The genera and species of Comatulidae (Comatulida: Crinoidea): taxonomic revisions and a molecular and morphological guide Author Summers, Mindi M. Author Messing, Charles G. Author Rouse, Greg W. text Zootaxa 2017 4268 2 151 190 journal article 33045 10.11646/zootaxa.4268.2.1 e114acb7-3683-4094-a53f-c9bc7ed84353 1175-5326 580173 0742D287-B82C-4014-A6AC-C357F259D5D7 Clarkcomanthus comanthipinnus ( Gislén, 1922 ) Fig. 13 O–R Actinometra cumingi : Bell, 1884 : 167 . Actinometra intricata (Lütken MS): Schmeltz, 1874 : 190 (part); AH Clark, 1908c : 441 (part). Actinometra mutabilis (Lütken MS): Hartlaub, 1891 : 98 ; AH Clark, 1908c : 444 . ? Actinometra polymorpha : Carpenter, 1879 : 53 (part; var. 4, pl. 1, fig. 16, pl. 2, fig. 7, pl. 3, fig. 3, pl. 6, figs. 23, 24); AH Clark, 1908c : 446 (part). Comanthus (Vania) parvicirra ß comanthipinna : Gislén, 1922 : 51 , figs. 44, 45. Comanthus parvicirra : AH Clark, 1931 : 631 (part). Comanthus parvicirrus : AM Clark & Rowe, 1971 : 6 , 16 (part); AM Clark, 1975 : 395 (part). Oxycomanthus comanthipinnus: Rowe et al. , 1986: 249 –250, figs. 3A, 8G, H. Oxycomanthus exilis: Rowe et al. , 1986: 251 –254, fig. 9A. Remarks. Clarkcomanthus exilis (Rowe et al. 1986) is synonymized under Cl. comanthipinnus ( Gislén, 1922 ) based on molecular results that indicate specimens from Australia identified morphologically as the two species are really the same species. Rowe et al. (1986) originally considered the two (as O. exilis and O. comanthipinnus ) as most closely related within Oxycomanthus . The two were distinguished based on transverse bars on cirri, comb on P2, and an aboral light arm stripe in Cl. comanthipinnus that are all lacking in Cl. exilis . Variation in cirral ornamentation may be the result of habitat differences; Cl. comanthipinnus is generally semicryptic with arms exposed, while Cl. exilis is primarily hidden under rubble during the day. In Papua New Guinea , we found most juveniles under rubble and no species exclusive to rubble habitats. Descriptions of both species from Japan , the type locality of Cl. comanthipinnus , agree with those from Australia ( Kogo 1998 , 2002 ).