) shows little intraspecific variation in size, shape, sculpture and colouration. Minor differences are as following: antenna with 33 flagellomeres; first flagellomere slightly shorter, 4.5 × as long as wide; ocellus diameter slightly shorter, 2.5 × longer than distance between lateral ocellus and eye; scutellum with lateral carina distinct only basally. Yellow colouration of the body is paler in
<typeStatus id="54958876FFA7AF267397FE7B56FA4EB6" box="[151,254,431,458]" pageId="5" pageNumber="43" type="holotype">holotype</typeStatus>
and somewhat brighter in

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@ -1,46 +1,47 @@
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@ -58,7 +59,7 @@
<subSubSection id="C32AFDA6FFAEFFAFFF62C409DA7E592E" box="[151,280,800,827]" pageId="1" pageNumber="568" type="description">
<paragraph id="8B8FAE2DFFAEFFAFFF62C409DA7E592E" blockId="1.[151,449,767,827]" box="[151,280,800,827]" pageId="1" pageNumber="568">
(
<figureCitation id="130BB2A8FFAEFFAFFF55C409DA69592E" box="[160,271,800,827]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="1.[151,226,2019,2043]" captionStartId-1="2.[151,226,1735,1759]" captionStartId-2="3.[151,226,1922,1946]" captionTargetBox-0="[302,1286,974,2000]" captionTargetBox-1="[151,1434,189,1716]" captionTargetBox-2="[167,1419,366,1834]" captionTargetId-0="figure-210@1.[302,1286,974,2000]" captionTargetId-1="figure-16@2.[151,1434,189,1716]" captionTargetId-2="figure-61@3.[151,1434,358,1885]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figure 1. Anatemnus reni sp. nov., holotype male: habitus, dorsal view." captionText-1="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." captionText-2="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="1" pageNumber="568">Figs. 13</figureCitation>
<figureCitation id="130BB2A8FFAEFFAFFF55C409DA69592E" box="[160,271,800,827]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="1.[151,226,2019,2043]" captionStartId-1="2.[151,226,1735,1759]" captionStartId-2="3.[151,226,1922,1946]" captionTargetBox-0="[302,1286,974,2000]" captionTargetBox-1="[151,1434,189,1716]" captionTargetBox-2="[167,1419,366,1834]" captionTargetId-0="figure-210@1.[302,1286,974,2000]" captionTargetId-1="figure-16@2.[151,1434,189,1716]" captionTargetId-2="figure-61@3.[151,1434,358,1885]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figure 1. Anatemnus reni sp. nov., holotype male: habitus, dorsal view." captionText-1="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." captionText-2="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi-0="http://doi.org/10.5281/zenodo.13315083" figureDoi-1="http://doi.org/10.5281/zenodo.13315087" figureDoi-2="http://doi.org/10.5281/zenodo.13315089" httpUri-0="https://zenodo.org/record/13315083/files/figure.png" httpUri-1="https://zenodo.org/record/13315087/files/figure.png" httpUri-2="https://zenodo.org/record/13315089/files/figure.png" pageId="1" pageNumber="568">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
@ -103,7 +104,7 @@ leg.
</materialsCitation>
</paragraph>
</subSubSection>
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<emphasis id="B944723FFFAEFFAFFF62C0CADB905DEE" bold="true" box="[151,246,2019,2043]" pageId="1" pageNumber="568">Figure 1</emphasis>
.
@ -114,7 +115,7 @@ leg.
, holotype male: habitus, dorsal view.
</paragraph>
</caption>
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<paragraph id="8B8FAE2DFFADFFACFF62C1EEDA515D2F" blockId="2.[151,1437,1734,1850]" pageId="2" pageNumber="569">
<emphasis id="B944723FFFADFFACFF62C1EEDB9A5CCA" bold="true" box="[151,252,1735,1759]" pageId="2" pageNumber="569">Figure 2</emphasis>
.
@ -179,7 +180,7 @@ and
.
</paragraph>
</subSubSection>
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<paragraph id="8B8FAE2DFFACFFADFF62C0ABDE275DE0" blockId="3.[151,1436,1921,2037]" pageId="3" pageNumber="570">
<emphasis id="B944723FFFACFFADFF62C0ABDB9F5D8F" bold="true" box="[151,249,1922,1946]" pageId="3" pageNumber="570">Figure 3</emphasis>
.
@ -194,14 +195,14 @@ and
<paragraph id="8B8FAE2DFFABFFAAFF03C7EDDF835B15" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
<emphasis id="B944723FFFABFFAAFF03C7EDDAE85ACB" bold="true" box="[246,398,196,222]" pageId="4" pageNumber="571">Description.</emphasis>
Body moderately flattened. Pedipalps and anterior half of carapace reddish-brown, legs, tergites and sternites yellowish-brown. Body setae straight, acuminate and short (
<figureCitation id="130BB2A8FFABFFAAFB7AC7CCDFB25B15" box="[1167,1236,229,256]" captionStart="Figure 1" captionStartId="1.[151,226,2019,2043]" captionTargetBox="[302,1286,974,2000]" captionTargetId="figure-210@1.[302,1286,974,2000]" captionTargetPageId="1" captionText="Figure 1. Anatemnus reni sp. nov., holotype male: habitus, dorsal view." pageId="4" pageNumber="571">Fig. 1</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFB7AC7CCDFB25B15" box="[1167,1236,229,256]" captionStart="Figure 1" captionStartId="1.[151,226,2019,2043]" captionTargetBox="[302,1286,974,2000]" captionTargetId="figure-210@1.[302,1286,974,2000]" captionTargetPageId="1" captionText="Figure 1. Anatemnus reni sp. nov., holotype male: habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13315083" httpUri="https://zenodo.org/record/13315083/files/figure.png" pageId="4" pageNumber="571">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C62ED9EC5BDF" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Each chelicera with four setae on hand (
<emphasis id="B944723FFFABFFAAFD29C62ED8675B37" box="[732,769,263,290]" italics="true" pageId="4" pageNumber="571">sbs</emphasis>
absent) (
<figureCitation id="130BB2A8FFABFFAAFC84C62ED8AF5B37" box="[881,969,263,290]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3b</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFC84C62ED8AF5B37" box="[881,969,263,290]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3b</figureCitation>
), surface smooth. Movable finger with one sub-distal seta. Trichobothria
<emphasis id="B944723FFFABFFAAFDDDC601D9245B56" box="[552,578,296,323]" italics="true" pageId="4" pageNumber="571">bs</emphasis>
and
@ -211,20 +212,20 @@ finely dentated,
and
<emphasis id="B944723FFFABFFAAFC52C601D8DD5B56" box="[935,955,296,323]" italics="true" pageId="4" pageNumber="571">ls</emphasis>
long and acute. Movable finger with 2-3 tiny teeth distally, while fixed finger with 4-5 moderately-sized pointed teeth. One lyrifissure present on fixed finger basally (
<figureCitation id="130BB2A8FFABFFAAFE66C645DA8B5B92" box="[403,493,364,391]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3b</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFE66C645DA8B5B92" box="[403,493,364,391]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3b</figureCitation>
); galea with a bifurcate tip and one small sub-median ramus (
<figureCitation id="130BB2A8FFABFFAAFB28C645DE5E5B92" box="[1245,1336,364,391]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3d</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFB28C645DE5E5B92" box="[1245,1336,364,391]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3d</figureCitation>
); rallum composed of four blades, only the distal one dentated anteriorly (
<figureCitation id="130BB2A8FFABFFAAFC6CC6A7D8885BBC" box="[921,1006,398,425]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3c</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFC6CC6A7D8885BBC" box="[921,1006,398,425]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3c</figureCitation>
); serrula exterior with 23-24 blades; lamina exterior present, moderately broad.
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C6F8DA025866" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Pedipalp. Mostly smooth; setae moderately long and acicular. Trochanter elongated, with two well-developed conical tubercles (
<figureCitation id="130BB2A8FFABFFAAFE06C6DAD92A581B" box="[499,588,499,526]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." pageId="4" pageNumber="571">Fig. 2d</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFE06C6DAD92A581B" box="[499,588,499,526]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." figureDoi="http://doi.org/10.5281/zenodo.13315087" httpUri="https://zenodo.org/record/13315087/files/figure.png" pageId="4" pageNumber="571">Fig. 2d</figureCitation>
,
<figureCitation id="130BB2A8FFABFFAAFDAEC6DAD91F581B" box="[603,633,499,526]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">3e</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFDAEC6DAD91F581B" box="[603,633,499,526]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">3e</figureCitation>
); both chelal fingers gaping (
<figureCitation id="130BB2A8FFABFFAAFC29C6DADF53581B" box="[988,1077,499,526]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">
<figureCitation id="130BB2A8FFABFFAAFC29C6DADF53581B" box="[988,1077,499,526]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">
Fig.
<quantity id="4CC803C8FFABFFAAFBE0C6DADF53581B" box="[1045,1077,499,526]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="571" unit="g" value="3.0">3g</quantity>
</figureCitation>
@ -232,7 +233,7 @@ Fig.
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C553D8A8594A" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Fixed chelal finger with eight trichobothria, whilst movable chelal finger with four trichobothria (
<figureCitation id="130BB2A8FFABFFAAFF55C5B5DB9158A2" box="[160,247,668,695]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">
<figureCitation id="130BB2A8FFABFFAAFF55C5B5DB9158A2" box="[160,247,668,695]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">
Fig.
<quantity id="4CC803C8FFABFFAAFF22C5B5DB9158A2" box="[215,247,668,695]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="571" unit="g" value="3.0">3g</quantity>
</figureCitation>
@ -278,29 +279,29 @@ than to tip of movable finger; with no microsetae present on both fingers. Venom
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C44FDAE95E1D" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Carapace (
<figureCitation id="130BB2A8FFABFFAAFE80C44FDAAA5994" box="[373,460,870,897]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." pageId="4" pageNumber="571">Fig. 2a</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFE80C44FDAAA5994" box="[373,460,870,897]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." figureDoi="http://doi.org/10.5281/zenodo.13315087" httpUri="https://zenodo.org/record/13315087/files/figure.png" pageId="4" pageNumber="571">Fig. 2a</figureCitation>
,
<figureCitation id="130BB2A8FFABFFAAFE2EC44FDA9D5994" box="[475,507,870,897]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">3a</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFE2EC44FDA9D5994" box="[475,507,870,897]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">3a</figureCitation>
). Smooth, 1.35 times as long as broad; anterior half of the carapace reddish-brown, remaining part yellowish-brown; without transverse furrow; with a pair of eyespots situated near anterior margin of carapace; with ca. 45 setae in total, including two on anterior and 10 on posterior margin; with two pairs of lyrifissures (indicated by black bars in
<figureCitation id="130BB2A8FFABFFAAFCB2C4E2D8F859F3" box="[839,926,971,998]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3a</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFCB2C4E2D8F859F3" box="[839,926,971,998]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3a</figureCitation>
). Manducatory process with one long seta and three short setae.
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C326DFD25EA4" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Abdomen. Pleural membrane longitudinally striate. Tergites divided except the first three, sternites unconspicuously divided. Tergal chaetotaxy (I-XI): 9: 10: 10: 7-8: 7-7: 8-7: 7-8: 8-9: 8-12: 8-10: 11 + 2T; all setae acicular. Sternal chaetotaxy (IV-XI): 7-8: 8-9: 9-9: 7-9: 8-9: 8-9: 10-10: 14 + 2T; setae uniseriate and acuminate; each tergite and sternite with 2-4 lyrifissures; male genital area as in the
<figureCitation id="130BB2A8FFABFFAAFB73C35DDFBA5E9A" box="[1158,1244,1140,1167]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." pageId="4" pageNumber="571">Fig. 2e</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFB73C35DDFBA5E9A" box="[1158,1244,1140,1167]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." figureDoi="http://doi.org/10.5281/zenodo.13315087" httpUri="https://zenodo.org/record/13315087/files/figure.png" pageId="4" pageNumber="571">Fig. 2e</figureCitation>
; glandular setae absent, but some tactile spots present; anus with four setae, not surrounded by sternite XI.
</paragraph>
<paragraph id="8B8FAE2DFFABFFAAFF00C391DB845F22" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">
Legs (
<figureCitation id="130BB2A8FFABFFAAFECBC39EDAAB5EC7" box="[318,461,1207,1234]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." pageId="4" pageNumber="571">
<figureCitation id="130BB2A8FFABFFAAFECBC39EDAAB5EC7" box="[318,461,1207,1234]" captionStart="Figure 2" captionStartId="2.[151,226,1735,1759]" captionTargetBox="[151,1434,189,1716]" captionTargetId="figure-16@2.[151,1434,189,1716]" captionTargetPageId="2" captionText="Figure 2. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view; b. left pedipalp, dorsal view; c. left pedipalpal chela, retro-lateral view; d. trochanter of left pedipalp, retro-lateral view; e. genital area, ventral view; f. tarsus of left leg IV, lateral view; g. left leg I, lateral view; h. left leg IV, lateral view. Scale bars: 0.50 mm (a and b) and 0.20 mm (c-h)." figureDoi="http://doi.org/10.5281/zenodo.13315087" httpUri="https://zenodo.org/record/13315087/files/figure.png" pageId="4" pageNumber="571">
Figs.
<quantity id="4CC803C8FFABFFAAFE74C39EDAC65EC7" box="[385,416,1207,1234]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="571" unit="g" value="2.0">2g</quantity>
, 2h
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,
<figureCitation id="130BB2A8FFABFFAAFE2CC39ED97E5EC7" box="[473,536,1207,1234]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">3i, 3j</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFE2CC39ED97E5EC7" box="[473,536,1207,1234]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">3i, 3j</figureCitation>
). The junction between femora and patellae I and II oblique; femur + patella of leg IV 2.79 times as long as broad; tactile seta on tarsus IV (
<figureCitation id="130BB2A8FFABFFAAFC8AC3F0D8B35EE1" box="[895,981,1241,1268]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." pageId="4" pageNumber="571">Fig. 3h</figureCitation>
<figureCitation id="130BB2A8FFABFFAAFC8AC3F0D8B35EE1" box="[895,981,1241,1268]" captionStart="Figure 3" captionStartId="3.[151,226,1922,1946]" captionTargetBox="[167,1419,366,1834]" captionTargetId="figure-61@3.[151,1434,358,1885]" captionTargetPageId="3" captionText="Figure 3. Anatemnus reni sp. nov., holotype male: a. carapace, dorsal view (the position of lyrifissures indicated by black bars); b. left chelicera, dorsal view; c. left rallum; d. left galea; e. left pedipalp without chela, dorsal view; f. left pedipalpal chela, retro-lateral view; g. distal part of left pedipalpal chela, retro-lateral view; h. tarsus of left leg IV, lateral view; i. left leg I, lateral view; j. left leg IV, lateral view. Scale bars: 0.25 mm (a, e-j) and 0.10 mm (b-d)." figureDoi="http://doi.org/10.5281/zenodo.13315089" httpUri="https://zenodo.org/record/13315089/files/figure.png" pageId="4" pageNumber="571">Fig. 3h</figureCitation>
) situated proximally (TS = 0.15); sub-terminal tarsal setae curved and apically acute; arolium undivided, clearly shorter than smooth and large claws.
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<paragraph id="8B8FAE2DFFABFFAAFF00C216DE195C17" blockId="4.[151,1437,195,1572]" pageId="4" pageNumber="571">Measurements (length/breadth or height in mm, ratios in parentheses). Body length 4.68. Pedipalps: trochanter 0.64/0.47 (1.36), femur 1.25/0.46 (2.72), patella 1.24/0.50 (2.48), chela (with pedicel) 1.91/0.57 (3.35), chela (without pedicel) 1.78 (3.12), hand length (with pedicel) 1.25 (2.19), hand length (without pedicel) 1.12 (1.96); movable finger length 0.73. Carapace 1.27/0.94 (1.35). Leg I: trochanter 0.23/0.21 (1.10), femur 0.37/0.25 (1.48), patella 0.59/0.24 (2.46), tibia 0.54/0.17 (3.18), tarsus 0.40/0.12 (3.33); leg IV: trochanter 0.39/0.23 (1.70), femur + patella 1.09/0.39 (2.79), tibia 0.76/0.23 (3.30), tarsus 0.48/0.13 (3.69).</paragraph>

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<paragraph id="8B8F369EFF850409CB6CFA963D01FA63" blockId="25.[138,735,1372,1430]" box="[358,515,1405,1430]" pageId="25" pageNumber="26">
(
<figureCitation id="130B2A1BFF850409CB67FA963ED3FA63" box="[365,465,1405,1430]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Figs. 19</figureCitation>
,
<figureCitation id="130B2A1BFF850409CBD6FA963EFEFA63" box="[476,508,1405,1430]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">20</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF850409CACEFA533FCEF9A4" blockId="25.[136,738,1464,1969]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEFA533EBDFA24" box="[196,447,1464,1489]" italics="true" pageId="25" pageNumber="26">Material examined</emphasis>
: Female, cl
<quantity id="4CC89B7BFF850409C879FA533DD9FA24" box="[627,731,1464,1489]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="26" unit="mm" value="7.5">7.5 mm</quantity>
, damaged with incomplete abdomen, ZRC Tonglin Cave, near Nan Cave, Kaiyuan County,
<collectingRegion id="49F4F87CFF850409C889FA133FFBF9C4" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF850409CB75F9F33ECEF9C7" box="[383,460,1560,1586]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, coll. B. Sket,
<date id="FF8E105EFF850409C88DF9F33FCAF9A4" pageId="25" pageNumber="26" value="1995-11-26">26 Nov 1995</date>
.
</paragraph>
<paragraph id="8B8F369EFF850409CACEF9B33DC3F8E4" blockId="25.[136,738,1464,1969]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEF9B33D63F984" box="[196,609,1624,1649]" italics="true" pageId="25" pageNumber="26">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF850409C87CF9B33D51F964" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heteradactyla">
<emphasis id="B944EA8CFF850409C87CF9B33E34F964" italics="true" pageId="25" pageNumber="26">Caridina heteradactyla</emphasis>
<bibRefCitation id="EFA14B6FFF850409CB48F9933D51F964" author="Liang X &amp; Yan S." box="[322,595,1656,1681]" pageId="25" pageNumber="26" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF850409C86DF9933DDEF964" box="[615,732,1656,1681]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF850409CAE1F9733E45F944" box="[235,327,1688,1713]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="25" pageNumber="26" unit="mm" value="5.6">5.6 mm</quantity>
, SFU, mountain stream at Sizhong County,
<collectingRegion id="49F4F87CFF850409CAE6F9533EBFF924" box="[236,445,1720,1745]" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
,
<collectingCountry id="F327760EFF850409CBC0F9533D16F927" box="[458,532,1720,1746]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, 1983.
<typeStatus id="548B883CFF850409C864F9533DDFF924" box="[622,733,1720,1745]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D36FD17FF850409CA82F9333FF9F904" box="[136,251,1752,1777]" count="1" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF850409CB20F9333E8DF904" box="[298,399,1752,1777]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="25" pageNumber="26" unit="mm" value="5.6">5.6 mm</quantity>
, SFU, mountain stream at Sizhong County,
<collectingRegion id="49F4F87CFF850409CB5BF9133D1DF8E4" box="[337,543,1784,1809]" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
,
<collectingCountry id="F327760EFF850409C823F9133D71F8E7" box="[553,627,1784,1810]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, 1983.
</paragraph>
<paragraph id="8B8F369EFF850409CACEF8F33BA2FEA3" blockId="25.[136,738,1464,1969]" lastBlockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEF8F33E4BF8C4" box="[196,329,1816,1841]" italics="true" pageId="25" pageNumber="26">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF850409CBC6F8F33D31F8C4" box="[460,563,1816,1841]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
) short, sharply downturned, reaching slightly beyond end of basal segment of antennular peduncle, dorsally with 24 teeth, of which, 12 are situated at carapace behind posterior margin of orbital margin, occupying 0.3 carapace length, armed ventrally with 7 small teeth. Antennal spine fused with suborbital angle. Pterygostomian angle rounded.
</paragraph>
<paragraph id="8B8F369EFF850409C963FEB63C8EFDE3" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
Eyes well developed (
<figureCitation id="130B2A1BFF850409CE77FEB63BECFE83" box="[1149,1262,349,374]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
). Stylocerite reaching slightly beyond end of basal segment of antennular peduncle. Antennular peduncle (
<figureCitation id="130B2A1BFF850409CF5FFE763C5CFE23" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20A</figureCitation>
) 0.43 times as long as carapace. Scaphocerite (
<figureCitation id="130B2A1BFF850409C93EFE363C9EFE03" box="[820,924,477,502]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20B</figureCitation>
) 2.7 times as long as wide, outer margin straight.
</paragraph>
<paragraph id="8B8F369EFF850409C963FDF63B03FBC3" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF850409CF1BFDF63A7CFDC3" box="[1297,1406,541,566]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19B</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF850409C9B0FDB63B31FD82" box="[954,1075,605,631]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19C</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with dense setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF850409CE28FD563B90FD23" box="[1058,1170,701,726]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19D</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long hooked setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF850409CE18FCF63B7CFCC3" box="[1042,1150,797,822]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19E</figureCitation>
) stout, terminating in triangular projection. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF850409C93FFCB63CA9FC83" box="[821,939,861,886]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19F</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF850409CE88FC963BEFFC62" box="[1154,1261,893,919]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20C</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior quarter of penultimate segment, ultimate segment slightly shorter than pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF850409C963FBD63C99F983" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
First pereiopod (
<figureCitation id="130B2A1BFF850409CE5AFBD63BC8FBA3" box="[1104,1226,1085,1110]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20D</figureCitation>
) short, stout, reaching distal margin of eye, merus 2.1 times as long as broad; carpus strong concave anteriorly, 1.3 times as long as high, chela 2.6 times as long as broad, fingers slightly longer than palm. Second pereiopod (
<figureCitation id="130B2A1BFF850409CE2BFB363B8FFB03" box="[1057,1165,1245,1270]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20E</figureCitation>
) reaching almost to end of second segment of antennular peduncle; carpus 3.9 times as long as high, chela shorter than carpus, 2.3 times as long as broad, fingers as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF850409CF06FAB63C46FA62" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20F, G</figureCitation>
) reaching beyond distal end of scaphocrite, propodus 10 times as long as broad, 4.8 times as long as dactylus (claw included); dactylus ending as 2 claws, with 5 spines on flexor margin. Fifth pereiopod (
<figureCitation id="130B2A1BFF850409C9F3FA163B64F9E3" box="[1017,1126,1533,1558]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20H</figureCitation>
) reaching distal end of antennular peduncle, propodus 10 times as long as wide, dactylus missing. Eggs1.2 ×
<quantity id="4CC89B7BFF850409CF1BF9D63A67F9A3" box="[1297,1381,1597,1622]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="mm" value="0.8">0.8mm</quantity>
in diameter.
</paragraph>
<paragraph id="8B8F369EFF850409C963F9963BCAF963" blockId="25.[813,1414,253,1974]" box="[873,1224,1661,1687]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409C963F9963CBFF963" box="[873,957,1661,1686]" italics="true" pageId="25" pageNumber="26">Habitat</emphasis>
: Stream in karst cave.
</paragraph>
<paragraph id="8B8F369EFF85040CC963F9763E1DFDA3" blockId="25.[813,1414,253,1974]" lastBlockId="28.[174,776,253,663]" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409C963F9763CD6F943" box="[873,980,1693,1718]" italics="true" pageId="25" pageNumber="26">Remarks</emphasis>
: The single female specimen found from the Tonglin Cave in Kaiyan is morphologically closest to
<taxonomicName id="4C304D1DFF850409C9A4F9363A87F903" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[942,1413,1757,1782]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF850409C9A4F9363B7BF903" box="[942,1145,1757,1782]" italics="true" pageId="25" pageNumber="26">C. heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF850409CE89F9363A87F903" author="Liang X &amp; Yan S." box="[1155,1413,1757,1782]" pageId="25" pageNumber="26" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">
Liang and Yan,
<quantity id="4CC89B7BFF850409CF43F9363A87F903" box="[1353,1413,1757,1782]" metricMagnitude="1" metricUnit="m" metricValue="5.0419" pageId="25" pageNumber="26" unit="in" value="1985.0">1985</quantity>
</bibRefCitation>
</taxonomicName>
in the downturned rostrum, the large number of postorbital teeth on carapace, the eggs and the long stylocerite, which reaches the end of basal segment of the antennular peduncle. It can be distinguished from
<taxonomicName id="4C304D1DFF850409CE1DF8963BE2F863" authorityName="Liang and Yan" authorityYear="1985" box="[1047,1248,1917,1942]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF850409CE1DF8963BE2F863" box="[1047,1248,1917,1942]" italics="true" pageId="25" pageNumber="26">C. heterodactyla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF850409CEE1F8963A2DF863" box="[1259,1327,1917,1942]" pageId="25" pageNumber="26" sensu="stricto">s. str.</taxonomicNameLabel>
by the much shorter length of rostrum (
<figureCitation id="130B2A1BFF850409CEB6F8763A28F843" box="[1212,1322,1949,1974]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
), which reaches only slightly beyond the end of the basal segment of antennular peduncle; and the shorter carpus of first pereiopod (1.3 times as long as high,
<figureCitation id="130B2A1BFF80040CCAA4FEB63E1EFE83" box="[174,284,349,374]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="28" pageNumber="29">Fig. 20D</figureCitation>
) (vs. 2.0-2.3 times as long as high; cf.
<bibRefCitation id="EFA14B6FFF80040CCAA4FE963EA4FE63" author="Liang X &amp; Yan S." box="[174,422,381,406]" pageId="28" pageNumber="29" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
: fig. 2(4)). We believe that the morphological differences indicate that the present specimen is probably an undescribed species. But with only a single incomplete female specimen available, we prefer to defer the naming of new species to a later stage when more specimens are available.
</paragraph>
<footnote id="E82B2A90FF850409CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="25" pageNumber="26">
<paragraph id="8B8F369EFF850409CA82F8363E9AF81A" blockId="25.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="25" pageNumber="26">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF86040ACAA4F8823E39F840" ID-DOI="http://doi.org/10.5281/zenodo.13315031" ID-Zenodo-Dep="13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="26" pageNumber="27" startId="26.[174,208,1897,1917]" targetBox="[290,1349,255,1859]" targetPageId="26" targetType="figure">
<paragraph id="8B8F369EFF86040ACAA4F8823E39F840" blockId="26.[174,1451,1897,1973]" pageId="26" pageNumber="27">
<emphasis id="B944EA8CFF86040ACAA4F8823FFBF888" bold="true" box="[174,249,1897,1917]" pageId="26" pageNumber="27">Fig. 19.</emphasis>
<taxonomicName id="4C304D1DFF86040ACB0DF8823DD1F888" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[263,723,1897,1917]" class="Malacostraca" family="Atyidae" genus="Caridina" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF86040ACB0DF8823E54F888" box="[263,342,1897,1917]" italics="true" pageId="26" pageNumber="27">Caridina</emphasis>
aff.
<emphasis id="B944EA8CFF86040ACB8BF8823D03F888" box="[385,513,1897,1917]" italics="true" pageId="26" pageNumber="27">heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF86040AC803F8823DD1F888" author="Liang X &amp; Yan S." box="[521,723,1897,1917]" pageId="26" pageNumber="27" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC).
</paragraph>
</caption>
<footnote id="E82B2A90FF86040ACE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="26" pageNumber="27">
<paragraph id="8B8F369EFF86040ACE96F8363AA9F81A" blockId="26.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="26" pageNumber="27">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF87040BCA82F8823D5EF840" ID-DOI="http://doi.org/10.5281/zenodo.13315033" ID-Zenodo-Dep="13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="27" pageNumber="28" startId="27.[136,171,1897,1917]" targetBox="[217,1364,251,1867]" targetPageId="27" targetType="figure">
<paragraph id="8B8F369EFF87040BCA82F8823D5EF840" blockId="27.[136,1415,1897,1973]" pageId="27" pageNumber="28">
<emphasis id="B944EA8CFF87040BCA82F8823FD7F888" bold="true" box="[136,213,1897,1917]" pageId="27" pageNumber="28">Fig. 20.</emphasis>
<taxonomicName id="4C304D1DFF87040BCAEFF8823DC2F888" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[229,704,1897,1917]" class="Malacostraca" family="Atyidae" genus="Caridina" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF87040BCAEFF8823E34F888" box="[229,310,1897,1917]" italics="true" pageId="27" pageNumber="28">Caridina</emphasis>
aff.
<emphasis id="B944EA8CFF87040BCB6FF8823EEAF888" box="[357,488,1897,1917]" italics="true" pageId="27" pageNumber="28">heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF87040BCBFAF8823DC2F888" author="Liang X &amp; Yan S." box="[496,704,1897,1917]" pageId="27" pageNumber="28" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC).
</paragraph>
</caption>
<footnote id="E82B2A90FF87040BCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="27" pageNumber="28">
<paragraph id="8B8F369EFF87040BCA82F8363E9AF81A" blockId="27.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="27" pageNumber="28">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF80040CCAE0FDB63FFEFD62" blockId="28.[174,776,253,663]" pageId="28" pageNumber="29">
<emphasis id="B944EA8CFF80040CCAE0FDB63E6DFD83" box="[234,367,605,630]" italics="true" pageId="28" pageNumber="29">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF80040CCB75FDB63D48FD83" box="[383,586,605,630]" country="China" name="Yunnan" pageId="28" pageNumber="29">Yunnan Province</collectingRegion>
(Tonglin Cave),
<collectingCountry id="F327760EFF80040CCAA4FD963FFAFD62" box="[174,248,637,663]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="447466CE41468E18C3E0BC213518E73A">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="6B2F3EB1AB40290A9D61DBEA204756B6">Cai, Yixiong</mods:namePart>
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<mods:namePart id="BE9DE23B7B1967AD47571347CF13EE86">Ng, Peter Kee Lin</mods:namePart>
<mods:affiliation id="0CBA8A0AA6D52695747C8E716502016D">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of &amp; National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<treatment id="03998788FF8C0402C9D0FED73E50F8C2" ID-DOI="http://doi.org/10.5281/zenodo.12826768" ID-Zenodo-Dep="12826768" LSID="urn:lsid:zoobank.org:act:D89160FE-F0C5-4F99-80AB-2F07FEDED2CB" httpUri="http://treatment.plazi.org/id/03998788FF8C0402C9D0FED73E50F8C2" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C32A6515FF8C0400C9D0FED73A26FEA3" box="[986,1316,316,342]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8F369EFF8C0400C9D0FED73A26FEA3" blockId="16.[861,1441,316,426]" box="[986,1316,316,342]" pageId="16" pageNumber="17">
<heading id="D0C781F2FF8C0400C9D0FED73A26FEA3" bold="true" box="[986,1316,316,342]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="17" reason="2">
<emphasis id="B944EA8CFF8C0400C9D0FED73A26FEA3" bold="true" box="[986,1316,316,342]" pageId="16" pageNumber="17">
<taxonomicName id="4C304D1DFF8C0400C9D0FED73BB8FEA3" ID-CoL="8VK8N" authorityName="Cai &amp; Ng" authorityYear="2018" box="[986,1210,316,342]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jiangkou" status="sp. nov.">
<emphasis id="B944EA8CFF8C0400C9D0FED73BB8FEA3" bold="true" box="[986,1210,316,342]" italics="true" pageId="16" pageNumber="17">Caridina jiangkou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8C0400CECBFED63A26FEA3" box="[1217,1316,317,342]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6515FF8C0402CE47FEB63E50F8C2" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8F369EFF8C0400CE47FEB63BB2FE83" blockId="16.[861,1441,316,426]" box="[1101,1200,349,374]" pageId="16" pageNumber="17">
(
<figureCitation id="130B2A1BFF8C0400CE5FFEB63BABFE83" box="[1109,1193,349,374]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF8C0400C957FE933BD8FE5F" blockId="16.[861,1441,316,426]" pageId="16" pageNumber="17">
<uri id="FFA13A9CFF8C0400C957FE933BD8FE5F" pageId="16" pageNumber="17">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8B8F369EFF8C0400C985FE3B3BD9FCFC" blockId="16.[850,1452,464,1961]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400C985FE3B3B90FE1C" box="[911,1170,464,489]" italics="true" pageId="16" pageNumber="17">Material examined</emphasis>
:
<materialsCitation id="3B583CC3FF8C0400CEA0FE3B3A1CFDBC" collectingDate="1988-05-15" collectionCode="IZCAS" country="China" location="Baishui Cave" pageId="16" pageNumber="17" specimenCode="DE 5036" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="548B883CFF8C0400CEA0FE3B3A26FE1C" box="[1194,1316,464,489]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF8C0400C958FE1B3CB8FDFC" box="[850,954,496,521]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="17" unit="mm" value="4.8">4.8 mm</quantity>
,
<collectionCode id="ED21AE5BFF8C0400C9D9FE1B3B29FDFF" box="[979,1067,496,522]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="16" pageNumber="17">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF8C0400CE30FE1B3BB1FDFC" box="[1082,1203,496,521]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="16" pageNumber="17">DE 5036</specimenCode>
,
<location id="8EEF6045FF8C0400CEC1FE1B3A84FDFC" LSID="urn:lsid:plazi:treatment:03998788FF8C0402C9D0FED73E50F8C2:8EEF6045FF8C0400CEC1FE1B3A84FDFC" box="[1227,1414,496,522]" country="China" name="Baishui Cave" pageId="16" pageNumber="17" stateProvince="Guizhou">Baishui Cave</location>
in
<taxonomicName id="4C304D1DFF8C0400C958FDFB3B26FDDF" authority="County" authorityName="County" box="[850,1060,528,554]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jiangkou">Jiangkou County</taxonomicName>
,
<collectingRegion id="49F4F87CFF8C0400CE38FDFB3A10FDDC" box="[1074,1298,528,554]" country="China" name="Guizhou" pageId="16" pageNumber="17">Guizhou Province</collectingRegion>
,
<collectingCountry id="F327760EFF8C0400CF2AFDFB3A6EFDDF" box="[1312,1388,528,554]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
, coll. zoological survey team,
<date id="FF8E105EFF8C0400CE70FDDB3A1CFDBC" box="[1146,1310,560,585]" pageId="16" pageNumber="17" value="1988-05-15">
<collectingDate id="EFCAE9B6FF8C0400CE70FDDB3A1CFDBC" box="[1146,1310,560,585]" pageId="16" pageNumber="17" value="1988-05-15">15 May 1988</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="548B883CFF8C0400CF21FDDB3AAAFDBC" box="[1323,1448,560,585]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF8C0400C958FDBB3CBCFD9C" box="[850,958,592,617]" count="1" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
, cl 4.0 mm, IZCAS DE 5037,
<specimenCount id="9D36FD17FF8C0400CF2AFDBB3A8EFD9C" box="[1312,1420,592,617]" count="1" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF8C0400C958FD9B3CB3FD7C" box="[850,945,624,649]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="16" pageNumber="17" unit="mm" value="6.6">6.6 mm</quantity>
, IZCAS DE 5038, data same as
<typeStatus id="548B883CFF8C0400CF35FD9B3AAAFD7C" box="[1343,1448,624,649]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF8C0400C958FD7B3CB8FD5C" box="[850,954,656,681]" count="2" pageId="16" pageNumber="17" type="male">2 males</specimenCount>
,
<quantity id="4CC89B7BFF8C0400C9C2FD7B3B58FD5C" box="[968,1114,656,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="16" pageNumber="17" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.7-4.9 mm</quantity>
,
<specimenCount id="9D36FD17FF8C0400CE63FD7B3BEAFD5C" box="[1129,1256,656,681]" count="4" pageId="16" pageNumber="17" type="female">4 females</specimenCount>
, cl
<quantity id="4CC89B7BFF8C0400CF1FFD7B3AA5FD5C" box="[1301,1447,656,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="6.2" metricValueMin="3.5" pageId="16" pageNumber="17" unit="mm" value="4.85" valueMax="6.2" valueMin="3.5">3.5-6.2 mm</quantity>
, ZRC 2018.0484, same data as
<typeStatus id="548B883CFF8C0400CED2FD5B3A41FD3C" box="[1240,1347,688,713]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF8C0400CF5AFD5B3AAAFD3C" box="[1360,1448,688,713]" count="1" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF8C0400C97AFD3B3CD2FD1C" box="[880,976,720,745]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="17" unit="mm" value="3.9">3.9 mm</quantity>
,
<specimenCount id="9D36FD17FF8C0400C9D7FD3B3B5AFD1C" box="[989,1112,720,745]" count="2" pageId="16" pageNumber="17" type="female">2 females</specimenCount>
,
<quantity id="4CC89B7BFF8C0400CE6FFD3B3BF1FD1C" box="[1125,1267,720,745]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.8" metricValueMin="4.2" pageId="16" pageNumber="17" unit="mm" value="5.0" valueMax="5.8" valueMin="4.2">4.2-5.8 mm</quantity>
, RMNH.CRUS. D.57292, data same as
<typeStatus id="548B883CFF8C0400CE64FD1B3BD5FCFC" box="[1134,1239,752,777]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF8C0400C985FCFB3A73FC3C" blockId="16.[850,1452,464,1961]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400C985FCFB3A29FCDC" box="[911,1323,784,809]" italics="true" pageId="16" pageNumber="17">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF8C0400CF4AFCFB3A10FCBC" authority="Liang and Yan, 1986" authorityName="Liang and Yan" authorityYear="1986" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="guizhouensis">
<emphasis id="B944EA8CFF8C0400CF4AFCFB3CFBFCBC" italics="true" pageId="16" pageNumber="17">Caridina guizhouensis</emphasis>
<bibRefCitation id="EFA14B6FFF8C0400CE0FFCDB3A10FCBC" author="Liang X &amp; Yan S." box="[1029,1298,816,841]" pageId="16" pageNumber="17" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
</taxonomicName>
<emphasis id="B944EA8CFF8C0400CF14FCDB3A25FCBC" box="[1310,1319,816,841]" italics="true" pageId="16" pageNumber="17">-</emphasis>
<typeStatus id="548B883CFF8C0400CF39FCDB3AA5FCBC" box="[1331,1447,816,841]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF8C0400C9C8FCBB3B25FC9C" box="[962,1063,848,873]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="17" unit="mm" value="3.4">3.4 mm</quantity>
, SFU 84-73-1, Maopo Village, Yuping County,
<collectingRegion id="49F4F87CFF8C0400CE1DFC9B3BF4FC7C" box="[1047,1270,880,906]" country="China" name="Guizhou" pageId="16" pageNumber="17">Guizhou Province</collectingRegion>
,
<collectingCountry id="F327760EFF8C0400CF09FC9B3A4DFC7F" box="[1283,1359,880,906]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
, 84-73- 3.
<typeStatus id="548B883CFF8C0400C978FC7B3CE3FC5C" box="[882,993,912,937]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
: 1 ovigerous female, SFU, cl
<quantity id="4CC89B7BFF8C0400CF43FC7B3AA5FC5C" box="[1353,1447,912,937]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="16" pageNumber="17" unit="mm" value="4.6">4.6 mm</quantity>
, eggs 0.95 ×
<quantity id="4CC89B7BFF8C0400C9ECFC5B3B53FC3C" box="[998,1105,944,969]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="mm" value="0.6">0.60 mm</quantity>
, same data as
<typeStatus id="548B883CFF8C0400CF0EFC5B3A6FFC3C" box="[1284,1389,944,969]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF8C0400C985FC3B3A27FB9C" blockId="16.[850,1452,464,1961]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400C985FC3B3B29FC1C" box="[911,1067,976,1001]" italics="true" pageId="16" pageNumber="17">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF8C0400CEC4FC3B3A46FC1C" box="[1230,1348,976,1001]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[136,170,1813,1833]" captionStartId-1="2.[176,211,1841,1861]" captionStartId-2="3.[136,171,1869,1889]" captionTargetBox-0="[136,1414,945,1784]" captionTargetBox-1="[169,1452,916,1812]" captionTargetBox-2="[141,1402,975,1815]" captionTargetId-0="figure-6@1.[125,1428,930,1787]" captionTargetId-1="figure-372@2.[156,1477,907,1826]" captionTargetId-2="figure-6@3.[136,1413,967,1843]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." captionText-1="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." captionText-2="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi-0="http://doi.org/10.5281/zenodo.13314993" figureDoi-1="http://doi.org/10.5281/zenodo.13314995" figureDoi-2="http://doi.org/10.5281/zenodo.13314997" httpUri-0="https://zenodo.org/record/13314993/files/figure.png" httpUri-1="https://zenodo.org/record/13314995/files/figure.png" httpUri-2="https://zenodo.org/record/13314997/files/figure.png" pageId="16" pageNumber="17">Fig. 13A</figureCitation>
) short, straight, reaching to distal end of basal segment of antennular peduncle rostral formula 0-5+4-7/1- 5. Antennal spine fused with suborbital angle. Pterygostomian angle broadly rounded.
</paragraph>
<paragraph id="8B8F369EFF8C0400C985FB9B3CBFFA7C" blockId="16.[850,1452,464,1961]" pageId="16" pageNumber="17">
Sixth abdominal somite 1.8 times as long as fifth abdominal somite, about half carapace length, shorter than telson. Telson (
<figureCitation id="130B2A1BFF8C0400CF05FB5B3AA6FB3C" box="[1295,1444,1200,1225]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="16" pageNumber="17">Fig. 13B, M</figureCitation>
) with prominent posteromedian projection, posterior margin rounded, lateral pair of spines longer than intermediate pairs of spiniform setae, sub-lateral pairs shortest. Inner margin of lateral spines and surface of intermediate pairs of setae covered with spinules.
</paragraph>
<paragraph id="8B8F369EFF8C0400C985FA7B3B4EF9FC" blockId="16.[850,1452,464,1961]" pageId="16" pageNumber="17">
Eyes (
<figureCitation id="130B2A1BFF8C0400C9D4FA7B3B49FA5C" box="[990,1099,1424,1449]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="16" pageNumber="17">Fig. 13A</figureCitation>
) well developed. Antennular peduncle about 0.7 times carapace length. Stylocerite reaching 0.75 basal segment length of antennular peduncle.
</paragraph>
<paragraph id="8B8F369EFF8C0402C985F9FB3E81FEC3" blockId="16.[850,1452,464,1961]" lastBlockId="18.[174,776,253,1847]" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
Mouthparts as figured. Mandible with blunt teeth at extremity of incisor process. Maxillula with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight. Maxilla with slender palp, upper endite subdivided, scaphognathite tapering with numerous long setae posteriorly. Palp of first maxilliped stout, terminating in triangular prjection. Endopod of second maxilliped with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF8C0400CEA2F8DB3A11F8BF" box="[1192,1299,1840,1866]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="16" pageNumber="17">Fig. 13C</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior quarter of penultimate segment, ultimate segment slightly shorter than pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<footnote id="E82B2A90FF8C0400CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="16" pageNumber="17">
<paragraph id="8B8F369EFF8C0400CE96F8363AA9F81A" blockId="16.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="16" pageNumber="17">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF8D0401CA82F8DA3E2FF840" ID-DOI="http://doi.org/10.5281/zenodo.13315019" ID-Zenodo-Dep="13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="17" pageNumber="18" startId="17.[136,171,1841,1861]" targetBox="[214,1327,245,1815]" targetPageId="17" targetType="figure">
<paragraph id="8B8F369EFF8D0401CA82F8DA3E2FF840" blockId="17.[136,1415,1841,1973]" pageId="17" pageNumber="18">
<emphasis id="B944EA8CFF8D0401CA82F8DA3FD7F8B0" bold="true" box="[136,213,1841,1861]" pageId="17" pageNumber="18">Fig. 13.</emphasis>
<taxonomicName id="4C304D1DFF8D0401CAEFF8DA3E8FF8B0" authorityName="Cai &amp; Ng" authorityYear="2018" box="[229,397,1841,1861]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jiangkou" status="sp. nov.">
<emphasis id="B944EA8CFF8D0401CAEFF8DA3E8FF8B0" box="[229,397,1841,1861]" italics="true" pageId="17" pageNumber="18">Caridina jiangkou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8D0401CB9CF8DA3EE5F8B0" box="[406,487,1841,1861]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484).
</paragraph>
</caption>
<footnote id="E82B2A90FF8D0401CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="17" pageNumber="18">
<paragraph id="8B8F369EFF8D0401CA82F8363E9AF81A" blockId="17.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="17" pageNumber="18">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF8E0402CAE0FED63E52FC83" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
First pereiopod (
<figureCitation id="130B2A1BFF8E0402CBDBFED63D49FEA3" box="[465,587,317,342]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13D</figureCitation>
) short, stout, reaching distal end of eyes, merus 2.1 times as long as broad; carpus strongly concave anteriorly, 1.2 times as long as high, chela 1.9 times as long as broad, fingers slightly shorter than palm. Second pereiopod (
<figureCitation id="130B2A1BFF8E0402CB9CFE363EFFFE03" box="[406,509,477,502]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13E</figureCitation>
) reaching beyond end of basal segment of antennular peduncle, carpus 4.0 times as long as high, chela shorter than carpus, 2.7 times as long as broad, fingers slightly shorter or as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF8E0402C8D2FDB63E02FD62" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13F, G</figureCitation>
) reaching beyond distal end of antennular, propodus 10 times as long as broad, 3.7 times as long as dactylus (claw included); dactylus ending as 2 claws, with 8 spines. Fifth pereiopod (
<figureCitation id="130B2A1BFF8E0402C8DCFD363FFAFCE3" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13H, I</figureCitation>
) reaching to end of antennular peduncle, propodus 11 times as long as wide, 4.0 times as long as dactylus; dactylus with 36 spinules on flexor margin.
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0FC963EF0FB83" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF8E0402C89AFC963DFCFC63" box="[656,766,893,918]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13J</figureCitation>
) reaching to half of endopod length, sub-elliptical, 2.4 times as long as wide, appendix interna near distal end of endopod, reaching slightly beyond distal end of endopod. Appendix masculine of male second pleopod (
<figureCitation id="130B2A1BFF8E0402CB8BFBF63EEEFBC3" box="[385,492,1053,1078]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13K</figureCitation>
) reaching 0.4 endopod length; appendix interna slender, reaching to 0.67 appendix masculina length.
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0FB963E1AFB43" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF8E0402C81BFB963D84FB63" box="[529,646,1149,1174]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[136,170,1813,1833]" captionStartId-1="2.[176,211,1841,1861]" captionStartId-2="3.[136,171,1869,1889]" captionTargetBox-0="[136,1414,945,1784]" captionTargetBox-1="[169,1452,916,1812]" captionTargetBox-2="[141,1402,975,1815]" captionTargetId-0="figure-6@1.[125,1428,930,1787]" captionTargetId-1="figure-372@2.[156,1477,907,1826]" captionTargetId-2="figure-6@3.[136,1413,967,1843]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." captionText-1="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." captionText-2="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi-0="http://doi.org/10.5281/zenodo.13314993" figureDoi-1="http://doi.org/10.5281/zenodo.13314995" figureDoi-2="http://doi.org/10.5281/zenodo.13314997" httpUri-0="https://zenodo.org/record/13314993/files/figure.png" httpUri-1="https://zenodo.org/record/13314995/files/figure.png" httpUri-2="https://zenodo.org/record/13314997/files/figure.png" pageId="18" pageNumber="19">Fig. 13L</figureCitation>
) with 13 spinules.
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0FB563D5DFB23" blockId="18.[174,776,253,1847]" box="[234,607,1213,1239]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402CAE0FB563E3CFB23" box="[234,318,1213,1238]" italics="true" pageId="18" pageNumber="19">Habitat</emphasis>
: Stream in a karst cave.
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0FB363E32FAA3" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402CAE0FB363E70FB03" box="[234,370,1245,1270]" italics="true" pageId="18" pageNumber="19">Etymology</emphasis>
:
<taxonomicName id="4C304D1DFF8E0402CB82FB363D73FB03" authorityName="Cai &amp; Ng" authorityYear="2018" box="[392,625,1245,1270]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="jiangkou" status="sp. nov.">
<emphasis id="B944EA8CFF8E0402CB82FB363D73FB03" box="[392,625,1245,1270]" italics="true" pageId="18" pageNumber="19">Caridina jiangkou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8E0402C877FB363DE4FB03" box="[637,742,1245,1270]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
is named after the
<typeStatus id="548B883CFF8E0402CB77FB163EB2FAE3" box="[381,432,1277,1302]" pageId="18" pageNumber="19">type</typeStatus>
locality
<taxonomicName id="4C304D1DFF8E0402C813FB163DE5FAE2" authority="County" authorityName="County" box="[537,743,1277,1303]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="jiangkou">Jiangkou County</taxonomicName>
in
<collectingRegion id="49F4F87CFF8E0402CAA4FAF63E84FAC3" box="[174,390,1309,1335]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou Province</collectingRegion>
. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0FAB63FECF903" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402CAE0FAB63E65FA83" box="[234,359,1373,1398]" italics="true" pageId="18" pageNumber="19">Remarks</emphasis>
:
<taxonomicName id="4C304D1DFF8E0402CB88FAB63D86FA83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[386,644,1373,1398]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="jiangkou" status="sp. nov.">
<emphasis id="B944EA8CFF8E0402CB88FAB63D86FA83" box="[386,644,1373,1398]" italics="true" pageId="18" pageNumber="19">Caridina jiangkou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8E0402C89EFAB63C06FA83" box="[660,772,1373,1398]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C304D1DFF8E0402CB3EFA963C01FA63" authority="Liang and Yan, 1986" authorityName="Liang and Yan" authorityYear="1986" box="[308,771,1405,1430]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guizhouensis">
<emphasis id="B944EA8CFF8E0402CB3EFA963EF4FA63" box="[308,502,1405,1430]" italics="true" pageId="18" pageNumber="19">C. guizhouensis</emphasis>
<bibRefCitation id="EFA14B6FFF8E0402C80BFA963C01FA63" author="Liang X &amp; Yan S." box="[513,771,1405,1430]" pageId="18" pageNumber="19" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
</taxonomicName>
, in the form of the rostrum and telson. However, it can be easily separated from the latter by the form of the endopod of the male first pleopod (2.5 times as long as wide,
<figureCitation id="130B2A1BFF8E0402CB71FA163EDFF9E3" box="[379,477,1533,1558]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13J</figureCitation>
) (vs. 3.0 times; cf.
<bibRefCitation id="EFA14B6FFF8E0402C8CCFA163E7EF9C3" author="Liang X &amp; Zhou J." pageId="18" pageNumber="19" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 2(7)); the longer appendix interna of the male second pereiopod (2/3 length of appendix masculine,
<figureCitation id="130B2A1BFF8E0402CBCDF9B63D2FF983" box="[455,557,1629,1654]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13K</figureCitation>
) (vs. 1/3; cf.
<bibRefCitation id="EFA14B6FFF8E0402C8CCF9B63E6DF963" author="Liang X &amp; Yan S." pageId="18" pageNumber="19" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
: fig. 2(8)); and the presence of spinules on the surface of setae of the telson (
<figureCitation id="130B2A1BFF8E0402C8D3F9763FE7F923" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[214,1327,245,1815]" captionTargetId="figure-96@17.[214,1335,233,1826]" captionTargetPageId="17" captionText="Fig. 13. Caridina jiangkou sp. nov., (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod, (I) dactylus of fifth pereiopod, (J) endopod of male first pleopod, (K) appendix masculina and appendix interna of male second pleopod, (L) uropodal diaeresis, M, posterior spine of telson. Scale bars: A, C-J = 1.0 mm; B = 0.2 mm; K = 0.1 mm. (male, cl 4.7 mm, paratype, ZRC 2018.0484)." figureDoi="http://doi.org/10.5281/zenodo.13315019" httpUri="https://zenodo.org/record/13315019/files/figure.png" pageId="18" pageNumber="19">Fig. 13M</figureCitation>
) (vs. absent; cf.
<bibRefCitation id="EFA14B6FFF8E0402CBC9F9563DD0F923" author="Liang X &amp; Yan S." box="[451,722,1725,1750]" pageId="18" pageNumber="19" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
: fig. 2(2)).
</paragraph>
<paragraph id="8B8F369EFF8E0402CAE0F9163E50F8C2" blockId="18.[174,776,253,1847]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402CAE0F9163E86F8E3" box="[234,388,1789,1814]" italics="true" pageId="18" pageNumber="19">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF8E0402CB91F9163D8EF8E3" box="[411,652,1789,1815]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou Province</collectingRegion>
(Baishui Cave),
<collectingCountry id="F327760EFF8E0402CB0EF8F63E4CF8C2" box="[260,334,1821,1847]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,295 @@
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<mods:title id="7B6FA1CA8CE389F423242425B557454F">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<heading id="D0C781F2FF8E0402C9E7FF163A12FEE3" bold="true" box="[1005,1296,253,279]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B944EA8CFF8E0402C9E7FF163A12FEE3" bold="true" box="[1005,1296,253,279]" pageId="18" pageNumber="19">
<taxonomicName id="4C304D1DFF8E0402C9E7FF163BA4FEE3" ID-CoL="8VK7V" authorityName="Cai &amp; Ng" authorityYear="2018" box="[1005,1190,253,279]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B944EA8CFF8E0402C9E7FF163BA4FEE3" bold="true" box="[1005,1190,253,279]" italics="true" pageId="18" pageNumber="19">Caridina guilin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8E0402CEA4FF163A12FEE3" box="[1198,1296,253,278]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6515FF8E0405CE3AFEF63DE0F842" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="description">
<paragraph id="8B8F369EFF8E0402CE3AFEF63BCFFEC3" blockId="18.[865,1436,253,363]" box="[1072,1229,285,310]" pageId="18" pageNumber="19">
(
<figureCitation id="130B2A1BFF8E0402CE3DFEF63B99FEC3" box="[1079,1179,285,310]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Figs. 14</figureCitation>
,
<figureCitation id="130B2A1BFF8E0402CEADFEF63BC4FEC3" box="[1191,1222,285,310]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF8E0402C96BFED33BDAFE9E" blockId="18.[865,1436,253,363]" pageId="18" pageNumber="19">
<uri id="FFA13A9CFF8E0402C96BFED33BDAFE9E" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="FF960C4BFF8E0402CE55FED33BDAFE9E" pageId="18" pageNumber="19">8126B546-4886-46AC-96F7- FDCCA77549F0</uuid>
</uri>
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FE7B3A30FDDC" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402C985FE7B3B84FE5C" box="[911,1158,400,425]" italics="true" pageId="18" pageNumber="19">Material examined</emphasis>
:
<materialsCitation id="3B583CC3FF8E0402CE96FE7B3B33FDFC" collectingDate="1977-12-01" collectionCode="IZCAS" collectorName="B. Sket" country="China" location="Guangxi Province" pageId="18" pageNumber="19" specimenCode="DE 5039" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="548B883CFF8E0402CE96FE7B3A15FE5C" box="[1180,1303,400,425]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: female, cl
<quantity id="4CC89B7BFF8E0402C958FE5B3CB6FE3C" box="[850,948,432,457]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="19" unit="mm" value="6.5">6.5 mm</quantity>
,
<collectionCode id="ED21AE5BFF8E0402C9C8FE5B3B10FE3F" box="[962,1042,432,458]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="19">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF8E0402CE16FE5B3B8FFE3C" box="[1052,1165,432,457]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="19">DE 5039</specimenCode>
, karst cave near Guilin city,
<collectingRegion id="49F4F87CFF8E0402C984FE3B3B70FE1C" box="[910,1138,464,490]" country="China" name="Guangxi" pageId="18" pageNumber="19">Guangxi Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF8E0402CEF2FE3B3A47FE1F" box="[1272,1349,464,490]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
, coll.
<collectorName id="26C55348FF8E0402CF9BFE3B3C8FFDFF" pageId="18" pageNumber="19">B. Sket</collectorName>
,
<date id="FF8E105EFF8E0402C996FE1B3B33FDFC" box="[924,1073,496,521]" pageId="18" pageNumber="19" value="1977-12-01">
<collectingDate id="EFCAE9B6FF8E0402C996FE1B3B33FDFC" box="[924,1073,496,521]" pageId="18" pageNumber="19" value="1977-12-01">1 Dec 1977</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="548B883CFF8E0402CE4AFE1B3BB1FDFC" box="[1088,1203,496,521]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D36FD17FF8E0402CECAFE1B3A18FDFC" box="[1216,1306,496,521]" count="1" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF8E0402CF4CFE1B3AAAFDFC" box="[1350,1448,496,521]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="18" pageNumber="19" unit="mm" value="4.3">4.3 mm</quantity>
, ZRC 2018.0485, same data as
<typeStatus id="548B883CFF8E0402CECFFDFB3A2CFDDC" box="[1221,1326,528,553]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FDDB3CC2FD1C" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402C985FDDB3A29FDBC" box="[911,1323,560,585]" italics="true" pageId="18" pageNumber="19">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF8E0402CF4AFDDB3A13FD9C" authority="Liang and Zhou, 1993" authorityName="Liang and Zhou" authorityYear="1993" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="B944EA8CFF8E0402CF4AFDDB3CE6FD9C" italics="true" pageId="18" pageNumber="19">Caridina cavernicola</emphasis>
<bibRefCitation id="EFA14B6FFF8E0402C9FBFDBB3A13FD9C" author="Liang X &amp; Zhou J." box="[1009,1297,592,617]" pageId="18" pageNumber="19" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF8E0402CF38FDBB3AA5FD9C" box="[1330,1447,592,617]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: female, cl
<quantity id="4CC89B7BFF8E0402C9D9FD9B3B31FD7C" box="[979,1075,624,649]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="18" pageNumber="19" unit="mm" value="7.3">7.3 mm</quantity>
, SFU 82A-15-1, Lenggu Cave, Duan County, coll.
<date id="FF8E105EFF8E0402CE5CFD7B3A04FD5C" box="[1110,1286,656,681]" pageId="18" pageNumber="19" value="1982-05-22">22 May 1982</date>
.
<typeStatus id="548B883CFF8E0402CF1FFD7B3A8FFD5C" box="[1301,1421,656,681]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D36FD17FF8E0402CF96FD7B3CA5FD3C" count="1" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
, cl 7.0 mm, SFU, 82A-15-2, same data as
<typeStatus id="548B883CFF8E0402C958FD3B3CB9FD1C" box="[850,955,720,745]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FD1B3A27FC7C" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
<emphasis id="B944EA8CFF8E0402C985FD1B3B29FCFC" box="[911,1067,752,777]" italics="true" pageId="18" pageNumber="19">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF8E0402CEC5FD1B3AA2FCFC" box="[1231,1440,752,777]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="1.[136,170,1813,1833]" captionStartId-1="2.[176,211,1841,1861]" captionStartId-2="3.[136,171,1869,1889]" captionStartId-3="5.[136,170,1895,1915]" captionStartId-4="6.[174,208,1895,1915]" captionTargetBox-0="[136,1414,945,1784]" captionTargetBox-1="[169,1452,916,1812]" captionTargetBox-2="[141,1402,975,1815]" captionTargetBox-3="[210,1335,267,1844]" captionTargetBox-4="[265,1379,257,1867]" captionTargetId-0="figure-6@1.[125,1428,930,1787]" captionTargetId-1="figure-372@2.[156,1477,907,1826]" captionTargetId-2="figure-6@3.[136,1413,967,1843]" captionTargetId-3="figure-6@5.[210,1339,249,1866]" captionTargetId-4="figure-52@6.[209,1420,249,1869]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." captionText-1="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." captionText-2="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." captionText-3="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." captionText-4="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi-0="http://doi.org/10.5281/zenodo.13314993" figureDoi-1="http://doi.org/10.5281/zenodo.13314995" figureDoi-2="http://doi.org/10.5281/zenodo.13314997" figureDoi-3="http://doi.org/10.5281/zenodo.13314999" figureDoi-4="http://doi.org/10.5281/zenodo.13315001" httpUri-0="https://zenodo.org/record/13314993/files/figure.png" httpUri-1="https://zenodo.org/record/13314995/files/figure.png" httpUri-2="https://zenodo.org/record/13314997/files/figure.png" httpUri-3="https://zenodo.org/record/13314999/files/figure.png" httpUri-4="https://zenodo.org/record/13315001/files/figure.png" pageId="18" pageNumber="19">Figs. 14A, 15A</figureCitation>
) straight, reaching to or beyond distal end of scaphocerite, rostral formula 6-8 + 20-22/7- 12. Antennal spine fused with suborbital angle. Pterygostomian angle broadly rounded.
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FC7B3BBAFB7C" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
Telson sexually dimorphic. Telson in male (
<figureCitation id="130B2A1BFF8E0402CF74FC7B3C84FC3C" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14B</figureCitation>
) with prominent posteromedian projection, posterior margin rounded, lateral pair of spines longer than intermediate pairs, sub-lateral pairs being shortest; telson in female (
<figureCitation id="130B2A1BFF8E0402CEF2FBFB3A64FBDC" box="[1272,1382,1040,1065]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="18" pageNumber="19">Fig. 15B</figureCitation>
) with prominent posteromedian projection, posterior margin acutely triangular, lateral pair of spines subequal to intermedian pairs.
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FB7B3B4DFABC" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
Eyes well developed (
<figureCitation id="130B2A1BFF8E0402CECEFB7B3A53FB5C" box="[1220,1361,1168,1193]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Figs. 14A</figureCitation>
,
<figureCitation id="130B2A1BFF8E0402CF69FB7B3A9FFB5C" box="[1379,1437,1168,1193]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="18" pageNumber="19">15A</figureCitation>
). Antennular peduncle about 0.6 times carapace length. Stylocerite reaching to 0.8 length or to end of basal segment of antennular peduncle. Scaphocerite (
<figureCitation id="130B2A1BFF8E0402C9F7FAFB3B6AFADC" box="[1021,1128,1296,1321]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14M</figureCitation>
) 3.2 times as long as wide, outer margin straight.
</paragraph>
<paragraph id="8B8F369EFF8E0402C985FABB3BB6F8BC" blockId="18.[850,1452,400,1961]" pageId="18" pageNumber="19">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF8E0402CF3CFABB3AA6FA9F" box="[1334,1444,1360,1386]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14C</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF8E0402C9C1FA7B3B37FA5C" box="[971,1077,1424,1449]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14D</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight. Maxilla with slender palp, upper endite subdivided, scaphognathite tapering with numerous long setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF8E0402C9D2F9DB3B3CF9BC" box="[984,1086,1584,1609]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14E</figureCitation>
) stout, terminating in triangular projection. Endopod of second maxilliped with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF8E0402C9EDF97B3B5AF95C" box="[999,1112,1680,1705]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Fig. 14F</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior quarter of penultimate segment length, ultimate segment distinctly shorter than pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF8E0405C985F8BB3EBBFCE3" blockId="18.[850,1452,400,1961]" lastBlockId="21.[136,738,253,1975]" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
First pereiopod (
<figureCitation id="130B2A1BFF8E0402CE59F8BB3BCCF89F" box="[1107,1230,1872,1898]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="18" pageNumber="19">Figs. 14G</figureCitation>
,
<figureCitation id="130B2A1BFF8E0402CED1F8BB3A0CF89F" box="[1243,1294,1872,1898]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="18" pageNumber="19">15C</figureCitation>
) short, stout, reaching end of basal segment of antennular peduncle, merus 1.5 times in female and 2.0 times in male as long as broad; carpus strongly concave anteriorly, as long as high, chela 2.0 times as long as broad, fingers 0.6 times as long as palm. Second pereiopod (
<figureCitation id="130B2A1BFF890405CB7CFEB63EF3FE83" box="[374,497,349,374]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Figs. 14H</figureCitation>
,
<figureCitation id="130B2A1BFF890405CBF5FEB63D33FE83" box="[511,561,349,374]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">15D</figureCitation>
) reaching end of second segment of antennular peduncle, carpus 3.0 times as long as high, chela slightly longer than carpus, 2.1-2.4 times as long as broad, fingers 0.7 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF890405C865FE363F9BFDE3" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Figs. 14I, J</figureCitation>
,
<figureCitation id="130B2A1BFF890405CAA3FE163FF3FDE3" box="[169,241,509,534]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">15E, I</figureCitation>
) reaching slightly beyond distal end of second segment of antennular peduncle, propodus 10 times as long as broad, 4.5 times as long as dactylus (including claw); dactylus with terminal claw, with 5 spines. Fifth pereiopod (
<figureCitation id="130B2A1BFF890405C845FD963DDBFD63" box="[591,729,637,662]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Fig. 14K, J</figureCitation>
) reaching to end of basal segment of antennular peduncle, propodus 14 times as long as wide, 5.8 times as long as dactylus; dactylus with 56 spinules on flexor margin.
</paragraph>
<footnote id="E82B2A90FF8E0402CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="18" pageNumber="19">
<paragraph id="8B8F369EFF8E0402CE96F8363AA9F81A" blockId="18.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="18" pageNumber="19">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF8F0403CA82F8A63DC2F840" ID-DOI="http://doi.org/10.5281/zenodo.13315021" ID-Zenodo-Dep="13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="19" pageNumber="20" startId="19.[136,170,1869,1889]" targetBox="[224,1332,259,1819]" targetPageId="19" targetType="figure">
<paragraph id="8B8F369EFF8F0403CA82F8A63DC2F840" blockId="19.[136,1415,1869,1973]" pageId="19" pageNumber="20">
<emphasis id="B944EA8CFF8F0403CA82F8A63FD6F894" bold="true" box="[136,212,1869,1889]" pageId="19" pageNumber="20">Fig. 14.</emphasis>
<taxonomicName id="4C304D1DFF8F0403CAEEF8A63E6FF894" authorityName="Cai &amp; Ng" authorityYear="2018" box="[228,365,1869,1889]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B944EA8CFF8F0403CAEEF8A63E6FF894" box="[228,365,1869,1889]" italics="true" pageId="19" pageNumber="20">Caridina guilin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8F0403CB7FF8A63EC0F894" box="[373,450,1869,1889]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
(A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039).
</paragraph>
</caption>
<footnote id="E82B2A90FF8F0403CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="19" pageNumber="20">
<paragraph id="8B8F369EFF8F0403CA82F8363E9AF81A" blockId="19.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="19" pageNumber="20">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF880404CAA4F8A63D07F840" ID-DOI="http://doi.org/10.5281/zenodo.13315023" ID-Zenodo-Dep="13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="20" pageNumber="21" startId="20.[174,209,1869,1889]" targetBox="[175,1454,306,1819]" targetPageId="20" targetType="figure">
<paragraph id="8B8F369EFF880404CAA4F8A63D07F840" blockId="20.[174,1451,1869,1973]" pageId="20" pageNumber="21">
<emphasis id="B944EA8CFF880404CAA4F8A63FFFF894" bold="true" box="[174,253,1869,1889]" pageId="20" pageNumber="21">Fig. 15.</emphasis>
<taxonomicName id="4C304D1DFF880404CB04F8A63E9CF894" authorityName="Cai &amp; Ng" authorityYear="2018" box="[270,414,1869,1889]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B944EA8CFF880404CB04F8A63E9CF894" box="[270,414,1869,1889]" italics="true" pageId="20" pageNumber="21">Caridina guilin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF880404CBADF8A63EF4F894" box="[423,502,1869,1889]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
(A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485).
</paragraph>
</caption>
<footnote id="E82B2A90FF880404CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="20" pageNumber="21">
<paragraph id="8B8F369EFF880404CE96F8363AA9F81A" blockId="20.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="20" pageNumber="21">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF890405CACFFCF63D91FC03" blockId="21.[136,738,253,1975]" pageId="21" pageNumber="22">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF890405C824FCF63D90FCC3" box="[558,658,797,822]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">Fig. 15E</figureCitation>
) short, reaching to 0.2 endopod length, sub-triangular, 1.7 times as long as wide, appendix interna at distal end of endopod, reaching remarkably beyond distal end of endopod. Appendix masculina of male second pleopod (
<figureCitation id="130B2A1BFF890405CB52FC563EC1FC22" box="[344,451,957,983]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">Fig. 15G</figureCitation>
) reaching half endopod length; slightly longer than appendix interna.
</paragraph>
<paragraph id="8B8F369EFF890405CACFFC163FF0FBC3" blockId="21.[136,738,253,1975]" pageId="21" pageNumber="22">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF890405CBCFFC163D37FBE3" box="[453,565,1021,1046]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">Fig. 15H</figureCitation>
) with 9 or 10 spinules.
</paragraph>
<paragraph id="8B8F369EFF890405CACFFBD63D3BFBA3" blockId="21.[136,738,253,1975]" box="[197,569,1085,1111]" pageId="21" pageNumber="22">
<emphasis id="B944EA8CFF890405CACFFBD63E1BFBA3" box="[197,281,1085,1110]" italics="true" pageId="21" pageNumber="22">Habitat</emphasis>
: Stream in a karst cave.
</paragraph>
<paragraph id="8B8F369EFF890405CACFFBB63FCDFB23" blockId="21.[136,738,253,1975]" pageId="21" pageNumber="22">
<emphasis id="B944EA8CFF890405CACFFBB63E40FB83" box="[197,322,1117,1142]" italics="true" pageId="21" pageNumber="22">Etymology</emphasis>
:
<taxonomicName id="4C304D1DFF890405CB59FBB63D02FB83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[339,512,1117,1142]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B944EA8CFF890405CB59FBB63D02FB83" box="[339,512,1117,1142]" italics="true" pageId="21" pageNumber="22">Caridina guilin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF890405C800FBB63D6BFB83" box="[522,617,1117,1142]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
is named after the
<typeStatus id="548B883CFF890405CB0DFB963E3CFB63" box="[263,318,1149,1174]" pageId="21" pageNumber="22">type</typeStatus>
locality, Guilin city in
<collectingRegion id="49F4F87CFF890405C87EFB963FFAFB43" country="China" name="Guangxi" pageId="21" pageNumber="22">Guangxi Province</collectingRegion>
. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF890405CACFFB363D7EF903" blockId="21.[136,738,253,1975]" pageId="21" pageNumber="22">
<emphasis id="B944EA8CFF890405CACFFB363E34FB03" box="[197,310,1245,1270]" italics="true" pageId="21" pageNumber="22">Remarks</emphasis>
:
<taxonomicName id="4C304D1DFF890405CB40FB363D01FB03" authorityName="Cai &amp; Ng" authorityYear="2018" box="[330,515,1245,1270]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B944EA8CFF890405CB40FB363D01FB03" box="[330,515,1245,1270]" italics="true" pageId="21" pageNumber="22">Caridina guilin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF890405C805FB363D76FB03" box="[527,628,1245,1270]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
bears a close resemblance to
<taxonomicName id="4C304D1DFF890405CBA9FB163E1DFAC3" authority="Liang and Zhou, 1993" authorityName="Liang and Zhou" authorityYear="1993" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="B944EA8CFF890405CBA9FB163D5AFAE3" box="[419,600,1277,1302]" italics="true" pageId="21" pageNumber="22">C. cavernicola</emphasis>
<bibRefCitation id="EFA14B6FFF890405C869FB163E1DFAC3" author="Liang X &amp; Zhou J." pageId="21" pageNumber="22" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
</taxonomicName>
(described from Duan County of
<collectingRegion id="49F4F87CFF890405CA82FAD63E6DFAA3" box="[136,367,1341,1367]" country="China" name="Guangxi" pageId="21" pageNumber="22">Guangxi Province</collectingRegion>
), in the long rostrum and the form of telson in female, which terminates in an acute triangle. The latter character is unusual and has so far not been reported in any other congener. However, the new species can easily be distinguished from
<taxonomicName id="4C304D1DFF890405CB7FFA363D24FA03" authorityName="Liang and Zhou" authorityYear="1993" box="[373,550,1501,1526]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="B944EA8CFF890405CB7FFA363D24FA03" box="[373,550,1501,1526]" italics="true" pageId="21" pageNumber="22">C. cavernicola</emphasis>
</taxonomicName>
by the smaller number of ventral teeth on the rostrum (7-12,
<figureCitation id="130B2A1BFF890405C8ADFA163FBFF9C3" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Figs. 14A</figureCitation>
,
<figureCitation id="130B2A1BFF890405CAC1F9F63FFFF9C3" box="[203,253,1565,1590]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">15A</figureCitation>
) (vs. 20-24; cf.
<bibRefCitation id="EFA14B6FFF890405CBCCF9F63DDCF9C3" author="Liang X &amp; Zhou J." box="[454,734,1565,1590]" pageId="21" pageNumber="22" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 2(1)); the shorter carpus of the first pereiopod which is as long as high (
<figureCitation id="130B2A1BFF890405CBBDF9B63D33F982" box="[439,561,1629,1655]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Figs. 14G</figureCitation>
,
<figureCitation id="130B2A1BFF890405C837F9B63D72F982" box="[573,624,1629,1655]" captionStart="Fig" captionStartId="20.[174,209,1869,1889]" captionTargetBox="[175,1454,306,1819]" captionTargetId="figure-76@20.[151,1471,283,1839]" captionTargetPageId="20" captionText="Fig. 15. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) first pereiopod, (D) second pereiopod, (E) third pereiopod, (F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod, (H) uropodal diaeresis, (I) dactylus of third pereiopod. Scale bars: A, C-E = 1.0 mm; B, F-I = 0.2 mm. (male, cl 4.3 mm, paratype, ZRC 2018.0485)." figureDoi="http://doi.org/10.5281/zenodo.13315023" httpUri="https://zenodo.org/record/13315023/files/figure.png" pageId="21" pageNumber="22">15C</figureCitation>
) (vs. 1.4- 1.7 and as long as high; cf.
<bibRefCitation id="EFA14B6FFF890405CBDEF9963DDCF963" author="Liang X &amp; Zhou J." box="[468,734,1661,1686]" pageId="21" pageNumber="22" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 2(4)); and the smaller number of spines (56) on the dactylus of the fifth pereiopod (
<figureCitation id="130B2A1BFF890405C836F9563DA6F923" box="[572,676,1725,1750]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Fig. 14J</figureCitation>
) (vs. 66-69; cf.
<bibRefCitation id="EFA14B6FFF890405CAF5F9363D05F903" author="Liang X &amp; Zhou J." box="[255,519,1757,1782]" pageId="21" pageNumber="22" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 2(7)).
</paragraph>
<paragraph id="8B8F369EFF890405CACFF9163D64F863" blockId="21.[136,738,253,1975]" pageId="21" pageNumber="22">
The adaptive significance of the unusual telson shape (
<figureCitation id="130B2A1BFF890405CB4CF8F63EB8F8C3" box="[326,442,1821,1846]" captionStart="Fig" captionStartId="19.[136,170,1869,1889]" captionTargetBox="[224,1332,259,1819]" captionTargetId="figure-89@19.[194,1356,249,1843]" captionTargetPageId="19" captionText="Fig. 14. Caridina guilin sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) distal portion of telson, (C) mandible, (D) maxilla, (E) first maxilliped, (F) third maxilliped, (G) first pereiopod, (H) second pereiopod, (I) third pereiopod, (J) dactylus of third pereiopod, (K) fifth pereiopod, (L) dactylus of fifth pereiopod. Scale bars: A = 2.0 mm; B, J = 0.2 mm; C-F, M = 1.0 mm; G-I, L = 0.1 mm; K = 0.5 mm. (female, cl 6.5 mm, holotype, IZCAS DE 5039)." figureDoi="http://doi.org/10.5281/zenodo.13315021" httpUri="https://zenodo.org/record/13315021/files/figure.png" pageId="21" pageNumber="22">Fig. 14B</figureCitation>
) is unknown, but may suggest that
<taxonomicName id="4C304D1DFF890405CB32F8D63EA7F8A3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[312,421,1853,1878]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="guilin">
<emphasis id="B944EA8CFF890405CB32F8D63EA7F8A3" box="[312,421,1853,1878]" italics="true" pageId="21" pageNumber="22">C. guilin</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D1DFF890405CBE4F8D63DA9F8A3" authorityName="Liang and Zhou" authorityYear="1993" box="[494,683,1853,1878]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="B944EA8CFF890405CBE4F8D63DA9F8A3" box="[494,683,1853,1878]" italics="true" pageId="21" pageNumber="22">C. cavernicola</emphasis>
</taxonomicName>
are related, although they occur in two separate cave systems some distance from each other.
</paragraph>
<paragraph id="8B8F369EFF890405CACFF8763DE0F842" blockId="21.[136,738,253,1975]" box="[197,738,1949,1975]" pageId="21" pageNumber="22">
<emphasis id="B944EA8CFF890405CACFF8763E4BF843" box="[197,329,1949,1974]" italics="true" pageId="21" pageNumber="22">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF890405CB52F8763D2BF843" box="[344,553,1949,1975]" country="China" name="Guangxi" pageId="21" pageNumber="22">Guangxi Province</collectingRegion>
(Guilin),
<collectingCountry id="F327760EFF890405C89EF8763DDCF842" box="[660,734,1949,1975]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,328 @@
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<mods:title id="EF360635953AD2024E729599377619C9">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<subSubSection id="C32A6515FF90041CC9E4FE573A12FE23" box="[1006,1296,444,470]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B8F369EFF90041CC9E4FE573A12FE23" blockId="12.[859,1443,444,555]" box="[1006,1296,444,470]" pageId="12" pageNumber="13">
<heading id="D0C781F2FF90041CC9E4FE573A12FE23" bold="true" box="[1006,1296,444,470]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B944EA8CFF90041CC9E4FE573A12FE23" bold="true" box="[1006,1296,444,470]" pageId="12" pageNumber="13">
<taxonomicName id="4C304D1DFF90041CC9E4FE573BA4FE23" ID-CoL="8VK4N" authorityName="Cai &amp; Ng" authorityYear="2018" box="[1006,1190,444,470]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF90041CC9E4FE573BA4FE23" bold="true" box="[1006,1190,444,470]" italics="true" pageId="12" pageNumber="13">Caridina beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CCEA7FE563A12FE23" box="[1197,1296,445,470]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B8F369EFF90041CCE39FE363BC8FE03" blockId="12.[859,1443,444,555]" box="[1075,1226,477,502]" pageId="12" pageNumber="13">
(
<figureCitation id="130B2A1BFF90041CCE30FE363BC1FE03" box="[1082,1219,477,502]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="13.[136,170,1898,1918]" captionStartId-1="14.[174,208,1925,1945]" captionStartId-2="15.[136,170,1898,1918]" captionTargetBox-0="[244,1338,260,1856]" captionTargetBox-1="[205,1322,249,1864]" captionTargetBox-2="[142,1414,261,1859]" captionTargetId-0="figure-58@13.[206,1344,249,1873]" captionTargetId-1="figure-6@14.[202,1347,249,1871]" captionTargetId-2="figure-6@15.[114,1423,249,1899]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." captionText-1="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" captionText-2="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi-0="http://doi.org/10.5281/zenodo.13315013" figureDoi-1="http://doi.org/10.5281/zenodo.13315015" figureDoi-2="http://doi.org/10.5281/zenodo.13315017" httpUri-0="https://zenodo.org/record/13315013/files/figure.png" httpUri-1="https://zenodo.org/record/13315015/files/figure.png" httpUri-2="https://zenodo.org/record/13315017/files/figure.png" pageId="12" pageNumber="13">Figs. 10-12</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF90041CC951FE133BD6FDDE" blockId="12.[859,1443,444,555]" pageId="12" pageNumber="13">
<uri id="FFA13A9CFF90041CC951FE133BD6FDDE" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="FF960C4BFF90041CCE5CFE133BD6FDDE" pageId="12" pageNumber="13">3CCBD229-9AEF-4016-A652- F932F38FD51B</uuid>
</uri>
</paragraph>
<paragraph id="8B8F369EFF90041CC985FDBB3BD9FC5C" blockId="12.[850,1452,592,1961]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CC985FDBB3B90FD9C" box="[911,1170,592,617]" italics="true" pageId="12" pageNumber="13">Material examined</emphasis>
:
<materialsCitation id="3B583CC3FF90041CCEA0FDBB3B94FD3C" collectingDate="1993-10-30" collectionCode="IZCAS" collectorName="Y. Cai" country="China" location="Goulou Cave" pageId="12" pageNumber="13" specimenCode="DE 5032" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="548B883CFF90041CCEA0FDBB3A26FD9C" box="[1194,1316,592,617]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF90041CC958FD9B3CB6FD7C" box="[850,948,624,649]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="13" unit="mm" value="5.8">5.8 mm</quantity>
,
<collectionCode id="ED21AE5BFF90041CC9C8FD9B3B10FD7F" box="[962,1042,624,650]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="12" pageNumber="13">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF90041CCE17FD9B3B8CFD7C" box="[1053,1166,624,649]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="12" pageNumber="13">DE 5032</specimenCode>
,
<location id="8EEF6045FF90041CCE96FD9B3A3FFD7F" LSID="urn:lsid:plazi:treatment:03998788FF900400C9E4FE573CF4FEE2:8EEF6045FF90041CCE96FD9B3A3FFD7F" box="[1180,1341,624,650]" country="China" name="Goulou Cave" pageId="12" pageNumber="13" stateProvince="Guangxi">Goulou Cave</location>
in
<taxonomicName id="4C304D1DFF90041CCF62FD9B3CAEFD5F" authority="County" authorityName="County" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County</taxonomicName>
,
<collectingRegion id="49F4F87CFF90041CC9B0FD7B3B99FD5C" box="[954,1179,656,682]" country="China" name="Guangxi" pageId="12" pageNumber="13">Guangxi Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF90041CCF2AFD7B3A6EFD5F" box="[1312,1388,656,682]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
, coll.
<collectorName id="26C55348FF90041CC958FD5B3C9EFD3F" box="[850,924,688,714]" pageId="12" pageNumber="13">Y. Cai</collectorName>
et al.,
<date id="FF8E105EFF90041CC9FDFD5B3B94FD3C" box="[1015,1174,688,714]" pageId="12" pageNumber="13" value="1993-10-30">
<collectingDate id="EFCAE9B6FF90041CC9FDFD5B3B94FD3C" box="[1015,1174,688,714]" pageId="12" pageNumber="13" value="1993-10-30">30 Oct 1993</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="548B883CFF90041CCEA9FD5B3A21FD3C" box="[1187,1315,688,713]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF90041CCF3BFD5B3A8BFD3C" box="[1329,1417,688,713]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CC958FD3B3CB4FD1C" box="[850,950,720,745]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="12" pageNumber="13" unit="mm" value="5.7">5.7 mm</quantity>
, IZCAS DE 5033,
<specimenCount id="9D36FD17FF90041CCEAEFD3B3A15FD1C" box="[1188,1303,720,745]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
, cl 5.0 mm, IZCAS DE 5034,
<specimenCount id="9D36FD17FF90041CCE33FD1B3BB3FCFC" box="[1081,1201,752,777]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CCEEBFD1B3A4AFCFC" box="[1249,1352,752,777]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="12" pageNumber="13" unit="mm" value="5.1">5.1 mm</quantity>
, IZCAS DE 5035, data same as
<typeStatus id="548B883CFF90041CCE92FCFB3A09FCDC" box="[1176,1291,784,809]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF90041CCF11FCFB3A85FCDC" box="[1307,1415,784,809]" count="3" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CC958FCDB3CF0FCBC" box="[850,1010,816,841]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="12" pageNumber="13" unit="mm" value="6.2" valueMax="6.7" valueMin="5.7">5.7-6.7 mm</quantity>
,
<specimenCount id="9D36FD17FF90041CCE09FCDB3B8FFCBC" box="[1027,1165,816,841]" count="5" pageId="12" pageNumber="13" type="female">5 females</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CCECBFCDB3A62FCBC" box="[1217,1376,816,841]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.8" metricValueMin="5.5" pageId="12" pageNumber="13" unit="mm" value="6.15" valueMax="6.8" valueMin="5.5">5.5-6.8 mm</quantity>
, ZRC 2018.0483, same data as
<typeStatus id="548B883CFF90041CCEA8FCBB3A13FC9C" box="[1186,1297,848,873]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF90041CCF2AFCBB3A8BFC9C" box="[1312,1417,848,873]" count="3" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CC958FC9B3CE1FC7C" box="[850,995,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="12" pageNumber="13" unit="mm" value="4.85" valueMax="5.5" valueMin="4.2">4.2-5.5 mm</quantity>
,
<specimenCount id="9D36FD17FF90041CC9F8FC9B3B60FC7C" box="[1010,1122,880,905]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CCE84FC9B3BEDFC7C" box="[1166,1263,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="13" unit="mm" value="5.8">5.8 mm</quantity>
, RMNH.CRUS. D.57291, data same as
<typeStatus id="548B883CFF90041CCE64FC7B3BD5FC5C" box="[1134,1239,912,937]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF90041CC985FC5B3B4BFBBC" blockId="12.[850,1452,592,1961]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CC985FC5B3A29FC3C" box="[911,1323,944,969]" italics="true" pageId="12" pageNumber="13">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF90041CCF4AFC5B3BB4FC1C" authority="Liang and Yan, 1986" authorityName="Liang and Yan" authorityYear="1986" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B944EA8CFF90041CCF4AFC5B3CAEFC1C" italics="true" pageId="12" pageNumber="13">Caridina cornuta</emphasis>
<materialsCitation id="3B583CC3FF90041CC9BDFC3B3BB4FC1C" box="[951,1206,976,1001]" collectingDate="1984-05-29" collectionCode="SFU" country="China" county="Zhijing County" location="Zhijing County" pageId="12" pageNumber="13" specimenCode="SFU 84-53, 1, 2" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<bibRefCitation id="EFA14B6FFF90041CC9BDFC3B3BB4FC1C" author="Liang X &amp; Yan S." box="[951,1206,976,1001]" pageId="12" pageNumber="13" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
</materialsCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF90041CCED9FC3B3A40FC1C" box="[1235,1346,976,1001]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D36FD17FF90041CCF5AFC3B3AAAFC1C" box="[1360,1448,976,1001]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CC967FC1B3CC8FBFC" box="[877,970,1008,1033]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="13" unit="mm" value="4.2">4.2 mm</quantity>
,
<specimenCode id="DB969EE5FF90041CC9DCFC1B3B60FBFC" box="[982,1122,1008,1034]" collectionCode="SFU" country="China" httpUri="http://grbio.org/cool/kb3t-9jg8" name="Shanghai Fisheries University" pageId="12" pageNumber="13">SFU 84-53</specimenCode>
-
<specimenCode id="DB969EE5FF90041CCE60FC1B3B70FBFC" box="[1130,1138,1008,1033]" collectionCode="SFU" country="China" httpUri="http://grbio.org/cool/kb3t-9jg8" name="Shanghai Fisheries University" pageId="12" pageNumber="13">1</specimenCode>
,
<specimenCount id="9D36FD17FF90041CCE74FC1B3BF5FBFC" box="[1150,1271,1008,1033]" count="2" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
, cl
<quantity id="4CC89B7BFF90041CCF17FC1B3AA5FBFC" box="[1309,1447,1008,1033]" metricMagnitude="-3" metricUnit="m" metricValue="5.199999999999999" metricValueMax="6.6" metricValueMin="3.8" pageId="12" pageNumber="13" unit="mm" value="5.199999999999999" valueMax="6.6" valueMin="3.8">3.8-6.6 mm</quantity>
,
<specimenCode id="DB969EE5FF90041CC958FBFB3CDDFBDC" box="[850,991,1040,1066]" collectionCode="SFU" country="China" httpUri="http://grbio.org/cool/kb3t-9jg8" name="Shanghai Fisheries University" pageId="12" pageNumber="13">SFU 84-53</specimenCode>
-
<specimenCode id="DB969EE5FF90041CC9EDFBFB3CF5FBDC" box="[999,1015,1040,1065]" collectionCode="SFU" country="China" httpUri="http://grbio.org/cool/kb3t-9jg8" name="Shanghai Fisheries University" pageId="12" pageNumber="13">2</specimenCode>
,
<collectingCounty id="62EE4E12FF90041CCE0FFBFB3BB8FBDF" box="[1029,1210,1040,1066]" pageId="12" pageNumber="13">Zhijing County</collectingCounty>
,
<collectingRegion id="49F4F87CFF90041CCEC2FBFB3AAAFBDC" box="[1224,1448,1040,1066]" country="China" name="Guizhou" pageId="12" pageNumber="13">Guizhou Province</collectingRegion>
,
<collectingCountry id="F327760EFF90041CC958FBDB3C9EFBBF" box="[850,924,1072,1098]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
,
<date id="FF8E105EFF90041CC9ADFBDB3B44FBBC" box="[935,1094,1072,1097]" pageId="12" pageNumber="13" value="1984-05-29">
<collectingDate id="EFCAE9B6FF90041CC9ADFBDB3B44FBBC" box="[935,1094,1072,1097]" pageId="12" pageNumber="13" value="1984-05-29">29 May 1984</collectingDate>
</date>
.
</paragraph>
<paragraph id="8B8F369EFF90041CC985FBBB3A27FADC" blockId="12.[850,1452,592,1961]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CC985FBBB3B20FB9C" box="[911,1058,1104,1129]" italics="true" pageId="12" pageNumber="13">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF90041CCEB3FBBB3A2FFB9C" box="[1209,1325,1104,1129]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10A</figureCitation>
) straight, slightly downwards, reaching near distal end of antennular peduncle, armed dorsally with 16 teeth, of which 4 are on carapace behind posterior margin of orbital margin, armed ventrally with 1 tooth. Antennal spine fused with suborbital angle. Pterygostomian angle broadly rounded.
</paragraph>
<paragraph id="8B8F369EFF90041CC985FADB3AA8F9FC" blockId="12.[850,1452,592,1961]" pageId="12" pageNumber="13">
Sixth abdominal somite twice as long as fifth abdominal somite, slightly more than half of carapace length. Telson (
<figureCitation id="130B2A1BFF90041CCEBBFA9B3A41FA7C" box="[1201,1347,1392,1417]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="12" pageNumber="13">Fig. 12A, B</figureCitation>
) slightly longer than sixth abdominal somite, with prominent posteromedian projection, posterior margin rounded, with lateral pair of spines longer than intermediate pairs, sub-lateral pairs being shortest.
</paragraph>
<paragraph id="8B8F369EFF90041CC985F9FB3BF0F95C" blockId="12.[850,1452,592,1961]" pageId="12" pageNumber="13">
Eyes (
<figureCitation id="130B2A1BFF90041CC9D4F9FB3B49F9DC" box="[990,1099,1552,1577]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10A</figureCitation>
) well developed. Antennular peduncle about half of carapace length. Stylocerite reaching to 0.8 basal segment length of antennular peduncle. Scaphocerite (
<figureCitation id="130B2A1BFF90041CCE87F99B3BFFF97F" box="[1165,1277,1648,1674]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="12" pageNumber="13">Fig. 12C</figureCitation>
) 3.0 times as long as wide, outer margin straight.
</paragraph>
<paragraph id="8B8F369EFF900400C985F95B3E38FDE3" blockId="12.[850,1452,592,1961]" lastBlockId="16.[174,776,253,1975]" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF90041CCF3DF95B3AA1F93C" box="[1335,1443,1712,1737]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10B</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF90041CC9D5F91B3B5BF8FF" box="[991,1113,1776,1802]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10C</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with dense setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF90041CCE42F8BB3BBAF89C" box="[1096,1208,1872,1897]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10D</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long hooked setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF8C0400CB99FF163EFDFEE3" box="[403,511,253,278]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="16" pageNumber="17">Fig. 10E</figureCitation>
) stout, terminating in triangular. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF8C0400C8DFFEF63FDCFEA3" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="16" pageNumber="17">Fig. 10F</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF8C0400CB9FFEB63D0FFE82" box="[405,525,349,375]" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="16" pageNumber="17">Fig. 10G</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior third of penultimate segment, ultimate segment shorter than pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<footnote id="E82B2A90FF90041CCE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="12" pageNumber="13">
<paragraph id="8B8F369EFF90041CCE96F8363AA9F81A" blockId="12.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="12" pageNumber="13">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF91041DCA82F8813D34F843" ID-DOI="http://doi.org/10.5281/zenodo.13315013" ID-Zenodo-Dep="13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="13" pageNumber="14" startId="13.[136,170,1898,1918]" targetBox="[244,1338,260,1856]" targetPageId="13" targetType="figure">
<paragraph id="8B8F369EFF91041DCA82F8813D34F843" blockId="13.[136,1415,1898,1974]" pageId="13" pageNumber="14">
<emphasis id="B944EA8CFF91041DCA82F8813FD1F88B" bold="true" box="[136,211,1898,1918]" pageId="13" pageNumber="14">Fig. 10.</emphasis>
<taxonomicName id="4C304D1DFF91041DCAEAF8813E6BF88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[224,361,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF91041DCAEAF8813E6BF88B" box="[224,361,1898,1918]" italics="true" pageId="13" pageNumber="14">Caridina beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF91041DCB65F8813EB9F88B" box="[367,443,1898,1918]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
(A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483).
</paragraph>
</caption>
<footnote id="E82B2A90FF91041DCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="13" pageNumber="14">
<paragraph id="8B8F369EFF91041DCA82F8363E9AF81A" blockId="13.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="13" pageNumber="14">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF92041ECAA4F86E3AA9F81A" ID-DOI="http://doi.org/10.5281/zenodo.13315015" ID-Zenodo-Dep="13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="14" pageNumber="15" startId="14.[174,208,1925,1945]" targetBox="[205,1322,249,1864]" targetPageId="14" targetType="figure">
<paragraph id="8B8F369EFF92041ECAA4F86E3A96F840" blockId="14.[174,1451,1924,2031]" pageId="14" pageNumber="15">
<emphasis id="B944EA8CFF92041ECAA4F86E3FF5F86C" bold="true" box="[174,247,1925,1945]" pageId="14" pageNumber="15">Fig. 11.</emphasis>
<taxonomicName id="4C304D1DFF92041ECB0FF86F3E8FF86C" authorityName="Cai &amp; Ng" authorityYear="2018" box="[261,397,1924,1945]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF92041ECB0FF86F3E8FF86C" box="[261,397,1924,1945]" italics="true" pageId="14" pageNumber="15">Caridina beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF92041ECB9EF86E3EDDF86C" box="[404,479,1925,1945]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483).
</paragraph>
<paragraph id="8B8F369EFF92041ECE96F8363AA9F81A" blockId="14.[174,1451,1924,2031]" box="[1180,1451,2013,2031]" pageId="14" pageNumber="15">© 2018 Academia Sinica, Taiwan</paragraph>
</caption>
<caption id="DF4F6616FF93041FCA82F8813CFEF843" ID-DOI="http://doi.org/10.5281/zenodo.13315017" ID-Zenodo-Dep="13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="15" pageNumber="16" startId="15.[136,170,1898,1918]" targetBox="[142,1414,261,1859]" targetPageId="15" targetType="figure">
<paragraph id="8B8F369EFF93041FCA82F8813CFEF843" blockId="15.[136,1413,1898,1974]" pageId="15" pageNumber="16">
<emphasis id="B944EA8CFF93041FCA82F8813FD1F88B" bold="true" box="[136,211,1898,1918]" pageId="15" pageNumber="16">Fig. 12.</emphasis>
<taxonomicName id="4C304D1DFF93041FCAE8F8813E69F88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[226,363,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF93041FCAE8F8813E69F88B" box="[226,363,1898,1918]" italics="true" pageId="15" pageNumber="16">Caridina beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF93041FCB78F8813EBCF88B" box="[370,446,1898,1918]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291).
</paragraph>
</caption>
<footnote id="E82B2A90FF93041FCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="15" pageNumber="16">
<paragraph id="8B8F369EFF93041FCA82F8363E9AF81A" blockId="15.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="15" pageNumber="16">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF8C0400CAE0FDF63EDDFB63" blockId="16.[174,776,253,1975]" pageId="16" pageNumber="17">
First pereiopod (
<figureCitation id="130B2A1BFF8C0400CBD8FDF63D48FDC3" box="[466,586,541,566]" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="16" pageNumber="17">Fig. 11A</figureCitation>
) short, stout, reaching distal end of eyes, merus 2.2 times as long as broad; carpus concave anteriorly, 1.5 times as long as high, chela 1.9 times as long as broad, fingers 0.7 times as long as palm. Second pereiopod (
<figureCitation id="130B2A1BFF8C0400CB3CFD563E9EFD23" box="[310,412,701,726]" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="16" pageNumber="17">Fig. 11B</figureCitation>
) reaching slightly beyond end of basal segment of antennular peduncle, carpus 4.0 times as long as high, chela shorter than carpus, 2.1 times as long as broad, fingers 1.5 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF8C0400C893FCD63FC1FC83" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="16" pageNumber="17">Fig. 11C, D</figureCitation>
) reaching beyond distal end of scaphocerite, propodus 10 times as long as broad, 4.7 times as long as dactylus (claw included); dactylus terminating as a claw, with 4 accessory spines increasing in length and size distally on flexor margin. Fifth pereiopod (
<figureCitation id="130B2A1BFF8C0400CBFBFC163D87FBE3" box="[497,645,1021,1046]" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="16" pageNumber="17">Fig. 11E, F</figureCitation>
) reaching beyond end of second segment of antennular peduncle, propodus 15 times as long as wide, 3.0 times as long as dactylus; dactylus with 46 spinules on flexor margin.
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0FB763E0EFA83" blockId="16.[174,776,253,1975]" pageId="16" pageNumber="17">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF8C0400C886FB763FE4FB23" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="16" pageNumber="17">Fig. 12D, E, F</figureCitation>
) short, sub-elliptical, 3.0 times as long as wide, inner margin straight, outer margin rounded, appendix interna at distal part of endopod, reaching beyond distal end of endopod. Appendix interna of male second pleopod (
<figureCitation id="130B2A1BFF8C0400C848FAD63DADFAA2" box="[578,687,1341,1367]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="16" pageNumber="17">Fig. 12G</figureCitation>
) small, slender.
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0FA963E1AFA43" blockId="16.[174,776,253,1975]" pageId="16" pageNumber="17">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF8C0400C805FA963D70FA63" box="[527,626,1405,1430]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
H) with 13 spinules.
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0FA563DDFFA23" blockId="16.[174,776,253,1975]" box="[234,733,1469,1495]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400CAE0FA563E3CFA23" box="[234,318,1469,1494]" italics="true" pageId="16" pageNumber="17">Habitat</emphasis>
: Subterrenean water in karst cave.
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0FA363E8DF9C3" blockId="16.[174,776,253,1975]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400CAE0FA363E69FA03" box="[234,363,1501,1526]" italics="true" pageId="16" pageNumber="17">Etymology</emphasis>
: The species is named after the
<typeStatus id="548B883CFF8C0400CAA4FA163FE2F9E3" box="[174,224,1533,1558]" pageId="16" pageNumber="17">type</typeStatus>
locality,
<taxonomicName id="4C304D1DFF8C0400CB42FA163EEFF9E2" authority="County." authorityName="County." box="[328,493,1533,1559]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County.</taxonomicName>
The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0F9D63E61F863" blockId="16.[174,776,253,1975]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400CAE0F9D63E54F9A3" box="[234,342,1597,1622]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
:
<taxonomicName id="4C304D1DFF8C0400CB62F9D63D14F9A3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[360,534,1597,1622]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF8C0400CB62F9D63D14F9A3" box="[360,534,1597,1622]" italics="true" pageId="16" pageNumber="17">Caridina beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF8C0400C82BF9D63D83F9A3" box="[545,641,1597,1622]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C304D1DFF8C0400CAA4F9B63D4FF983" authority="Liang and Yan, 1986" authorityName="Liang and Yan" authorityYear="1986" box="[174,589,1629,1654]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B944EA8CFF8C0400CAA4F9B63E31F983" box="[174,307,1629,1654]" italics="true" pageId="16" pageNumber="17">C. cornuta</emphasis>
<bibRefCitation id="EFA14B6FFF8C0400CB34F9B63D4FF983" author="Liang X &amp; Yan S." box="[318,589,1629,1654]" pageId="16" pageNumber="17" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49F4F87CFF8C0400C895F9B63E1FF963" country="China" name="Guizhou" pageId="16" pageNumber="17">Guizhou Province</collectingRegion>
, in the form of the rostrum. However, it can be distinguished from
<taxonomicName id="4C304D1DFF8C0400CBE3F9763D67F943" authorityName="Liang and Yan" authorityYear="1986" box="[489,613,1693,1718]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B944EA8CFF8C0400CBE3F9763D67F943" box="[489,613,1693,1718]" italics="true" pageId="16" pageNumber="17">C. cornuta</emphasis>
</taxonomicName>
by the telson possessing a posteromedian projection (
<figureCitation id="130B2A1BFF8C0400C89CF9563C02F923" box="[662,768,1725,1750]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="16" pageNumber="17">Fig. 12B</figureCitation>
) (vs. with no projection in
<taxonomicName id="4C304D1DFF8C0400CBD7F9363D58F903" authorityName="Liang and Yan" authorityYear="1986" box="[477,602,1757,1782]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B944EA8CFF8C0400CBD7F9363D58F903" box="[477,602,1757,1782]" italics="true" pageId="16" pageNumber="17">C. cornuta</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="EFA14B6FFF8C0400C89AF9363E2BF8E3" author="Liang X &amp; Yan S." pageId="16" pageNumber="17" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
: fig. 3(2)); and the sub-elliptical endopod of the male pleopod does not have a projection at the base (
<figureCitation id="130B2A1BFF8C0400CB46F8D63EB4F8A3" box="[332,438,1853,1878]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="16" pageNumber="17">Fig. 12F</figureCitation>
) (vs. subrectangular with a basal projection in
<taxonomicName id="4C304D1DFF8C0400CB9DF8B63D14F883" authorityName="Liang and Yan" authorityYear="1986" box="[407,534,1885,1910]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B944EA8CFF8C0400CB9DF8B63D14F883" box="[407,534,1885,1910]" italics="true" pageId="16" pageNumber="17">C. cornuta</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="EFA14B6FFF8C0400C845F8B63FECF863" author="Liang X &amp; Yan S." pageId="16" pageNumber="17" pagination="196 - 206" refId="ref16020" refString="Liang X, Yan S. 1986. Study on Caridina (Decapoda, Caridea) from Guizhou Province, China. Oceanol Limnol Sinica Suppl 1986: 196 - 206, figures 1 - 5. (in Chinese with English abstract)" type="journal article" year="1986">Liang and Yan, 1986</bibRefCitation>
: fig. 3(7)).
</paragraph>
<paragraph id="8B8F369EFF8C0400CAE0F8763CF4FEE2" blockId="16.[174,776,253,1975]" box="[234,1014,253,1975]" lastBlockId="16.[850,1014,253,279]" pageId="16" pageNumber="17">
<emphasis id="B944EA8CFF8C0400CAE0F8763E87F843" box="[234,389,1949,1974]" italics="true" pageId="16" pageNumber="17">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF8C0400CB96F8763D8CF843" box="[412,654,1949,1975]" country="China" name="Guangxi" pageId="16" pageNumber="17">Guangxi Province</collectingRegion>
(Goulou Cave),
<collectingCountry id="F327760EFF8C0400C9A2FF163CF0FEE2" box="[936,1010,253,279]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,313 @@
<document id="D36625560F12970973AA6B121F6BC441" ID-DOI="10.6620/ZS.2018.57-27" ID-ISSN="1810-522X" ID-PMC="PMC6517715" ID-PubMed="31966267" ID-Zenodo-Dep="13314991" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721916104468" checkinUser="felipe" docAuthor="Cai, Yixiong &amp; Ng, Peter Kee Lin" docDate="2018" docId="03998788FF94041CCB3BFA373CF4FE62" docLanguage="en" docName="ZoolStud.57.27.1-33.pdf" docOrigin="Zoological Studies 57 (27)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Caridina spinicrus Cai &amp; Ng, 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="FFA0FFF0FF9C0410CA0AFFEB3F02FFF5" masterDocTitle="Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="9" updateTime="1723578273083" updateUser="ExternalLinkService">
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<mods:title id="A56EB1B39D2C290B0B3435406122D743">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="1B21DCC3C707281FBE3D729414B6E838">Cai, Yixiong</mods:namePart>
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<mods:namePart id="F8BA3119301C1FD736FE4518DB58F2F0">Ng, Peter Kee Lin</mods:namePart>
<mods:affiliation id="2097D2E9D185FE0AC58CA9FD2C850F03">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of &amp; National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<treatment id="03998788FF94041CCB3BFA373CF4FE62" ID-DOI="http://doi.org/10.5281/zenodo.12826764" ID-Zenodo-Dep="12826764" LSID="urn:lsid:zoobank.org:act:EF868C18-D01C-4920-A9C4-51DFACB8D42C" httpUri="http://treatment.plazi.org/id/03998788FF94041CCB3BFA373CF4FE62" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
<subSubSection id="C32A6515FF940418CB3BFA373D81FA03" box="[305,643,1500,1526]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8F369EFF940418CB3BFA373D81FA03" blockId="8.[183,766,1500,1610]" box="[305,643,1500,1526]" pageId="8" pageNumber="9">
<heading id="D0C781F2FF940418CB3BFA373D81FA03" bold="true" box="[305,643,1500,1526]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
<emphasis id="B944EA8CFF940418CB3BFA373D81FA03" bold="true" box="[305,643,1500,1526]" pageId="8" pageNumber="9">
<taxonomicName id="4C304D1DFF940418CB3BFA373D18FA03" ID-CoL="8VKFL" authorityName="Cai &amp; Ng" authorityYear="2018" box="[305,538,1500,1526]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF940418CB3BFA373D18FA03" bold="true" box="[305,538,1500,1526]" italics="true" pageId="8" pageNumber="9">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF940418C82BFA363D81FA03" box="[545,643,1501,1526]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6515FF94041CCB94FA163CF4FE62" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
<paragraph id="8B8F369EFF940418CB94FA163D15F9E3" blockId="8.[183,766,1500,1610]" box="[414,535,1533,1558]" pageId="8" pageNumber="9">
(
<figureCitation id="130B2A1BFF940418CBAFFA163D12F9E3" box="[421,528,1533,1558]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[136,170,1898,1918]" captionStartId-1="10.[174,208,1898,1918]" captionStartId-2="11.[136,170,1898,1918]" captionTargetBox-0="[252,1337,267,1858]" captionTargetBox-1="[301,1322,254,1855]" captionTargetBox-2="[204,1333,269,1863]" captionTargetId-0="figure-6@9.[196,1354,249,1870]" captionTargetId-1="figure-58@10.[275,1351,249,1870]" captionTargetId-2="figure-58@11.[195,1354,249,1870]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." captionText-1="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." captionText-2="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi-0="http://doi.org/10.5281/zenodo.13315007" figureDoi-1="http://doi.org/10.5281/zenodo.13315009" figureDoi-2="http://doi.org/10.5281/zenodo.13315011" httpUri-0="https://zenodo.org/record/13315007/files/figure.png" httpUri-1="https://zenodo.org/record/13315009/files/figure.png" httpUri-2="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Figs. 7-9</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF940418CABDF9F33D37F9BF" blockId="8.[183,766,1500,1610]" pageId="8" pageNumber="9">
<uri id="FFA13A9CFF940418CABDF9F33D37F9BF" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="FF960C4BFF940418CBB0F9F33D37F9BF" pageId="8" pageNumber="9">EF868C18-D01C-4920-A9C4- 51DFACB8D42C</uuid>
</uri>
</paragraph>
<paragraph id="8B8F369EFF940418CAE0F99B3D4DF8DC" blockId="8.[174,776,1648,1961]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0F99B3EEFF97C" box="[234,493,1648,1673]" italics="true" pageId="8" pageNumber="9">Material examined</emphasis>
:
<materialsCitation id="3B583CC3FF940418C80FF99B3FF2F8FC" collectingDate="1993-10-30" collectionCode="IZCAS" collectorName="Y. Cai" country="China" location="Goulou Cave" pageId="8" pageNumber="9" specimenCode="DE 5031" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="548B883CFF940418C80FF99B3D7DF97C" box="[517,639,1648,1673]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF940418CAA4F97B3E11F95C" box="[174,275,1680,1705]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="9" unit="mm" value="5.5">5.5 mm</quantity>
,
<collectionCode id="ED21AE5BFF940418CB28F97B3E76F95F" box="[290,372,1680,1706]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="9">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF940418CB8AF97B3EF7F95C" box="[384,501,1680,1705]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="9">DE 5031</specimenCode>
, artificial pond inside
<location id="8EEF6045FF940418CAA4F95B3E4CF93F" LSID="urn:lsid:plazi:treatment:03998788FF94041CCB3BFA373CF4FE62:8EEF6045FF940418CAA4F95B3E4CF93F" box="[174,334,1712,1738]" country="China" name="Goulou Cave" pageId="8" pageNumber="9" stateProvince="Guangxi">Goulou Cave</location>
in
<taxonomicName id="4C304D1DFF940418CB7DF95B3D19F93F" authority="County" authorityName="County" box="[375,539,1712,1738]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County</taxonomicName>
,
<collectingRegion id="49F4F87CFF940418C822F95B3C01F93C" box="[552,771,1712,1738]" country="China" name="Guangxi" pageId="8" pageNumber="9">Guangxi Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF940418CB3CF93B3E87F91C" box="[310,389,1744,1770]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
, coll.
<collectorName id="26C55348FF940418CBECF93B3D38F91C" box="[486,570,1744,1770]" pageId="8" pageNumber="9">Y. Cai</collectorName>
et al.,
<date id="FF8E105EFF940418C8A0F93B3FF2F8FC" pageId="8" pageNumber="9" value="1993-10-30">
<collectingDate id="EFCAE9B6FF940418C8A0F93B3FF2F8FC" pageId="8" pageNumber="9" value="1993-10-30">30 Oct 1993</collectingDate>
</date>
</materialsCitation>
;
<typeStatus id="548B883CFF940418CAF5F91B3E81F8FC" box="[255,387,1776,1801]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF940418CB98F91B3EF9F8FC" box="[402,507,1776,1801]" count="2" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
, cl 5.5-6.0 mm, ZRC 2018.0482, same data as
<typeStatus id="548B883CFF940418CBE8F8FB3D49F8DC" box="[482,587,1808,1833]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0F8DB3A14FE83" blockId="8.[174,776,1648,1961]" lastBlockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0F8DB3E83F8BC" box="[234,385,1840,1865]" italics="true" pageId="8" pageNumber="9">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF940418C814F8DB3D87F8BC" box="[542,645,1840,1865]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
) straight, reaching anterior half of second segment of antennular peduncle, dorsally with 17 teeth, of which 7 on carapace behind posterior margin of orbital margin, occupying 0.14 carapace length, armed ventrally with only 1 small tooth. Antennal spine sharp, completely fused with suborbital angle. Pterygostomian angle rounded.
</paragraph>
<paragraph id="8B8F369EFF940418C985FE963B34FE03" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Telson (
<figureCitation id="130B2A1BFF940418CE0DFE963B73FE63" box="[1031,1137,381,406]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9B</figureCitation>
) with posteromedian projection, posterior margin rounded, lateral pair of spines longer than intermediate pairs, sub-lateral pair being shortest.
</paragraph>
<paragraph id="8B8F369EFF940418C985FE163CBCFD43" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Eyes well developed (
<figureCitation id="130B2A1BFF940418CEA3FE163A0FFDE3" box="[1193,1293,509,534]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
). Antennular peduncle 0.50 times as long as carapace. Stylocerite reaching 0.7 basal segment length of antennular peduncle. Scaphocerite (
<figureCitation id="130B2A1BFF940418CF49FDB63AA1FD83" box="[1347,1443,605,630]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8H</figureCitation>
) 3.0 times as long as wide, outer margin slightly concave.
</paragraph>
<paragraph id="8B8F369EFF940418C985FD563CDDFB43" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF940418CF1DFD563A6CFD23" box="[1303,1390,701,726]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7B</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF940418C950FD163CB8FCE2" box="[858,954,765,791]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7C</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF940418CF76FCD63C75FC83" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7D</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF940418CF73FC763C77FC23" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7E</figureCitation>
) ending as triangular projection. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF940418CE4DFC363BA7FC03" box="[1095,1189,989,1014]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7E</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF940418CF42FC163AA6FBE2" box="[1352,1444,1021,1047]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7G</figureCitation>
) reaching beyond distal end of scaphocerite, ending in single terminal claw; exopod reaching 0.5 times penultimate segment length, ultimate segment as long as pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF940418C985FB563AA9F842" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
First pereiopod (
<figureCitation id="130B2A1BFF940418CE53FB563BDAFB23" box="[1113,1240,1213,1238]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8A, B</figureCitation>
) reaching end of basal segment of antennular peduncle, merus 2.7 times as long as broad; carpus concave anteriorly, 1.7 times as long as high, chela 2.5 times as long as broad, fingers 1.3 times as long as palm. Second pereiopod (
<figureCitation id="130B2A1BFF940418CE47FAB63BACFA82" box="[1101,1198,1373,1399]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8C</figureCitation>
) reaching to end of antennular peduncle, carpus 5.2 times as long as high, chela slightly shorter than carpus, 2.7 times as long as broad, fingers 1.7 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF940418CEA2FA363A0BFA03" box="[1192,1289,1501,1526]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8D</figureCitation>
) long, stout, reaching beyond distal end of scaphocerite. Merus 4.0 times as long as broad; carpus 2.4 times as long as broad, ventral surface armed with a row of 16 closely packed strong spines. Propodus strongly curved inwards, 9.0 times as long as broad, 4.0 times as long as dactylus (claw included), ventral surface armed with row of 23 strong spines, similar to those of at carpus, but more widely lined anteriorly; dactylus terminating as a claw, with 5 accessory spines on flexor margin. Fourth pereiopod (
<figureCitation id="130B2A1BFF940418C9EAF8D63B8BF8A3" box="[992,1161,1853,1879]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9C, D, E</figureCitation>
) reaching to distal end of antennular peduncle, similar to third pereiopod in form, with strong spines on ventral surfaces of carpus and propodus. Fifth pereiopod (
<figureCitation id="130B2A1BFF940418CF23F8763AA6F842" box="[1321,1444,1949,1975]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9F, G</figureCitation>
)
</paragraph>
<footnote id="E82B2A90FF940418CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">
<paragraph id="8B8F369EFF940418CE96F8363AA9F81A" blockId="8.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF950419CADDFC2C3FF1FC2E" blockId="9.[215,243,967,987]" box="[215,243,967,987]" pageId="9" pageNumber="10">(B)</paragraph>
<caption id="DF4F6616FF950419CA82F8813B74F843" ID-DOI="http://doi.org/10.5281/zenodo.13315007" ID-Zenodo-Dep="13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,170,1898,1918]" targetBox="[252,1337,267,1858]" targetPageId="9" targetType="figure">
<paragraph id="8B8F369EFF950419CA82F8813B74F843" blockId="9.[136,1415,1898,1974]" pageId="9" pageNumber="10">
<emphasis id="B944EA8CFF950419CA82F8813FC5F88B" bold="true" box="[136,199,1898,1918]" pageId="9" pageNumber="10">Fig. 7.</emphasis>
<taxonomicName id="4C304D1DFF950419CADDF8813E80F88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[215,386,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF950419CADDF8813E80F88B" box="[215,386,1898,1918]" italics="true" pageId="9" pageNumber="10">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF950419CB83F8813ED4F88B" box="[393,470,1898,1918]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
(A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF950419CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="9" pageNumber="10">
<paragraph id="8B8F369EFF950419CA82F8363E9AF81A" blockId="9.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="9" pageNumber="10">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF96041ACAA4F8813D18F843" ID-DOI="http://doi.org/10.5281/zenodo.13315009" ID-Zenodo-Dep="13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,208,1898,1918]" targetBox="[301,1322,254,1855]" targetPageId="10" targetType="figure">
<paragraph id="8B8F369EFF96041ACAA4F8813D18F843" blockId="10.[174,1453,1898,1974]" pageId="10" pageNumber="11">
<emphasis id="B944EA8CFF96041ACAA4F8813FE9F88B" bold="true" box="[174,235,1898,1918]" pageId="10" pageNumber="11">Fig. 8.</emphasis>
<taxonomicName id="4C304D1DFF96041ACAFDF8813EA3F88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[247,417,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF96041ACAFDF8813EA3F88B" box="[247,417,1898,1918]" italics="true" pageId="10" pageNumber="11">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF96041ACBADF8813EF0F88B" box="[423,498,1898,1918]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF96041ACE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">
<paragraph id="8B8F369EFF96041ACE96F8363AA9F81A" blockId="10.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF97041BCA82F8813FFEF843" ID-DOI="http://doi.org/10.5281/zenodo.13315011" ID-Zenodo-Dep="13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,170,1898,1918]" targetBox="[204,1333,269,1863]" targetPageId="11" targetType="figure">
<paragraph id="8B8F369EFF97041BCA82F8813FFEF843" blockId="11.[136,1414,1898,1974]" pageId="11" pageNumber="12">
<emphasis id="B944EA8CFF97041BCA82F8813FC5F88B" bold="true" box="[136,199,1898,1918]" pageId="11" pageNumber="12">Fig. 9.</emphasis>
<taxonomicName id="4C304D1DFF97041BCADFF8813E7DF88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[213,383,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF97041BCADFF8813E7DF88B" box="[213,383,1898,1918]" italics="true" pageId="11" pageNumber="12">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF97041BCB8DF8813ED0F88B" box="[391,466,1898,1918]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF97041BCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="11" pageNumber="12">
<paragraph id="8B8F369EFF97041BCA82F8363E9AF81A" blockId="11.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="11" pageNumber="12">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF90041CCAA4FF163E0AFE43" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">reaching distal end of antennular peduncle; merus 4.0 times as long as broad; carpus 4.0 times as long as broad; propodus slightly curved inwards, 10 times as long as broad, 3.8 times as long as dactylus; dactylus with 50 spinules on flexor margin.</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FE563DA8FCE3" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF90041CC893FE563DFCFE23" box="[665,766,445,470]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) reaching to 0.67 endopod length, subrectangular, distal end rounded, inner margin slightly curved backwards, 3.0 times as long as wide; appendix interna situated at posterior 0.2 endopod length, extending to end of endopod. Appendix masculina of male second pleopod (
<figureCitation id="130B2A1BFF90041CCBFFFD963D7AFD62" box="[501,632,637,663]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8F, G</figureCitation>
) narrowing down distally, with 2 rows of spines at inner surface, reaching to 0.75 of endopod length; appendix interna of male second pleopod small, reaching to 0.4 appendix masculina length.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FCF63C07FCC3" blockId="12.[174,775,253,1974]" box="[234,773,797,822]" pageId="12" pageNumber="13">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF90041CCBD2FCF63D32FCC3" box="[472,560,797,822]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7H</figureCitation>
) with 13 spinules.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FCD63E8FFC23" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FCD63E4AFCA3" box="[234,328,829,854]" italics="true" pageId="12" pageNumber="13">Habitat</emphasis>
:
<taxonomicName id="4C304D1DFF90041CCB55FCD63D52FCA3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[351,592,829,854]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCB55FCD63D52FCA3" box="[351,592,829,854]" italics="true" pageId="12" pageNumber="13">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC854FCD63DCAFCA3" box="[606,712,829,854]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
was collected together with
<taxonomicName id="4C304D1DFF90041CCBE9FCB63D4DFC83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[483,591,861,886]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCBE9FCB63D4DFC83" box="[483,591,861,886]" italics="true" pageId="12" pageNumber="13">C. beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC851FCB63DC0FC83" box="[603,706,861,886]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
from an artificial concrete pond inside Goulou Cave in
<taxonomicName id="4C304D1DFF90041CCADEFC763E98FC43" authority="County." authorityName="County." box="[212,410,925,951]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County.</taxonomicName>
The water comes from a subterranean river.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FC363D08FBA3" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FC363E64FC03" box="[234,358,989,1014]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
:
<taxonomicName id="4C304D1DFF90041CCB7CFC363D48FC03" authorityName="Cai &amp; Ng" authorityYear="2018" box="[374,586,989,1014]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCB7CFC363D48FC03" box="[374,586,989,1014]" italics="true" pageId="12" pageNumber="13">Caridina spinicrus</emphasis>
</taxonomicName>
is named for its distinctive spiny third and fourth pereiopods, from the Latin for spiny and legs (crura). The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FBB63E56F983" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FBB63E54FB83" box="[234,342,1117,1142]" italics="true" pageId="12" pageNumber="13">Remarks</emphasis>
: With regard to the spiny third and fourth pereiopods, only one species,
<taxonomicName id="4C304D1DFF90041CC891FB963EE8FB43" authority="Gurney, 1984" authorityName="Gurney" authorityYear="1984" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CC891FB963E3BFB43" italics="true" pageId="12" pageNumber="13">Caridina crurispinata</emphasis>
<bibRefCitation id="EFA14B6FFF90041CCB49FB763EE8FB43" author="Gurney AR" box="[323,490,1181,1207]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
</taxonomicName>
, described from a cave in
<collectingCountry id="F327760EFF90041CCAC7FB563E64FB23" box="[205,358,1213,1238]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
, shares such features. Compared with
<taxonomicName id="4C304D1DFF90041CCAE0FB363EA2FB03" authorityName="Gurney" authorityYear="1984" box="[234,416,1245,1270]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CCAE0FB363EA2FB03" box="[234,416,1245,1270]" italics="true" pageId="12" pageNumber="13">C. crurispinata</emphasis>
</taxonomicName>
, the spines on the third and fourth pereiopods in
<taxonomicName id="4C304D1DFF90041CCBC9FB163D5CFAE3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[451,606,1277,1302]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCBC9FB163D5CFAE3" box="[451,606,1277,1302]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC861FB163DD0FAE3" box="[619,722,1277,1302]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
are more prominent (
<figureCitation id="130B2A1BFF90041CCB75FAF63EDCFAC3" box="[383,478,1309,1334]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8D</figureCitation>
;
<figureCitation id="130B2A1BFF90041CCBE6FAF63D8DFAC3" box="[492,655,1309,1335]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="12" pageNumber="13">Fig. 9C, D, E</figureCitation>
) (vs. less prominent; cf.
<bibRefCitation id="EFA14B6FFF90041CCB6BFAD63D09FAA3" author="Gurney AR" box="[353,523,1341,1367]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 6C). Apart from this,
<taxonomicName id="4C304D1DFF90041CCAE4FAB63E85FA83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[238,391,1373,1398]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCAE4FAB63E85FA83" box="[238,391,1373,1398]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C304D1DFF90041CC8E1FAB63E3AFA63" authorityName="Gurney" authorityYear="1984" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CC8E1FAB63E3AFA63" italics="true" pageId="12" pageNumber="13">C. crurispinata</emphasis>
</taxonomicName>
by the straight rostrum which reaches to the anterior half of the second segment of antennular peduncle (
<figureCitation id="130B2A1BFF90041CCBDFFA563D39FA23" box="[469,571,1469,1494]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
) (vs. gradually curving upwards and reaching beyond the antennal scale; cf.
<bibRefCitation id="EFA14B6FFF90041CCB2BFA163EC4F9E3" author="Gurney AR" box="[289,454,1533,1559]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 4B); and the presence of an appendix interna on the endopod of the male first pleopod (
<figureCitation id="130B2A1BFF90041CCB68F9D63EC1F9A3" box="[354,451,1597,1622]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) (vs. absent; cf.
<bibRefCitation id="EFA14B6FFF90041CC8AEF9D63FECF983" author="Gurney AR" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 7B).
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0F9963BB7FEA3" blockId="12.[174,775,253,1974]" lastBlockId="12.[850,1451,253,407]" pageId="12" pageNumber="13">
Compared with other Chinese species,
<taxonomicName id="4C304D1DFF90041CC8E1F9963E15F943" authorityName="Cai &amp; Ng" authorityYear="2018" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CC8E1F9963E15F943" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
also superficially resembles the sympatric
<taxonomicName id="4C304D1DFF90041CCAA4F9563E12F923" authorityName="Cai &amp; Ng" authorityYear="2018" box="[174,272,1725,1750]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCAA4F9563E12F923" box="[174,272,1725,1750]" italics="true" pageId="12" pageNumber="13">C. beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CCB13F9563E75F923" box="[281,375,1725,1750]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
However,
<taxonomicName id="4C304D1DFF90041CCBF2F9563D87F923" authorityName="Cai &amp; Ng" authorityYear="2018" box="[504,645,1725,1750]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCBF2F9563D87F923" box="[504,645,1725,1750]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
can easily be separated by the rostrum not reaching beyond the end of the second segment of the antennular peduncle (
<figureCitation id="130B2A1BFF90041CCB3CF8F63E9AF8C3" box="[310,408,1821,1846]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
) (vs. beyond the end of the second segment of the antennular peduncle;
<figureCitation id="130B2A1BFF90041CC8D0F8D63FE1F883" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10A</figureCitation>
); the more slender first pereiopod having the dactylus 2.5 times as long as broad (
<figureCitation id="130B2A1BFF90041CC874F8963DFDF863" box="[638,767,1917,1942]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8A, B</figureCitation>
) (vs. 1.5 times;
<figureCitation id="130B2A1BFF90041CCB51F8763EC3F843" box="[347,449,1949,1974]" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="12" pageNumber="13">Fig. 11A</figureCitation>
); and the male first pleopod having an appendix interna that reaches to the distal end of the endopod (
<figureCitation id="130B2A1BFF90041CCE95FEF63BF9FEC3" box="[1183,1275,285,310]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) (vs. distinctly reaching beyond;
<figureCitation id="130B2A1BFF90041CCE2CFED63B7BFEA3" box="[1062,1145,317,342]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
D-F).
</paragraph>
<paragraph id="8B8F369EFF90041CC985FEB63CF4FE62" blockId="12.[850,1451,253,407]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CC985FEB63B28FE83" box="[911,1066,349,374]" italics="true" pageId="12" pageNumber="13">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF90041CCE4BFEB63A31FE83" box="[1089,1331,349,375]" country="China" name="Guangxi" pageId="12" pageNumber="13">Guangxi Province</collectingRegion>
(Goulou Cave),
<collectingCountry id="F327760EFF90041CC9A2FE963CF0FE62" box="[936,1010,381,407]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,341 @@
<document id="987675CA88FA4B97D77F2E41FE8292D1" ID-DOI="10.6620/ZS.2018.57-27" ID-ISSN="1810-522X" ID-PMC="PMC6517715" ID-PubMed="31966267" ID-Zenodo-Dep="13314991" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721916104468" checkinUser="felipe" docAuthor="Cai, Yixiong &amp; Ng, Peter Kee Lin" docDate="2018" docId="03998788FF980418CB53FBD73FFEFA42" docLanguage="en" docName="ZoolStud.57.27.1-33.pdf" docOrigin="Zoological Studies 57 (27)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Caridina alu Cai &amp; Ng, 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="FFA0FFF0FF9C0410CA0AFFEB3F02FFF5" masterDocTitle="Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="5" updateTime="1723578273083" updateUser="ExternalLinkService">
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<mods:titleInfo id="2AE6C413DFF634A8D1BA53BE89647CB9">
<mods:title id="2C42E91D064AE47E61051479F8B4594B">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="CA7898BB27FE749FCD6CE1C01E79C245" type="personal">
<mods:role id="8FB2D29D5C4CA317B8CA1535376DE84E">
<mods:roleTerm id="8BACF68C5ADD9E736F11E61125483A92">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2E52A0B2BB79A7D824AD450177F1019">Cai, Yixiong</mods:namePart>
<mods:affiliation id="7397891C625D29C508181251F7E5A5A7">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name id="DD21DBB613382CE974D76D8452E22D07" type="personal">
<mods:role id="6FAE69CEADC1A2A99DF4DB26E2558929">
<mods:roleTerm id="2B3C2FF70911263E621EBA39F96BB877">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE5B26E93BBF7388D351102ACEF0DD35">Ng, Peter Kee Lin</mods:namePart>
<mods:affiliation id="663C88914B35DAF9840581D40AE849E2">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of &amp; National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6AEAFD6C59F22966C5D98EDC0B79F5EA">text</mods:typeOfResource>
<mods:relatedItem id="F3CE6549BF4E2A0B3D9EA18C0C6D4515" type="host">
<mods:titleInfo id="19C97EB75EBE93892D92D9A0BBDF1A92">
<mods:title id="AD9108C8A7B8E2903C2A14C6CDD3C686">Zoological Studies</mods:title>
</mods:titleInfo>
<mods:part id="A2181AE2E2A35F884ACA9BB0CFC27B70">
<mods:date id="6C2B56C0D2784E72253406F2D4739EDC">2018</mods:date>
<mods:detail id="B18F48EE3E0C12BC6FA263238F9A7732" type="pubDate">
<mods:number id="D343C455A71AE2E73EC257EB200F6E90">2018-06-26</mods:number>
</mods:detail>
<mods:detail id="F7BD45A9508C8213321E19F016A56108" type="volume">
<mods:number id="3B3F73C8B89357E0BE903BDD3EEF32B1">57</mods:number>
</mods:detail>
<mods:detail id="9594A260D695294A93D606EF4E77385B" type="issue">
<mods:number id="D5E1D468C006A74CAD6DDFB1E634D3D9">27</mods:number>
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<mods:start id="6AB25FC7258AB54C8B9F3AF7835F8833">1</mods:start>
<mods:end id="F69C75D2CEB1BF6FA24AB5BAD802BC28">33</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="48414D337BDFA244236578597C0A19B7">journal article</mods:classification>
<mods:identifier id="DF3F4A76FB9984600333C33458D80522" type="DOI">10.6620/ZS.2018.57-27</mods:identifier>
<mods:identifier id="473E5471941607D336869D9210D8617E" type="ISSN">1810-522X</mods:identifier>
<mods:identifier id="37D9169A72488763C21668693BCA0F8A" type="PMC">PMC6517715</mods:identifier>
<mods:identifier id="FC5AD8C00E30A1ADF3FA5BE61896893F" type="PubMed">31966267</mods:identifier>
<mods:identifier id="A728EC041CD933486EB6DAB029B76B46" type="Zenodo-Dep">13314991</mods:identifier>
</mods:mods>
<treatment id="03998788FF980418CB53FBD73FFEFA42" ID-DOI="http://doi.org/10.5281/zenodo.12826762" ID-Zenodo-Dep="12826762" LSID="urn:lsid:zoobank.org:act:F7760EDF-36A2-48DD-9B49-409B6B51A1CD" httpUri="http://treatment.plazi.org/id/03998788FF980418CB53FBD73FFEFA42" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<subSubSection id="C32A6515FF980414CB53FBD73D59FBA3" box="[345,603,1084,1110]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B8F369EFF980414CB53FBD73D59FBA3" blockId="4.[183,766,1084,1194]" box="[345,603,1084,1110]" pageId="4" pageNumber="5">
<heading id="D0C781F2FF980414CB53FBD73D59FBA3" bold="true" box="[345,603,1084,1110]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="5" reason="2">
<emphasis id="B944EA8CFF980414CB53FBD73D59FBA3" bold="true" box="[345,603,1084,1110]" pageId="4" pageNumber="5">
<taxonomicName id="4C304D1DFF980414CB53FBD73EF0FBA3" ID-CoL="8VK3X" authorityName="Cai &amp; Ng" authorityYear="2018" box="[345,498,1084,1110]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alu" status="sp. nov.">
<emphasis id="B944EA8CFF980414CB53FBD73EF0FBA3" bold="true" box="[345,498,1084,1110]" italics="true" pageId="4" pageNumber="5">Caridina alu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF980414CBF3FBD63D59FBA3" box="[505,603,1085,1110]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6515FF980418CB94FBB63FFEFA42" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="description">
<paragraph id="8B8F369EFF980414CB94FBB63D15FB83" blockId="4.[183,766,1084,1194]" box="[414,535,1117,1142]" pageId="4" pageNumber="5">
(
<figureCitation id="130B2A1BFF980414CBAFFBB63D12FB83" box="[421,528,1117,1142]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="5.[136,170,1895,1915]" captionStartId-1="6.[174,208,1895,1915]" captionStartId-2="7.[136,170,1870,1890]" captionTargetBox-0="[210,1335,267,1844]" captionTargetBox-1="[265,1379,257,1867]" captionTargetBox-2="[196,1328,250,1830]" captionTargetId-0="figure-6@5.[210,1339,249,1866]" captionTargetId-1="figure-52@6.[209,1420,249,1869]" captionTargetId-2="figure-88@7.[196,1353,216,1852]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." captionText-1="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." captionText-2="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi-0="http://doi.org/10.5281/zenodo.13314999" figureDoi-1="http://doi.org/10.5281/zenodo.13315001" figureDoi-2="http://doi.org/10.5281/zenodo.13315005" httpUri-0="https://zenodo.org/record/13314999/files/figure.png" httpUri-1="https://zenodo.org/record/13315001/files/figure.png" httpUri-2="https://zenodo.org/record/13315005/files/figure.png" pageId="4" pageNumber="5">Figs. 4-6</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF980414CABDFB933D31FB5F" blockId="4.[183,766,1084,1194]" pageId="4" pageNumber="5">
<uri id="FFA13A9CFF980414CABDFB933D31FB5F" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
<uuid id="FF960C4BFF980414CBBDFB933D31FB5F" pageId="4" pageNumber="5">F7760EDF-36A2-48DD-9B49- 409B6B51A1CD</uuid>
</uri>
</paragraph>
<paragraph id="8B8F369EFF980414CAE0FB3B3D8CF9BC" blockId="4.[174,776,1232,1961]" pageId="4" pageNumber="5">
<emphasis id="B944EA8CFF980414CAE0FB3B3EEFFB1C" box="[234,493,1232,1257]" italics="true" pageId="4" pageNumber="5">Material examined</emphasis>
:
<materialsCitation id="3B583CC3FF980414C80FFB3B3E5AFA7C" collectingDate="1995-11-24" collectionCode="IZCAS" collectorName="Kos, B &amp; B. Sket &amp; Velkovrh." country="China" county="Luxi County" location="Alufa Village" pageId="4" pageNumber="5" specimenCode="DE 5029" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="548B883CFF980414C80FFB3B3D7DFB1C" box="[517,639,1232,1257]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF980414CAA4FB1B3E11FAFC" box="[174,275,1264,1289]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="5" unit="mm" value="6.5">6.5 mm</quantity>
,
<collectionCode id="ED21AE5BFF980414CB28FB1B3E77FAFF" box="[290,373,1264,1290]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="5">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF980414CB8BFB1B3EF4FAFC" box="[385,502,1264,1289]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="5">DE 5029</specimenCode>
,
<collectingMethod id="52714E89FF980414C80FFB1B3C07FAFC" box="[517,773,1264,1289]" pageId="4" pageNumber="5">caught by fish traps</collectingMethod>
in subterranean river,
<taxonomicName id="4C304D1DFF980414CBC2FAFB3D40FADF" authority="Cave" authorityName="Cave" box="[456,578,1296,1322]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alu">Alu Cave</taxonomicName>
, behind a hotel at
<location id="8EEF6045FF980414CAC5FADB3E71FABC" LSID="urn:lsid:plazi:treatment:03998788FF980418CB53FBD73FFEFA42:8EEF6045FF980414CAC5FADB3E71FABC" box="[207,371,1328,1353]" country="China" county="Luxi County" name="Alufa Village" pageId="4" pageNumber="5" stateProvince="Yunnan">Alufa Village</location>
,
<collectingCounty id="62EE4E12FF980414CB8BFADB3D18FABF" box="[385,538,1328,1354]" pageId="4" pageNumber="5">Luxi County</collectingCounty>
,
<collectingRegion id="49F4F87CFF980414C822FADB3C01FABC" box="[552,771,1328,1353]" country="China" name="Yunnan" pageId="4" pageNumber="5">Yunnan Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF980414CB22FABB3E75FA9F" box="[296,375,1360,1386]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
, coll.
<collectorName id="26C55348FF980414CBCFFABB3D1DFA9C" box="[453,543,1360,1385]" pageId="4" pageNumber="5">Kos, B</collectorName>
. Sket &amp;
<collectorName id="26C55348FF980414C89BFABB3C05FA9C" box="[657,775,1360,1385]" pageId="4" pageNumber="5">Velkovrh.</collectorName>
<date id="FF8E105EFF980414CAA4FA9B3E5AFA7C" box="[174,344,1392,1417]" pageId="4" pageNumber="5" value="1995-11-24">
<collectingDate id="EFCAE9B6FF980414CAA4FA9B3E5AFA7C" box="[174,344,1392,1417]" pageId="4" pageNumber="5" value="1995-11-24">24 Nov 1995</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="548B883CFF980414CB6DFA9B3EEEFA7C" box="[359,492,1392,1417]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF980414CBF1FA9B3D72FA7C" box="[507,624,1392,1417]" count="1" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
, cl 4.0 mm, IZCAS DE 5030, same data as
<typeStatus id="548B883CFF980414C83EFA7B3D9CFA5C" box="[564,670,1424,1449]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF980414C8A6FA7B3C01FA5C" box="[684,771,1424,1449]" count="1" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
, cl 6.0 mm,
<specimenCount id="9D36FD17FF980414CB4AFA5B3EB6FA3C" box="[320,436,1456,1481]" count="1" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF980414CBE8FA5B3D44FA3C" box="[482,582,1456,1481]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="5" unit="mm" value="4.8">4.8 mm</quantity>
, RMNH.CRUS. D.57290;
<specimenCount id="9D36FD17FF980414CB36FA3B3E9EFA1C" box="[316,412,1488,1513]" count="1" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF980414CBD0FA3B3D41FA1C" box="[474,579,1488,1513]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="5" unit="mm" value="6.2">6.2 mm</quantity>
,
<specimenCount id="9D36FD17FF980414C856FA3B3DDAFA1C" box="[604,728,1488,1513]" count="1" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF980414CAA4FA1B3E0DF9FC" box="[174,271,1520,1545]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="5" unit="mm" value="5.6">5.6 mm</quantity>
, UL, same data as
<typeStatus id="548B883CFF980414C80FFA1B3D73F9FC" box="[517,625,1520,1545]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF980414C875FA1B3DE7F9FC" box="[639,741,1520,1545]" count="2" pageId="4" pageNumber="5" type="male">2 males</specimenCount>
, cl
<quantity id="4CC89B7BFF980414CAA4F9FB3E3BF9DC" box="[174,313,1552,1577]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.3" metricValueMin="5.8" pageId="4" pageNumber="5" unit="mm" value="6.05" valueMax="6.3" valueMin="5.8">5.8-6.3 mm</quantity>
,
<specimenCount id="9D36FD17FF980414CB4EF9FB3EBFF9DC" box="[324,445,1552,1577]" count="4" pageId="4" pageNumber="5" type="female">4 females</specimenCount>
, 3.0-
<quantity id="4CC89B7BFF980414CBFEF9FB3D50F9DC" box="[500,594,1552,1577]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="5" unit="mm" value="4.7">4.7 mm</quantity>
, cl
<quantity id="4CC89B7BFF980414C872F9FB3C01F9DC" box="[632,771,1552,1577]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="6.2" metricValueMin="4.2" pageId="4" pageNumber="5" unit="mm" value="5.2" valueMax="6.2" valueMin="4.2">4.2-6.2 mm</quantity>
, ZRC 2018.0481, same data as
<typeStatus id="548B883CFF980414C82AF9DB3D8BF9BC" box="[544,649,1584,1609]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF980414CAE0F9BB3DB5F87C" blockId="4.[174,776,1232,1961]" pageId="4" pageNumber="5">
<emphasis id="B944EA8CFF980414CAE0F9BB3D85F99C" box="[234,647,1616,1641]" italics="true" pageId="4" pageNumber="5">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF980414C896F9BB3D48F97C" authority="Liang and Zhou, 1993" authorityName="Liang and Zhou" authorityYear="1993" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sphyrapoda">
<emphasis id="B944EA8CFF980414C896F9BB3E3BF97C" italics="true" pageId="4" pageNumber="5">Caridina sphyrapoda</emphasis>
<bibRefCitation id="EFA14B6FFF980414CB48F99B3D48F97C" author="Liang X &amp; Zhou J." box="[322,586,1648,1673]" pageId="4" pageNumber="5" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF980414C85BF99B3DB8F97C" box="[593,698,1648,1673]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF980414CAC6F97B3E2EF95C" box="[204,300,1680,1705]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="5" unit="mm" value="5.5">5.5 mm</quantity>
, SFU 85a-8-1, Longdong river, Laibing County,
<collectingRegion id="49F4F87CFF980414CB1DF95B3EF8F93C" box="[279,506,1712,1738]" country="China" name="Guangxi" pageId="4" pageNumber="5">Guangxi Province</collectingRegion>
,
<collectingCountry id="F327760EFF980414C802F95B3D57F93F" box="[520,597,1712,1738]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
, coll. J. Zhou,
<date id="FF8E105EFF980414CAA4F93B3E5AF91C" box="[174,344,1744,1769]" pageId="4" pageNumber="5" value="1985-05-24">24 May 1985</date>
.
<typeStatus id="548B883CFF980414CB6CF93B3ED8F91C" box="[358,474,1744,1769]" pageId="4" pageNumber="5" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D36FD17FF980414CBE2F93B3D58F91C" box="[488,602,1744,1769]" count="1" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF980414C88CF93B3DE8F91C" box="[646,746,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="5" unit="mm" value="3.8">3.8 mm</quantity>
, 1 ovigerous female, cl
<quantity id="4CC89B7BFF980414CBA1F91B3D09F8FC" box="[427,523,1776,1801]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="5" unit="mm" value="5.8">5.8 mm</quantity>
, SFU 85a-8-2, same data as
<typeStatus id="548B883CFF980414CB1BF8FB3E78F8DC" box="[273,378,1808,1833]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
. Others:
<specimenCount id="9D36FD17FF980414CBE2F8FB3D4EF8DC" box="[488,588,1808,1833]" count="3" pageId="4" pageNumber="5" type="male">3 males</specimenCount>
, cl
<quantity id="4CC89B7BFF980414C87CF8FB3C01F8DC" box="[630,771,1808,1833]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.3" metricValueMin="4.5" pageId="4" pageNumber="5" unit="mm" value="5.4" valueMax="6.3" valueMin="4.5">4.5-6.3 mm</quantity>
,
<specimenCount id="9D36FD17FF980414CAA4F8DB3E32F8BC" box="[174,304,1840,1865]" count="6" pageId="4" pageNumber="5" type="female">6 females</specimenCount>
, cl
<quantity id="4CC89B7BFF980414CB6AF8DB3EF4F8BC" box="[352,502,1840,1865]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.6" metricValueMin="4.7" pageId="4" pageNumber="5" unit="mm" value="5.65" valueMax="6.6" valueMin="4.7">4.7-6.6 mm</quantity>
, IZCAS, stream near Kuiyang Town, Yinlin County,
<collectingRegion id="49F4F87CFF980414C828F8BB3C00F89C" box="[546,770,1872,1898]" country="China" name="Guangxi" pageId="4" pageNumber="5">Guangxi Province</collectingRegion>
, coll. Y. Cai, H. Tang and H. Zhen,
<date id="FF8E105EFF980414C837F89B3DB1F87C" box="[573,691,1904,1929]" pageId="4" pageNumber="5" value="1993-11">Nov 1993</date>
.
</paragraph>
<paragraph id="8B8F369EFF980414CAE0F87B3A27FE03" blockId="4.[174,776,1232,1961]" lastBlockId="4.[850,1452,253,1974]" pageId="4" pageNumber="5">
<emphasis id="B944EA8CFF980414CAE0F87B3E80F85C" box="[234,386,1936,1961]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
: Rostrum long (
<figureCitation id="130B2A1BFF980414C86FF87B3DFAF85C" box="[613,760,1936,1961]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4A, B</figureCitation>
), straight, reaching slightly beyond distal end of scaphocerite, armed dorsally with 16-24 teeth, of which 5-8 are situated on carapace behind posterior margin of orbital margin, occupying about half carapace length, armed ventrally with 4-12 (mode 7-10) teeth. Orbit narrow; antennal spine sharp, completely fused with suborbital angle. Pterygostomian angle broadly rounded.
</paragraph>
<paragraph id="8B8F369EFF980414C985FE163CAAFD23" blockId="4.[850,1452,253,1974]" pageId="4" pageNumber="5">
Sixth abdominal somite twice as long as fifth abdominal somite, about half carapace length, slightly shorter than telson. Telson (
<figureCitation id="130B2A1BFF980414CF31FDD63AA0FDA2" box="[1339,1442,573,599]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5G</figureCitation>
) without posteromedian projection, posterior margin rounded, lateral pair of spines longer than sub-lateral pairs, subequal to intermediate pairs of spines.
</paragraph>
<paragraph id="8B8F369EFF980414C985FD363A1FFC23" blockId="4.[850,1452,253,1974]" pageId="4" pageNumber="5">
Eyes degenerated (
<figureCitation id="130B2A1BFF980414CEADFD363A35FD03" box="[1191,1335,733,758]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4A, B</figureCitation>
), small, pigment at centre of cornea. Antennular peduncle (
<figureCitation id="130B2A1BFF980414C954FCF63CC6FCC3" box="[862,964,797,822]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5A</figureCitation>
) 0.54 times as long as carapace. Antennular flagellum extraordinary long, more than twice total length. Stylocerite reaching 0.74 basal segment length of antennular peduncle or to end of segment. Scaphocerite (
<figureCitation id="130B2A1BFF980414CE9AFC763BEAFC43" box="[1168,1256,925,950]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5B</figureCitation>
) 3 times as long as wide, outer margin slightly concave.
</paragraph>
<paragraph id="8B8F369EFF980414C985FC363A2BFA23" blockId="4.[850,1452,253,1974]" pageId="4" pageNumber="5">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF980414CF48FC363AA6FC02" box="[1346,1444,989,1015]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4C</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF980414C9C4FBF63B28FBC3" box="[974,1066,1053,1078]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4D</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with dense setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF980414C9B0FB963B15FB63" box="[954,1047,1149,1174]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4E</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long hooked setae posteriorly. Palp of first maxilliped stout (
<figureCitation id="130B2A1BFF980414CE10FB363B76FB03" box="[1050,1140,1245,1270]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="4" pageNumber="5">Fig. 4F</figureCitation>
), terminating in projection. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF980414CEC3FB163A21FAE2" box="[1225,1315,1277,1303]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5C</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF980414C951FAD63CBDFAA3" box="[859,959,1341,1366]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5D</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior 0.25 penultimate segment length, ultimate segment shorter than penultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF980418C985FA363E52FE83" blockId="4.[850,1452,253,1974]" lastBlockId="8.[174,776,253,1463]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
First pereiopod (
<figureCitation id="130B2A1BFF980414CE44FA363BA7FA03" box="[1102,1189,1501,1526]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5E</figureCitation>
) short, stout, reaching slightly beyond end of basal segment of antennular peduncle, merus 2.5 times as long as broad; carpus strong concave anteriorly, 1.3 times as long as high, chela 1.7 times as long as broad, fingers slightly longer than palm. Second pereiopod (
<figureCitation id="130B2A1BFF980414C950F9763CB6F943" box="[858,948,1693,1718]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="4" pageNumber="5">Fig. 5F</figureCitation>
) reaching to end of antennular peduncle, carpus 2.6 times as long as high, chela slightly shorter than carpus, 2.6 times as long as broad, fingers 1.5 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF980414C950F8F63CD9F8C3" box="[858,987,1821,1846]" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="4" pageNumber="5">Fig. 6A, B</figureCitation>
) long, reaching slightly beyond distal end of scaphocerite, propodus 10 times as long as broad, 4.3 times as long as dactylus (claw included); dactylus ending in 2 claws, with 2-4 accessory spines on flexor margin. Fifth pereiopod (
<figureCitation id="130B2A1BFF940418CABCFF163E3BFEE3" box="[182,313,253,279]" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="8" pageNumber="9">Fig. 6C, D</figureCitation>
) reaching distal end of scaphocerite, propodus 13 times as long as wide, 4.9 times as long as dactylus; dactylus with 25 spinules on flexor margin.
</paragraph>
<footnote id="E82B2A90FF980414CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">
<paragraph id="8B8F369EFF980414CE96F8363AA9F81A" blockId="4.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF990415CA82F88C3D7CF846" ID-DOI="http://doi.org/10.5281/zenodo.13314999" ID-Zenodo-Dep="13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,170,1895,1915]" targetBox="[210,1335,267,1844]" targetPageId="5" targetType="figure">
<paragraph id="8B8F369EFF990415CA82F88C3D7CF846" blockId="5.[136,1414,1895,1971]" pageId="5" pageNumber="6">
<emphasis id="B944EA8CFF990415CA82F88C3FC5F88E" bold="true" box="[136,199,1895,1915]" pageId="5" pageNumber="6">Fig. 4.</emphasis>
<taxonomicName id="4C304D1DFF990415CADCF88C3E4AF88E" authorityName="Cai &amp; Ng" authorityYear="2018" box="[214,328,1895,1915]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alu" status="sp. nov.">
<emphasis id="B944EA8CFF990415CADCF88C3E4AF88E" box="[214,328,1895,1915]" italics="true" pageId="5" pageNumber="6">Caridina alu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF990415CB5AF88C3E9EF88E" box="[336,412,1895,1915]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029).
</paragraph>
</caption>
<footnote id="E82B2A90FF990415CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="5" pageNumber="6">
<paragraph id="8B8F369EFF990415CA82F8363E9AF81A" blockId="5.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="5" pageNumber="6">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF9A0416CAA4F88C3E23F846" ID-DOI="http://doi.org/10.5281/zenodo.13315001" ID-Zenodo-Dep="13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="6" pageNumber="7" startId="6.[174,208,1895,1915]" targetBox="[265,1379,257,1867]" targetPageId="6" targetType="figure">
<paragraph id="8B8F369EFF9A0416CAA4F88C3E23F846" blockId="6.[174,1453,1895,1971]" pageId="6" pageNumber="7">
<emphasis id="B944EA8CFF9A0416CAA4F88C3FEEF88E" bold="true" box="[174,236,1895,1915]" pageId="6" pageNumber="7">Fig. 5.</emphasis>
<taxonomicName id="4C304D1DFF9A0416CAF0F88C3E6EF88E" authorityName="Cai &amp; Ng" authorityYear="2018" box="[250,364,1895,1915]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alu" status="sp. nov.">
<emphasis id="B944EA8CFF9A0416CAF0F88C3E6EF88E" box="[250,364,1895,1915]" italics="true" pageId="6" pageNumber="7">Caridina alu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9A0416CB79F88C3EBDF88E" box="[371,447,1895,1915]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481).
</paragraph>
</caption>
<footnote id="E82B2A90FF9A0416CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="6" pageNumber="7">
<paragraph id="8B8F369EFF9A0416CE96F8363AA9F81A" blockId="6.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="6" pageNumber="7">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF9B0417CA82F8A53D2DF843" ID-DOI="http://doi.org/10.5281/zenodo.13315005" ID-Zenodo-Dep="13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,170,1870,1890]" targetBox="[196,1328,250,1830]" targetPageId="7" targetType="figure">
<paragraph id="8B8F369EFF9B0417CA82F8A53D2DF843" blockId="7.[136,1415,1870,1974]" pageId="7" pageNumber="8">
<emphasis id="B944EA8CFF9B0417CA82F8A53FC4F897" bold="true" box="[136,198,1870,1890]" pageId="7" pageNumber="8">Fig. 6.</emphasis>
<taxonomicName id="4C304D1DFF9B0417CADEF8A53E44F897" authorityName="Cai &amp; Ng" authorityYear="2018" box="[212,326,1870,1890]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alu" status="sp. nov.">
<emphasis id="B944EA8CFF9B0417CADEF8A53E44F897" box="[212,326,1870,1890]" italics="true" pageId="7" pageNumber="8">Caridina alu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9B0417CB44F8A53E9BF897" box="[334,409,1870,1890]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
(A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481).
</paragraph>
</caption>
<footnote id="E82B2A90FF9B0417CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="7" pageNumber="8">
<paragraph id="8B8F369EFF9B0417CA82F8363E9AF81A" blockId="7.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="7" pageNumber="8">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF940418CAE0FE963E2CFD43" blockId="8.[174,776,253,1463]" pageId="8" pageNumber="9">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF940418C891FE963FC4FE42" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="8" pageNumber="9">Fig. 6E, G</figureCitation>
) short, reaching to 0.67 endopod length, subrectangular, 3.3 times as long as wide, appendix interna situated at posterior 0.25 endopod length, extending to end of endopod. Appendix masculina of male second pleopod (
<figureCitation id="130B2A1BFF940418C8D3FDF63FF6FDA3" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="8" pageNumber="9">Fig. 6F, H</figureCitation>
) slightly curved outwards, reaching 0.75 endopod length; appendix interna of male second pleopod elongated, reaching to middle of appendix masculina.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0FD563E1AFD03" blockId="8.[174,776,253,1463]" pageId="8" pageNumber="9">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF940418C80DFD563D70FD23" box="[519,626,701,726]" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="8" pageNumber="9">Fig. 6H</figureCitation>
) with 9-11 spinules.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0FD163DE4FCE3" blockId="8.[174,776,253,1463]" box="[234,742,765,791]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0FD163E3CFCE3" box="[234,318,765,790]" italics="true" pageId="8" pageNumber="9">Habitat</emphasis>
: Subterranean river in a karst cave.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0FCF63E8DFC83" blockId="8.[174,776,253,1463]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0FCF63E69FCC3" box="[234,363,797,822]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
: The species is named after the
<typeStatus id="548B883CFF940418CAA4FCD63FE1FCA3" box="[174,227,829,854]" pageId="8" pageNumber="9">type</typeStatus>
locality,
<taxonomicName id="4C304D1DFF940418CB5FFCD63ED1FCA2" authority="Cave." authorityName="Cave." box="[341,467,829,855]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alu">Alu Cave.</taxonomicName>
The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0FC963E61FA83" blockId="8.[174,776,253,1463]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0FC963E58FC63" box="[234,346,893,918]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
:
<taxonomicName id="4C304D1DFF940418CB64FC963D0BFC63" authorityName="Cai &amp; Ng" authorityYear="2018" box="[366,521,893,918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alu" status="sp. nov.">
<emphasis id="B944EA8CFF940418CB64FC963D0BFC63" box="[366,521,893,918]" italics="true" pageId="8" pageNumber="9">Caridina alu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF940418C81FFC963D78FC63" box="[533,634,893,918]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C304D1DFF940418CAA4FC763DC6FC43" authority="Liang and Zhou, 1993" authorityName="Liang and Zhou" authorityYear="1993" box="[174,708,925,950]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sphyropoda">
<emphasis id="B944EA8CFF940418CAA4FC763EAAFC43" box="[174,424,925,950]" italics="true" pageId="8" pageNumber="9">Caridina sphyropoda</emphasis>
<bibRefCitation id="EFA14B6FFF940418CBB9FC763DC6FC43" author="Liang X &amp; Zhou J." box="[435,708,925,950]" pageId="8" pageNumber="9" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49F4F87CFF940418CAA4FC563E88FC23" box="[174,394,957,983]" country="China" name="Guangxi" pageId="8" pageNumber="9">Guangxi Province</collectingRegion>
in the form of its rostrum, but can be differentiated by the arrangement of its teeth: in
<taxonomicName id="4C304D1DFF940418CB2AFC163E69FBE3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[288,363,1021,1046]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alu">
<emphasis id="B944EA8CFF940418CB2AFC163E69FBE3" box="[288,363,1021,1046]" italics="true" pageId="8" pageNumber="9">C. alu</emphasis>
</taxonomicName>
, the postorbital teeth are widely placed, occupying half the length of the carapace (
<figureCitation id="130B2A1BFF940418CABCFBD63E43FBA3" box="[182,321,1085,1110]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, B</figureCitation>
) (vs. postorbital teeth are closely placed, occupying only a third of the carapace length; cf.
<bibRefCitation id="EFA14B6FFF940418CB3CFB963D50FB63" author="Liang X &amp; Zhou J." box="[310,594,1149,1174]" pageId="8" pageNumber="9" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 4(1)); the eyes are degenerated (
<figureCitation id="130B2A1BFF940418CBCFFB763D42FB43" box="[453,576,1181,1206]" captionStart="Fig" captionStartId="5.[136,170,1895,1915]" captionTargetBox="[210,1335,267,1844]" captionTargetId="figure-6@5.[210,1339,249,1866]" captionTargetPageId="5" captionText="Fig. 4. Caridina alu sp. nov. (A, B) cephalothorax and cephalic appendages, lateral view, (C) mandible, (D) maxillula, (E) maxilla, (F) first maxilliped. Scale bars: A, B = 2.0 mm; C, D = 0.5 mm; E, F = 1.0 mm; G, H = 0.2 mm. (A, C-F, female, cl 5.0 mm, paratype, ZRC 2018.0481; B, holotype, cl 6.5 mm, IZCAS DE 5029)." figureDoi="http://doi.org/10.5281/zenodo.13314999" httpUri="https://zenodo.org/record/13314999/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, B</figureCitation>
) (vs. developed; cf.
<bibRefCitation id="EFA14B6FFF940418CADFFB563EE5FB23" author="Liang X &amp; Zhou J." box="[213,487,1213,1238]" pageId="8" pageNumber="9" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 4(1)); the endopod is rectangular (
<figureCitation id="130B2A1BFF940418CB6AFB363EBEFB02" box="[352,444,1245,1271]" captionStart="Fig" captionStartId="7.[136,170,1870,1890]" captionTargetBox="[196,1328,250,1830]" captionTargetId="figure-88@7.[196,1353,216,1852]" captionTargetPageId="7" captionText="Fig. 6. Caridina alu sp. nov. (A) third pereiopod, (B) dactylus of third pereiopod, (C) fifth pereiopod, (D) dactylus of fifth pereiopod, (E, G) endopod of male first pleopod, (F, H) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm; B, D = 0.2 mm; E-H = 0.5 mm. (A-D, female, cl 5.0 paratype, ZRC 2018.0481; E, F, male, cl 5.8 mm, holotype, IZCAS DE 5029; G, H, male, cl 6.3 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315005" httpUri="https://zenodo.org/record/13315005/files/figure.png" pageId="8" pageNumber="9">Fig. 6G</figureCitation>
) (vs. sigmoid; cf.
<bibRefCitation id="EFA14B6FFF940418C89AFB363E46FAE3" author="Liang X &amp; Zhou J." pageId="8" pageNumber="9" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 4(8)), and the telson does not possess a posteromedian projection (
<figureCitation id="130B2A1BFF940418C865FAF63DCEFAC2" box="[623,716,1309,1335]" captionStart="Fig" captionStartId="6.[174,208,1895,1915]" captionTargetBox="[265,1379,257,1867]" captionTargetId="figure-52@6.[209,1420,249,1869]" captionTargetPageId="6" captionText="Fig. 5. Caridina alu sp. nov. (A) antennular peduncle, (B) scaphocerite, (C) second maxilliped, (D) third maxilliped, (E) first pereiopod, (F) second pereiopod, (G) distal portion of telsom. Scale bars: A-F = 1.0 mm; G, H = 0.3 mm. (female, cl 5.0 mm, paratype, ZRC 2018.0481)." figureDoi="http://doi.org/10.5281/zenodo.13315001" httpUri="https://zenodo.org/record/13315001/files/figure.png" pageId="8" pageNumber="9">Fig. 5G</figureCitation>
) (vs. projection distinctly present; cf.
<bibRefCitation id="EFA14B6FFF940418C831FAD63FECFA83" author="Liang X &amp; Zhou J." pageId="8" pageNumber="9" pagination="231 - 239" refId="ref16064" refString="Liang X, Zhou J. 1993. Study on new atyid shrimps (Decapoda, Caridea) from Guangxi, China. Acta Hydrobiol Sinica 17 (3): 231 - 239, figures 1 - 4. (in Chinese with English abstract)" type="journal article" year="1993">Liang and Zhou, 1993</bibRefCitation>
: fig. 4(2)).
</paragraph>
<paragraph id="8B8F369EFF940418CAE0FA963FFEFA42" blockId="8.[174,776,253,1463]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0FA963E79FA63" box="[234,379,1405,1430]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF940418CB85FA963D6BFA63" box="[399,617,1405,1430]" country="China" name="Yunnan" pageId="8" pageNumber="9">Yunnan Province</collectingRegion>
(
<taxonomicName id="4C304D1DFF940418C876FA963DF9FA62" authority="Cave" authorityName="Cave" box="[636,763,1405,1431]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alu">Alu Cave</taxonomicName>
),
<collectingCountry id="F327760EFF940418CAA4FA763FFAFA42" box="[174,248,1437,1463]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,378 @@
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<mods:title id="7AD7763E6F6B20AD7C28542BC94A0662">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<heading id="D0C781F2FF9D0411CB01FD773D5CFD43" bold="true" box="[267,606,668,694]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
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<emphasis id="B944EA8CFF9D0411CB01FD773EF6FD43" bold="true" box="[267,500,668,694]" italics="true" pageId="1" pageNumber="2">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9D0411CBF1FD763D5CFD43" box="[507,606,669,694]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8F369EFF9D0411CB72FD563EF3FD23" blockId="1.[151,722,668,779]" box="[376,497,701,726]" pageId="1" pageNumber="2">
(
<figureCitation id="130B2A1BFF9D0411CB75FD563EEAFD23" box="[383,488,701,726]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[136,170,1813,1833]" captionStartId-1="2.[176,211,1841,1861]" captionStartId-2="3.[136,171,1869,1889]" captionTargetBox-0="[136,1414,945,1784]" captionTargetBox-1="[169,1452,916,1812]" captionTargetBox-2="[141,1402,975,1815]" captionTargetId-0="figure-6@1.[125,1428,930,1787]" captionTargetId-1="figure-372@2.[156,1477,907,1826]" captionTargetId-2="figure-6@3.[136,1413,967,1843]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." captionText-1="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." captionText-2="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi-0="http://doi.org/10.5281/zenodo.13314993" figureDoi-1="http://doi.org/10.5281/zenodo.13314995" figureDoi-2="http://doi.org/10.5281/zenodo.13314997" httpUri-0="https://zenodo.org/record/13314993/files/figure.png" httpUri-1="https://zenodo.org/record/13314995/files/figure.png" httpUri-2="https://zenodo.org/record/13314997/files/figure.png" pageId="1" pageNumber="2">Figs. 1-3</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF9D0411CA9DFD333D05FCFE" blockId="1.[151,722,668,779]" pageId="1" pageNumber="2">
<uri id="FFA13A9CFF9D0411CA9DFD333D05FCFE" pageId="1" pageNumber="2">
urn:lsid:zoobank.org:act:
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<paragraph id="8B8F369EFF9D0411CACFFCDB3DDDFCBC" blockId="1.[136,738,816,874]" box="[197,735,816,841]" pageId="1" pageNumber="2">
<emphasis id="B944EA8CFF9D0411CACFFCDB3EC5FCBC" box="[197,455,816,841]" italics="true" pageId="1" pageNumber="2">M a t e r i a l e x a m i n e d</emphasis>
: H o l o t y p e: m a l e, c l
</paragraph>
<paragraph id="8B8F369EFF9D0411CA82FCBB3B2DFD83" blockId="1.[136,738,816,874]" lastBlockId="1.[813,1415,253,886]" pageId="1" pageNumber="2">
8.0 mm, IZCAS DE 5028, stream at end of cave, Panon Cave, Huaoyan Village,
<taxonomicName id="4C304D1DFF9D0411CEA1FF163A80FEE2" authority="County" authorityName="County" box="[1195,1410,253,279]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
,
<collectingRegion id="49F4F87CFF9D0411C927FEF63CFAFEC3" box="[813,1016,285,310]" country="China" name="Hunan" pageId="1" pageNumber="2">Hunan Province</collectingRegion>
southern
<collectingCountry id="F327760EFF9D0411CE77FEF63BC9FEC2" box="[1149,1227,285,311]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
, coll. Cao and B. Sket,
<date id="FF8E105EFF9D0411C99DFED63B33FEA3" box="[919,1073,317,342]" pageId="1" pageNumber="2" value="1997-04-13">13 Apr 1997</date>
.
<typeStatus id="548B883CFF9D0411CE34FED63BB9FEA3" box="[1086,1211,317,342]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF9D0411CEC2FED63A2FFEA3" box="[1224,1325,317,342]" count="2" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
, cl 6.6- 9.0 mm;
<specimenCount id="9D36FD17FF9D0411C9AAFEB63B11FE83" box="[928,1043,349,374]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411CE48FEB63BA4FE83" box="[1090,1190,349,374]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="2" unit="mm" value="6.5">6.5 mm</quantity>
, ZRC 2018.0480, same data as
<typeStatus id="548B883CFF9D0411C9D7FE963B45FE63" box="[989,1095,381,406]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF9D0411CE5FFE963BC1FE63" box="[1109,1219,381,406]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, cl 7.0 mm, UL, same data as
<typeStatus id="548B883CFF9D0411C9E7FE763B63FE43" box="[1005,1121,413,438]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D36FD17FF9D0411CE7BFE763BEBFE43" box="[1137,1257,413,438]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411CF10FE763A83FE43" box="[1306,1409,413,438]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="2" unit="mm" value="6.8">6.8 mm</quantity>
, RMNH.CRUS.D.57289, Cave of Flying Tiger, Hoyan Village,
<taxonomicName id="4C304D1DFF9D0411C9E3FE363BC6FE02" authority="County" authorityName="County" box="[1001,1220,477,503]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
,
<collectingRegion id="49F4F87CFF9D0411CEDBFE363A27FE03" box="[1233,1317,477,502]" country="China" name="Hunan" pageId="1" pageNumber="2">Hunan</collectingRegion>
, coll. L. Deharveng and A. Bedos,
<date id="FF8E105EFF9D0411CE63FE163A07FDE3" box="[1129,1285,509,534]" pageId="1" pageNumber="2" value="1995-08-14">14 Aug 1995</date>
;
<specimenCount id="9D36FD17FF9D0411CF1BFE163A64FDE3" box="[1297,1382,509,534]" count="1" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411C927FDF63C88FDC3" box="[813,906,541,566]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="2" unit="mm" value="8.8">8.8 mm</quantity>
, ZRC 2018.0489,
<specimenCount id="9D36FD17FF9D0411CE68FDF63BCEFDC3" box="[1122,1228,541,566]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411CEF9FDF63A52FDC3" box="[1267,1360,541,566]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="2" unit="mm" value="7.4">7.4 mm</quantity>
, UL, Parking Cave, near Hoyan Village, coll. Canto and Delprat,
<date id="FF8E105EFF9D0411C99BFDB63B2EFD83" box="[913,1068,605,630]" pageId="1" pageNumber="2" value="1995-08-25">25 Aug 1995</date>
.
</paragraph>
<paragraph id="8B8F369EFF9D0411C963FD963CA8FC83" blockId="1.[813,1415,253,886]" pageId="1" pageNumber="2">
<emphasis id="B944EA8CFF9D0411C963FD963A07FD63" box="[873,1285,637,662]" italics="true" pageId="1" pageNumber="2">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF9D0411CF10FD963BF8FD43" authority="Bouvier, 1919" authorityName="Bouvier" authorityYear="1919" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="cavaleriei" subSpecies="cavaleriei">
<emphasis id="B944EA8CFF9D0411CF10FD963B37FD43" italics="true" pageId="1" pageNumber="2">Caridina cavaleriei cavaleriei</emphasis>
<bibRefCitation id="EFA14B6FFF9D0411CE48FD763BF8FD43" author="Bouvier EL" box="[1090,1274,669,694]" pageId="1" pageNumber="2" pagination="330 - 336" refId="ref14415" refString="Bouvier EL. 1919. Quelques especes nouvelles de Caridines, Bull Musee nat Hist Natur 1919: 330 - 336." type="journal article" year="1919">Bouvier, 1919</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF9D0411CF16FD763A86FD42" box="[1308,1412,669,695]" pageId="1" pageNumber="2" type="syntype">Syntype</typeStatus>
<specimenCount id="9D36FD17FF9D0411C927FD563C98FD23" box="[813,922,701,726]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, MNHN Na 689, Gan-chouen-fou, Kouy- Tcheou (
<collectingRegion id="49F4F87CFF9D0411C99FFD363B01FD02" box="[917,1027,733,759]" country="China" name="Guizhou" pageId="1" pageNumber="2">Guizhou</collectingRegion>
),
<collectingCountry id="F327760EFF9D0411CE1DFD363B60FD02" box="[1047,1122,733,759]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
.
<taxonomicName id="4C304D1DFF9D0411CE65FD363CFFFCE3" authority="NK Ng and Cai, 2000" authorityName="NK Ng and Cai" authorityYear="2000" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dentifrons">
<emphasis id="B944EA8CFF9D0411CE65FD363A54FD03" box="[1135,1366,733,758]" italics="true" pageId="1" pageNumber="2">Caridina dentifrons</emphasis>
NK
<bibRefCitation id="EFA14B6FFF9D0411C927FD163CFFFCE3" author="Ng NK &amp; Cai Y." box="[813,1021,765,791]" pageId="1" pageNumber="2" pagination="167 - 175" refId="ref16203" refString="Ng NK, Cai Y. 2000. Two new species of atyid shrimps from southern China (Crustacea: Decapoda: Caridea). Raffles Bull Zool 48 (1): 167 - 175, figures 1 - 6." type="journal article" year="2000">Ng and Cai, 2000</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF9D0411CE1CFD163B90FCE3" box="[1046,1170,765,790]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD17FF9D0411CE97FD163BFCFCE3" box="[1181,1278,765,790]" count="2" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411CF2FFD163A80FCE3" box="[1317,1410,765,790]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="2" unit="mm" value="4.7">4.7 mm</quantity>
,
<specimenCount id="9D36FD17FF9D0411C927FCF63C95FCC3" box="[813,919,797,822]" count="1" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF9D0411C9B4FCF63B1EFCC3" box="[958,1052,797,822]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="2" unit="mm" value="5.2">5.2 mm</quantity>
, ZRC 1997.593, Baijin Village, Huishui County,
<collectingRegion id="49F4F87CFF9D0411C9F9FCD63BC9FCA3" box="[1011,1227,829,855]" country="China" name="Guizhou" pageId="1" pageNumber="2">Guizhou Province</collectingRegion>
,
<collectingCountry id="F327760EFF9D0411CED2FCD63A20FCA2" box="[1240,1314,829,855]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
, coll.
<date id="FF8E105EFF9D0411CF62FCD63CA4FC83" pageId="1" pageNumber="2" value="1983-05-16">16 May 1983</date>
.
</paragraph>
<caption id="DF4F6616FF9D0411CA82F8FE3E68F840" ID-DOI="http://doi.org/10.5281/zenodo.13314993" ID-Zenodo-Dep="13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="1" pageNumber="2" startId="1.[136,170,1813,1833]" targetBox="[136,1414,945,1784]" targetPageId="1" targetType="figure">
<paragraph id="8B8F369EFF9D0411CA82F8FE3E68F840" blockId="1.[136,1415,1813,1973]" pageId="1" pageNumber="2">
<emphasis id="B944EA8CFF9D0411CA82F8FE3FC5F8DC" bold="true" box="[136,199,1813,1833]" pageId="1" pageNumber="2">Fig. 1.</emphasis>
<taxonomicName id="4C304D1DFF9D0411CADFF8FE3E81F8DC" authorityName="Cai &amp; Ng" authorityYear="2018" box="[213,387,1813,1833]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF9D0411CADFF8FE3E81F8DC" box="[213,387,1813,1833]" italics="true" pageId="1" pageNumber="2">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9D0411CB83F8FE3ED7F8DC" box="[393,469,1813,1833]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
(A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village,
<taxonomicName id="4C304D1DFF9D0411CB41F8A63EF4F894" authority="County" authorityName="County" box="[331,502,1869,1889]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village,
<taxonomicName id="4C304D1DFF9D0411CF22F8A63FCDF888" authority="County" authorityName="County" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village,
<taxonomicName id="4C304D1DFF9D0411CE23F8823BDAF888" authority="County" authorityName="County" box="[1065,1240,1897,1917]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village,
<taxonomicName id="4C304D1DFF9D0411C9C6F86E3B7AF86C" authority="County" authorityName="County" box="[972,1144,1925,1945]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, Hunan Province, paratype, RMNHCRUS.D.57289).
</paragraph>
</caption>
<footnote id="E82B2A90FF9D0411CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="1" pageNumber="2">
<paragraph id="8B8F369EFF9D0411CA82F8363E9AF81A" blockId="1.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="1" pageNumber="2">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF9E0412CAE0FF163E75FDA3" blockId="2.[174,775,253,854]" pageId="2" pageNumber="3">
<emphasis id="B944EA8CFF9E0412CAE0FF163E87FEE3" box="[234,389,253,278]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
: Rostrum long (
<figureCitation id="130B2A1BFF9E0412C866FF163DBBFEE3" box="[620,697,253,278]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A-D), reaching from end of antennular peduncle to end of scaphocerite, anterior half distinctly upturned, armed dorsally with 18-21 slender teeth, of which 6 or 7 teeth situated on carapace behind posterior margin of orbit, occupied 0.25-0.33 carapace length, armed ventrally with 12-17 teeth, of which posterior 4 or 5 teeth are wider separated than others. Antennal spine sharp, completely fused with suborbital angle; pterygostomian margin broadly rounded.
</paragraph>
<paragraph id="8B8F369EFF9E0412CAE0FDB63E85FCC3" blockId="2.[174,775,253,854]" pageId="2" pageNumber="3">
Sixth abdominal somite 1.8 times as long as fifth abdominal somite, as long as telson. Telson (
<figureCitation id="130B2A1BFF9E0412CABCFD763E38FD43" box="[182,314,669,694]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2K, L</figureCitation>
) without posteriomedian projection, lateral pair of spines on posterior margin longer than intermediate pairs, surface of proximal half covered by fine setae and spinules. Sublateral pair of spines shortest.
</paragraph>
<paragraph id="8B8F369EFF9E0412CAE0FCD63A97FE43" blockId="2.[174,775,253,854]" lastBlockId="2.[850,1452,253,854]" pageId="2" pageNumber="3">
Eyes degenerated (
<figureCitation id="130B2A1BFF9E0412CBE5FCD63D3AFCA3" box="[495,568,829,854]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A-D), eye stalk absent, cornea pigmentation variable, from totally absent to with small black spot. Antennular peduncle slender (
<figureCitation id="130B2A1BFF9E0412CE31FED63B9BFEA3" box="[1083,1177,317,342]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="2" pageNumber="3">Fig. 1E</figureCitation>
), about half carapace length. Stylocerite sharp, long, reaching near end of basal segment of antennular peduncle. Scaphocerite 2.8 times as long as wide (
<figureCitation id="130B2A1BFF9E0412CF3AFE763A84FE43" box="[1328,1414,413,438]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="2" pageNumber="3">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8B8F369EFF9E0413C985FE563D7CFEE3" blockId="2.[850,1452,253,854]" lastBlockId="3.[136,738,253,918]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Mouthparts as figured. Mandible with blunt teeth at extremity of incisor process (
<figureCitation id="130B2A1BFF9E0412CF30FE363A9DFE03" box="[1338,1439,477,502]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
). Maxillula with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with dense setae and teeth (
<figureCitation id="130B2A1BFF9E0412CF4CFDD63AA2FDA3" box="[1350,1440,573,598]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2B</figureCitation>
). Maxilla with slender palp, upper endite subdivided, scaphognathite tapering with numerous long hooked setae posteriorly (
<figureCitation id="130B2A1BFF9E0412CE97FD763A03FD42" box="[1181,1281,669,695]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2C</figureCitation>
). Palp of first maxilliped stout, terminating in sub-acute angle (
<figureCitation id="130B2A1BFF9E0412C950FD363CB7FD03" box="[858,949,733,758]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2D</figureCitation>
). Endopod of second maxilliped with fused dactylus and propodal segments (
<figureCitation id="130B2A1BFF9E0412CEFDFD163A55FCE3" box="[1271,1367,765,790]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3">Fig. 2E</figureCitation>
). Third maxilliped ending in single terminal claw; exopod reaching posterior quarter of penultimate segment (
<figureCitation id="130B2A1BFF9F0413CA9AFF163E0BFEE2" box="[144,265,253,279]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="3" pageNumber="4">Fig. 2F, G</figureCitation>
). Epipods on first 4 pereiopods.
</paragraph>
<caption id="DF4F6616FF9E0412CABAF8DA3BF6F840" ID-DOI="http://doi.org/10.5281/zenodo.13314995" ID-Zenodo-Dep="13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="2" pageNumber="3" startId="2.[176,211,1841,1861]" targetBox="[169,1452,916,1812]" targetPageId="2" targetType="figure">
<paragraph id="8B8F369EFF9E0412CABAF8DA3BF6F840" blockId="2.[174,1453,1841,1973]" pageId="2" pageNumber="3">
<emphasis id="B944EA8CFF9E0412CABAF8DA3FF0F8B0" bold="true" box="[176,242,1841,1861]" pageId="2" pageNumber="3">Fig. 2.</emphasis>
<taxonomicName id="4C304D1DFF9E0412CB0BF8DA3EB5F8B0" authorityName="Cai &amp; Ng" authorityYear="2018" box="[257,439,1841,1861]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF9E0412CB0BF8DA3EB5F8B0" box="[257,439,1841,1861]" italics="true" pageId="2" pageNumber="3">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9E0412CBB5F8DA3D0DF8B0" box="[447,527,1841,1861]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village,
<taxonomicName id="4C304D1DFF9E0412C9BEF86E3B61F86C" authority="County" authorityName="County" box="[948,1123,1925,1945]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village,
<taxonomicName id="4C304D1DFF9E0412C934F84A3CEAF840" authority="County" authorityName="County" box="[830,1000,1953,1973]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, paratype, ZRC 2018.0489).
</paragraph>
</caption>
<footnote id="E82B2A90FF9E0412CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">
<paragraph id="8B8F369EFF9E0412CE96F8363AA9F81A" blockId="2.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF9F0413CACFFEF63B5FFEE3" blockId="3.[136,738,253,918]" lastBlockId="3.[813,1414,253,918]" pageId="3" pageNumber="4">
First pereiopod very stout (
<figureCitation id="130B2A1BFF9F0413C803FEF63D64FEC3" box="[521,614,285,310]" captionStart="Fig" captionStartId="3.[136,171,1869,1889]" captionTargetBox="[141,1402,975,1815]" captionTargetId="figure-6@3.[136,1413,967,1843]" captionTargetPageId="3" captionText="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi="http://doi.org/10.5281/zenodo.13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
), reaching to middle of basal segment of antennular peduncle; merus 3.0 times as long as wide, carpus 1.9 times as long as high; chela 2.7 times as long as broad, fingers 1.2 times as long as palm. Second pereiopod (
<figureCitation id="130B2A1BFF9F0413CB18FE563E6EFE23" box="[274,364,445,470]" captionStart="Fig" captionStartId="3.[136,171,1869,1889]" captionTargetBox="[141,1402,975,1815]" captionTargetId="figure-6@3.[136,1413,967,1843]" captionTargetPageId="3" captionText="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi="http://doi.org/10.5281/zenodo.13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
) reaching to middle of second segment of antennular peduncle; merus as long as carpus; carpus longer than chela, 5.8 times as long as high; chela 3.1 times as long as broad; fingers 1.5 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF9F0413CA9AFDB63E1AFD83" box="[144,280,605,631]" captionStart="Fig" captionStartId="3.[136,171,1869,1889]" captionTargetBox="[141,1402,975,1815]" captionTargetId="figure-6@3.[136,1413,967,1843]" captionTargetPageId="3" captionText="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi="http://doi.org/10.5281/zenodo.13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="3" pageNumber="4">Fig. 3C, D</figureCitation>
) reaching with dactylus beyond the distal end of scaphocerite, propodus 12 times as long as wide, 4.3 times as long as dactylus (claw included); dactylus ending in 2 claws, with 7 or 8 accessory spines increasing in length and size distally on flexor margin. Fifth pereiopod (
<figureCitation id="130B2A1BFF9F0413CA9AFCF63E38FCC2" box="[144,314,797,823]" captionStart="Fig" captionStartId="3.[136,171,1869,1889]" captionTargetBox="[141,1402,975,1815]" captionTargetId="figure-6@3.[136,1413,967,1843]" captionTargetPageId="3" captionText="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi="http://doi.org/10.5281/zenodo.13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="3" pageNumber="4">Fig. 3E, F, G</figureCitation>
) reaching slightly beyond end of antennular peduncle but not beyond distal end of scaphocerite, propodus 15 times as long as wide, 3.2 times as long as dactylus; dactylus with 42-49 spinules on flexor margin.
</paragraph>
<paragraph id="8B8F369EFF9F0413C963FEF63B30FDE3" blockId="3.[813,1414,253,918]" pageId="3" pageNumber="4">
Endopod of male first pleopod sub-elliptical (
<figureCitation id="130B2A1BFF9F0413C93EFED63CAFFEA3" box="[820,941,317,342]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="3" pageNumber="4">Fig. 2H, I</figureCitation>
), inner margin straight, reaching 0.4 exopod length, distal half curved posteriorly, 2.2-2.5 times as long as wide, appendix interna reaching to distal end of endopod. Appendix masculina of male second pleopod short (
<figureCitation id="130B2A1BFF9F0413CEAAFE563BFDFE23" box="[1184,1279,445,470]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="3" pageNumber="4">Fig. 2J</figureCitation>
), reaching to middle of endopod; appendix interna of male second pleopod stout.
</paragraph>
<paragraph id="8B8F369EFF9F0413C963FDF63C60FDA3" blockId="3.[813,1414,253,918]" pageId="3" pageNumber="4">
Uropodal diaeresis with 11 or 12 spinules (
<figureCitation id="130B2A1BFF9F0413CF52FDF63C55FDA3" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="3" pageNumber="4">Fig. 2M</figureCitation>
).
</paragraph>
<paragraph id="8B8F369EFF9F0413C963FDB63BD7FD83" blockId="3.[813,1414,253,918]" box="[873,1237,605,631]" pageId="3" pageNumber="4">
<emphasis id="B944EA8CFF9F0413C963FDB63CBFFD83" box="[873,957,605,630]" italics="true" pageId="3" pageNumber="4">Habitat</emphasis>
: Stream in karst caves.
</paragraph>
<paragraph id="8B8F369EFF9F0413C963FD963A21FD23" blockId="3.[813,1414,253,918]" pageId="3" pageNumber="4">
<emphasis id="B944EA8CFF9F0413C963FD963CEDFD63" box="[873,1007,637,662]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
:
<taxonomicName id="4C304D1DFF9F0413CE0FFD963BF0FD63" authorityName="Cai &amp; Ng" authorityYear="2018" box="[1029,1266,637,662]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF9F0413CE0FFD963BF0FD63" box="[1029,1266,637,662]" italics="true" pageId="3" pageNumber="4">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9F0413CEF4FD963A67FD63" box="[1278,1381,637,662]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is named after its
<typeStatus id="548B883CFF9F0413C9E5FD763B20FD43" box="[1007,1058,669,694]" pageId="3" pageNumber="4">type</typeStatus>
locality in
<taxonomicName id="4C304D1DFF9F0413CEA3FD763A84FD42" authority="County." authorityName="County." box="[1193,1414,669,695]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="longshan">Longshan County.</taxonomicName>
The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF9F0414C963FD363D0FFD23" blockId="3.[813,1414,253,918]" lastBlockId="4.[174,775,253,1047]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B944EA8CFF9F0413C963FD363CDCFD03" box="[873,990,733,758]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
:
<taxonomicName id="4C304D1DFF9F0413C9FFFD363BE9FD03" authorityName="Cai &amp; Ng" authorityYear="2018" box="[1013,1259,733,758]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF9F0413C9FFFD363BE9FD03" box="[1013,1259,733,758]" italics="true" pageId="3" pageNumber="4">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9F0413CEF3FD363A66FD03" box="[1273,1380,733,758]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically similar to
<taxonomicName id="4C304D1DFF9F0413CE65FD163CE4FCC3" authority="Bouvier, 1919" authorityName="Bouvier" authorityYear="1919" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="cavaleriei" subSpecies="cavaleriei">
<emphasis id="B944EA8CFF9F0413CE65FD163A87FCE3" box="[1135,1413,765,790]" italics="true" pageId="3" pageNumber="4">C. cavaleriei cavaleriei</emphasis>
<bibRefCitation id="EFA14B6FFF9F0413C927FCF63CE4FCC3" author="Bouvier EL" box="[813,998,797,822]" pageId="3" pageNumber="4" pagination="330 - 336" refId="ref14415" refString="Bouvier EL. 1919. Quelques especes nouvelles de Caridines, Bull Musee nat Hist Natur 1919: 330 - 336." type="journal article" year="1919">Bouvier, 1919</bibRefCitation>
</taxonomicName>
, a subspecies known only from the original specimens from
<collectingRegion id="49F4F87CFF9F0413CE96FCD63A80FCA3" box="[1180,1410,829,855]" country="China" name="Guizhou" pageId="3" pageNumber="4">Guizhou Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF9F0413C9A3FCB63CFBFC82" box="[937,1017,861,887]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
. It can be distinguished by its strongly upturned rostrum (
<figureCitation id="130B2A1BFF9F0413CE78FC963BB0FC63" box="[1138,1202,893,918]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
A-D) (vs. straight in
<taxonomicName id="4C304D1DFF980414CAC6FF163ED8FEE3" authorityName="Bouvier" authorityYear="1919" box="[204,474,253,278]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="cavaleriei" subSpecies="cavaleriei">
<emphasis id="B944EA8CFF980414CAC6FF163ED8FEE3" box="[204,474,253,278]" italics="true" pageId="4" pageNumber="5">C. cavaleriei cavaleriei</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="EFA14B6FFF980414C805FF163DBAFEE3" author="Bouvier EL" box="[527,696,253,278]" pageId="4" pageNumber="5" pagination="1 - 370" refId="ref14437" refString="Bouvier EL. 1925. Recherches sur la morphologie, la geographique des crevettes des la famille des Atyides, Ency Entomol 4 (A): 1 - 370, figs. 1 - 761." type="journal article" year="1925">Bouvier, 1925</bibRefCitation>
: figure 544) and degenerated eyes (
<figureCitation id="130B2A1BFF980414C80EFEF63D40FEC3" box="[516,578,285,310]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
A-D) (vs. clearly developed in
<taxonomicName id="4C304D1DFF980414CB5FFED63D69FEA3" authorityName="Bouvier" authorityYear="1919" box="[341,619,317,342]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="cavaleriei" subSpecies="cavaleriei">
<emphasis id="B944EA8CFF980414CB5FFED63D69FEA3" box="[341,619,317,342]" italics="true" pageId="4" pageNumber="5">C. cavaleriei cavaleriei</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="EFA14B6FFF980414C8AEFED63FF7FE83" author="Bouvier EL" pageId="4" pageNumber="5" pagination="1 - 370" refId="ref14437" refString="Bouvier EL. 1925. Recherches sur la morphologie, la geographique des crevettes des la famille des Atyides, Ency Entomol 4 (A): 1 - 370, figs. 1 - 761." type="journal article" year="1925">Bouvier, 1925</bibRefCitation>
: fig. 544). With the long and upturned rostrum,
<taxonomicName id="4C304D1DFF980414CB14FE963D02FE63" authorityName="Cai &amp; Ng" authorityYear="2018" box="[286,512,381,406]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF980414CB14FE963D02FE63" box="[286,512,381,406]" italics="true" pageId="4" pageNumber="5">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF980414C801FE963D6FFE63" box="[523,621,381,406]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
morphology resembles that of
<taxonomicName id="4C304D1DFF980414CB99FE763FECFE23" authority="NK Ng and Cai, 2000" authorityName="NK Ng and Cai" authorityYear="2000" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dentifrons">
<emphasis id="B944EA8CFF980414CB99FE763D36FE43" box="[403,564,413,438]" italics="true" pageId="4" pageNumber="5">C. dentifrons</emphasis>
NK
<bibRefCitation id="EFA14B6FFF980414C865FE763FECFE23" author="Ng NK &amp; Cai Y." pageId="4" pageNumber="5" pagination="167 - 175" refId="ref16203" refString="Ng NK, Cai Y. 2000. Two new species of atyid shrimps from southern China (Crustacea: Decapoda: Caridea). Raffles Bull Zool 48 (1): 167 - 175, figures 1 - 6." type="journal article" year="2000">Ng and Cai, 2000</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49F4F87CFF980414CB36FE563EA7FE22" box="[316,421,445,471]" country="China" name="Guizhou" pageId="4" pageNumber="5">Guizhou</collectingRegion>
. It could be easily separated by its larger number of ventral rostral teeth (12- 17 vs.
<quantity id="4CC89B7BFF980414CB0DFE163E53FDE3" box="[263,337,509,534]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="2.032" metricValueMin="1.016" pageId="4" pageNumber="5" unit="in" value="6.0" valueMax="8.0" valueMin="4.0">4-8 in</quantity>
<taxonomicName id="4C304D1DFF980414CB56FE163EFCFDE3" authorityName="NK Ng and Cai" authorityYear="2000" box="[348,510,509,534]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dentifrons">
<emphasis id="B944EA8CFF980414CB56FE163EFCFDE3" box="[348,510,509,534]" italics="true" pageId="4" pageNumber="5">C. dentifrons</emphasis>
</taxonomicName>
; cf. NK
<bibRefCitation id="EFA14B6FFF980414C867FE163FF2FDC3" author="Ng NK &amp; Cai Y." pageId="4" pageNumber="5" pagination="167 - 175" refId="ref16203" refString="Ng NK, Cai Y. 2000. Two new species of atyid shrimps from southern China (Crustacea: Decapoda: Caridea). Raffles Bull Zool 48 (1): 167 - 175, figures 1 - 6." type="journal article" year="2000">Ng and Cai, 2000</bibRefCitation>
: fig. 1A-C), degenerated eyes (
<figureCitation id="130B2A1BFF980414C888FDF63DC4FDC3" box="[642,710,541,566]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
A-D) (vs. clearly developed in
<taxonomicName id="4C304D1DFF980414CBEBFDD63D7FFDA3" authorityName="NK Ng and Cai" authorityYear="2000" box="[481,637,573,598]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dentifrons">
<emphasis id="B944EA8CFF980414CBEBFDD63D7FFDA3" box="[481,637,573,598]" italics="true" pageId="4" pageNumber="5">C. dentifrons</emphasis>
</taxonomicName>
; cf. NK
<bibRefCitation id="EFA14B6FFF980414C8EFFDD63E6BFD83" author="Ng NK &amp; Cai Y." pageId="4" pageNumber="5" pagination="167 - 175" refId="ref16203" refString="Ng NK, Cai Y. 2000. Two new species of atyid shrimps from southern China (Crustacea: Decapoda: Caridea). Raffles Bull Zool 48 (1): 167 - 175, figures 1 - 6." type="journal article" year="2000">Ng and Cai, 2000</bibRefCitation>
: fig. 1A-C) and dactylus of third pereiopod with 7 or 8 accessory spines on flexor margin (
<figureCitation id="130B2A1BFF980414CB1BFD763E6EFD43" box="[273,364,669,694]" captionStart="Fig" captionStartId="3.[136,171,1869,1889]" captionTargetBox="[141,1402,975,1815]" captionTargetId="figure-6@3.[136,1413,967,1843]" captionTargetPageId="3" captionText="Fig. 3. Caridina longshan sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location)." figureDoi="http://doi.org/10.5281/zenodo.13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="4" pageNumber="5">Fig. 3D</figureCitation>
) (vs. 6 spines in
<taxonomicName id="4C304D1DFF980414C84AFD763DDBFD43" authorityName="NK Ng and Cai" authorityYear="2000" box="[576,729,669,694]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dentifrons">
<emphasis id="B944EA8CFF980414C84AFD763DDBFD43" box="[576,729,669,694]" italics="true" pageId="4" pageNumber="5">C. dentifrons</emphasis>
</taxonomicName>
; cf. NK
<bibRefCitation id="EFA14B6FFF980414CAD0FD563EADFD23" author="Ng NK &amp; Cai Y." box="[218,431,701,727]" pageId="4" pageNumber="5" pagination="167 - 175" refId="ref16203" refString="Ng NK, Cai Y. 2000. Two new species of atyid shrimps from southern China (Crustacea: Decapoda: Caridea). Raffles Bull Zool 48 (1): 167 - 175, figures 1 - 6." type="journal article" year="2000">Ng and Cai, 2000</bibRefCitation>
: fig. 3E)
</paragraph>
<caption id="DF4F6616FF9F0413CA82F8A63DE2F840" ID-DOI="http://doi.org/10.5281/zenodo.13314997" ID-Zenodo-Dep="13314997" httpUri="https://zenodo.org/record/13314997/files/figure.png" pageId="3" pageNumber="4" startId="3.[136,171,1869,1889]" targetBox="[141,1402,975,1815]" targetPageId="3" targetType="figure">
<paragraph id="8B8F369EFF9F0413CA82F8A63DE2F840" blockId="3.[136,1415,1869,1973]" pageId="3" pageNumber="4">
<emphasis id="B944EA8CFF9F0413CA82F8A63FCBF894" bold="true" box="[136,201,1869,1889]" pageId="3" pageNumber="4">Fig. 3.</emphasis>
<taxonomicName id="4C304D1DFF9F0413CAD3F8A63E89F894" authorityName="Cai &amp; Ng" authorityYear="2018" box="[217,395,1869,1889]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="longshan" status="sp. nov.">
<emphasis id="B944EA8CFF9F0413CAD3F8A63E89F894" box="[217,395,1869,1889]" italics="true" pageId="3" pageNumber="4">Caridina longshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF9F0413CB99F8A63EE3F894" box="[403,481,1869,1889]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
(A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) propodus and dactylus of fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A-C, E, F = 1.0 mm; D, G = 0.2 mm. (A-E, G, female, female, cl 6.5 mm, Panon Cave, Huaoyan Village,
<taxonomicName id="4C304D1DFF9F0413C996F86E3B4AF86C" authority="County" authorityName="County" box="[924,1096,1925,1945]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="longshan">Longshan County</taxonomicName>
, Hunan Province, paratype, ZRC 2018.0480; F, wanting leg of another specimen, same location).
</paragraph>
</caption>
<footnote id="E82B2A90FF9F0413CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="3" pageNumber="4">
<paragraph id="8B8F369EFF9F0413CA82F8363E9AF81A" blockId="3.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="3" pageNumber="4">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF980414CAE0FD363DDDFC23" blockId="4.[174,775,253,1047]" pageId="4" pageNumber="5">
Two characters of this species are unusual: the pigmentation in the cornea of eyes and the presence of spinules on the surface of distal spines (serrated setae in
<bibRefCitation id="EFA14B6FFF980414CB96FCD63DDCFCA3" author="Garms A &amp; Watling L." box="[412,734,829,855]" pageId="4" pageNumber="5" pagination="167 - 198" refId="ref15133" refString="Garms A, Watling L. 2013. The crustacean integument: setae, setules, and other ornamentation. In: Watling L, Martin T. 2013. (ed.). Functional morphology and diversity. Oxford University Press, pp. 167 - 198." type="book chapter" year="2013">Garms and Watling 2013</bibRefCitation>
) of telson (
<figureCitation id="130B2A1BFF980414CB0EFCB63E79FC83" box="[260,379,861,886]" captionStart="Fig" captionStartId="2.[176,211,1841,1861]" captionTargetBox="[169,1452,916,1812]" captionTargetId="figure-372@2.[156,1477,907,1826]" captionTargetPageId="2" captionText="Fig. 2. Caridina longshan sp. nov. (A) mandible, (B) maxillula, (C) maxilla, (D) first maxilliped, (E) second maxilliped, (F) third maxilliped, (G) distal portion of third maxilliped, (H) male first pleopod, (I) endopod of male first pleopod, (J) male second pleopod, (K) distal portion of telson, (L) sublateral spine, right, (M) uropodal diaeresis. Scale bars: A-F = 1.0 mm; G-J, M = 0.5 mm; K = 0.2 mm; L = 0.1 mm. (A-G, K, L, female, cl 6.5 mm, Cave of Panon, Huaoyan Village, Longshan County, Hunan Province, Paratype, ZRC 2018.0480; H, J, cl 8.8 mm, male, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489)." figureDoi="http://doi.org/10.5281/zenodo.13314995" httpUri="https://zenodo.org/record/13314995/files/figure.png" pageId="4" pageNumber="5">Fig. 2K, L</figureCitation>
). The condition of the eye cornea pigmentation ranges from totally absent to just a black spot (
<figureCitation id="130B2A1BFF980414CB37FC763E7DFC43" box="[317,383,925,950]" captionStart="Fig" captionStartId="1.[136,170,1813,1833]" captionTargetBox="[136,1414,945,1784]" captionTargetId="figure-6@1.[125,1428,930,1787]" captionTargetPageId="1" captionText="Fig. 1. Caridina longshan sp. nov. (A-D) cephalothorax and cephalic appendages, lateral view, (E) antennular peduncle, ventral view, (F) scaphocerite, (G) preanal carina. Scale bars: A-D, F = 2.0 mm; E = 1.0 mm; G = 0.5 mm. (A, E-G, male, cl 8.8 mm, Parking Cave, near Hoyan Village, Longshan County, paratype, ZRC 2018.0489; B. female, cl 7.4 mm, Parking Cave, near Hoyan Village, Longshan County, Hunan Province, paratype, UL; C, female, cl 6.5 mm, Panon Cave, Huaoyan Village, Longshan County, Hunan Province, paratype, ZRC 2018.0480; D, female, cl 6.8 mm, Cave of flying Tiger, Hoyan Village, Longshan County, Hunan Province, paratype, RMNHCRUS.D.57289)." figureDoi="http://doi.org/10.5281/zenodo.13314993" httpUri="https://zenodo.org/record/13314993/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
A-D). The adaptive significance of the serrated setae of the telson is not known.
</paragraph>
<paragraph id="8B8F369EFF980414CAE0FC363FFEFBE2" blockId="4.[174,775,253,1047]" pageId="4" pageNumber="5">
<emphasis id="B944EA8CFF980414CAE0FC363E7EFC03" box="[234,380,989,1014]" italics="true" pageId="4" pageNumber="5">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF980414CB9BFC363D63FC03" box="[401,609,989,1014]" country="China" name="Hunan" pageId="4" pageNumber="5">Hunan Province</collectingRegion>
(Longshan),
<collectingCountry id="F327760EFF980414CAA4FC163FFAFBE2" box="[174,248,1021,1047]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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@ -53,7 +56,7 @@
<subSubSection id="C30FCAA3FFC85333E3E5FAE0FE83136B" box="[151,289,1353,1380]" pageId="1" pageNumber="28" type="description">
<paragraph id="8BAA9928FFC85333E3E5FAE0FE83136B" blockId="1.[151,412,1321,1380]" box="[151,289,1353,1380]" pageId="1" pageNumber="28">
(
<figureCitation id="132E85ADFFC85333E3D2FAE0FEBA136B" box="[160,280,1353,1380]" captionStart="Figures 2-19" captionStartId="2.[151,237,1871,1895]" captionTargetBox="[181,1405,189,1825]" captionTargetId="figure-14@2.[181,1406,189,1853]" captionTargetPageId="2" captionText="Figures 2-19. Opilio rossicus sp.n.: 2 male body, dorsal view; 3 male body, lateral view; 4 female body, dorsal view; 5 eye mound, dorsal view (male or female); 6 eye mound, lateral view (male or female); 7 male femur I, dorsal view; 8 - male right pedipalp, mesal view; 9 male right pedipalp, ectal view; 10- male right chelicera, ectal view; 11 male right chelicera, mesal view; 12 - female right pedipalp, mesal view; 13 - female right pedipalp, ectal view; 14 - female right chelicera, mesal view; 15 female right chelicera, ectal view; 16 penis, dorsal view; 17 penis, lateral view; 18 glans of penis, dorsal view; 20 glans of penis, lateral view." pageId="1" pageNumber="28">Figs. 2-19</figureCitation>
<figureCitation id="132E85ADFFC85333E3D2FAE0FEBA136B" box="[160,280,1353,1380]" captionStart="Figures 2-19" captionStartId="2.[151,237,1871,1895]" captionTargetBox="[181,1405,189,1825]" captionTargetId="figure-14@2.[181,1406,189,1853]" captionTargetPageId="2" captionText="Figures 2-19. Opilio rossicus sp.n.: 2 male body, dorsal view; 3 male body, lateral view; 4 female body, dorsal view; 5 eye mound, dorsal view (male or female); 6 eye mound, lateral view (male or female); 7 male femur I, dorsal view; 8 - male right pedipalp, mesal view; 9 male right pedipalp, ectal view; 10- male right chelicera, ectal view; 11 male right chelicera, mesal view; 12 - female right pedipalp, mesal view; 13 - female right pedipalp, ectal view; 14 - female right chelicera, mesal view; 15 female right chelicera, ectal view; 16 penis, dorsal view; 17 penis, lateral view; 18 glans of penis, dorsal view; 20 glans of penis, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13314894" httpUri="https://zenodo.org/record/13314894/files/figure.png" pageId="1" pageNumber="28">Figs. 2-19</figureCitation>
)
</paragraph>
</subSubSection>
@ -61,8 +64,10 @@
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<emphasis id="B961453AFFC85333E3E5FA24FEA613A8" bold="true" box="[151,260,1421,1447]" pageId="1" pageNumber="28">Material</emphasis>
:
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<typeStatus id="54AE278AFFC85333E26FFA24FE2813A7" box="[285,394,1421,1448]" pageId="1" pageNumber="28" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1352A1FFC85333E2E9FA27FE1113A7" box="[411,435,1422,1448]" count="1" pageId="1" pageNumber="28" type="male"></specimenCount>
</materialsCitation>
,
<materialsCitation id="3B7D9375FFC85333E2BBFA27FBB813C5" collectingDate="2009-09-10" collectionCode="V" collectorName="Perepechaenko" country="Russia" county="Orlov district" location="Manich-Gudilo Lake Valley" municipality="Manich" pageId="1" pageNumber="28" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Rostov" typeStatus="paratype">
<typeStatus id="54AE278AFFC85333E2BBFA27FD9913A7" box="[457,571,1422,1448]" pageId="1" pageNumber="28" type="paratype">Paratypes</typeStatus>
@ -101,7 +106,7 @@ genus, with 3-4 denticles on each side. Legs short, femora I short (1.75), cylin
<paragraph id="8BAA9928FFC85333E3E5F914FB381150" blockId="1.[151,1436,1725,1887]" pageId="1" pageNumber="28">Pedipalps small, femur dorsally with black-tipped denticles, ventrally and laterally with large denticles, patella dorsally and ventrally with denticles, tibia dorsally and ventrally with several denticles. Pedipalp lengths 1.4+0.25+0.5+0.95 = 3.1. Chelicera small, basal segment dorsally with some denticles. Cheliceral basal segment length 1.1, distal segment 1.1. Penis small with wide base (1.65 long, glans 0.2, stylus 0.08) narrows to glans, wings small size, glans of the penis oval in form with 2 pairs of setae.</paragraph>
<paragraph id="8BAA9928FFC85333E3E5F82EFC9011EA" blockId="1.[151,1436,1927,2021]" pageId="1" pageNumber="28">Female differs from male by being larger in size, more oval form of the body and less developed denticles on the body and extremities. Female body length 4.8, wide 2.5. Pedipalp lengths 0.9+0.4+0.6+2.4 = 4.3. Cheliceral basal segment length 2.1, distal segment 1.25.</paragraph>
</subSubSection>
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<emphasis id="B961453AFFCB5330E3E5F8E6FE8F1168" bold="true" box="[151,301,1871,1895]" pageId="2" pageNumber="29">Figures 2-19.</emphasis>
<taxonomicName id="4C15E2ABFFCB5330E244F8E6FE781168" authorityName="Snegovaya" authorityYear="2016" box="[310,474,1871,1895]" class="Arachnida" family="Phalangiidae" genus="Opilio" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="rossicus" status="sp. nov.">

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Genus
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;
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<bibRefCitation id="EFF04B2AFFB84A20FD76FA27FEF9FA11" author="Tiwari KK" pageId="2" pageNumber="3" pagination="295 - 300" refId="ref9673" refString="Tiwari KK. 1952. On a new species of the rare cymothoid genus Agarna Schi. &amp; Mein., parasitic on the clupeid fish Nematalosa nasus (Bl.) in the Bay of Bengal. Records of the Indian Museum 50: 295 - 300." type="journal article" year="1952">Tiwari 1952: 295-300</bibRefCitation>
</treatmentCitation>
, pl. iv, text fig. 1-2;
<bibRefCitation id="EFF04B2AFFB84A20FDE1FA0DFDB9FA11" author="Pillai NK" box="[516,632,1462,1482]" pageId="2" pageNumber="3" pagination="1 - 21" refId="ref9194" refString="Pillai NK. 1954. A preliminary note on the Tanaidacea and Isopoda of Travancore. Bulletin of the Central Research Institute, University of Kerala, India, Natural Sciences 3: 1 - 21." type="journal article" year="1954">Pillai, 1954</bibRefCitation>
;
<bibRefCitation id="EFF04B2AFFB84A20FD61FA0DFE9CFA3E" author="Bowman TE &amp; Tareen IU" pageId="2" pageNumber="3" pagination="1 - 30" refId="ref8495" refString="Bowman TE, Tareen IU. 1983. Cymothoidae from Fishes of Kuwait (Arabian Gulf) (Crustacea, Isopoda). Smith Contr Zool 382: 1 - 30." type="journal article" year="1983">Bowman and Tareen 1983</bibRefCitation>
, 21;
<bibRefCitation id="EFF04B2AFFB84A20FE6DFA6AFDC4FA3E" author="Aneesh PT" box="[392,517,1489,1509]" pageId="2" pageNumber="3" refId="ref8134" refString="Aneesh PT. 2014. Studies on parasitic crustaceans infesting the fishes of Malabar Coast. PhD dissertation, Kannur University, 147 pp." type="book" year="2014">Aneesh 2014</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B6550FFB84A20FF0FF9B7FD3CF9FE" box="[234,765,1548,1573]" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BDE36DBFFB84A20FF0FF9B7FD3CF9FE" blockId="2.[234,765,1548,1573]" box="[234,765,1548,1573]" pageId="2" pageNumber="3">
<emphasis id="B915EAC9FFB84A20FF0FF9B7FE44F9FE" box="[234,389,1548,1573]" italics="true" pageId="2" pageNumber="3">
<typeStatus id="54DA8879FFB84A20FF0FF9B7FEE2F9FE" box="[234,291,1548,1573]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C614D58FFB84A20FE71F9B7FD3CF9FE" authority="(Haller, 1880)" baseAuthorityName="Haller" baseAuthorityYear="1880" box="[404,765,1548,1573]" class="Malacostraca" family="Cymothoidae" genus="Agarna" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cumulus">
<emphasis id="B915EAC9FFB84A20FE71F9B7FD94F9FE" box="[404,597,1548,1573]" italics="true" pageId="2" pageNumber="3">Agarna cumulus</emphasis>
(Haller, 1880)
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<figureCitation id="135DE2A7FFD5FFF3FCF7FD86FC3FFDFB" box="[889,957,533,560]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="2" pageNumber="575">Fig. 1</figureCitation>
)
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@ -200,7 +201,7 @@ M 1.21, widest somewhat in front of the ¾ length level, narrower than pronotum
<specimenCode id="DBC05659FFD5FFF3FD17FBA3FD4DFB80" box="[665,719,1072,1099]" collectionCode="HW" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="575">1.18</specimenCode>
, M 1.17) (
<figureCitation id="135DE2A7FFD5FFF3FCDEFBA3FC24FB80" box="[848,934,1072,1099]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="2" pageNumber="575">Fig. 1a</figureCitation>
<figureCitation id="135DE2A7FFD5FFF3FCDEFBA3FC24FB80" box="[848,934,1072,1099]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="2" pageNumber="575">Fig. 1a</figureCitation>
). Eyes completely missing, each replaced by a darkened arcuate slit. Frontal furrows regularly arcuate or sub-arcuate, anteriorly deepened, ending at the level between 1
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and 2
@ -224,12 +225,12 @@ male = 1.08: 1.00: 1.24: 1.04: 1.04: 1.04: 1.00: 0.96: 1.00: 0.94: 1.20. Antenno
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<paragraph id="8BD9FE22FFD5FFF3FF79FA8EFC16FA2B" blockId="2.[151,1437,600,2044]" pageId="2" pageNumber="575">
Pronotum cordiform, rounded, transverse (PW/PL R 1.121.21, M 1.16), widest somewhat below the ¼ length level, somewhat constricted towards base (PW/PB R 1.531.625, M 1.57), lateral margins anteriorly rounded, posteriorly narrowing, sinuate (
<figureCitation id="135DE2A7FFD5FFF3FD74FAF3FCCDFAB0" box="[762,847,1376,1403]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="2" pageNumber="575">Fig. 1a</figureCitation>
<figureCitation id="135DE2A7FFD5FFF3FD74FAF3FCCDFAB0" box="[762,847,1376,1403]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="2" pageNumber="575">Fig. 1a</figureCitation>
). Hind angles sharp to almost rectangular, acute, laterally dragged, fore angles rounded, protruding forward. Base almost straight. Basal fossettes small and deep. Midline basally deepened and broadened, reaching pronotal base. Lateral furrows anteriorly wider, posteriorly narrow. Two pairs of marginal pronotal setae present.
</paragraph>
<paragraph id="8BD9FE22FFD5FFF3FF79FA74FBC4F907" blockId="2.[151,1437,600,2044]" pageId="2" pageNumber="575">
Elytra rounded, oval (EL/EW R 1.601.82, M 1.69), widest at the middle, in average less than three times as long as pronotum (male: EL/PL 3.085; females: R 2.802.915, M 2.86), considerably wider than pronotum (EW/PW R 1.451.60, M 1.55) (
<figureCitation id="135DE2A7FFD5FFF3FD10F9B8FD76F98D" box="[670,756,1579,1606]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="2" pageNumber="575">Fig. 1a</figureCitation>
<figureCitation id="135DE2A7FFD5FFF3FD10F9B8FD76F98D" box="[670,756,1579,1606]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="2" pageNumber="575">Fig. 1a</figureCitation>
). Shoulders protruding forward, obtuse-angled, rounded. Lateral furrows moderately wide anteriorly, gradually narrowing posteriorly. Striae poorly punctate, the three inner ones deepened, from the fourth being weakly noticeable. Striae 57 posteriorly not completely disappeared, barely visible. Fifth striae at the beginning conspicuously impressed. Eighth striae being the weakest. Inner interstriae convex, the remaining flattened. Elytral apex rounded.
</paragraph>
<paragraph id="8BD9FE22FFD5FFF3FF79F940FC64F837" blockId="2.[151,1437,600,2044]" pageId="2" pageNumber="575">
@ -259,7 +260,7 @@ and 3
<superScript id="7C13536AFFD5FFF3FAB5F8C7FAC8F8AE" attach="left" box="[1339,1354,1876,1893]" fontSize="7" pageId="2" pageNumber="575">rd</superScript>
striae, much closer to elytral suture than to elytral apex. Post-humeral setae 1 and 2 mutually somewhat more distanced than the distances 23 and 34 (the latter two distances being the same). Third and fourth post-humeral setae on eighth striae, out of elytral marginal furrow. The distance between fore and median groups of the umbilicate series around twice as long as the length of fore series.
</paragraph>
<caption id="DF19AEAAFFD4FFF2FF19F8BCFE3FF869" pageId="3" pageNumber="576" startId="3.[151,226,1839,1863]" targetBox="[188,1427,237,1724]" targetPageId="3" targetType="figure">
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<paragraph id="8BD9FE22FFD4FFF2FF19F8BCFE3FF869" blockId="3.[151,1436,1838,1954]" pageId="3" pageNumber="576">
<emphasis id="B9122230FFD4FFF2FF19F8BCFE82F88C" bold="true" box="[151,256,1839,1863]" pageId="3" pageNumber="576">Figure 1.</emphasis>
<taxonomicName id="4C6685A1FFD4FFF2FE85F8BDFDEDF88D" authorityName="Ćurĉić &amp; Pavićević &amp; Vesović &amp; Petković" authorityYear="2016" box="[267,623,1838,1863]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="bozidari" status="sp. nov." subGenus="Neoduvalius">
@ -277,7 +278,7 @@ from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbi
</caption>
<paragraph id="8BD9FE22FFD3FFF5FF79FF57FE02FE88" blockId="4.[151,1437,195,1841]" pageId="4" pageNumber="577">
Legs of medium length (
<figureCitation id="135DE2A7FFD3FFF5FDBBFF50FD12FF15" box="[565,656,195,222]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1a</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FDBBFF50FD12FF15" box="[565,656,195,222]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1a</figureCitation>
). Hind tarsi relatively long and narrow (male: EL/HTL 2.76; females: R 2.61, M 2.61). First metatarsomere much longer than 23 combined, slightly shorter than 24 combined. A weakly expressed fissure on each protibia at the outer side. First protasomere length/width in
<typeStatus id="54DD4080FFD3FFF5FF19FEBBFF7CFE88" box="[151,254,296,323]" pageId="4" pageNumber="577" type="holotype">holotype</typeStatus>
male 1.33.
@ -285,7 +286,7 @@ male 1.33.
<paragraph id="8BD9FE22FFD3FFF5FF79FED9FDF4FD9A" blockId="4.[151,1437,195,1841]" pageId="4" pageNumber="577">
<materialsCitation id="3B0EF47FFFD3FFF5FF79FED9FDF2FD9A" collectionCode="EL" pageId="4" pageNumber="577" specimenCode="AEL 2.40" specimenCount="1" typeStatus="holotype">
Aedeagus (
<figureCitation id="135DE2A7FFD3FFF5FEF1FED9FDBDFEAE" box="[383,575,330,357]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Figs. 1b and 1c</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FEF1FED9FDBDFEAE" box="[383,575,330,357]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Figs. 1b and 1c</figureCitation>
)
<quantity id="4C9E53C7FFD3FFF5FDDBFED9FD42FEAE" box="[597,704,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="577" unit="mm" value="1.13">1.13 mm</quantity>
long in
@ -295,33 +296,33 @@ long in
/
<specimenCode id="DBC05659FFD3FFF5FA9CFED9FA0DFEAE" box="[1298,1423,330,357]" collectionCode="AEL" pageId="4" pageNumber="577">AEL 2.40</specimenCode>
). Medial lobe in lateral view basally curved, anteriorly straight, medially convex, with a narrow apex moderately dragged upwards (
<figureCitation id="135DE2A7FFD3FFF5FD8CFE1DFDDBFE62" box="[514,601,398,425]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1b</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FD8CFE1DFDDBFE62" box="[514,601,398,425]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1b</figureCitation>
). Basal bulb in lateral view relatively small, elongately rounded (
<figureCitation id="135DE2A7FFD3FFF5FAE2FE1DFF34FE01" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1b</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FAE2FE1DFF34FE01" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1b</figureCitation>
). Medial lobe in dorsal view wide, sub-parallel, gradually narrowing towards a rounded top, while basal bulb rounded (
<figureCitation id="135DE2A7FFD3FFF5FECBFE42FE19FE27" box="[325,411,465,492]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1c</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FECBFE42FE19FE27" box="[325,411,465,492]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1c</figureCitation>
). Aedeagus dorsally weakly sclerotized (
<figureCitation id="135DE2A7FFD3FFF5FC0AFE42FC5AFE27" box="[900,984,465,492]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1c</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FC0AFE42FC5AFE27" box="[900,984,465,492]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1c</figureCitation>
), while ventrally well chitinized (
<figureCitation id="135DE2A7FFD3FFF5FAE2FE42FF34FDC5" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1b</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FAE2FE42FF34FDC5" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1b</figureCitation>
). Copulatory piece situated in an inner sac, gutter-formed, weakly chitinized, barely visible, thus the details could not be observed (
<figureCitation id="135DE2A7FFD3FFF5FD83FD86FDE7FDFB" box="[525,613,533,560]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1c</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FD83FD86FDE7FDFB" box="[525,613,533,560]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1c</figureCitation>
). Parameres slender, somewhat widened basally, each carrying four short and narrow setae (
<figureCitation id="135DE2A7FFD3FFF5FE3EFDA5FDE4FD9A" box="[432,614,566,593]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Figs. 1b and 1c</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FE3EFDA5FDE4FD9A" box="[432,614,566,593]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Figs. 1b and 1c</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9FE22FFD3FFF5FF78FDCBFC77FDB8" blockId="4.[151,1437,195,1841]" box="[246,1013,600,627]" pageId="4" pageNumber="577">
Male abdominal sternite IX (urite) large, sub-triangular (
<figureCitation id="135DE2A7FFD3FFF5FC01FDCBFC67FDB8" box="[911,997,600,627]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1d</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FC01FDCBFC67FDB8" box="[911,997,600,627]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1d</figureCitation>
).
</paragraph>
<paragraph id="8BD9FE22FFD3FFF5FF79FDE9FD8DFD13" blockId="4.[151,1437,195,1841]" pageId="4" pageNumber="577">
Both gonocoxites and gonosubcoxites IX as presented in
<figureCitation id="135DE2A7FFD3FFF5FC3CFDE9FB88FD5E" box="[946,1034,634,661]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1e</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FC3CFDE9FB88FD5E" box="[946,1034,634,661]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1e</figureCitation>
. Gonocoxites IX short and wide, curved, gradually narrowing distally, each with a pointed apex, basally joined with rounded gonosubcoxites IX of moderate length (
<figureCitation id="135DE2A7FFD3FFF5FE25FD2EFE7DFD13" box="[427,511,701,728]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." pageId="4" pageNumber="577">Fig. 1e</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FE25FD2EFE7DFD13" box="[427,511,701,728]" captionStart="Figure 1" captionStartId="3.[151,226,1839,1863]" captionTargetBox="[188,1427,237,1724]" captionTargetId="figure-7@3.[151,1436,189,1802]" captionTargetPageId="3" captionText="Figure 1. Duvalius (Neoduvalius) bozidari sp. n. from the Jovanjska Pećina Cave, village of Jovanje, near Valjevo, western Serbia: a - holotype male, habitus (dorsal view); b - holotype male, aedeagus (lateral view); c - holotype male, aedeagus (dorsal view); d - holotype male, abdominal sternite IX (urite); e - paratype female, genitalia. Scales = 2.00 mm (a) and 0.20 mm (be)." figureDoi="http://doi.org/10.5281/zenodo.13315066" httpUri="https://zenodo.org/record/13315066/files/figure.png" pageId="4" pageNumber="577">Fig. 1e</figureCitation>
).
</paragraph>
</subSubSection>
@ -441,7 +442,7 @@ discal seta-apex of elytra (M 1.315
is endogean and inhabits Mt. Maljen (Sedlo, Divĉibare,
<quantity id="4C9E53C7FFD3FFF5FDEAF9B8FD6BF98D" box="[612,745,1579,1606]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="4" pageNumber="577" unit="m" value="800.0" valueMax="900.0" valueMin="700.0">700900 m</quantity>
a.s.l.) (
<figureCitation id="135DE2A7FFD3FFF5FCCFF9B8FC0AF98D" box="[833,904,1579,1606]" captionStart="Figure 2" captionStartId="5.[151,226,973,997]" captionTargetBox="[375,1251,193,954]" captionTargetId="figure-7@5.[334,1251,189,954]" captionTargetPageId="5" captionText="Figure 2. The distribution of Duvalius (Neoduvalius) bozidari sp. n. (black circles) and D. (N.) guidononveilleri Janák &amp; Moravec, 2008 (black star) in Serbia." pageId="4" pageNumber="577">Fig. 2</figureCitation>
<figureCitation id="135DE2A7FFD3FFF5FCCFF9B8FC0AF98D" box="[833,904,1579,1606]" captionStart="Figure 2" captionStartId="5.[151,226,973,997]" captionTargetBox="[375,1251,193,954]" captionTargetId="figure-7@5.[334,1251,189,954]" captionTargetPageId="5" captionText="Figure 2. The distribution of Duvalius (Neoduvalius) bozidari sp. n. (black circles) and D. (N.) guidononveilleri Janák &amp; Moravec, 2008 (black star) in Serbia." figureDoi="http://doi.org/10.5281/zenodo.13315068" httpUri="https://zenodo.org/record/13315068/files/figure.png" pageId="4" pageNumber="577">Fig. 2</figureCitation>
) (
<bibRefCitation id="EFF783D3FFD3FFF5FC2EF9B8FB3BF98D" author="Janak, J. &amp; Moravec, P." box="[928,1209,1579,1606]" pageId="4" pageNumber="577" pagination="1 - 19" refId="ref3594" refString="Janak, J. &amp; Moravec, P. (2008) Drei neue Duvalius - Arten aus Bulgarien und Serbien (Coleoptera: Carabidae: Trechinae). Klapalekiana, 44, 1 - 19." type="journal article" year="2008">Janák &amp; Moravec 2008</bibRefCitation>
).

View file

@ -1,48 +1,49 @@
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@ -64,7 +65,7 @@
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(
<figureCitation id="136F44AFFFBC370BFF2FF9167662F95C" box="[160,265,1745,1772]" captionStart-0="Figures 1-2" captionStart-1="Figures 3-7" captionStartId-0="1.[151,237,1949,1973]" captionStartId-1="2.[151,237,1561,1585]" captionTargetBox-0="[151,1417,1113,1900]" captionTargetBox-1="[170,1422,199,1518]" captionTargetId-0="figure-379@1.[151,1436,1100,1930]" captionTargetId-1="figure-17@2.[151,1436,189,1554]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." captionText-1="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="0" pageNumber="34">Figs. 1-7</figureCitation>
<figureCitation id="136F44AFFFBC370BFF2FF9167662F95C" box="[160,265,1745,1772]" captionStart-0="Figures 1-2" captionStart-1="Figures 3-7" captionStartId-0="1.[151,237,1949,1973]" captionStartId-1="2.[151,237,1561,1585]" captionTargetBox-0="[151,1417,1113,1900]" captionTargetBox-1="[170,1422,199,1518]" captionTargetId-0="figure-379@1.[151,1436,1100,1930]" captionTargetId-1="figure-17@2.[151,1436,189,1554]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." captionText-1="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13314868" figureDoi-1="http://doi.org/10.5281/zenodo.13314870" httpUri-0="https://zenodo.org/record/13314868/files/figure.png" httpUri-1="https://zenodo.org/record/13314870/files/figure.png" pageId="0" pageNumber="34">Figs. 1-7</figureCitation>
)
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@ -162,7 +163,7 @@ short, thick; dorsal shield usually bearing 9 pairs setae; coxal plates in three
</paragraph>
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Dorsal and ventral shields present. Dorsal shield (
<figureCitation id="136F44AFFFBD370AFD7FFDD2745DFD80" box="[752,822,533,560]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." pageId="1" pageNumber="35">Fig. 1</figureCitation>
<figureCitation id="136F44AFFFBD370AFD7FFDD2745DFD80" box="[752,822,533,560]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13314868" httpUri="https://zenodo.org/record/13314868/files/figure.png" pageId="1" pageNumber="35">Fig. 1</figureCitation>
) elongate (ratio L/
<geoCoordinate id="EE603EEDFFBD370AFB92FDD2731EFD80" box="[1053,1141,533,560]" degrees="1.35" direction="west" orientation="longitude" pageId="1" pageNumber="35" precision="555" value="-1.35">W 1.35</geoCoordinate>
-1.50), usually bearing 9 pairs of setae (
@ -193,7 +194,7 @@ i
</paragraph>
<paragraph id="8BEB582AFFBD370AFF79FD18778FFC74" blockId="1.[151,1437,500,1032]" pageId="1" pageNumber="35">
Coxal plates and genital field incorporated into ventral shield (
<figureCitation id="136F44AFFFBD370AFC6CFD187346FD4A" box="[995,1069,735,762]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." pageId="1" pageNumber="35">Fig. 2</figureCitation>
<figureCitation id="136F44AFFFBD370AFC6CFD187346FD4A" box="[995,1069,735,762]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13314868" httpUri="https://zenodo.org/record/13314868/files/figure.png" pageId="1" pageNumber="35">Fig. 2</figureCitation>
). Coxal plates in three groups. Capitular bay V-shaped; coxal plates I fused to each other medially but suture line between them present. Sclerites, bearing setae
<emphasis id="B9208438FFBD370AFE35FCE37688FC8D" box="[442,483,804,829]" italics="true" pageId="1" pageNumber="35">Hv,</emphasis>
located on coxal plates II posterolaterally. Anterior coxal group with narrow posterior apodemes directed laterally. Glandularia
@ -202,12 +203,12 @@ on the coxal plate IV shifted from suture line between coxal plates III+IV. Geni
</paragraph>
<paragraph id="8BEB582AFFBD370AFF79FC0C7318FBB8" blockId="1.[151,1437,500,1032]" pageId="1" pageNumber="35">
Capitulum (
<figureCitation id="136F44AFFFBD370AFE0CFC0C76A0FC56" box="[387,459,971,998]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="1" pageNumber="35">Fig. 3</figureCitation>
<figureCitation id="136F44AFFFBD370AFE0CFC0C76A0FC56" box="[387,459,971,998]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="1" pageNumber="35">Fig. 3</figureCitation>
) with moderately long anchoral process and two pairs of subequal anterior setae; rostrum relatively short. Chelicera with large basal segment and short chela (
<figureCitation id="136F44AFFFBD370AFB93FC2A730EFBB8" box="[1052,1125,1005,1032]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="1" pageNumber="35">Fig. 4</figureCitation>
<figureCitation id="136F44AFFFBD370AFB93FC2A730EFBB8" box="[1052,1125,1005,1032]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="1" pageNumber="35">Fig. 4</figureCitation>
).
</paragraph>
<caption id="DF2B08A2FFBD370AFF18F85A723DF805" box="[151,1366,1948,1973]" pageId="1" pageNumber="35" startId="1.[151,237,1949,1973]" targetBox="[151,1417,1113,1900]" targetPageId="1" targetType="figure">
<caption id="DF2B08A2FFBD370AFF18F85A723DF805" ID-DOI="http://doi.org/10.5281/zenodo.13314868" ID-Zenodo-Dep="13314868" box="[151,1366,1948,1973]" httpUri="https://zenodo.org/record/13314868/files/figure.png" pageId="1" pageNumber="35" startId="1.[151,237,1949,1973]" targetBox="[151,1417,1113,1900]" targetPageId="1" targetType="figure">
<paragraph id="8BEB582AFFBD370AFF18F85A723DF805" blockId="1.[151,1366,1948,1973]" box="[151,1366,1948,1973]" pageId="1" pageNumber="35">
<emphasis id="B9208438FFBD370AFF18F85A767CF805" bold="true" box="[151,279,1949,1973]" pageId="1" pageNumber="35">Figures 1-2</emphasis>
.
@ -218,7 +219,7 @@ Capitulum (
, female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm.
</paragraph>
</caption>
<caption id="DF2B08A2FFBE3709FF18F9DE747BF9E2" pageId="2" pageNumber="36" startId="2.[151,237,1561,1585]" targetBox="[170,1422,199,1518]" targetPageId="2" targetType="figure">
<caption id="DF2B08A2FFBE3709FF18F9DE747BF9E2" ID-DOI="http://doi.org/10.5281/zenodo.13314870" ID-Zenodo-Dep="13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="2" pageNumber="36" startId="2.[151,237,1561,1585]" targetBox="[170,1422,199,1518]" targetPageId="2" targetType="figure">
<paragraph id="8BEB582AFFBE3709FF18F9DE747BF9E2" blockId="2.[151,1436,1560,1618]" pageId="2" pageNumber="36">
<emphasis id="B9208438FFBE3709FF18F9DE7670F981" bold="true" box="[151,283,1561,1585]" pageId="2" pageNumber="36">Figures 3-7</emphasis>
.
@ -231,22 +232,22 @@ Capitulum (
</caption>
<paragraph id="8BEB582AFFBE3709FF18F95A7460F832" blockId="2.[151,1436,1693,2023]" pageId="2" pageNumber="36">
Pedipalp (
<figureCitation id="136F44AFFFBE3709FE98F95A760CF908" box="[279,359,1693,1720]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="2" pageNumber="36">Fig. 5</figureCitation>
<figureCitation id="136F44AFFFBE3709FE98F95A760CF908" box="[279,359,1693,1720]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="2" pageNumber="36">Fig. 5</figureCitation>
) compact: P-1 short, with single short dorsodistal seta; P2 with long ventrodistal projection, bearing numerous small teeth, with three subequal proximal and two unequal distal short dorsal setae; P3 ventrally denticulate, with single dorsoproximal spine and two thin dorsodistal setae; P4 slender, with obtuse ventral protrusion bearing peg-like seta and thin seta near middle of segment, second ventral seta located on half way between anterior ventral seta and end of segment and three thin distal setae; P5 thin, with two hook-like distal spines and six unequal distal setae. Surface P-2 - P-4 striated. P-3 ventral teeth much larger than teeth on P-2 ventrodistal projection.
</paragraph>
<paragraph id="8BEB582AFFBE3709FF79F84D7242F814" blockId="2.[151,1436,1693,2023]" box="[246,1321,1929,1956]" pageId="2" pageNumber="36">
Legs without swimming setae. Shape and arrangement of setae on legs I as shown in
<figureCitation id="136F44AFFFBE3709FB51F84E724DF814" box="[1246,1318,1929,1956]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="2" pageNumber="36">Fig. 6</figureCitation>
<figureCitation id="136F44AFFFBE3709FB51F84E724DF814" box="[1246,1318,1929,1956]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="2" pageNumber="36">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="8BEB582AFFBE3708FF18F86C7601FEB0" blockId="2.[151,1436,1693,2023]" lastBlockId="3.[151,1438,195,458]" lastPageId="3" lastPageNumber="37" pageId="2" pageNumber="36">
I-Leg-1 with long distoventral seta, I-Leg-2 and I-Leg-3 with sword-like distoventral seta each. I-Leg-5-6 bearing only thin setae. All legs claws (
<figureCitation id="136F44AFFFBE3709FDE2F80B75DFF857" box="[621,692,1996,2023]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="2" pageNumber="36">Fig. 7</figureCitation>
<figureCitation id="136F44AFFFBE3709FDE2F80B75DFF857" box="[621,692,1996,2023]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="2" pageNumber="36">Fig. 7</figureCitation>
) with three clawlets, central and internal clawlets nearly equal in length and longer than external clawlet; lamella with slightly convex ventral margin. All claws slightly indented dorsally.
</paragraph>
<paragraph id="8BEB582AFFBF3708FF79FECF750AFE7A" blockId="3.[151,1438,195,458]" pageId="3" pageNumber="37">Measurements (n=7). Idiosoma L 455-510; dorsal shields L 405-485, W 275-320; acetabular plates L 78-85, W 65-72; genital acetabula (ac-1-3) L 25-30, 30-35, 25-30; capitulum L 125-140, rostrum L 24-30; cheliceral segments: base L 150-162, chela L 40-48; pedipalp segments (P-1-5) L: 21-30, 70-73, 81-90, 96- 102, 35-37; leg segments L: I-Leg-1-6: 30-35, 45-60, 60-63, 77-84, 78-95, 70-90; II-Leg-1-6: 35-42, 42-55, 54- 65, 84-95, 84-90, 72-90; III-Leg-1-6: 35-48, 48-54, 72-75, 102-115, 108-120, 95-102; IV-Leg-1-6: 78-90, 65- 72, 95-108, 135-145, 150-156, 108-125.</paragraph>
</subSubSection>
<caption id="DF2B08A2FFBF3708FF18FBD77382FB98" box="[151,1257,1040,1064]" pageId="3" pageNumber="37" startId="3.[151,226,1040,1064]" targetBox="[198,1369,506,991]" targetPageId="3" targetType="figure">
<caption id="DF2B08A2FFBF3708FF18FBD77382FB98" ID-DOI="http://doi.org/10.5281/zenodo.13314872" ID-Zenodo-Dep="13314872" box="[151,1257,1040,1064]" httpUri="https://zenodo.org/record/13314872/files/figure.png" pageId="3" pageNumber="37" startId="3.[151,226,1040,1064]" targetBox="[198,1369,506,991]" targetPageId="3" targetType="figure">
<paragraph id="8BEB582AFFBF3708FF18FBD77382FB98" blockId="3.[151,1257,1040,1064]" box="[151,1257,1040,1064]" pageId="3" pageNumber="37">
<emphasis id="B9208438FFBF3708FF18FBD7779DFB98" bold="true" box="[151,246,1040,1064]" pageId="3" pageNumber="37">Figure 8</emphasis>
.
@ -274,15 +275,15 @@ but differs in the following characters (character states of female
given in parenthesis, data from
<bibRefCitation id="EFC525DBFFBF3708FA87FB5372E4FB1F" author="Cook, D. R." box="[1288,1423,1172,1199]" pageId="3" pageNumber="37" pagination="1 - 356" refId="ref1700" refString="Cook, D. R. (1988) Water mites from Chile. Memoirs of the American Entomological Institute, 42: 1 - 356. Tuzovskij P. V. (1987) Morfologiya i postembrional'noye razvitiye vodyanykh kleshchey [Morphology and" type="journal article" year="1988">Cook 1988</bibRefCitation>
): the excretory pore located dorsocaudally,
<figureCitation id="136F44AFFFBF3708FD09FB7175A0FB61" box="[646,715,1206,1233]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." pageId="3" pageNumber="37">Fig. 1</figureCitation>
<figureCitation id="136F44AFFFBF3708FD09FB7175A0FB61" box="[646,715,1206,1233]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13314868" httpUri="https://zenodo.org/record/13314868/files/figure.png" pageId="3" pageNumber="37">Fig. 1</figureCitation>
(ventrocaudally); the acetabular plates with distinct outlines,
<figureCitation id="136F44AFFFBF3708FF18FB1F77B5FB43" box="[151,222,1240,1267]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." pageId="3" pageNumber="37">Fig. 2</figureCitation>
<figureCitation id="136F44AFFFBF3708FF18FB1F77B5FB43" box="[151,222,1240,1267]" captionStart="Figures 1-2" captionStartId="1.[151,237,1949,1973]" captionTargetBox="[151,1417,1113,1900]" captionTargetId="figure-379@1.[151,1436,1100,1930]" captionTargetPageId="1" captionText="Figures 1-2. Corticacarus magellanicus sp. n., female: 1 - dorsal view; 2 - ventral view. Scale bar: 1-2 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13314868" httpUri="https://zenodo.org/record/13314868/files/figure.png" pageId="3" pageNumber="37">Fig. 2</figureCitation>
(without outlines); P-3 with comparatively large teeth,
<figureCitation id="136F44AFFFBF3708FCF1FB1F74AEFB43" box="[894,965,1240,1267]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="3" pageNumber="37">Fig. 5</figureCitation>
<figureCitation id="136F44AFFFBF3708FCF1FB1F74AEFB43" box="[894,965,1240,1267]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="3" pageNumber="37">Fig. 5</figureCitation>
(with very small teeth,
<figureCitation id="136F44AFFFBF3708FB67FB1F725BFB43" box="[1256,1328,1240,1267]" captionStart="Figure 8" captionStartId="3.[151,226,1040,1064]" captionTargetBox="[198,1369,506,991]" captionTargetId="figure-235@3.[151,1436,471,1022]" captionTargetPageId="3" captionText="Figure 8. Corticacarus argentinensis Cook, 1988, female: 8 pedipalp, lateral view (after Cook 1988)." pageId="3" pageNumber="37">Fig. 8</figureCitation>
<figureCitation id="136F44AFFFBF3708FB67FB1F725BFB43" box="[1256,1328,1240,1267]" captionStart="Figure 8" captionStartId="3.[151,226,1040,1064]" captionTargetBox="[198,1369,506,991]" captionTargetId="figure-235@3.[151,1436,471,1022]" captionTargetPageId="3" captionText="Figure 8. Corticacarus argentinensis Cook, 1988, female: 8 pedipalp, lateral view (after Cook 1988)." figureDoi="http://doi.org/10.5281/zenodo.13314872" httpUri="https://zenodo.org/record/13314872/files/figure.png" pageId="3" pageNumber="37">Fig. 8</figureCitation>
); the leg claws with dorsal indentation,
<figureCitation id="136F44AFFFBF3708FE71FB3D7528FAA5" box="[510,579,1274,1301]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." pageId="3" pageNumber="37">Fig. 7</figureCitation>
<figureCitation id="136F44AFFFBF3708FE71FB3D7528FAA5" box="[510,579,1274,1301]" captionStart="Figures 3-7" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[170,1422,199,1518]" captionTargetId="figure-17@2.[151,1436,189,1554]" captionTargetPageId="2" captionText="Figures 3-7. Corticacarus magellanicus sp. n., female: 3 capitulum, lateral view; 4 chelicera, lateral view; 5 pedipalp, lateral view; 6 Leg I; 7 claw. All bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13314870" httpUri="https://zenodo.org/record/13314870/files/figure.png" pageId="3" pageNumber="37">Fig. 7</figureCitation>
(without dorsal indentation).
</paragraph>
</subSubSection>

View file

@ -0,0 +1,189 @@
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<mods:title id="8BCB31E192876FAFD3A08B15BF807AB8">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="CE9CFA0A667DDE13B695EBD4E8E0EA7B">
<mods:date id="98CD54FFD751C416ADF3EEE5731557E7">2024</mods:date>
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<mods:number id="A93705176A6E5F7951C9C971CA479590">201</mods:number>
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<mods:start id="D80201BDE06DCB55E0C76AFA41259731">358</mods:start>
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<mods:identifier id="ECB5F10E4A135868EB09115A923ECC3D" type="DOI">10.1093/zooldnnean/zlad146</mods:identifier>
<mods:identifier id="ED9E32679DABA45BC15CFFC379F303E6" type="ISSN">0024-4082</mods:identifier>
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<treatment id="C27387A7FFE0FFCF9A050201FB0A7162" ID-DOI="http://doi.org/10.5281/zenodo.13315095" ID-Zenodo-Dep="13315095" LSID="urn:lsid:plazi:treatment:C27387A7FFE0FFCF9A050201FB0A7162" httpUri="http://treatment.plazi.org/id/C27387A7FFE0FFCF9A050201FB0A7162" lastPageNumber="376" pageId="18" pageNumber="376">
<subSubSection id="02C0653AFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" pageId="18" pageNumber="376" type="nomenclature">
<paragraph id="4A6536B1FFE0FFCF9A050201FD767058" blockId="18.[113,762,1852,2009]" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9A050201FD767058" bold="true" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
2.
<taxonomicName id="8DDA4D32FFE0FFCF9AFC0201FDAB7059" authority="(Mulsant and Rey)" baseAuthorityName="Mulsant and Rey" box="[142,552,1852,1879]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9AFC0201FEE17059" bold="true" box="[142,354,1852,1879]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
(Mulsant and Rey)
</taxonomicName>
stat. res.,comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE0FFCF9B290246FB0A7162" pageId="18" pageNumber="376" type="description">
<paragraph id="4A6536B1FFE0FFCF9B290246FD92709A" blockId="18.[113,762,1852,2009]" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B290246FD92709A" box="[347,529,1915,1941]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B290246FD92709A" bold="true" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B140246FE30709A" box="[358,435,1915,1941]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="18" pageNumber="376">Figs 2B</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BCF0246FE5C709A" box="[445,479,1915,1941]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="18" pageNumber="376">6G</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9B9B0246FD85709A" box="[489,518,1915,1941]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="18" pageNumber="376">8B</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03029FFD7970D6" blockId="18.[113,762,1852,2009]" pageId="18" pageNumber="376">
<pageTitle id="0A45EED6FFE0FFCF9A03029FFD7970D6" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03029FFED670B6" box="[113,341,1954,1977]" italics="true" pageId="18" pageNumber="376">Eurynotus asperipennis</emphasis>
Mulsant and Rey, 1854a: 23. Mulsant and Rey 1854b: 169; Lacordadre 1859: 239; Gebden 1938: 414;
</pageTitle>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805ADFC1B77C8" blockId="18.[810,1459,144,199]" pageId="18" pageNumber="376">
Koch 1954a: 280; 1954b: 2;
<bibRefCitation id="2E4B4B40FFE0FFCF9E2205ADFAC477A7" author="Banaszkdewdcz M" box="[1104,1351,144,168]" pageId="18" pageNumber="376" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
; Kamdńskd 2016: 235.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805D4FB0E7577" blockId="18.[810,1460,233,632]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995805D4FC13760F" box="[810,912,233,256]" italics="true" pageId="18" pageNumber="376">Diagnosis:</emphasis>
Lacks sclerdte dn bursa copulatrdx (present dn
<taxonomicName id="8DDA4D32FFE0FFCF9FED05D4FC1E762F" authorityName="Koch" authorityYear="1954" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9FED05D4FC1E762F" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF99920434FBE9762F" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[992,1130,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF99920434FBE9762F" box="[992,1130,264,288]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%.
<taxonomicName id="8DDA4D32FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
</taxonomicName>
has two puncture
<typeStatus id="95618813FFE0FFCF99E30415FC4B764F" box="[913,968,296,320]" pageId="18" pageNumber="376">types</typeStatus>
on ventrdte V (sdngle
<typeStatus id="95618813FFE0FFCF9EC80415FB6B764F" box="[1210,1256,296,320]" pageId="18" pageNumber="376">type</typeStatus>
dn
<taxonomicName id="8DDA4D32FFE0FFCF9F7D0415FAE77630" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1295,1380,296,319]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F7D0415FAE77630" box="[1295,1380,296,319]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF9FED0415FC127650" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9FED0415FC127650" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%, and golden setae on the tarsd (dark/black dn
<taxonomicName id="8DDA4D32FFE0FFCF9F2F047AFA2E7651" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1373,1453,327,350]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F2F047AFA2E7651" box="[1373,1453,327,350]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9958045AFC1C7671" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[810,927,358,382]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9958045AFC1C7671" box="[810,927,358,382]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%. Wdthout close dnspectdon,
<taxonomicName id="8DDA4D32FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE0FFCF990C04B8FC707692" box="[894,1011,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="18" pageNumber="376" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">1518 mm</quantity>
; wddth
<quantity id="8D229B54FFE0FFCF9E3004B8FB307692" box="[1090,1203,389,413]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="18" pageNumber="376" unit="mm" value="8.75" valueMax="10.0" valueMin="7.5">7.510 mm</quantity>
% specdmens may appear sdmdlar to smaller specdmens of
<taxonomicName id="8DDA4D32FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE0FFCF9F030498FCD576D4" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922 mm</quantity>
; wddth
<quantity id="8D229B54FFE0FFCF99ED04F9FC7D76D3" box="[927,1022,452,476]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
%; however,
<taxonomicName id="8DDA4D32FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
has golden tarsal setae (black or dark dn
<taxonomicName id="8DDA4D32FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%, a weaker cdrcumocular depres* sdon, the pronotal process ds more pronounced, and the male tdbdae are also at most only sldghtly moddfied dn
<taxonomicName id="8DDA4D32FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
, whdle dn
<taxonomicName id="8DDA4D32FFE0FFCF9958077FFC387556" box="[810,955,578,601]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9958077FFC387556" box="[810,955,578,601]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
they are more strongly curved. See the ddagnosds of
<taxonomicName id="8DDA4D32FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
below for more detadls.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995807A7FC6D720A" blockId="18.[810,1460,666,1285]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995807A7FC3E75BD" box="[810,957,666,690]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE0FFCF9E5B07A7FB1A75BD" box="[1065,1177,666,690]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922mm</quantity>
, wddth
<quantity id="8D229B54FFE0FFCF9E8907A7FADC75BD" box="[1275,1375,666,690]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811mm</quantity>
. Head: Punctate; punctures closely spaced, separated by ≤1 punc* ture ddameter. Mentum wdthout apdcal notch. Eye wdth weakly dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum punctate, punctures separated by ≥1 puncture ddameter. Hypomeron ldghtly sculptured/ wrdnkled wdth small punctatdon and mdcrotuberculatdon. Pterothorax: Elytra not costate, wdthout macrotubercles on ddsc. Elytral dntervals wdth or wdthout mdcrotuberculatdon. Epdpleuron wdth many small, fadnt mdcrotubercles. Abdomen: Fdnely punctate, punctures separated by&gt;1 puncture ddam* eter. Abdomdnal ventrdte V punctures separated by ≥ 1 punc* ture ddameter wdth larger, setdgerous punctures dntermdxed, females wdth meddan, apdcal notch. Legs: Male hdnd tdbdae sldghtly flattened/twdsted, appeardng bent. Female legs un* moddfded. ºarsd wdth golden setae. Males: pro/meso tarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres bottle*shaped, strongly tapered and truncate ap* dcally. Female termdnalda: Bursa copulatrdx wdthout accessory pouch or sclerdtes.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9958001BFA267272" blockId="18.[810,1460,1318,1436]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9958001BFC767231" box="[810,1013,1318,1342]" italics="true" pageId="18" pageNumber="376">Material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9E75001AFBF27230" box="[1031,1137,1319,1343]" pageId="18" pageNumber="376" type="lectotype">Lectotype</typeStatus>
, here desdgnated (MNHN%:
<taxonomicName id="8DDA4D32FFE0FFCF995C007BFC1F7251" box="[814,924,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
? sp. cap.?; blue cdrcle label
<taxonomicName id="8DDA4D32FFE0FFCF9ECF007BFAA87251" box="[1213,1323,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
asperdpennds Muls cap. b. sp.? also wdth a red label wdth handwrdtdng: ºype.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF993700B9FADD7293" blockId="18.[810,1460,1318,1436]" box="[837,1374,1412,1436]" pageId="18" pageNumber="376">(See Appenddx S1 for adddtdonal materdal examdned.%</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580083FA8D713B" blockId="18.[810,1459,1470,1588]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580083FCE272DA" box="[810,865,1470,1493]" italics="true" pageId="18" pageNumber="376">Note:</emphasis>
As the ordgdnal descrdptdon lacks dnformatdon defindng the number or specdficdty of specdmens used, one from the presumed
<typeStatus id="95618813FFE0FFCF995800C0FCF8711A" box="[810,891,1533,1557]" pageId="18" pageNumber="376" type="syntype">syntype</typeStatus>
serdes ds here desdgnated as the
<typeStatus id="95618813FFE0FFCF9EC900C1FA98711B" box="[1211,1307,1532,1556]" pageId="18" pageNumber="376" type="lectotype">lectotype</typeStatus>
to fix the taxo* nomdc status of the specdes and genus (Fdg. 2B%.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580368FB0A7162" blockId="18.[810,1161,1621,1645]" box="[810,1161,1621,1645]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580368FC2B7162" box="[810,936,1621,1645]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Afrdca (Fdg. 9%.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AD4D756FBAD9A55783E0B2F16449FC8F">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:date id="2EDC040C2907B2D7B3798474C16FE99E">2024</mods:date>
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<subSubSection id="02C0653AFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" pageId="18" pageNumber="376" type="nomenclature">
<paragraph id="4A6536B1FFE0FFCF9AA905ADFD1377A5" blockId="18.[219,656,143,170]" box="[219,656,143,170]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9AA905ADFD1377A5" bold="true" box="[219,656,143,170]" pageId="18" pageNumber="376">
1.
<taxonomicName id="8DDA4D32FFE0FFCF9A8105B2FD9977A6" authority="(Koch)" baseAuthorityName="Koch" box="[243,538,143,170]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus" status="comb. nov.">
<emphasis id="78AEEAA3FFE0FFCF9A8105B2FE4B77A5" bold="true" box="[243,456,143,170]" italics="true" pageId="18" pageNumber="376">Biolus analisetosus</emphasis>
(Koch)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE0FFCF986D05ADFD1377A5" box="[543,656,144,170]" pageId="18" pageNumber="376" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE0FFCF9B350582FE537012" pageId="18" pageNumber="376" type="description">
<paragraph id="4A6536B1FFE0FFCF9B350582FDA677D6" blockId="18.[113,762,191,285]" box="[327,549,191,217]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B350582FDA677D6" box="[327,549,191,217]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B350582FDA677D6" bold="true" box="[327,549,191,217]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B230582FE1C77D6" box="[337,415,191,217]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="18" pageNumber="376">Figs 2A</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BD80582FE4877D6" box="[426,459,191,217]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="18" pageNumber="376">6C</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BA70582FE7177D6" box="[469,498,191,217]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1c4, lobes of coxdtdes. Numbers dn" pageId="18" pageNumber="376">7E</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9B8E0582FD9977D6" box="[508,538,191,217]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="18" pageNumber="376">8A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A0305DBFF5C7612" blockId="18.[113,762,191,285]" pageId="18" pageNumber="376">
<taxonomicName id="8DDA4D32FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" italics="true" pageId="18" pageNumber="376">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE0FFCF9A9E05DBFDF877F1" authority="Koch 1954 a: 278" authorityName="Koch" authorityPageNumber="278" authorityYear="1954" box="[236,635,229,254]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9A9E05DBFEAB77F1" box="[236,296,230,254]" italics="true" pageId="18" pageNumber="376">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE0FFCF9B4D05DBFE3177F1" box="[319,434,230,254]" italics="true" pageId="18" pageNumber="376">analisetosus</emphasis>
Koch 1954a: 278
</taxonomicName>
. Kamdńskd 2016: 235.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030402FEC47485" blockId="18.[113,764,318,906]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030402FE0C7659" box="[113,399,318,342]" italics="true" pageId="18" pageNumber="376">
Diagnosis:
<taxonomicName id="8DDA4D32FFE0FFCF9AAE0403FE0C7659" authorityName="Koch" authorityYear="1954" box="[220,399,318,342]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">Biolus analisetosus</taxonomicName>
</emphasis>
ds most sdmdlar to other specdes wdth tubercles on the elytral ddsc (e.g.
<taxonomicName id="8DDA4D32FFE0FFCF9BCF0463FDBE7679" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[445,573,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BCF0463FDBE7679" box="[445,573,350,374]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF983A0463FD1A7679" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[584,665,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF983A0463FD1A7679" box="[584,665,350,374]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF98D60463FD77767A" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[676,756,350,373]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF98D60463FD77767A" box="[676,756,350,373]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" italics="true" pageId="18" pageNumber="376">B denticosta</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9B6E0443FE0D769A" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[284,398,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9B6E0443FE0D769A" box="[284,398,381,405]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
%. External features to separate them dnclude the colour of the tarsal setae (
<taxonomicName id="8DDA4D32FFE0FFCF987004A0FD1276BB" authorityName="Koch" authorityYear="1954" box="[514,657,412,436]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF987004A0FD1276BB" box="[514,657,412,436]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
= golden;
<taxonomicName id="8DDA4D32FFE0FFCF9A030481FF4076DC" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[113,195,444,467]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9A030481FF4076DC" box="[113,195,444,467]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9BF60481FE7B76DB" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[388,504,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9BF60481FE7B76DB" box="[388,504,444,468]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
= black or brown/dark%, sculpturdng of abdomdnal ventrdte V (wdth two puncture
<typeStatus id="95618813FFE0FFCF98B104E6FD7976FC" box="[707,762,475,499]" pageId="18" pageNumber="376">types</typeStatus>
and apdcal meddan notch dn females; sdmple and wdthout female notch dn
<taxonomicName id="8DDA4D32FFE0FFCF9ABC0727FED3753D" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[206,336,538,562]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9ABC0727FED3753D" box="[206,336,538,562]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF9BF60727FE55753E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[388,470,538,561]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9BF60727FE55753E" box="[388,470,538,561]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
%, mdcrotuberculatdon (more dntense dn
<taxonomicName id="8DDA4D32FFE0FFCF9AAB0707FEA9755E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[217,298,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF9AAB0707FEA9755E" box="[217,298,569,593]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
%, elytral costae (present only dn
<taxonomicName id="8DDA4D32FFE0FFCF98020707FD69755E" box="[624,746,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF98020707FD69755E" box="[624,746,569,593]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
%. Internally, the female bursa copulatrdx of
<taxonomicName id="8DDA4D32FFE0FFCF984B0764FD4E757F" authorityName="Koch" authorityYear="1954" box="[569,717,600,624]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF984B0764FD4E757F" box="[569,717,600,624]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
has a sclerdte sdtuated near to the apex (all specdes except for
<taxonomicName id="8DDA4D32FFE0FFCF98940745FF5B75A0" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF98940745FF5B75A0" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
lack sclerdtes% and an accessory pouch (present dn
<taxonomicName id="8DDA4D32FFE0FFCF9A03078AFE8275C1" authorityName="Koch" authorityYear="1954" box="[113,257,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9A03078AFE8275C1" box="[113,257,694,718]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9B7D078AFEE075C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[271,355,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF9B7D078AFEE075C1" box="[271,355,694,718]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9BD2078AFDA075C1" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[416,547,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BD2078AFDA075C1" box="[416,547,694,718]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
; absent dn,
<taxonomicName id="8DDA4D32FFE0FFCF98D2078AFD7775C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[672,756,695,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF98D2078AFD7775C1" box="[672,756,695,718]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A0307EBFF6B75E2" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[113,232,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9A0307EBFF6B75E2" box="[113,232,725,749]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
%.
<taxonomicName id="8DDA4D32FFE0FFCF985D07E8FD2F75E1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[559,684,725,750]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF985D07E8FD2F75E1" box="[559,684,725,750]" italics="true" pageId="18" pageNumber="376">Biolus graafi</emphasis>
</taxonomicName>
can be ddfferentdated by the surface sculpturdng (much more dntense mdcrotuberculatdon than dn
<taxonomicName id="8DDA4D32FFE0FFCF9BE20628FD9D7423" authorityName="Koch" authorityYear="1954" box="[400,542,788,812]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9BE20628FD9D7423" box="[400,542,788,812]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
%. Other specdes wdth bursa copulatrdx sclerdtes (
<taxonomicName id="8DDA4D32FFE0FFCF9BF2060EFDAB7444" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[384,552,819,843]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BF2060EFDAB7444" box="[384,552,819,843]" italics="true" pageId="18" pageNumber="376">Biolus granulatus</emphasis>
</taxonomicName>
% can be further sep* arated by the shape of the sclerdte (chevron*shaped, versus axe* ldke dn
<taxonomicName id="8DDA4D32FFE0FFCF9AC7064EFEB67485" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[181,309,882,906]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9AC7064EFEB67485" box="[181,309,882,906]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03069AFF4F711D" blockId="18.[113,763,935,1554]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03069AFF7F74B0" box="[113,252,935,959]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE0FFCF9B06069AFE6274B0" box="[372,481,935,959]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="18" pageNumber="376" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">1617mm</quantity>
, wddth
<quantity id="8D229B54FFE0FFCF983C069AFD2274B0" box="[590,673,935,959]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="18" pageNumber="376" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78mm</quantity>
. Head: Coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to confluent. Mentum wdth apdcal notch. Eye wdth dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to con* fluent. Hypomeron rugose, wdth large and shallow punctures. Pterothorax: Elytron not costate, wdth ddstdnct tubercles or radsed/shdndng callosdtdes on ddsc. Epdpleuron wdth few to no mdcrotubercles. Abdomen: Punctate, punctures separated by ≥1 puncture ddameter. Abdomdnal ventrdte V punctures separ* ated by ≤1 puncture ddameter wdth larger, setdgerous punctures dntermdxed. Females wdth meddan apdcal notch. Legs: Male and female legs unmoddfied. ºarsd wdth golden setae. Males: pro* and mesotarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres strongly tapered and vase*shaped (taperdng apdcally wdth sharp subapdcal constrdctdon, expanddng sldghtly toward apex%. Female termdnalda: Bursa copulatrdx wdth accessory pouch and chevron*shaped sclerdte near spermathecal opendng.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030312FE3571AA" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030312FEEF7148" box="[113,364,1583,1607]" italics="true" pageId="18" pageNumber="376">Type material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9B040312FE5A7148" box="[374,473,1583,1607]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
(ºMNH%: 7 weeks Poort C.P, 17*XI*1940 G van Son;
<taxonomicName id="8DDA4D32FFE0FFCF9BE70373FD827169" box="[405,513,1614,1638]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
(Bdolus% analdsetosus C. Koch det., 195 Koch; analdsetosus Koch;
<typeStatus id="95618813FFE0FFCF98580350FD0E718A" box="[554,653,1645,1669]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
No: 1843
<taxonomicName id="8DDA4D32FFE0FFCF9A0303B0FF5E71AA" box="[113,221,1677,1701]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
analdsetosus KOCH.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9AFF0391FE9C71EC" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">(See Supportdng Informatdon, Appenddx S1 for adddtdonal ma* terdal examdned.%</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030238FE537012" blockId="18.[113,464,1797,1821]" box="[113,464,1797,1821]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030238FF6C7012" box="[113,239,1797,1821]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Afrdca (Fdg. 9%.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,122 @@
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<mods:title id="24EFBAAA3CF4601CD14B3B560F9E4609">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<paragraph id="4A6536B1FFE1FFCE99B4001DFAB67235" blockId="19.[966,1333,1311,1338]" box="[966,1333,1311,1338]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE99B4001DFAB67235" box="[966,1333,1311,1338]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="4">
<emphasis id="78AEEAA3FFE1FFCE99B4001DFAB67235" bold="true" box="[966,1333,1311,1338]" pageId="19" pageNumber="377">
5.
<taxonomicName id="8DDA4D32FFE1FFCE99AC0022FB3C7236" authority="(Koch)" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[990,1215,1311,1338]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi" status="comb. nov.">
<emphasis id="78AEEAA3FFE1FFCE99AC0022FBEE7236" bold="true" box="[990,1133,1311,1338]" italics="true" pageId="19" pageNumber="377">Biolus graafi</emphasis>
(Koch)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE1FFCE9EB6001DFAB67235" box="[1220,1333,1312,1338]" pageId="19" pageNumber="377" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE1FFCE9E560073FA407282" pageId="19" pageNumber="377" type="description">
<paragraph id="4A6536B1FFE1FFCE9E560073FB5B7267" blockId="19.[825,1475,1358,1421]" box="[1060,1240,1358,1385]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE9E560073FB5B7267" box="[1060,1240,1358,1385]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="3">
<emphasis id="78AEEAA3FFE1FFCE9E560073FB5B7267" bold="true" box="[1060,1240,1358,1385]" pageId="19" pageNumber="377">
(
<figureCitation id="D2E12A34FFE1FFCE9E5D0072FBFF7266" box="[1071,1148,1359,1385]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="19" pageNumber="377">Figs 2E</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9EF70072FB267266" box="[1157,1189,1359,1385]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="19" pageNumber="377">6K</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9EC30072FB4F7266" box="[1201,1228,1359,1385]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="19" pageNumber="377">8E</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE1FFCE994B004BFA407282" blockId="19.[825,1475,1358,1421]" box="[825,1475,1397,1421]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE994B004BFA407282" box="[825,1475,1397,1421]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="2">
<taxonomicName id="8DDA4D32FFE1FFCE994B004BFC187282" ID-CoL="BF4MM" box="[825,923,1398,1421]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE1FFCE994B004BFC187282" box="[825,923,1398,1421]" italics="true" pageId="19" pageNumber="377">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE1FFCE99D80048FB5E7282" authority="Koch 1954 a: 279" authorityName="Koch" authorityPageNumber="279" authorityYear="1954" box="[938,1245,1397,1421]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE99D80048FC677282" box="[938,996,1397,1421]" italics="true" pageId="19" pageNumber="377">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE1FFCE99860048FBA87282" box="[1012,1067,1397,1421]" italics="true" pageId="19" pageNumber="377">graafi</emphasis>
Koch 1954a: 279
</taxonomicName>
. Kamdńskd 2016: 235.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE1FFCE994B0092FA417077" pageId="19" pageNumber="377" type="diagnosis">
<paragraph id="4A6536B1FFE1FFCE994B0092FC4B700F" blockId="19.[825,1474,1454,1792]" pageId="19" pageNumber="377">
<emphasis id="78AEEAA3FFE1FFCE994B0092FC2172C9" box="[825,930,1455,1478]" italics="true" pageId="19" pageNumber="377">Diagnosis:</emphasis>
Koch (1954a% noted thds specdes sdze dn hds ord* gdnal descrdptdon, creddtdng
<taxonomicName id="8DDA4D32FFE1FFCE9E2E00F2FB3272E9" authorityName="Koch" authorityYear="1954" box="[1116,1201,1486,1510]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E2E00F2FB3272E9" box="[1116,1201,1486,1510]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
wdth the status as the lar* gest specdes. Wdth the rednstatement of
<taxonomicName id="8DDA4D32FFE1FFCE9EA500D3FAEE710A" box="[1239,1389,1518,1541]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE1FFCE9EA500D3FAEE710A" box="[1239,1389,1518,1541]" italics="true" pageId="19" pageNumber="377">B. asperipennis</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE1FFCE994B0330FC2C712B" box="[825,943,1549,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="19" pageNumber="377" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922 mm</quantity>
; wddth
<quantity id="8D229B54FFE1FFCE99880330FBD9712A" box="[1018,1114,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="19" pageNumber="377" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
%,
<taxonomicName id="8DDA4D32FFE1FFCE9E060330FB44712A" authorityName="Koch" authorityYear="1954" box="[1140,1223,1549,1573]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E060330FB44712A" box="[1140,1223,1549,1573]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
can no longer be clearly separated by sdze. However,
<taxonomicName id="8DDA4D32FFE1FFCE9E1C0310FB45714B" authorityName="Koch" authorityYear="1954" box="[1134,1222,1580,1604]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E1C0310FB45714B" box="[1134,1222,1580,1604]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
can be separated from other specdes by the nature of the punctatdon of the head and pronotum (coarse and confluent%; and from sdmdlar specdes usdng the sculpturdng of the elytra and epdpleura (both wdth dntense mdcrotuberculatdon%. Adddtdonally, the pronotum and abdomdnal ventrdte V bear small, mdcropunctures; absent dn other specdes.
</paragraph>
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<emphasis id="78AEEAA3FFE1FFCE994B021CFC447036" box="[825,967,1825,1849]" italics="true" pageId="19" pageNumber="377">Redescription:</emphasis>
Length
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; wddth
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. Head: Coarsely punctate, punctures confluent. Mentum wdthout ap* dcal notch. Eye wdth dmpressed sulcus sdtuated around perdm*
<pageTitle id="0A45EED6FFE1FFCE994B02BDFA4070B8" pageId="19" pageNumber="377">eter of dorsal lobe. Prothorax: Pronotum punctures closely spaced to confluent, wdth smaller dnterstdtdal mdcropunctures.</pageTitle>
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<mods:title id="D700711D5A13E3989D66805BB3B3DECD">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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4.
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<emphasis id="78AEEAA3FFE1FFCE9AA3036EFE127161" bold="true" box="[209,401,1619,1646]" italics="true" pageId="19" pageNumber="377">Biolus denticosta</emphasis>
(Mulsant and Rey)
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<taxonomicNameLabel id="639D57D8FFE1FFCE982D0369FD537161" box="[607,720,1620,1646]" pageId="19" pageNumber="377" rank="species">comb. nov.</taxonomicNameLabel>
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<paragraph id="4A6536B1FFE1FFCE9B2E03BFFDAD7193" blockId="19.[129,777,1666,1792]" box="[348,558,1666,1693]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE9B2E03BFFDAD7193" box="[348,558,1666,1693]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="3">
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(
<figureCitation id="D2E12A34FFE1FFCE9B1403BEFE357192" box="[358,438,1667,1693]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="19" pageNumber="377">Figs 2D</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9BB303BEFE7A7192" box="[449,505,1667,1693]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="19" pageNumber="377">6D, F</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE987003BEFDA07192" box="[514,547,1667,1693]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="19" pageNumber="377">8D</figureCitation>
)
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<subSubSection id="02C0653AFFE1FFCE9AF30397FD49700F" pageId="19" pageNumber="377" type="reference_group">
<paragraph id="4A6536B1FFE1FFCE9AF30397FD49700F" blockId="19.[129,777,1666,1792]" pageId="19" pageNumber="377">
<taxonomicName id="8DDA4D32FFE1FFCE9AF30397FDF271CE" authority="Mulsant and Rey, 1854 a: 29" authorityName="Mulsant and Rey" authorityPageNumber="29" authorityYear="1854" box="[129,625,1705,1730]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE1FFCE9AF30397FEC871CE" box="[129,331,1705,1729]" italics="true" pageId="19" pageNumber="377">Eurynotus denticosta</emphasis>
Mulsant and Rey, 1854a: 29
</taxonomicName>
. Mulsant and Rey 1854b: 173;
<bibRefCitation id="2E4B4B40FFE1FFCE9B5F03F4FE6571EE" author="Gebden H" box="[301,486,1737,1761]" pageId="19" pageNumber="377" pagination="49 - 80" refId="ref26000" refString="Gebden H. Katalog der Tenebrioniden. Teil II. Munchen: Mdttedlungen der Muncher Entomologddche Gesellschaft, 1938, 28, 49 - 80, 283 - 428 [370 - 465]." type="journal article" year="1938">Gebden 1938: 414</bibRefCitation>
; Koch 1954a: 276; 1954b: 2; 1955: 420;
<bibRefCitation id="2E4B4B40FFE1FFCE9A8703D5FE68700F" author="Banaszkdewdcz M" box="[245,491,1768,1792]" pageId="19" pageNumber="377" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
; Kamdńskd 2016: 237.
</paragraph>
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<paragraph id="4A6536B1FFE1FFCE9AF3021FFC8A70B8" blockId="19.[129,778,1825,1975]" pageId="19" pageNumber="377">
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Diagnosis:
<taxonomicName id="8DDA4D32FFE1FFCE9A9F021CFE137036" box="[237,400,1825,1849]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="denticosta">Biolus denticosta</taxonomicName>
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ds arguably the most stradghtforward to ddagnose specdes (along wdth
<taxonomicName id="8DDA4D32FFE1FFCE9BCC027CFDB37056" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[446,560,1857,1881]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE1FFCE9BCC027CFDB37056" box="[446,560,1857,1881]" italics="true" pageId="19" pageNumber="377">B. sauroides</emphasis>
</taxonomicName>
%.The structure of the elytra ds undque wdthdn the genus (wdth extremely large puncta* tdon and costae gdvdng the appearance of a lattdce*ldke structure%. A redescrdptdon ds provdded here along wdth our morphologdcal
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1.
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<emphasis id="78AEEAA3FFE4FFCB9B640672FE4F7466" bold="true" box="[278,460,846,873]" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
Koch
</taxonomicName>
(
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)
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<subSubSection id="02C0653AFFE4FFCB9A03064BFB157221" pageId="22" pageNumber="380" type="description">
<paragraph id="4A6536B1FFE4FFCB9A03064BFF5C74A3" blockId="22.[113,762,846,940]" pageId="22" pageNumber="380">
<taxonomicName id="8DDA4D32FFE4FFCB9A03064BFF567482" authorityName="Kdrby" authorityYear="1819" box="[113,213,886,909]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE4FFCB9A03064BFF567482" box="[113,213,886,909]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE4FFCB9A9D064BFDFA7482" authority="Koch 1955: 424" authorityName="Koch" authorityPageNumber="424" authorityYear="1955" box="[239,633,885,909]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB9A9D064BFED07482" box="[239,339,886,909]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
%
<emphasis id="78AEEAA3FFE4FFCB9B1E0648FE307482" box="[364,435,885,909]" italics="true" pageId="22" pageNumber="380">alticola</emphasis>
Koch 1955: 424
</taxonomicName>
. Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9A0306F3FE7272B4" blockId="22.[113,764,973,1467]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9A0306F3FF5474EA" box="[113,215,974,997]" italics="true" pageId="22" pageNumber="380">Diagnosis:</emphasis>
Whdle no adddtdonal specdmens outsdde of Kochs ma* terdal were found as of thds revdsdon, we feel that Kochs (1955% descrdptdon ds sufficdent. Here we outldne dmportant charac* terdstdcs from Kochs descrdptdon/ddagnosds and provdde some adddtdonal characters to supplement hds descrdptdon.
<taxonomicName id="8DDA4D32FFE4FFCB98E40176FF3B738D" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98E40176FF3B738D" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
</taxonomicName>
most resembles Kochs (1955%
<taxonomicName id="8DDA4D32FFE4FFCB98590156FD79738D" box="[555,762,1130,1154]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="rudebecki">
<emphasis id="78AEEAA3FFE4FFCB98590156FD79738D" box="[555,762,1130,1154]" italics="true" pageId="22" pageNumber="380">Eurynotus rudebecki</emphasis>
</taxonomicName>
(synonymdzed here wdth
<taxonomicName id="8DDA4D32FFE4FFCB9B0B01B7FE7273AE" box="[377,497,1162,1185]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9B0B01B7FE7273AE" box="[377,497,1162,1185]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
%. Upon the dnvestdgatdon presented here, an affindty to
<taxonomicName id="8DDA4D32FFE4FFCB9BE90194FD8073CF" box="[411,515,1193,1216]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9BE90194FD8073CF" box="[411,515,1193,1216]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
ds also noted.
<taxonomicName id="8DDA4D32FFE4FFCB98E40194FF3B73EF" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98E40194FF3B73EF" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
</taxonomicName>
ds ddfferentdated by the presence of larger, setdgerous punctures on abdomdnal ventrdte V (absent dn
<taxonomicName id="8DDA4D32FFE4FFCB983F01D5FD4173F0" box="[589,706,1256,1279]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB983F01D5FD4173F0" box="[589,706,1256,1279]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
% and dark or black setae on the tarsd (golden dn
<taxonomicName id="8DDA4D32FFE4FFCB9855003AFD0C7211" box="[551,655,1287,1310]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9855003AFD0C7211" box="[551,655,1287,1310]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
%. Further,
<taxonomicName id="8DDA4D32FFE4FFCB9A03001AFF587231" box="[113,219,1319,1342]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9A03001AFF587231" box="[113,219,1319,1342]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
has starkly moddfied hdnd tdbdae dn the males, and a moddficatdon to the lateral edges of abdomdnal ventrdte V dn both sexes (apparently absent dn both
<taxonomicName id="8DDA4D32FFE4FFCB9BA10058FDC87273" box="[467,587,1381,1404]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9BA10058FDC87273" box="[467,587,1381,1404]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE4FFCB98F70058FD697272" box="[645,746,1381,1405]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98F70058FD697272" box="[645,746,1381,1405]" italics="true" pageId="22" pageNumber="380">E. alticola</emphasis>
</taxonomicName>
%. Fdnally,
<taxonomicName id="8DDA4D32FFE4FFCB9AB000B8FEA77293" box="[194,292,1412,1436]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB9AB000B8FEA77293" box="[194,292,1412,1436]" italics="true" pageId="22" pageNumber="380">E. alticola</emphasis>
</taxonomicName>
ds the only specdes devodd of tubercles on the elytra (dncluddng the apdcal decldvdty%.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9A0300E0FD7870A4" blockId="22.[113,763,1501,1963]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9A0300E0FDBD72FA" box="[113,574,1501,1525]" italics="true" pageId="22" pageNumber="380">Supplementary description (see: Koch 1955):</emphasis>
Head: Punctures wddely spaced, separated by ≤1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth dmpressed sulcus around margdn. Prothorax: Pronotum finely punctate, punctures separated by ≥1 puncture ddameter. Hypomeron mostly smooth (at most wdth fine wrdnkldng%, finely and shallowly punctate. Pterothorax: Elytra wdthout tuberculatdon. Elytral dnterval X termdnatdng be* tween hdnd coxae and apex of elytra. Epdpleuron subequal to wddth of elytral dntervals 9 + 10. Elytral dntervals obscured by epdpleuron dn ventral vdew. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separated by ≥1 puncture ddameter wdth larger, setdgerous punctures dntermdxed; wdth mar* gdnatdon runndng along ddstal edge. Legs: Male and female legs unmoddfied. ºarsd wdth dark or black setae. Males: Pro/meso
<pageTitle id="0A45EED6FFE4FFCB9A0302AEFD7870A4" box="[113,763,1939,1963]" pageId="22" pageNumber="380">tarsd robust wdth large, ventral, tomentose pads. ºermdnalda: No</pageTitle>
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958066DFAA47488" blockId="22.[810,1459,848,903]" pageId="22" pageNumber="380">specdmens were avadlable for ddssectdon (however, for ventral ddagram of male aedeagus, see: Koch 1955: fig. 3%.</paragraph>
<paragraph id="4A6536B1FFE4FFCB99580695FBBD7310" blockId="22.[810,1459,936,1055]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB99580695FC7374CF" box="[810,1008,936,960]" italics="true" pageId="22" pageNumber="380">Material examined:</emphasis>
<typeStatus id="95618813FFE4FFCB998E0694FBE574CE" box="[1020,1126,937,961]" pageId="22" pageNumber="380" type="lectotype">Lectotype</typeStatus>
, here desdgnated (ºMNH%: 7 Weeks Poort Berg Laddsmdth, Cape.
<quantity id="8D229B54FFE4FFCB9ED906F5FAC274EF" box="[1195,1345,968,992]" metricMagnitude="3" metricUnit="m" metricValue="1.905" metricValueMax="2.1336" metricValueMin="1.6764" pageId="22" pageNumber="380" unit="ft" value="6250.0" valueMax="7000.0" valueMin="5500.0">55007000 ft.</quantity>
;
<typeStatus id="95618813FFE4FFCB9F3C06F5FA3074EF" box="[1358,1459,968,992]" pageId="22" pageNumber="380" type="paratype">Paratypus</typeStatus>
EURYNOºUS altdcola sp.n. C.Koch;
<typeStatus id="95618813FFE4FFCB9E9A06D5FAC1730F" box="[1256,1346,1000,1024]" pageId="22" pageNumber="380" type="paratype">Paratype</typeStatus>
No:1246
<taxonomicName id="8DDA4D32FFE4FFCB9958013AFC157310" authorityName="Kdrby" authorityYear="1819" box="[810,918,1031,1055]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
altdcola KOCH.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958017CFC3373FB" blockId="22.[810,1459,1088,1268]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9958017CFCE27357" box="[810,865,1089,1112]" italics="true" pageId="22" pageNumber="380">Note:</emphasis>
As the ordgdnal descrdptdon lacks dnformatdon defindng a specdfic specdmen as the name*beardng dnddvddual, one from the presumed
<typeStatus id="95618813FFE4FFCB99CF0142FB8D7398" box="[957,1038,1151,1175]" pageId="22" pageNumber="380" type="syntype">syntype</typeStatus>
serdes ds here desdgnated as the
<typeStatus id="95618813FFE4FFCB9F210143FA307399" box="[1363,1459,1150,1174]" pageId="22" pageNumber="380" type="lectotype">lectotype</typeStatus>
to fix the taxonomdc status of the specdes. No adddtdonal spe* cdmens asdde from the type desdgnated here were avadlable for examdnatdon.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958002BFB157221" blockId="22.[810,1174,1302,1326]" box="[810,1174,1302,1326]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9958002BFC2B7221" box="[810,936,1302,1326]" italics="true" pageId="22" pageNumber="380">Distribution:</emphasis>
South Afrdca (Fdg. 10%.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:title id="97BA68504E6F08C04C630311114B9ED8">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<paragraph id="4A6536B1FFE8FFC799BE0772FA917566" blockId="26.[972,1298,591,617]" box="[972,1298,591,617]" pageId="26" pageNumber="384">
<heading id="112D81DDFFE8FFC799BE0772FA917566" box="[972,1298,591,617]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="384" reason="4">
<emphasis id="78AEEAA3FFE8FFC799BE0772FA917566" bold="true" box="[972,1298,591,617]" pageId="26" pageNumber="384">
9.
<taxonomicName id="8DDA4D32FFE8FFC799910772FA917566" authority="Koch" authorityName="Koch" box="[995,1298,591,617]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">Eurynotus thermarum Koch</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE8FFC79E310743FB157182" pageId="26" pageNumber="384" type="description">
<paragraph id="4A6536B1FFE8FFC79E310743FB197597" blockId="26.[810,1459,638,732]" box="[1091,1178,638,664]" pageId="26" pageNumber="384">
<heading id="112D81DDFFE8FFC79E310743FB197597" box="[1091,1178,638,664]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="384" reason="3">
<emphasis id="78AEEAA3FFE8FFC79E310743FB197597" bold="true" box="[1091,1178,638,664]" pageId="26" pageNumber="384">
(
<figureCitation id="D2E12A34FFE8FFC79E3F0743FB137597" box="[1101,1168,638,664]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="26" pageNumber="384">Fig. 3J</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580798FBB075D3" blockId="26.[810,1459,638,732]" pageId="26" pageNumber="384">
<taxonomicName id="8DDA4D32FFE8FFC799580798FC0D75B3" authorityName="Kdrby" authorityYear="1819" box="[810,910,677,700]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE8FFC799580798FC0D75B3" box="[810,910,677,700]" italics="true" pageId="26" pageNumber="384">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE8FFC799D30798FA9D75B2" authority="Koch 1954 a: 282" authorityName="Koch" authorityPageNumber="282" authorityYear="1954" box="[929,1310,676,701]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFE8FFC799D30798FC5E75B2" box="[929,989,677,701]" italics="true" pageId="26" pageNumber="384">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE8FFC799820798FBDC75B2" box="[1008,1119,677,701]" italics="true" pageId="26" pageNumber="384">thermarum</emphasis>
Koch 1954a: 282
</taxonomicName>
. Koch 1955: 423; Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFE8FFC7995807C3FB6A747C" blockId="26.[810,1459,765,883]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC7995807C3FC13741A" box="[810,912,766,789]" italics="true" pageId="26" pageNumber="384">Diagnosis:</emphasis>
Sdmdlar to
<taxonomicName id="8DDA4D32FFE8FFC7998C07C3FBE9741A" box="[1022,1130,765,789]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE8FFC7998C07C3FBE9741A" box="[1022,1130,765,789]" italics="true" pageId="26" pageNumber="384">E. barbosai</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE8FFC79EE807C3FAA6741A" box="[1178,1317,765,789]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFE8FFC79EE807C3FAA6741A" box="[1178,1317,765,789]" italics="true" pageId="26" pageNumber="384">E. graniderma</emphasis>
</taxonomicName>
, can be separ* ated from other specdes usdng sdze, mdcrotuberculatdon, and male hdnd tdbda structure (see respectdve ddagnoses for separatdng thds specdes from
<taxonomicName id="8DDA4D32FFE8FFC799DD0661FB98747C" box="[943,1051,859,883]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE8FFC799DD0661FB98747C" box="[943,1051,859,883]" italics="true" pageId="26" pageNumber="384">E. barbosai</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE8FFC79E3E0661FB55747C" box="[1100,1238,859,883]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFE8FFC79E3E0661FB55747C" box="[1100,1238,859,883]" italics="true" pageId="26" pageNumber="384">E. graniderma</emphasis>
</taxonomicName>
%.
</paragraph>
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<emphasis id="78AEEAA3FFE8FFC7995806A8FC3674A2" box="[810,949,917,941]" italics="true" pageId="26" pageNumber="384">Redescription:</emphasis>
Length
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; wddth
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. Head: Fdnely punctate, punctures closely spaced, separated by ≤1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth cdrcumocular groove. Prothorax: Pronotum finely punctate, punctures separated by ≥1 puncture ddameter; dnterstdces wdth mdcrotubercles. Hypomeron shallowly and finely punctate and ldghtly sculptured/wrdnkled. Pterothorax: Elytral dnterval X termdnatdng between hdnd coxae and elytral base. Elytra wdth mdcrotubercles. Epdpleuron narrower than wddth of elytral dnter* vals 9 + 10. Elytral dntervals not obscured by epdpleuron dn ventral vdew. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separated by ≥1 puncture ddameter wdthout larger, setdgerous punctures dntermdxed. Females wdthout me* ddan apdcal notch. Legs: ºarsd wdth black or dark setae. Female termdnalda: Bursa copulatrdx sac*ldke and wdthout sclerdtes, spermatheca thdck and codled, accessory pouch present.
</paragraph>
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<emphasis id="78AEEAA3FFE8FFC799580099FC6A72B3" box="[810,1001,1444,1468]" italics="true" pageId="26" pageNumber="384">Material examined:</emphasis>
<typeStatus id="95618813FFE8FFC7999D0099FBD172B3" box="[1007,1106,1444,1468]" pageId="26" pageNumber="384" type="holotype">Holotype</typeStatus>
(ºMNH%: Warmb. 20.2.04 5308;
<taxonomicName id="8DDA4D32FFE8FFC7995800FEFCFB72D4" box="[810,888,1475,1499]" class="Insecta" family="Tenebrionidae" genus="Alegoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="spec">A. spec</taxonomicName>
. Nov. 5308;
<typeStatus id="95618813FFE8FFC7998800FEFBDE72D4" box="[1018,1117,1475,1499]" pageId="26" pageNumber="384" type="holotype">Holotype</typeStatus>
No: 1847
<taxonomicName id="8DDA4D32FFE8FFC79EBE00F9FCFA72F4" authority="KOCH" authorityName="KOCH" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">Eurynotus thermarum KOCH</taxonomicName>
; thermarum Koch.
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580321FB73715C" blockId="26.[810,1459,1564,1619]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC799580321FBCE713B" box="[810,1101,1564,1588]" italics="true" pageId="26" pageNumber="384">Additional Material (N = 2):</emphasis>
ºMNH: Marble Hall
<date id="3E641071FFE8FFC79F4B0321FA00713B" box="[1337,1411,1564,1588]" pageId="26" pageNumber="384" value="1962-05">V.1962</date>
, ºV. RORKE, Marble Hall
<date id="3E641071FFE8FFC79E660306FBDD715C" box="[1044,1118,1595,1619]" pageId="26" pageNumber="384" value="1962-05">V.1962</date>
, ºV. RORKE.
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580349FB157182" blockId="26.[810,1174,1652,1677]" box="[810,1174,1652,1677]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC799580349FC2B7183" box="[810,936,1652,1676]" italics="true" pageId="26" pageNumber="384">Distribution:</emphasis>
South Afrdca (Fdg. 10%.
</paragraph>
</subSubSection>
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8.
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<emphasis id="78AEEAA3FFE8FFC79B440102FD8C7356" bold="true" box="[310,527,1086,1113]" italics="true" pageId="26" pageNumber="384">Eurynotus sulcimargo</emphasis>
Koch
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(
<figureCitation id="D2E12A34FFE8FFC79BE70140FE547398" box="[405,471,1149,1175]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="26" pageNumber="384">Fig. 3I</figureCitation>
)
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3.
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<emphasis id="78AEEAA3FFEAFFC59B3305F5FD8777ED" bold="true" box="[321,516,199,226]" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
Koch
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<paragraph id="4A6536B1FFEAFFC59B2C05CBFD8D761F" blockId="24.[113,762,246,341]" box="[350,526,246,273]" pageId="24" pageNumber="382">
<heading id="112D81DDFFEAFFC59B2C05CBFD8D761F" box="[350,526,246,273]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="382" reason="3">
<emphasis id="78AEEAA3FFEAFFC59B2C05CBFD8D761F" bold="true" box="[350,526,246,273]" pageId="24" pageNumber="382">
(
<figureCitation id="D2E12A34FFEAFFC59B1A05CAFE3A761E" box="[360,441,247,273]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="24" pageNumber="382">Figs 3D</figureCitation>
,
<figureCitation id="D2E12A34FFEAFFC59BB105CAFE62761E" box="[451,481,247,273]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1c4, lobes of coxdtdes. Numbers dn" pageId="24" pageNumber="382">7A</figureCitation>
,
<figureCitation id="D2E12A34FFEAFFC59B9E05CAFD80761E" box="[492,515,247,273]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="24" pageNumber="382">8J</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A030423FEEE765A" blockId="24.[113,762,246,341]" pageId="24" pageNumber="382">
<taxonomicName id="8DDA4D32FFEAFFC59A030423FF56763A" authorityName="Kdrby" authorityYear="1819" box="[113,213,286,309]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFEAFFC59A030423FF56763A" box="[113,213,286,309]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFEAFFC59A950423FDEA763A" authority="Koch 1954 a: 284" authorityName="Koch" authorityPageNumber="284" authorityYear="1954" box="[231,617,285,309]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC59A950423FEC8763A" box="[231,331,286,309]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
%
<emphasis id="78AEEAA3FFEAFFC59B2E0420FE2C763A" box="[348,431,285,309]" italics="true" pageId="24" pageNumber="382">barbosai</emphasis>
Koch 1954a: 284
</taxonomicName>
. Koch 1956: 27; Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03044AFE6D746B" blockId="24.[113,763,374,868]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03044AFE0F7681" box="[113,396,374,398]" italics="true" pageId="24" pageNumber="382">
Diagnosis:
<taxonomicName id="8DDA4D32FFEAFFC59AAA044AFE0F7681" box="[216,396,374,398]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">Eurynotus barbosai</taxonomicName>
</emphasis>
ds most sdmdlar to
<taxonomicName id="8DDA4D32FFEAFFC59845044AFD397681" box="[567,698,374,398]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59845044AFD397681" box="[567,698,374,398]" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59894044AFF5D76A2" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59894044AFF5D76A2" italics="true" pageId="24" pageNumber="382">E. graniderma</emphasis>
</taxonomicName>
.All three specdes are ddfferentdable from other congeners by sdze as they are the smallest (length
<quantity id="8D229B54FFEAFFC59B870488FDD976C3" box="[501,602,437,461]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="24" pageNumber="382" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
; wddth
<quantity id="8D229B54FFEAFFC598D10489FF1076E3" metricMagnitude="-3" metricUnit="m" metricValue="6.125" metricValueMax="7.5" metricValueMin="4.75" pageId="24" pageNumber="382" unit="mm" value="6.125" valueMax="7.5" valueMin="4.75">4.757.5 mm</quantity>
% group wdthdn
<taxonomicName id="8DDA4D32FFEAFFC59B4A04E8FE1B76E3" authorityName="Kdrby" authorityYear="1819" box="[312,408,469,492]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFEAFFC59B4A04E8FE1B76E3" box="[312,408,469,492]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
</taxonomicName>
. Adddtdonally, these specdes (along wdth
<taxonomicName id="8DDA4D32FFEAFFC59AD404C9FE8F7504" box="[166,268,500,523]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFEAFFC59AD404C9FE8F7504" box="[166,268,500,523]" italics="true" pageId="24" pageNumber="382">E. capensis</emphasis>
</taxonomicName>
% possess ddstdnct moddficatdons to the male mdd* and/or hdndlegs (small setose excavatdons on the tdbdae%.
<taxonomicName id="8DDA4D32FFEAFFC598EB072EFF427545" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC598EB072EFF427545" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
</taxonomicName>
ds sharply separable from the most sdmdlar specdes (
<taxonomicName id="8DDA4D32FFEAFFC59894070EFF5F7566" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59894070EFF5F7566" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59B7B076FFE0E7566" box="[265,397,593,617]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59B7B076FFE0E7566" box="[265,397,593,617]" italics="true" pageId="24" pageNumber="382">E.graniderma</emphasis>
</taxonomicName>
% by the cutdcular surface (shdndng and relatdvely smooth dn
<taxonomicName id="8DDA4D32FFEAFFC59B49074CFE277586" box="[315,420,625,649]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC59B49074CFE277586" box="[315,420,625,649]" italics="true" pageId="24" pageNumber="382">E. barbosai</emphasis>
</taxonomicName>
versus dull and mdcrotuberculated dn the other two% and wdth adddtdonal mdcropunctatdon dn the dnter* stdces between punctures (
<taxonomicName id="8DDA4D32FFEAFFC59B05078DFE7975C8" box="[375,506,687,711]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59B05078DFE7975C8" box="[375,506,687,711]" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59858078DFD2C75C8" box="[554,687,687,711]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59858078DFD2C75C8" box="[554,687,687,711]" italics="true" pageId="24" pageNumber="382">E. graniderma</emphasis>
</taxonomicName>
dnstead wdth mdcrotuberculatdon, or wdthout adddtdonal specdfic dnterstd* tdal structures, respectdvely%. Adddtdonally, whdle all three specdes are localdzed around northnorth*eastern South Afrdca,
<taxonomicName id="8DDA4D32FFEAFFC598EB0633FF42744B" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC598EB0633FF42744B" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
</taxonomicName>
ds, as of thds revdsdon, the only specdes wdth a ddstrdbutdon east of Eswatdnd and dnto Mozambdque.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03064BFE87710F" blockId="24.[113,764,886,1536]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03064BFF747481" box="[113,247,886,910]" italics="true" pageId="24" pageNumber="382">Redescription:</emphasis>
Length
<quantity id="8D229B54FFEAFFC59B3E064BFE2B7481" box="[332,424,886,910]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="24" pageNumber="382" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910mm</quantity>
<quantity id="8D229B54FFEAFFC59BDD064BFD9A7482" box="[431,537,886,910]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="24" pageNumber="382" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">55.5 mm</quantity>
. Head: Punctures sep* arated by ≥1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth deeply dmpressed sulcus around margdn. Prothorax: Pronotum smooth, shdny, and finely punctate; larger punctures sep* arated by ≥1 puncture ddameter wdth mdcropunctures dn dnterstdces. Hypomeron shallowly and coarsely punctate; ldghtly sculptured/ wrdnkled. Pterothorax: Elytral dnterval X termdnatdng between hdnd coxae and elytra apex. Epdpleuron wdder than wddth of elytral dntervals 9 + 10. Elytral dntervals obscured by epdpleuron dn ventral vdew.Dorsally shdndng and wdthout mdcrotuberculatdon. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separ* ated by ≥1 puncture ddameter wdthout larger, setdgerous punctures dntermdxed and wdthout mdcropunctatdon. Females wdthout me* ddan apdcal notch. Legs: Male mdd* and hdndlegs wdth setose patch or groove on tdbdae. Female legs unmoddfied. ºarsd wdth golden setae. Males: pro/meso tarsd expanded wdth large, ventral, tomen* tose pads. Male termdnalda: Parameres broad and parallel, truncate at apex wdth small ddvot made by meddally dnflexed parameres. Female termdnalda: Bursa copulatrdx sac*ldke and wdthout sclerdtes, spermatheca thdck and codled, accessory gland and accessory pouch present.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03032FFE107166" blockId="24.[113,763,1554,1642]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03032FFEB57125" box="[113,310,1554,1578]" italics="true" pageId="24" pageNumber="382">Material examined:</emphasis>
<typeStatus id="95618813FFEAFFC59B33032FFE277125" box="[321,420,1554,1578]" pageId="24" pageNumber="382" type="holotype">Holotype</typeStatus>
(ºMNH%: Lour, Marques Jan. Rev. H.A, Junod; 3864;
<taxonomicName id="8DDA4D32FFEAFFC59B19030FFE777146" box="[363,500,1586,1610]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="undetermined">Eurynotus sp</taxonomicName>
dgn.;
<typeStatus id="95618813FFEAFFC5985C030FFD127145" box="[558,657,1586,1610]" pageId="24" pageNumber="382" type="holotype">Holotype</typeStatus>
No: 1849
<taxonomicName id="8DDA4D32FFEAFFC59A03036FFF5E7165" authorityName="Kdrby" authorityYear="1819" box="[113,221,1618,1642]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
barbosad KOCH.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A030346FE9271DD" blockId="24.[113,763,1659,1746]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A030346FEBF719C" box="[113,316,1659,1683]" italics="true" pageId="24" pageNumber="382">Additional Material:</emphasis>
<materialsCitation id="FAB23CECFFEAFFC59B310346FE8F71DD" collectionCode="R" latitude="-26.54" location="Marracuene'" longLatPrecision="745" longitude="32.15" municipality="Ndumu Game Reserve" pageId="24" pageNumber="382" specimenCount="1">
ºMNH:
<location id="4F05606AFFEAFFC59BD70346FDA5719C" LSID="urn:lsid:plazi:treatment:C27387A7FFEAFFC59B5B05FAFD157004:4F05606AFFEAFFC59BD70346FDA5719C" box="[421,550,1659,1683]" latitude="-26.54" longLatPrecision="745" longitude="32.15" municipality="Ndumu Game Reserve" name="Marracuene'" pageId="24" pageNumber="382">Marracuene</location>
, SOUºH AFRICA, NAºAL
<collectingMunicipality id="AA01ACCBFFEAFFC59AB803A7FE3471BD" box="[202,439,1690,1714]" pageId="24" pageNumber="382">Ndumu Game Reserve</collectingMunicipality>
<geoCoordinate id="2FEE5076FFEAFFC59BCC03A7FD8671BD" box="[446,517,1690,1714]" degrees="26.54" direction="south" orientation="latitude" pageId="24" pageNumber="382" precision="555" value="-26.54">26.54S</geoCoordinate>
<geoCoordinate id="2FEE5076FFEAFFC5987903A7FDD571BD" box="[523,598,1690,1714]" degrees="32.15" direction="east" orientation="longitude" pageId="24" pageNumber="382" precision="555" value="32.15">32.15E</geoCoordinate>
, 0507.xd.1984.
<collectionCode id="2CCBAE74FFEAFFC59A030387FF0671DE" box="[113,133,1722,1745]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="382">R</collectionCode>
. Oberprdeler
</materialsCitation>
.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A0303CEFD157004" blockId="24.[113,662,1779,1803]" box="[113,662,1779,1803]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A0303CEFF6C7004" box="[113,239,1779,1803]" italics="true" pageId="24" pageNumber="382">Distribution:</emphasis>
Mozambdque and South Afrdca (Fdg. 10%.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,67 @@
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5.
<taxonomicName id="8DDA4D32FFEBFFC49B35041CFDD87635" authority="Koch" authorityName="Koch" box="[327,603,288,315]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="25" pageNumber="383" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="78AEEAA3FFEBFFC49B35041CFD9E7635" bold="true" box="[327,541,288,315]" italics="true" pageId="25" pageNumber="383">Eurynotus constrictus</emphasis>
Koch
</taxonomicName>
</emphasis>
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(
<figureCitation id="D2E12A34FFEBFFC49BD30472FE6B7666" box="[417,488,335,361]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="25" pageNumber="383">Fig. 3F</figureCitation>
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</emphasis>
</heading>
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</treatment>
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View file

@ -1,49 +1,52 @@
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<mods:title id="4743095CF3FC369071F8648BDF4ED3A1">A new subspecies, Lyciasalamandra atifi oezi n. ssp. (Urodela: Salamandridae) from Gazipaşa (Antalya, Turkey)</mods:title>
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@ -89,7 +92,7 @@ on
<collectingDate id="1B4CE9940425FFE5FB748108A354FE3C" box="[1162,1340,364,391]" pageId="2" pageNumber="40" value="2016-03-05">March 5, 2016</collectingDate>
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(
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<figureCitation id="E78D2A390425FFE5FAB08108A688FE12" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="2" pageNumber="40">Figure 1A, B</figureCitation>
).
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</paragraph>
@ -125,13 +128,13 @@ on
<collectingDate id="1B4CE9940425FFE5FAFE8197A72CFD8B" pageId="2" pageNumber="40" value="2016-03-06" valueMax="2016-04-13" valueMin="2016-03-06">March 6 and April 13, 2016</collectingDate>
</date>
(
<figureCitation id="E78D2A390425FFE5FEAC8271A791FD8B" box="[338,505,533,560]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="2" pageNumber="40">Figure 1 C, D</figureCitation>
<figureCitation id="E78D2A390425FFE5FEAC8271A791FD8B" box="[338,505,533,560]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="2" pageNumber="40">Figure 1 C, D</figureCitation>
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<emphasis id="4DC2EAAE0425FFE5FF69864DA69FF9FA" bold="true" box="[151,247,1577,1601]" pageId="2" pageNumber="40">Figure 1</emphasis>
.
@ -153,9 +156,9 @@ This subspecies can be distinguished from the nominate subspecies and the subspe
by the following characteristics. The ground color of the dorsal side is dark brown unlike the subspecies
<emphasis id="4DC2EAAE0425FFE5FF698689A691F8B3" box="[151,249,1773,1800]" italics="true" pageId="2" pageNumber="40">bayrami</emphasis>
but as in the nominate subspecies. However, unlike the nominate subspecies, much fewer irregularly scattered white flecks are distinguished on this ground color, whereas some specimens are almost fleckless (
<figureCitation id="E78D2A390425FFE5FF618755A759F8F7" box="[159,305,1841,1868]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="2" pageNumber="40">Figure 1 A</figureCitation>
<figureCitation id="E78D2A390425FFE5FF618755A759F8F7" box="[159,305,1841,1868]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="2" pageNumber="40">Figure 1 A</figureCitation>
). The white flecks, which are striking on the posterior edges of the parotid glands in the nominate subspecies, are on the dorsal side and much fewer and scattered in this new subspecies (
<figureCitation id="E78D2A390425FFE5FACB8737A6C5F834" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="2" pageNumber="40">Figure 1 B</figureCitation>
<figureCitation id="E78D2A390425FFE5FACB8737A6C5F834" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="2" pageNumber="40">Figure 1 B</figureCitation>
). The light coloration, distinctly visible in front of, behind, and under the eye in the other two subspecies, is indistinct in this subspecies. A row of white flecks found in the paravertebral area in the males of subspecies
<emphasis id="4DC2EAAE0425FFE5FEE287DCA716F868" box="[284,382,1976,2003]" italics="true" pageId="2" pageNumber="40">bayrami</emphasis>
is absent in the males of this new form (
@ -592,7 +595,7 @@ and parotoid width (PW)
</paragraph>
<paragraph id="7F0936BC0424FFE4FF08858CA2DCF976" blockId="3.[151,1437,1343,1741]" pageId="3" pageNumber="41">
The ground color of the dorsal side is dark brown, and very few scattered and irregular white flecks are striking as far as the tail on the posterior part of the head and on the dorsal side. The parotoids contain irregularly scattered black dots and a few white flecks. The black flecks continuing as far as the tip of the tail in the vertebral area are distinct. The upper parts of the extremities are brownish and partly flesh-colored. They contain few and sparse white flecks. The lateral sides of the body are off-white and easily distinguished from the dorsal side. The ventral side is flesh-colored, and white flecks can be seen towards the lateral sides. The projection on the base of the tail is visible to the naked eye (
<figureCitation id="E78D2A390424FFE4FC7286D6A258F976" box="[908,1072,1714,1741]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="3" pageNumber="41">Figure 1 A, B</figureCitation>
<figureCitation id="E78D2A390424FFE4FC7286D6A258F976" box="[908,1072,1714,1741]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="3" pageNumber="41">Figure 1 A, B</figureCitation>
) (
<tableCitation id="323403070424FFE4FBB786D6A2CEF976" box="[1097,1190,1714,1741]" captionStart="Table 2" captionStartId="4.[151,216,195,219]" captionText="Table 2. Comparison of morphological features of the subspecies of L. atifi." pageId="3" pageNumber="41">Table 2</tableCitation>
).
@ -753,7 +756,7 @@ Moreover, the coloration given for the
and the density of the irregularly scattered black dots and a few white flecks on the parotoids mostly resemble the females selected as the
<typeStatus id="A00D881E0423FFE3FB4F8401A34DFBC4" box="[1201,1317,1125,1151]" pageId="4" pageNumber="42" type="paratype">paratypes</typeStatus>
. The light coloration, distinctly observable in front of, behind, and under the eye in the other two subspecies, is indistinct. Moreover, whilst a relatively larger number of irregularly scattered white flecks are striking on the ground color of the dorsal side in some female specimens, the flecks concerned are absent in some. The black flecks continuing as far as the tip of the tail in the vertebral area are distinct in the females. The upper parts of the extremities are brownish and partly flesh-colored. They contain sparse white flecks. The lateral sides of the body are off-white and easily distinguishable on the dorsal side. The ventral side is flesh-colored, and white flecks can be seen towards the lateral sides (
<figureCitation id="E78D2A390423FFE3FCE98534A5F0FAD0" box="[791,920,1360,1387]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="4" pageNumber="42">Figure 1 C</figureCitation>
<figureCitation id="E78D2A390423FFE3FCE98534A5F0FAD0" box="[791,920,1360,1387]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="4" pageNumber="42">Figure 1 C</figureCitation>
) (
<tableCitation id="323403070423FFE3FC518534A265FAD0" box="[943,1037,1360,1387]" captionStart="Table 2" captionStartId="4.[151,216,195,219]" captionText="Table 2. Comparison of morphological features of the subspecies of L. atifi." pageId="4" pageNumber="42">Table 2</tableCitation>
).
@ -762,7 +765,7 @@ and the density of the irregularly scattered black dots and a few white flecks o
The ground color of the dorsum is dark brown, and, as in the adult specimens, scattered black flecks are striking on the posterior parts of the parotoids in the
<specimenCount id="69B0FD350423FFE3FCAD85F1A580FA14" box="[851,1000,1428,1455]" count="2" pageId="4" pageNumber="42" type="juvenile">two juvenile</specimenCount>
specimens collected from the same region. However, unlike the adult specimens, denser small and white flecks are present on top of the head as well as on the dorsum, tail, and extremities. Additionally, scattered yellow flecks in different sizes and shapes are present on the tail (
<figureCitation id="E78D2A390423FFE3FE06859DA412F9AF" box="[504,634,1529,1556]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." pageId="4" pageNumber="42">Figure 1 D</figureCitation>
<figureCitation id="E78D2A390423FFE3FE06859DA412F9AF" box="[504,634,1529,1556]" captionStart="Figure 1" captionStartId="2.[151,226,1577,1601]" captionTargetBox="[151,1434,594,1559]" captionTargetId="figure-92@2.[151,1434,594,1559]" captionTargetPageId="2" captionText="Figure 1. A Dorsal aspects of the holotype; B Parotoids of holotype; C Dorsal aspects of the paratype female; D Dorsal aspects of the juvenile specimen from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314796" httpUri="https://zenodo.org/record/13314796/files/figure.png" pageId="4" pageNumber="42">Figure 1 D</figureCitation>
) (
<tableCitation id="323403070423FFE3FD6C859DA487F9AF" box="[658,751,1529,1556]" captionStart="Table 2" captionStartId="4.[151,216,195,219]" captionText="Table 2. Comparison of morphological features of the subspecies of L. atifi." pageId="4" pageNumber="42">Table 2</tableCitation>
).
@ -770,7 +773,7 @@ specimens collected from the same region. However, unlike the adult specimens, d
<paragraph id="7F0936BC0423FFE3FF698659A5B0F964" blockId="4.[151,1436,1597,1759]" pageId="4" pageNumber="42">
<emphasis id="4DC2EAAE0423FFE3FF698659A78BF9EC" bold="true" box="[151,483,1597,1623]" pageId="4" pageNumber="42">Habitat and Distribution:</emphasis>
The specimens were collected from the open areas in the forest around the Doğanca Village, Gazipaşa (
<figureCitation id="E78D2A390423FFE3FE0D863AA437F9C2" box="[499,607,1630,1657]" captionStart="Figure 2" captionStartId="5.[151,226,692,716]" captionTargetBox="[151,1435,189,674]" captionTargetId="figure-17@5.[151,1435,189,674]" captionTargetPageId="5" captionText="Figure 2. Habitats of Lyciasalamandra atifi oezi n. ssp. from the Doğanca Village (Gazipaşa, Antalya)." pageId="4" pageNumber="42">Figure 2</figureCitation>
<figureCitation id="E78D2A390423FFE3FE0D863AA437F9C2" box="[499,607,1630,1657]" captionStart="Figure 2" captionStartId="5.[151,226,692,716]" captionTargetBox="[151,1435,189,674]" captionTargetId="figure-17@5.[151,1435,189,674]" captionTargetPageId="5" captionText="Figure 2. Habitats of Lyciasalamandra atifi oezi n. ssp. from the Doğanca Village (Gazipaşa, Antalya)." figureDoi="http://doi.org/10.5281/zenodo.13314800" httpUri="https://zenodo.org/record/13314800/files/figure.png" pageId="4" pageNumber="42">Figure 2</figureCitation>
).
<specimenCount id="69B0FD350423FFE3FD86863AA4A5F9C2" box="[632,717,1630,1657]" count="1" pageId="4" pageNumber="42" type="male">One ♂</specimenCount>
,