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<document id="91163752CD0A7D959BB49454699916CB" ID-DOI="10.1093/zoolinnean/zlad151" ID-ISSN="0024-4082" ID-Zenodo-Dep="14243740" ID-ZooBank="044A402-2A0F-4135-9410-7DE081CB11C4Corresponding" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1732840371232" checkinUser="plazi" docAuthor="Gąsiorek, Piotr, Blagden, Brian, Morek, Witold &amp; Michalczyk, Łukasz" docDate="2024" docId="03AF87C4A632FFB9AC7168DDFB248FA7" docLanguage="en" docName="zlad151.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad151" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Adropion Pilato 1987" docType="treatment" docVersion="1" lastPageNumber="12" masterDocId="FF96FFBCA639FFB2AD5E6C5BFFA28A0F" masterDocTitle="What is a strong synapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="12" updateTime="1734709154915" updateUser="julia">
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<mods:title id="FFCB7A20F867AE1F68DB369FC6DEBD04">What is a strong synapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="E612F69C9BC5A4103A91B352231A314B">Gąsiorek, Piotr</mods:namePart>
<mods:affiliation id="F28A23AD261E77AB68B74F8BA32A9DE6">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="0C060EE0866D5ABCB857565341420B56">Blagden, Brian</mods:namePart>
<mods:affiliation id="B5AABD4C6C64BA4C69C289587F387A80">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:affiliation id="D0B43F421A0165DEA4FE187789B7EDCB">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="E4732F773A3A5EDD8E18BD6538FD1765">Michalczyk, Łukasz</mods:namePart>
<mods:affiliation id="6D403FE0E2EEC99FA482889FCE729BB5">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:date id="18253F9523A926E78ED53ECD481E32BF">2024</mods:date>
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<paragraph id="8BB936D2A632FFB9AC7168DDFD9E8EAE" blockId="11.[112,764,1158,1441]" box="[303,572,1158,1185]" pageId="11" pageNumber="12">
<emphasis id="B972EAC0A632FFB9AC7168DDFD9E8EAE" bold="true" box="[303,572,1158,1185]" pageId="11" pageNumber="12">
Composition of
<taxonomicName id="4C064D51A632FFB9AC8268DDFD9E8EAE" authorityName="Pilato" authorityYear="1987" box="[476,572,1158,1185]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AC8268DDFD9E8EAE" bold="true" box="[476,572,1158,1185]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
</taxonomicName>
</emphasis>
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The genus
<taxonomicName id="4C064D51A632FFB9ADB868F6FEE68ECA" authorityName="Pilato" authorityYear="1987" box="[230,324,1197,1221]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9ADB868F6FEE68ECA" box="[230,324,1197,1221]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
</taxonomicName>
was inferred as polyphyletic by
<bibRefCitation id="EF974B23A632FFB9AFC568F6FF578EEA" author="Bertolani R &amp; Guidetti R &amp; Marchioro T" pageId="11" pageNumber="12" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
Bertolani
<emphasis id="B972EAC0A632FFB9AD2F6896FF018EEB" box="[113,163,1228,1252]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014)
</bibRefCitation>
.
<bibRefCitation id="EF974B23A632FFB9AC5F6897FDFD8EEA" author="Gasiorek P &amp; Michalczyk L" box="[257,607,1228,1253]" pageId="11" pageNumber="12" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
confirmed the non-monophyly of
<taxonomicName id="4C064D51A632FFB9AC1D68B7FE038F0B" authorityName="Pilato" authorityYear="1987" box="[323,417,1260,1284]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AC1D68B7FE038F0B" box="[323,417,1260,1284]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
</taxonomicName>
and established
<taxonomicName id="4C064D51A632FFB9AF0468B7FD668F0B" authorityName="Gasiorek &amp; Michalczyk" authorityYear="2020" box="[602,708,1260,1284]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="22" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AF0468B7FD668F0B" box="[602,708,1260,1284]" italics="true" pageId="11" pageNumber="12">Guidettion</emphasis>
</taxonomicName>
(the latter having only three bar-like macroplacoids in the pharynx). However, the genus is still heterogeneous in terms of the number of macroplacoids: the majority of species exhibit three placoids, whereas there is a small group of species with only two macroplacoids, i.e.
<taxonomicName id="4C064D51A632FFB9AC6369D2FE0B8FAF" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[317,425,1416,1440]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="belgicae">
<emphasis id="B972EAC0A632FFB9AC6369D2FE0B8FAF" box="[317,425,1416,1440]" italics="true" pageId="11" pageNumber="12">A. belgicae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C064D51A632FFB9ACE969D2FDD58FAF" authorityName="Gąsiorek &amp; Blagden &amp; Morek &amp; Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek &amp; Blagden &amp; Morek &amp; Michalczyk" baseAuthorityYear="1991" box="[439,631,1416,1440]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
<emphasis id="B972EAC0A632FFB9ACE969D2FDD58FAF" box="[439,631,1416,1440]" italics="true" pageId="11" pageNumber="12">A. diphasconiellum</emphasis>
</taxonomicName>
<emphasis id="B972EAC0A632FFB9AF2169D2FC0189FE" bold="true" box="[639,931,985,1441]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A24157BBA632FFB9AF2169D2FC0189FE" box="[639,931,985,1441]" pageId="11" pageNumber="12">comb. nov. et stat. rev.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C064D51A632FFB9AE856F82FB8089FE" box="[987,1058,985,1009]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gani">
<emphasis id="B972EAC0A632FFB9AE856F82FB8089FE" box="[987,1058,985,1009]" italics="true" pageId="11" pageNumber="12">A. gani</emphasis>
</taxonomicName>
. The adoption of molecular phylogenetic methods in tardigrade research has shown that the number and shape of placoids in the pharynx is indicative of common ancestry (e.g.
<bibRefCitation id="EF974B23A632FFB9AEE3686CFB2E8E40" author="Gasiorek P &amp; Stec D &amp; Morek W" box="[957,1164,1079,1103]" pageId="11" pageNumber="12" pagination="78 - 132" refId="ref36279" refString="Gasiorek P, Stec D, Morek W et al. Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology 2019; 88: 78 - 132. https: // doi. org / 10.1163 / 18759866 - 20191350" type="journal article" year="2019">
Gąsiorek
<emphasis id="B972EAC0A632FFB9A97E6863FBF38E40" box="[1056,1105,1079,1103]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2019
</bibRefCitation>
,
<bibRefCitation id="EF974B23A632FFB9A9C6686CFAC08E40" author="Guidetti R &amp; Giovannini I &amp; Del Papa V" box="[1176,1378,1079,1103]" pageId="11" pageNumber="12" pagination="1099 - 117" refId="ref36442" refString="Guidetti R, Giovannini I, Del Papa V et al. Phylogeny of the asexual lineage Murrayidae (Macrobiotoidea, Eutardigrada) with the description of Paramurrayon gen. nov. and Paramurrayon meieri sp. nov. Invertebrate Systematics 2022; 36: 1099 - 117. https: // doi. org / 10.1071 / IS 22031" type="journal article" year="2022">
Guidetti
<emphasis id="B972EAC0A632FFB9A9A86863FA838E40" box="[1270,1313,1079,1103]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2022
</bibRefCitation>
). Thus, it is plausible that
<taxonomicName id="4C064D51A632FFB9AEA6680CFBC58E61" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1016,1127,1110,1134]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="belgicae">
<emphasis id="B972EAC0A632FFB9AEA6680CFBC58E61" box="[1016,1127,1110,1134]" italics="true" pageId="11" pageNumber="12">A. belgicae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D51A632FFB9A9FC680CFAC68E61" authorityName="Gąsiorek &amp; Blagden &amp; Morek &amp; Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek &amp; Blagden &amp; Morek &amp; Michalczyk" baseAuthorityYear="1991" box="[1186,1380,1110,1134]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
<emphasis id="B972EAC0A632FFB9A9FC680CFAC68E61" box="[1186,1380,1110,1134]" italics="true" pageId="11" pageNumber="12">A. diphasconiellum</emphasis>
</taxonomicName>
<emphasis id="B972EAC0A632FFB9A831680CFC7D8E81" bold="true" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A24157BBA632FFB9A831680CFC7D8E81" pageId="11" pageNumber="12">comb. nov. et stat. rev.</taxonomicNameLabel>
</emphasis>
are closely related and represent a separate evolutionary lineage. If reciprocal monophyly is confirmed molecularly, they should be erected as a new genus. However,
<taxonomicName id="4C064D51A632FFB9AE72688FFCD78EE4" box="[812,885,1236,1259]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gani">
<emphasis id="B972EAC0A632FFB9AE72688FFCD78EE4" box="[812,885,1236,1259]" italics="true" pageId="11" pageNumber="12">A. gani</emphasis>
</taxonomicName>
is designated as
<emphasis id="B972EAC0A632FFB9A974688FFAA78EE4" bold="true" box="[1066,1285,1235,1259]" italics="true" pageId="11" pageNumber="12">nomen inquirendum</emphasis>
, because having two rod-like macroplacoids arranged in parentheses (i.e. as in the members of the genus
<taxonomicName id="4C064D51A632FFB9A9156949FB0D8F25" box="[1099,1199,1298,1322]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9A9156949FB0D8F25" box="[1099,1199,1298,1322]" italics="true" pageId="11" pageNumber="12">Pilatobius</emphasis>
</taxonomicName>
) it does not fit the current diagnosis of
<taxonomicName id="4C064D51A632FFB9AEB66969FBE48F45" authorityName="Pilato" authorityYear="1987" box="[1000,1094,1330,1354]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AEB66969FBE48F45" box="[1000,1094,1330,1354]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
</taxonomicName>
. Moreover, recently three species of
<taxonomicName id="4C064D51A632FFB9AE12690AFC088F66" authorityName="Pilato" authorityYear="1987" box="[844,938,1361,1385]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AE12690AFC088F66" box="[844,938,1361,1385]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
</taxonomicName>
, each with three placoids, were transferred to
<taxonomicName id="4C064D51A632FFB9AE74692BFB0B8F86" authority="(Zawierucha et al. 2023)" baseAuthorityName="Zawierucha" baseAuthorityYear="2023" box="[810,1193,1392,1417]" class="Eutardigrada" family="Hypsibiidae" genus="Kararehius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="B972EAC0A632FFB9AE74692BFC3A8F87" box="[810,920,1392,1416]" italics="true" pageId="11" pageNumber="12">Kararehius</emphasis>
(
<bibRefCitation id="EF974B23A632FFB9AEF5692BFB3F8F86" author="Zawierucha K &amp; Stec D &amp; Dearden PK" box="[939,1181,1392,1417]" pageId="11" pageNumber="12" refId="ref40518" refString="Zawierucha K, Stec D, Dearden PK et al. Two new tardigrade genera from New Zealand's Southern Alp glaciers display morphological stasis and parallel evolution. Molecular Phylogenetics and Evolution 2023; 178: 107634. https: // doi. org / 10.1016 / j. ympev. 2022.107634" type="journal volume" year="2023">
Zawierucha
<emphasis id="B972EAC0A632FFB9A971692AFBC38F87" box="[1071,1121,1392,1416]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2023
</bibRefCitation>
)
</taxonomicName>
. Their systematic affinity remains to be verified genetically.
</paragraph>
</subSubSection>
</treatment>
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