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<mods:title id="7343B5B314C30E282524135A9A23BDA7">The genera and species of Comatulidae (Comatulida: Crinoidea): taxonomic revisions and a molecular and morphological guide</mods:title>
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<treatment id="039EDF70FFA6A179FF66DD6311D5FE55" ID-DOI="http://doi.org/10.5281/zenodo.6009142" ID-GBIF-Taxon="130809849" ID-Zenodo-Dep="6009142" LSID="urn:lsid:plazi:treatment:039EDF70FFA6A179FF66DD6311D5FE55" httpUri="http://treatment.plazi.org/id/039EDF70FFA6A179FF66DD6311D5FE55" lastPageId="25" lastPageNumber="176" pageId="24" pageNumber="175">
<subSubSection id="C32D3DEDFFA6A178FF66DD6310D7F950" pageId="24" pageNumber="175" type="nomenclature">
<paragraph id="8B886E66FFA6A178FF66DD6311ECF9B1" blockId="24.[151,467,1618,1677]" box="[151,467,1618,1644]" pageId="24" pageNumber="175">
<heading id="D0C0D90AFFA6A178FF66DD6311ECF9B1" bold="true" box="[151,467,1618,1644]" fontSize="11" level="1" pageId="24" pageNumber="175" reason="1">
<emphasis id="B943B274FFA6A178FF66DD6311ECF9B1" bold="true" box="[151,467,1618,1644]" pageId="24" pageNumber="175">
<taxonomicName id="4C3715E5FFA6A178FF66DD6310CAF9B1" box="[151,245,1618,1644]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="genus">
<emphasis id="B943B274FFA6A178FF66DD6310CAF9B1" bold="true" box="[151,245,1618,1644]" italics="true" pageId="24" pageNumber="175">Cenolia</emphasis>
</taxonomicName>
AH
<bibRefCitation id="EFA61397FFA6A178FEC1DD6311ECF9B1" author="Clark" box="[304,467,1618,1644]" pageId="24" pageNumber="175" refString="Clark, A. H. (1916 b) The first New Zealand crinoid. Proceedings of the Biological Society of Washington, 29, 48." type="journal article" year="1916" yearSuffix="b">Clark, 1916b</bibRefCitation>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B886E66FFA6A178FF66DD4510D7F950" blockId="24.[151,467,1618,1677]" box="[151,232,1652,1677]" pageId="24" pageNumber="175">
<tableCitation id="C6B55BDDFFA6A178FF66DD4510D7F950" box="[151,232,1652,1677]" captionStart="TABLE 1" captionStartId="2.[125,197,537,558]" captionText="TABLE 1. Diagnostic morphological characteristics of genera of Comatulidae." pageId="24" pageNumber="175">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA6A178FF66DD8411E5F8E6" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="8B886E66FFA6A178FF66DD8412F3F913" blockId="24.[151,1436,1717,2030]" box="[151,716,1717,1742]" pageId="24" pageNumber="175">
<emphasis id="B943B274FFA6A178FF66DD84110DF913" bold="true" box="[151,306,1717,1742]" pageId="24" pageNumber="175">Type species.</emphasis>
<taxonomicName id="4C3715E5FFA6A178FEC9DD8712F8F913" authority="Muller, 1846" authorityName="Muller" authorityYear="1846" box="[312,711,1717,1742]" class="Crinoidea" family="Comasteridae" genus="Comatula" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="trichoptera">
<emphasis id="B943B274FFA6A178FEC9DD871213F913" box="[312,556,1718,1742]" italics="true" pageId="24" pageNumber="175">Comatula trichoptera</emphasis>
<bibRefCitation id="EFA61397FFA6A178FDC2DD8412F8F913" author="Muller" box="[563,711,1717,1742]" pageId="24" pageNumber="175" refString="Muller, J. (1846) Nachtrag zu der Abhandlungen uber die Comatulen. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglichen. Preussische Akademie der Wissenschaften zu Berlin, 1846, 177 - 179." type="journal article" year="1846">Müller, 1846</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B886E66FFA6A178FF36DDEB11E5F8E6" blockId="24.[151,1436,1717,2030]" pageId="24" pageNumber="175">
<emphasis id="B943B274FFA6A178FF36DDEB11D0F92E" bold="true" box="[199,495,1754,1779]" pageId="24" pageNumber="175">Other included taxa (5).</emphasis>
<taxonomicName id="4C3715E5FFA6A178FE08DDEB13A9F92F" authority="Messing, 2003" authorityName="Messing" authorityYear="2003" box="[505,918,1754,1779]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="amezianeae">
<emphasis id="B943B274FFA6A178FE08DDEB12DFF92F" box="[505,736,1754,1778]" italics="true" pageId="24" pageNumber="175">Cenolia amezianeae</emphasis>
Messing, 2003
</taxonomicName>
,
<taxonomicName id="4C3715E5FFA6A178FC55DDEB1133F8CB" authority="Rowe, Hoggett, Birtles &amp; Vail, 1986" authorityName="Rowe, Hoggett, Birtles &amp; Vail" authorityYear="1986" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="glebosus">
<emphasis id="B943B274FFA6A178FC55DDEB1456F92F" box="[932,1129,1754,1778]" italics="true" pageId="24" pageNumber="175">Cenolia glebosus</emphasis>
<bibRefCitation id="EFA61397FFA6A178FB82DDEA1133F8CB" author="Rowe" pageId="24" pageNumber="175" refString="Rowe, F. W. E., Hoggett, A. K., Birtles, R. A. &amp; Vail, L. L. (1986) Revision of some comasterid genera from Australia (Echinodermata: Crinoidea), with descriptions of two new genera and nine new species. Zoological Journal of the Linnean Society, 86, 197 - 277." type="journal article" year="1986">Rowe, Hoggett, Birtles &amp; Vail, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3715E5FFA6A178FEE6DDCF1266F8CB" box="[279,601,1790,1814]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="novaezealandiae">
<emphasis id="B943B274FFA6A178FEE6DDCF1266F8CB" box="[279,601,1790,1814]" italics="true" pageId="24" pageNumber="175">Comanthus novaezealandiae</emphasis>
</taxonomicName>
AH
<bibRefCitation id="EFA61397FFA6A178FD7EDDCC131CF8CB" author="Clark" box="[655,803,1789,1814]" pageId="24" pageNumber="175" refString="Clark, A. H. (1918 b) A new crinoid from New Zealand, and another from Tasmania. Proceedings of the Biological Society of Washington, 31, 41 - 44." type="journal article" year="1918" yearSuffix="b">Clark, 1918b</bibRefCitation>
,
<taxonomicName id="4C3715E5FFA6A178FCDFDDCF1466F8CB" box="[814,1113,1790,1814]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="spanoschistum">
<emphasis id="B943B274FFA6A178FCDFDDCF1466F8CB" box="[814,1113,1790,1814]" italics="true" pageId="24" pageNumber="175">Comanthus spanoschistum</emphasis>
</taxonomicName>
HL Clark, 1916,
<taxonomicName id="4C3715E5FFA6A178FAEDDDCF1135F8E7" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="tasmaniae">
<emphasis id="B943B274FFA6A178FAEDDDCF1135F8E7" italics="true" pageId="24" pageNumber="175">Comanthus tasmaniae</emphasis>
</taxonomicName>
AH
<bibRefCitation id="EFA61397FFA6A178FEB1DC1311E9F8E6" author="Clark" box="[320,470,1826,1851]" pageId="24" pageNumber="175" refString="Clark, A. H. (1918 b) A new crinoid from New Zealand, and another from Tasmania. Proceedings of the Biological Society of Washington, 31, 41 - 44." type="journal article" year="1918" yearSuffix="b">Clark, 1918b</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA6A178FF36DC741234F833" pageId="24" pageNumber="175" type="diagnosis">
<paragraph id="8B886E66FFA6A178FF36DC741234F833" blockId="24.[151,1436,1717,2030]" pageId="24" pageNumber="175">
<emphasis id="B943B274FFA6A178FF36DC74117EF883" bold="true" box="[199,321,1861,1886]" pageId="24" pageNumber="175">Diagnosis.</emphasis>
Mouth excentric in fully developed individuals; up to 41 arms; centrodorsal circular to pentagonal, cirri always present; IBr2 united by synarthry; IIBr and subsequent brachitaxes mostly 4(3+4); first syzygy at 3+4 on all undivided arms; distal intersyzygial interval 4, or 46 (
<taxonomicName id="4C3715E5FFA6A178FCB0DCBF141BF87B" box="[833,1060,1934,1958]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="amezianeae">
<emphasis id="B943B274FFA6A178FCB0DCBF141BF87B" box="[833,1060,1934,1958]" italics="true" pageId="24" pageNumber="175">Cenolia amezianeae</emphasis>
</taxonomicName>
); distalmost pinnule comb on P2, or P19 (
<taxonomicName id="4C3715E5FFA6A178FF01DC8311AAF817" box="[240,405,1970,1994]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="24" pageNumber="175" phylum="Echinodermata" rank="species" species="amezianeae">
<emphasis id="B943B274FFA6A178FF01DC8311AAF817" box="[240,405,1970,1994]" italics="true" pageId="24" pageNumber="175">C. amezianeae</emphasis>
</taxonomicName>
); comb arising gradually and tapering to a blunt point; teeth paired, equally-sized, confluent and/or forming transverse ridges.
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA7A179FF36DBA61430FF2A" pageId="25" pageNumber="176" type="distribution">
<paragraph id="8B886E66FFA7A179FF36DBA61430FF2A" blockId="25.[151,1436,151,392]" pageId="25" pageNumber="176">
<emphasis id="B943B274FFA7A179FF36DBA6115FFF6D" bold="true" box="[199,352,151,176]" pageId="25" pageNumber="176">Distribution.</emphasis>
<collectingRegion id="49F3A084FFA7A179FE9CDBA611EDFF6D" box="[365,466,151,176]" country="Sri Lanka" name="Southern" pageId="25" pageNumber="176">Southern</collectingRegion>
and eastern
<collectingCountry id="F3202EF6FFA7A179FD83DBA612E6FF6D" box="[626,729,151,176]" name="Australia" pageId="25" pageNumber="176">Australia</collectingCountry>
(Trigg I., WA, to Mooloolaba, QLD, including
<collectingRegion id="49F3A084FFA7A179FAD0DBA615AEFF6D" box="[1313,1425,151,176]" country="Australia" name="Tasmania" pageId="25" pageNumber="176">Tasmania</collectingRegion>
),
<collectingRegion id="49F3A084FFA7A179FF66DB8D10CFFF08" box="[151,240,188,213]" country="United Kingdom" name="Norfolk" pageId="25" pageNumber="176">Norfolk</collectingRegion>
I., Lord Howe I.,
<collectingCountry id="F3202EF6FFA7A179FE23DB8C124FFF08" box="[466,624,188,213]" name="New Zealand" pageId="25" pageNumber="176">
New
<collectingRegion id="49F3A084FFA7A179FDE2DB8D124FFF08" box="[531,624,188,213]" country="Netherlands" name="Zeeland" pageId="25" pageNumber="176">Zealand</collectingRegion>
</collectingCountry>
, and (
<taxonomicName id="4C3715E5FFA7A179FD4CDB8D1359FF09" box="[701,870,188,212]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="176" phylum="Echinodermata" rank="species" species="amezianeae">
<emphasis id="B943B274FFA7A179FD4CDB8D1359FF09" box="[701,870,188,212]" italics="true" pageId="25" pageNumber="176">C. amezianeae</emphasis>
</taxonomicName>
)
<collectingCountry id="F3202EF6FFA7A179FC8ADB8C1411FF08" box="[891,1070,188,213]" name="New Caledonia" pageId="25" pageNumber="176">New Caledonia</collectingCountry>
and
<collectingCountry id="F3202EF6FFA7A179FB9FDB8C14F5FF09" box="[1134,1226,189,212]" name="Vanuatu" pageId="25" pageNumber="176">Vanuatu</collectingCountry>
(AH
<bibRefCitation id="EFA61397FFA7A179FAE3DB8D15A8FF09" author="Clark" box="[1298,1431,188,213]" pageId="25" pageNumber="176" refString="Clark, A. H. (1931) A monograph of the existing crinoids 1 (3). Bulletin of the United States National Museum, 82, 1 - 916." type="journal article" year="1931">Clark 1931</bibRefCitation>
; Messing 2003; Rowe
<emphasis id="B943B274FFA7A179FE7DDBD011FBFF25" box="[396,452,224,248]" italics="true" pageId="25" pageNumber="176">et al.</emphasis>
1986;
<bibRefCitation id="EFA61397FFA7A179FDE3DBD112C5FF25" author="Rowe" box="[530,762,223,248]" pageId="25" pageNumber="176" refString="Rowe, F. W. E. &amp; Gates, J. (1995) Echinodermata. Zoological Catalogue of Australia. CSIRO, Melbourne, 510 pp." type="book" year="1995">Rowe &amp; Gates 1995</bibRefCitation>
). Depth range:
<quantity id="4CCFC383FFA7A179FC54DBD11434FF2A" box="[933,1035,224,248]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="3.1" metricValueMin="0.0" pageId="25" pageNumber="176" unit="m" value="155.0" valueMax="310.0" valueMin="0.0">
0
<specimenCount id="9D31A5EFFFA7A179FC30DBD11434FF2A" box="[961,1035,224,248]" pageId="25" pageNumber="176" type="generic">310 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA7A179FF36DA3511D5FE55" pageId="25" pageNumber="176" type="discussion">
<paragraph id="8B886E66FFA7A179FF36DA3511D5FE55" blockId="25.[151,1436,151,392]" pageId="25" pageNumber="176">
<materialsCitation id="3B5F643BFFA7A179FF36DA3511D5FE55" ID-GBIF-Occurrence="1503125126" collectorName="Remarks. A" country="Vanuatu" location="southern New Caledonia" pageId="25" pageNumber="176" specimenCount="1">
<collectorName id="26C20BB0FFA7A179FF36DA351166FEC1" box="[199,345,260,285]" pageId="25" pageNumber="176">
<emphasis id="B943B274FFA7A179FF36DA351104FEC0" bold="true" box="[199,315,260,285]" pageId="25" pageNumber="176">Remarks.</emphasis>
A
</collectorName>
revision of all current
<taxonomicName id="4C3715E5FFA7A179FD9BDA3512FCFEC1" box="[618,707,260,284]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="176" phylum="Echinodermata" rank="genus">
<emphasis id="B943B274FFA7A179FD9BDA3512FCFEC1" box="[618,707,260,284]" italics="true" pageId="25" pageNumber="176">Cenolia</emphasis>
</taxonomicName>
species is currently underway (Wilson
<emphasis id="B943B274FFA7A179FB61DA341517FEC1" box="[1168,1320,260,284]" italics="true" pageId="25" pageNumber="176">et al. in prep</emphasis>
).
<emphasis id="B943B274FFA7A179FAB2DA35111DFEE2" italics="true" pageId="25" pageNumber="176">
<taxonomicName id="4C3715E5FFA7A179FAB2DA351121FEE2" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="176" phylum="Echinodermata" rank="species" species="amezianeae">Cenolia amezianeae</taxonomicName>
,
</emphasis>
known from
<location id="8EE838BDFFA7A179FE39DA1612D7FE9D" LSID="urn:lsid:plazi:treatment:039EDF70FFA6A179FF66DD6311D5FE55:8EE838BDFFA7A179FE39DA1612D7FE9D" box="[456,744,295,320]" country="Vanuatu" name="southern New Caledonia" pageId="25" pageNumber="176">southern New Caledonia</location>
and
<collectingCountry id="F3202EF6FFA7A179FCD9DA1913BBFEE2" box="[808,900,296,319]" name="Vanuatu" pageId="25" pageNumber="176">Vanuatu</collectingCountry>
(Messing 2003), is the only tropical
<taxonomicName id="4C3715E5FFA7A179FAB2DA1915A3FE9D" box="[1347,1436,296,320]" class="Crinoidea" family="Comasteridae" genus="Cenolia" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="176" phylum="Echinodermata" rank="genus">
<emphasis id="B943B274FFA7A179FAB2DA1915A3FE9D" box="[1347,1436,296,320]" italics="true" pageId="25" pageNumber="176">Cenolia</emphasis>
</taxonomicName>
species, the only one with pinnule combs distal to P2, and is the only
<taxonomicName id="4C3715E5FFA7A179FC2ADA7D145BFEB8" box="[987,1124,332,357]" class="Crinoidea" family="Comatulidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="176" phylum="Echinodermata" rank="tribe" tribe="Comasterini">Comasterini</taxonomicName>
with a distal intersyzygial interval of 46 (rarely 3, 7, 9).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>