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<mods:title id="5641B62E8AA7521674B89D66B355DAD0">The genera and species of Comatulidae (Comatulida: Crinoidea): taxonomic revisions and a molecular and morphological guide</mods:title>
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<mods:namePart id="09EA05773F25B0838B3335A66431D32B">Summers, Mindi M.</mods:namePart>
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<treatment id="039EDF70FFA3A17EFF66DDAB1447FDEC" ID-DOI="http://doi.org/10.5281/zenodo.6009166" ID-GBIF-Taxon="130809851" ID-Zenodo-Dep="6009166" LSID="urn:lsid:plazi:treatment:039EDF70FFA3A17EFF66DDAB1447FDEC" httpUri="http://treatment.plazi.org/id/039EDF70FFA3A17EFF66DDAB1447FDEC" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180">
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<taxonomicName id="4C3715E5FFA3A17DFF66DDAB12DEF968" authority="Gislen, 1922" authorityName="Gislen" authorityYear="1922" box="[151,737,1690,1717]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA3A17DFF66DDAB12DEF968" bold="true" box="[151,737,1690,1717]" pageId="29" pageNumber="180">
<emphasis id="B943B274FFA3A17DFF66DDAB1212F969" bold="true" box="[151,557,1690,1716]" italics="true" pageId="29" pageNumber="180">Clarkcomanthus comanthipinnus</emphasis>
(
<bibRefCitation id="EFA61397FFA3A17DFDCDDDAA12E6F968" author="Gislen" box="[572,729,1691,1717]" pageId="29" pageNumber="180" refString="Gislen, T. (1922) The crinoids from Dr. S. Bock's expedition to Japan 1914. Nova Acta Societatis Scientiarum Upsaliensis, 5, 1 - 183." type="journal article" year="1922">Gislén, 1922</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
<paragraph id="8B886E66FFA3A17DFF66DD8C1124F90B" blockId="29.[151,737,1690,1750]" box="[151,283,1725,1750]" pageId="29" pageNumber="180">
<figureCitation id="130C72E3FFA3A17DFF66DD8C10DCF90B" box="[151,227,1725,1750]" captionStart="FIGURE 13" captionStartId="27.[151,250,1675,1697]" captionTargetBox="[252,1326,193,1632]" captionTargetId="figure@27.[252,1326,193,1632]" captionTargetPageId="27" captionText="FIGURE 13. Clarkcomanthus. (A J) Clarkcomanthus albinotus— (A) live, oral view following extraction from rubble; (B) semicryptic under ledge; (C H) aboral views; (I J) short combs tapering near tip, with J showing paired confluent comb teeth. (K L) Clarkcomanthus luteofuscum, aboral surface showing rounded centrodorsal with row of cirri. (M N) Clarkcomanthus cf. littoralis — (M) aboral surface showing centrodorsal with no cirri; (N) comb with separated confluent teeth, smaller near tip. (O R) Clarkcomanthus comanthipinnus— (O, P) live, oral view following extraction from rubble; (Q) aboral surfaces showing rounded centrodorsal with few cirri at interradials; (R) comb tapering to distal point, composed of separated nonconfluent teeth. (S T) Clarkcomanthus mirus— (S) aboral surface showing centrodorsal with cirrus buds at interradials; (T) comb tapering to blunt tip, composed of separated nonconfluent teeth. Madang, Papua New Guinea: A. MNHN-IE- 2013 - 8060; B, C. MNHN-IE- 2013 - 8122; D. MNHN-IE- 2013 - 8068; K. MNHN-IE- 2013 - 8184; O. MNHN-IE- 2013 - 8082; P. MNHN-IE- 2013 - 8151. Raja Ampat, Indonesia: E. SIO-BIC E 6186; F. SIO-BIC E 6187; G, I. SIO-BIC E 6190; L. SIO-BIC E 6210; M, N. SIO-BIC E 6193. South Island, near Lizard Island, Australia: H, J. AM J 17360; holotype. Lizard Island, Australia: Q-R. AM J 16992, holotype; S, T. AM J 17202, holotype." httpUri="https://zenodo.org/record/580186/files/figure.png" pageId="29" pageNumber="180">Fig. 13</figureCitation>
OR
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA3A17DFF66DC37138AF869" pageId="29" pageNumber="180" type="reference_group">
<paragraph id="8B886E66FFA3A17DFF66DC371222F8C6" blockId="29.[151,1436,1797,2034]" box="[151,541,1797,1819]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DC371222F8C6" box="[151,541,1797,1819]" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DC371226F8C6" ID-CoL="fcfad083-5626-4c80-bbf4-09ee0c1d50c0" authority="Bell, 1884: 167" authorityName="Bell" authorityPageNumber="167" authorityYear="1884" box="[151,537,1797,1819]" class="Crinoidea" family="Comatulidae" genus="Actinometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="cumingi">
<emphasis id="B943B274FFA3A17DFF66DC371153F8C6" box="[151,364,1798,1819]" italics="true" pageId="29" pageNumber="180">Actinometra cumingi</emphasis>
:
<treatmentCitation id="0A964877FFA3A17DFE88DC341226F8C6" author="Bell" box="[377,537,1797,1819]" page="167" pageId="29" pageNumber="180" year="1884">
<bibRefCitation id="EFA61397FFA3A17DFE88DC3411D9F8C6" author="Bell" box="[377,486,1797,1819]" pageId="29" pageNumber="180" refString="Bell, F. J. (1884) Echinodermata. In: Coppinger, R. W. (Ed.), Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. &quot; Alert &quot;, 1881 - 2. Taylor &amp; Francis, London, pp. 117 - 177." type="book chapter" year="1884">Bell, 1884</bibRefCitation>
: 167
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</taxonomicName>
.
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</paragraph>
<paragraph id="8B886E66FFA3A17DFF66DC141472F8E7" blockId="29.[151,1436,1797,2034]" box="[151,1101,1828,1850]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DC141472F8E7" box="[151,1101,1828,1850]" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DC14114EF8E7" box="[151,369,1829,1850]" class="Crinoidea" family="Comatulidae" genus="Actinometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="intricata">
<emphasis id="B943B274FFA3A17DFF66DC14114EF8E7" box="[151,369,1829,1850]" italics="true" pageId="29" pageNumber="180">Actinometra intricata</emphasis>
</taxonomicName>
(Lütken MS):
<treatmentCitation id="0A964877FFA3A17DFDFBDC1512E4F8E7" author="Schmeltz" box="[522,731,1828,1850]" page="190" pageId="29" pageNumber="180" year="1874">
<bibRefCitation id="EFA61397FFA3A17DFDFBDC151294F8E7" author="Schmeltz" box="[522,683,1828,1850]" pageId="29" pageNumber="180" refString="Schmeltz, J. D. E. (1874) Museum Godeffroy. Catalog V. Nebst einer Beilage enthaltend topographische und zoologische Notizen. L. Friedrichsen &amp; Co., Hamburg, 215 pp." type="book" year="1874">Schmeltz, 1874</bibRefCitation>
: 190
</treatmentCitation>
(part); AH
<treatmentCitation id="0A964877FFA3A17DFCA3DC151438F8E7" author="Clark" box="[850,1031,1828,1850]" page="441" pageId="29" pageNumber="180" year="1908">
<bibRefCitation id="EFA61397FFA3A17DFCA3DC1513E7F8E7" author="Clark" box="[850,984,1828,1850]" pageId="29" pageNumber="180" refString="Clark, A. H. (1908 c) The nomenclature of the recent crinoids. Proceedings of the United States National Museum, 34, 435 - 542. https: // doi. org / 10.5479 / si. 00963801.34 - 1623.435" type="journal article" year="1908" yearSuffix="c">Clark, 1908c</bibRefCitation>
: 441
</treatmentCitation>
(part).
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</paragraph>
<paragraph id="8B886E66FFA3A17DFF66DC7213FBF884" blockId="29.[151,1436,1797,2034]" box="[151,964,1859,1881]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DC7213FBF884" box="[151,964,1859,1881]" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DC721147F884" ID-CoL="d101e272-8efa-40f9-9fbc-d2f1ce40cbab" box="[151,376,1859,1881]" class="Crinoidea" family="Comatulidae" genus="Actinometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="mutabilis">
<emphasis id="B943B274FFA3A17DFF66DC721147F884" box="[151,376,1859,1881]" italics="true" pageId="29" pageNumber="180">Actinometra mutabilis</emphasis>
</taxonomicName>
(Lütken MS):
<treatmentCitation id="0A964877FFA3A17DFDE1DC7212EEF884" author="Hartlaub" box="[528,721,1859,1881]" page="98" pageId="29" pageNumber="180" year="1891">
<bibRefCitation id="EFA61397FFA3A17DFDE1DC721294F884" author="Hartlaub" box="[528,683,1859,1881]" pageId="29" pageNumber="180" refString="Hartlaub, C. (1891) Beitrag zur Kenntniss der Comatulidenfauna des Indischen Archipels. Nova Acta der Kaiserlische Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher, 58, 5 - 120." type="journal article" year="1891">Hartlaub, 1891</bibRefCitation>
: 98
</treatmentCitation>
; AH
<treatmentCitation id="0A964877FFA3A17DFCF7DC721380F884" author="Clark" box="[774,959,1859,1881]" page="444" pageId="29" pageNumber="180" year="1908">
<bibRefCitation id="EFA61397FFA3A17DFCF7DC7213B3F884" author="Clark" box="[774,908,1859,1881]" pageId="29" pageNumber="180" refString="Clark, A. H. (1908 c) The nomenclature of the recent crinoids. Proceedings of the United States National Museum, 34, 435 - 542. https: // doi. org / 10.5479 / si. 00963801.34 - 1623.435" type="journal article" year="1908" yearSuffix="c">Clark, 1908c</bibRefCitation>
: 444
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B886E66FFA3A17DFF66DC531141F84B" blockId="29.[151,1436,1797,2034]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DC531141F84B" infered="true" pageId="29" pageNumber="180">
<emphasis id="B943B274FFA3A17DFF66DC53109FF8AA" box="[151,160,1890,1911]" italics="true" pageId="29" pageNumber="180">?</emphasis>
<taxonomicName id="4C3715E5FFA3A17DFF51DC53124AF8AA" ID-CoL="8G2RS" authority="Carpenter, 1879: 53" authorityName="Carpenter" authorityPageNumber="53" authorityYear="1879" box="[160,629,1889,1911]" class="Crinoidea" family="Comatulidae" genus="Actinometra" higherTaxonomySource="CoL" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="polymorpha">
<emphasis id="B943B274FFA3A17DFF51DC53119FF8AA" box="[160,416,1889,1911]" italics="true" pageId="29" pageNumber="180">Actinometra polymorpha</emphasis>
:
<treatmentCitation id="0A964877FFA3A17DFE5CDC50124AF8AA" author="Carpenter" box="[429,629,1889,1911]" page="53" pageId="29" pageNumber="180" year="1879">
<bibRefCitation id="EFA61397FFA3A17DFE5CDC50126DF8AA" author="Carpenter" box="[429,594,1889,1911]" pageId="29" pageNumber="180" refString="Carpenter, P. H. (1879) On the genus Actinometra, Muller, with a morphological account of a new species, A. polymorpha, from the Philippine Islands. Part I. Transactions of the Linnean Society of London, 2, 1 - 122." type="journal article" year="1879">Carpenter, 1879</bibRefCitation>
: 53
</treatmentCitation>
</taxonomicName>
(part; var. 4, pl. 1, fig. 16, pl. 2, fig. 7, pl. 3, fig. 3, pl. 6, figs. 23, 24); AH
<treatmentCitation id="0A964877FFA3A17DFAACDC501107F84B" author="Clark" page="446" pageId="29" pageNumber="180" year="1908">
<bibRefCitation id="EFA61397FFA3A17DFAACDC501137F84B" author="Clark" pageId="29" pageNumber="180" refString="Clark, A. H. (1908 c) The nomenclature of the recent crinoids. Proceedings of the United States National Museum, 34, 435 - 542. https: // doi. org / 10.5479 / si. 00963801.34 - 1623.435" type="journal article" year="1908" yearSuffix="c">Clark, 1908c</bibRefCitation>
: 446
</treatmentCitation>
(part).
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</paragraph>
<paragraph id="8B886E66FFA3A17DFF66DCAF138AF869" blockId="29.[151,1436,1797,2034]" box="[151,949,1950,1973]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DCAF138AF869" box="[151,949,1950,1973]" pageId="29" pageNumber="180">
<emphasis id="B943B274FFA3A17DFF66DCAF1244F869" box="[151,635,1950,1973]" italics="true" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DCAF11F6F868" box="[151,457,1950,1973]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="parvicirra" subGenus="Vania">Comanthus (Vania) parvicirra</taxonomicName>
ß comanthipinna
</emphasis>
:
<treatmentCitation id="0A964877FFA3A17DFD79DCAF130DF869" author="Gislen" box="[648,818,1950,1972]" page="51" pageId="29" pageNumber="180" year="1922">
<bibRefCitation id="EFA61397FFA3A17DFD79DCAF1332F869" author="Gislen" box="[648,781,1950,1972]" pageId="29" pageNumber="180" refString="Gislen, T. (1922) The crinoids from Dr. S. Bock's expedition to Japan 1914. Nova Acta Societatis Scientiarum Upsaliensis, 5, 1 - 183." type="journal article" year="1922">Gislén, 1922</bibRefCitation>
: 51
</treatmentCitation>
, figs. 44, 45.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA3A17DFF66DC8C129CF80E" box="[151,675,1981,2003]" pageId="29" pageNumber="180" type="description">
<paragraph id="8B886E66FFA3A17DFF66DC8C129CF80E" blockId="29.[151,1436,1797,2034]" box="[151,675,1981,2003]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DC8C129CF80E" box="[151,675,1981,2003]" pageId="29" pageNumber="180">
<emphasis id="B943B274FFA3A17DFF66DC8C11BEF80E" box="[151,385,1981,2003]" italics="true" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DC8C1144F80E" box="[151,379,1981,2003]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="parvicirra">Comanthus parvicirra</taxonomicName>
:
</emphasis>
AH
<treatmentCitation id="0A964877FFA3A17DFE43DC8C1262F80E" author="Clark" box="[434,605,1981,2003]" page="631" pageId="29" pageNumber="180" year="1931">
<bibRefCitation id="EFA61397FFA3A17DFE43DC8C1212F80E" author="Clark" box="[434,557,1981,2003]" pageId="29" pageNumber="180" refString="Clark, A. H. (1931) A monograph of the existing crinoids 1 (3). Bulletin of the United States National Museum, 82, 1 - 916." type="journal article" year="1931">Clark, 1931</bibRefCitation>
: 631
</treatmentCitation>
(part).
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</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA3A17EFF66DCED12D3FF11" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180" type="reference_group">
<paragraph id="8B886E66FFA3A17DFF66DCED1401F82F" blockId="29.[151,1436,1797,2034]" box="[151,1086,2012,2034]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="AB274948FFA3A17DFF66DCED1401F82F" box="[151,1086,2012,2034]" pageId="29" pageNumber="180">
<emphasis id="B943B274FFA3A17DFF66DCED11B4F82F" box="[151,395,2012,2034]" italics="true" pageId="29" pageNumber="180">
<taxonomicName id="4C3715E5FFA3A17DFF66DCED11BAF82F" box="[151,389,2012,2034]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="parvicirrus">Comanthus parvicirrus</taxonomicName>
:
</emphasis>
AM
<treatmentCitation id="0A964877FFA3A17DFE31DCED1290F82F" author="Clark" box="[448,687,2012,2034]" page="6" pageId="29" pageNumber="180" year="1971">
<bibRefCitation id="EFA61397FFA3A17DFE31DCED12A8F82F" author="Clark" box="[448,663,2012,2034]" pageId="29" pageNumber="180" refString="Clark, A. M. &amp; Rowe, F. W. E. (1971) Shallow-water Indo-West Pacific Echinoderms. Trustees of the British Museum (Natural History), London, 238 pp." type="book" year="1971">Clark &amp; Rowe, 1971</bibRefCitation>
: 6
</treatmentCitation>
, 16 (part); AM
<treatmentCitation id="0A964877FFA3A17DFCBFDCED13C7F82F" author="Clark" box="[846,1016,2012,2034]" page="395" pageId="29" pageNumber="180" year="1975">
<bibRefCitation id="EFA61397FFA3A17DFCBFDCED13F7F82F" author="Clark" box="[846,968,2012,2034]" pageId="29" pageNumber="180" refString="Clark, A. M. (1975) The Swain Reefs expedition: Crinoidea. Records of the Australian Museum, 29, 391 - 406. https: // doi. org / 10.3853 / j. 0067 - 1975.29.1975.189" type="journal article" year="1975">Clark, 1975</bibRefCitation>
: 395
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B886E66FFA0A17EFF66DBA912D3FF11" blockId="30.[151,938,152,205]" pageId="30" pageNumber="181">
<taxonomicName id="4C3715E5FFA0A17EFF66DBA912F4FF73" authority="Rowe et al., 1986: 249" authorityName="Rowe et al." authorityPageNumber="249" authorityYear="1986" box="[151,715,152,174]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA0A17EFF66DBA911E1FF73" box="[151,478,152,174]" italics="true" pageId="30" pageNumber="181">Oxycomanthus comanthipinnus:</emphasis>
Rowe
<emphasis id="B943B274FFA0A17EFDD4DBA81266FF73" box="[549,601,152,174]" italics="true" pageId="30" pageNumber="181">et al.</emphasis>
, 1986: 249
</taxonomicName>
250, figs. 3A, 8G, H.
<taxonomicName id="4C3715E5FFA0A17EFF66DB871261FF11" authority="Rowe et al., 1986: 251" authorityName="Rowe et al." authorityPageNumber="251" authorityYear="1986" box="[151,606,182,205]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="exilis">
<emphasis id="B943B274FFA0A17EFF66DB87114EFF11" box="[151,369,182,204]" italics="true" pageId="30" pageNumber="181">Oxycomanthus exilis:</emphasis>
Rowe
<emphasis id="B943B274FFA0A17EFE48DB8911D3FF11" box="[441,492,182,205]" italics="true" pageId="30" pageNumber="181">et al.</emphasis>
, 1986: 251
</taxonomicName>
254, fig. 9A.
</paragraph>
</subSubSection>
<subSubSection id="C32D3DEDFFA0A17EFF66DBC81447FDEC" pageId="30" pageNumber="181" type="discussion">
<paragraph id="8B886E66FFA0A17EFF66DBC81447FDEC" blockId="30.[151,1436,249,562]" pageId="30" pageNumber="181">
<emphasis id="B943B274FFA0A17EFF66DBC81134FECF" bold="true" box="[151,267,249,274]" pageId="30" pageNumber="181">Remarks.</emphasis>
<taxonomicName id="4C3715E5FFA0A17EFEE4DBC812C4FECF" authority="Rowe et al. 1986" authorityName="Rowe et al." authorityYear="1986" box="[277,763,249,274]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="exilis">
<emphasis id="B943B274FFA0A17EFEE4DBC8122BFECC" box="[277,532,249,273]" italics="true" pageId="30" pageNumber="181">Clarkcomanthus exilis</emphasis>
(Rowe
<emphasis id="B943B274FFA0A17EFD85DBCB1290FECC" box="[628,687,249,273]" italics="true" pageId="30" pageNumber="181">et al.</emphasis>
1986)
</taxonomicName>
is synonymized under
<taxonomicName id="4C3715E5FFA0A17EFBFCDBC815A3FECC" authority="Gislen, 1922" authorityName="Gislen" authorityYear="1922" box="[1037,1436,249,274]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA0A17EFBFCDBC814D3FECC" box="[1037,1260,249,273]" italics="true" pageId="30" pageNumber="181">Cl. comanthipinnus</emphasis>
(
<bibRefCitation id="EFA61397FFA0A17EFB0FDBC815ABFECC" author="Gislen" box="[1278,1428,249,274]" pageId="30" pageNumber="181" refString="Gislen, T. (1922) The crinoids from Dr. S. Bock's expedition to Japan 1914. Nova Acta Societatis Scientiarum Upsaliensis, 5, 1 - 183." type="journal article" year="1922">Gislén, 1922</bibRefCitation>
)
</taxonomicName>
based on molecular results that indicate specimens from
<collectingCountry id="F3202EF6FFA0A17EFCFCDA2C134CFEEB" box="[781,883,285,310]" name="Australia" pageId="30" pageNumber="181">Australia</collectingCountry>
identified morphologically as the two species are really the same species. Rowe
<emphasis id="B943B274FFA0A17EFE05DA731211FE84" box="[500,558,321,345]" italics="true" pageId="30" pageNumber="181">et al.</emphasis>
(1986) originally considered the two (as
<taxonomicName id="4C3715E5FFA0A17EFBF4DA70145BFE84" box="[1029,1124,321,345]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="exilis">
<emphasis id="B943B274FFA0A17EFBF4DA70145BFE84" box="[1029,1124,321,345]" italics="true" pageId="30" pageNumber="181">O. exilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3715E5FFA0A17EFB6DDA70154CFE84" box="[1180,1395,321,345]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA0A17EFB6DDA70154CFE84" box="[1180,1395,321,345]" italics="true" pageId="30" pageNumber="181">O. comanthipinnus</emphasis>
</taxonomicName>
) as most closely related within
<taxonomicName id="4C3715E5FFA0A17EFE3CDA54124CFEA0" box="[461,627,357,381]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="genus">
<emphasis id="B943B274FFA0A17EFE3CDA54124CFEA0" box="[461,627,357,381]" italics="true" pageId="30" pageNumber="181">Oxycomanthus</emphasis>
</taxonomicName>
. The two were distinguished based on transverse bars on cirri, comb on P2, and an aboral light arm stripe in
<taxonomicName id="4C3715E5FFA0A17EFDA6DAB81307FE7C" box="[599,824,393,417]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA0A17EFDA6DAB81307FE7C" box="[599,824,393,417]" italics="true" pageId="30" pageNumber="181">Cl. comanthipinnus</emphasis>
</taxonomicName>
that are all lacking in
<taxonomicName id="4C3715E5FFA0A17EFBA0DAB81486FE7C" box="[1105,1209,393,417]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="exilis">
<emphasis id="B943B274FFA0A17EFBA0DAB81486FE7C" box="[1105,1209,393,417]" italics="true" pageId="30" pageNumber="181">Cl. exilis</emphasis>
</taxonomicName>
. Variation in cirral ornamentation may be the result of habitat differences;
<taxonomicName id="4C3715E5FFA0A17EFCD0DA9C143FFE18" box="[801,1024,429,453]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">
<emphasis id="B943B274FFA0A17EFCD0DA9C143FFE18" box="[801,1024,429,453]" italics="true" pageId="30" pageNumber="181">Cl. comanthipinnus</emphasis>
</taxonomicName>
is generally semicryptic with arms exposed, while
<taxonomicName id="4C3715E5FFA0A17EFEB8DAE01190FE34" box="[329,431,465,489]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="exilis">
<emphasis id="B943B274FFA0A17EFEB8DAE01190FE34" box="[329,431,465,489]" italics="true" pageId="30" pageNumber="181">Cl. exilis</emphasis>
</taxonomicName>
is primarily hidden under rubble during the day. In
<collectingCountry id="F3202EF6FFA0A17EFBF7DAE314DAFE37" box="[1030,1253,465,490]" name="Papua New Guinea" pageId="30" pageNumber="181">
<collectingRegion id="49F3A084FFA0A17EFBF7DAE31475FE34" box="[1030,1098,466,489]" country="Indonesia" name="Papua Barat" pageId="30" pageNumber="181">Papua</collectingRegion>
New Guinea
</collectingCountry>
, we found most juveniles under rubble and no species exclusive to rubble habitats. Descriptions of both species from
<collectingCountry id="F3202EF6FFA0A17EFADBDAC71552FDD0" box="[1322,1389,502,525]" name="Japan" pageId="30" pageNumber="181">Japan</collectingCountry>
, the
<typeStatus id="548CD0C4FFA0A17EFF66D92B10F8FDEF" box="[151,199,538,562]" pageId="30" pageNumber="181">type</typeStatus>
locality of
<emphasis id="B943B274FFA0A17EFEB6D9281215FDEC" box="[327,554,537,561]" italics="true" pageId="30" pageNumber="181">
<taxonomicName id="4C3715E5FFA0A17EFEB6D9281219FDEC" box="[327,550,537,561]" class="Crinoidea" family="Comasteridae" genus="Oxycomanthus" kingdom="Animalia" order="Comatulida" pageId="30" pageNumber="181" phylum="Echinodermata" rank="species" species="comanthipinnus">Cl. comanthipinnus</taxonomicName>
,
</emphasis>
agree with those from
<collectingCountry id="F3202EF6FFA0A17EFCDCD92813ABFDEF" box="[813,916,537,562]" name="Australia" pageId="30" pageNumber="181">Australia</collectingCountry>
(
<bibRefCitation id="EFA61397FFA0A17EFC53D928141BFDEC" author="Kogo" box="[930,1060,537,561]" pageId="30" pageNumber="181" refString="Kogo, I. (1998) Crinoids from Japan and its adjacent waters. Special Publication of the Osaka Museum of National History, 30, 1 - 148." type="journal article" year="1998">Kogo 1998</bibRefCitation>
,
<bibRefCitation id="EFA61397FFA0A17EFBDED9281453FDEC" author="Kogo" box="[1071,1132,537,561]" pageId="30" pageNumber="181" refString="Kogo, I. (2002) Report on the crinoids collected from the Nansei Islands, southern Japan, during a cruise of the training vessel Toyoshio Maru in 1999 (Crinoidea). Bulletin of the Osaka Museum of Natural History, 56, 1 - 44." type="journal article" year="2002">2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>