treatments-xml/data/03/BE/CF/03BECF3DD81CF87DD80DFD9BFC522D5C.xml

<document id="BFD81C40159052337A5EB5D7D55FD6C3" ID-CLB-Dataset="21859" ID-DOI="10.5852/ejt.2016.245" ID-GBIF-Dataset="84ee7aa9-b217-40fb-9a7d-d94dd8c906de" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850674" ID-ZooBank="E42F1AE2-B8E6-43C8-896C-75A87814642A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590447021153" checkinUser="valdenar" docAuthor="Pan, Zhixiang &amp; Zhang, Feng" docDate="2016" docId="03BECF3DD81CF87DD80DFD9BFC522D5C" docLanguage="en" docName="ejt-2016-245.pdf.imf" docOrigin="European Journal of Taxonomy 245" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Willowsia Shoebotham 1917" docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FF87B745D80EF86BDA0DFF95FFB02E54" masterDocTitle="Contribution to the Willowsia species having body scales of the long basal rib type: four new species and a redescription of W. qui (Collembola: Entomobryidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="19" updateTime="1698841320807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AAD1D801F4FD7ACC1D3D97964A9413B8">Contribution to the Willowsia species having body scales of the long basal rib type: four new species and a redescription of W. qui (Collembola: Entomobryidae)</mods:title>
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<mods:namePart id="A555DB80FDA71BCEA9DA39314B624FAD">Pan, Zhixiang</mods:namePart>
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<mods:affiliation id="2468432FDD7462558C1CF79907E5B37F">School of Life Sciences, Taizhou University, Jiaojiang, Taizhou 318000, Zhejiang Province, P. R. China. &amp; Email: pzx 1118 @ hotmail. com &amp; urn: lsid: zoobank. org: author: C 66099 CD- 2681 - 453 F-BCC 3 - 3 B 627 AB 31 DC 5</mods:affiliation>
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<mods:namePart id="881902B5C1087B00C285C11CACC77994">Zhang, Feng</mods:namePart>
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<mods:affiliation id="EC6C25582EE0278F1666B94DB1B7032C">Department of Entomology, College of Plant Protection, Nanjing Agricultural University, 1 Weigang, Nanjing 210095, P. R. China. &amp; urn: lsid: zoobank. org: author: 904 C 4445 - AAEA- 40 BC-B 418 - 502053 EF 39 B 8 &amp; Corresponding author: xtmtd. zf @ gmail. com</mods:affiliation>
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<treatment id="03BECF3DD81CF87DD80DFD9BFC522D5C" ID-DOI="http://doi.org/10.5281/zenodo.3854742" ID-GBIF-Taxon="164283285" ID-Zenodo-Dep="3854742" LSID="urn:lsid:plazi:treatment:03BECF3DD81CF87DD80DFD9BFC522D5C" httpUri="http://treatment.plazi.org/id/03BECF3DD81CF87DD80DFD9BFC522D5C" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
<subSubSection id="C30D2DA0D81CF879D80DFD9BFB832C7C" box="[512,1075,526,553]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BA87E2BD81CF879D80DFD9BFB832C7C" blockId="18.[512,1075,526,588]" box="[512,1075,526,553]" pageId="18" pageNumber="19">
<heading id="D0E0C947D81CF879D80DFD9BFB832C7C" box="[512,1075,526,553]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis id="B963A239D81CF879D80DFD9BFD172C7C" bold="true" box="[512,679,526,553]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C1705A8D81CF879D80DFD9BFDC82C7D" authorityName="Shoebotham" authorityYear="1917" box="[512,632,526,553]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Willowsia</taxonomicName>
qui
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<bibRefCitation id="EF8603DAD81CF879D8A3FD9BFB832C7C" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[686,1075,526,552]" pageId="18" pageNumber="19" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D2DA0D81CF879D8A5FDA4FC3B2C1F" box="[680,907,561,588]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA87E2BD81CF879D8A5FDA4FC3B2C1F" blockId="18.[512,1075,526,588]" box="[680,907,561,588]" pageId="18" pageNumber="19">
<figureCitation id="132C62AED81CF879D8A5FDA4FC992C18" box="[680,809,561,588]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="19.[189,231,1967,1993]" captionStartId-1="20.[189,232,1892,1918]" captionStartId-2="21.[189,232,1965,1991]" captionTargetBox-0="[188,1399,1351,1922]" captionTargetBox-1="[195,1395,309,1852]" captionTargetBox-2="[202,1381,423,1908]" captionTargetId-0="figure@19.[183,1402,1349,1926]" captionTargetId-1="figure@20.[189,1398,307,1862]" captionTargetId-2="figure@21.[189,1398,423,1911]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="Fig. 13. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Habitus. B. Abd. III. Scale bars: A = 500 μm; B = 20 μm." captionText-1="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." captionText-2="Fig. 15. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Thoracic chaetotaxy. B–C. Abdominal chaetotaxy. B. Abd. I–IV. C. Abd. V. Scale bars: 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3850722" figureDoi-1="http://doi.org/10.5281/zenodo.3850726" figureDoi-2="http://doi.org/10.5281/zenodo.3850730" httpUri-0="https://zenodo.org/record/3850722/files/figure.png" httpUri-1="https://zenodo.org/record/3850726/files/figure.png" httpUri-2="https://zenodo.org/record/3850730/files/figure.png" pageId="18" pageNumber="19">Figs 13–15</figureCitation>
; 
<tableCitation id="C6954B90D81CF879D938FDA4FC3B2C1F" box="[821,907,561,587]" captionStart="Table 3" captionStartId="19.[189,258,268,294]" captionText="Table 3. Comparison between W. qui Zhang, Chen &amp; Deharveng, 2011 and W. japonica Folsom, 1897." pageId="18" pageNumber="19">Table 3</tableCitation>
</paragraph>
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<subSubSection id="C30D2DA0D81CF879DAB0FDE7FDF82D71" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BA87E2BD81CF879DAB0FDE7FE872CD8" blockId="18.[189,311,626,652]" box="[189,311,626,652]" pageId="18" pageNumber="19">
<heading id="D0E0C947D81CF879DAB0FDE7FE872CD8" bold="true" box="[189,311,626,652]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B963A239D81CF879DAB0FDE7FE872CD8" bold="true" box="[189,311,626,652]" pageId="18" pageNumber="19">Diagnosis</emphasis>
</heading>
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Very narrow lateral stripes on Th. II–Abd. I and transverse bands on postero-lateral margin of Abd. II–III; cephalic mac S 
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absent but S 
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' present; 7 clypeal chaetae; Th. II with 2 medio-medial, 3 mediolateral, and 12–13 posterior mac; chaeta a1 as mic on Th. III; Abd. I with 4 mac; Abd. III with 2 central mac; Abd. IV with 6 central mac.
</paragraph>
</subSubSection>
<subSubSection id="C30D2DA0D81CF879DAB0FCD9FE9A2AFB" pageId="18" pageNumber="19" type="materials_examined">
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<emphasis id="B963A239D81CF879DAB0FCD9FE1B2D32" bold="true" box="[189,427,844,870]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
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<collectingCountry id="F3003EBBD81CF879DAB0FCEEFEAF2DC1" box="[189,287,891,917]" name="China" pageId="18" pageNumber="19">CHINA</collectingCountry>
: 
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and 
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on slides, 
<collectingRegion id="49D3B0C9D81CF879D8CBFCEEFC152DC1" box="[710,933,891,917]" country="China" name="Zhejiang" pageId="18" pageNumber="19">Zhejiang Province</collectingRegion>
, Taishun County, Wuyanling National Forest Park, Shuangkengkou, 
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, 
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, altitude 
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, 
<date id="FFA958EBD81CF879D9E7FC0BFB302DED" box="[1002,1152,926,953]" pageId="18" pageNumber="19" value="2005-08-04">4 Aug. 2005</date>
, J.X. Chen team leg. (sample # C9279); 2 ³³, 
<specimenCount id="9D11B5A2D81CF879DBF8FC54FD932D88" box="[501,547,961,988]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
, on slides, 
<collectingRegion id="49D3B0C9D81CF879D8BCFC54FCCD2D8F" box="[689,893,961,987]" country="China" name="Jiangxi" pageId="18" pageNumber="19">Jiangxi Province</collectingRegion>
, Nanchang, Shangfengjing Village, under leaf litter of bamboo, 
<geoCoordinate id="EE2318ECD81CF879DBCDFC71FD882DAB" box="[448,568,996,1023]" degrees="28.792" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="55" value="28.792">28.792° N</geoCoordinate>
, 
<geoCoordinate id="EE2318ECD81CF879D848FC71FD772DAB" box="[581,711,996,1023]" degrees="115.808" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="55" value="115.808">115.808° E</geoCoordinate>
, altitude 
<quantity id="4CEFD3CED81CF879D939FC71FCCE2DAB" box="[820,894,996,1023]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="18" pageNumber="19" unit="m" value="118.0">118 m</quantity>
, 
<date id="FFA958EBD81CF879D986FC71FB9A2DAA" box="[907,1066,996,1023]" pageId="18" pageNumber="19" value="2012-08-11">11 Aug. 2012</date>
, Z.X. Pan and Q.P. Ren leg. (slide # 4383_1–3, sample # 4383); 
<specimenCount id="9D11B5A2D81CF879D85FFB9DFDCD2A76" box="[594,637,1032,1058]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
on slide, 
<collectingRegion id="49D3B0C9D81CF879D8E0FB92FC1B2A75" box="[749,939,1031,1057]" country="China" name="Anhui" pageId="18" pageNumber="19">Anhui Province</collectingRegion>
, Yi County, Xidi Village, under leaves, 
<geoCoordinate id="EE2318ECD81CF879DAB0FBBEFE872A11" box="[189,311,1067,1094]" degrees="29.905" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="55" value="29.905">29.905° N</geoCoordinate>
, 
<geoCoordinate id="EE2318ECD81CF879DB48FBBEFE772A11" box="[325,455,1067,1093]" degrees="117.992" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="55" value="117.992">117.992° E</geoCoordinate>
, altitude 
<quantity id="4CEFD3CED81CF879D837FBBEFD3A2A11" box="[570,650,1067,1093]" metricMagnitude="2" metricUnit="m" metricValue="2.23" pageId="18" pageNumber="19" unit="m" value="223.0">223 m</quantity>
, 
<date id="FFA958EBD81CF879D894FBBEFC8B2A11" box="[665,827,1067,1093]" pageId="18" pageNumber="19" value="2012-11-18">18 Nov. 2012</date>
, F. Zhang, Z.-X. Pan and Z.-H. Li leg. (slide # HC13_1, sample # HC13); 
<specimenCount id="9D11B5A2D81CF879DBFBFBDBFD932A3C" box="[502,547,1102,1128]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
on slide, 
<collectingRegion id="49D3B0C9D81CF879D89BFBDBFCE52A3C" box="[662,853,1102,1128]" country="China" name="Anhui" pageId="18" pageNumber="19">Anhui Province</collectingRegion>
, Tangkou Town, Fuxi Village, bamboo forest, 
<geoCoordinate id="EE2318ECD81CF879DAB0FBE4FE9A2AD8" box="[189,298,1137,1164]" degrees="30.08" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="30.08">30.08° N</geoCoordinate>
, 
<geoCoordinate id="EE2318ECD81CF879DB34FBE4FE0A2AD8" box="[313,442,1137,1164]" degrees="118.148" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="55" value="118.148">118.148° E</geoCoordinate>
, altitude 
<quantity id="4CEFD3CED81CF879D821FBE4FDCC2AD8" box="[556,636,1137,1164]" metricMagnitude="2" metricUnit="m" metricValue="6.89" pageId="18" pageNumber="19" unit="m" value="689.0">689 m</quantity>
, 
<date id="FFA958EBD81CF879D887FBE4FCAF2AD8" box="[650,799,1137,1164]" pageId="18" pageNumber="19" value="2015-07-15">15 Jul. 2015</date>
, F. Zhang leg. (slide # 15AH10, sample # 15AH_ monkey).
</paragraph>
</subSubSection>
<subSubSection id="C30D2DA0D81CF878DAB0FB43FE8A2B45" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
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<emphasis id="B963A239D81CF879DAB0FB43FEFF2AA4" bold="true" box="[189,335,1238,1264]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA87E2BD81CF879DAB0FA91FE492B4B" blockId="18.[189,505,1284,1311]" box="[189,505,1284,1311]" pageId="18" pageNumber="19">
Body length up to 
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.
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<paragraph id="8BA87E2BD81CF879DAB0FAD3FE7A2844" blockId="18.[189,1399,1348,1552]" pageId="18" pageNumber="19">
Ground colour pale in alcohol. Ant. I̅II with distal patches and III–IV dark. A transverse band between eye patches. Narrow stripes present on lateral margin of Th. II–Abd. I. Narrow transverse bands on posterior margin of Abd. II–III. Small patches posteriorly on Abd. IV–VI (
<figureCitation id="132C62AED81CF879DE4FFA19FB1C2BF2" box="[1090,1196,1420,1446]" captionStart="Fig" captionStartId="19.[189,231,1967,1993]" captionTargetBox="[188,1399,1351,1922]" captionTargetId="figure@19.[183,1402,1349,1926]" captionTargetPageId="19" captionText="Fig. 13. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Habitus. B. Abd. III. Scale bars: A = 500 μm; B = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3850722" httpUri="https://zenodo.org/record/3850722/files/figure.png" pageId="18" pageNumber="19">Fig. 13A</figureCitation>
). Scales pointed, heavily striate with basal ribs longer than distal ones, and only present dorsally on head and terga. Some chaetae of the row along the posterior margin of each tergum transformed into scales and larger than other scales (
<figureCitation id="132C62AED81CF879DB5AFA63FE0D2844" box="[343,445,1526,1552]" captionStart="Fig" captionStartId="19.[189,231,1967,1993]" captionTargetBox="[188,1399,1351,1922]" captionTargetId="figure@19.[183,1402,1349,1926]" captionTargetPageId="19" captionText="Fig. 13. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Habitus. B. Abd. III. Scale bars: A = 500 μm; B = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3850722" httpUri="https://zenodo.org/record/3850722/files/figure.png" pageId="18" pageNumber="19">Fig. 13B</figureCitation>
).
</paragraph>
<paragraph id="8BA87E2BD81CF879DAB0F9A2FBE728CC" blockId="18.[189,1400,1591,1688]" pageId="18" pageNumber="19">Based on 5 measured specimens, antenna 2.48–2.54 times as long as cephalic diagonal. Antennal segment ratio as I: II: III: IV = 1: 1.69–2.12: 1.69–2.00: 2.65–3.15. Smooth spiny mic at base of antennae: 3 dorsal and 2 ventral on Ant. I; 1 internal, 1 external and 1 ventral on Ant. II.</paragraph>
<paragraph id="8BA87E2BD81CF879DAB0F92AFEE52932" blockId="18.[189,1399,1727,1894]" pageId="18" pageNumber="19">
Labral intrusion slightly curved. Seven clypeal chaetae ciliate and arranged in two rows (
<figureCitation id="132C62AED81CF879DF0DF92AFADA288D" box="[1280,1386,1727,1753]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="18" pageNumber="19">Fig. 14B</figureCitation>
). Dorsal cephalic chaetotaxy with 4 antennal (An), 4 median (M) and 7 sutural (S) mac; mac S 
<subScript id="17937C6ED81CF879DF09F97AFABC28A9" attach="left" box="[1284,1292,1775,1789]" fontSize="6" pageId="18" pageNumber="19">0</subScript>
' present; mac S 
<subScript id="17937C6ED81CF879DB04F887FEA22974" attach="left" box="[265,274,1810,1824]" fontSize="6" pageId="18" pageNumber="19">0</subScript>
and S 
<subScript id="17937C6ED81CF879DB6CF887FEDA2974" attach="left" box="[353,362,1810,1824]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
absent; mac Ps 
<subScript id="17937C6ED81CF879D825F887FD812974" attach="left" box="[552,561,1810,1824]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
rarely absent (
<figureCitation id="132C62AED81CF879D8E5F890FCE32974" box="[744,851,1797,1824]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="18" pageNumber="19">Fig. 14C</figureCitation>
). Tip of lateral process of labial palp slightly beyond apex of labial papilla. Labial chaeta R/M = 0.65–0.76. Cephalic groove with 7(6) ciliate chaetae on each side.
</paragraph>
<paragraph id="8BA87E2BD81CF878DAB0F818FBC82A74" blockId="18.[189,1399,1933,1995]" lastBlockId="19.[189,1399,853,1056]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Metatrochanteral organ with 10̅14 smooth spine-like chaetae; 8̅ 
<quantity id="4CEFD3CED81CF879D9B6F818FC4329F3" box="[955,1011,1933,1959]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="18" pageNumber="19" unit="in" value="11.0">11 in</quantity>
L-shaped arms and 2̅3 between arms (
<figureCitation id="132C62AED81CF879DB05F825FEC1299F" box="[264,369,1968,1995]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="18" pageNumber="19">Fig. 14D</figureCitation>
). Ungual basal paired inner teeth with tip reaching 52% from base, median one at 75% and distal one at 88%. Tenent hairs slightly longer than unguis (
<figureCitation id="132C62AED81DF878D9B5FCC0FB902D3B" box="[952,1056,853,879]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14E</figureCitation>
). Abd. IV 3.47–3.70 times as long as Abd. III along dorsal midline. Ventral tube anteriorly with 12–15 ciliate chaetae, 5–7 of them larger than others (
<figureCitation id="132C62AED81DF878DBA8FC0EFDBC2DE2" box="[421,524,923,950]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14F</figureCitation>
); posteriorly with 6–8 smooth chaetae; each lateral flap with 11 chaetae (
<figureCitation id="132C62AED81DF878DAC8FC2AFE9F2D8D" box="[197,303,959,985]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14G</figureCitation>
). Manubrium dorsally with 6+6 thick chaetae, 3 large and 3 (
<quantity id="4CEFD3CED81DF878D9F4FC2AFB922D8D" box="[1017,1058,959,985]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="20" unit="in" value="1.0">1 in</quantity>
distal plaque) median in size (
<figureCitation id="132C62AED81DF878DAC8FC77FE9F2DA8" box="[197,303,994,1020]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14H</figureCitation>
); ventrally with 8(7)+8(7) distal chaetae (
<figureCitation id="132C62AED81DF878D91AFC77FCC52DA8" box="[791,885,994,1020]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14I</figureCitation>
). Manubrial plaque with 2 pseudopores and 4 ciliate chaetae (
<figureCitation id="132C62AED81DF878DB86FB90FE452A4B" box="[395,501,1029,1055]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[195,1395,309,1852]" captionTargetId="figure@20.[189,1398,307,1862]" captionTargetPageId="20" captionText="Fig. 14. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Ant. III organ. B. Clypeal chaetae. C. Dorsal cephalic chaetotaxy. D. Trochanteral organ. E. Hind claw. F. Anterior face of ventral tube. G. Posterior face and lateral flap of ventral tube. H. Manubrium and base of dens, dorsal view. I. Distal chaetae of ventral face of manubrium. J. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="19" pageNumber="20">Fig. 14H</figureCitation>
). Distal smooth part 1.60̅1.81 times as long as mucro.
</paragraph>
<caption id="DF682EA3D81DF878DAB0FE99FADD2F72" box="[189,1389,267,294]" pageId="19" pageNumber="20">
<paragraph id="8BA87E2BD81DF878DAB0FE99FADD2F72" blockId="19.[189,1389,267,294]" box="[189,1389,267,294]" pageId="19" pageNumber="20">
<emphasis id="B963A239D81DF878DAB0FE99FEAF2F71" bold="true" box="[189,287,267,294]" pageId="19" pageNumber="20">Table 3.</emphasis>
Comparison between 
<emphasis id="B963A239D81DF878D825FE99FDC02F71" box="[552,624,268,293]" italics="true" pageId="19" pageNumber="20">W. qui</emphasis>
<bibRefCitation id="EF8603DAD81DF878D877FE9EFC4E2F72" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[634,1022,267,294]" pageId="19" pageNumber="20" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
and 
<taxonomicName id="4C1705A8D81DF878DE35FE99FAD92F72" authority="Folsom, 1897" authorityName="Folsom" authorityYear="1897" box="[1080,1385,267,294]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B963A239D81DF878DE35FE99FB732F71" box="[1080,1219,268,293]" italics="true" pageId="19" pageNumber="20">W. japonica</emphasis>
Folsom, 1897
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA87E2BD81DF878DAC5FEFCFAF12D56" pageId="19" pageNumber="20">
<table id="F9178C8BD81D0794DAC9FEC2FAF12D57" box="[196,1345,343,771]" gridcols="3" gridrows="11" pageId="19" pageNumber="20">
<tr id="35277C69D81D0794DAC9FEC2FAF12FC1" box="[196,1345,343,405]" gridrow="0" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FEC2FD9D2FC1" box="[196,557,343,405]" gridcol="0" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="B963A239D81DF878DAC5FEFCFEE42FD7" bold="true" box="[200,340,361,387]" pageId="19" pageNumber="20">Characters</emphasis>
</th>
<th id="76F61515D81D0794D8AEFEC2FB982FC1" box="[675,1064,343,405]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="B963A239D81DF878D932FECDFC3D2F25" bold="true" box="[831,909,343,370]" italics="true" pageId="19" pageNumber="20">W. qui</emphasis>
<bibRefCitation id="EF8603DAD81DF878D8AEFEEEFB982FC1" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[675,1064,379,405]" pageId="19" pageNumber="20" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
</th>
<th id="76F61515D81D0794DE91FEC2FAF12FC1" box="[1180,1345,343,405]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">
<taxonomicName id="4C1705A8D81DF878DEAAFECDFAF02FC1" authority="Folsom, 1897" authorityName="Folsom" authorityYear="1897" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B963A239D81DF878DEAAFECDFA862F25" bold="true" box="[1191,1334,343,370]" italics="true" pageId="19" pageNumber="20">W. japonica</emphasis>
Folsom, 1897
</taxonomicName>
</th>
</tr>
<tr id="35277C69D81D0794DAC9FE33FAF12F94" box="[196,1345,422,448]" gridrow="1" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FE33FD9D2F94" box="[196,557,422,448]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">Stripes dorsally on body</th>
<td id="76F61515D81D0794D8AEFE33FB982F94" box="[675,1064,422,448]" gridcol="1" gridrow="1" pageId="19" pageNumber="20">very narrow</td>
<td id="76F61515D81D0794DE91FE33FAF12F94" box="[1180,1345,422,448]" gridcol="2" gridrow="1" pageId="19" pageNumber="20">normal</td>
</tr>
<tr id="35277C69D81D0794DAC9FE5CFAF12FB0" box="[196,1345,457,484]" gridrow="2" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FE5CFD9D2FB0" box="[196,557,457,484]" gridcol="0" gridrow="2" pageId="19" pageNumber="20">Small patch dorsally on Th. III</th>
<td id="76F61515D81D0794D8AEFE5CFB982FB0" box="[675,1064,457,484]" gridcol="1" gridrow="2" pageId="19" pageNumber="20">absent</td>
<td id="76F61515D81D0794DE91FE5CFAF12FB0" box="[1180,1345,457,484]" gridcol="2" gridrow="2" pageId="19" pageNumber="20">present</td>
</tr>
<tr id="35277C69D81D0794DAC9FE79FAF12C53" box="[196,1345,492,519]" gridrow="3" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FE79FD9D2C53" box="[196,557,492,519]" gridcol="0" gridrow="3" pageId="19" pageNumber="20">Scale width</th>
<td id="76F61515D81D0794D8AEFE79FB982C53" box="[675,1064,492,519]" gridcol="1" gridrow="3" pageId="19" pageNumber="20">broad</td>
<td id="76F61515D81D0794DE91FE79FAF12C53" box="[1180,1345,492,519]" gridcol="2" gridrow="3" pageId="19" pageNumber="20">narrow</td>
</tr>
<tr id="35277C69D81D0794DAC9FD85FAF12C7F" box="[196,1345,528,555]" gridrow="4" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FD85FD9D2C7F" box="[196,557,528,555]" gridcol="0" gridrow="4" pageId="19" pageNumber="20">Scale surface sculpture</th>
<td id="76F61515D81D0794D8AEFD85FB982C7F" box="[675,1064,528,555]" gridcol="1" gridrow="4" pageId="19" pageNumber="20">long basal rib type</td>
<td id="76F61515D81D0794DE91FD85FAF12C7F" box="[1180,1345,528,555]" gridcol="2" gridrow="4" pageId="19" pageNumber="20">short rib type</td>
</tr>
<tr id="35277C69D81D0794DAC9FDA6FAF12C1A" box="[196,1345,563,590]" gridrow="5" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FDA6FD9D2C1A" box="[196,557,563,590]" gridcol="0" gridrow="5" pageId="19" pageNumber="20">Smooth part of dens/mucro</th>
<td id="76F61515D81D0794D8AEFDA6FB982C1A" box="[675,1064,563,590]" gridcol="1" gridrow="5" pageId="19" pageNumber="20">1.6–1.8</td>
<td id="76F61515D81D0794DE91FDA6FAF12C1A" box="[1180,1345,563,590]" gridcol="2" gridrow="5" pageId="19" pageNumber="20">2.2–2.9</td>
</tr>
<tr id="35277C69D81D0794DAC9FDC2FAF12C25" box="[196,1345,599,625]" gridrow="6" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FDC2FAF12C25" box="[196,1345,599,625]" colspan="3" colspanRight="2" gridcol="0" gridrow="6" pageId="19" pageNumber="20">Chaetotaxy of Th. II</th>
</tr>
<tr id="35277C69D81D0794DAC9FDEFFAF12CC1" box="[196,1345,634,661]" gridrow="7" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FDEFFD9D2CC1" box="[196,557,634,661]" gridcol="0" gridrow="7" pageId="19" pageNumber="20">m1 and m5</th>
<td id="76F61515D81D0794D8AEFDEFFB982CC1" box="[675,1064,634,661]" gridcol="1" gridrow="7" pageId="19" pageNumber="20">mac</td>
<td id="76F61515D81D0794DE91FDEFFAF12CC1" box="[1180,1345,634,661]" gridcol="2" gridrow="7" pageId="19" pageNumber="20">mic</td>
</tr>
<tr id="35277C69D81D0794DAC9FD0AFAF12CEE" box="[196,1345,671,698]" gridrow="8" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FD0AFD9D2CEE" box="[196,557,671,698]" gridcol="0" gridrow="8" pageId="19" pageNumber="20">mac m4i and m4p</th>
<td id="76F61515D81D0794D8AEFD0AFB982CEE" box="[675,1064,671,698]" gridcol="1" gridrow="8" pageId="19" pageNumber="20">present</td>
<td id="76F61515D81D0794DE91FD0AFAF12CEE" box="[1180,1345,671,698]" gridcol="2" gridrow="8" pageId="19" pageNumber="20">absent</td>
</tr>
<tr id="35277C69D81D0794DAC9FD51FAF12C8B" box="[196,1345,708,735]" gridrow="9" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FD51FD9D2C8B" box="[196,557,708,735]" gridcol="0" gridrow="9" pageId="19" pageNumber="20">Mac on Abd. I</th>
<td id="76F61515D81D0794D8AEFD51FB982C8B" box="[675,1064,708,735]" gridcol="1" gridrow="9" pageId="19" pageNumber="20">4</td>
<td id="76F61515D81D0794DE91FD51FAF12C8B" box="[1180,1345,708,735]" gridcol="2" gridrow="9" pageId="19" pageNumber="20">3</td>
</tr>
<tr id="35277C69D81D0794DAC9FD7DFAF12D57" box="[196,1345,744,771]" gridrow="10" pageId="19" pageNumber="20">
<th id="76F61515D81D0794DAC9FD7DFD9D2D57" box="[196,557,744,771]" gridcol="0" gridrow="10" pageId="19" pageNumber="20">Central mac on Abd. IV</th>
<td id="76F61515D81D0794D8AEFD7DFB982D57" box="[675,1064,744,771]" gridcol="1" gridrow="10" pageId="19" pageNumber="20">6 (A5 as meso)</td>
<td id="76F61515D81D0794DE91FD7DFAF12D57" box="[1180,1345,744,771]" gridcol="2" gridrow="10" pageId="19" pageNumber="20">7 (A5 as mac)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA87E2BD81DF878DAB0FBD3FE8A2B45" blockId="19.[189,1399,1094,1297]" pageId="19" pageNumber="20">
Th. II with 2(3) (m1, m2) medio-medial, 3 (m4, m4i, m4p) medio-lateral and 12–13 posterior mac; mac m2i present in one specimen; m5 and p4–6 as mac; mac p2a often absent. Th. III with 14 mac; a1 as mic; m6e as mesochaeta (
<figureCitation id="132C62AED81DF878DBEBFB18FDFD2AFC" box="[486,589,1165,1192]" captionStart="Fig" captionStartId="21.[189,232,1965,1991]" captionTargetBox="[202,1381,423,1908]" captionTargetId="figure@21.[189,1398,423,1911]" captionTargetPageId="21" captionText="Fig. 15. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Thoracic chaetotaxy. B–C. Abdominal chaetotaxy. B. Abd. I–IV. C. Abd. V. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850730" httpUri="https://zenodo.org/record/3850730/files/figure.png" pageId="19" pageNumber="20">Fig. 15A</figureCitation>
). Abd. I with 4 (m2̅4, m4p) mac. Abd. II with 3 (a2, m3, m3e) central and 1 (m5) lateral mac. Abd. III with 2 (a2, m3) central and 3 (am6, pm6, p6) lateral mac; a3 as mac in one specimen. Abd. IV with 6 central mac, about 12 lateral mac and at least 9 sens; A5 as mesochaeta (
<figureCitation id="132C62AED81DF878DAC8FB63FE9C2B45" box="[197,300,1270,1297]" captionStart="Fig" captionStartId="21.[189,232,1965,1991]" captionTargetBox="[202,1381,423,1908]" captionTargetId="figure@21.[189,1398,423,1911]" captionTargetPageId="21" captionText="Fig. 15. Willowsia qui Zhang, Chen &amp; Deharveng, 2011. A. Thoracic chaetotaxy. B–C. Abdominal chaetotaxy. B. Abd. I–IV. C. Abd. V. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3850730" httpUri="https://zenodo.org/record/3850730/files/figure.png" pageId="19" pageNumber="20">Fig. 15B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF682EA3D81DF878DAB0F83AFEF329B9" ID-DOI="http://doi.org/10.5281/zenodo.3850722" ID-Zenodo-Dep="3850722" httpUri="https://zenodo.org/record/3850722/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,231,1967,1993]" targetBox="[188,1399,1351,1922]" targetPageId="19">
<paragraph id="8BA87E2BD81DF878DAB0F83AFEF329B9" blockId="19.[189,1399,1967,2029]" pageId="19" pageNumber="20">
<emphasis id="B963A239D81DF878DAB0F83AFEA7299D" bold="true" box="[189,279,1967,1993]" pageId="19" pageNumber="20">Fig. 13.</emphasis>
<emphasis id="B963A239D81DF878DB13F83AFE0B299D" box="[286,443,1967,1993]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C1705A8D81DF878DB13F83AFE3F299D" authorityName="Shoebotham" authorityYear="1917" box="[286,399,1967,1993]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Willowsia</taxonomicName>
qui
</emphasis>
<bibRefCitation id="EF8603DAD81DF878DBCCF83AFC8F299D" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[449,831,1967,1993]" pageId="19" pageNumber="20" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
. 
<emphasis id="B963A239D81DF878D944F83AFCEE299D" bold="true" box="[841,862,1967,1993]" pageId="19" pageNumber="20">A</emphasis>
. Habitus. 
<emphasis id="B963A239D81DF878D9D9F825FC57299E" bold="true" box="[980,999,1968,1994]" pageId="19" pageNumber="20">B</emphasis>
. Abd. III. Scale bars: A = 500 μm; B = 20 μm.
</paragraph>
</caption>
<caption id="DF682EA3D81AF87FDAB0F8F1FB3929BD" ID-DOI="http://doi.org/10.5281/zenodo.3850726" ID-Zenodo-Dep="3850726" httpUri="https://zenodo.org/record/3850726/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1892,1918]" targetBox="[195,1395,309,1852]" targetPageId="20">
<paragraph id="8BA87E2BD81AF87FDAB0F8F1FB3929BD" blockId="20.[189,1399,1892,2025]" pageId="20" pageNumber="21">
<emphasis id="B963A239D81AF87FDAB0F8F1FEA5292A" bold="true" box="[189,277,1892,1918]" pageId="20" pageNumber="21">Fig. 14.</emphasis>
<emphasis id="B963A239D81AF87FDB10F8F0FE0B292A" box="[285,443,1893,1919]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C1705A8D81AF87FDB10F8F0FE21292B" authorityName="Shoebotham" authorityYear="1917" box="[285,401,1893,1919]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Willowsia</taxonomicName>
qui
</emphasis>
<bibRefCitation id="EF8603DAD81AF87FDBCFF8F1FCF4292B" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[450,836,1892,1919]" pageId="20" pageNumber="21" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
. 
<emphasis id="B963A239D81AF87FD940F8F1FCD2292A" bold="true" box="[845,866,1892,1918]" pageId="20" pageNumber="21">A</emphasis>
. Ant. III organ. 
<emphasis id="B963A239D81AF87FDE13F8F0FB81292B" bold="true" box="[1054,1073,1893,1919]" pageId="20" pageNumber="21">B</emphasis>
. Clypeal chaetae. 
<emphasis id="B963A239D81AF87FDF0BF8F1FAAB292A" bold="true" box="[1286,1307,1892,1918]" pageId="20" pageNumber="21">C</emphasis>
. Dorsal cephalic chaetotaxy. 
<emphasis id="B963A239D81AF87FDBBEF81DFE7729F6" bold="true" box="[435,455,1928,1954]" pageId="20" pageNumber="21">D</emphasis>
. Trochanteral organ. 
<emphasis id="B963A239D81AF87FD8C9F81DFD6729F6" bold="true" box="[708,727,1928,1954]" pageId="20" pageNumber="21">E</emphasis>
. Hind claw. 
<emphasis id="B963A239D81AF87FD961F81DFCCB29F6" bold="true" box="[876,891,1928,1954]" pageId="20" pageNumber="21">F</emphasis>
. Anterior face of ventral tube. 
<emphasis id="B963A239D81AF87FDEE5F81DFB4E29F6" bold="true" box="[1256,1278,1928,1954]" pageId="20" pageNumber="21">G</emphasis>
. Posterior face and lateral flap of ventral tube. 
<emphasis id="B963A239D81AF87FD860F83EFD342991" bold="true" box="[621,644,1963,1989]" pageId="20" pageNumber="21">H</emphasis>
. Manubrium and base of dens, dorsal view. 
<emphasis id="B963A239D81AF87FDE99F83EFB2F2991" bold="true" box="[1172,1183,1963,1989]" pageId="20" pageNumber="21">I</emphasis>
. Distal chaetae of ventral face of manubrium. 
<emphasis id="B963A239D81AF87FD808F85BFDA329BC" bold="true" box="[517,531,1998,2024]" pageId="20" pageNumber="21">J</emphasis>
. Mucro. Scale bars: A–B, D–J = 20 μm; C = 100 μm.
</paragraph>
</caption>
<caption id="DF682EA3D81BF87EDAB0F838FCEE29BF" ID-DOI="http://doi.org/10.5281/zenodo.3850730" ID-Zenodo-Dep="3850730" httpUri="https://zenodo.org/record/3850730/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1965,1991]" targetBox="[202,1381,423,1908]" targetPageId="21">
<paragraph id="8BA87E2BD81BF87EDAB0F838FCEE29BF" blockId="21.[189,1398,1965,2027]" pageId="21" pageNumber="22">
<emphasis id="B963A239D81BF87EDAB0F838FEAC2993" bold="true" box="[189,284,1965,1991]" pageId="21" pageNumber="22">Fig. 15.</emphasis>
<emphasis id="B963A239D81BF87EDB2AF838FE7B2993" box="[295,459,1965,1991]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C1705A8D81BF87EDB2AF838FE2B2993" authorityName="Shoebotham" authorityYear="1917" box="[295,411,1965,1991]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Willowsia</taxonomicName>
qui
</emphasis>
<bibRefCitation id="EF8603DAD81BF87EDBDBF838FCDC2993" author="Zhang F. &amp; Chen J. X. &amp; Deharveng L." box="[470,876,1965,1991]" pageId="21" pageNumber="22" pagination="1 - 20" refId="ref9231" refString="Zhang F., Chen J. X. &amp; Deharveng L. 2011. New insight into the systematics of the Willowsia complex (Collembola: Entomobryidae). Annales de la Societe Entomologique de France 47: 1 - 20. http: // dx. doi. org / 10.1080 / 00379271.2011.10697692" type="journal article" year="2011">Zhang, Chen &amp; Deharveng, 2011</bibRefCitation>
. 
<emphasis id="B963A239D81BF87ED977F838FC3F2993" bold="true" box="[890,911,1965,1991]" pageId="21" pageNumber="22">A</emphasis>
. Thoracic chaetotaxy. 
<emphasis id="B963A239D81BF87EDEA8F838FB6F2993" bold="true" box="[1189,1247,1965,1991]" pageId="21" pageNumber="22">B–C</emphasis>
. Abdominal chaetotaxy. 
<emphasis id="B963A239D81BF87EDB47F845FEED29BE" bold="true" box="[330,349,2000,2026]" pageId="21" pageNumber="22">B</emphasis>
. Abd. I–IV. 
<emphasis id="B963A239D81BF87EDBFCF845FDB629BE" bold="true" box="[497,518,2000,2026]" pageId="21" pageNumber="22">C</emphasis>
. Abd. V. Scale bars: 100 μm.
</paragraph>
</caption>
<subSubSection id="C30D2DA0D818F87DDAB0FE98FE7C2F91" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="8BA87E2BD818F87DDAB0FE98FE912F73" blockId="22.[189,289,269,295]" box="[189,289,269,295]" pageId="22" pageNumber="23">
<heading id="D0E0C947D818F87DDAB0FE98FE912F73" bold="true" box="[189,289,269,295]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B963A239D818F87DDAB0FE98FE912F73" bold="true" box="[189,289,269,295]" pageId="22" pageNumber="23">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA87E2BD818F87DDAB0FEAEFC4E2F01" blockId="22.[189,1022,315,342]" box="[189,1022,315,342]" pageId="22" pageNumber="23">Found in litter, debris and rotten logs of evergreen broad-leaved forest.</paragraph>
<paragraph id="8BA87E2BD818F87DDAB0FEE8FEC32FC3" blockId="22.[189,371,381,407]" box="[189,371,381,407]" pageId="22" pageNumber="23">
<heading id="D0E0C947D818F87DDAB0FEE8FEC32FC3" bold="true" box="[189,371,381,407]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B963A239D818F87DDAB0FEE8FEC32FC3" bold="true" box="[189,371,381,407]" pageId="22" pageNumber="23">DNA barcodes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA87E2BD818F87DDAB0FE3EFE7C2F91" blockId="22.[189,460,427,453]" box="[189,460,427,453]" pageId="22" pageNumber="23">
<accessionNumber id="9444E3C8D818F87DDAB0FE3EFE8C2F91" box="[189,316,427,453]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU833221" pageId="22" pageNumber="23">KU833221</accessionNumber>
, 
<accessionNumber id="9444E3C8D818F87DDB4AFE3EFE782F91" box="[327,456,427,453]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU833226" pageId="22" pageNumber="23">KU833226</accessionNumber>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D2DA0D818F87DDAB0FE78FC522D5C" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BA87E2BD818F87DDAB0FE78FE9F2C53" blockId="22.[189,303,493,519]" box="[189,303,493,519]" pageId="22" pageNumber="23">
<heading id="D0E0C947D818F87DDAB0FE78FE9F2C53" bold="true" box="[189,303,493,519]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B963A239D818F87DDAB0FE78FE9F2C53" bold="true" box="[189,303,493,519]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA87E2BD818F87DDAB0FD89FC522D5C" blockId="22.[189,1399,539,777]" pageId="22" pageNumber="23">
We found a 
<typeStatus id="54ACC089D818F87DDB41FD89FE062C62" box="[332,438,540,566]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
, which was actually 
<taxonomicName id="4C1705A8D818F87DD8A1FD89FC412C61" authority="(Folsom, 1897)" baseAuthorityName="Folsom" baseAuthorityYear="1897" box="[684,1009,539,565]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B963A239D818F87DD8A1FD89FC882C61" box="[684,824,540,565]" italics="true" pageId="22" pageNumber="23">W. japonica</emphasis>
(Folsom, 1897)
</taxonomicName>
and incorrectly designated as 
<emphasis id="B963A239D818F87DDF54FD89FF532C0C" italics="true" pageId="22" pageNumber="23">W. qui</emphasis>
by the original authors, resulting in some mistakes in the original description. Based on the types and fresh material, 
<emphasis id="B963A239D818F87DDBAEFDF6FE5B2C28" box="[419,491,611,636]" italics="true" pageId="22" pageNumber="23">W. qui</emphasis>
differs from 
<taxonomicName id="4C1705A8D818F87DD881FDF6FCA82C28" baseAuthorityName="Folsom" baseAuthorityYear="1897" box="[652,792,611,636]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B963A239D818F87DD881FDF6FCA82C28" box="[652,792,611,636]" italics="true" pageId="22" pageNumber="23">W. japonica</emphasis>
</taxonomicName>
in narrower stripes on lateral margin of thorax and posterior margin of Abd. II–III, broad body scales, short distal smooth part of dens, and chaetotaxy of Th. II and Abd. I and IV (
<tableCitation id="C6954B90D818F87DDBFDFD3DFDF92C96" box="[496,585,680,706]" captionStart="Table 3" captionStartId="19.[189,258,268,294]" captionText="Table 3. Comparison between W. qui Zhang, Chen &amp; Deharveng, 2011 and W. japonica Folsom, 1897." pageId="22" pageNumber="23">Table 3</tableCitation>
). Mesochaeta A5 is as long as macrochaetae, but thinner, with small socket and pointed tip. The K2P genetic distance (
<geoCoordinate id="EE2318ECD818F87DD908FD5EFCF52CB1" box="[773,837,715,741]" degrees="0.184" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="0.184">0.184</geoCoordinate>
<geoCoordinate id="EE2318ECD818F87DD948FD5EFC242CB1" box="[837,916,715,741]" degrees="0.192" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-0.192">–0.192</geoCoordinate>
) of the COI (one sequence of 
<taxonomicName id="4C1705A8D818F87DDEE1FD59FAC72CB1" baseAuthorityName="Folsom" baseAuthorityYear="1897" box="[1260,1399,716,741]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B963A239D818F87DDEE1FD59FAC72CB1" box="[1260,1399,716,741]" italics="true" pageId="22" pageNumber="23">W. japonica</emphasis>
</taxonomicName>
from GenBank: 
<accessionNumber id="9444E3C8D818F87DDB72FD7AFDB42D5D" box="[383,516,751,777]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM978378" pageId="22" pageNumber="23">KM978378</accessionNumber>
) also indicates two independent species.
</paragraph>
</subSubSection>
</treatment>
</document>