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<mods:title id="829C193FB8FF841CD961E32ED5AF0DEB">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="10827CCEBF8118D0C61FB762A54E7D68">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="3C63C887B6BFDFC381B2F2014117F068">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="1E6EE760D7BD059F8EC2FB65C88B4C2D" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A397CB277F5448AB218BC45811106DDC">Blagden, Brian</mods:namePart>
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<mods:affiliation id="A91E430822EBAA5D1A5EF630FEF4F72B">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="940BAF08DA28E6F97F145EBEF4587163">Morek, Witold</mods:namePart>
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<mods:affiliation id="221A1E6E1F4075BAC27936323F104329">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="DB8610A194260DC80E63A1652192EE4E">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="E1DE90C0C38D876414BFFF94E8B57DF6">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:title id="45102A48B955F93994B8FFB808188E96">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="57443F8B3E188282A41886A2CF71BF25">
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<mods:date id="0005ABA6E960213171C79A07B0D1A0A0">2024</mods:date>
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<mods:number id="4DFCCA31506B6B3334CBE20FC94C182B">2023-11-14</mods:number>
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<mods:identifier id="7A6FF9A50A1270BC364B797003C3EA8B" type="DOI">10.1093/zoolinnean/zlad151</mods:identifier>
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<mods:identifier id="A725F17EC33B36B272B047CDC49C651D" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="5F536A8C7480D7B22C5D8973FF78CB91" type="Zenodo-Dep">14243740</mods:identifier>
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<mods:identifier id="4D21760BF6D64EC789916F45ED981897" type="ZooBank">044A402-2A0F-4135-9410-7DE081CB11C4Corresponding</mods:identifier>
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<subSubSection id="C31C6559A600FF8BAC4D68C7FDFA8EB8" box="[275,600,1180,1207]" pageId="57" pageNumber="58" type="nomenclature">
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<paragraph id="8BB936D2A600FF8BAC4D68C7FDFA8EB8" blockId="57.[275,600,1180,1207]" box="[275,600,1180,1207]" pageId="57" pageNumber="58">
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<taxonomicName id="4C064D51A600FF8BAC4D68C7FDFA8EB8" ID-CoL="9WNCD" authority="Bertolani et al., 2014" authorityName="Bertolani" authorityYear="2014" box="[275,600,1180,1207]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
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<emphasis id="B972EAC0A600FF8BAC4D68C7FE438EB8" bold="true" box="[275,481,1180,1207]" pageId="57" pageNumber="58">
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<emphasis id="B972EAC0A600FF8BAC4D68C7FEDB8EB8" bold="true" box="[275,377,1180,1207]" italics="true" pageId="57" pageNumber="58">Pilatobius</emphasis>
|
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<bibRefCitation id="EF974B23A600FF8BAC2368C6FE438EB8" author="Bertolani R & Guidetti R & Marchioro T" box="[381,481,1181,1207]" pageId="57" pageNumber="58" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">Bertolani</bibRefCitation>
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<emphasis id="B972EAC0A600FF8BACB868C5FDBA8EB8" box="[486,536,1181,1207]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
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,
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<emphasis id="B972EAC0A600FF8BAF7F68C6FDFA8EB8" bold="true" box="[545,600,1181,1207]" pageId="57" pageNumber="58">2014</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C31C6559A600FF8BAD2F6882FD7B8D17" pageId="57" pageNumber="58" type="diagnosis">
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<paragraph id="8BB936D2A600FF8BAD2F6882FD5F8C67" blockId="57.[113,765,1241,1641]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6882FE8D8EFE" box="[113,303,1241,1265]" italics="true" pageId="57" pageNumber="58">Amended diagnosis:</emphasis>
|
||||
Cuticle uniformly sculptured throughout the dorsal part of trunk; dorsolateral gibbosities may be present. The AISMs in the shape of semilunar hooks. Buccal tube followed by a flexible annulated pharyngeal tube; the pharyngeal annulation is semi-complex (the annuli are single laterally and forking dorsoventrally). A DABT is present. A spherical pharynx with two macroplacoids and a septulum. Macroplacoids arranged rhomboidally (◊). Claws are of the
|
||||
<taxonomicName id="4C064D51A600FF8BACD169EFFE498FC3" authorityName="Ehrenberg" authorityYear="1848" box="[399,491,1460,1484]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BACD169EFFE498FC3" box="[399,491,1460,1484]" italics="true" pageId="57" pageNumber="58">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A600FF8BACAE69EFFDBF8FC3" box="[496,541,1460,1484]" pageId="57" pageNumber="58">type</typeStatus>
|
||||
, i.e. asymmetrical both with respect to the sequence of primary and secondary branches (2-1-2-1) and with respect to their size, with external and posterior claws always evidently larger than the internal and anterior claws. Pseudolunulae below claws I–IV may be present; cuticular bars of various
|
||||
<typeStatus id="54BD8870A600FF8BADE06A0AFF518C66" box="[190,243,1617,1641]" pageId="57" pageNumber="58">types</typeStatus>
|
||||
(internal, anterior, posterior, and orthogonal) present.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6AD8FD7B8D17" blockId="57.[113,763,1667,1816]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6AD8FED38C94" box="[113,369,1667,1691]" italics="true" pageId="57" pageNumber="58">
|
||||
Composition:
|
||||
<taxonomicName id="4C064D51A600FF8BADA06AD8FED38C94" baseAuthorityName="Maucci" baseAuthorityYear="1952" box="[254,369,1667,1691]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="aculeatus">P. aculeatus</taxonomicName>
|
||||
</emphasis>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BACDB6AD8FE718C94" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[389,467,1667,1691]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A600FF8BACDB6AD8FE718C94" box="[389,467,1667,1691]" italics="true" pageId="57" pageNumber="58">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAF6F6AD8FD3A8C94" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[561,664,1667,1691]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A600FF8BAF6F6AD8FD3A8C94" box="[561,664,1667,1691]" italics="true" pageId="57" pageNumber="58">P. bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAFF26AD8FD588C94" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[684,762,1667,1691]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A600FF8BAFF26AD8FD588C94" box="[684,762,1667,1691]" italics="true" pageId="57" pageNumber="58">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group,
|
||||
<typeStatus id="54BD8870A600FF8BADE76AF9FF458CB5" box="[185,231,1698,1722]" pageId="57" pageNumber="58">type</typeStatus>
|
||||
species),
|
||||
<taxonomicName id="4C064D51A600FF8BAC136AF8FE0D8CB5" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[333,431,1698,1722]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A600FF8BAC136AF8FE0D8CB5" box="[333,431,1698,1722]" italics="true" pageId="57" pageNumber="58">P. granifer</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BACE16AF8FDAA8CB5" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[447,520,1699,1722]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BACE16AF8FDAA8CB5" box="[447,520,1699,1722]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAF3F6AF8FD038CB5" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[609,673,1698,1722]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A600FF8BAF3F6AF8FD038CB5" box="[609,673,1698,1722]" italics="true" pageId="57" pageNumber="58">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAFEC6AF8FD598CB5" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[690,763,1699,1722]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAFEC6AF8FD598CB5" box="[690,763,1699,1722]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAD9B6A99FE878CD6" baseAuthorityName="Binda & Pilato" baseAuthorityYear="1971" box="[197,293,1730,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="patanei">
|
||||
<emphasis id="B972EAC0A600FF8BAD9B6A99FE878CD6" box="[197,293,1730,1753]" italics="true" pageId="57" pageNumber="58">P. patanei</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAC666A9AFE248CD6" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[312,390,1729,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A600FF8BAC666A9AFE248CD6" box="[312,390,1729,1753]" italics="true" pageId="57" pageNumber="58">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BACBC6A99FDE98CD6" box="[482,587,1730,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="procerus">
|
||||
<emphasis id="B972EAC0A600FF8BACBC6A99FDE98CD6" box="[482,587,1730,1753]" italics="true" pageId="57" pageNumber="58">P. procerus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAF006A99FD058CD6" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[606,679,1730,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAF006A99FD058CD6" box="[606,679,1730,1753]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAD2F6ABAFEA18CF7" baseAuthorityName="Robotti" baseAuthorityYear="1970" box="[113,259,1761,1784]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="ramazzottii">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6ABAFEA18CF7" box="[113,259,1761,1784]" italics="true" pageId="57" pageNumber="58">P. ramazzottii</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAC406ABAFEC58CF7" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[286,359,1761,1784]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAC406ABAFEC58CF7" box="[286,359,1761,1784]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAC8C6ABAFD9E8CF7" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[466,572,1761,1784]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAC8C6ABAFD9E8CF7" box="[466,572,1761,1784]" italics="true" pageId="57" pageNumber="58">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAF086ABAFD3D8CF7" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[598,671,1761,1784]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAF086ABAFD3D8CF7" box="[598,671,1761,1784]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A600FF8BAD2F6B5AFF518D17" box="[113,243,1792,1816]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6B5AFF518D17" box="[113,243,1792,1816]" italics="true" pageId="57" pageNumber="58">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAC5D6B5BFEF38D17" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[259,337,1792,1816]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A600FF8BAC5D6B5BFEF38D17" box="[259,337,1792,1816]" italics="true" pageId="57" pageNumber="58">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group), and
|
||||
<taxonomicName id="4C064D51A600FF8BAC8D6B5AFD8C8D17" box="[467,558,1792,1816]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="ziliense">
|
||||
<emphasis id="B972EAC0A600FF8BAC8D6B5AFD8C8D17" box="[467,558,1792,1816]" italics="true" pageId="57" pageNumber="58">P. ziliense</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A600FF8BAF616B5AFD2A8D17" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[575,648,1793,1816]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A600FF8BAF616B5AFD2A8D17" box="[575,648,1793,1816]" italics="true" pageId="57" pageNumber="58">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A600FF88AD2F6B62FA9789D8" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="discussion">
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6B62FE5B8DA0" blockId="57.[113,762,1849,1967]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6B62FF6D8D5E" box="[113,207,1849,1873]" italics="true" pageId="57" pageNumber="58">Remarks:</emphasis>
|
||||
We consider the genus
|
||||
<taxonomicName id="4C064D51A600FF8BAC866B62FD9B8D5E" box="[472,569,1849,1873]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BAC866B62FD9B8D5E" box="[472,569,1849,1873]" italics="true" pageId="57" pageNumber="58">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
as artificial (nonmonophyletic) in its present composition. Additional DNA data should contribute to disentangling its phylogenetic affinities, and the topology of the pilatobiin tree.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BAEF86CCBFA958AA5" blockId="57.[809,1461,144,1961]" box="[934,1335,144,170]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAEF86CCBFA958AA5" bold="true" box="[934,1335,144,170]" pageId="57" pageNumber="58">Problems with hypsibiid classification</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BAE746CEDFB87898F" blockId="57.[809,1461,144,1961]" pageId="57" pageNumber="58">
|
||||
This study provides redescriptions of some crucial hypsibiid species and descriptions of new species that were included in the paper devoted to hypsibiid phylogeny (
|
||||
<bibRefCitation id="EF974B23A600FF8BA9ED6CAEFCFF8B23" author="Gasiorek P & Michalczyk L" pageId="57" pageNumber="58" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk 2020</bibRefCitation>
|
||||
). We provide a good starting point for sound taxonomic research on the genera
|
||||
<taxonomicName id="4C064D51A600FF8BA97E6D6FFB518B43" authority=", Guidettion" authorityName="Guidettion" box="[1056,1267,308,332]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BA97E6D6FFBDE8B43" box="[1056,1148,308,332]" italics="true" pageId="57" pageNumber="58">Adropion</emphasis>
|
||||
,
|
||||
<emphasis id="B972EAC0A600FF8BA9D56D6FFB518B43" box="[1163,1267,308,332]" italics="true" pageId="57" pageNumber="58">Guidettion</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A600FF8BA8716D6FFA308B43" authorityName="Pilato" authorityYear="1987" box="[1327,1426,308,332]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BA8716D6FFA308B43" box="[1327,1426,308,332]" italics="true" pageId="57" pageNumber="58">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
, at the same time stressing that without a reliable redescription of
|
||||
<taxonomicName id="4C064D51A600FF8BAE726D28FC5C8B85" baseAuthorityName="Arcidiacono" baseAuthorityYear="1962" box="[812,1022,371,394]" class="Eutardigrada" family="Hypsibiidae" genus="Astatumen" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="trinacriae">
|
||||
<emphasis id="B972EAC0A600FF8BAE726D28FC5C8B85" box="[812,1022,371,394]" italics="true" pageId="57" pageNumber="58">Astatumen trinacriae</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingRegion id="49C2F830A600FF8BA91F6D29FBDB8B85" box="[1089,1145,370,394]" country="Italy" name="Sicilia" pageId="57" pageNumber="58">Sicily</collectingRegion>
|
||||
, a trustworthy species identification within that genus remains impossible. A higher-level classification of
|
||||
<taxonomicName id="4C064D51A600FF8BAE906DEAFBE88BC6" authorityName="Pilato" authorityYear="1969" box="[974,1098,433,457]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
illustrates the conflict between the morphological vs.phylogenetic approaches to tardigrade systematics, which had been ripening over the last few years and has recently erupted independently in several contributions. Although the currently recognized four hypsibiid subfamilies (
|
||||
<bibRefCitation id="EF974B23A600FF8BA8146E75FC068869" author="Pilato G & Binda MG" pageId="57" pageNumber="58" pagination="1 - 54" refId="ref38815" refString="Pilato G, Binda MG. Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification. Zootaxa 2010; 2404: 1 - 54. https: // doi. org / 10.11646 / zootaxa. 2404.1.1" type="journal article" year="2010">Pilato and Binda 2010</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A600FF8BAEEC6E15FB2B8869" author="Bertolani R & Guidetti R & Marchioro T" box="[946,1161,590,614]" pageId="57" pageNumber="58" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A600FF8BA9456E15FBEF8869" box="[1051,1101,590,614]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
) have been recovered independently as clades (
|
||||
<bibRefCitation id="EF974B23A600FF8BA95D6E36FAE6888A" author="Gasiorek P & Michalczyk L" box="[1027,1348,621,645]" pageId="57" pageNumber="58" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk 2020</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A600FF8BA80E6E36FCFC88AB" author="Tumanov DV" pageId="57" pageNumber="58" pagination="1 - 37" refId="ref40150" refString="Tumanov DV. Integrative redescription of Hypsibius pallidoides Pilato et al., 2011 (Eutardigrada: Hypsibioidea) with the erection of a new genus and discussion on the phylogeny of Hypsibiidae. European Journal of Taxonomy 2020; 681: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.681" type="journal article" year="2020">Tumanov 2020</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A600FF8BAE326ED6FB0588AB" author="Tumanov DV & Tsvetkova AY" box="[876,1191,652,676]" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A600FF8BA9EB6ED7FA0088AB" author="Zawierucha K & Stec D & Dearden PK" box="[1205,1442,652,676]" pageId="57" pageNumber="58" refId="ref40518" refString="Zawierucha K, Stec D, Dearden PK et al. Two new tardigrade genera from New Zealand's Southern Alp glaciers display morphological stasis and parallel evolution. Molecular Phylogenetics and Evolution 2023; 178: 107634. https: // doi. org / 10.1016 / j. ympev. 2022.107634" type="journal volume" year="2023">
|
||||
Zawierucha
|
||||
<emphasis id="B972EAC0A600FF8BA8686ED6FAC588AB" box="[1334,1383,652,676]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
), their current morphological diagnoses are meaningless owing to the erroneous inclusion of some taxonomic features within each clade (originally,
|
||||
<taxonomicName id="4C064D51A600FF8BAE836EB1FBF5890D" box="[989,1111,746,770]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
were defined as having an elongated pharyngeal tube,
|
||||
<taxonomicName id="4C064D51A600FF8BA9536F51FB05892D" authorityName="Rudescu" authorityYear="1964" box="[1037,1191,778,802]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
– without the DABT, and
|
||||
<taxonomicName id="4C064D51A600FF8BAE756F72FC66894E" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[811,964,809,833]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
– as having macroplacoids in the pharynx; all these characteristics have recently been demonstrated as diagnostically questionable).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BAE1B6FDCFBEF8CC1" blockId="57.[809,1461,144,1961]" pageId="57" pageNumber="58">
|
||||
Recently,
|
||||
<bibRefCitation id="EF974B23A600FF8BAEEE6FDCFAB38990" author="Tumanov DV & Tsvetkova AY" box="[944,1297,903,927]" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
proposed that
|
||||
<taxonomicName id="4C064D51A600FF8BAE746FFDFC6689B1" authorityName="Rudescu" authorityYear="1964" box="[810,964,934,958]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A600FF8BAEA86FFDFBD289B1" box="[1014,1136,934,958]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
should be elevated to the ranks of separate families within
|
||||
<taxonomicName id="4C064D51A600FF8BA9696F9DFB6189D1" authorityName="Pilato" authorityYear="1969" box="[1079,1219,966,990]" pageId="57" pageNumber="63" rank="superFamily" superFamily="Hypsibioidea">Hypsibioidea</taxonomicName>
|
||||
. However, in this paper, we have maintained the previous classification of broadly understood
|
||||
<taxonomicName id="4C064D51A600FF8BAE34685FFC4A8E13" authorityName="Pilato" authorityYear="1969" box="[874,1000,1028,1052]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
, for several reasons. The main arguments for dividing
|
||||
<taxonomicName id="4C064D51A600FF8BAED4687FFBA48E33" authorityName="Pilato" authorityYear="1969" box="[906,1030,1060,1084]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
were their morphological heterogeneity and the lack of monophyly in the molecular analyses of
|
||||
<bibRefCitation id="EF974B23A600FF8BA8116818FBB28E75" author="Tumanov DV & Tsvetkova AY" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
. However, the first reason is highly subjective (we do not consider traditionally defined
|
||||
<taxonomicName id="4C064D51A600FF8BA84268D9FA3A8E95" authorityName="Pilato" authorityYear="1969" box="[1308,1432,1154,1178]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
as more diverse than, e.g.
|
||||
<taxonomicName id="4C064D51A600FF8BA94D68FAFB0D8EB6" authorityName="Thulin" authorityYear="1928" box="[1043,1199,1185,1209]" class="Eutardigrada" family="Macrobiotidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Macrobiotidae</taxonomicName>
|
||||
), and the elevation of two of the subfamilies to the rank of families does not change the fact that none of the four hypsibiid groups has a coherent diagnosis. Moreover, neither
|
||||
<taxonomicName id="4C064D51A600FF8BAEB868A4FB228F18" authorityName="Rudescu" authorityYear="1964" box="[998,1152,1279,1303]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
nor
|
||||
<taxonomicName id="4C064D51A600FF8BA9F268A4FA848F18" box="[1196,1318,1279,1303]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
differ consistently from the clade
|
||||
<taxonomicName id="4C064D51A600FF8BA9556945FB068F39" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[1035,1188,1310,1334]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C064D51A600FF8BA99F6945FA9F8F39" authorityName="Pilato" authorityYear="1969" box="[1217,1341,1310,1334]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Hypsibiinae">Hypsibiinae</taxonomicName>
|
||||
[
|
||||
<bibRefCitation id="EF974B23A600FF8BA80E6944FBAE8F59" author="Tumanov DV & Tsvetkova AY" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
proposed a ‘transfer’ of
|
||||
<taxonomicName id="4C064D51A600FF8BA9A16965FA3A8F59" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[1279,1432,1342,1366]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
to
|
||||
<taxonomicName id="4C064D51A600FF8BAE746906FC0A8F7A" authorityName="Pilato" authorityYear="1969" box="[810,936,1373,1397]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
, which was redundant, because diphasconins had already been included in this family at that time] and between each other, which would justify the taxonomic elevation of their status. The lack of hypsibiid monophyly can be explained by the highly fragmented (e.g. only the 18S rRNA marker available for some taxa), incomplete, and taxonomically biased (many unusual hypsibiid morphotypes are still not sequenced) genetic dataset at our disposal. In short, based on the available data, we do not recognize the validity of the proposal put forward by
|
||||
<bibRefCitation id="EF974B23A600FF8BAE756A2CFBDD8C80" author="Tumanov DV & Tsvetkova AY" box="[811,1151,1655,1679]" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
. Nevertheless, we do not exclude the possibility of such moves in the future. Currently, the evidence is simply not there.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF88AE1B6A8EFD5E8830" blockId="57.[809,1461,144,1961]" lastBlockId="58.[127,780,144,1985]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||||
In concordance with
|
||||
<bibRefCitation id="EF974B23A600FF8BA9756A8EFA208CE2" author="Tumanov DV & Tsvetkova AY" box="[1067,1410,1749,1773]" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
, the presence of the DABT (= drop-like thickening
|
||||
<emphasis id="B972EAC0A600FF8BA9A96AAEFA8B8D03" box="[1271,1321,1781,1804]" italics="true" pageId="57" pageNumber="58">sensu</emphasis>
|
||||
<bibRefCitation id="EF974B23A600FF8BA8706AAFFCFF8D23" author="Guidetti R & Bertolani R & Rebecchi L" pageId="57" pageNumber="58" pagination="24 - 35" refId="ref36384" refString="Guidetti R, Bertolani R, Rebecchi L. Comparative analysis of the tardigrade feeding apparatus: adaptive convergence and evolutionary pattern of the piercing stylet system. Journal of Limnology, 2013; 72: 24 - 35. https: // doi. org / 10.4081 / jlimnol. 2013. s 1. e 4" type="journal article" year="2013">
|
||||
Guidetti
|
||||
<emphasis id="B972EAC0A600FF8BA8D96AAEFA168D03" box="[1415,1460,1780,1804]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||||
2013
|
||||
</bibRefCitation>
|
||||
/drop-like apodeme
|
||||
<emphasis id="B972EAC0A600FF8BA96B6B4FFBC58D24" box="[1077,1127,1812,1835]" italics="true" pageId="57" pageNumber="58">sensu</emphasis>
|
||||
<bibRefCitation id="EF974B23A600FF8BA9316B4FFA058D23" author="Tumanov DV & Tsvetkova AY" box="[1135,1447,1812,1836]" pageId="57" pageNumber="58" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
) can no longer serve as the main distinguishing criterion between
|
||||
<taxonomicName id="4C064D51A600FF8BAE746B09FC278D65" box="[810,901,1874,1898]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BAE746B09FC278D65" box="[810,901,1874,1898]" italics="true" pageId="57" pageNumber="58">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A600FF8BAEEB6B09FBBE8D65" authorityName="Plate" authorityYear="1888" box="[949,1052,1874,1898]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BAEEB6B09FBBE8D65" box="[949,1052,1874,1898]" italics="true" pageId="57" pageNumber="58">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
(and,
|
||||
<emphasis id="B972EAC0A600FF8BA9056B08FB168D65" box="[1115,1204,1874,1898]" italics="true" pageId="57" pageNumber="58">ipso facto</emphasis>
|
||||
, between
|
||||
<taxonomicName id="4C064D51A600FF8BA8446B09FA118D65" authorityName="Rudescu" authorityYear="1964" box="[1306,1459,1874,1898]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A600FF8BAE046B29FC558D85" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[858,1015,1906,1930]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
). We revealed that this structure can take various forms, and it should be analysed under SEM to determine its shape accurately. In this regard,
|
||||
<bibRefCitation id="EF974B23A603FF88ACBF6CCBFD148AA7" author="Dastych H & Ryan PG & Watkins BP" box="[481,694,144,168]" pageId="58" pageNumber="59" pagination="57 - 66" refId="ref35792" refString="Dastych H, Ryan PG, Watkins BP. Notes on Tardigrada from western Dronning Maud Land (Antarctica) with description of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 1990; 10: 57 - 66." type="journal article" year="1990">
|
||||
Dastych
|
||||
<emphasis id="B972EAC0A603FF88AF676CCAFDCA8AA7" box="[569,616,144,168]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
(1990)
|
||||
</bibRefCitation>
|
||||
have already noted that the straightforward distinction between the lack (
|
||||
<taxonomicName id="4C064D51A603FF88ADD56C94FF448AE8" box="[139,230,207,231]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88ADD56C94FF448AE8" box="[139,230,207,231]" italics="true" pageId="58" pageNumber="59">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
) and presence (
|
||||
<taxonomicName id="4C064D51A603FF88ACC26C94FDA18AE8" authorityName="Plate" authorityYear="1888" box="[412,515,207,231]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88ACC26C94FDA18AE8" box="[412,515,207,231]" italics="true" pageId="58" pageNumber="59">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
) of an apodeme (Pilato 1987) is contravened by the slight thickening of the buccal tube wall in
|
||||
<taxonomicName id="4C064D51A603FF88AD866D56FD878B2A" authority="Binda, 1971" authorityName="Binda" authorityYear="1971" box="[216,549,269,293]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="higginsi">
|
||||
<emphasis id="B972EAC0A603FF88AD866D56FE378B2A" box="[216,405,269,293]" italics="true" pageId="58" pageNumber="59">Diphascon higginsi</emphasis>
|
||||
Binda, 1971
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A603FF88AF666D56FEB58B4A" authority="Jennings, 1976" authorityName="Jennings" authorityYear="1976" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="puniceum">
|
||||
<emphasis id="B972EAC0A603FF88AF666D56FCAB8B2A" box="[568,777,269,293]" italics="true" pageId="58" pageNumber="59">Diphascon puniceum</emphasis>
|
||||
Jennings,
|
||||
<quantity id="4CFE9B37A603FF88ADBD6D76FEB58B4A" box="[227,279,301,325]" metricMagnitude="1" metricUnit="m" metricValue="5.0190399999999995" pageId="58" pageNumber="59" unit="in" value="1976.0">1976</quantity>
|
||||
</taxonomicName>
|
||||
in
|
||||
<bibRefCitation id="EF974B23A603FF88AC676D76FE498B4A" author="Jennings PG" box="[313,491,301,325]" pageId="58" pageNumber="59" pagination="1 - 25" refId="ref36838" refString="Jennings PG. The Tardigrada of Signy Island, South Orkney Islands, with a note on the Rotifera. British Antarctic Survey Bulletin 1976 b; 44: 1 - 25." type="journal article" year="1976">Jennings (1976b)</bibRefCitation>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A603FF88AF7A6D76FF168B6B" authority="Dastych et al., 1990" authorityName="Dastych" authorityYear="1990" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="sanae">
|
||||
<emphasis id="B972EAC0A603FF88AF7A6D76FDD58B4B" box="[548,631,301,324]" italics="true" pageId="58" pageNumber="59">D. sanae</emphasis>
|
||||
<bibRefCitation id="EF974B23A603FF88AF226D76FF168B6B" author="Dastych H & Ryan PG & Watkins BP" pageId="58" pageNumber="59" pagination="57 - 66" refId="ref35792" refString="Dastych H, Ryan PG, Watkins BP. Notes on Tardigrada from western Dronning Maud Land (Antarctica) with description of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 1990; 10: 57 - 66." type="journal article" year="1990">
|
||||
Dastych
|
||||
<emphasis id="B972EAC0A603FF88AF8B6D76FCA78B4A" box="[725,773,301,325]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
, 1990
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. The apodeme is reduced and flattened in
|
||||
<taxonomicName id="4C064D51A603FF88AF336D16FD768B6B" authorityName="Binda" authorityYear="1971" box="[621,724,332,356]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="higginsi">
|
||||
<emphasis id="B972EAC0A603FF88AF336D16FD768B6B" box="[621,724,332,356]" italics="true" pageId="58" pageNumber="59">D. higginsi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133D2A57A603FF88AFBA6D17FF198B8C" captionStart="Figure 2" captionStartId="5.[113,178,1715,1739]" captionTargetBox="[115,1454,150,1684]" captionTargetId="graphics-78@5.[115,1181,156,1684]" captionTargetPageId="5" captionText="Figure 2. Phylogenetic relationships within Hypsibiidae in the Bayesian and maximum likelihood analyses (identical topologies; the scale refers to the Bayesian tree and represents substitutions per position; posterior probability values are provided above the nodes, whereas bootstrap values can be found below the nodes; *, maximal support; #, no support).Calohypsibius ornatus constitutes an outgroup. Newly added species and populations are in bold." figureDoi="http://doi.org/10.5281/zenodo.14243746" httpUri="https://zenodo.org/record/14243746/files/figure.png" pageId="58" pageNumber="59">Fig. 2A, B</figureCitation>
|
||||
), which has contributed to confounding views regarding the affinity of the species (
|
||||
<bibRefCitation id="EF974B23A603FF88ACCA6DD0FD638BAC" author="Ramazzotti G & Maucci W" box="[404,705,395,419]" pageId="58" pageNumber="59" pagination="1 - 1012" refId="ref39289" refString="Ramazzotti G, Maucci W. Il Phylum Tardigrada. III edizione riveduta e aggiornata. Memorie dell'Istituto Italiano di Idrobiologia 1983; 41: 1 - 1012." type="journal article" year="1983">Ramazzotti and Maucci 1983</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A603FF88AF906DD0FE8A8BCD" author="Pilato G & Binda MG" pageId="58" pageNumber="59" pagination="45 - 55" refId="ref38678" refString="Pilato G, Binda MG. Precisazioni e rettifiche alla descrizione di alcune specie di tardigradi (terza nota). Animalia 1988; 15: 45 - 55." type="journal article" year="1988">Pilato and Binda 1988</bibRefCitation>
|
||||
). Furthermore, the apodeme in the
|
||||
<taxonomicName id="4C064D51A603FF88AFF46DF0FCAB8BCD" baseAuthorityName="Marcus" baseAuthorityYear="1936" box="[682,777,427,450]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="pingue">
|
||||
<emphasis id="B972EAC0A603FF88AFF46DF0FCAB8BCD" box="[682,777,427,450]" italics="true" pageId="58" pageNumber="59">D. pingue</emphasis>
|
||||
</taxonomicName>
|
||||
group is not a solid drop-shaped thickening, as it would appear in LCM. Instead, when observed in SEM, it is a pair of looped filaments with mostly empty lateral portions (possibly joined by one or more irregular cuticular septa in the middle;
|
||||
<figureCitation id="133D2A57A603FF88AFDB6E7CFD4E8830" box="[645,748,551,575]" captionStart="Figure 3" captionStartId="6.[129,194,1333,1357]" captionTargetBox="[131,1470,146,1304]" captionTargetId="figure-244@6.[129,1473,144,1306]" captionTargetPageId="6" captionText="Figure 3. Various Diphasconinae, morphology of the DABT (SEM): A, Diphascon higginsi Binda, 1971 (dorsal view); B, Diphascon higginsi (lateral view); C, Diphascon pingue (Marcus, 1936) (lateral view); D, Diphascon pinguiforme Pilato & Binda, 1997/8 (dorsolateral view). Scale bars: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.14243748" httpUri="https://zenodo.org/record/14243748/files/figure.png" pageId="58" pageNumber="59">Fig. 3C, D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A603FF88ADC26E1CFED38956" blockId="58.[127,780,144,1985]" pageId="58" pageNumber="59">
|
||||
As noted above,
|
||||
<bibRefCitation id="EF974B23A603FF88AC0D6E1CFDA88850" author="Tumanov DV" box="[339,522,583,607]" pageId="58" pageNumber="59" pagination="1 - 37" refId="ref40150" refString="Tumanov DV. Integrative redescription of Hypsibius pallidoides Pilato et al., 2011 (Eutardigrada: Hypsibioidea) with the erection of a new genus and discussion on the phylogeny of Hypsibiidae. European Journal of Taxonomy 2020; 681: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.681" type="journal article" year="2020">Tumanov (2020)</bibRefCitation>
|
||||
described a remarkable case of paraphyly: the genus
|
||||
<taxonomicName id="4C064D51A603FF88AC9F6E3DFDEE8871" authorityName="Tumanov" authorityYear="2020" box="[449,588,614,638]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AC9F6E3DFDEE8871" box="[449,588,614,638]" italics="true" pageId="58" pageNumber="59">Notahypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
, characterized by a hypothetically plesiomorphic rigid buccal tube (
|
||||
<bibRefCitation id="EF974B23A603FF88AFF06EDEFECC88B2" author="Gasiorek P & Michalczyk L" pageId="58" pageNumber="59" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk 2020</bibRefCitation>
|
||||
), has been placed within
|
||||
<taxonomicName id="4C064D51A603FF88AFFC6EFEFCA188B2" box="[674,771,677,701]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AFFC6EFEFCA188B2" box="[674,771,677,701]" italics="true" pageId="58" pageNumber="59">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
, which exhibits an advanced, elongated posterior portion to its buccopharyngeal apparatus referred to as the pharyngeal tube (
|
||||
<bibRefCitation id="EF974B23A603FF88ADD56F58FEFE8914" author="Bertolani R & Guidetti R & Marchioro T" box="[139,348,771,795]" pageId="58" pageNumber="59" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A603FF88ADAF6F58FE808914" box="[241,290,771,795]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
). This topology is maintained in the new phylogeny presented herein (
|
||||
<figureCitation id="133D2A57A603FF88ACF46F79FE408935" box="[426,482,802,826]" captionStart="Figure 2" captionStartId="5.[113,178,1715,1739]" captionTargetBox="[115,1454,150,1684]" captionTargetId="graphics-78@5.[115,1181,156,1684]" captionTargetPageId="5" captionText="Figure 2. Phylogenetic relationships within Hypsibiidae in the Bayesian and maximum likelihood analyses (identical topologies; the scale refers to the Bayesian tree and represents substitutions per position; posterior probability values are provided above the nodes, whereas bootstrap values can be found below the nodes; *, maximal support; #, no support).Calohypsibius ornatus constitutes an outgroup. Newly added species and populations are in bold." figureDoi="http://doi.org/10.5281/zenodo.14243746" httpUri="https://zenodo.org/record/14243746/files/figure.png" pageId="58" pageNumber="59">Fig. 2</figureCitation>
|
||||
), which also includes several recently described taxa.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A603FF88ADC26F3AFE2A8C66" blockId="58.[127,780,144,1985]" pageId="58" pageNumber="59">
|
||||
The issue of the paraphyletic taxa in classification, which affects many tardigrade lineages, is a topic long discussed in both botany and zoology (
|
||||
<bibRefCitation id="EF974B23A603FF88AC456FC4FE1489B8" author="Schwenk K" box="[283,438,927,951]" pageId="58" pageNumber="59" pagination="69 - 82" refId="ref39574" refString="Schwenk K. Comparative biology and the importance of cladistic classification: a case study from the sensory biology of squamate reptiles. Biological Journal of the Linnean Society 1994; 52: 69 - 82. https: // doi. org / 10.1111 / j. 1095 - 8312.1994. tb 00979. x" type="journal article" year="1994">Schwenk 1994</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A603FF88AC946FC4FDCC89B8" author="Brummitt RK" box="[458,622,927,951]" pageId="58" pageNumber="59" pagination="371 - 84" refId="ref35232" refString="Brummitt RK. In defence of paraphyletic taxa. In: van der Maesen LJG, van der Burgt XM, van Medenbach de Rooy JM (eds.), The Biodiversity of African Plants, Proceedings of the 14 th Association for the Taxonomic Study of the Flora of Tropical Africa Congress, 22 - 27 August 1994, Wageningen, The Netherlands. Springer, Dordrecht, 1996, 371 - 84. https: // doi. org / 10.1007 / 978 - 94 - 009 - 0285 - 5 _ 49" type="book chapter" year="1996">Brummitt 1996</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A603FF88AFDF6FC4FEA789D8" author="Horandl E & Stuessy TF" pageId="58" pageNumber="59" pagination="1641 - 53" refId="ref36651" refString="Horandl E, Stuessy TF. Paraphyletic groups as natural units of biological classification. Taxon 2010; 59: 1641 - 53. https: // doi. org / 10.1002 / tax. 596001" type="journal article" year="2010">Hörandl and Stuessy 2010</bibRefCitation>
|
||||
). Putting aside the fact that a natural consequence of anacladogenesis is the presence of paraphyletic taxa, we are of the opinion that natural classification should aim at distinguishing onlymonophyletictaxa.Besidesthe
|
||||
<emphasis id="B972EAC0A603FF88ACBF6846FD758E3A" box="[481,727,1053,1077]" italics="true" pageId="58" pageNumber="59">
|
||||
<taxonomicName id="4C064D51A603FF88ACBF6846FDCB8E3A" authorityName="Tumanov" authorityYear="2020" box="[481,617,1053,1077]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">Notahypsibius</taxonomicName>
|
||||
–
|
||||
<taxonomicName id="4C064D51A603FF88AF266846FD758E3A" box="[632,727,1053,1077]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">Pilatobius</taxonomicName>
|
||||
</emphasis>
|
||||
case, which we resolve, in part, by establishing
|
||||
<taxonomicName id="4C064D51A603FF88AF4B6866FDCE8E5B" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[533,620,1085,1108]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AF4B6866FDCE8E5B" box="[533,620,1085,1108]" italics="true" pageId="58" pageNumber="59">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A603FF88AFC56866FCA68E5B" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[667,772,1085,1108]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AFC56866FCA68E5B" box="[667,772,1085,1108]" italics="true" pageId="58" pageNumber="59">Fontourion</emphasis>
|
||||
</taxonomicName>
|
||||
, a similar puzzling state is found in
|
||||
<taxonomicName id="4C064D51A603FF88AC996800FDE08E7C" authorityName="Pilato" authorityYear="1969" box="[455,578,1115,1139]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="subFamily" subFamily="Hypsibiinae">Hypsibiinae</taxonomicName>
|
||||
, in which
|
||||
<taxonomicName id="4C064D51A603FF88AFC06800FE948E9C" authority="(Pilato et al. 2006)" baseAuthorityName="Pilato" baseAuthorityYear="2006" class="Eutardigrada" family="Hypsibiidae" genus="Borealibius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AFC06800FCAA8E7C" box="[670,776,1115,1139]" italics="true" pageId="58" pageNumber="59">Borealibius</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A603FF88ADD56820FE898E9C" author="Pilato G & Guidetti R & Rebecchi L" box="[139,299,1147,1171]" pageId="58" pageNumber="59" pagination="595 - 603" refId="ref38984" refString="Pilato G, Guidetti R, Rebecchi L et al. Geonemy, ecology, reproductive biology and morphology of the tardigrade Hypsibius zetlandicus (Eutardigrada: Hypsibiidae) with erection of Borealibius gen. n. Polar Biology 2006; 29: 595 - 603. https: // doi. org / 10.1007 / s 00300 - 005 - 0094 - 1" type="journal article" year="2006">
|
||||
Pilato
|
||||
<emphasis id="B972EAC0A603FF88AD976820FF578E9C" box="[201,245,1147,1171]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
2006
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A603FF88AC3C6820FDCC8E9C" authority="(Dastych 2019)" baseAuthorityName="Dastych" baseAuthorityYear="2019" box="[354,622,1147,1171]" class="Eutardigrada" family="Hypsibiidae" genus="Cryobiotus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AC3C6820FE688E9C" box="[354,458,1147,1171]" italics="true" pageId="58" pageNumber="59">Cryobiotus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A603FF88AC846820FDC18E9C" author="Dastych H" box="[474,611,1147,1171]" pageId="58" pageNumber="59" pagination="95 - 111" refId="ref35714" refString="Dastych H. Cryobiotus roswithae gen. n., sp. n., a new genus and species of glacier-dwelling tardigrades from northern Norway (Tardigrada, Panarthropoda). Entomologie heute 2019; 31: 95 - 111." type="journal article" year="2019">Dastych 2019</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, i.e. genera with clear autapomorphic conditions, are probably (although genetic data are scarce) embedded within the ‘plesiomorphic’
|
||||
<taxonomicName id="4C064D51A603FF88AFF568E2FCAB8EDE" authorityName="Ehrenberg" authorityYear="1848" box="[683,777,1209,1233]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88AFF568E2FCAB8EDE" box="[683,777,1209,1233]" italics="true" pageId="58" pageNumber="59">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
(see also
|
||||
<bibRefCitation id="EF974B23A603FF88ADBC6882FDBA8EFE" author="Tumanov DV & Tsvetkova AY" box="[226,536,1241,1265]" pageId="58" pageNumber="59" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="4C064D51A603FF88AF6F6882FEA88F1F" authority="Cuenot, 1929" authorityName="Cuenot" authorityYear="1929" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="scabropygus">
|
||||
<emphasis id="B972EAC0A603FF88AF6F6882FCAB8EFE" box="[561,777,1241,1265]" italics="true" pageId="58" pageNumber="59">Hypsibius scabropygus</emphasis>
|
||||
<bibRefCitation id="EF974B23A603FF88ADDF68A3FEA88F1F" author="Cuenot L" box="[129,266,1272,1296]" pageId="58" pageNumber="59" pagination="121 - 5" refId="ref35447" refString="Cuenot L. Description d'un tardigrade nouveau de la faune francaise. Archives d'anatomie microscopique 1929; 25: 121 - 5." type="journal article" year="1929">Cuénot, 1929</bibRefCitation>
|
||||
</taxonomicName>
|
||||
is typically recovered as sister to all other hypsibiins and should be erected together with a few other
|
||||
<taxonomicName id="4C064D51A603FF88AF01694CFCAB8F20" box="[607,777,1303,1327]" pageId="58" pageNumber="59">
|
||||
<emphasis id="B972EAC0A603FF88AF01694CFD1F8F20" box="[607,701,1303,1327]" italics="true" pageId="58" pageNumber="59">Hypsibius</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
as a separate genus (
|
||||
<bibRefCitation id="EF974B23A603FF88AC15696CFDB28F40" author="Gasiorek P & Stec D & Morek W" box="[331,528,1335,1359]" pageId="58" pageNumber="59" pagination="45 - 75" refId="ref36224" refString="Gasiorek P, Stec D, Morek W et al. An integrative redescription of Hypsibius dujardini (Doyere, 1840), the nominal taxon for Hypsibioidea (Tardigrada: Eutardigrada). Zootaxa 2018; 4415: 45 - 75. https: // doi. org / 10.11646 / zootaxa. 4415.1.2" type="journal article" year="2018">
|
||||
Gąsiorek
|
||||
<emphasis id="B972EAC0A603FF88ACF4696CFE7B8F40" box="[426,473,1335,1359]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
). An analogous problem was recently shown in
|
||||
<taxonomicName id="4C064D51A603FF88AC03690DFE508F61" authorityName="Thulin" authorityYear="1928" box="[349,498,1366,1390]" class="Eutardigrada" family="Macrobiotidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="family">Macrobiotidae</taxonomicName>
|
||||
, in which the strongly modified genus
|
||||
<taxonomicName id="4C064D51A603FF88ADB7692EFDCB8F82" authority="(Bertolani & Biserov, 1996)" baseAuthorityName="Bertolani & Biserov" baseAuthorityYear="1996" box="[233,617,1397,1421]" class="Eutardigrada" family="Macrobiotidae" genus="Xerobiotus" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88ADB7692EFEED8F82" box="[233,335,1397,1421]" italics="true" pageId="58" pageNumber="59">Xerobiotus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A603FF88AC01692EFDFC8F82" author="Bertolani R & Biserov VI" box="[351,606,1397,1421]" pageId="58" pageNumber="59" pagination="299 - 304" refId="ref34889" refString="Bertolani R, Biserov VI. Leg and claw adaptations in soil tardigrades, with erection of two new genera of Eutardigrada, Macrobiotidae: Pseudohexapodibius and Xerobiotus. Invertebrate Biology 1996; 115: 299 - 304. https: // doi. org / 10.2307 / 3227019" type="journal article" year="1996">Bertolani & Biserov, 1996</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
is nested within
|
||||
<taxonomicName id="4C064D51A603FF88ADDE69CEFF558FA2" box="[128,247,1429,1453]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88ADDE69CEFF558FA2" box="[128,247,1429,1453]" italics="true" pageId="58" pageNumber="59">Macrobiotus</emphasis>
|
||||
</taxonomicName>
|
||||
and thus should be considered a species group within
|
||||
<taxonomicName id="4C064D51A603FF88ADDE69EFFE098FC3" authority="(Stec et al. 2021)" baseAuthorityName="Stec" baseAuthorityYear="2021" box="[128,427,1460,1484]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A603FF88ADDE69EFFF558FC3" box="[128,247,1460,1484]" italics="true" pageId="58" pageNumber="59">Macrobiotus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A603FF88AC5669EFFE028FC3" author="Stec D & Vecchi M & Calhim S" box="[264,416,1460,1484]" pageId="58" pageNumber="59" refId="ref39741" refString="Stec D, Vecchi M, Calhim S et al. New multilocus phylogeny reorganises the family Macrobiotidae (Eutardigrada) and unveils complex morphological evolution of the Macrobiotus hufelandi group. Molecular Phylogenetics and Evolution 2021; 160: 106987. https: // doi. org / 10.1016 / j. ympev. 2020.106987" type="journal volume" year="2021">
|
||||
Stec
|
||||
<emphasis id="B972EAC0A603FF88AC6769EEFECA8FC3" box="[313,360,1460,1484]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. We do not exclude the possibility that some of these taxa are only superficially artificial, because DNA sequences are lacking for the vast majority of species, and that the phyletic relationships might eventually turn out to be different from what we currently know, but this issue seems topical and requires further study.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A603FF88ADC26A2BFB798BCD" blockId="58.[127,780,144,1985]" lastBlockId="58.[824,1476,144,1516]" pageId="58" pageNumber="59">
|
||||
In fact, the only hypsibiid subfamily that seems well defined morphologically from the remaining clades are the
|
||||
<taxonomicName id="4C064D51A603FF88AFD46AD4FCAA8CA8" authorityName="Pilato" authorityYear="1969" box="[650,776,1679,1703]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="subFamily" subFamily="Hypsibiinae">Hypsibiinae</taxonomicName>
|
||||
, whose morphology of the buccopharyngeal apparatus includes plesiomorphic eutardigrade characters [a likely ancestral
|
||||
<typeStatus id="54BD8870A603FF88AF866A95FCA58CE9" box="[728,775,1742,1766]" pageId="58" pageNumber="59">type</typeStatus>
|
||||
, comprising only a rigid buccal tube and granular or bar-like macroplacoids without signs of thinning and forming single, thin and long macroplacoids typical of many itaquasconins and some diphasconins; however, see the alternative hypothesis postulated by
|
||||
<bibRefCitation id="EF974B23A603FF88AC626B30FDBD8D8C" author="Bertolani R & Guidetti R & Marchioro T" box="[316,543,1899,1923]" pageId="58" pageNumber="59" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A603FF88ACFE6B30FE728D8C" box="[416,464,1899,1923]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
(2014)
|
||||
</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="EF974B23A603FF88AF0C6B30FCA68D8C" author="Tumanov DV" box="[594,772,1899,1923]" pageId="58" pageNumber="59" pagination="1 - 37" refId="ref40150" refString="Tumanov DV. Integrative redescription of Hypsibius pallidoides Pilato et al., 2011 (Eutardigrada: Hypsibioidea) with the erection of a new genus and discussion on the phylogeny of Hypsibiidae. European Journal of Taxonomy 2020; 681: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.681" type="journal article" year="2020">Tumanov (2020)</bibRefCitation>
|
||||
, according to which the presence of only the buccal tube within the buccopharyngeal apparatus is considered apomorphic]; and claws that are relatively homogeneous within
|
||||
<taxonomicName id="4C064D51A603FF88A84A6CCBFA328AA7" authorityName="Pilato" authorityYear="1969" box="[1300,1424,144,168]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="family">Hypsibiidae</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EF974B23A603FF88AE676CF4FC748AC8" author="Guil N & Machordom A & Guidetti R" box="[825,982,175,199]" pageId="58" pageNumber="59" pagination="1 - 26" refId="ref36502" refString="Guil N, Machordom A, Guidetti R. High level of phenotypic homoplasy amongst eutardigrades (Tardigrada) based on morphological and total evidence phylogenetic analyses. Zoological Journal of the Linnean Society 2013; 169: 1 - 26. https: // doi. org / 10.1111 / zoj. 12046" type="journal article" year="2013">
|
||||
Guil
|
||||
<emphasis id="B972EAC0A603FF88AE336CEBFC3E8AC8" box="[877,924,175,199]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
2013
|
||||
</bibRefCitation>
|
||||
). This means, and is the main point of our criticism, that narrow definitions of newly erected supraspecific taxa, which focus on autapomorphies (always termed subjectively as ‘strong’ or ‘weak’) and neglect the topology of the evolutionary tree, will inevitably lead to creation of paraphyletic taxa characterized by advanced characters and nested among groups of a plesiomorphic nature (
|
||||
<figureCitation id="133D2A57A603FF88A9766D30FBCD8B8C" box="[1064,1135,363,387]" captionStart="Figure 44" captionStartId="56.[130,195,572,596]" captionTargetBox="[130,776,144,544]" captionTargetId="graphics-841@56.[162,743,225,546]" captionTargetPageId="56" captionText="Figure 44.Evolution scheme for a tardigrade clade, where the blue triangle and red rectrangle signify newly erected genera (‘apical taxa’ in the paper by Tumanov and Tsvetkova 2023) with ‘strong apomorphies’, i.e. the set of well-defined, advanced morphological characters, distinguished from the previous broadly defined genus, whose members (marked with black circles) are still classified as the original genus, which became artificial but cannot be divided further because the lineages group species with mainly (or solely) plesiomorphic characters." figureDoi="http://doi.org/10.5281/zenodo.14243843" httpUri="https://zenodo.org/record/14243843/files/figure.png" pageId="58" pageNumber="59">Fig. 44</figureCitation>
|
||||
). Therefore, we advocate for the phylogenetic approach and the rule of reciprocal monophyly (
|
||||
<bibRefCitation id="EF974B23A603FF88AE1A6DF1FC688BCD" author="Hennig W" box="[836,970,426,450]" pageId="58" pageNumber="59" refId="ref36590" refString="Hennig W. Phylogenetic Systematics. Urbana: University of Illinois Press, 1966." type="book" year="1966">Hennig 1966</bibRefCitation>
|
||||
) in tardigrade systematics.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A603FF88AE0B6D92FA9789D8" blockId="58.[824,1476,144,1516]" pageId="58" pageNumber="59">
|
||||
The paraphyletic status of some tardigrade taxa is not the only problem affecting hypsibiid classification. The addition of
|
||||
<taxonomicName id="4C064D51A603FF88AEC86E53FC49882F" baseAuthorityName="Thulin" baseAuthorityYear="1928" box="[918,1003,520,544]" class="Eutardigrada" family="Hypsibiidae" genus="Arctodiphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="tenue">
|
||||
<emphasis id="B972EAC0A603FF88AEC86E53FC49882F" box="[918,1003,520,544]" italics="true" pageId="58" pageNumber="59">A. tenue</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<emphasis id="B972EAC0A603FF88A9776E53FB62882F" box="[1065,1216,520,544]" italics="true" pageId="58" pageNumber="59">Arctodiphascon</emphasis>
|
||||
<taxonomicName id="4C064D51A603FF88A9926E52FCDA8830" authority="(Sun et al., 2020)" baseAuthorityName="Sun" baseAuthorityYear="2020" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="wuyingensis">
|
||||
<emphasis id="B972EAC0A603FF88A9926E52FA9C882F" box="[1228,1342,521,544]" italics="true" pageId="58" pageNumber="59">wuyingensis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A603FF88A80A6E53FCCF8830" author="Sun X-L & Zhang J-Y & Wang N" pageId="58" pageNumber="59" pagination="185 - 94" refId="ref39841" refString="Sun X-L, Zhang J-Y, Wang N et al. A new species of Diphascon (Tardigrada: Hypsibiidae) from Northern China supported by integrated taxonomy. Zootaxa 2020; 4722: 185 - 94. https: // doi. org / 10.11646 / zootaxa. 4722.2.5" type="journal article" year="2020">
|
||||
Sun
|
||||
<emphasis id="B972EAC0A603FF88A8D66E52FA1F882F" box="[1416,1469,520,544]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||||
, 2020
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
blurred the distinction between
|
||||
<taxonomicName id="4C064D51A603FF88A9AA6E7CFA2F8830" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[1268,1421,551,575]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A603FF88AE676E1CFC718850" authorityName="Rudescu" authorityYear="1964" box="[825,979,583,607]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
further (
|
||||
<bibRefCitation id="EF974B23A603FF88A9646E1CFADE8850" author="Tumanov DV & Tsvetkova AY" box="[1082,1404,583,607]" pageId="58" pageNumber="59" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
). This means that some
|
||||
<taxonomicName id="4C064D51A603FF88AEAB6E3DFB2D8872" box="[1013,1167,614,638]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="undetermined">
|
||||
<emphasis id="B972EAC0A603FF88AEAB6E3DFBFF8871" box="[1013,1117,614,638]" italics="true" pageId="58" pageNumber="59">Diphascon</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
with a DABT might, in fact, belong to the
|
||||
<taxonomicName id="4C064D51A603FF88AEBC6EDEFBDC8892" authorityName="Rudescu" authorityYear="1964" box="[994,1150,645,669]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="subFamily" subFamily="Itaquasconinae">Itaquasconinae</taxonomicName>
|
||||
, ordinarily apodemeless (the apodeme is small and may be reduced in
|
||||
<taxonomicName id="4C064D51A603FF88A9AF6EFEFA2088B3" box="[1265,1410,677,700]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="wuyingensis">
|
||||
<emphasis id="B972EAC0A603FF88A9AF6EFEFA2088B3" box="[1265,1410,677,700]" italics="true" pageId="58" pageNumber="59">A. wuyingensis</emphasis>
|
||||
</taxonomicName>
|
||||
). The intricacy of this situation stems from the lack of a morphological apomorphy that would allow
|
||||
<taxonomicName id="4C064D51A603FF88A93E6EBFFB4F88F4" box="[1120,1261,740,763]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="58" pageNumber="59" phylum="Tardigrada" rank="species" species="wuyingensis">
|
||||
<emphasis id="B972EAC0A603FF88A93E6EBFFB4F88F4" box="[1120,1261,740,763]" italics="true" pageId="58" pageNumber="59">A. wuyingensis</emphasis>
|
||||
</taxonomicName>
|
||||
to be placed in a new genus (
|
||||
<emphasis id="B972EAC0A603FF88AED96F58FBBC8914" box="[903,1054,771,795]" italics="true" pageId="58" pageNumber="59">Arctodiphascon</emphasis>
|
||||
was erected with clear evidence of non-monophyly;
|
||||
<bibRefCitation id="EF974B23A603FF88AEE26F79FB508935" author="Tumanov DV & Tsvetkova AY" box="[956,1266,802,826]" pageId="58" pageNumber="59" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
). However, this discrepancy might be resolved when the species is studied in more detail, because
|
||||
<bibRefCitation id="EF974B23A603FF88AE876F3AFB2F8976" author="Tumanov DV" box="[985,1165,865,889]" pageId="58" pageNumber="59" pagination="1 - 37" refId="ref40150" refString="Tumanov DV. Integrative redescription of Hypsibius pallidoides Pilato et al., 2011 (Eutardigrada: Hypsibioidea) with the erection of a new genus and discussion on the phylogeny of Hypsibiidae. European Journal of Taxonomy 2020; 681: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.681" type="journal article" year="2020">Tumanov (2020)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="EF974B23A603FF88A99D6F3AFC268997" author="Tumanov DV & Tsvetkova AY" pageId="58" pageNumber="59" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
pointed out that traits previously neglected in hypsibiid taxonomy, such as ornamentation of eggs deposited in exuviae, might prove useful in monophyletic classification.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A603FF88AE0B6F85FAE18FE4" pageId="58" pageNumber="59" type="materials_examined">
|
||||
<paragraph id="8BB936D2A603FF88AE0B6F85FAE18FE4" blockId="58.[824,1476,144,1516]" pageId="58" pageNumber="59">
|
||||
To sum up, the phylogenetic and morphological diversity of hypsibiids must be researched to a greater extent in order to define accurately the four main clades that are currently recognized as subfamilies. Based on our analyses, we postulate that the morphology of the
|
||||
<collectionCode id="ED17AE17A603FF88AEE46807FC5F8E7B" box="[954,1021,1116,1140]" pageId="58" pageNumber="59">DABT</collectionCode>
|
||||
might bear an important evolutionary value. The morphological characterization of pedal bars, summarized in this paper, aims at unifying the anatomical terminology to aid taxonomic descriptions and comparisons. Integrative revisions of
|
||||
<typeStatus id="54BD8870A603FF88AE0B6882FC208EFE" box="[853,898,1241,1265]" pageId="58" pageNumber="59">type</typeStatus>
|
||||
species were, once again, demonstrated as pivotal to the decluttering of tardigrade classification. It appears that, in many instances, the evolutionary significance of particular phenotypic characters in tardigrade systematics is misunderstood. As a result, subjective assessments of morphology as the only basis for erection of new higher taxa, i.e. finding (‘strong’) apomorphies, lead to the erection of taxa defined by characters (phenetic systematics), and not by factual evolutionary affinities (i.e. reciprocal monophyly; e.g. see
|
||||
<bibRefCitation id="EF974B23A603FF88A95C6988FA908FE4" author="de Queiroz K & Gauthier J" box="[1026,1330,1491,1515]" pageId="58" pageNumber="59" pagination="307 - 22" refId="ref35864" refString="de Queiroz K, Gauthier J. Phylogeny as a central principle in taxonomy: phylogenetic definitions of taxon names. Systematic Zoology 1990; 39: 307 - 22. https: // doi. org / 10.2307 / 2992353" type="journal article" year="1990">de Queiroz and Gauthier 1990</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
218
data/03/AF/87/03AF87C4A601FF8AAE2D6D5DFC478FFA.xml
Normal file
218
data/03/AF/87/03AF87C4A601FF8AAE2D6D5DFC478FFA.xml
Normal file
|
|
@ -0,0 +1,218 @@
|
|||
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<mods:title id="9F050FB1CB8F6D4EE58993AB5C94ED4A">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="32C6D6B9F94CAF3C73D966504748F34C">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="295E782465B81AF614444F1D817A4F7E">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="96BF15BAA4A751F8A4BA04151E4E70DC" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5012BAD82E690E20713CEAF554EEB1E4">Blagden, Brian</mods:namePart>
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<mods:affiliation id="F8134B593FD90982EA77A9DAEF02FE14">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:affiliation id="19F6658C89829A3DE215A4BA18A749C2">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:affiliation id="F96EBCA7CB5CE3BB8651089F6A47B25B">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:date id="9E42708547D517F571D68CBBC5BF3C84">2024</mods:date>
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<taxonomicName id="4C064D51A601FF8AAE2D6D5DFA8A8B2F" authority="Gasiorek, Morek & Michalczyk" authorityName="Gasiorek, Morek & Michalczyk" authorityYear="2024" box="[883,1320,262,288]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
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<emphasis id="B972EAC0A601FF8AAE2D6D5DFC7B8B2F" bold="true" box="[883,985,262,288]" italics="true" pageId="56" pageNumber="57">Degmion</emphasis>
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Gąsiorek, Morek & Michalczyk
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||||
</taxonomicName>
|
||||
gen. nov.
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||||
</emphasis>
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</heading>
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|
||||
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ZooBank registration:
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urn:lsid:zoobank.org:act:
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||||
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|
||||
<subSubSection id="C31C6559A601FF8AAE676DDDFBD6895B" pageId="56" pageNumber="57" type="diagnosis">
|
||||
<paragraph id="8BB936D2A601FF8AAE676DDDFBD6895B" blockId="56.[825,1476,389,852]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AAE676DDDFC3D8B92" box="[825,927,390,413]" italics="true" pageId="56" pageNumber="57">Diagnosis:</emphasis>
|
||||
Cuticle distinctly sculptured in the caudal portion of the trunk; more anterior portions with a weaker sculpturing or smooth. Apophyses for the insertion of the stylet muscles (AISMs) in the shape of semilunar hooks. Buccal tube followed by a flexible annulated pharyngeal tube; the pharyngeal annulation is semi-complex (annuli single laterally and forking dorsoventrally). A DABT is present. There is a spherical pharynx with two macroplacoids and a septulum. Macroplacoids are arranged rhomboidally (◊). Claws of the
|
||||
<taxonomicName id="4C064D51A601FF8AA9246EDBFB788897" authorityName="Ehrenberg" authorityYear="1848" box="[1146,1242,640,664]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AA9246EDBFB788897" box="[1146,1242,640,664]" italics="true" pageId="56" pageNumber="57">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A601FF8AA9BE6EDBFAAD8897" box="[1248,1295,640,664]" pageId="56" pageNumber="57">type</typeStatus>
|
||||
, i.e. asymmetrical both with respect to the sequence of primary and secondary branches (2-1-2-1) and with respect to their size, with external and posterior claws always evidently larger than the internal and anterior claws. Pseudolunulae below claws I–IV may be present; only basal cuticular bars below claws IV may be present, with the remaining
|
||||
<typeStatus id="54BD8870A601FF8AAEFB6F67FC7E895B" box="[933,988,828,852]" pageId="56" pageNumber="57">types</typeStatus>
|
||||
of bars absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A601FF8AAE676F2DFAF089E4" pageId="56" pageNumber="57" type="discussion">
|
||||
<paragraph id="8BB936D2A601FF8AAE676F2DFAF089E4" blockId="56.[825,1474,885,1003]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AAE676F2DFBFE8982" box="[825,1116,885,909]" italics="true" pageId="56" pageNumber="57">
|
||||
Composition:
|
||||
<taxonomicName id="4C064D51A601FF8AAE966F2DFBFE8982" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[968,1116,885,909]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="burti" status="comb. nov.">Degmion burti</taxonomicName>
|
||||
</emphasis>
|
||||
<emphasis id="B972EAC0A601FF8AA9396F2EFB418982" bold="true" box="[1127,1251,885,909]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA9396F2EFB418982" box="[1127,1251,885,909]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AA9AA6F2DFA608982" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1268,1474,885,909]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="nodulosum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA9AA6F2DFA608982" box="[1268,1474,885,909]" italics="true" pageId="56" pageNumber="57">Degmion nodulosum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AAE676FCEFC0D89A2" bold="true" box="[825,943,917,941]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAE676FCEFC0D89A2" box="[825,943,917,941]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AAEE66FCEFB2589A3" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[952,1159,917,940]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="nuominense" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AAEE66FCEFB2589A3" box="[952,1159,917,940]" italics="true" pageId="56" pageNumber="57">Degmion nuominense</emphasis>
|
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</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AA9D26FCEFAA389A2" bold="true" box="[1164,1281,917,941]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA9D26FCEFAA389A2" box="[1164,1281,917,941]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AA8556FCEFA6089A2" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1291,1474,917,941]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="oculatum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA8556FCEFA6089A2" box="[1291,1474,917,941]" italics="true" pageId="56" pageNumber="57">Degmion oculatum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AAE676FEFFC0D89C3" bold="true" box="[825,943,948,972]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAE676FEFFC0D89C3" box="[825,943,948,972]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<typeStatus id="54BD8870A601FF8AAEE26FEFFC4F89C3" box="[956,1005,948,972]" pageId="56" pageNumber="57">type</typeStatus>
|
||||
species),
|
||||
<taxonomicName id="4C064D51A601FF8AA9116FEEFAE389C3" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1103,1345,948,972]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="opisthoglyptum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA9116FEEFAE389C3" box="[1103,1345,948,972]" italics="true" pageId="56" pageNumber="57">Degmion opisthoglyptum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AA8196FEFFA1F89C3" bold="true" box="[1351,1469,948,972]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA8196FEFFA1F89C3" box="[1351,1469,948,972]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A601FF8AAE3B6F8FFBED89E4" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[869,1103,979,1003]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugocaudatum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AAE3B6F8FFBED89E4" box="[869,1103,979,1003]" italics="true" pageId="56" pageNumber="57">Degmion rugocaudatum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AA90B6F88FAF089E4" bold="true" box="[1109,1362,979,1003]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA90B6F88FB6989E4" box="[1109,1227,979,1003]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
et nom. inq.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A601FF8AAE676856FAA28E4B" pageId="56" pageNumber="57" type="etymology">
|
||||
<paragraph id="8BB936D2A601FF8AAE676856FAA28E4B" blockId="56.[825,1474,1037,1092]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AAE676856FC0A8E2A" box="[825,936,1037,1061]" italics="true" pageId="56" pageNumber="57">Etymology:</emphasis>
|
||||
The name honours Dr. Peter Degma, a renowned Slovakian tardigradologist and a great friend.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A601FF8AAE67683EFC478FFA" pageId="56" pageNumber="57" type="diagnosis">
|
||||
<paragraph id="8BB936D2A601FF8AAE67683EFB3C8EB3" blockId="56.[825,1474,1125,1212]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AAE67683EFBB68E72" box="[825,1044,1125,1149]" italics="true" pageId="56" pageNumber="57">Differential diagnosis:</emphasis>
|
||||
Within the
|
||||
<taxonomicName id="4C064D51A601FF8AA9FD683EFABC8E72" box="[1187,1310,1125,1149]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AA871683DFA258E72" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1327,1415,1126,1149]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA871683DFA258E72" box="[1327,1415,1126,1149]" italics="true" pageId="56" pageNumber="57">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AA8CA683EFCC58E92" bold="true" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA8CA683EFCC58E92" pageId="56" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AAE2A68DEFC778E92" box="[884,981,1157,1181]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AAE2A68DEFC778E92" box="[884,981,1157,1181]" italics="true" pageId="56" pageNumber="57">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A601FF8AA94F68DEFBDE8E93" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1041,1148,1157,1180]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA94F68DEFBDE8E93" box="[1041,1148,1157,1180]" italics="true" pageId="56" pageNumber="57">Fontourion</emphasis>
|
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</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AA9DC68DEFB478E92" bold="true" box="[1154,1253,1157,1181]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AA9DC68DEFB478E92" box="[1154,1253,1157,1181]" pageId="56" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
possess a pharyngeal tube (absent in
|
||||
<taxonomicName id="4C064D51A601FF8AAE8668FFFBC18EB3" authorityName="Tumanov" authorityYear="2020" box="[984,1123,1188,1212]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AAE8668FFFBC18EB3" box="[984,1123,1188,1212]" italics="true" pageId="56" pageNumber="57">Notahypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
) but:
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A601FF8AAE0A68B8FB758F98" blockId="56.[852,1475,1251,1525]" pageId="56" pageNumber="57">
|
||||
•
|
||||
<taxonomicName id="4C064D51A601FF8AAEDA68B8FC7E8EF5" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[900,988,1251,1274]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AAEDA68B8FC7E8EF5" box="[900,988,1251,1274]" italics="true" pageId="56" pageNumber="57">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AAEBA68B8FBE58EF4" bold="true" box="[996,1095,1251,1275]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAEBA68B8FBE58EF4" box="[996,1095,1251,1275]" pageId="56" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
has the dorsal sculpturing restricted to the caudal part of the trunk or the caudal sculpturing is much better developed than that found anteriorly and lacks dorsolateral gibbosities, whereas
|
||||
<taxonomicName id="4C064D51A601FF8AA842691AFADF8F56" box="[1308,1405,1345,1369]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AA842691AFADF8F56" box="[1308,1405,1345,1369]" italics="true" pageId="56" pageNumber="57">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
either exhibits gibbosities and/or its dorsal cuticle is uniformly sculptured throughout the trunk;
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A601FF8AAE0A69C4FC478FFA" blockId="56.[852,1475,1251,1525]" pageId="56" pageNumber="57">
|
||||
•
|
||||
<taxonomicName id="4C064D51A601FF8AAEDA69C4FC7E8FB9" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[900,988,1439,1462]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AAEDA69C4FC7E8FB9" box="[900,988,1439,1462]" italics="true" pageId="56" pageNumber="57">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AAEB969C4FBEF8FB8" bold="true" box="[999,1101,1439,1463]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAEB969C4FBEF8FB8" box="[999,1101,1439,1463]" pageId="56" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
always exhibits an obvious caudal sculpturing, whereas the cuticle is smooth in
|
||||
<taxonomicName id="4C064D51A601FF8AA80669E4FA618FD9" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1368,1475,1471,1494]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AA80669E4FA618FD9" box="[1368,1475,1471,1494]" italics="true" pageId="56" pageNumber="57">Fontourion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A601FF8AAEDA6986FC478FFA" bold="true" box="[900,997,1501,1525]" pageId="56" pageNumber="57">
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAEDA6986FC478FFA" box="[900,997,1501,1525]" pageId="56" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
244
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Normal file
244
data/03/AF/87/03AF87C4A601FF8BAE396A6FFD2A8E72.xml
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|
|
@ -0,0 +1,244 @@
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<mods:title id="469D056874A49829DC3FBD147B38C473">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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|
||||
<taxonomicName id="4C064D51A601FF8AAE396A6FFA968C41" authority="Gasiorek, Morek & Michalczyk" authorityName="Gasiorek, Morek & Michalczyk" authorityYear="2024" box="[871,1332,1588,1614]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A601FF8AAE396A6FFC448C41" bold="true" box="[871,998,1588,1614]" italics="true" pageId="56" pageNumber="57">Fontourion</emphasis>
|
||||
Gąsiorek, Morek & Michalczyk
|
||||
</taxonomicName>
|
||||
gen. nov.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A601FF8AAE676A00FB018C9D" pageId="56" pageNumber="57" type="description">
|
||||
<paragraph id="8BB936D2A601FF8AAE676A00FB018C9D" blockId="56.[825,1474,1588,1682]" pageId="56" pageNumber="57">
|
||||
ZooBank registration:
|
||||
<uri id="FF973AD0A601FF8AA9686A01FB018C9D" pageId="56" pageNumber="57">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFA00C07A601FF8AA8686A01FB018C9D" pageId="56" pageNumber="57">ACDC00E5- 6B25-4823-BFE8-79CEA2A9457E</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A601FF8BAE676AEFFD098B71" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="diagnosis">
|
||||
<paragraph id="8BB936D2A601FF8BAE676AEFFD928AE9" blockId="56.[824,1475,1715,1990]" lastBlockId="57.[113,763,144,231]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AAE676AEFFC3D8CC4" box="[825,927,1716,1739]" italics="true" pageId="56" pageNumber="57">Diagnosis:</emphasis>
|
||||
Cuticle smooth. The AISMs are in the shape of semilunar hooks. Buccal tube is followed by a flexible annulated pharyngeal tube; the pharyngeal annulation is semi-complex (annuli single laterally and forking dorsoventrally). A DABT is present. There is a spherical pharynx, with two macroplacoids and a septulum. Macroplacoids are arranged rhomboidally (◊). Claws are of the
|
||||
<taxonomicName id="4C064D51A601FF8AAEF46B34FBA88D88" authorityName="Ehrenberg" authorityYear="1848" box="[938,1034,1903,1927]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AAEF46B34FBA88D88" box="[938,1034,1903,1927]" italics="true" pageId="56" pageNumber="57">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A601FF8AA94D6B2BFBE08D87" box="[1043,1090,1904,1928]" pageId="56" pageNumber="57">type</typeStatus>
|
||||
, i.e. asymmetrical both with respect to the sequence of primary and secondary branches (2-1-2-1) and with respect to their size, with external and posterior claws always evidently larger than the internal and anterior claws. Pseudolunulae below claws I–IV may be present; only internal and posterior cuticular bars may be present.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6D53FD098B71" blockId="57.[113,763,264,382]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6D53FE768B2F" box="[113,468,264,288]" italics="true" pageId="57" pageNumber="58">
|
||||
Composition:
|
||||
<taxonomicName id="4C064D51A600FF8BAC526D52FE768B2F" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[268,468,264,288]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="boreale" status="comb. nov.">Fontourion boreale</taxonomicName>
|
||||
</emphasis>
|
||||
<emphasis id="B972EAC0A600FF8BACB56D53FDD08B2F" bold="true" box="[491,626,264,288]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BACB56D53FDD08B2F" box="[491,626,264,288]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAFD16D52FF1D8B30" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="2024" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="brevipes" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAFD16D52FF1D8B30" italics="true" pageId="57" pageNumber="58">Fontourion brevipes</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAD986D7CFE9C8B30" bold="true" box="[198,318,295,319]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAD986D7CFE9C8B30" box="[198,318,295,319]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAC146D73FDA68B30" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[330,516,295,319]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="glaciale" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAC146D73FDA68B30" box="[330,516,295,319]" italics="true" pageId="57" pageNumber="58">Fontourion glaciale</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAF556D7CFD218B30" bold="true" box="[523,643,295,319]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAF556D7CFD218B30" box="[523,643,295,319]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAFD16D73FF788B50" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="2024" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="islandicum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAFD16D73FF788B50" italics="true" pageId="57" pageNumber="58">Fontourion islandicum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BADBD6D1CFEFE8B50" bold="true" box="[227,348,327,351]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BADBD6D1CFEFE8B50" box="[227,348,327,351]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAC346D1CFD958B51" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1911" box="[362,567,327,350]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="recamieri" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAC346D1CFD958B51" box="[362,567,327,350]" italics="true" pageId="57" pageNumber="58">Fontourion recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAF1E6D1CFD188B50" bold="true" box="[576,698,327,351]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAF1E6D1CFD188B50" box="[576,698,327,351]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<typeStatus id="54BD8870A600FF8BAF976D1CFD588B50" box="[713,762,327,351]" pageId="57" pageNumber="58">type</typeStatus>
|
||||
species), and
|
||||
<taxonomicName id="4C064D51A600FF8BADA46D3CFE0A8B71" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[250,424,358,382]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="secchii" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BADA46D3CFE0A8B71" box="[250,424,358,382]" italics="true" pageId="57" pageNumber="58">Fontourion secchii</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BACF36D3DFD098B71" bold="true" box="[429,683,358,382]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BACF36D3DFD868B71" box="[429,548,358,382]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
et nom. inq.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A600FF8BAD2F6DC4FE318BD8" pageId="57" pageNumber="58" type="etymology">
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6DC4FE318BD8" blockId="57.[113,762,415,471]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6DC4FF438BB8" box="[113,225,415,439]" italics="true" pageId="57" pageNumber="58">Etymology:</emphasis>
|
||||
The name honours Dr. Paulo Fontoura, an esteemed Portuguese tardigradologist.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A600FF8BAD2F6DA3FD7F8904" pageId="57" pageNumber="58" type="diagnosis">
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6DA3FE238840" blockId="57.[113,763,504,591]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6DA3FEE4881F" box="[113,326,504,528]" italics="true" pageId="57" pageNumber="58">Differential diagnosis:</emphasis>
|
||||
Within
|
||||
<taxonomicName id="4C064D51A600FF8BACC26DA3FDB5881F" box="[412,535,504,528]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAF7C6DA2FD2F881F" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[546,653,505,528]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAF7C6DA2FD2F881F" box="[546,653,505,528]" italics="true" pageId="57" pageNumber="58">Fontourion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAFCD6DA3FD57881F" bold="true" box="[659,757,504,528]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAFCD6DA3FD57881F" box="[659,757,504,528]" pageId="57" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C064D51A600FF8BAD2F6E43FF6B8820" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[113,201,536,559]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6E43FF6B8820" box="[113,201,536,559]" italics="true" pageId="57" pageNumber="58">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAD906E4CFE8C8820" bold="true" box="[206,302,535,559]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAD906E4CFE8C8820" box="[206,302,535,559]" pageId="57" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A600FF8BAC3C6E4CFE618820" box="[354,451,535,559]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BAC3C6E4CFE618820" box="[354,451,535,559]" italics="true" pageId="57" pageNumber="58">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
possess a pharyngeal tube (absent in
|
||||
<taxonomicName id="4C064D51A600FF8BADE46E6CFEE78840" authorityName="Tumanov" authorityYear="2020" box="[186,325,567,591]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BADE46E6CFEE78840" box="[186,325,567,591]" italics="true" pageId="57" pageNumber="58">Notahypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
) but:
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BADD26E2DFDAD88E4" blockId="57.[140,762,629,779]" pageId="57" pageNumber="58">
|
||||
•
|
||||
<taxonomicName id="4C064D51A600FF8BADE26E2DFE858882" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[188,295,630,653]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BADE26E2DFE858882" box="[188,295,630,653]" italics="true" pageId="57" pageNumber="58">Fontourion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAC6C6E2EFE3A8882" bold="true" box="[306,408,629,653]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAC6C6E2EFE3A8882" box="[306,408,629,653]" pageId="57" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
has a smooth dorsal cuticle and lacks dorsolateral gibbosities, whereas
|
||||
<taxonomicName id="4C064D51A600FF8BAF0A6ECEFD1788A2" box="[596,693,661,685]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A600FF8BAF0A6ECEFD1788A2" box="[596,693,661,685]" italics="true" pageId="57" pageNumber="58">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
either exhibits gibbosities and/or its dorsal cuticle is uniformly sculptured throughout the trunk;
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A600FF8BADD26EA8FD7F8904" blockId="57.[140,762,629,779]" box="[140,733,755,779]" pageId="57" pageNumber="58">
|
||||
• for the distinction with
|
||||
<taxonomicName id="4C064D51A600FF8BACF26EA8FDA68905" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[428,516,755,778]" class="Eutardigrada" family="Hypsibiidae" genus="Degmion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BACF26EA8FDA68905" box="[428,516,755,778]" italics="true" pageId="57" pageNumber="58">Degmion</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAF546EA8FDC98904" bold="true" box="[522,619,755,779]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAF546EA8FDC98904" box="[522,619,755,779]" pageId="57" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, see above.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A600FF8BAD2F6F10FD2A8E72" pageId="57" pageNumber="58" type="discussion">
|
||||
<paragraph id="8BB936D2A600FF8BAD2F6F10FD2A8E72" blockId="57.[113,763,843,1150]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B972EAC0A600FF8BAD2F6F10FF6D896C" box="[113,207,843,867]" italics="true" pageId="57" pageNumber="58">Remarks:</emphasis>
|
||||
Large pores used by
|
||||
<bibRefCitation id="EF974B23A600FF8BACF96F10FDFD896C" author="Buda J & Olszanowski Z & Wierzgon M" box="[423,607,843,867]" pageId="57" pageNumber="58" pagination="425 - 53" refId="ref35328" refString="Buda J, Olszanowski Z, Wierzgon M et al. Tardigrades and oribatid mites in bryophytes from geothermally active lava fields (Krafla, Iceland) and the description of Pilatobius islandicus sp. nov. (Eutardigrada). Polish Polar Research 2018; 39: 425 - 53. https: // doi. org / 10.24425 / 118755" type="journal article" year="2018">
|
||||
Buda
|
||||
<emphasis id="B972EAC0A600FF8BACBF6F17FDB2896C" box="[481,528,843,867]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||||
(2018)
|
||||
</bibRefCitation>
|
||||
to characterize
|
||||
<taxonomicName id="4C064D51A600FF8BAD2C6F30FF53898C" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="2024" box="[114,241,875,899]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="islandicum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAD2C6F30FF53898C" box="[114,241,875,899]" italics="true" pageId="57" pageNumber="58">F. islandicum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BADA86F30FECE898C" bold="true" box="[246,364,875,899]" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BADA86F30FECE898C" box="[246,364,875,899]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
are muscle attachment points that have no taxonomic value. With the exception of
|
||||
<taxonomicName id="4C064D51A600FF8BAF1A6FD0FD0F89AD" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="2024" box="[580,685,906,930]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="brevipes" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A600FF8BAF1A6FD0FD0F89AD" box="[580,685,906,930]" italics="true" pageId="57" pageNumber="58">F. brevipes</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A600FF8BAFE96FD1FF0289CE" bold="true" pageId="57" pageNumber="58">
|
||||
<taxonomicNameLabel id="A24157BBA600FF8BAFE96FD1FF0289CE" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, the species in the
|
||||
<taxonomicName id="4C064D51A600FF8BAC396FF1FE6289CE" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1911" box="[359,448,938,961]" class="Eutardigrada" family="Hypsibiidae" genus="Fontourion" kingdom="Animalia" order="Parachela" pageId="57" pageNumber="58" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A600FF8BAC396FF1FE6289CE" box="[359,448,938,961]" italics="true" pageId="57" pageNumber="58">recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
complex are extremely similar (
|
||||
<bibRefCitation id="EF974B23A600FF8BAD226F92FEFD89EE" author="Zawierucha K & Buda J & Novotna Jaromerska T" box="[124,351,969,993]" pageId="57" pageNumber="58" pagination="141 - 65" refId="ref40452" refString="Zawierucha K, Buda J, Novotna Jaromerska T et al. Integrative approach reveals new species of water bears (Pilatobius, Grevenius, and Acutuncus) from Arctic cryoconite holes, with the discovery of hidden lineages of Hypsibius. Zoologischer Anzeiger 2020; 289: 141 - 65. https: // doi. org / 10.1016 / j. jcz. 2020.09.004" type="journal article" year="2020">
|
||||
Zawierucha
|
||||
<emphasis id="B972EAC0A600FF8BADA46F92FE8589EE" box="[250,295,969,993]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||||
2020
|
||||
</bibRefCitation>
|
||||
), and we are of the opinion that the morphometric differences used to delineate them could result from discrepancies in the morphometry conducted by the various researchers. The species are clearly delineated genetically (
|
||||
<figureCitation id="133D2A57A600FF8BAFEF687CFD4B8E30" box="[689,745,1063,1087]" captionStart="Figure 1" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[113,1460,148,1477]" captionTargetPageId="3" captionText="Figure 1. Schematic depiction of six types of cuticular pedal bars and other leg structures found in the Hypsibiidae. Note that the edges of convex pulvini may overlap with fully sclerotized cuticular bars but are generally faint and weakly defined when observed under light microscopy. Historically, these edges of pulvini were sometimes described as ‘bars’ [e.g. see Dastych (1988) for Adropion belgicae or Gąsiorek et al. (2016) for Mesocrista]." figureDoi="http://doi.org/10.5281/zenodo.14243744" httpUri="https://zenodo.org/record/14243744/files/figure.png" pageId="57" pageNumber="58">Fig. 1</figureCitation>
|
||||
), thus until proper redescriptions are performed, species identification in this lineage should rely on DNA barcoding.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
978
data/03/AF/87/03AF87C4A609FF86AE8B6A86FECE8D4D.xml
Normal file
978
data/03/AF/87/03AF87C4A609FF86AE8B6A86FECE8D4D.xml
Normal file
|
|
@ -0,0 +1,978 @@
|
|||
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<subSubSection id="C31C6559A609FF82AE8B6A86FA858CF8" box="[981,1319,1757,1784]" pageId="48" pageNumber="49" type="nomenclature">
|
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<paragraph id="8BB936D2A609FF82AE8B6A86FA858CF8" blockId="48.[981,1319,1757,1784]" box="[981,1319,1757,1784]" pageId="48" pageNumber="49">
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<taxonomicName id="4C064D51A609FF82AE8B6A86FA858CF8" ID-CoL="9WRBG" authority="(Ito, 1995)" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[981,1319,1757,1784]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A609FF82AE8B6A86FA858CF8" bold="true" box="[981,1319,1757,1784]" pageId="48" pageNumber="49">
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<emphasis id="B972EAC0A609FF82AE8B6A86FB0C8CF7" bold="true" box="[981,1198,1757,1784]" italics="true" pageId="48" pageNumber="49">Pilatobius sexbullatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A9E36A85FABE8CF8" author="Ito M" box="[1213,1308,1757,1784]" pageId="48" pageNumber="49" pagination="18 - 39" refId="ref36771" refString="Ito M. Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan, II. Family Hypsibiidae. Proceedings of the Japanese Society of Systematic Zoology 1995; 53: 18 - 39." type="journal article" year="1995">Ito, 1995</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A609FF83AE396B47FD908D7D" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="reference_group">
|
||||
<paragraph id="8BB936D2A609FF82AE396B47FA228D59" blockId="48.[871,1428,1820,1878]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B972EAC0A609FF82AE396B47FA228D59" bold="true" pageId="48" pageNumber="49">
|
||||
<taxonomicName id="4C064D51A609FF82AE396B47FB7B8D38" authorityName="Ito" authorityYear="1995" box="[871,1241,1820,1847]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="sexbullatum" subGenus="Diphascon">
|
||||
<emphasis id="B972EAC0A609FF82AE396B47FC768D38" bold="true" box="[871,980,1820,1847]" italics="true" pageId="48" pageNumber="49">Diphascon</emphasis>
|
||||
(
|
||||
<emphasis id="B972EAC0A609FF82AEBD6B47FBF28D38" bold="true" box="[995,1104,1820,1847]" italics="true" pageId="48" pageNumber="49">Diphascon</emphasis>
|
||||
)
|
||||
<emphasis id="B972EAC0A609FF82A9016B47FB7B8D38" bold="true" box="[1119,1241,1820,1847]" italics="true" pageId="48" pageNumber="49">sexbullatum</emphasis>
|
||||
</taxonomicName>
|
||||
; Narusawa-mura, Minamitsuru-gun, Mount Fuji,
|
||||
<collectingCountry id="F3117642A609FF82A99A6B67FAA08D59" box="[1220,1282,1852,1878]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A609FF82A8556B67FAD98D5A" author="Ito M" box="[1291,1403,1851,1878]" pageId="48" pageNumber="49" pagination="18 - 39" refId="ref36771" refString="Ito M. Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan, II. Family Hypsibiidae. Proceedings of the Japanese Society of Systematic Zoology 1995; 53: 18 - 39." type="journal article" year="1995">Ito (1995)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A609FF82AE0B6B20FAD18DBB" blockId="48.[853,1445,1914,1972]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B972EAC0A609FF82AE0B6B20FAD18DBB" bold="true" pageId="48" pageNumber="49">
|
||||
<taxonomicName id="4C064D51A609FF82AE0B6B20FBBE8D9A" authorityName="Ito" authorityYear="1995" box="[853,1052,1914,1941]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="sexbullatum" subGenus="Diphascon">
|
||||
<emphasis id="B972EAC0A609FF82AE0B6B20FCCF8D9A" bold="true" box="[853,877,1915,1941]" italics="true" pageId="48" pageNumber="49">D.</emphasis>
|
||||
(
|
||||
<emphasis id="B972EAC0A609FF82AE226B20FC318D9A" bold="true" box="[892,915,1915,1941]" italics="true" pageId="48" pageNumber="49">D.</emphasis>
|
||||
)
|
||||
<emphasis id="B972EAC0A609FF82AEFC6B21FBBE8D9A" bold="true" box="[930,1052,1914,1941]" italics="true" pageId="48" pageNumber="49">sexbullatum</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<collectingRegion id="49C2F830A609FF82A9766B21FB2B8D9B" box="[1064,1161,1914,1940]" country="Mongolia" name="Arhangay" pageId="48" pageNumber="49">Arhangaj</collectingRegion>
|
||||
Ajmak, Chubsugul Ajmak,
|
||||
<collectingCountry id="F3117642A609FF82AED66BC2FC538DBC" box="[904,1009,1945,1971]" name="Mongolia" pageId="48" pageNumber="49">Mongolia</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A609FF82AEA46BC1FACC8DBC" author="Kaczmarek L & Michalczyk L" box="[1018,1390,1945,1972]" pageId="48" pageNumber="49" pagination="11 - 23" refId="ref36955" refString="Kaczmarek L, Michalczyk L. The Tardigrada fauna of Mongolia (Central Asia) with a description of Isohypsibius altai sp. nov. (Eutardigrada: Hypsibiidae). Zoological Studies 2006; 45: 11 - 23." type="journal article" year="2006">Kaczmarek and Michalczyk (2006)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<caption id="DF79665AA608FF83AD2F6CD4FE2A8ACC" pageId="49" pageNumber="50" startId="49.[113,168,143,167]" targetBox="[113,1451,217,1720]" targetIsTable="true" targetPageId="49" targetType="table">
|
||||
<paragraph id="8BB936D2A608FF83AD2F6CD4FE2A8ACC" blockId="49.[113,1442,142,195]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AD2F6CD4FF6E8AA9" bold="true" box="[113,204,142,167]" pageId="49" pageNumber="50">Table 13.</emphasis>
|
||||
Measurements (in micrometres) of selected morphological structures of
|
||||
<taxonomicName id="4C064D51A608FF83AE226CD4FB6B8AA8" authority="(Bartos, 1935)" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[892,1225,142,167]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="49" pageNumber="50" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A608FF83AE226CD4FC748AA8" box="[892,982,143,167]" italics="true" pageId="49" pageNumber="50">Pilatobius</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A608FF83AEA86CD4FB9B8AA9" box="[1014,1081,143,166]" italics="true" pageId="49" pageNumber="50">rugosus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A608FF83A9166CD4FB1D8AA9" author="Bartos E" box="[1096,1215,142,167]" pageId="49" pageNumber="50" pagination="257 - 60" refId="ref34712" refString="Bartos E. Vier neue Hypsibius - Arten aus der Tschechoslovakei. Zoologischer Anzeiger 1935; 110: 257 - 60." type="journal article" year="1935">Bartoš, 1935</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
from Poland (PL.120) mounted in Hoyer’s medium.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A608FF83AD2F6C82FA3D8CB7" pageId="49" pageNumber="50">
|
||||
<table id="F906C472A608004DAD2F6C82FA098CB7" box="[113,1451,217,1720]" gridcols="8" gridrows="44" pageId="49" pageNumber="50">
|
||||
<tr id="35363490A608004DAD2F6C82FA098AFE" box="[113,1451,217,241]" gridrow="0" pageId="49" pageNumber="50" rowspan-3="1" rowspan-5="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F6C82FE428AFE" box="[113,480,217,241]" gridcol="0" gridrow="0" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AD2F6C82FF768AFE" bold="true" box="[113,212,217,241]" pageId="49" pageNumber="50">Character</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA608004DAF776C82FDE38AFE" box="[553,577,217,241]" gridcol="1" gridrow="0" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AF776C82FD998AFF" bold="true" box="[553,571,217,240]" italics="true" pageId="49" pageNumber="50">N</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA608004DAFCD6C82FD568AFE" box="[659,756,217,241]" gridcol="2" gridrow="0" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AFCD6C82FD708AFE" bold="true" box="[659,722,217,241]" pageId="49" pageNumber="50">Range</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA608004DA9536C82FBE78AFE" box="[1037,1093,217,241]" gridcol="4" gridrow="0" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9536C82FBE78AFE" bold="true" box="[1037,1093,217,241]" pageId="49" pageNumber="50">Mean</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA608004DA8516C82FA8D8AFE" box="[1295,1327,217,241]" gridcol="6" gridrow="0" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8516C82FA8C8AFE" bold="true" box="[1295,1326,217,241]" pageId="49" pageNumber="50">SD</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6D5EFA098B12" box="[113,1451,261,285]" gridrow="1" pageId="49" pageNumber="50" rowspan-0="1" rowspan-1="1">
|
||||
<td id="76E75DECA608004DAFCD6D5EFD568B12" box="[659,756,261,285]" gridcol="2" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AFCD6D5EFD118B12" bold="true" box="[659,691,261,285]" pageId="49" pageNumber="50">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DAE1B6D5EFC1E8B12" box="[837,956,261,285]" gridcol="3" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE1B6D5EFCFB8B13" bold="true" box="[837,857,261,284]" italics="true" pageId="49" pageNumber="50">pt</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536D5EFBE78B12" box="[1037,1093,261,285]" gridcol="4" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9536D5EFB8F8B12" bold="true" box="[1037,1069,261,285]" pageId="49" pageNumber="50">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9D66D5EFB1F8B12" box="[1160,1213,261,285]" gridcol="5" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66D5EFB3E8B13" bold="true" box="[1160,1180,261,284]" italics="true" pageId="49" pageNumber="50">pt</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516D5EFA8D8B12" box="[1295,1327,261,285]" gridcol="6" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8516D5EFA8D8B12" bold="true" box="[1295,1327,261,285]" pageId="49" pageNumber="50">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8DC6D5EFA098B12" box="[1410,1451,261,285]" gridcol="7" gridrow="1" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6D5EFA348B13" bold="true" box="[1410,1430,261,284]" italics="true" pageId="49" pageNumber="50">pt</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6D6EFA098B42" box="[113,1451,309,333]" gridrow="2" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6D6EFE428B42" box="[113,480,309,333]" gridcol="0" gridrow="2" pageId="49" pageNumber="50">Body length</th>
|
||||
<td id="76E75DECA608004DAF776D6EFDE38B42" box="[553,577,309,333]" gridcol="1" gridrow="2" pageId="49" pageNumber="50">23</td>
|
||||
<td id="76E75DECA608004DAFCD6D6EFD568B42" box="[659,756,309,333]" gridcol="2" gridrow="2" pageId="49" pageNumber="50">218–321</td>
|
||||
<td id="76E75DECA608004DAE1B6D6EFC1E8B42" box="[837,956,309,333]" gridcol="3" gridrow="2" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE146D6EFC158B42" box="[842,951,309,333]" italics="true" pageId="49" pageNumber="50">1005–1413</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536D6EFBE78B42" box="[1037,1093,309,333]" gridcol="4" gridrow="2" pageId="49" pageNumber="50">279</td>
|
||||
<td id="76E75DECA608004DA9D66D6EFB1F8B42" box="[1160,1213,309,333]" gridcol="5" gridrow="2" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66D6EFB1A8B42" box="[1160,1208,309,333]" italics="true" pageId="49" pageNumber="50">1229</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516D6EFA8D8B42" box="[1295,1327,309,333]" gridcol="6" gridrow="2" pageId="49" pageNumber="50">29</td>
|
||||
<td id="76E75DECA608004DA8DC6D6EFA098B42" box="[1410,1451,309,333]" gridcol="7" gridrow="2" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6D6EFA048B42" box="[1410,1446,309,333]" italics="true" pageId="49" pageNumber="50">103</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6D0CFA098B60" box="[113,1451,343,367]" gridrow="3" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F6D0CFE428B60" box="[113,480,343,367]" gridcol="0" gridrow="3" pageId="49" pageNumber="50">Buccopharyngeal tube</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6D23FA098B9E" box="[113,1451,376,401]" gridrow="4" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6D23FE428B9E" box="[113,480,376,401]" gridcol="0" gridrow="4" pageId="49" pageNumber="50">Buccal tube length</th>
|
||||
<td id="76E75DECA608004DAF776D23FDE38B9E" box="[553,577,376,401]" gridcol="1" gridrow="4" pageId="49" pageNumber="50">18</td>
|
||||
<td id="76E75DECA608004DAFCD6D23FD568B9E" box="[659,756,376,401]" gridcol="2" gridrow="4" pageId="49" pageNumber="50">20.0–24.9</td>
|
||||
<td id="76E75DECA608004DAE1B6D23FC1E8B9E" box="[837,956,376,401]" gridcol="3" gridrow="4" pageId="49" pageNumber="50">–</td>
|
||||
<td id="76E75DECA608004DA9536D23FBE78B9E" box="[1037,1093,376,401]" gridcol="4" gridrow="4" pageId="49" pageNumber="50">22.7</td>
|
||||
<td id="76E75DECA608004DA9D66D23FB1F8B9E" box="[1160,1213,376,401]" gridcol="5" gridrow="4" pageId="49" pageNumber="50">–</td>
|
||||
<td id="76E75DECA608004DA8516D23FA8D8B9E" box="[1295,1327,376,401]" gridcol="6" gridrow="4" pageId="49" pageNumber="50">1.5</td>
|
||||
<td id="76E75DECA608004DA8DC6D23FA098B9E" box="[1410,1451,376,401]" gridcol="7" gridrow="4" pageId="49" pageNumber="50">–</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6DC1FA098BBD" box="[113,1451,410,434]" gridrow="5" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6DC1FE428BBD" box="[113,480,410,434]" gridcol="0" gridrow="5" pageId="49" pageNumber="50">Pharyngeal tube length</th>
|
||||
<td id="76E75DECA608004DAF776DC1FDE38BBD" box="[553,577,410,434]" gridcol="1" gridrow="5" pageId="49" pageNumber="50">15</td>
|
||||
<td id="76E75DECA608004DAFCD6DC1FD568BBD" box="[659,756,410,434]" gridcol="2" gridrow="5" pageId="49" pageNumber="50">27.5–32.8</td>
|
||||
<td id="76E75DECA608004DAE1B6DC1FC1E8BBD" box="[837,956,410,434]" gridcol="3" gridrow="5" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE1B6DC1FC198BBD" box="[837,955,410,434]" italics="true" pageId="49" pageNumber="50">114.3–150.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536DC1FBE78BBD" box="[1037,1093,410,434]" gridcol="4" gridrow="5" pageId="49" pageNumber="50">30.2</td>
|
||||
<td id="76E75DECA608004DA9D66DC1FB1F8BBD" box="[1160,1213,410,434]" gridcol="5" gridrow="5" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66DC1FB1F8BBD" box="[1160,1213,410,434]" italics="true" pageId="49" pageNumber="50">132.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516DC1FA8D8BBD" box="[1295,1327,410,434]" gridcol="6" gridrow="5" pageId="49" pageNumber="50">1.7</td>
|
||||
<td id="76E75DECA608004DA8DC6DC1FA098BBD" box="[1410,1451,410,434]" gridcol="7" gridrow="5" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6DC1FA098BBD" box="[1410,1451,410,434]" italics="true" pageId="49" pageNumber="50">10.4</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6DE7FA098BDB" box="[113,1451,444,468]" gridrow="6" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6DE7FE428BDB" box="[113,480,444,468]" gridcol="0" gridrow="6" pageId="49" pageNumber="50">Buccopharyngeal tube length</th>
|
||||
<td id="76E75DECA608004DAF776DE7FDE38BDB" box="[553,577,444,468]" gridcol="1" gridrow="6" pageId="49" pageNumber="50">15</td>
|
||||
<td id="76E75DECA608004DAFCD6DE7FD568BDB" box="[659,756,444,468]" gridcol="2" gridrow="6" pageId="49" pageNumber="50">49.5–56.7</td>
|
||||
<td id="76E75DECA608004DAE1B6DE7FC1E8BDB" box="[837,956,444,468]" gridcol="3" gridrow="6" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE1B6DE7FC1E8BDB" box="[837,956,444,468]" italics="true" pageId="49" pageNumber="50">214.3–250.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536DE7FBE78BDB" box="[1037,1093,444,468]" gridcol="4" gridrow="6" pageId="49" pageNumber="50">53.2</td>
|
||||
<td id="76E75DECA608004DA9D66DE7FB1F8BDB" box="[1160,1213,444,468]" gridcol="5" gridrow="6" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66DE7FB1F8BDB" box="[1160,1213,444,468]" italics="true" pageId="49" pageNumber="50">232.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516DE7FA8D8BDB" box="[1295,1327,444,468]" gridcol="6" gridrow="6" pageId="49" pageNumber="50">2.3</td>
|
||||
<td id="76E75DECA608004DA8DC6DE7FA098BDB" box="[1410,1451,444,468]" gridcol="7" gridrow="6" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6DE7FA098BDB" box="[1410,1451,444,468]" italics="true" pageId="49" pageNumber="50">10.4</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6D85FA098BF9" box="[113,1451,478,502]" gridrow="7" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6D85FE428BF9" box="[113,480,478,502]" gridcol="0" gridrow="7" pageId="49" pageNumber="50">Buccal/pharyngeal tube length ratio</th>
|
||||
<td id="76E75DECA608004DAF776D85FDE38BF9" box="[553,577,478,502]" gridcol="1" gridrow="7" pageId="49" pageNumber="50">15</td>
|
||||
<td id="76E75DECA608004DAFCD6D85FD568BF9" box="[659,756,478,502]" gridcol="2" gridrow="7" pageId="49" pageNumber="50">67%–88%</td>
|
||||
<td id="76E75DECA608004DAE1B6D85FC1E8BF9" box="[837,956,478,502]" gridcol="3" gridrow="7" pageId="49" pageNumber="50">–</td>
|
||||
<td id="76E75DECA608004DA9536D85FBE78BF9" box="[1037,1093,478,502]" gridcol="4" gridrow="7" pageId="49" pageNumber="50">76%</td>
|
||||
<td id="76E75DECA608004DA9D66D85FB1F8BF9" box="[1160,1213,478,502]" gridcol="5" gridrow="7" pageId="49" pageNumber="50">–</td>
|
||||
<td id="76E75DECA608004DA8516D85FA8D8BF9" box="[1295,1327,478,502]" gridcol="6" gridrow="7" pageId="49" pageNumber="50">6%</td>
|
||||
<td id="76E75DECA608004DA8DC6D85FA098BF9" box="[1410,1451,478,502]" gridcol="7" gridrow="7" pageId="49" pageNumber="50">–</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6E5BFA098817" box="[113,1451,512,536]" gridrow="8" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6E5BFE428817" box="[113,480,512,536]" gridcol="0" gridrow="8" pageId="49" pageNumber="50">Stylet support insertion point</th>
|
||||
<td id="76E75DECA608004DAF776E5BFDE38817" box="[553,577,512,536]" gridcol="1" gridrow="8" pageId="49" pageNumber="50">18</td>
|
||||
<td id="76E75DECA608004DAFCD6E5BFD568817" box="[659,756,512,536]" gridcol="2" gridrow="8" pageId="49" pageNumber="50">12.5–15.5</td>
|
||||
<td id="76E75DECA608004DAE1B6E5BFC1E8817" box="[837,956,512,536]" gridcol="3" gridrow="8" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6E5BFC128817" box="[849,944,512,536]" italics="true" pageId="49" pageNumber="50">60.1–63.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536E5BFBE78817" box="[1037,1093,512,536]" gridcol="4" gridrow="8" pageId="49" pageNumber="50">14.1</td>
|
||||
<td id="76E75DECA608004DA9D66E5BFB1F8817" box="[1160,1213,512,536]" gridcol="5" gridrow="8" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66E5BFB138817" box="[1160,1201,512,536]" italics="true" pageId="49" pageNumber="50">62.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516E5BFA8D8817" box="[1295,1327,512,536]" gridcol="6" gridrow="8" pageId="49" pageNumber="50">0.9</td>
|
||||
<td id="76E75DECA608004DA8DC6E5BFA098817" box="[1410,1451,512,536]" gridcol="7" gridrow="8" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6E5BFA3D8817" box="[1410,1439,512,536]" italics="true" pageId="49" pageNumber="50">1.0</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6E79FA098835" box="[113,1451,546,570]" gridrow="9" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6E79FE428835" box="[113,480,546,570]" gridcol="0" gridrow="9" pageId="49" pageNumber="50">Buccal tube external width</th>
|
||||
<td id="76E75DECA608004DAF776E79FDE38835" box="[553,577,546,570]" gridcol="1" gridrow="9" pageId="49" pageNumber="50">17</td>
|
||||
<td id="76E75DECA608004DAFCD6E79FD568835" box="[659,756,546,570]" gridcol="2" gridrow="9" pageId="49" pageNumber="50">1.3–1.8</td>
|
||||
<td id="76E75DECA608004DAE1B6E79FC1E8835" box="[837,956,546,570]" gridcol="3" gridrow="9" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE036E79FC068835" box="[861,932,546,570]" italics="true" pageId="49" pageNumber="50">5.9–8.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536E79FBE78835" box="[1037,1093,546,570]" gridcol="4" gridrow="9" pageId="49" pageNumber="50">1.6</td>
|
||||
<td id="76E75DECA608004DA9D66E79FB1F8835" box="[1160,1213,546,570]" gridcol="5" gridrow="9" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66E79FB078835" box="[1160,1189,546,570]" italics="true" pageId="49" pageNumber="50">7.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516E79FA8D8835" box="[1295,1327,546,570]" gridcol="6" gridrow="9" pageId="49" pageNumber="50">0.1</td>
|
||||
<td id="76E75DECA608004DA8DC6E79FA098835" box="[1410,1451,546,570]" gridcol="7" gridrow="9" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6E79FA3D8835" box="[1410,1439,546,570]" italics="true" pageId="49" pageNumber="50">0.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6E18FA098853" box="[113,1451,579,604]" gridrow="10" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6E18FE428853" box="[113,480,579,604]" gridcol="0" gridrow="10" pageId="49" pageNumber="50">Buccal tube internal width</th>
|
||||
<td id="76E75DECA608004DAF776E18FDE38853" box="[553,577,579,604]" gridcol="1" gridrow="10" pageId="49" pageNumber="50">17</td>
|
||||
<td id="76E75DECA608004DAFCD6E18FD568853" box="[659,756,579,604]" gridcol="2" gridrow="10" pageId="49" pageNumber="50">0.4–0.7</td>
|
||||
<td id="76E75DECA608004DAE1B6E18FC1E8853" box="[837,956,579,604]" gridcol="3" gridrow="10" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE036E1FFC018853" box="[861,931,580,604]" italics="true" pageId="49" pageNumber="50">2.0–3.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536E18FBE78853" box="[1037,1093,579,604]" gridcol="4" gridrow="10" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA9D66E18FB1F8853" box="[1160,1213,579,604]" gridcol="5" gridrow="10" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66E1FFB078853" box="[1160,1189,580,604]" italics="true" pageId="49" pageNumber="50">2.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516E18FA8D8853" box="[1295,1327,579,604]" gridcol="6" gridrow="10" pageId="49" pageNumber="50">0.1</td>
|
||||
<td id="76E75DECA608004DA8DC6E18FA098853" box="[1410,1451,579,604]" gridcol="7" gridrow="10" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6E1FFA3D8853" box="[1410,1439,580,604]" italics="true" pageId="49" pageNumber="50">0.4</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6E3EFA098872" box="[113,1451,613,637]" gridrow="11" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F6E3EFE428872" box="[113,480,613,637]" gridcol="0" gridrow="11" pageId="49" pageNumber="50">Placoid lengths</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6EDCFA098890" box="[113,1451,647,671]" gridrow="12" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6EDCFE428890" box="[113,480,647,671]" gridcol="0" gridrow="12" pageId="49" pageNumber="50">Macroplacoid 1</th>
|
||||
<td id="76E75DECA608004DAF776EDCFDE38890" box="[553,577,647,671]" gridcol="1" gridrow="12" pageId="49" pageNumber="50">21</td>
|
||||
<td id="76E75DECA608004DAFCD6EDCFD568890" box="[659,756,647,671]" gridcol="2" gridrow="12" pageId="49" pageNumber="50">4.7–6.2</td>
|
||||
<td id="76E75DECA608004DAE1B6EDCFC1E8890" box="[837,956,647,671]" gridcol="3" gridrow="12" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6EDCFC128890" box="[849,944,647,671]" italics="true" pageId="49" pageNumber="50">21.7–26.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536EDCFBE78890" box="[1037,1093,647,671]" gridcol="4" gridrow="12" pageId="49" pageNumber="50">5.5</td>
|
||||
<td id="76E75DECA608004DA9D66EDCFB1F8890" box="[1160,1213,647,671]" gridcol="5" gridrow="12" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66EDCFB138890" box="[1160,1201,647,671]" italics="true" pageId="49" pageNumber="50">24.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516EDCFA8D8890" box="[1295,1327,647,671]" gridcol="6" gridrow="12" pageId="49" pageNumber="50">0.4</td>
|
||||
<td id="76E75DECA608004DA8DC6EDCFA098890" box="[1410,1451,647,671]" gridcol="7" gridrow="12" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6EDCFA3D8890" box="[1410,1439,647,671]" italics="true" pageId="49" pageNumber="50">1.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6EF2FA0988CE" box="[113,1451,681,705]" gridrow="13" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6EF2FE4288CE" box="[113,480,681,705]" gridcol="0" gridrow="13" pageId="49" pageNumber="50">Macroplacoid 2</th>
|
||||
<td id="76E75DECA608004DAF776EF2FDE388CE" box="[553,577,681,705]" gridcol="1" gridrow="13" pageId="49" pageNumber="50">21</td>
|
||||
<td id="76E75DECA608004DAFCD6EF2FD5688CE" box="[659,756,681,705]" gridcol="2" gridrow="13" pageId="49" pageNumber="50">3.2–4.8</td>
|
||||
<td id="76E75DECA608004DAE1B6EF2FC1E88CE" box="[837,956,681,705]" gridcol="3" gridrow="13" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6EF2FC1288CE" box="[849,944,681,705]" italics="true" pageId="49" pageNumber="50">15.4–22.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536EF2FBE788CE" box="[1037,1093,681,705]" gridcol="4" gridrow="13" pageId="49" pageNumber="50">4.0</td>
|
||||
<td id="76E75DECA608004DA9D66EF2FB1F88CE" box="[1160,1213,681,705]" gridcol="5" gridrow="13" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66EF2FB1388CE" box="[1160,1201,681,705]" italics="true" pageId="49" pageNumber="50">17.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516EF2FA8D88CE" box="[1295,1327,681,705]" gridcol="6" gridrow="13" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6EF2FA0988CE" box="[1410,1451,681,705]" gridcol="7" gridrow="13" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6EF2FA3D88CE" box="[1410,1439,681,705]" italics="true" pageId="49" pageNumber="50">2.1</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6E90FA0988EC" box="[113,1451,715,739]" gridrow="14" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6E90FE4288EC" box="[113,480,715,739]" gridcol="0" gridrow="14" pageId="49" pageNumber="50">Septulum</th>
|
||||
<td id="76E75DECA608004DAF776E90FDE388EC" box="[553,577,715,739]" gridcol="1" gridrow="14" pageId="49" pageNumber="50">21</td>
|
||||
<td id="76E75DECA608004DAFCD6E90FD5688EC" box="[659,756,715,739]" gridcol="2" gridrow="14" pageId="49" pageNumber="50">1.5–3.0</td>
|
||||
<td id="76E75DECA608004DAE1B6E90FC1E88EC" box="[837,956,715,739]" gridcol="3" gridrow="14" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE036E90FC0D88EC" box="[861,943,715,739]" italics="true" pageId="49" pageNumber="50">7.8–12.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536E90FBE788EC" box="[1037,1093,715,739]" gridcol="4" gridrow="14" pageId="49" pageNumber="50">2.3</td>
|
||||
<td id="76E75DECA608004DA9D66E90FB1F88EC" box="[1160,1213,715,739]" gridcol="5" gridrow="14" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66E90FB1388EC" box="[1160,1201,715,739]" italics="true" pageId="49" pageNumber="50">10.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516E90FA8D88EC" box="[1295,1327,715,739]" gridcol="6" gridrow="14" pageId="49" pageNumber="50">0.3</td>
|
||||
<td id="76E75DECA608004DA8DC6E90FA0988EC" box="[1410,1451,715,739]" gridcol="7" gridrow="14" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6E90FA3D88EC" box="[1410,1439,715,739]" italics="true" pageId="49" pageNumber="50">1.3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6EB6FA09890A" box="[113,1451,749,773]" gridrow="15" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6EB6FE42890A" box="[113,480,749,773]" gridcol="0" gridrow="15" pageId="49" pageNumber="50">Macroplacoid row</th>
|
||||
<td id="76E75DECA608004DAF776EB6FDE3890A" box="[553,577,749,773]" gridcol="1" gridrow="15" pageId="49" pageNumber="50">21</td>
|
||||
<td id="76E75DECA608004DAFCD6EB6FD56890A" box="[659,756,749,773]" gridcol="2" gridrow="15" pageId="49" pageNumber="50">8.8–11.5</td>
|
||||
<td id="76E75DECA608004DAE1B6EB6FC1E890A" box="[837,956,749,773]" gridcol="3" gridrow="15" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6EB6FC12890A" box="[849,944,749,773]" italics="true" pageId="49" pageNumber="50">42.4–51.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536EB6FBE7890A" box="[1037,1093,749,773]" gridcol="4" gridrow="15" pageId="49" pageNumber="50">10.4</td>
|
||||
<td id="76E75DECA608004DA9D66EB6FB1F890A" box="[1160,1213,749,773]" gridcol="5" gridrow="15" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66EB6FB13890A" box="[1160,1201,749,773]" italics="true" pageId="49" pageNumber="50">46.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516EB6FA8D890A" box="[1295,1327,749,773]" gridcol="6" gridrow="15" pageId="49" pageNumber="50">0.8</td>
|
||||
<td id="76E75DECA608004DA8DC6EB6FA09890A" box="[1410,1451,749,773]" gridcol="7" gridrow="15" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6EB6FA3D890A" box="[1410,1439,749,773]" italics="true" pageId="49" pageNumber="50">2.5</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6F54FA098928" box="[113,1451,783,807]" gridrow="16" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F6F54FE428928" box="[113,480,783,807]" gridcol="0" gridrow="16" pageId="49" pageNumber="50">Claw 1 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6F6BFA098946" box="[113,1451,816,841]" gridrow="17" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6F6BFE428946" box="[113,480,816,841]" gridcol="0" gridrow="17" pageId="49" pageNumber="50">External base</th>
|
||||
<td id="76E75DECA608004DAF776F6BFDE38946" box="[553,577,816,841]" gridcol="1" gridrow="17" pageId="49" pageNumber="50">7</td>
|
||||
<td id="76E75DECA608004DAFCD6F6BFD568946" box="[659,756,816,841]" gridcol="2" gridrow="17" pageId="49" pageNumber="50">2.8–4.2</td>
|
||||
<td id="76E75DECA608004DAE1B6F6BFC1E8946" box="[837,956,816,841]" gridcol="3" gridrow="17" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6F6AFC128946" box="[849,944,817,841]" italics="true" pageId="49" pageNumber="50">14.0–17.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536F6BFBE78946" box="[1037,1093,816,841]" gridcol="4" gridrow="17" pageId="49" pageNumber="50">3.4</td>
|
||||
<td id="76E75DECA608004DA9D66F6BFB1F8946" box="[1160,1213,816,841]" gridcol="5" gridrow="17" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66F6AFB138946" box="[1160,1201,817,841]" italics="true" pageId="49" pageNumber="50">15.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516F6BFA8D8946" box="[1295,1327,816,841]" gridcol="6" gridrow="17" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6F6BFA098946" box="[1410,1451,816,841]" gridcol="7" gridrow="17" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6F6AFA3D8946" box="[1410,1439,817,841]" italics="true" pageId="49" pageNumber="50">2.0</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6F09FA098965" box="[113,1451,850,874]" gridrow="18" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6F09FE428965" box="[113,480,850,874]" gridcol="0" gridrow="18" pageId="49" pageNumber="50">External primary branch</th>
|
||||
<td id="76E75DECA608004DAF776F09FDE38965" box="[553,577,850,874]" gridcol="1" gridrow="18" pageId="49" pageNumber="50">2</td>
|
||||
<td id="76E75DECA608004DAFCD6F09FD568965" box="[659,756,850,874]" gridcol="2" gridrow="18" pageId="49" pageNumber="50">7.7–8.3</td>
|
||||
<td id="76E75DECA608004DAE1B6F09FC1E8965" box="[837,956,850,874]" gridcol="3" gridrow="18" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6F09FC128965" box="[849,944,850,874]" italics="true" pageId="49" pageNumber="50">35.2–35.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536F09FBE78965" box="[1037,1093,850,874]" gridcol="4" gridrow="18" pageId="49" pageNumber="50">8.0</td>
|
||||
<td id="76E75DECA608004DA9D66F09FB1F8965" box="[1160,1213,850,874]" gridcol="5" gridrow="18" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66F09FB138965" box="[1160,1201,850,874]" italics="true" pageId="49" pageNumber="50">35.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516F09FA8D8965" box="[1295,1327,850,874]" gridcol="6" gridrow="18" pageId="49" pageNumber="50">0.4</td>
|
||||
<td id="76E75DECA608004DA8DC6F09FA098965" box="[1410,1451,850,874]" gridcol="7" gridrow="18" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6F08FA298965" box="[1410,1419,851,874]" italics="true" pageId="49" pageNumber="50">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6F2FFA098983" box="[113,1451,884,908]" gridrow="19" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6F2FFE428983" box="[113,480,884,908]" gridcol="0" gridrow="19" pageId="49" pageNumber="50">External secondary branch</th>
|
||||
<td id="76E75DECA608004DAF776F2FFDE38983" box="[553,577,884,908]" gridcol="1" gridrow="19" pageId="49" pageNumber="50">7</td>
|
||||
<td id="76E75DECA608004DAFCD6F2FFD568983" box="[659,756,884,908]" gridcol="2" gridrow="19" pageId="49" pageNumber="50">5.7–7.0</td>
|
||||
<td id="76E75DECA608004DAE1B6F2FFC1E8983" box="[837,956,884,908]" gridcol="3" gridrow="19" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6F2FFC128983" box="[849,944,884,908]" italics="true" pageId="49" pageNumber="50">26.0–31.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536F2FFBE78983" box="[1037,1093,884,908]" gridcol="4" gridrow="19" pageId="49" pageNumber="50">6.2</td>
|
||||
<td id="76E75DECA608004DA9D66F2FFB1F8983" box="[1160,1213,884,908]" gridcol="5" gridrow="19" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66F2FFB138983" box="[1160,1201,884,908]" italics="true" pageId="49" pageNumber="50">28.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516F2FFA8D8983" box="[1295,1327,884,908]" gridcol="6" gridrow="19" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6F2FFA098983" box="[1410,1451,884,908]" gridcol="7" gridrow="19" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6F2FFA3D8983" box="[1410,1439,884,908]" italics="true" pageId="49" pageNumber="50">2.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6FCDFA0989A1" box="[113,1451,918,942]" gridrow="20" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6FCDFE4289A1" box="[113,480,918,942]" gridcol="0" gridrow="20" pageId="49" pageNumber="50">Internal base</th>
|
||||
<td id="76E75DECA608004DAF776FCDFDE389A1" box="[553,577,918,942]" gridcol="1" gridrow="20" pageId="49" pageNumber="50">8</td>
|
||||
<td id="76E75DECA608004DAFCD6FCDFD5689A1" box="[659,756,918,942]" gridcol="2" gridrow="20" pageId="49" pageNumber="50">3.0–4.6</td>
|
||||
<td id="76E75DECA608004DAE1B6FCDFC1E89A1" box="[837,956,918,942]" gridcol="3" gridrow="20" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6FCDFC1289A1" box="[849,944,918,942]" italics="true" pageId="49" pageNumber="50">14.0–18.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536FCDFBE789A1" box="[1037,1093,918,942]" gridcol="4" gridrow="20" pageId="49" pageNumber="50">3.6</td>
|
||||
<td id="76E75DECA608004DA9D66FCDFB1F89A1" box="[1160,1213,918,942]" gridcol="5" gridrow="20" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66FCDFB1389A1" box="[1160,1201,918,942]" italics="true" pageId="49" pageNumber="50">16.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516FCDFA8D89A1" box="[1295,1327,918,942]" gridcol="6" gridrow="20" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6FCDFA0989A1" box="[1410,1451,918,942]" gridcol="7" gridrow="20" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6FCDFA3D89A1" box="[1410,1439,918,942]" italics="true" pageId="49" pageNumber="50">2.3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6FE3FA0989DF" box="[113,1451,952,976]" gridrow="21" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6FE3FE4289DF" box="[113,480,952,976]" gridcol="0" gridrow="21" pageId="49" pageNumber="50">Internal primary branch</th>
|
||||
<td id="76E75DECA608004DAF776FE3FDE389DF" box="[553,577,952,976]" gridcol="1" gridrow="21" pageId="49" pageNumber="50">4</td>
|
||||
<td id="76E75DECA608004DAFCD6FE3FD5689DF" box="[659,756,952,976]" gridcol="2" gridrow="21" pageId="49" pageNumber="50">6.4–7.0</td>
|
||||
<td id="76E75DECA608004DAE1B6FE3FC1E89DF" box="[837,956,952,976]" gridcol="3" gridrow="21" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6FE3FC0D89DF" box="[849,943,952,976]" italics="true" pageId="49" pageNumber="50">28.6–32.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536FE3FBE789DF" box="[1037,1093,952,976]" gridcol="4" gridrow="21" pageId="49" pageNumber="50">6.7</td>
|
||||
<td id="76E75DECA608004DA9D66FE3FB1F89DF" box="[1160,1213,952,976]" gridcol="5" gridrow="21" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66FE3FB1389DF" box="[1160,1201,952,976]" italics="true" pageId="49" pageNumber="50">30.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516FE3FA8D89DF" box="[1295,1327,952,976]" gridcol="6" gridrow="21" pageId="49" pageNumber="50">0.3</td>
|
||||
<td id="76E75DECA608004DA8DC6FE3FA0989DF" box="[1410,1451,952,976]" gridcol="7" gridrow="21" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6FE3FA3D89DF" box="[1410,1439,952,976]" italics="true" pageId="49" pageNumber="50">2.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6F81FA0989FD" box="[113,1451,986,1010]" gridrow="22" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6F81FE4289FD" box="[113,480,986,1010]" gridcol="0" gridrow="22" pageId="49" pageNumber="50">Internal secondary branch</th>
|
||||
<td id="76E75DECA608004DAF776F81FDE389FD" box="[553,577,986,1010]" gridcol="1" gridrow="22" pageId="49" pageNumber="50">4</td>
|
||||
<td id="76E75DECA608004DAFCD6F81FD5689FD" box="[659,756,986,1010]" gridcol="2" gridrow="22" pageId="49" pageNumber="50">4.5–5.7</td>
|
||||
<td id="76E75DECA608004DAE1B6F81FC1E89FD" box="[837,956,986,1010]" gridcol="3" gridrow="22" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6F81FC0D89FD" box="[849,943,986,1010]" italics="true" pageId="49" pageNumber="50">21.6–24.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536F81FBE789FD" box="[1037,1093,986,1010]" gridcol="4" gridrow="22" pageId="49" pageNumber="50">5.1</td>
|
||||
<td id="76E75DECA608004DA9D66F81FB1F89FD" box="[1160,1213,986,1010]" gridcol="5" gridrow="22" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66F81FB1389FD" box="[1160,1201,986,1010]" italics="true" pageId="49" pageNumber="50">22.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516F81FA8D89FD" box="[1295,1327,986,1010]" gridcol="6" gridrow="22" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6F81FA0989FD" box="[1410,1451,986,1010]" gridcol="7" gridrow="22" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6F81FA3D89FD" box="[1410,1439,986,1010]" italics="true" pageId="49" pageNumber="50">1.8</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6FA0FA098E1B" box="[113,1451,1019,1044]" gridrow="23" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F6FA0FE428E1B" box="[113,480,1019,1044]" gridcol="0" gridrow="23" pageId="49" pageNumber="50">Claw 2 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6846FA098E3A" box="[113,1451,1053,1077]" gridrow="24" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6846FE428E3A" box="[113,480,1053,1077]" gridcol="0" gridrow="24" pageId="49" pageNumber="50">External base</th>
|
||||
<td id="76E75DECA608004DAF776846FDE38E3A" box="[553,577,1053,1077]" gridcol="1" gridrow="24" pageId="49" pageNumber="50">11</td>
|
||||
<td id="76E75DECA608004DAFCD6846FD568E3A" box="[659,756,1053,1077]" gridcol="2" gridrow="24" pageId="49" pageNumber="50">3.1–4.2</td>
|
||||
<td id="76E75DECA608004DAE1B6846FC1E8E3A" box="[837,956,1053,1077]" gridcol="3" gridrow="24" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6846FC128E3A" box="[849,944,1053,1077]" italics="true" pageId="49" pageNumber="50">13.3–18.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536846FBE78E3A" box="[1037,1093,1053,1077]" gridcol="4" gridrow="24" pageId="49" pageNumber="50">3.7</td>
|
||||
<td id="76E75DECA608004DA9D66846FB1F8E3A" box="[1160,1213,1053,1077]" gridcol="5" gridrow="24" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66846FB138E3A" box="[1160,1201,1053,1077]" italics="true" pageId="49" pageNumber="50">16.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516846FA8D8E3A" box="[1295,1327,1053,1077]" gridcol="6" gridrow="24" pageId="49" pageNumber="50">0.4</td>
|
||||
<td id="76E75DECA608004DA8DC6846FA098E3A" box="[1410,1451,1053,1077]" gridcol="7" gridrow="24" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6846FA3D8E3A" box="[1410,1439,1053,1077]" italics="true" pageId="49" pageNumber="50">1.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6864FA098E58" box="[113,1451,1087,1111]" gridrow="25" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6864FE428E58" box="[113,480,1087,1111]" gridcol="0" gridrow="25" pageId="49" pageNumber="50">External primary branch</th>
|
||||
<td id="76E75DECA608004DAF776864FDE38E58" box="[553,577,1087,1111]" gridcol="1" gridrow="25" pageId="49" pageNumber="50">6</td>
|
||||
<td id="76E75DECA608004DAFCD6864FD568E58" box="[659,756,1087,1111]" gridcol="2" gridrow="25" pageId="49" pageNumber="50">7.3–9.4</td>
|
||||
<td id="76E75DECA608004DAE1B6864FC1E8E58" box="[837,956,1087,1111]" gridcol="3" gridrow="25" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6864FC128E58" box="[849,944,1087,1111]" italics="true" pageId="49" pageNumber="50">33.9–36.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536864FBE78E58" box="[1037,1093,1087,1111]" gridcol="4" gridrow="25" pageId="49" pageNumber="50">8.3</td>
|
||||
<td id="76E75DECA608004DA9D66864FB1F8E58" box="[1160,1213,1087,1111]" gridcol="5" gridrow="25" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66864FB138E58" box="[1160,1201,1087,1111]" italics="true" pageId="49" pageNumber="50">35.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516864FA8D8E58" box="[1295,1327,1087,1111]" gridcol="6" gridrow="25" pageId="49" pageNumber="50">0.8</td>
|
||||
<td id="76E75DECA608004DA8DC6864FA098E58" box="[1410,1451,1087,1111]" gridcol="7" gridrow="25" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6864FA3D8E58" box="[1410,1439,1087,1111]" italics="true" pageId="49" pageNumber="50">1.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F683AFA098E76" box="[113,1451,1121,1145]" gridrow="26" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F683AFE428E76" box="[113,480,1121,1145]" gridcol="0" gridrow="26" pageId="49" pageNumber="50">External secondary branch</th>
|
||||
<td id="76E75DECA608004DAF77683AFDE38E76" box="[553,577,1121,1145]" gridcol="1" gridrow="26" pageId="49" pageNumber="50">11</td>
|
||||
<td id="76E75DECA608004DAFCD683AFD568E76" box="[659,756,1121,1145]" gridcol="2" gridrow="26" pageId="49" pageNumber="50">5.9–7.7</td>
|
||||
<td id="76E75DECA608004DAE1B683AFC1E8E76" box="[837,956,1121,1145]" gridcol="3" gridrow="26" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F683AFC128E76" box="[849,944,1121,1145]" italics="true" pageId="49" pageNumber="50">27.9–36.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA953683AFBE78E76" box="[1037,1093,1121,1145]" gridcol="4" gridrow="26" pageId="49" pageNumber="50">6.8</td>
|
||||
<td id="76E75DECA608004DA9D6683AFB1F8E76" box="[1160,1213,1121,1145]" gridcol="5" gridrow="26" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D6683AFB138E76" box="[1160,1201,1121,1145]" italics="true" pageId="49" pageNumber="50">29.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA851683AFA8D8E76" box="[1295,1327,1121,1145]" gridcol="6" gridrow="26" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA8DC683AFA098E76" box="[1410,1451,1121,1145]" gridcol="7" gridrow="26" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC683AFA3D8E76" box="[1410,1439,1121,1145]" italics="true" pageId="49" pageNumber="50">2.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F68D8FA098E94" box="[113,1451,1155,1179]" gridrow="27" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F68D8FE428E94" box="[113,480,1155,1179]" gridcol="0" gridrow="27" pageId="49" pageNumber="50">Internal base</th>
|
||||
<td id="76E75DECA608004DAF7768D8FDE38E94" box="[553,577,1155,1179]" gridcol="1" gridrow="27" pageId="49" pageNumber="50">9</td>
|
||||
<td id="76E75DECA608004DAFCD68D8FD568E94" box="[659,756,1155,1179]" gridcol="2" gridrow="27" pageId="49" pageNumber="50">3.0–3.9</td>
|
||||
<td id="76E75DECA608004DAE1B68D8FC1E8E94" box="[837,956,1155,1179]" gridcol="3" gridrow="27" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F68D8FC128E94" box="[849,944,1155,1179]" italics="true" pageId="49" pageNumber="50">13.7–18.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA95368D8FBE78E94" box="[1037,1093,1155,1179]" gridcol="4" gridrow="27" pageId="49" pageNumber="50">3.6</td>
|
||||
<td id="76E75DECA608004DA9D668D8FB1F8E94" box="[1160,1213,1155,1179]" gridcol="5" gridrow="27" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D668D8FB138E94" box="[1160,1201,1155,1179]" italics="true" pageId="49" pageNumber="50">16.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA85168D8FA8D8E94" box="[1295,1327,1155,1179]" gridcol="6" gridrow="27" pageId="49" pageNumber="50">0.3</td>
|
||||
<td id="76E75DECA608004DA8DC68D8FA098E94" box="[1410,1451,1155,1179]" gridcol="7" gridrow="27" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC68D8FA3D8E94" box="[1410,1439,1155,1179]" italics="true" pageId="49" pageNumber="50">1.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F68FEFA098EB2" box="[113,1451,1189,1213]" gridrow="28" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F68FEFE428EB2" box="[113,480,1189,1213]" gridcol="0" gridrow="28" pageId="49" pageNumber="50">Internal primary branch</th>
|
||||
<td id="76E75DECA608004DAF7768FEFDE38EB2" box="[553,577,1189,1213]" gridcol="1" gridrow="28" pageId="49" pageNumber="50">6</td>
|
||||
<td id="76E75DECA608004DAFCD68FEFD568EB2" box="[659,756,1189,1213]" gridcol="2" gridrow="28" pageId="49" pageNumber="50">5.6–7.6</td>
|
||||
<td id="76E75DECA608004DAE1B68FEFC1E8EB2" box="[837,956,1189,1213]" gridcol="3" gridrow="28" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F68FEFC128EB2" box="[849,944,1189,1213]" italics="true" pageId="49" pageNumber="50">26.7–33.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA95368FEFBE78EB2" box="[1037,1093,1189,1213]" gridcol="4" gridrow="28" pageId="49" pageNumber="50">6.7</td>
|
||||
<td id="76E75DECA608004DA9D668FEFB1F8EB2" box="[1160,1213,1189,1213]" gridcol="5" gridrow="28" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D668FEFB138EB2" box="[1160,1201,1189,1213]" italics="true" pageId="49" pageNumber="50">30.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA85168FEFA8D8EB2" box="[1295,1327,1189,1213]" gridcol="6" gridrow="28" pageId="49" pageNumber="50">0.7</td>
|
||||
<td id="76E75DECA608004DA8DC68FEFA098EB2" box="[1410,1451,1189,1213]" gridcol="7" gridrow="28" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC68FEFA3D8EB2" box="[1410,1439,1189,1213]" italics="true" pageId="49" pageNumber="50">3.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F689DFA098ED0" box="[113,1451,1222,1247]" gridrow="29" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F689DFE428ED0" box="[113,480,1222,1247]" gridcol="0" gridrow="29" pageId="49" pageNumber="50">Internal secondary branch</th>
|
||||
<td id="76E75DECA608004DAF77689DFDE38ED0" box="[553,577,1222,1247]" gridcol="1" gridrow="29" pageId="49" pageNumber="50">5</td>
|
||||
<td id="76E75DECA608004DAFCD689DFD568ED0" box="[659,756,1222,1247]" gridcol="2" gridrow="29" pageId="49" pageNumber="50">4.8–5.4</td>
|
||||
<td id="76E75DECA608004DAE1B689DFC1E8ED0" box="[837,956,1222,1247]" gridcol="3" gridrow="29" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F689CFC128ED0" box="[849,944,1223,1247]" italics="true" pageId="49" pageNumber="50">22.0–23.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA953689DFBE78ED0" box="[1037,1093,1222,1247]" gridcol="4" gridrow="29" pageId="49" pageNumber="50">5.1</td>
|
||||
<td id="76E75DECA608004DA9D6689DFB1F8ED0" box="[1160,1213,1222,1247]" gridcol="5" gridrow="29" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D6689CFB138ED0" box="[1160,1201,1223,1247]" italics="true" pageId="49" pageNumber="50">22.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA851689DFA8D8ED0" box="[1295,1327,1222,1247]" gridcol="6" gridrow="29" pageId="49" pageNumber="50">0.3</td>
|
||||
<td id="76E75DECA608004DA8DC689DFA098ED0" box="[1410,1451,1222,1247]" gridcol="7" gridrow="29" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC689CFA3D8ED0" box="[1410,1439,1223,1247]" italics="true" pageId="49" pageNumber="50">0.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F68B3FA098F0E" box="[113,1451,1256,1281]" gridrow="30" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F68B3FE428F0E" box="[113,480,1256,1281]" gridcol="0" gridrow="30" pageId="49" pageNumber="50">Claw 3 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6951FA098F2D" box="[113,1451,1290,1314]" gridrow="31" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6951FE428F2D" box="[113,480,1290,1314]" gridcol="0" gridrow="31" pageId="49" pageNumber="50">External base</th>
|
||||
<td id="76E75DECA608004DAF776951FDE38F2D" box="[553,577,1290,1314]" gridcol="1" gridrow="31" pageId="49" pageNumber="50">11</td>
|
||||
<td id="76E75DECA608004DAFCD6951FD568F2D" box="[659,756,1290,1314]" gridcol="2" gridrow="31" pageId="49" pageNumber="50">3.0–4.3</td>
|
||||
<td id="76E75DECA608004DAE1B6951FC1E8F2D" box="[837,956,1290,1314]" gridcol="3" gridrow="31" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6951FC0D8F2D" box="[849,943,1290,1314]" italics="true" pageId="49" pageNumber="50">13.1–20.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536951FBE78F2D" box="[1037,1093,1290,1314]" gridcol="4" gridrow="31" pageId="49" pageNumber="50">3.6</td>
|
||||
<td id="76E75DECA608004DA9D66951FB1F8F2D" box="[1160,1213,1290,1314]" gridcol="5" gridrow="31" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66951FB138F2D" box="[1160,1201,1290,1314]" italics="true" pageId="49" pageNumber="50">17.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516951FA8D8F2D" box="[1295,1327,1290,1314]" gridcol="6" gridrow="31" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6951FA098F2D" box="[1410,1451,1290,1314]" gridcol="7" gridrow="31" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6951FA3D8F2D" box="[1410,1439,1290,1314]" italics="true" pageId="49" pageNumber="50">2.1</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6977FA098F4B" box="[113,1451,1324,1348]" gridrow="32" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6977FE428F4B" box="[113,480,1324,1348]" gridcol="0" gridrow="32" pageId="49" pageNumber="50">External primary branch</th>
|
||||
<td id="76E75DECA608004DAF776977FDE38F4B" box="[553,577,1324,1348]" gridcol="1" gridrow="32" pageId="49" pageNumber="50">4</td>
|
||||
<td id="76E75DECA608004DAFCD6977FD568F4B" box="[659,756,1324,1348]" gridcol="2" gridrow="32" pageId="49" pageNumber="50">7.0–9.0</td>
|
||||
<td id="76E75DECA608004DAE1B6977FC1E8F4B" box="[837,956,1324,1348]" gridcol="3" gridrow="32" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6977FC128F4B" box="[849,944,1324,1348]" italics="true" pageId="49" pageNumber="50">33.3–33.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536977FBE78F4B" box="[1037,1093,1324,1348]" gridcol="4" gridrow="32" pageId="49" pageNumber="50">8.2</td>
|
||||
<td id="76E75DECA608004DA9D66977FB1F8F4B" box="[1160,1213,1324,1348]" gridcol="5" gridrow="32" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66977FB138F4B" box="[1160,1201,1324,1348]" italics="true" pageId="49" pageNumber="50">33.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516977FA8D8F4B" box="[1295,1327,1324,1348]" gridcol="6" gridrow="32" pageId="49" pageNumber="50">0.9</td>
|
||||
<td id="76E75DECA608004DA8DC6977FA098F4B" box="[1410,1451,1324,1348]" gridcol="7" gridrow="32" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6976FA298F4B" box="[1410,1419,1325,1348]" italics="true" pageId="49" pageNumber="50">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6915FA098F69" box="[113,1451,1358,1382]" gridrow="33" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6915FE428F69" box="[113,480,1358,1382]" gridcol="0" gridrow="33" pageId="49" pageNumber="50">External secondary branch</th>
|
||||
<td id="76E75DECA608004DAF776915FDE38F69" box="[553,577,1358,1382]" gridcol="1" gridrow="33" pageId="49" pageNumber="50">11</td>
|
||||
<td id="76E75DECA608004DAFCD6915FD568F69" box="[659,756,1358,1382]" gridcol="2" gridrow="33" pageId="49" pageNumber="50">5.5–7.2</td>
|
||||
<td id="76E75DECA608004DAE1B6915FC1E8F69" box="[837,956,1358,1382]" gridcol="3" gridrow="33" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6915FC0D8F69" box="[849,943,1358,1382]" italics="true" pageId="49" pageNumber="50">26.2–32.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536915FBE78F69" box="[1037,1093,1358,1382]" gridcol="4" gridrow="33" pageId="49" pageNumber="50">6.6</td>
|
||||
<td id="76E75DECA608004DA9D66915FB1F8F69" box="[1160,1213,1358,1382]" gridcol="5" gridrow="33" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66915FB138F69" box="[1160,1201,1358,1382]" italics="true" pageId="49" pageNumber="50">28.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516915FA8D8F69" box="[1295,1327,1358,1382]" gridcol="6" gridrow="33" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA8DC6915FA098F69" box="[1410,1451,1358,1382]" gridcol="7" gridrow="33" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6915FA3D8F69" box="[1410,1439,1358,1382]" italics="true" pageId="49" pageNumber="50">2.3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F692BFA098F87" box="[113,1451,1392,1416]" gridrow="34" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F692BFE428F87" box="[113,480,1392,1416]" gridcol="0" gridrow="34" pageId="49" pageNumber="50">Internal base</th>
|
||||
<td id="76E75DECA608004DAF77692BFDE38F87" box="[553,577,1392,1416]" gridcol="1" gridrow="34" pageId="49" pageNumber="50">11</td>
|
||||
<td id="76E75DECA608004DAFCD692BFD568F87" box="[659,756,1392,1416]" gridcol="2" gridrow="34" pageId="49" pageNumber="50">3.0–4.0</td>
|
||||
<td id="76E75DECA608004DAE1B692BFC1E8F87" box="[837,956,1392,1416]" gridcol="3" gridrow="34" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F692BFC0D8F87" box="[849,943,1392,1416]" italics="true" pageId="49" pageNumber="50">14.3–18.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA953692BFBE78F87" box="[1037,1093,1392,1416]" gridcol="4" gridrow="34" pageId="49" pageNumber="50">3.5</td>
|
||||
<td id="76E75DECA608004DA9D6692BFB1F8F87" box="[1160,1213,1392,1416]" gridcol="5" gridrow="34" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D6692BFB138F87" box="[1160,1201,1392,1416]" italics="true" pageId="49" pageNumber="50">15.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA851692BFA8D8F87" box="[1295,1327,1392,1416]" gridcol="6" gridrow="34" pageId="49" pageNumber="50">0.4</td>
|
||||
<td id="76E75DECA608004DA8DC692BFA098F87" box="[1410,1451,1392,1416]" gridcol="7" gridrow="34" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC692BFA3D8F87" box="[1410,1439,1392,1416]" italics="true" pageId="49" pageNumber="50">1.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F69CAFA098FA5" box="[113,1451,1425,1450]" gridrow="35" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F69CAFE428FA5" box="[113,480,1425,1450]" gridcol="0" gridrow="35" pageId="49" pageNumber="50">Internal primary branch</th>
|
||||
<td id="76E75DECA608004DAF7769CAFDE38FA5" box="[553,577,1425,1450]" gridcol="1" gridrow="35" pageId="49" pageNumber="50">5</td>
|
||||
<td id="76E75DECA608004DAFCD69CAFD568FA5" box="[659,756,1425,1450]" gridcol="2" gridrow="35" pageId="49" pageNumber="50">6.7–7.3</td>
|
||||
<td id="76E75DECA608004DAE1B69CAFC1E8FA5" box="[837,956,1425,1450]" gridcol="3" gridrow="35" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F69C9FC0D8FA5" box="[849,943,1426,1450]" italics="true" pageId="49" pageNumber="50">29.7–34.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA95369CAFBE78FA5" box="[1037,1093,1425,1450]" gridcol="4" gridrow="35" pageId="49" pageNumber="50">7.0</td>
|
||||
<td id="76E75DECA608004DA9D669CAFB1F8FA5" box="[1160,1213,1425,1450]" gridcol="5" gridrow="35" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D669C9FB138FA5" box="[1160,1201,1426,1450]" italics="true" pageId="49" pageNumber="50">32.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA85169CAFA8D8FA5" box="[1295,1327,1425,1450]" gridcol="6" gridrow="35" pageId="49" pageNumber="50">0.3</td>
|
||||
<td id="76E75DECA608004DA8DC69CAFA098FA5" box="[1410,1451,1425,1450]" gridcol="7" gridrow="35" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC69C9FA3D8FA5" box="[1410,1439,1426,1450]" italics="true" pageId="49" pageNumber="50">3.3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F69E8FA098FC3" box="[113,1451,1459,1484]" gridrow="36" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F69E8FE428FC3" box="[113,480,1459,1484]" gridcol="0" gridrow="36" pageId="49" pageNumber="50">Internal secondary branch</th>
|
||||
<td id="76E75DECA608004DAF7769E8FDE38FC3" box="[553,577,1459,1484]" gridcol="1" gridrow="36" pageId="49" pageNumber="50">5</td>
|
||||
<td id="76E75DECA608004DAFCD69E8FD568FC3" box="[659,756,1459,1484]" gridcol="2" gridrow="36" pageId="49" pageNumber="50">4.4–5.9</td>
|
||||
<td id="76E75DECA608004DAE1B69E8FC1E8FC3" box="[837,956,1459,1484]" gridcol="3" gridrow="36" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F69EFFC128FC3" box="[849,944,1460,1484]" italics="true" pageId="49" pageNumber="50">21.0–25.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA95369E8FBE78FC3" box="[1037,1093,1459,1484]" gridcol="4" gridrow="36" pageId="49" pageNumber="50">5.1</td>
|
||||
<td id="76E75DECA608004DA9D669E8FB1F8FC3" box="[1160,1213,1459,1484]" gridcol="5" gridrow="36" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D669EFFB138FC3" box="[1160,1201,1460,1484]" italics="true" pageId="49" pageNumber="50">23.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA85169E8FA8D8FC3" box="[1295,1327,1459,1484]" gridcol="6" gridrow="36" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA8DC69E8FA098FC3" box="[1410,1451,1459,1484]" gridcol="7" gridrow="36" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC69EFFA3D8FC3" box="[1410,1439,1460,1484]" italics="true" pageId="49" pageNumber="50">1.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F698EFA098FE2" box="[113,1451,1493,1517]" gridrow="37" pageId="49" pageNumber="50" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76E75DECA608004DAD2F698EFE428FE2" box="[113,480,1493,1517]" gridcol="0" gridrow="37" pageId="49" pageNumber="50">Claw 4 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F69ACFA098C00" box="[113,1451,1527,1551]" gridrow="38" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F69ACFE428C00" box="[113,480,1527,1551]" gridcol="0" gridrow="38" pageId="49" pageNumber="50">Anterior base</th>
|
||||
<td id="76E75DECA608004DAF7769ACFDE38C00" box="[553,577,1527,1551]" gridcol="1" gridrow="38" pageId="49" pageNumber="50">8</td>
|
||||
<td id="76E75DECA608004DAFCD69ACFD568C00" box="[659,756,1527,1551]" gridcol="2" gridrow="38" pageId="49" pageNumber="50">2.8–4.4</td>
|
||||
<td id="76E75DECA608004DAE1B69ACFC1E8C00" box="[837,956,1527,1551]" gridcol="3" gridrow="38" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F69ACFC0D8C00" box="[849,943,1527,1551]" italics="true" pageId="49" pageNumber="50">14.3–18.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA95369ACFBE78C00" box="[1037,1093,1527,1551]" gridcol="4" gridrow="38" pageId="49" pageNumber="50">3.6</td>
|
||||
<td id="76E75DECA608004DA9D669ACFB1F8C00" box="[1160,1213,1527,1551]" gridcol="5" gridrow="38" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D669ACFB138C00" box="[1160,1201,1527,1551]" italics="true" pageId="49" pageNumber="50">16.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA85169ACFA8D8C00" box="[1295,1327,1527,1551]" gridcol="6" gridrow="38" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC69ACFA098C00" box="[1410,1451,1527,1551]" gridcol="7" gridrow="38" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC69ACFA3D8C00" box="[1410,1439,1527,1551]" italics="true" pageId="49" pageNumber="50">1.3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6A42FA098C3E" box="[113,1451,1561,1585]" gridrow="39" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6A42FE428C3E" box="[113,480,1561,1585]" gridcol="0" gridrow="39" pageId="49" pageNumber="50">Anterior primary branch</th>
|
||||
<td id="76E75DECA608004DAF776A42FDE38C3E" box="[553,577,1561,1585]" gridcol="1" gridrow="39" pageId="49" pageNumber="50">5</td>
|
||||
<td id="76E75DECA608004DAFCD6A42FD568C3E" box="[659,756,1561,1585]" gridcol="2" gridrow="39" pageId="49" pageNumber="50">6.8–8.8</td>
|
||||
<td id="76E75DECA608004DAE1B6A42FC1E8C3E" box="[837,956,1561,1585]" gridcol="3" gridrow="39" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6A42FC128C3E" box="[849,944,1561,1585]" italics="true" pageId="49" pageNumber="50">30.9–35.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536A42FBE78C3E" box="[1037,1093,1561,1585]" gridcol="4" gridrow="39" pageId="49" pageNumber="50">7.5</td>
|
||||
<td id="76E75DECA608004DA9D66A42FB1F8C3E" box="[1160,1213,1561,1585]" gridcol="5" gridrow="39" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66A42FB138C3E" box="[1160,1201,1561,1585]" italics="true" pageId="49" pageNumber="50">32.6</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516A42FA8D8C3E" box="[1295,1327,1561,1585]" gridcol="6" gridrow="39" pageId="49" pageNumber="50">0.8</td>
|
||||
<td id="76E75DECA608004DA8DC6A42FA098C3E" box="[1410,1451,1561,1585]" gridcol="7" gridrow="39" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6A42FA3D8C3E" box="[1410,1439,1561,1585]" italics="true" pageId="49" pageNumber="50">2.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6A60FA098C5C" box="[113,1451,1595,1619]" gridrow="40" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6A60FE428C5C" box="[113,480,1595,1619]" gridcol="0" gridrow="40" pageId="49" pageNumber="50">Anterior secondary branch</th>
|
||||
<td id="76E75DECA608004DAF776A60FDE38C5C" box="[553,577,1595,1619]" gridcol="1" gridrow="40" pageId="49" pageNumber="50">7</td>
|
||||
<td id="76E75DECA608004DAFCD6A60FD568C5C" box="[659,756,1595,1619]" gridcol="2" gridrow="40" pageId="49" pageNumber="50">4.4–6.3</td>
|
||||
<td id="76E75DECA608004DAE1B6A60FC1E8C5C" box="[837,956,1595,1619]" gridcol="3" gridrow="40" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6A60FC128C5C" box="[849,944,1595,1619]" italics="true" pageId="49" pageNumber="50">19.2–29.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536A60FBE78C5C" box="[1037,1093,1595,1619]" gridcol="4" gridrow="40" pageId="49" pageNumber="50">5.5</td>
|
||||
<td id="76E75DECA608004DA9D66A60FB1F8C5C" box="[1160,1213,1595,1619]" gridcol="5" gridrow="40" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66A60FB138C5C" box="[1160,1201,1595,1619]" italics="true" pageId="49" pageNumber="50">24.6</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516A60FA8D8C5C" box="[1295,1327,1595,1619]" gridcol="6" gridrow="40" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA8DC6A60FA098C5C" box="[1410,1451,1595,1619]" gridcol="7" gridrow="40" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6A60FA3D8C5C" box="[1410,1439,1595,1619]" italics="true" pageId="49" pageNumber="50">3.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6A06FA098C7A" box="[113,1451,1629,1653]" gridrow="41" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6A06FE428C7A" box="[113,480,1629,1653]" gridcol="0" gridrow="41" pageId="49" pageNumber="50">Posterior base</th>
|
||||
<td id="76E75DECA608004DAF776A06FDE38C7A" box="[553,577,1629,1653]" gridcol="1" gridrow="41" pageId="49" pageNumber="50">7</td>
|
||||
<td id="76E75DECA608004DAFCD6A06FD568C7A" box="[659,756,1629,1653]" gridcol="2" gridrow="41" pageId="49" pageNumber="50">3.4–5.0</td>
|
||||
<td id="76E75DECA608004DAE1B6A06FC1E8C7A" box="[837,956,1629,1653]" gridcol="3" gridrow="41" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6A06FC128C7A" box="[849,944,1629,1653]" italics="true" pageId="49" pageNumber="50">16.1–20.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536A06FBE78C7A" box="[1037,1093,1629,1653]" gridcol="4" gridrow="41" pageId="49" pageNumber="50">4.0</td>
|
||||
<td id="76E75DECA608004DA9D66A06FB1F8C7A" box="[1160,1213,1629,1653]" gridcol="5" gridrow="41" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66A06FB138C7A" box="[1160,1201,1629,1653]" italics="true" pageId="49" pageNumber="50">18.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516A06FA8D8C7A" box="[1295,1327,1629,1653]" gridcol="6" gridrow="41" pageId="49" pageNumber="50">0.5</td>
|
||||
<td id="76E75DECA608004DA8DC6A06FA098C7A" box="[1410,1451,1629,1653]" gridcol="7" gridrow="41" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6A06FA3D8C7A" box="[1410,1439,1629,1653]" italics="true" pageId="49" pageNumber="50">1.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6A25FA098C98" box="[113,1451,1662,1687]" gridrow="42" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6A25FE428C98" box="[113,480,1662,1687]" gridcol="0" gridrow="42" pageId="49" pageNumber="50">Posterior primary branch</th>
|
||||
<td id="76E75DECA608004DAF776A25FDE38C98" box="[553,577,1662,1687]" gridcol="1" gridrow="42" pageId="49" pageNumber="50">6</td>
|
||||
<td id="76E75DECA608004DAFCD6A25FD568C98" box="[659,756,1662,1687]" gridcol="2" gridrow="42" pageId="49" pageNumber="50">8.2–10.0</td>
|
||||
<td id="76E75DECA608004DAE1B6A25FC1E8C98" box="[837,956,1662,1687]" gridcol="3" gridrow="42" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6A24FC128C98" box="[849,944,1663,1687]" italics="true" pageId="49" pageNumber="50">37.8–45.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536A25FBE78C98" box="[1037,1093,1662,1687]" gridcol="4" gridrow="42" pageId="49" pageNumber="50">9.1</td>
|
||||
<td id="76E75DECA608004DA9D66A25FB1F8C98" box="[1160,1213,1662,1687]" gridcol="5" gridrow="42" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66A24FB138C98" box="[1160,1201,1663,1687]" italics="true" pageId="49" pageNumber="50">40.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516A25FA8D8C98" box="[1295,1327,1662,1687]" gridcol="6" gridrow="42" pageId="49" pageNumber="50">0.7</td>
|
||||
<td id="76E75DECA608004DA8DC6A25FA098C98" box="[1410,1451,1662,1687]" gridcol="7" gridrow="42" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A8DC6A24FA3D8C98" box="[1410,1439,1663,1687]" italics="true" pageId="49" pageNumber="50">3.0</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A608004DAD2F6AFBFA098CB7" box="[113,1451,1696,1720]" gridrow="43" pageId="49" pageNumber="50">
|
||||
<th id="76E75DECA608004DAD2F6AFBFE428CB7" box="[113,480,1696,1720]" gridcol="0" gridrow="43" pageId="49" pageNumber="50">Posterior secondary branch</th>
|
||||
<td id="76E75DECA608004DAF776AFBFDE38CB7" box="[553,577,1696,1720]" gridcol="1" gridrow="43" pageId="49" pageNumber="50">7</td>
|
||||
<td id="76E75DECA608004DAFCD6AFBFD568CB7" box="[659,756,1696,1720]" gridcol="2" gridrow="43" pageId="49" pageNumber="50">6.3–8.0</td>
|
||||
<td id="76E75DECA608004DAE1B6AFBFC1E8CB7" box="[837,956,1696,1720]" gridcol="3" gridrow="43" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE0F6AFBFC128CB7" box="[849,944,1696,1720]" italics="true" pageId="49" pageNumber="50">27.5–36.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA9536AFBFBE78CB7" box="[1037,1093,1696,1720]" gridcol="4" gridrow="43" pageId="49" pageNumber="50">7.3</td>
|
||||
<td id="76E75DECA608004DA9D66AFBFB1F8CB7" box="[1160,1213,1696,1720]" gridcol="5" gridrow="43" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83A9D66AFBFB138CB7" box="[1160,1201,1696,1720]" italics="true" pageId="49" pageNumber="50">31.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA608004DA8516AFBFA8D8CB7" box="[1295,1327,1696,1720]" gridcol="6" gridrow="43" pageId="49" pageNumber="50">0.6</td>
|
||||
<td id="76E75DECA608004DA8DC6AFBFA098CB7" box="[1410,1451,1696,1720]" gridcol="7" gridrow="43" pageId="49" pageNumber="50">3.7</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A608FF83ADA96B52FDD78D2C" blockId="49.[247,629,1801,1827]" box="[247,629,1801,1827]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83ADA96B52FDD78D2C" bold="true" box="[247,629,1801,1827]" pageId="49" pageNumber="50">Proabably also the following record:</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A608FF83AD286B62FD908D7D" blockId="49.[118,758,1848,1906]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AD286B62FD908D7D" bold="true" pageId="49" pageNumber="50">
|
||||
<taxonomicName id="4C064D51A608FF83AD286B62FF1E8D5C" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[118,188,1848,1875]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="49" pageNumber="50" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A608FF83AD286B62FF1E8D5C" bold="true" box="[118,188,1848,1875]" italics="true" pageId="49" pageNumber="50">D. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
; Pektu-San Mountain,
|
||||
<collectingRegion id="49C2F830A608FF83ACF36B62FD098D5D" box="[429,683,1848,1875]" country="North Korea" name="Yanggang-do" pageId="49" pageNumber="50">Ryanggang-do Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F3117642A608FF83AFEA6B62FEDE8D7D" name="North Korea" pageId="49" pageNumber="50">North Korea</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A608FF83ACDB6B03FD8F8D7D" author="Dastych H" box="[389,557,1880,1906]" pageId="49" pageNumber="50" pagination="125 - 45" refId="ref35500" refString="Dastych H. North Korean Tardigrada. Acta Zoologica Cracoviensia 1974; 19: 125 - 45." type="journal article" year="1974">Dastych (1974)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A608FF81AD2F6BDDFE628DA2" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="materials_examined">
|
||||
<paragraph id="8BB936D2A608FF83AD2F6BDDFE788DB1" blockId="49.[113,762,1926,1982]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AD2F6BDDFE988D91" box="[113,314,1926,1950]" italics="true" pageId="49" pageNumber="50">Material examined:</emphasis>
|
||||
13 individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A608FF83AC2C6BFDFE688DB1" box="[370,458,1958,1982]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="49" pageNumber="50">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A608FF83AE746B51FAA88D6E" blockId="49.[810,1459,1801,1889]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE746B51FBA48D2D" box="[810,1030,1802,1826]" italics="true" pageId="49" pageNumber="50">Comparative material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA608FF83A9546B51FAA48D6E" collectingDate="2016-09-14" collectorName="Barczyk" country="Russia" elevation="525" latitude="53.26889" location="Baikal Lake" longLatPrecision="17" longitude="107.55139" municipality="Olchon" pageId="49" pageNumber="50" specimenCount="1">
|
||||
<collectingCountry id="F3117642A608FF83A9546B51FBED8D2D" box="[1034,1103,1802,1826]" name="Russia" pageId="49" pageNumber="50">Russia</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A608FF83A9096B51FB6C8D2D" LSID="urn:lsid:plazi:treatment:03AF87C4A609FF86AE8B6A86FECE8D4D:8ED96009A608FF83A9096B51FB6C8D2D" box="[1111,1230,1802,1826]" country="Russia" latitude="53.26889" longLatPrecision="17" longitude="107.55139" municipality="Olchon" name="Baikal Lake" pageId="49" pageNumber="50">Baikal Lake</location>
|
||||
,
|
||||
<collectingMunicipality id="6BDDACA8A608FF83A9896B51FA878D2D" box="[1239,1317,1802,1826]" pageId="49" pageNumber="50">Olchon</collectingMunicipality>
|
||||
(
|
||||
<geoCoordinate id="EE325015A608FF83A86C6B51FA0D8D2D" box="[1330,1455,1801,1826]" degrees="53" direction="north" minutes="16" orientation="latitude" pageId="49" pageNumber="50" precision="15" seconds="08" value="53.26889">
|
||||
53°16
|
||||
<emphasis id="B972EAC0A608FF83A82C6B52FADA8D2D" box="[1394,1400,1801,1826]" italics="true" pageId="49" pageNumber="50">ʹ</emphasis>
|
||||
08″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A608FF83AE746B72FC0D8D4E" box="[810,943,1832,1857]" degrees="107" direction="east" minutes="33" orientation="longitude" pageId="49" pageNumber="50" precision="15" seconds="05" value="107.55139">
|
||||
107°33
|
||||
<emphasis id="B972EAC0A608FF83AE2D6B73FCDB8D4E" box="[883,889,1832,1857]" italics="true" pageId="49" pageNumber="50">ʹ</emphasis>
|
||||
05″E
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A608FF83AE9F6B72FBE58D4E" box="[961,1095,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="49" pageNumber="50" unit="m" value="525.0">
|
||||
<quantity id="4CFE9B37A608FF83AE9F6B72FBA98D4E" box="[961,1035,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="49" pageNumber="50" unit="m" value="525.0">525 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), shrubland, moss from soil,
|
||||
<date id="FFB81012A608FF83A8C76B72FC768D6E" pageId="49" pageNumber="50" value="2016-09-14">
|
||||
<collectingDate id="EFFCE9FAA608FF83A8C76B72FC768D6E" pageId="49" pageNumber="50" value="2016-09-14">14 September 2016</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A608FF83AE836B12FB8C8D6E" box="[989,1070,1865,1889]" pageId="49" pageNumber="50">Barczyk</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA608FF83A9356B12FB5D8D6E" box="[1131,1279,1865,1889]" count="1" pageId="49" pageNumber="50" type="generic">one specimen</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A608FF81AE746BD9FEF38C3A" blockId="49.[810,1459,1922,1978]" lastBlockId="51.[112,764,1283,1966]" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
|
||||
<emphasis id="B972EAC0A608FF83AE746BD9FBA78D95" box="[810,1029,1922,1946]" italics="true" pageId="49" pageNumber="50">Amended description:</emphasis>
|
||||
Body small (
|
||||
<tableCitation id="C6840369A608FF83A9FA6BD9FAA68D95" box="[1188,1284,1922,1946]" captionStart="Table 14" captionStartId="52.[129,184,143,167]" captionTargetPageId="52" captionText="Table 14. Measurements (in micrometres) of selected morphological structures of Pilatobius sexbullatus (Ito, 1995) from Mongolia (MN.008) mounted in Hoyer’s medium." pageId="49" pageNumber="50">Table 14</tableCitation>
|
||||
) and corpulent (
|
||||
<figureCitation id="133D2A57A608FF83AE6B6BFAFC2F8DB6" box="[821,909,1953,1977]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="49" pageNumber="50">Fig. 40A</figureCitation>
|
||||
). A single median line of six large, poorly delineated hemispherical gibbosities: three at the level of legs I–III, two between legs I–II and II–III, and one in the segment behind the third leg pair (
|
||||
<figureCitation id="133D2A57A60BFF80AC1C6BC3FE398DBF" box="[322,411,1944,1968]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="50" pageNumber="51">Fig. 40A</figureCitation>
|
||||
). Cuticle greatly sculptured on the entire dorsal surface by sub-round polygons of variable size: the largest present on the gibbosities and in the cephalic region, and the smallest on the lateral sides of the trunk (
|
||||
<figureCitation id="133D2A57A60BFF80A8606BC3FA138DA0" box="[1342,1457,1944,1968]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="50" pageNumber="51">Fig. 40A, B</figureCitation>
|
||||
). Cribriform areas not visible under PCM. Legs short, plump, and barely delimited from the trunk (
|
||||
<figureCitation id="133D2A57A60AFF81ACAC6979FDEB8F34" box="[498,585,1314,1339]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="51" pageNumber="52">Fig. 40A</figureCitation>
|
||||
). Eyes present in living animals, but usually quickly dissolving in Hoyer’s medium. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A60AFF81AC85693AFD9E8F76" box="[475,572,1377,1401]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60AFF81AC85693AFD9E8F76" box="[475,572,1377,1401]" italics="true" pageId="51" pageNumber="52">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A60AFF81AF1C6939FDD38F75" box="[578,625,1378,1402]" pageId="51" pageNumber="52">type</typeStatus>
|
||||
, with a short pharyngeal tube (
|
||||
<figureCitation id="133D2A57A60AFF81AC7769DBFE208F96" box="[297,386,1408,1433]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="51" pageNumber="52">Fig. 40C</figureCitation>
|
||||
). The OCA not visible under PCM. Furcae of the
|
||||
<taxonomicName id="4C064D51A60AFF81ADA369FBFEFF8FB7" authorityName="Ehrenberg" authorityYear="1848" box="[253,349,1440,1464]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60AFF81ADA369FBFEFF8FB7" box="[253,349,1440,1464]" italics="true" pageId="51" pageNumber="52">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A60AFF81AC3C69FBFE308FB7" box="[354,402,1440,1464]" pageId="51" pageNumber="52">type</typeStatus>
|
||||
. The DABT large; pharynx circular, with large pharyngeal apophyses (
|
||||
<figureCitation id="133D2A57A60AFF81ACB069E4FDEC8FD8" box="[494,590,1471,1495]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="51" pageNumber="52">Fig. 40C</figureCitation>
|
||||
). Macroplacoid length sequence 2 <1, the first with delicate constriction at twofifths of its length. Septulum large, but clearly shorter than the second macroplacoid.
|
||||
</paragraph>
|
||||
<caption id="DF79665AA60BFF80ADDF6A82FF5D8D26" ID-DOI="http://doi.org/10.5281/zenodo.14243830" ID-Zenodo-Dep="14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="50" pageNumber="51" startId="50.[129,194,1753,1777]" targetBox="[129,1473,144,1725]" targetPageId="50" targetType="figure">
|
||||
<paragraph id="8BB936D2A60BFF80ADDF6A82FF5D8D26" blockId="50.[129,1444,1753,1833]" pageId="50" pageNumber="51">
|
||||
<emphasis id="B972EAC0A60BFF80ADDF6A82FF478CFE" bold="true" box="[129,229,1753,1777]" pageId="50" pageNumber="51">Figure 38.</emphasis>
|
||||
General morphology of
|
||||
<taxonomicName id="4C064D51A60BFF80AC936A82FCB88CFE" authority="(Bartos, 1935)" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[461,794,1753,1777]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="50" pageNumber="51" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A60BFF80AC936A82FD858CFE" box="[461,551,1753,1777]" italics="true" pageId="50" pageNumber="51">Pilatobius</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A60BFF80AF196A81FD288CFE" box="[583,650,1754,1777]" italics="true" pageId="50" pageNumber="51">rugosus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60BFF80AFC76A82FCB28CFE" author="Bartos E" box="[665,784,1753,1777]" pageId="50" pageNumber="51" pagination="257 - 60" refId="ref34712" refString="Bartos E. Vier neue Hypsibius - Arten aus der Tschechoslovakei. Zoologischer Anzeiger 1935; 110: 257 - 60." type="journal article" year="1935">Bartoš, 1935</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(all but C, PCM): A, specimen
|
||||
<emphasis id="B972EAC0A60BFF80A9196A81FB228CFE" box="[1095,1152,1754,1777]" italics="true" pageId="50" pageNumber="51">in toto</emphasis>
|
||||
(ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A60BFF80A9946AADFAA68D02" box="[1226,1284,1782,1805]" italics="true" pageId="50" pageNumber="51">in toto</emphasis>
|
||||
. Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA60AFF81AD2F68C9FA078EC9" ID-DOI="http://doi.org/10.5281/zenodo.14243832" ID-Zenodo-Dep="14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="51" pageNumber="52" startId="51.[113,176,1170,1194]" targetBox="[114,1458,144,1142]" targetPageId="51" targetType="figure">
|
||||
<paragraph id="8BB936D2A60AFF81AD2F68C9FA078EC9" blockId="51.[113,1445,1170,1222]" pageId="51" pageNumber="52">
|
||||
<emphasis id="B972EAC0A60AFF81AD2F68C9FF768EA5" bold="true" box="[113,212,1170,1194]" pageId="51" pageNumber="52">Figure 39.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A60AFF81AC7568C9FE418EA5" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[299,483,1170,1194]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A60AFF81AC7568C9FE218EA5" box="[299,387,1170,1194]" italics="true" pageId="51" pageNumber="52">Pilatobius</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A60AFF81ACFF68C8FE418EA5" box="[417,483,1171,1194]" italics="true" pageId="51" pageNumber="52">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars, and white arrowhead indicates anterior bar. Scale bars are in micrometres, identical in panels A, B and in C, D.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A60AFF81ADD36A67FE628DA2" blockId="51.[112,764,1283,1966]" pageId="51" pageNumber="52">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A60AFF81AC786A67FE298C5B" authorityName="Ehrenberg" authorityYear="1848" box="[294,395,1596,1620]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60AFF81AC786A67FE298C5B" box="[294,395,1596,1620]" italics="true" pageId="51" pageNumber="52">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A60AFF81ACC86A66FE658C5A" box="[406,455,1597,1621]" pageId="51" pageNumber="52">type</typeStatus>
|
||||
, with very short basal portions; those of the external claws also slightly broadened (
|
||||
<figureCitation id="133D2A57A60AFF81AD236A20FF7B8C9C" box="[125,217,1659,1683]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41A</figureCitation>
|
||||
). The most remarkable are the bases of the posterior claws, which are extremely short, with margins evidently pointed upwards (
|
||||
<figureCitation id="133D2A57A60AFF81ACDA6AE1FE7F8CDD" box="[388,477,1722,1746]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41B</figureCitation>
|
||||
). Primary branches of the internal and anterior claws much longer than the secondary branches (
|
||||
<figureCitation id="133D2A57A60AFF81ADB86AA3FE938D1E" box="[230,305,1784,1809]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41</figureCitation>
|
||||
). Both branches of the external and posterior claws elongated. Accessory points barely divergent (
|
||||
<figureCitation id="133D2A57A60AFF81AD236B6CFF648D40" box="[125,198,1847,1871]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41</figureCitation>
|
||||
). Pseudolunulae absent.
|
||||
<specimenCount id="9D00FD5BA60AFF81AC8B6B63FDE78D5F" box="[469,581,1848,1872]" count="2" pageId="51" pageNumber="52" type="generic" typeStatus="types">Two types</specimenCount>
|
||||
of cuticular bars present: (i) internal, long thickenings at the bases of claws IIII (
|
||||
<figureCitation id="133D2A57A60AFF81ADFE6B2DFF598D81" box="[160,251,1910,1934]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41A</figureCitation>
|
||||
); and (ii) posterior, short rods (
|
||||
<figureCitation id="133D2A57A60AFF81AF3F6B2DFD1E8D81" box="[609,700,1910,1934]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="51" pageNumber="52">Fig. 40A</figureCitation>
|
||||
); the latter may be absent (
|
||||
<figureCitation id="133D2A57A60AFF81AC096BCEFE128DA2" box="[343,432,1941,1965]" captionStart="Figure 41" captionStartId="53.[113,178,1751,1775]" captionTargetBox="[114,1458,1225,1722]" captionTargetId="figure-135@53.[114,1458,1225,1722]" captionTargetPageId="53" captionText="Figure 41. Claws of Pilatobius sexbullatus (PCM): A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="51" pageNumber="52">Fig. 41B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A60AFF86AE746958FECE8D4D" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="discussion">
|
||||
<paragraph id="8BB936D2A60AFF86AE746958FECE8D4D" blockId="51.[808,1460,1283,1965]" lastBlockId="52.[129,777,1803,1858]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
|
||||
<emphasis id="B972EAC0A60AFF81AE746958FC2A8F14" box="[810,904,1283,1307]" italics="true" pageId="51" pageNumber="52">Remarks:</emphasis>
|
||||
Specimens depicted by
|
||||
<bibRefCitation id="EF974B23A60AFF81A9276958FABE8F14" author="Dastych H" box="[1145,1308,1283,1307]" pageId="51" pageNumber="52" pagination="125 - 45" refId="ref35500" refString="Dastych H. North Korean Tardigrada. Acta Zoologica Cracoviensia 1974; 19: 125 - 45." type="journal article" year="1974">Dastych (1974)</bibRefCitation>
|
||||
fit the description of
|
||||
<taxonomicName id="4C064D51A60AFF81AE256978FBA38F35" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[891,1025,1314,1338]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60AFF81AE256978FBA38F35" box="[891,1025,1314,1338]" italics="true" pageId="51" pageNumber="52">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
. If their conspecificity is confirmed with DNA barcodes, all published records of this species (from
|
||||
<collectingCountry id="F3117642A60AFF81A8316919FA0D8F56" box="[1391,1455,1346,1369]" name="South Korea" pageId="51" pageNumber="52">Korea</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F3117642A60AFF81AE77693AFC2C8F76" box="[809,910,1377,1401]" name="Mongolia" pageId="51" pageNumber="52">Mongolia</collectingCountry>
|
||||
, and
|
||||
<collectingCountry id="F3117642A60AFF81AEE0693AFBA18F76" box="[958,1027,1377,1401]" name="Russia" pageId="51" pageNumber="52">Russia</collectingCountry>
|
||||
, apart from the
|
||||
<emphasis id="B972EAC0A60AFF81A9C4693AFAB38F76" box="[1178,1297,1377,1401]" italics="true" pageId="51" pageNumber="52">locus typicus</emphasis>
|
||||
in
|
||||
<collectingCountry id="F3117642A60AFF81A870693AFACB8F76" box="[1326,1385,1377,1401]" name="Japan" pageId="51" pageNumber="52">Japan</collectingCountry>
|
||||
) would strongly indicate an Eastern Palaearctic species. However, it must be noted that Dastych examined the
|
||||
<typeStatus id="54BD8870A60AFF81A9FE69FBFB758FB7" box="[1184,1239,1440,1464]" pageId="51" pageNumber="52">types</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C064D51A60AFF81A9A669FAFA958FB7" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[1272,1335,1440,1464]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A60AFF81A9A669FAFA958FB7" box="[1272,1335,1440,1464]" italics="true" pageId="51" pageNumber="52">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
, confirming the similarity of the Korean and American individuals [importantly, an unidentified
|
||||
<taxonomicName id="4C064D51A60AFF81A9596985FB148FF9" box="[1031,1206,1502,1526]" pageId="51" pageNumber="52">
|
||||
<emphasis id="B972EAC0A60AFF81A9596985FBCA8FF9" box="[1031,1128,1502,1526]" italics="true" pageId="51" pageNumber="52">Pilatobius</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A60AFF81A9996985FA8D8FF9" authorityName="Plate" authorityYear="1888" box="[1223,1327,1502,1526]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60AFF81A9996985FA8D8FF9" box="[1223,1327,1502,1526]" italics="true" pageId="51" pageNumber="52">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
at that time) was recorded from the Polish Tatras by
|
||||
<bibRefCitation id="EF974B23A60AFF81A98869A5FADC8C19" author="Dastych H" box="[1238,1406,1534,1558]" pageId="51" pageNumber="52" pagination="1 - 232" refId="ref35518" refString="Dastych H. Niesporczaki (Tardigrada) Tatrzanskiego Parku Narodowego. Monografie Fauny Polski 1980; 9: 1 - 232." type="journal article" year="1980">Dastych (1980)</bibRefCitation>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A60AFF81AE746A45FCCB8C3A" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[810,873,1565,1589]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A60AFF81AE746A45FCCB8C3A" box="[810,873,1565,1589]" italics="true" pageId="51" pageNumber="52">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
was indicated as the taxon morphologically closest to that single specimen; the detailed drawing of that specimen shows a strong similarity to
|
||||
<taxonomicName id="4C064D51A60AFF81AEAA6A06FBD48C7B" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[1012,1142,1628,1652]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60AFF81AEAA6A06FBD48C7B" box="[1012,1142,1628,1652]" italics="true" pageId="51" pageNumber="52">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
]. The dorsal cuticular thickenings of
|
||||
<taxonomicName id="4C064D51A60AFF81AE266A27FC1A8C9C" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[888,952,1659,1683]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A60AFF81AE266A27FC1A8C9C" box="[888,952,1659,1683]" italics="true" pageId="51" pageNumber="52">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
are likely to be an extremely flattened row of the dorsolateral gibbosities typically found in
|
||||
<taxonomicName id="4C064D51A60AFF81A9BD6AC0FACA8CBD" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[1251,1384,1690,1714]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60AFF81A9BD6AC0FACA8CBD" box="[1251,1384,1690,1714]" italics="true" pageId="51" pageNumber="52">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
(slides made by Schuster and Grigarick usually contain strongly compressed tardigrades, which can cause artefacts; P. Gąsiorek, pers. obs.). Therefore, a putative synonymy of
|
||||
<taxonomicName id="4C064D51A60AFF81A99A6AA2FAA08D1F" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[1220,1282,1784,1808]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A60AFF81A99A6AA2FAA08D1F" box="[1220,1282,1784,1808]" italics="true" pageId="51" pageNumber="52">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A60AFF81A86F6AA2FA108D1F" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[1329,1458,1784,1808]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60AFF81A86F6AA2FA108D1F" box="[1329,1458,1784,1808]" italics="true" pageId="51" pageNumber="52">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
(with a potential younger synonym being
|
||||
<taxonomicName id="4C064D51A60AFF81A98D6B42FAF78D3F" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[1235,1365,1816,1840]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60AFF81A98D6B42FAF78D3F" box="[1235,1365,1816,1840]" italics="true" pageId="51" pageNumber="52">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
owing to an inadequate description of
|
||||
<taxonomicName id="4C064D51A60AFF81A90B6B63FB368D40" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[1109,1172,1847,1871]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A60AFF81A90B6B63FB368D40" box="[1109,1172,1847,1871]" italics="true" pageId="51" pageNumber="52">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
) should be considered. The zoogeographical implication of such action would be a single Holarctic species. All members of the
|
||||
<taxonomicName id="4C064D51A60AFF81A9F46B2DFB5A8D81" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[1194,1272,1910,1934]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="51" pageNumber="52" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A60AFF81A9F46B2DFB5A8D81" box="[1194,1272,1910,1934]" italics="true" pageId="51" pageNumber="52">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group have paired dorsal gibbosities, thus they are easily distinguishable from
|
||||
<taxonomicName id="4C064D51A60DFF86ADDF6B57FEA78D2C" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[129,261,1803,1827]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60DFF86ADDF6B57FEA78D2C" box="[129,261,1803,1827]" italics="true" pageId="52" pageNumber="53">P. sexbullatus</emphasis>
|
||||
</taxonomicName>
|
||||
with its autapomorphic condition of a single median line of gibbosities.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
446
data/03/AF/87/03AF87C4A60DFF8AAC626B39FA3C8AE9.xml
Normal file
446
data/03/AF/87/03AF87C4A60DFF8AAC626B39FA3C8AE9.xml
Normal file
|
|
@ -0,0 +1,446 @@
|
|||
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<mods:titleInfo id="10F46DC513218794730E636EFB4B77C6">
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<mods:title id="CAF3DFEAA128A43C3D6D612EDAA8A080">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="243E2EFB799890940CB69F9E69E97FFE">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="5A76CC50D243E4D1C70792539A57F253">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
<mods:nameIdentifier id="AA2322A5398A21D0BC51A6D342F80D90" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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</mods:name>
|
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</mods:role>
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<mods:namePart id="721F6BF456FD6A32207AA40AFB5D711C">Blagden, Brian</mods:namePart>
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<mods:affiliation id="813FEB0062B38227583C382E52BC3742">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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</mods:name>
|
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<mods:namePart id="28559A97CF68C4786FA07DBE0C34013A">Morek, Witold</mods:namePart>
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<mods:affiliation id="4D52141DCA8A2AD537D1979847EB8D65">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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</mods:role>
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<mods:namePart id="AA4E34D805A6EF5DED762E40507DE700">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="8ED3C94DAECD95285775885F6C8BBACC">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:title id="5D27C5BAF6CB6A08882510632FCD763F">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="7710EEE52692501FC1B9D8C8FF40708A">
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<mods:date id="6F639BC4596A534D9E9541FC6452180A">2024</mods:date>
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<mods:number id="36DDB1709B5BCF9D7509D46C7876D166">2023-11-14</mods:number>
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<mods:detail id="AC8EEF3D9CDCCAEFB2FA468F441520C8" type="volume">
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<mods:number id="08CF630733D5015B5769B195606BADB7">202</mods:number>
|
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<mods:detail id="DAD34E193F3499BE74868B52A2851CF7" type="issue">
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<mods:number id="294A27247AE7C1DB8137C9947E98BE62">1</mods:number>
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<mods:url id="40E35F180E32441DEA75CF4AC4580BF9">http://dx.doi.org/10.1093/zoolinnean/zlad151</mods:url>
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<mods:classification id="6EA18F522BB74D58C7F34F8E5D3D4277">journal article</mods:classification>
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<mods:identifier id="6A036F8D716D5924BC9B6108B0CEB1AE" type="DOI">10.1093/zoolinnean/zlad151</mods:identifier>
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<mods:identifier id="CF483B9B99F3A800A26B74F18AE8391D" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="79B9B43BDCE5F0F202C09472ABBD3918" type="Zenodo-Dep">14243740</mods:identifier>
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<mods:identifier id="2D3B46B4CD0D39B26265FFE73FD42FAE" type="ZooBank">044A402-2A0F-4135-9410-7DE081CB11C4Corresponding</mods:identifier>
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<treatment id="03AF87C4A60DFF8AAC626B39FA3C8AE9" LSID="urn:lsid:plazi:treatment:03AF87C4A60DFF8AAC626B39FA3C8AE9" httpUri="http://treatment.plazi.org/id/03AF87C4A60DFF8AAC626B39FA3C8AE9" lastPageId="56" lastPageNumber="57" pageId="52" pageNumber="53">
|
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<subSubSection id="C31C6559A60DFF86AC626B39FDED8D73" box="[316,591,1889,1916]" pageId="52" pageNumber="53" type="nomenclature">
|
||||
<paragraph id="8BB936D2A60DFF86AC626B39FDED8D73" blockId="52.[129,777,1889,1984]" box="[316,591,1889,1916]" pageId="52" pageNumber="53">
|
||||
<emphasis id="B972EAC0A60DFF86AC626B39FDED8D73" bold="true" box="[316,591,1889,1916]" pageId="52" pageNumber="53">
|
||||
Composition of
|
||||
<taxonomicName id="4C064D51A60DFF86ACB76B3AFDED8D73" box="[489,591,1889,1916]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60DFF86ACB76B3AFDED8D73" bold="true" box="[489,591,1889,1916]" italics="true" pageId="52" pageNumber="53">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A60DFF8AADDF6BD3FA3C8AE9" lastPageId="56" lastPageNumber="57" pageId="52" pageNumber="53" type="description">
|
||||
<paragraph id="8BB936D2A60DFF87ADDF6BD3FD588DBE" blockId="52.[129,777,1889,1984]" lastBlockId="53.[113,762,1851,1969]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
|
||||
The genus
|
||||
<taxonomicName id="4C064D51A60DFF86ADA46BD3FEF98DAF" box="[250,347,1928,1952]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60DFF86ADA46BD3FEF98DAF" box="[250,347,1928,1952]" italics="true" pageId="52" pageNumber="53">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
comprises species exclusively with two macroplacoids, arranged in parentheses, and a septulum in the pharynx. With the exception of the
|
||||
<taxonomicName id="4C064D51A60DFF86A9B96B57FAF88D2C" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1255,1370,1804,1827]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A60DFF86A9B96B57FAF88D2C" box="[1255,1370,1804,1827]" italics="true" pageId="52" pageNumber="53">P. recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
complex,
|
||||
<taxonomicName id="4C064D51A60DFF86AE676B71FC388D4D" box="[825,922,1834,1858]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60DFF86AE676B71FC388D4D" box="[825,922,1834,1858]" italics="true" pageId="52" pageNumber="53">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
members typically exhibit varying degrees of dorsal cuticular sculpturing (
|
||||
<figureCitation id="133D2A57A60DFF86A97E6B11FBDF8D6D" box="[1056,1149,1866,1890]" captionStart="Figure 33" captionStartId="43.[113,178,1794,1818]" captionTargetBox="[114,1458,144,1766]" captionTargetId="figure-68@43.[114,1458,144,1766]" captionTargetPageId="43" captionText="Figure 33. General morphology of Pilatobius bullatus (Murray, 1905) in Murray (1905a) (PCM, topotypes): A, dorsal view; B, ventral view; C, close up of dorsal gibbosities and sculpturing.Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243820" httpUri="https://zenodo.org/record/14243820/files/figure.png" pageId="52" pageNumber="53">Figs 33C</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A60DFF86A9D76B11FB148D6D" box="[1161,1206,1866,1890]" captionStart="Figure 35" captionStartId="46.[129,194,1337,1361]" captionTargetBox="[129,1473,144,1309]" captionTargetId="figure-279@46.[129,1473,144,1309]" captionTargetPageId="46" captionText="Figure 35. General morphology of Pilatobius oculatus (Murray, 1906) in Murray (1906b) (PCM): A, neotype in ventral view; B, cuticular wrinkling at the level of third pair of legs (III), with muscle attachments visible as large white ovals, note marginal sculpturing; C, caudal sculpturing, note smaller muscle attachments.Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243824" httpUri="https://zenodo.org/record/14243824/files/figure.png" pageId="52" pageNumber="53">35C</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A60DFF86A99C6B11FB498D6D" box="[1218,1259,1866,1890]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="52" pageNumber="53">38B</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A60DFF86A9A66B11FA838D6D" box="[1272,1313,1866,1890]" captionStart="Figure 40" captionStartId="53.[114,179,1145,1169]" captionTargetBox="[114,1458,144,1117]" captionTargetId="figure-95@53.[114,1458,144,1117]" captionTargetPageId="53" captionText="Figure 40. General morphology of Pilatobius sexbullatus (Ito, 1995) (PCM): A, specimen in toto (dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="52" pageNumber="53">40B</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A60DFF86A8706B11FA318D6D" box="[1326,1427,1866,1890]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="52" pageNumber="53">42D, F, G</figureCitation>
|
||||
; see below). The punctiform microplacoid mentioned in the original description of
|
||||
<taxonomicName id="4C064D51A60DFF86AE8C6BD2FBD68DAF" authorityName="CNHM" baseAuthorityName="Maucci" baseAuthorityYear="1987" box="[978,1140,1928,1952]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="52" pageNumber="53" phylum="Tardigrada" rank="species" species="opisthoglyptus">
|
||||
<emphasis id="B972EAC0A60DFF86AE8C6BD2FBD68DAF" box="[978,1140,1928,1952]" italics="true" pageId="52" pageNumber="53">P. opisthoglyptus</emphasis>
|
||||
</taxonomicName>
|
||||
is most probably a mistake, because the interval between the second macroplacoid and the septulum seems too small to leave space for additional structures (
|
||||
<figureCitation id="133D2A57A60CFF87AD226B01FF718D7D" box="[124,211,1882,1906]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="53" pageNumber="54">Fig. 42A</figureCitation>
|
||||
). The claws of this species are stumpy, with short primary branches; those of the internal claws with a clear hump, and accessory points held tightly adjacent to the claw (
|
||||
<figureCitation id="133D2A57A60CFF87AFC96BC2FD4B8DBE" box="[663,745,1945,1969]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="53" pageNumber="54">Fig. 42B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF79665AA60CFF87AD2C6822FB298EA2" ID-DOI="http://doi.org/10.5281/zenodo.14243834" ID-Zenodo-Dep="14243834" httpUri="https://zenodo.org/record/14243834/files/figure.png" pageId="53" pageNumber="54" startId="53.[114,179,1145,1169]" targetBox="[114,1458,144,1117]" targetPageId="53" targetType="figure">
|
||||
<paragraph id="8BB936D2A60CFF87AD2C6822FB298EA2" blockId="53.[113,1439,1145,1197]" pageId="53" pageNumber="54">
|
||||
<emphasis id="B972EAC0A60CFF87AD2C6822FF748E9E" bold="true" box="[114,214,1145,1169]" pageId="53" pageNumber="54">Figure 40.</emphasis>
|
||||
General morphology of
|
||||
<taxonomicName id="4C064D51A60CFF87ACE06822FC9C8E9E" authority="(Ito, 1995) (PCM)" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[446,830,1145,1169]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60CFF87ACE06822FD238E9E" box="[446,641,1145,1169]" italics="true" pageId="53" pageNumber="54">Pilatobius sexbullatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60CFF87AFD16822FD418E9E" author="Ito M" box="[655,739,1145,1169]" pageId="53" pageNumber="54" pagination="18 - 39" refId="ref36771" refString="Ito M. Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan, II. Family Hypsibiidae. Proceedings of the Japanese Society of Systematic Zoology 1995; 53: 18 - 39." type="journal article" year="1995">Ito, 1995</bibRefCitation>
|
||||
) (PCM)
|
||||
</taxonomicName>
|
||||
: A, specimen
|
||||
<emphasis id="B972EAC0A60CFF87AE9E6821FC5B8E9E" box="[960,1017,1146,1169]" italics="true" pageId="53" pageNumber="54">in toto</emphasis>
|
||||
(dorsal view); B, cuticular sculpturing of the centromedial region with three gibbosities; C, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A60CFF87AE7168CDFCCA8EA2" box="[815,872,1174,1197]" italics="true" pageId="53" pageNumber="54">in toto</emphasis>
|
||||
. Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA60CFF87AD2F6A8CFDAB8D04" ID-DOI="http://doi.org/10.5281/zenodo.14243836" ID-Zenodo-Dep="14243836" httpUri="https://zenodo.org/record/14243836/files/figure.png" pageId="53" pageNumber="54" startId="53.[113,178,1751,1775]" targetBox="[114,1458,1225,1722]" targetPageId="53" targetType="figure">
|
||||
<paragraph id="8BB936D2A60CFF87AD2F6A8CFDAB8D04" blockId="53.[113,1373,1751,1803]" pageId="53" pageNumber="54">
|
||||
<emphasis id="B972EAC0A60CFF87AD2F6A8CFF748CE0" bold="true" box="[113,214,1751,1775]" pageId="53" pageNumber="54">Figure 41.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A60CFF87AC6E6A8CFDE18CE0" authority="(PCM)" baseAuthorityName="PCM" box="[304,579,1751,1775]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A60CFF87AC6E6A8CFE518CE0" box="[304,499,1751,1775]" italics="true" pageId="53" pageNumber="54">Pilatobius sexbullatus</emphasis>
|
||||
(PCM)
|
||||
</taxonomicName>
|
||||
: A, claws II (black arrowhead indicates internal bars); B, claws IV (empty arrowhead indicates pseudolunule). Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A60CFF85AE1B6B60FCC68D68" blockId="53.[810,1459,1851,1969]" lastBlockId="55.[810,1460,1777,1958]" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
|
||||
Two general morphotypes of the pharyngeal structures can be distinguished within
|
||||
<taxonomicName id="4C064D51A60CFF87A93C6B01FB618D7D" box="[1122,1219,1882,1906]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60CFF87A93C6B01FB618D7D" box="[1122,1219,1882,1906]" italics="true" pageId="53" pageNumber="54">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
: (i) short and robust macroplacoids, partly overlapping with the septulum (
|
||||
<figureCitation id="133D2A57A60CFF87A8086B22FC9A8DBF" captionStart="Figure 43" captionStartId="55.[113,178,1671,1695]" captionTargetBox="[114,1458,144,1643]" captionTargetId="figure-125@55.[114,1458,144,1643]" captionTargetPageId="55" captionText="Figure 43. Various Pilatobiinae, morphology of pharyngeal structures (SEM): A, P.oculatus; B, P.bullatus; C, P.ramazzottii; D, P.recamieri (Richters, 1911); E, P.patanei (Binda & Pilato, 1971), lateral view; F, P.patanei, dorsal view. Scale bars: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14243840" httpUri="https://zenodo.org/record/14243840/files/figure.png" pageId="53" pageNumber="54">Fig. 43A, B</figureCitation>
|
||||
); and (ii) long and bar-like macroplacoids, followed by a clear gap between the septulum (
|
||||
<figureCitation id="133D2A57A60EFF85ACD36AAAFDA28D06" box="[397,512,1777,1801]" captionStart="Figure 43" captionStartId="55.[113,178,1671,1695]" captionTargetBox="[114,1458,144,1643]" captionTargetId="figure-125@55.[114,1458,144,1643]" captionTargetPageId="55" captionText="Figure 43. Various Pilatobiinae, morphology of pharyngeal structures (SEM): A, P.oculatus; B, P.bullatus; C, P.ramazzottii; D, P.recamieri (Richters, 1911); E, P.patanei (Binda & Pilato, 1971), lateral view; F, P.patanei, dorsal view. Scale bars: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14243840" httpUri="https://zenodo.org/record/14243840/files/figure.png" pageId="55" pageNumber="56">Fig. 43C–F</figureCitation>
|
||||
). The first macroplacoid is typically subdivided at 40–50% of its length by a deep constriction. The second has the constriction positioned (sub)terminally (
|
||||
<figureCitation id="133D2A57A60EFF85AD226B14FEAE8D68" box="[124,268,1871,1895]" captionStart="Figure 43" captionStartId="55.[113,178,1671,1695]" captionTargetBox="[114,1458,144,1643]" captionTargetId="figure-125@55.[114,1458,144,1643]" captionTargetPageId="55" captionText="Figure 43. Various Pilatobiinae, morphology of pharyngeal structures (SEM): A, P.oculatus; B, P.bullatus; C, P.ramazzottii; D, P.recamieri (Richters, 1911); E, P.patanei (Binda & Pilato, 1971), lateral view; F, P.patanei, dorsal view. Scale bars: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14243840" httpUri="https://zenodo.org/record/14243840/files/figure.png" pageId="55" pageNumber="56">Fig. 43A–D, F</figureCitation>
|
||||
). Only rarely are the constrictions poorly developed (
|
||||
<figureCitation id="133D2A57A60EFF85ADE86B35FEA98D88" box="[182,267,1902,1927]" captionStart="Figure 43" captionStartId="55.[113,178,1671,1695]" captionTargetBox="[114,1458,144,1643]" captionTargetId="figure-125@55.[114,1458,144,1643]" captionTargetPageId="55" captionText="Figure 43. Various Pilatobiinae, morphology of pharyngeal structures (SEM): A, P.oculatus; B, P.bullatus; C, P.ramazzottii; D, P.recamieri (Richters, 1911); E, P.patanei (Binda & Pilato, 1971), lateral view; F, P.patanei, dorsal view. Scale bars: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14243840" httpUri="https://zenodo.org/record/14243840/files/figure.png" pageId="55" pageNumber="56">Fig. 43E</figureCitation>
|
||||
). The morphometry of pharyngeal structures is, together with the existence of the cuticular sculpture, of crucial importance to the taxonomy of
|
||||
<taxonomicName id="4C064D51A60EFF85A9236AAAFB7C8D06" box="[1149,1246,1777,1801]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60EFF85A9236AAAFB7C8D06" box="[1149,1246,1777,1801]" italics="true" pageId="55" pageNumber="56">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
, thus redescriptions of ‘old’ taxa and descriptions of new species should provide extensive measurements of these structures and their relative positions.
|
||||
</paragraph>
|
||||
<caption id="DF79665AA60FFF84ADDF6B68FE898DB4" ID-DOI="http://doi.org/10.5281/zenodo.14243838" ID-Zenodo-Dep="14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="54" pageNumber="55" startId="54.[129,194,1843,1867]" targetBox="[254,1348,144,1815]" targetPageId="54" targetType="figure">
|
||||
<paragraph id="8BB936D2A60FFF84ADDF6B68FE898DB4" blockId="54.[129,1440,1843,1979]" pageId="54" pageNumber="55">
|
||||
<emphasis id="B972EAC0A60FFF84ADDF6B68FF478D44" bold="true" box="[129,229,1843,1867]" pageId="54" pageNumber="55">Figure 42.</emphasis>
|
||||
Various
|
||||
<taxonomicName id="4C064D51A60FFF84AC686B68FE0A8D44" box="[310,424,1843,1867]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
, morphological details (PCM): A,
|
||||
<taxonomicName id="4C064D51A60FFF84AFAC6B6FFB828D44" authority="(Maucci, 1987)" authorityName="CNHM" baseAuthorityName="Maucci" baseAuthorityYear="1987" box="[754,1056,1843,1867]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="opisthoglyptus">
|
||||
<emphasis id="B972EAC0A60FFF84AFAC6B6FFC248D44" box="[754,902,1843,1867]" italics="true" pageId="54" pageNumber="55">P. opisthoglyptus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60FFF84AECB6B68FBB78D44" author="Maucci W & North American Tardigrada & Yellowstone National Park & Bertolani R & Biology of Tardigrades & Selected Symposia & Monographs U" box="[917,1045,1843,1867]" pageId="54" pageNumber="55" pagination="187 - 210" refId="ref37741" refString="Maucci W. A contribution to the knowledge of the North American Tardigrada with emphasis on the fauna of Yellowstone National Park (Wyoming). In: Bertolani R (ed.), Biology of Tardigrades. Selected Symposia and Monographs U. Z. I., 1987; 1: 187 - 210." type="journal article" year="1987">Maucci, 1987</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, paratype, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A60FFF84A8D06B6FFF068D68" italics="true" pageId="54" pageNumber="55">in toto</emphasis>
|
||||
(empty white arrowhead indicates septulum); B,
|
||||
<taxonomicName id="4C064D51A60FFF84AF2A6B0BFCAA8D68" authorityName="CNHM" baseAuthorityName="Maucci" baseAuthorityYear="1987" box="[628,776,1871,1895]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="opisthoglyptus">
|
||||
<emphasis id="B972EAC0A60FFF84AF2A6B0BFCAA8D68" box="[628,776,1871,1895]" italics="true" pageId="54" pageNumber="55">P. opisthoglyptus</emphasis>
|
||||
</taxonomicName>
|
||||
, paratype, claws I; C,
|
||||
<taxonomicName id="4C064D51A60FFF84AE8B6B0BFAA28D68" authority="(Ramazzotti, 1957)" authorityName="CNHM" baseAuthorityName="Ramazzotti" baseAuthorityYear="1957" box="[981,1280,1871,1895]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="nodulosus">
|
||||
<emphasis id="B972EAC0A60FFF84AE8B6B0BFBE08D68" box="[981,1090,1871,1895]" italics="true" pageId="54" pageNumber="55">P. nodulosus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60FFF84A90E6B14FB548D68" author="Ramazzotti G" box="[1104,1270,1871,1895]" pageId="54" pageNumber="55" pagination="188 - 91" refId="ref39255" refString="Ramazzotti G. Due nuove specie di tardigradi extra-Europei. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 1957; 96: 188 - 91." type="journal article" year="1957">Ramazzotti, 1957</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, claws I; D,
|
||||
<taxonomicName id="4C064D51A60FFF84ADDF6B37FF4C8D8C" authorityName="CNHM" baseAuthorityName="Ramazzotti" baseAuthorityYear="1957" box="[129,238,1899,1923]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="nodulosus">
|
||||
<emphasis id="B972EAC0A60FFF84ADDF6B37FF4C8D8C" box="[129,238,1899,1923]" italics="true" pageId="54" pageNumber="55">P. nodulosus</emphasis>
|
||||
</taxonomicName>
|
||||
, claws IV; E,
|
||||
<taxonomicName id="4C064D51A60FFF84AC346B37FD778D8C" authority="(Bertolani & Rebecchi, 1996)" baseAuthorityName="Bertolani & Rebecchi" baseAuthorityYear="1996" box="[362,725,1899,1923]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="secchii">
|
||||
<emphasis id="B972EAC0A60FFF84AC346B37FE158D8C" box="[362,439,1899,1923]" italics="true" pageId="54" pageNumber="55">P. secchii</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60FFF84AC9B6B30FD698D8C" author="Bertolani R & Rebecchi L" box="[453,715,1899,1923]" pageId="54" pageNumber="55" pagination="3 - 12" refId="ref34938" refString="Bertolani R, Rebecchi L. The tardigrades of Emilia (Italy). II. Monte Rondinaio. AmultihabitatstudyonahighaltitudevalleyoftheNorthern Apennines. Zoological Journal of the Linnean Society 1996; 116: 3 - 12. https: // doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02329. x" type="journal article" year="1996">Bertolani & Rebecchi, 1996</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
nom. inq., claws II; F,
|
||||
<taxonomicName id="4C064D51A60FFF84AEF66B37FB638D8C" authority="(Robotti, 1970)" baseAuthorityName="Robotti" baseAuthorityYear="1970" box="[936,1217,1899,1923]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="ramazzottii">
|
||||
<emphasis id="B972EAC0A60FFF84AEF66B37FB878D8C" box="[936,1061,1900,1923]" italics="true" pageId="54" pageNumber="55">P. ramazzottii</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60FFF84A96D6B30FB158D8C" author="Robotti C" box="[1075,1207,1899,1923]" pageId="54" pageNumber="55" pagination="251 - 5" refId="ref39407" refString="Robotti C. Hypsibius (D.) ramazzottii spec. nov. e Macrobiotus aviglianae spec. nov. Primo contributo alla conoscenza dei tardigradi del Piemonte. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 1970; 110: 251 - 5." type="journal article" year="1970">Robotti, 1970</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, caudal sculpturing; G,
|
||||
<taxonomicName id="4C064D51A60FFF84ADDF6BD3FED18D90" authority="(Greven, 1972)" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[129,371,1927,1951]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A60FFF84ADDF6BD3FF798D90" box="[129,219,1927,1951]" italics="true" pageId="54" pageNumber="55">P. granifer</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60FFF84ADB76BDCFECB8D90" author="Greven H" box="[233,361,1927,1951]" pageId="54" pageNumber="55" pagination="368 - 81" refId="ref36334" refString="Greven H. Tardigraden des nordlichen Sauerlandes. Zoologischer Anzeiger 1972; 189: 368 - 81." type="journal article" year="1972">Greven, 1972</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA60EFF85AD2F6ADCFBD28CB3" ID-DOI="http://doi.org/10.5281/zenodo.14243840" ID-Zenodo-Dep="14243840" httpUri="https://zenodo.org/record/14243840/files/figure.png" pageId="55" pageNumber="56" startId="55.[113,178,1671,1695]" targetBox="[114,1458,144,1643]" targetPageId="55" targetType="figure">
|
||||
<paragraph id="8BB936D2A60EFF85AD2F6ADCFBD28CB3" blockId="55.[113,1406,1671,1724]" pageId="55" pageNumber="56">
|
||||
<emphasis id="B972EAC0A60EFF85AD2F6ADCFF748C90" bold="true" box="[113,214,1671,1695]" pageId="55" pageNumber="56">Figure 43.</emphasis>
|
||||
Various
|
||||
<taxonomicName id="4C064D51A60EFF85AC796ADCFE3B8C90" box="[295,409,1671,1695]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
, morphology of pharyngeal structures (SEM): A,
|
||||
<taxonomicName id="4C064D51A60EFF85AE316AD3FC6F8C90" box="[879,973,1671,1695]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A60EFF85AE316AD3FC6F8C90" box="[879,973,1671,1695]" italics="true" pageId="55" pageNumber="56">P. oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
; B,
|
||||
<taxonomicName id="4C064D51A60EFF85AEAF6AD3FBEF8C90" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[1009,1101,1671,1695]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A60EFF85AEAF6AD3FBEF8C90" box="[1009,1101,1671,1695]" italics="true" pageId="55" pageNumber="56">P. bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
; C,
|
||||
<taxonomicName id="4C064D51A60EFF85A92A6AD3FB538C90" baseAuthorityName="Robotti" baseAuthorityYear="1970" box="[1140,1265,1672,1695]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="ramazzottii">
|
||||
<emphasis id="B972EAC0A60EFF85A92A6AD3FB538C90" box="[1140,1265,1672,1695]" italics="true" pageId="55" pageNumber="56">P. ramazzottii</emphasis>
|
||||
</taxonomicName>
|
||||
; D,
|
||||
<taxonomicName id="4C064D51A60EFF85A8496AD3FEB28CB4" authority="(Richters, 1911)" baseAuthorityName="Richters" baseAuthorityYear="1911" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A60EFF85A8496AD3FADC8C90" box="[1303,1406,1672,1695]" italics="true" pageId="55" pageNumber="56">P. recamieri</emphasis>
|
||||
(Richters, 1911)
|
||||
</taxonomicName>
|
||||
; E,
|
||||
<taxonomicName id="4C064D51A60EFF85AC6D6AFFFDC58CB4" authority="(Binda & Pilato, 1971)" baseAuthorityName="Binda & Pilato" baseAuthorityYear="1971" box="[307,615,1699,1723]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="patanei">
|
||||
<emphasis id="B972EAC0A60EFF85AC6D6AFFFE288CB4" box="[307,394,1700,1723]" italics="true" pageId="55" pageNumber="56">P. patanei</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A60EFF85ACC76AF8FDFE8CB4" author="Binda MG & Pilato G" box="[409,604,1699,1723]" pageId="55" pageNumber="56" pagination="766 - 74" refId="ref35123" refString="Binda MG, Pilato G. Nuove osservazioni sui tardigradi delle Isole Eolie. Bollettino delle sedute dell'Accademia Gioenia di Scienze naturali di Catania, Serie IV 1971; 10: 766 - 74." type="journal article" year="1971">Binda & Pilato, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, lateral view; F,
|
||||
<taxonomicName id="4C064D51A60EFF85AFA36AFFFCF68CB4" baseAuthorityName="Binda & Pilato" baseAuthorityYear="1971" box="[765,852,1700,1723]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="species" species="patanei">
|
||||
<emphasis id="B972EAC0A60EFF85AFA36AFFFCF68CB4" box="[765,852,1700,1723]" italics="true" pageId="55" pageNumber="56">P. patanei</emphasis>
|
||||
</taxonomicName>
|
||||
, dorsal view. Scale bars: 5 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A60EFF8AAE1B6B35FDFD8CA4" blockId="55.[810,1460,1777,1958]" lastBlockId="56.[127,779,868,1989]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
|
||||
Shortly after
|
||||
<taxonomicName id="4C064D51A60EFF85AE836B35FB9F8D89" box="[989,1085,1902,1926]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="55" pageNumber="56" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A60EFF85AE836B35FB9F8D89" box="[989,1085,1902,1926]" italics="true" pageId="55" pageNumber="56">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
was established (
|
||||
<bibRefCitation id="EF974B23A60EFF85A8536B35FCFF8DA9" author="Bertolani R & Guidetti R & Marchioro T" pageId="55" pageNumber="56" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A60EFF85A8256B34FA168D89" box="[1403,1460,1902,1926]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
), it was revealed that the genus is paraphyletic, because
|
||||
<taxonomicName id="4C064D51A601FF8AADDF6F3FFEA88973" authorityName="Tumanov" authorityYear="2020" box="[129,266,868,892]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AADDF6F3FFEA88973" box="[129,266,868,892]" italics="true" pageId="56" pageNumber="57">Notahypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
, which exhibits only a rigid buccal tube, is embedded among the pilatobiins (
|
||||
<bibRefCitation id="EF974B23A601FF8AAC9D6FDFFDC28993" author="Tumanov DV" box="[451,608,900,924]" pageId="56" pageNumber="57" pagination="1 - 37" refId="ref40150" refString="Tumanov DV. Integrative redescription of Hypsibius pallidoides Pilato et al., 2011 (Eutardigrada: Hypsibioidea) with the erection of a new genus and discussion on the phylogeny of Hypsibiidae. European Journal of Taxonomy 2020; 681: 1 - 37. https: // doi. org / 10.5852 / ejt. 2020.681" type="journal article" year="2020">Tumanov 2020</bibRefCitation>
|
||||
). The new phylogeny (
|
||||
<figureCitation id="133D2A57A601FF8AAD896FF8FEBB89B4" box="[215,281,931,955]" captionStart="Figure 2" captionStartId="5.[113,178,1715,1739]" captionTargetBox="[115,1454,150,1684]" captionTargetId="graphics-78@5.[115,1181,156,1684]" captionTargetPageId="5" captionText="Figure 2. Phylogenetic relationships within Hypsibiidae in the Bayesian and maximum likelihood analyses (identical topologies; the scale refers to the Bayesian tree and represents substitutions per position; posterior probability values are provided above the nodes, whereas bootstrap values can be found below the nodes; *, maximal support; #, no support).Calohypsibius ornatus constitutes an outgroup. Newly added species and populations are in bold." figureDoi="http://doi.org/10.5281/zenodo.14243746" httpUri="https://zenodo.org/record/14243746/files/figure.png" pageId="56" pageNumber="57">Fig. 2</figureCitation>
|
||||
) corroborates the previous findings, placing
|
||||
<taxonomicName id="4C064D51A601FF8AADDF6F99FEA989D5" authorityName="Tumanov" authorityYear="2020" box="[129,267,962,986]" class="Eutardigrada" family="Hypsibiidae" genus="Notahypsibius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AADDF6F99FEA989D5" box="[129,267,962,986]" italics="true" pageId="56" pageNumber="57">Notahypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
as sister to the
|
||||
<taxonomicName id="4C064D51A601FF8AACFE6F98FDAF89D5" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[416,525,963,986]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A601FF8AACFE6F98FDAF89D5" box="[416,525,963,986]" italics="true" pageId="56" pageNumber="57">P. recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
group, whereas the clade
|
||||
<taxonomicName id="4C064D51A601FF8AADDE6FB9FF4589F5" box="[128,231,994,1018]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A601FF8AADDE6FB9FF4589F5" box="[128,231,994,1018]" italics="true" pageId="56" pageNumber="57">P. oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
group +
|
||||
<taxonomicName id="4C064D51A601FF8AAC166FB9FE6989F5" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[328,459,994,1018]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AAC166FB9FEF889F6" box="[328,346,994,1017]" italics="true" pageId="56" pageNumber="57">P.</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A601FF8AAC206FB9FE6989F5" box="[382,459,994,1018]" italics="true" pageId="56" pageNumber="57">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
is sister to that clade. In an attempt to make the systematics of
|
||||
<taxonomicName id="4C064D51A601FF8AAC85685AFDF18E16" box="[475,595,1025,1049]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
closer to the natural state, we propose the division of the genus into four coherent morphological groups, in a similar manner to the way in which we approached the reorganization of
|
||||
<taxonomicName id="4C064D51A601FF8AAC866804FDEE8E78" authorityName="Thulin" authorityYear="1928" box="[472,588,1119,1143]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AAC866804FDEE8E78" box="[472,588,1119,1143]" italics="true" pageId="56" pageNumber="57">Isohypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A601FF8AAF206804FEF98E99" authority="(Gasiorek et al. 2019)" baseAuthorityName="Gasiorek" baseAuthorityYear="2019" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A601FF8AAF206804FCA88E78" box="[638,778,1119,1143]" italics="true" pageId="56" pageNumber="57">Doryphoribius</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AADD56825FEED8E99" author="Gasiorek P & Stec D & Morek W" box="[139,335,1150,1174]" pageId="56" pageNumber="57" pagination="78 - 132" refId="ref36279" refString="Gasiorek P, Stec D, Morek W et al. Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology 2019; 88: 78 - 132. https: // doi. org / 10.1163 / 18759866 - 20191350" type="journal article" year="2019">
|
||||
Gąsiorek
|
||||
<emphasis id="B972EAC0A601FF8AADB46824FEBA8E99" box="[234,280,1150,1174]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
||||
2019
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Hence, the
|
||||
<taxonomicName id="4C064D51A601FF8AAC8B6824FD8F8E99" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[469,557,1151,1174]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A601FF8AAC8B6824FD8F8E99" box="[469,557,1151,1174]" italics="true" pageId="56" pageNumber="57">recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
group comprises species with a smooth cuticle that lack dorsal gibbosities, whereas the
|
||||
<taxonomicName id="4C064D51A601FF8AADF568E6FF5A8EDA" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[171,248,1213,1237]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AADF568E6FF5A8EDA" box="[171,248,1213,1237]" italics="true" pageId="56" pageNumber="57">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group (=
|
||||
<emphasis id="B972EAC0A601FF8AAC3368E6FE508EDA" box="[365,498,1213,1237]" italics="true" pageId="56" pageNumber="57">
|
||||
<taxonomicName id="4C064D51A601FF8AAC3368E6FE6F8EDA" box="[365,461,1213,1237]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="genus" sensu="stricto">Pilatobius</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA601FF8AAC8968E5FE508EDA" box="[471,498,1214,1237]" pageId="56" pageNumber="57" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
owing to the inclusion of
|
||||
<taxonomicName id="4C064D51A601FF8AADDF6886FF478EFB" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[129,229,1244,1268]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AADDF6886FF478EFB" box="[129,229,1244,1268]" italics="true" pageId="56" pageNumber="57">P. bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
, its
|
||||
<typeStatus id="54BD8870A601FF8AAC516886FE9E8EFA" box="[271,316,1245,1269]" pageId="56" pageNumber="57">type</typeStatus>
|
||||
species) comprises species with a sculptured cuticle and dorsal gibbosities. Neither the
|
||||
<taxonomicName id="4C064D51A601FF8AAF7468A7FDDB8F1B" box="[554,633,1276,1300]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A601FF8AAF7468A7FDDB8F1B" box="[554,633,1276,1300]" italics="true" pageId="56" pageNumber="57">oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
group nor the
|
||||
<taxonomicName id="4C064D51A601FF8AADDF6947FF6B8F3C" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[129,201,1308,1331]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AADDF6947FF6B8F3C" box="[129,201,1308,1331]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group exhibit dorsolateral gibbosities; however, the cuticular sculpturing is developed differently in each of these species complexes: in the
|
||||
<taxonomicName id="4C064D51A601FF8AAC656901FE288F7D" box="[315,394,1370,1394]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A601FF8AAC656901FE288F7D" box="[315,394,1370,1394]" italics="true" pageId="56" pageNumber="57">oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
group, the cuticle is noticeably sculptured only caudally, although some small sculpturing may appear more anteriorly; whereas in the
|
||||
<taxonomicName id="4C064D51A601FF8AAC9269C2FDB68FBF" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[460,532,1433,1456]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AAC9269C2FDB68FBF" box="[460,532,1433,1456]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group, the cuticle is intensely sculptured over the whole dorsum. Two of these groups (the
|
||||
<taxonomicName id="4C064D51A601FF8AADEC698CFF5D8FE0" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[178,255,1495,1519]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AADEC698CFF5D8FE0" box="[178,255,1495,1519]" italics="true" pageId="56" pageNumber="57">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AAC55698CFE1F8FE0" box="[267,445,1495,1519]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="variety" species="oculatus" variety="recamieri">
|
||||
<emphasis id="B972EAC0A601FF8AAC55698CFEF88FE0" box="[267,346,1495,1519]" italics="true" pageId="56" pageNumber="57">oculatus</emphasis>
|
||||
,
|
||||
<emphasis id="B972EAC0A601FF8AAC3B6983FE1F8FE0" box="[357,445,1496,1519]" italics="true" pageId="56" pageNumber="57">recamieri</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A601FF8AACA96983FD9D8FE0" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[503,575,1496,1519]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AACA96983FD9D8FE0" box="[503,575,1496,1519]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
groups) are erected as new genera (see below) to reflect the phylogeny of
|
||||
<taxonomicName id="4C064D51A601FF8AAFCC69ADFCA98C01" box="[658,779,1526,1550]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="subFamily" subFamily="Pilatobiinae">Pilatobiinae</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133D2A57A601FF8AADD56A4DFF668C21" box="[139,196,1558,1582]" captionStart="Figure 2" captionStartId="5.[113,178,1715,1739]" captionTargetBox="[115,1454,150,1684]" captionTargetId="graphics-78@5.[115,1181,156,1684]" captionTargetPageId="5" captionText="Figure 2. Phylogenetic relationships within Hypsibiidae in the Bayesian and maximum likelihood analyses (identical topologies; the scale refers to the Bayesian tree and represents substitutions per position; posterior probability values are provided above the nodes, whereas bootstrap values can be found below the nodes; *, maximal support; #, no support).Calohypsibius ornatus constitutes an outgroup. Newly added species and populations are in bold." figureDoi="http://doi.org/10.5281/zenodo.14243746" httpUri="https://zenodo.org/record/14243746/files/figure.png" pageId="56" pageNumber="57">Fig. 2</figureCitation>
|
||||
). Unfortunately, owing to our stringent selection criteria for the composition of the genetic dataset, we were forced to eliminate
|
||||
<taxonomicName id="4C064D51A601FF8AADBD6A0EFEF48C63" authorityName="CNHM" baseAuthorityName="Ramazzotti" baseAuthorityYear="1957" box="[227,342,1620,1644]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="nodulosus">
|
||||
<emphasis id="B972EAC0A601FF8AADBD6A0EFEF48C63" box="[227,342,1620,1644]" italics="true" pageId="56" pageNumber="57">P. nodulosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A601FF8AAC3B6A0EFE0F8C63" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[357,429,1621,1644]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AAC3B6A0EFE0F8C63" box="[357,429,1621,1644]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A601FF8AAF5E6A0EFDFF8C63" baseAuthorityName="Binda & Pilato" baseAuthorityYear="1971" box="[512,605,1621,1644]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="patanei">
|
||||
<emphasis id="B972EAC0A601FF8AAF5E6A0EFDFF8C63" box="[512,605,1621,1644]" italics="true" pageId="56" pageNumber="57">P.patanei</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A601FF8AAF336A0FFD188C63" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[621,698,1620,1644]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AAF336A0FFD188C63" box="[621,698,1620,1644]" italics="true" pageId="56" pageNumber="57">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group), and
|
||||
<taxonomicName id="4C064D51A601FF8AADF56A2FFE8D8C84" baseAuthorityName="Robotti" baseAuthorityYear="1970" box="[171,303,1652,1675]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="ramazzottii">
|
||||
<emphasis id="B972EAC0A601FF8AADF56A2FFE8D8C84" box="[171,303,1652,1675]" italics="true" pageId="56" pageNumber="57">P. ramazzottii</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C064D51A601FF8AAC606A2FFE248C84" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[318,390,1652,1675]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AAC606A2FFE248C84" box="[318,390,1652,1675]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group) from our analysis, because only 18S rRNA barcodes were available in GenBank.
|
||||
</paragraph>
|
||||
<caption id="DF79665AA601FF8AADDC6E67FED7893B" ID-DOI="http://doi.org/10.5281/zenodo.14243843" ID-Zenodo-Dep="14243843" httpUri="https://zenodo.org/record/14243843/files/figure.png" pageId="56" pageNumber="57" startId="56.[130,195,572,596]" targetBox="[130,776,144,544]" targetPageId="56" targetType="figure">
|
||||
<paragraph id="8BB936D2A601FF8AADDC6E67FED7893B" blockId="56.[129,754,572,820]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B972EAC0A601FF8AADDC6E67FF44885B" bold="true" box="[130,230,572,596]" pageId="56" pageNumber="57">Figure 44.</emphasis>
|
||||
Evolution scheme for a tardigrade clade, where the blue triangle and red rectrangle signify newly erected genera (‘apical taxa’ in the paper by
|
||||
<bibRefCitation id="EF974B23A601FF8AAC1C6E2EFDFC8882" author="Tumanov DV & Tsvetkova AY" box="[322,606,628,653]" pageId="56" pageNumber="57" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
) with ‘strong apomorphies’, i.e. the set of well-defined, advanced morphological characters, distinguished from the previous broadly defined genus, whose members (marked with black circles) are still classified as the original genus, which became artificial but cannot be divided further because the lineages group species with mainly (or solely) plesiomorphic characters.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A601FF8AADC26AE8FA3C8AE9" blockId="56.[127,779,868,1989]" lastBlockId="56.[825,1475,144,231]" pageId="56" pageNumber="57">
|
||||
In agreement with the doubts raised by
|
||||
<bibRefCitation id="EF974B23A601FF8AAF3D6AE9FEBF8CE5" author="Ramazzotti G & Maucci W" pageId="56" pageNumber="57" pagination="1 - 1012" refId="ref39289" refString="Ramazzotti G, Maucci W. Il Phylum Tardigrada. III edizione riveduta e aggiornata. Memorie dell'Istituto Italiano di Idrobiologia 1983; 41: 1 - 1012." type="journal article" year="1983">Ramazzotti and Maucci (1983)</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="4C064D51A601FF8AAC756A89FDF48CE5" authority="(Mihelcic, 1959)" baseAuthorityName="Mihelcic" baseAuthorityYear="1959" box="[299,598,1745,1770]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="elongatus">
|
||||
<emphasis id="B972EAC0A601FF8AAC756A89FE3C8CE5" box="[299,414,1746,1770]" italics="true" pageId="56" pageNumber="57">P. elongatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AACEE6A89FDE98CE6" author="Mihelcic F" box="[432,587,1745,1770]" pageId="56" pageNumber="57" pagination="254 - 61" refId="ref37940" refString="Mihelcic F. Zwei neue Tardigraden aus der Gattung Hypsibius Thulin aus Osttirol (Osterreich): Systematisches zur Gattung Hypsibius Thulin. Zoologischer Anzeiger 1959; 163: 254 - 61." type="journal article" year="1959">Mihelčič, 1959</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
sp. dub. is designated as
|
||||
<emphasis id="B972EAC0A601FF8AAD886AAAFEDC8D07" bold="true" box="[214,382,1777,1800]" italics="true" pageId="56" pageNumber="57">synonym novum</emphasis>
|
||||
of
|
||||
<taxonomicName id="4C064D51A601FF8AACC26AA9FE5C8D06" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[412,510,1777,1801]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AACC26AA9FE5C8D06" box="[412,510,1777,1801]" italics="true" pageId="56" pageNumber="57">P. bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
owing to the lack of separating characters.Given that the alleged rows of ‘minor’ gibbosities in
|
||||
<taxonomicName id="4C064D51A601FF8AADC26B6BFDAB8D47" authority="(Iharos, 1964)" baseAuthorityName="Iharos" baseAuthorityYear="1964" box="[156,521,1840,1864]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bisbullatus">
|
||||
<emphasis id="B972EAC0A601FF8AADC26B6BFECE8D47" box="[156,364,1840,1864]" italics="true" pageId="56" pageNumber="57">Pilatobius bisbullatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AAC236B6BFE5C8D47" author="Iharos G" box="[381,510,1840,1864]" pageId="56" pageNumber="57" pagination="57 - 67" refId="ref36710" refString="Iharos G. Neuere Beitrage zur Kenntnis der Tardigraden-Fauna Ungarns, V. Opuscula Zoologica, Budapest 1964; 5: 57 - 67." type="journal article" year="1964">Iharos, 1964</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
seem to be only folds of cuticle between otherwise typical gibbosities and are an artefact resulting from shrinkage of the specimens in the mounting medium, we designate the species as invalid. Other species (
|
||||
<taxonomicName id="4C064D51A601FF8AADD56BF5FE028DCA" authority="(Mihelcic, 1951)" baseAuthorityName="Mihelcic" baseAuthorityYear="1951" box="[139,416,1965,1989]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="gerdae">
|
||||
<emphasis id="B972EAC0A601FF8AADD56BF5FF468DCA" box="[139,228,1965,1989]" italics="true" pageId="56" pageNumber="57">P. gerdae</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AADA76BF6FE378DCA" author="Mihelcic F" box="[249,405,1965,1989]" pageId="56" pageNumber="57" pagination="57 - 103" refId="ref37895" refString="Mihelcic F. Beitrag zur Systematik de Tardigraden. Archivio Zoologico Italiano 1951; 36: 57 - 103." type="journal article" year="1951">Mihelčič, 1951</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
sp. dub.,
|
||||
<taxonomicName id="4C064D51A601FF8AAF546BF5FCDA8AA7" authority="(Mihelcic, 1951)" baseAuthorityName="Mihelcic" baseAuthorityYear="1951" box="[522,888,144,1989]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="nonbullatus">
|
||||
<emphasis id="B972EAC0A601FF8AAF546BF5FD378DCA" box="[522,661,1965,1989]" italics="true" pageId="56" pageNumber="57">P. nonbullatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AAFF46BF6FCCF8AA7" author="Mihelcic F" box="[682,877,144,1989]" pageId="56" pageNumber="57" pagination="57 - 103" refId="ref37895" refString="Mihelcic F. Beitrag zur Systematik de Tardigraden. Archivio Zoologico Italiano 1951; 36: 57 - 103." type="journal article" year="1951">Mihelčič, 1951</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
sp. dub. (both in
|
||||
<taxonomicName id="4C064D51A601FF8AA9416CCBFBCE8AA7" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[1055,1132,144,168]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="bullatus">
|
||||
<emphasis id="B972EAC0A601FF8AA9416CCBFBCE8AA7" box="[1055,1132,144,168]" italics="true" pageId="56" pageNumber="57">bullatus</emphasis>
|
||||
</taxonomicName>
|
||||
group),
|
||||
<taxonomicName id="4C064D51A601FF8AA9E16CCAFA608AA7" authority="(Mihelcic, 1955)" baseAuthorityName="Mihelcic" baseAuthorityYear="1955" box="[1215,1474,144,168]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="latipes">
|
||||
<emphasis id="B972EAC0A601FF8AA9E16CCAFAB38AA7" box="[1215,1297,144,168]" italics="true" pageId="56" pageNumber="57">P.latipes</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A601FF8AA87F6CCBFA158AA7" author="Mihelcic F" box="[1313,1463,144,168]" pageId="56" pageNumber="57" pagination="240 - 4" refId="ref37915" refString="Mihelcic F. Zur Okologie und Verbreitung der Gattung Hypsibius (Tardigrada). Bonner Zoologische Beitrage 1955; 6: 240 - 4." type="journal article" year="1955">Mihelčič, 1955</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
sp. dub. (
|
||||
<taxonomicName id="4C064D51A601FF8AAEFD6CEBFC498AC8" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[931,1003,176,199]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="56" pageNumber="57" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A601FF8AAEFD6CEBFC498AC8" box="[931,1003,176,199]" italics="true" pageId="56" pageNumber="57">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group)) designated as dubious by
|
||||
<bibRefCitation id="EF974B23A601FF8AA8316CF4FC218AE9" author="Dastych H" pageId="56" pageNumber="57" pagination="1 - 25" refId="ref35653" refString="Dastych H. Checklisten der Fauna Osterreichs, No. 8: Tardigrada. In: Winkler H, Stuessy T (eds.), Biosystematics and Ecology Series, No. 31. Osterreichischen Akademie der Wissenschaften, 2015, 1 - 25. https: // doi. org / 10.1553 / 0 x 00327 e 9 a" type="book chapter" year="2015">Dastych (2015)</bibRefCitation>
|
||||
are also considered as such and rejected in this paper.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1172
data/03/AF/87/03AF87C4A615FF9CAC4C6976FAFD8C41.xml
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1172
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416
data/03/AF/87/03AF87C4A617FF82AE9A6A36FB5E8CB1.xml
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416
data/03/AF/87/03AF87C4A617FF82AE9A6A36FB5E8CB1.xml
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|
|
@ -0,0 +1,416 @@
|
|||
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<mods:titleInfo id="19BA4AC1CC2EBD823D83E7DC16317576">
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<mods:title id="57C05C8EB5CD6CEFDE6803116C73FE13">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:affiliation id="A05B1EEE6DCDFAA0DEF3841906798C99">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="03A2E3194B262A34DD34AE9E156FF955" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7B35E05575168F97839C6B932CE1CCEF">Blagden, Brian</mods:namePart>
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<mods:affiliation id="741BECAB1A1121305EDDB9B24B9C70B7">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="DD592F5B84CB139EEC40FF9EF3A801E3">Morek, Witold</mods:namePart>
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<mods:affiliation id="F5F160D54DCB6D762F59559432C2049D">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="3B7711F67DD8E3ABB20BE1C9FD4ABD31">Michalczyk, Łukasz</mods:namePart>
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<treatment id="03AF87C4A617FF82AE9A6A36FB5E8CB1" LSID="urn:lsid:plazi:treatment:03AF87C4A617FF82AE9A6A36FB5E8CB1" httpUri="http://treatment.plazi.org/id/03AF87C4A617FF82AE9A6A36FB5E8CB1" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
|
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<subSubSection id="C31C6559A617FF9CAE9A6A36FA9A8C88" box="[964,1336,1645,1672]" pageId="46" pageNumber="47" type="nomenclature">
|
||||
<paragraph id="8BB936D2A617FF9CAE9A6A36FA9A8C88" blockId="46.[964,1336,1645,1672]" box="[964,1336,1645,1672]" pageId="46" pageNumber="47">
|
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<heading id="D0F181BEA617FF9CAE9A6A36FA9A8C88" bold="true" box="[964,1336,1645,1672]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
|
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<taxonomicName id="4C064D51A617FF9CAE9A6A36FA9A8C88" ID-CoL="9WRBD" authority="(Bartos, 1935)" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[964,1336,1645,1672]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="46" pageNumber="47" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A617FF9CAE9A6A36FA9A8C88" bold="true" box="[964,1336,1645,1672]" pageId="46" pageNumber="47">
|
||||
<emphasis id="B972EAC0A617FF9CAE9A6A36FB888C87" bold="true" box="[964,1066,1645,1672]" italics="true" pageId="46" pageNumber="47">Pilatobius</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A617FF9CA9156A35FB358C87" bold="true" box="[1099,1175,1646,1672]" italics="true" pageId="46" pageNumber="47">rugosus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A617FF9CA9F86A36FA8F8C88" author="Bartos E" box="[1190,1325,1645,1671]" pageId="46" pageNumber="47" pagination="257 - 60" refId="ref34712" refString="Bartos E. Vier neue Hypsibius - Arten aus der Tschechoslovakei. Zoologischer Anzeiger 1935; 110: 257 - 60." type="journal article" year="1935">Bartoš, 1935</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A617FF9CAE046AF7FA308D6C" pageId="46" pageNumber="47" type="reference_group">
|
||||
<paragraph id="8BB936D2A617FF9CAE046AF7FA908CE9" blockId="46.[858,1441,1708,1766]" pageId="46" pageNumber="47">
|
||||
<emphasis id="B972EAC0A617FF9CAE046AF7FA908CE9" bold="true" pageId="46" pageNumber="47">
|
||||
<taxonomicName id="4C064D51A617FF9CAE046AF7FB378CC8" authorityName="Bartos" authorityYear="1935" box="[858,1173,1708,1735]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="46" pageNumber="47" phylum="Tardigrada" rank="species" species="rugosus" subGenus="Diphascon">
|
||||
<emphasis id="B972EAC0A617FF9CAE046AF7FC1C8CC8" bold="true" box="[858,958,1708,1735]" italics="true" pageId="46" pageNumber="47">Hypsibius</emphasis>
|
||||
(
|
||||
<emphasis id="B972EAC0A617FF9CAE906AF7FB998CC8" bold="true" box="[974,1083,1708,1735]" italics="true" pageId="46" pageNumber="47">Diphascon</emphasis>
|
||||
)
|
||||
<emphasis id="B972EAC0A617FF9CA9176AF6FB378CC8" bold="true" box="[1097,1173,1709,1735]" italics="true" pageId="46" pageNumber="47">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<emphasis id="B972EAC0A617FF9CA9FF6AF7FABD8CC9" bold="true" box="[1185,1311,1708,1735]" italics="true" pageId="46" pageNumber="47">locus typicus</emphasis>
|
||||
: Karlik near Mošovce,
|
||||
<collectingCountry id="F3117642A617FF9CA96C6A90FB2F8CEA" box="[1074,1165,1739,1765]" name="Slovakia" pageId="46" pageNumber="47">Slovakia</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A617FF9CA9C86A90FA8F8CEA" author="Bartos E" box="[1174,1325,1739,1765]" pageId="46" pageNumber="47" pagination="257 - 60" refId="ref34712" refString="Bartos E. Vier neue Hypsibius - Arten aus der Tschechoslovakei. Zoologischer Anzeiger 1935; 110: 257 - 60." type="journal article" year="1935">Bartoš (1935)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A617FF9CAE3A6B50FA358D2B" blockId="46.[868,1431,1802,1829]" box="[868,1431,1802,1829]" pageId="46" pageNumber="47">
|
||||
<heading id="D0F181BEA617FF9CAE3A6B50FA358D2B" bold="true" box="[868,1431,1802,1829]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
|
||||
<emphasis id="B972EAC0A617FF9CAE3A6B50FA358D2B" bold="true" box="[868,1431,1802,1829]" pageId="46" pageNumber="47">
|
||||
<taxonomicName id="4C064D51A617FF9CAE3A6B50FC6F8D2A" authorityName="Bartos" authorityYear="1935" box="[868,973,1802,1829]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="46" pageNumber="47" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A617FF9CAE3A6B50FCD58D2A" bold="true" box="[868,887,1803,1829]" italics="true" pageId="46" pageNumber="47">D</emphasis>
|
||||
.
|
||||
<emphasis id="B972EAC0A617FF9CAEDF6B50FC6F8D2A" bold="true" box="[897,973,1803,1829]" italics="true" pageId="46" pageNumber="47">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
; Tatra Mountains,
|
||||
<collectingCountry id="F3117642A617FF9CA9CC6B51FB428D2B" box="[1170,1248,1802,1828]" name="Poland" pageId="46" pageNumber="47">Poland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A617FF9CA9B46B51FA308D2B" author="Dastych H" box="[1258,1426,1802,1828]" pageId="46" pageNumber="47" pagination="1 - 232" refId="ref35518" refString="Dastych H. Niesporczaki (Tardigrada) Tatrzanskiego Parku Narodowego. Monografie Fauny Polski 1980; 9: 1 - 232." type="journal article" year="1980">Dastych (1980)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A617FF9CAE376B12FA308D6C" blockId="46.[873,1426,1865,1891]" box="[873,1426,1865,1891]" pageId="46" pageNumber="47">
|
||||
<heading id="D0F181BEA617FF9CAE376B12FA308D6C" bold="true" box="[873,1426,1865,1891]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
|
||||
<emphasis id="B972EAC0A617FF9CAE376B12FA308D6C" bold="true" box="[873,1426,1865,1891]" pageId="46" pageNumber="47">
|
||||
<taxonomicName id="4C064D51A617FF9CAE376B12FC7F8D6C" ID-CoL="7T86X" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[873,989,1865,1891]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="46" pageNumber="47" phylum="Tardigrada" rank="species" species="rugosum">
|
||||
<emphasis id="B972EAC0A617FF9CAE376B12FCDE8D6C" bold="true" box="[873,892,1865,1891]" italics="true" pageId="46" pageNumber="47">D</emphasis>
|
||||
.
|
||||
<emphasis id="B972EAC0A617FF9CAED86B12FC7F8D6C" bold="true" box="[902,989,1865,1891]" italics="true" pageId="46" pageNumber="47">rugosum</emphasis>
|
||||
</taxonomicName>
|
||||
; various locales,
|
||||
<collectingCountry id="F3117642A617FF9CA9D36B12FB798D6C" box="[1165,1243,1865,1891]" name="Poland" pageId="46" pageNumber="47">Poland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A617FF9CA9BB6B12FA2F8D6C" author="Dastych H" box="[1253,1421,1865,1891]" pageId="46" pageNumber="47" pagination="1 - 255" refId="ref35583" refString="Dastych H. The Tardigrada of Poland. Monografie Fauny Polski 1988; 16: 1 - 255." type="journal article" year="1988">Dastych (1988)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A617FF9CAE676BD2FB008DCE" pageId="46" pageNumber="47" type="materials_examined">
|
||||
<paragraph id="8BB936D2A617FF9CAE676BD2FB008DCE" blockId="46.[825,1474,1929,1985]" pageId="46" pageNumber="47">
|
||||
<emphasis id="B972EAC0A617FF9CAE676BD2FBA08DAE" box="[825,1026,1929,1953]" italics="true" pageId="46" pageNumber="47">Material examined:</emphasis>
|
||||
40 individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A617FF9CA9646BF2FB338DCE" box="[1082,1169,1961,1985]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="46" pageNumber="47">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF79665AA616FF9DAD2F6B0AFD618DAE" ID-DOI="http://doi.org/10.5281/zenodo.14243826" ID-Zenodo-Dep="14243826" httpUri="https://zenodo.org/record/14243826/files/figure.png" pageId="47" pageNumber="48" startId="47.[113,178,1873,1897]" targetBox="[140,1433,144,1846]" targetPageId="47" targetType="figure">
|
||||
<paragraph id="8BB936D2A616FF9DAD2F6B0AFD618DAE" blockId="47.[113,1451,1873,1953]" pageId="47" pageNumber="48">
|
||||
<emphasis id="B972EAC0A616FF9DAD2F6B0AFF748D66" bold="true" box="[113,214,1873,1897]" pageId="47" pageNumber="48">Figure 36.</emphasis>
|
||||
Detailed morphology of buccopharyngeal apparatus of
|
||||
<taxonomicName id="4C064D51A616FF9DAFBD6B0AFC2E8D66" baseAuthorityName="Murray" baseAuthorityYear="1906" box="[739,908,1873,1897]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="47" pageNumber="48" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A616FF9DAFBD6B0AFC2E8D66" box="[739,908,1873,1897]" italics="true" pageId="47" pageNumber="48">Pilatobius oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
(all but A, SEM): A, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A616FF9DA8026B09FA378D66" box="[1372,1429,1874,1897]" italics="true" pageId="47" pageNumber="48">in toto</emphasis>
|
||||
(PCM); B,
|
||||
<emphasis id="B972EAC0A616FF9DAD836B36FEAB8D8A" box="[221,265,1901,1925]" italics="true" pageId="47" pageNumber="48">idem</emphasis>
|
||||
; C, DABT and the transition zone between the buccal and pharyngeal tube; D, vanishing of annulation in the posterior part of the pharyngeal tube; E, pharynx. Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA609FF82ADDF6F4BFB8A8926" ID-DOI="http://doi.org/10.5281/zenodo.14243828" ID-Zenodo-Dep="14243828" box="[129,1064,784,809]" httpUri="https://zenodo.org/record/14243828/files/figure.png" pageId="48" pageNumber="49" startId="48.[129,194,784,808]" targetBox="[129,1473,144,756]" targetPageId="48" targetType="figure">
|
||||
<paragraph id="8BB936D2A609FF82ADDF6F4BFB8A8926" blockId="48.[129,1064,784,809]" box="[129,1064,784,809]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B972EAC0A609FF82ADDF6F4BFF478927" bold="true" box="[129,229,784,808]" pageId="48" pageNumber="49">Figure 37.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A609FF82AC616F4BFD9A8927" authority="(PCM)" baseAuthorityName="PCM" box="[319,568,784,808]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="oculatus">
|
||||
<emphasis id="B972EAC0A609FF82AC616F4BFE4A8927" box="[319,488,784,808]" italics="true" pageId="48" pageNumber="49">Pilatobius oculatus</emphasis>
|
||||
(PCM)
|
||||
</taxonomicName>
|
||||
: A, claws I; B, claws IV (neotype). Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C31C6559A609FF82ADDF6F02FDE28C0D" pageId="48" pageNumber="49" type="description">
|
||||
<paragraph id="8BB936D2A609FF82ADDF6F02FEC28F08" blockId="48.[127,779,856,1538]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B972EAC0A609FF82ADDF6F02FF55897F" box="[129,247,857,880]" italics="true" pageId="48" pageNumber="49">Description:</emphasis>
|
||||
Body small to medium in size (
|
||||
<tableCitation id="C6840369A609FF82AF666F03FD32897F" box="[568,656,856,880]" captionStart="Table 13" captionStartId="49.[113,168,143,167]" captionTargetPageId="49" captionText="Table 13. Measurements (in micrometres) of selected morphological structures of Pilatobius cf. rugosus (Bartoš, 1935) from Poland (PL.120) mounted in Hoyer’s medium." pageId="48" pageNumber="49">Table 13</tableCitation>
|
||||
), corpulent (
|
||||
<figureCitation id="133D2A57A609FF82ADD56F2CFF43899F" box="[139,225,887,912]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38A</figureCitation>
|
||||
) and white. Cuticle uniformly sculptured on the entire dorsal surface by polygons of various sizes placed tightly adjacent to each other (
|
||||
<figureCitation id="133D2A57A609FF82AC156FEDFE1C89C1" box="[331,446,950,974]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38B, C</figureCitation>
|
||||
); legs smooth. Cribriform areas not visible under PCM. Legs short, plump, and barely delimited from the trunk (
|
||||
<figureCitation id="133D2A57A609FF82AC6E6FAEFE2B8E02" box="[304,393,1013,1037]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38A</figureCitation>
|
||||
). Eyes present in living animals, but usually quickly dissolving in Hoyer’s medium. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A609FF82AC6D6868FE368E44" box="[307,404,1075,1099]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A609FF82AC6D6868FE368E44" box="[307,404,1075,1099]" italics="true" pageId="48" pageNumber="49">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A609FF82ACC2686FFE698E43" box="[412,459,1076,1100]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
, with a short pharyngeal tube (
|
||||
<figureCitation id="133D2A57A609FF82ADD56808FF408E64" box="[139,226,1107,1131]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38D</figureCitation>
|
||||
). The OCA not visible under PCM (
|
||||
<figureCitation id="133D2A57A609FF82AF066808FD0D8E64" box="[600,687,1107,1131]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38D</figureCitation>
|
||||
). Furcae of the
|
||||
<taxonomicName id="4C064D51A609FF82AD946829FE888E85" authorityName="Ehrenberg" authorityYear="1848" box="[202,298,1138,1162]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A609FF82AD946829FE888E85" box="[202,298,1138,1162]" italics="true" pageId="48" pageNumber="49">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A609FF82AC6A6828FEC68E84" box="[308,356,1139,1163]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
. The DABT large and slightly elongate. Pharynx circular, with large pharyngeal apophyses. Macroplacoid length sequence 2 <1, the first with a delicate central constriction (
|
||||
<figureCitation id="133D2A57A609FF82ADE1688BFEB98EE7" box="[191,283,1232,1256]" captionStart="Figure 38" captionStartId="50.[129,194,1753,1777]" captionTargetBox="[129,1473,144,1725]" captionTargetId="figure-75@50.[129,1473,144,1725]" captionTargetPageId="50" captionText="Figure 38. General morphology of Pilatobius cf. rugosus (Bartoš, 1935) (all but C, PCM): A, specimen in toto (ventral view); B, cuticular sculpturing of the caudal region; C, cuticular sculpturing of the caudal region (SEM); D, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243830" httpUri="https://zenodo.org/record/14243830/files/figure.png" pageId="48" pageNumber="49">Fig. 38D</figureCitation>
|
||||
). Septulum large, but clearly shorter than the second macroplacoid.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A609FF82ADC26954FDE28C0D" blockId="48.[127,779,856,1538]" pageId="48" pageNumber="49">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A609FF82AC7A6954FE268F28" authorityName="Ehrenberg" authorityYear="1848" box="[292,388,1295,1319]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A609FF82AC7A6954FE268F28" box="[292,388,1295,1319]" italics="true" pageId="48" pageNumber="49">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A609FF82ACD46954FE1B8F28" box="[394,441,1295,1319]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
, with external and internal claws of similar size (
|
||||
<figureCitation id="133D2A57A609FF82AC406975FEC68F49" box="[286,356,1326,1350]" captionStart="Figure 39" captionStartId="51.[113,176,1170,1194]" captionTargetBox="[114,1458,144,1142]" captionTargetId="figure-459@51.[114,1458,144,1142]" captionTargetPageId="51" captionText="Figure 39.Claws of Pilatobius cf.rugosus: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars,and white arrowhead indicates anterior bar.Scale bars are in micrometres, identical in panels A, B and in C, D." figureDoi="http://doi.org/10.5281/zenodo.14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="48" pageNumber="49">Fig. 39</figureCitation>
|
||||
). Accessory points slightly divergent. All claw bases distinctly narrowed, but especially the anterior claws, which have calyx-like bases (
|
||||
<figureCitation id="133D2A57A609FF82ACEB6936FD8D8F8B" box="[437,559,1389,1413]" captionStart="Figure 39" captionStartId="51.[113,176,1170,1194]" captionTargetBox="[114,1458,144,1142]" captionTargetId="figure-459@51.[114,1458,144,1142]" captionTargetPageId="51" captionText="Figure 39.Claws of Pilatobius cf.rugosus: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars,and white arrowhead indicates anterior bar.Scale bars are in micrometres, identical in panels A, B and in C, D." figureDoi="http://doi.org/10.5281/zenodo.14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="48" pageNumber="49">Fig. 39B, D</figureCitation>
|
||||
). Pseudolunulae absent. Cuticular bars present and of
|
||||
<specimenCount id="9D00FD5BA609FF82ACBC69D7FDFA8FAA" box="[482,600,1420,1445]" count="3" pageId="48" pageNumber="49" type="generic" typeStatus="types">three types</specimenCount>
|
||||
: (i) internal, long thickenings terminating at the claw I–III bases (
|
||||
<figureCitation id="133D2A57A609FF82AF2B69F0FD6F8FCB" box="[629,717,1451,1476]" captionStart="Figure 39" captionStartId="51.[113,176,1170,1194]" captionTargetBox="[114,1458,144,1142]" captionTargetId="figure-459@51.[114,1458,144,1142]" captionTargetPageId="51" captionText="Figure 39.Claws of Pilatobius cf.rugosus: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars,and white arrowhead indicates anterior bar.Scale bars are in micrometres, identical in panels A, B and in C, D." figureDoi="http://doi.org/10.5281/zenodo.14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="48" pageNumber="49">Fig. 39A</figureCitation>
|
||||
); (ii) anterior and (iii) posterior short rods (
|
||||
<figureCitation id="133D2A57A609FF82AF456990FDD08FEC" box="[539,626,1483,1507]" captionStart="Figure 39" captionStartId="51.[113,176,1170,1194]" captionTargetBox="[114,1458,144,1142]" captionTargetId="figure-459@51.[114,1458,144,1142]" captionTargetPageId="51" captionText="Figure 39.Claws of Pilatobius cf.rugosus: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars,and white arrowhead indicates anterior bar.Scale bars are in micrometres, identical in panels A, B and in C, D." figureDoi="http://doi.org/10.5281/zenodo.14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="48" pageNumber="49">Fig. 39B</figureCitation>
|
||||
). All cuticular bars are undetectable in SEM (
|
||||
<figureCitation id="133D2A57A609FF82ACE469B1FD8D8C0D" box="[442,559,1514,1538]" captionStart="Figure 39" captionStartId="51.[113,176,1170,1194]" captionTargetBox="[114,1458,144,1142]" captionTargetId="figure-459@51.[114,1458,144,1142]" captionTargetPageId="51" captionText="Figure 39.Claws of Pilatobius cf.rugosus: A, claws II (PCM); B, claws IV (PCM); C, claws III (SEM); D, claws IV (SEM). Black arrowheads indicate internal and posterior bars,and white arrowhead indicates anterior bar.Scale bars are in micrometres, identical in panels A, B and in C, D." figureDoi="http://doi.org/10.5281/zenodo.14243832" httpUri="https://zenodo.org/record/14243832/files/figure.png" pageId="48" pageNumber="49">Fig. 39C, D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A609FF82ADDF6A78FB5E8CB1" pageId="48" pageNumber="49" type="discussion">
|
||||
<paragraph id="8BB936D2A609FF82ADDF6A78FB5E8CB1" blockId="48.[128,778,1571,1971]" lastBlockId="48.[825,1476,856,1726]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B972EAC0A609FF82ADDF6A78FF408C34" box="[129,226,1571,1595]" italics="true" pageId="48" pageNumber="49">Remarks:</emphasis>
|
||||
Currently, there are two discriminative traits separating
|
||||
<taxonomicName id="4C064D51A609FF82ADE26A1FFEBD8C54" authorityName="Bartos" authorityYear="1935" box="[188,287,1604,1627]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82ADE26A1FFEBD8C54" box="[188,287,1604,1627]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A609FF82AC086A1FFD788C54" authority="(Schuster & Grigarick, 1965)" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[342,730,1603,1627]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A609FF82AC086A1FFE3B8C54" box="[342,409,1603,1627]" italics="true" pageId="48" pageNumber="49">P. iltisi</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82ACF46A18FD6F8C54" author="Schuster RO & Grigarick AA" box="[426,717,1603,1627]" pageId="48" pageNumber="49" pagination="1 - 67" refId="ref39541" refString="Schuster RO, Grigarick AA. Tardigrada from Western North America with emphasis on the fauna of California. University of California Publications in Zoology 1965; 76: 1 - 67." type="journal article" year="1965">Schuster & Grigarick, 1965</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
: the presence of deep transverse cuticular folds in
|
||||
<taxonomicName id="4C064D51A609FF82AF336A38FD128C75" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[621,688,1634,1658]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="iltisi">
|
||||
<emphasis id="B972EAC0A609FF82AF336A38FD128C75" box="[621,688,1634,1658]" italics="true" pageId="48" pageNumber="49">P. iltisi</emphasis>
|
||||
</taxonomicName>
|
||||
and, according to
|
||||
<bibRefCitation id="EF974B23A609FF82ADAA6AD9FE298C96" author="Bartos E" box="[244,395,1665,1690]" pageId="48" pageNumber="49" pagination="257 - 60" refId="ref34712" refString="Bartos E. Vier neue Hypsibius - Arten aus der Tschechoslovakei. Zoologischer Anzeiger 1935; 110: 257 - 60." type="journal article" year="1935">Bartoš (1935)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="EF974B23A609FF82ACE16ADAFCA18C96" author="Schuster RO & Grigarick AA" box="[447,771,1665,1689]" pageId="48" pageNumber="49" pagination="1 - 67" refId="ref39541" refString="Schuster RO, Grigarick AA. Tardigrada from Western North America with emphasis on the fauna of California. University of California Publications in Zoology 1965; 76: 1 - 67." type="journal article" year="1965">Schuster and Grigarick (1965)</bibRefCitation>
|
||||
, the diameter of the polygonal granules that constitute the sculpturing increasing posteriorly in
|
||||
<taxonomicName id="4C064D51A609FF82AF406A9AFD258CD7" authorityName="Bartos" authorityYear="1935" box="[542,647,1729,1752]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AF406A9AFD258CD7" box="[542,647,1729,1752]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
but not in
|
||||
<emphasis id="B972EAC0A609FF82ADDF6ABBFF698CF8" box="[129,203,1759,1783]" italics="true" pageId="48" pageNumber="49">
|
||||
<taxonomicName id="4C064D51A609FF82ADDF6ABBFF658CF8" baseAuthorityName="Schuster & Grigarick" baseAuthorityYear="1965" box="[129,199,1759,1783]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="iltisi">P. iltisi</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
These cuticular folds could be a result of the preparation technique [see ‘Remarks’ on
|
||||
<taxonomicName id="4C064D51A609FF82ACAB6B5BFD5B8D18" authority="(Ito, 1995)" authorityName="PCM" baseAuthorityName="Ito" baseAuthorityYear="1995" box="[501,761,1791,1815]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="sexbullatus">
|
||||
<emphasis id="B972EAC0A609FF82ACAB6B5BFDDF8D18" box="[501,637,1791,1815]" italics="true" pageId="48" pageNumber="49">P. sexbullatus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82AFD36AA4FD4F8D18" author="Ito M" box="[653,749,1791,1815]" pageId="48" pageNumber="49" pagination="18 - 39" refId="ref36771" refString="Ito M. Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan, II. Family Hypsibiidae. Proceedings of the Japanese Society of Systematic Zoology 1995; 53: 18 - 39." type="journal article" year="1995">Ito, 1995</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
]. The dorsal sculpturing in our examined population does not agree with the original description but conforms to specimens identified as
|
||||
<taxonomicName id="4C064D51A609FF82AC5B6B05FEC48D7A" authorityName="Bartos" authorityYear="1935" box="[261,358,1886,1909]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AC5B6B05FEC48D7A" box="[261,358,1886,1909]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EF974B23A609FF82ACD46B06FD908D7A" author="Dastych H" box="[394,562,1885,1909]" pageId="48" pageNumber="49" pagination="1 - 255" refId="ref35583" refString="Dastych H. The Tardigrada of Poland. Monografie Fauny Polski 1988; 16: 1 - 255." type="journal article" year="1988">Dastych (1988)</bibRefCitation>
|
||||
, thus indicating that resampling in the Turiec Basin is needed to verify the true morphotype of
|
||||
<taxonomicName id="4C064D51A609FF82AC786BC7FE2A8DBC" authorityName="Bartos" authorityYear="1935" box="[294,392,1948,1971]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AC786BC7FE2A8DBC" box="[294,392,1948,1971]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
. General claw morphology is similar in
|
||||
<taxonomicName id="4C064D51A609FF82AE066F02FC1D897F" authorityName="Bartos" authorityYear="1935" box="[856,959,857,880]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AE066F02FC1D897F" box="[856,959,857,880]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A609FF82AE8E6F02FAA5897F" authority="(Richters, 1911)" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[976,1287,856,880]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="recamieri">
|
||||
<emphasis id="B972EAC0A609FF82AE8E6F02FBEA897F" box="[976,1096,857,880]" italics="true" pageId="48" pageNumber="49">P. recamieri</emphasis>
|
||||
(Richters, 1911)
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EF974B23A609FF82A8166F03FC32899F" author="Gasiorek P & Zawierucha K & Stec D" pageId="48" pageNumber="49" pagination="2239 - 52" refId="ref36170" refString="Gasiorek P, Zawierucha K, Stec D et al. Integrative redescription of a common Arctic water bear Pilatobius recamieri (Richters, 1911). Polar Biology 2017; 40: 2239 - 52. https: // doi. org / 10.1007 / s 00300 - 017 - 2137 - 9" type="journal article" year="2017">
|
||||
Gąsiorek
|
||||
<emphasis id="B972EAC0A609FF82A8EF6F02FCF08980" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
), and
|
||||
<taxonomicName id="4C064D51A609FF82AE856F23FA20899F" authority="(Bertolani & Rebecchi, 1996)" baseAuthorityName="Bertolani & Rebecchi" baseAuthorityYear="1996" box="[987,1410,887,912]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="secchii">
|
||||
<emphasis id="B972EAC0A609FF82AE856F23FB978980" box="[987,1077,887,911]" italics="true" pageId="48" pageNumber="49">P. secchii</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A9176F2CFAD58980" author="Bertolani R & Rebecchi L" box="[1097,1399,887,912]" pageId="48" pageNumber="49" pagination="3 - 12" refId="ref34938" refString="Bertolani R, Rebecchi L. The tardigrades of Emilia (Italy). II. Monte Rondinaio. AmultihabitatstudyonahighaltitudevalleyoftheNorthern Apennines. Zoological Journal of the Linnean Society 1996; 116: 3 - 12. https: // doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02329. x" type="journal article" year="1996">Bertolani & Rebecchi, 1996</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
nom. inq. (
|
||||
<figureCitation id="133D2A57A609FF82AE2D6FCCFC6989A0" box="[883,971,919,943]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42E</figureCitation>
|
||||
). Sculpture of
|
||||
<taxonomicName id="4C064D51A609FF82A9336FC3FA0889A0" authority="(Robotti, 1970)" baseAuthorityName="Robotti" baseAuthorityYear="1970" box="[1133,1450,919,943]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="ramazzottii">
|
||||
<emphasis id="B972EAC0A609FF82A9336FC3FB5989A1" box="[1133,1275,919,943]" italics="true" pageId="48" pageNumber="49">P. ramazzottii</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A8556FCCFA3D89A0" author="Robotti C" box="[1291,1439,919,943]" pageId="48" pageNumber="49" pagination="251 - 5" refId="ref39407" refString="Robotti C. Hypsibius (D.) ramazzottii spec. nov. e Macrobiotus aviglianae spec. nov. Primo contributo alla conoscenza dei tardigradi del Piemonte. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 1970; 110: 251 - 5." type="journal article" year="1970">Robotti, 1970</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
is similar to that of
|
||||
<taxonomicName id="4C064D51A609FF82AEA86FECFBFA89C1" authorityName="Bartos" authorityYear="1935" box="[1014,1112,951,974]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AEA86FECFBFA89C1" box="[1014,1112,951,974]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
; however, the polygons are larger and closer to each other (
|
||||
<figureCitation id="133D2A57A609FF82A9146F8EFB0389E1" box="[1098,1185,981,1006]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42F</figureCitation>
|
||||
). Within the
|
||||
<taxonomicName id="4C064D51A609FF82A86D6F8DFADC89E2" authorityName="Bartos" authorityYear="1935" box="[1331,1406,982,1005]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82A86D6F8DFADC89E2" box="[1331,1406,982,1005]" italics="true" pageId="48" pageNumber="49">rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
group (species with the entire dorsum covered by sculpturing and lacking dorsal gibbosities), four other species are gathered:
|
||||
<taxonomicName id="4C064D51A609FF82AE67686FFBFC8E43" authority="(Greven, 1972)" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[825,1118,1076,1100]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A609FF82AE67686FFC078E43" box="[825,933,1076,1100]" italics="true" pageId="48" pageNumber="49">P. granifer</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82AEE3686FFBF18E43" author="Greven H" box="[957,1107,1076,1100]" pageId="48" pageNumber="49" pagination="368 - 81" refId="ref36334" refString="Greven H. Tardigraden des nordlichen Sauerlandes. Zoologischer Anzeiger 1972; 189: 368 - 81." type="journal article" year="1972">Greven, 1972</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A609FF82A92F686FFA3B8E43" authority="(Mihelcic, 1955)" baseAuthorityName="Mihelcic" baseAuthorityYear="1955" box="[1137,1433,1075,1100]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="latipes">
|
||||
<emphasis id="B972EAC0A609FF82A92F686FFB728E44" box="[1137,1232,1075,1099]" italics="true" pageId="48" pageNumber="49">P. latipes</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A9B66868FA2F8E44" author="Mihelcic F" box="[1256,1421,1075,1099]" pageId="48" pageNumber="49" pagination="240 - 4" refId="ref37915" refString="Mihelcic F. Zur Okologie und Verbreitung der Gattung Hypsibius (Tardigrada). Bonner Zoologische Beitrage 1955; 6: 240 - 4." type="journal article" year="1955">Mihelčič, 1955</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
sp. dub.,
|
||||
<taxonomicName id="4C064D51A609FF82AE2A680FFB108E64" authority="(Pilato et al., 2014)" baseAuthorityName="Pilato" baseAuthorityYear="2014" box="[884,1202,1107,1131]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="procerus">
|
||||
<emphasis id="B972EAC0A609FF82AE2A680FFC7D8E64" box="[884,991,1108,1131]" italics="true" pageId="48" pageNumber="49">P. procerus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82AEAF6808FB058E64" author="Pilato G & Sabella G & Lisi O" box="[1009,1191,1107,1131]" pageId="48" pageNumber="49" pagination="173 - 84" refId="ref39095" refString="Pilato G, Sabella G, Lisi O. Two new tardigrade species from Sicily. Zootaxa 2014; 3754: 173 - 84. https: // doi. org / 10.11646 / zootaxa. 3754.2.6" type="journal article" year="2014">
|
||||
Pilato
|
||||
<emphasis id="B972EAC0A609FF82A96B680FFBC48E64" box="[1077,1126,1107,1131]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
|
||||
, 2014
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A609FF82A9B3680FFCD98E85" authority="(Lisi et al., 2014)" baseAuthorityName="Lisi" baseAuthorityYear="2014" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="ziliense">
|
||||
<emphasis id="B972EAC0A609FF82A9B3680FFAEC8E64" box="[1261,1358,1107,1131]" italics="true" pageId="48" pageNumber="49">P. ziliense</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A8006808FCCC8E85" author="Lisi O & Sabella G & Pilato G" pageId="48" pageNumber="49" pagination="459 - 68" refId="ref37490" refString="Lisi O, Sabella G, Pilato G. Mixibius parvus sp. nov. and Diphascon (Diphascon) ziliense sp. nov., two new species of Eutardigrada from Sicily. Zootaxa 2014; 3802: 459 - 68. https: // doi. org / 10.11646 / zootaxa. 3802.4.3" type="journal article" year="2014">
|
||||
Lisi
|
||||
<emphasis id="B972EAC0A609FF82A8D2680FFA1F8E64" box="[1420,1469,1107,1131]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
|
||||
, 2014
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Of these,
|
||||
<bibRefCitation id="EF974B23A609FF82AEB96829FB2D8E85" author="Dastych H" box="[999,1167,1138,1162]" pageId="48" pageNumber="49" pagination="1 - 25" refId="ref35653" refString="Dastych H. Checklisten der Fauna Osterreichs, No. 8: Tardigrada. In: Winkler H, Stuessy T (eds.), Biosystematics and Ecology Series, No. 31. Osterreichischen Akademie der Wissenschaften, 2015, 1 - 25. https: // doi. org / 10.1553 / 0 x 00327 e 9 a" type="book chapter" year="2015">Dastych (2015)</bibRefCitation>
|
||||
has already expressed doubts regarding the validity of
|
||||
<taxonomicName id="4C064D51A609FF82A90D68C9FB138EA6" baseAuthorityName="Mihelcic" baseAuthorityYear="1955" box="[1107,1201,1169,1193]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="latipes">
|
||||
<emphasis id="B972EAC0A609FF82A90D68C9FB138EA6" box="[1107,1201,1169,1193]" italics="true" pageId="48" pageNumber="49">P. latipes</emphasis>
|
||||
</taxonomicName>
|
||||
, with which we concur.
|
||||
<taxonomicName id="4C064D51A609FF82AE6768E9FC008EC6" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[825,930,1201,1225]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A609FF82AE6768E9FC008EC6" box="[825,930,1201,1225]" italics="true" pageId="48" pageNumber="49">P. granifer</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C064D51A609FF82A96B68E9FB398EC6" authorityName="Bartos" authorityYear="1935" box="[1077,1179,1202,1225]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82A96B68E9FB398EC6" box="[1077,1179,1202,1225]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
by a completely dissimilar
|
||||
<typeStatus id="54BD8870A609FF82AE67688AFCCA8EE6" box="[825,872,1233,1257]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
of sculpturing [irregular small polygons in
|
||||
<taxonomicName id="4C064D51A609FF82A863688AFA038EE7" authorityName="Bartos" authorityYear="1935" box="[1341,1441,1233,1256]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82A863688AFA038EE7" box="[1341,1441,1233,1256]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
vs. large polygons resembling the caudal sculpturing of some species of the
|
||||
<taxonomicName id="4C064D51A609FF82AEB46954FA608F28" authority="(Doyere, 1840)" baseAuthorityName="Doyere" baseAuthorityYear="1840" box="[1002,1474,1295,1319]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||||
<emphasis id="B972EAC0A609FF82AEB46954FAA68F28" box="[1002,1284,1295,1319]" italics="true" pageId="48" pageNumber="49">Ramazzottius oberhaeuseri</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A609FF82A8416954FA148F28" author="Doyere ML" box="[1311,1462,1295,1319]" pageId="48" pageNumber="49" pagination="269 - 362" refId="ref35945" refString="Doyere ML. Memorie sur les Tardigrades. Annales des sciences naturelles Paris 1840; 14: 269 - 362." type="journal article" year="1840">Doyère, 1840</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
complex, but clearly larger and with a conspicuous light centre in
|
||||
<taxonomicName id="4C064D51A609FF82AE086915FC1F8F69" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[854,957,1358,1382]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A609FF82AE086915FC1F8F69" box="[854,957,1358,1382]" italics="true" pageId="48" pageNumber="49">P. granifer</emphasis>
|
||||
</taxonomicName>
|
||||
; see
|
||||
<figureCitation id="133D2A57A609FF82AEA96916FBF68F69" box="[1015,1108,1357,1382]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42G</figureCitation>
|
||||
]. Unfortunately, other differences cannot be given, because the description of
|
||||
<taxonomicName id="4C064D51A609FF82A85B6935FAC88F8A" baseAuthorityName="Greven" baseAuthorityYear="1972" box="[1285,1386,1389,1413]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="granifer">
|
||||
<emphasis id="B972EAC0A609FF82A85B6935FAC88F8A" box="[1285,1386,1389,1413]" italics="true" pageId="48" pageNumber="49">P. granifer</emphasis>
|
||||
</taxonomicName>
|
||||
is insufficient, and the
|
||||
<typeStatus id="54BD8870A609FF82AEBC69D6FBB38FAA" box="[994,1041,1421,1445]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
material lost (H. Greven, pers. comm.). In our opinion, both
|
||||
<taxonomicName id="4C064D51A609FF82A94169F7FB2E8FCC" box="[1055,1164,1452,1475]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="procerus">
|
||||
<emphasis id="B972EAC0A609FF82A94169F7FB2E8FCC" box="[1055,1164,1452,1475]" italics="true" pageId="48" pageNumber="49">P. procerus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A609FF82A99C69F7FA868FCC" box="[1218,1316,1451,1475]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="ziliense">
|
||||
<emphasis id="B972EAC0A609FF82A99C69F7FA868FCC" box="[1218,1316,1451,1475]" italics="true" pageId="48" pageNumber="49">P. ziliense</emphasis>
|
||||
</taxonomicName>
|
||||
are poorly differentiated from
|
||||
<taxonomicName id="4C064D51A609FF82AEAE6997FBF78FEC" authorityName="Bartos" authorityYear="1935" box="[1008,1109,1484,1507]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82AEAE6997FBF78FEC" box="[1008,1109,1484,1507]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
and require further investigation.
|
||||
<taxonomicName id="4C064D51A609FF82AE6769B0FC148C0D" authorityName="CNHM" baseAuthorityName="Ramazzotti" baseAuthorityYear="1957" box="[825,950,1514,1538]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="nodulosus">
|
||||
<emphasis id="B972EAC0A609FF82AE6769B0FC148C0D" box="[825,950,1514,1538]" italics="true" pageId="48" pageNumber="49">P. nodulosus</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from
|
||||
<taxonomicName id="4C064D51A609FF82A9F569B0FAAD8C0D" authorityName="Bartos" authorityYear="1935" box="[1195,1295,1515,1538]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="48" pageNumber="49" phylum="Tardigrada" rank="species" species="rugosus">
|
||||
<emphasis id="B972EAC0A609FF82A9F569B0FAAD8C0D" box="[1195,1295,1515,1538]" italics="true" pageId="48" pageNumber="49">P. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
by its dissimilar
|
||||
<typeStatus id="54BD8870A609FF82AE676A51FCCA8C2D" box="[825,872,1546,1570]" pageId="48" pageNumber="49">type</typeStatus>
|
||||
of sculpturing: very large polygons in the caudal zone (
|
||||
<figureCitation id="133D2A57A609FF82AE1A6A72FC0B8C4E" box="[836,937,1577,1601]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42D</figureCitation>
|
||||
), with the size of the polygons subsequently decreasing anteriorly. The internal and anterior claws of the former species are also more massive (
|
||||
<figureCitation id="133D2A57A609FF82A9AF6A3CFAD78C70" box="[1265,1397,1639,1664]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42C, D</figureCitation>
|
||||
), rectangular bars are present below claws (
|
||||
<figureCitation id="133D2A57A609FF82A9956ADCFAE68C91" box="[1227,1348,1671,1695]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42C, D</figureCitation>
|
||||
), and claws IV have evident pseudolunulae (
|
||||
<figureCitation id="133D2A57A609FF82A9CE6AFDFB498CB1" box="[1168,1259,1702,1726]" captionStart="Figure 42" captionStartId="54.[129,194,1843,1867]" captionTargetBox="[254,1348,144,1815]" captionTargetId="figure-5@54.[254,1348,144,1815]" captionTargetPageId="54" captionText="Figure 42. Various Pilatobiinae, morphological details (PCM): A, P.opisthoglyptus (Maucci, 1987), paratype, buccopharyngeal apparatus in toto (empty white arrowhead indicates septulum); B, P.opisthoglyptus, paratype, claws I; C, P.nodulosus (Ramazzotti, 1957), claws I; D, P.nodulosus, claws IV; E, P.secchii (Bertolani & Rebecchi, 1996) nom.inq., claws II; F, P.ramazzottii (Robotti, 1970), caudal sculpturing; G, P.granifer (Greven, 1972), paratype, cephalic sculpturing (scale unknown). Filled incised white arrowheads indicate bars at the limb bases. Scale bars:10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243838" httpUri="https://zenodo.org/record/14243838/files/figure.png" pageId="48" pageNumber="49">Fig. 42D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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<mods:title id="913FBECC53DECA5DE9F6121D6944965E">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="0A6ED2767E9B47A567A8846F5A18E186">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="4037AF0A7584D40FAC318DB545E1B5FA">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="09B2CFE899870238C936FD3EDE1CC149" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AC72B46A2C74C93C84CDEBA485239E21">Blagden, Brian</mods:namePart>
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<mods:affiliation id="D5916B0D33E9360C3E0C62D647BEEA76">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="FE26721F8271B2DF0326E7E67FBF8998">Morek, Witold</mods:namePart>
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<mods:affiliation id="A8897947B7A089BB2CB36637B60D6AEC">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="C5138809701D29C7E185814C604658D3">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="042409A5C85720A61D253619AD9C44AC">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:date id="8F039F4874778192D9EE5B5C0E8DF827">2024</mods:date>
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<taxonomicName id="4C064D51A61AFF91ADCF686BFD288E45" authority="Gasiorek & Michalczyk" authorityName="Gasiorek & Michalczyk" authorityYear="2024" box="[145,650,1072,1099]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="35" pageNumber="36" phylum="Tardigrada" rank="species" species="minutissimus" status="sp. nov.">
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<emphasis id="B972EAC0A61AFF91ADCF686BFE2C8E45" bold="true" box="[145,398,1072,1099]" italics="true" pageId="35" pageNumber="36">Raribius minutissimus</emphasis>
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Gąsiorek & Michalczyk
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|
||||
<taxonomicNameLabel id="A24157BBA61AFF91AFD0686BFD788E45" box="[654,730,1072,1098]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<emphasis id="B972EAC0A61AFF91AD286835FD578E86" bold="true" box="[118,757,1134,1161]" pageId="35" pageNumber="36">
|
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<taxonomicName id="4C064D51A61AFF91AD286835FF6F8E86" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[118,205,1134,1161]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="35" pageNumber="36" phylum="Tardigrada" rank="genus" status="sp. nov.">
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<emphasis id="B972EAC0A61AFF91AD286835FF6F8E86" bold="true" box="[118,205,1134,1161]" italics="true" pageId="35" pageNumber="36">Raribius</emphasis>
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</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA61AFF91AD8C6834FE838E86" box="[210,289,1135,1161]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
;
|
||||
<collectingCountry id="F3117642A61AFF91AC746835FE288E87" box="[298,394,1134,1160]" name="United Kingdom" pageId="35" pageNumber="36">Scotland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A61AFF91ACCA6835FD528E87" author="Gasiorek P & Michalczyk L" box="[404,752,1134,1161]" pageId="35" pageNumber="36" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A61AFF91AD2F68CEFE4A8EC3" blockId="35.[113,762,1134,1228]" pageId="35" pageNumber="36">
|
||||
ZooBank registration:
|
||||
<uri id="FF973AD0A61AFF91AC2968CEFE4A8EC3" pageId="35" pageNumber="36">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFA00C07A61AFF91AF2A68CEFE4A8EC3" pageId="35" pageNumber="36">EB412AD2- 2AD0-4F57-BA97-07D3D287D33B</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A61AFF91AD2F68B5FC618F72" pageId="35" pageNumber="36" type="materials_examined">
|
||||
<paragraph id="8BB936D2A61AFF91AD2F68B5FE788F2A" blockId="35.[113,762,1262,1317]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B972EAC0A61AFF91AD2F68B5FE9B8F09" box="[113,313,1262,1286]" italics="true" pageId="35" pageNumber="36">Material examined:</emphasis>
|
||||
Six individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A61AFF91AC2C6956FE688F2A" box="[370,458,1293,1317]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="35" pageNumber="36">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A61AFF91AD2F691CFBAF8E88" blockId="35.[113,763,1350,1406]" lastBlockId="35.[810,1461,946,1159]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B972EAC0A61AFF91AD2F691CFEA18F51" box="[113,259,1350,1374]" italics="true" pageId="35" pageNumber="36">Type material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA61AFF91AC52691DFBAF8E88" collectingDate="2016-04" collectingDateMax="2016-09-31" collectingDateMin="2016-04" collectionCode="GB" collectorName="Blagden" country="United Kingdom" elevation="114" latitude="57.094997" location="Drum Castle" longLatPrecision="17" longitude="-2.3377776" municipality="Drumoak" pageId="35" pageNumber="36" specimenCode="GB.043.10, GB.043.08-9, GB.059.01-2" specimenCount="2" typeStatus="holotype">
|
||||
<typeStatus id="54BD8870A61AFF91AC52691DFECD8F51" box="[268,367,1350,1374]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
|
||||
(slide
|
||||
<specimenCode id="DBA09EA9A61AFF91ACE4691CFD8E8F50" box="[442,556,1351,1375]" pageId="35" pageNumber="36">GB.043.10</specimenCode>
|
||||
) and
|
||||
<specimenCount id="9D00FD5BA61AFF91AF35691DFD2D8F51" box="[619,655,1350,1374]" pageId="35" pageNumber="36">five</specimenCount>
|
||||
<typeStatus id="54BD8870A61AFF91AFC6691CFD598F50" box="[664,763,1351,1375]" pageId="35" pageNumber="36">paratypes</typeStatus>
|
||||
(slides
|
||||
<specimenCode id="DBA09EA9A61AFF91AD9F693DFEEE8F71" box="[193,332,1382,1406]" pageId="35" pageNumber="36">GB.043.08–9</specimenCode>
|
||||
, 11, and
|
||||
<specimenCode id="DBA09EA9A61AFF91ACE5693DFDEA8F71" box="[443,584,1382,1406]" pageId="35" pageNumber="36">GB.059.01–2</specimenCode>
|
||||
) from
|
||||
<collectingCountry id="F3117642A61AFF91AFC5693DFD558F71" box="[667,759,1382,1406]" name="United Kingdom" pageId="35" pageNumber="36">Scotland</collectingCountry>
|
||||
,
|
||||
<collectingMunicipality id="6BDDACA8A61AFF91AE746FE8FC3289C4" box="[810,912,947,971]" pageId="35" pageNumber="36">Drumoak</collectingMunicipality>
|
||||
,
|
||||
<location id="8ED96009A61AFF91AEC26FEFFB8189C4" LSID="urn:lsid:plazi:treatment:03AF87C4A61AFF97ADCF686BFBC28DB9:8ED96009A61AFF91AEC26FEFFB8189C4" box="[924,1059,947,971]" country="United Kingdom" latitude="57.094997" longLatPrecision="17" longitude="-2.3377776" municipality="Drumoak" name="Drum Castle" pageId="35" pageNumber="36">Drum Castle</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A61AFF91A96D6FE8FB0089C4" box="[1075,1186,946,971]" degrees="57" direction="north" minutes="5" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="42" value="57.094997">
|
||||
57°5
|
||||
<emphasis id="B972EAC0A61AFF91A93B6FE9FBC989C4" box="[1125,1131,946,971]" italics="true" pageId="35" pageNumber="36">ʹ</emphasis>
|
||||
42″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A61AFF91A9F06FEFFABC89C4" box="[1198,1310,946,972]" degrees="2" direction="west" minutes="20" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="16" value="-2.3377776">
|
||||
2°20
|
||||
<emphasis id="B972EAC0A61AFF91A9836FE9FB4189C4" box="[1245,1251,946,971]" italics="true" pageId="35" pageNumber="36">ʹ</emphasis>
|
||||
16″W
|
||||
</geoCoordinate>
|
||||
,
|
||||
<quantity id="4CFE9B37A61AFF91A8756FEFFACD89C4" box="[1323,1391,948,972]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="35" pageNumber="36" unit="m" value="114.0">
|
||||
<elevation id="002BD1E1A61AFF91A8756FEFFACD89C4" box="[1323,1391,948,972]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="35" pageNumber="36" unit="m" value="114.0">114 m</elevation>
|
||||
</quantity>
|
||||
a.s.l.), leaf litter (mixed
|
||||
<taxonomicName id="4C064D51A61AFF91AE866F88FBC089E4" box="[984,1122,979,1003]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="robur">
|
||||
<emphasis id="B972EAC0A61AFF91AE866F88FBC089E4" box="[984,1122,979,1003]" italics="true" pageId="35" pageNumber="36">Quercus robur</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C064D51A61AFF91A9DE6F88FAAD89E4" box="[1152,1295,979,1003]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="B972EAC0A61AFF91A9DE6F88FAAD89E4" box="[1152,1295,979,1003]" italics="true" pageId="35" pageNumber="36">Rhododendron</emphasis>
|
||||
</taxonomicName>
|
||||
) collected from soil,
|
||||
<date id="FFB81012A61AFF91AE056FA9FBD38E05" box="[859,1137,1010,1034]" pageId="35" pageNumber="36" value="2016-04" valueMax="2016-09-31" valueMin="2016-04-00">
|
||||
<collectingDate id="EFFCE9FAA61AFF91AE056FA9FBD38E05" box="[859,1137,1010,1034]" pageId="35" pageNumber="36" value="2016-04" valueMax="2016-09-31" valueMin="2016-04-00">April and September 2016</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A61AFF91A9226FA9FB6D8E05" box="[1148,1231,1010,1034]" pageId="35" pageNumber="36">Blagden</collectorName>
|
||||
coll., are deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University, Kraków. Hologenophores were retrieved in a good condition and included in the type material owing to an overall paucity of individuals.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A61AFF91AE7468F2FC618F72" blockId="35.[810,1459,1193,1405]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B972EAC0A61AFF91AE7468F2FBB18ECE" box="[810,1043,1193,1217]" italics="true" pageId="35" pageNumber="36">Comparative material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA61AFF91A97B68F2FB5F8F10" collectingDate="1971-02" country="Poland" county="Tatra Mountains" location="Polana pod Woloszynem" municipality="Woloszynem" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Tatra Mountains" typeStatus="paratype">
|
||||
<taxonomicName id="4C064D51A61AFF91A97B68F2FB548ECE" authorityName="NEC" baseAuthorityName="Weglarska" baseAuthorityYear="1973" box="[1061,1270,1193,1217]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="35" pageNumber="36" phylum="Tardigrada" rank="species" species="pawlowskii">
|
||||
<emphasis id="B972EAC0A61AFF91A97B68F2FB548ECE" box="[1061,1270,1193,1217]" italics="true" pageId="35" pageNumber="36">Raribius pawlowskii</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<collectingCountry id="F3117642A61AFF91A85168F2FAFB8ECE" box="[1295,1369,1193,1217]" name="Poland" pageId="35" pageNumber="36">Poland</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A61AFF91A83068F2FC408EEF" LSID="urn:lsid:plazi:treatment:03AF87C4A61AFF97ADCF686BFBC28DB9:8ED96009A61AFF91A83068F2FC408EEF" country="Poland" county="Tatra Mountains" municipality="Woloszynem" name="Polana pod Woloszynem" pageId="35" pageNumber="36" stateProvince="Tatra Mountains">Polana pod Wołoszynem</location>
|
||||
,
|
||||
<collectingRegion id="49C2F830A61AFF91AEB06892FB3E8EEF" box="[1006,1180,1224,1248]" pageId="35" pageNumber="36">Tatra Mountains</collectingRegion>
|
||||
, lower forest zone, mosses from rocks,
|
||||
<date id="FFB81012A61AFF91AEF768BCFBE18F0F" box="[937,1091,1255,1280]" pageId="35" pageNumber="36" value="1971-02">
|
||||
<collectingDate id="EFFCE9FAA61AFF91AEF768BCFBE18F0F" box="[937,1091,1255,1280]" pageId="35" pageNumber="36" value="1971-02">February 1971</collectingDate>
|
||||
</date>
|
||||
, a
|
||||
<typeStatus id="54BD8870A61AFF91A93C68B3FB198F0F" box="[1122,1211,1256,1280]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
|
||||
in the collection of the Zoological Museum, Jagiellonian University
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA61AFF91A852695CFC1D8F72" country="Italy" county="University of Modena" location="Treviso province" municipality="Department of Life Sciences" pageId="35" pageNumber="36" specimenCount="1" stateProvince="vicinity of Rugolo" typeStatus="paratype">
|
||||
<collectingCountry id="F3117642A61AFF91A852695CFA9B8F10" box="[1292,1337,1287,1311]" name="Italy" pageId="35" pageNumber="36">Italy</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49C2F830A61AFF91A819695CFCD48F31" pageId="35" pageNumber="36">vicinity of Rugolo</collectingRegion>
|
||||
,
|
||||
<location id="8ED96009A61AFF91AEDC697DFB938F31" LSID="urn:lsid:plazi:treatment:03AF87C4A61AFF97ADCF686BFBC28DB9:8ED96009A61AFF91AEDC697DFB938F31" box="[898,1073,1318,1342]" country="Italy" county="University of Modena" municipality="Department of Life Sciences" name="Treviso province" pageId="35" pageNumber="36" stateProvince="vicinity of Rugolo">Treviso province</location>
|
||||
, detailed data unspecified, collection of the Department of Life Sciences, University of Modena and Reggio Emilia
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF79665AA61AFF91AD2F6BF9FAB68DB4" ID-DOI="http://doi.org/10.5281/zenodo.14243801" ID-Zenodo-Dep="14243801" box="[113,1300,1954,1979]" httpUri="https://zenodo.org/record/14243801/files/figure.png" pageId="35" pageNumber="36" startId="35.[113,178,1954,1978]" targetBox="[114,1458,1456,1927]" targetPageId="35" targetType="figure">
|
||||
<paragraph id="8BB936D2A61AFF91AD2F6BF9FAB68DB4" blockId="35.[113,1300,1954,1979]" box="[113,1300,1954,1979]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B972EAC0A61AFF91AD2F6BF9FF748DB4" bold="true" box="[113,214,1954,1979]" pageId="35" pageNumber="36">Figure 24.</emphasis>
|
||||
General morphology of
|
||||
<taxonomicName id="4C064D51A61AFF91ACE06BF8FDDF8DB4" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[446,637,1955,1979]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="35" pageNumber="36" phylum="Tardigrada" rank="species" species="carpathica" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A61AFF91ACE06BF8FDDF8DB4" box="[446,637,1955,1979]" italics="true" pageId="35" pageNumber="36">Platicrista carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A61AFF91AFDC6BF8FD6F8DB4" bold="true" box="[642,717,1955,1979]" pageId="35" pageNumber="36">
|
||||
<taxonomicNameLabel id="A24157BBA61AFF91AFDC6BF8FD6F8DB4" box="[642,717,1955,1979]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype in ventral view, PCM). Scale bar is in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA61DFF96ADDF6E9CFBD288D0" ID-DOI="http://doi.org/10.5281/zenodo.14243804" ID-Zenodo-Dep="14243804" box="[129,1136,711,736]" httpUri="https://zenodo.org/record/14243804/files/figure.png" pageId="36" pageNumber="37" startId="36.[129,194,711,735]" targetBox="[129,1473,144,684]" targetPageId="36" targetType="figure">
|
||||
<paragraph id="8BB936D2A61DFF96ADDF6E9CFBD288D0" blockId="36.[129,1136,711,736]" box="[129,1136,711,736]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B972EAC0A61DFF96ADDF6E9CFF4788D0" bold="true" box="[129,229,711,735]" pageId="36" pageNumber="37">Figure 25.</emphasis>
|
||||
Caudal wrinkling of
|
||||
<taxonomicName id="4C064D51A61DFF96ACF76E9CFDCB88D0" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[425,617,711,735]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="36" pageNumber="37" phylum="Tardigrada" rank="species" species="carpathica" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A61DFF96ACF76E9CFDCB88D0" box="[425,617,711,735]" italics="true" pageId="36" pageNumber="37">Platicrista carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A61DFF96AF306E93FD1B88EF" bold="true" box="[622,697,712,736]" pageId="36" pageNumber="37">
|
||||
<taxonomicNameLabel id="A24157BBA61DFF96AF306E93FD1B88EF" box="[622,697,712,736]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(paratype, PCM). Scale bar is in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA61DFF96ADDF6ADDFE578CD9" ID-DOI="http://doi.org/10.5281/zenodo.14243806" ID-Zenodo-Dep="14243806" httpUri="https://zenodo.org/record/14243806/files/figure.png" pageId="36" pageNumber="37" startId="36.[129,194,1670,1694]" targetBox="[131,1470,789,1638]" targetPageId="36" targetType="figure">
|
||||
<paragraph id="8BB936D2A61DFF96ADDF6ADDFE578CD9" blockId="36.[128,1474,1670,1750]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B972EAC0A61DFF96ADDF6ADDFF478C91" bold="true" box="[129,229,1670,1694]" pageId="36" pageNumber="37">Figure 26.</emphasis>
|
||||
Detailed morphology of
|
||||
<taxonomicName id="4C064D51A61DFF96AC8C6ADDFD308C91" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[466,658,1670,1694]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="36" pageNumber="37" phylum="Tardigrada" rank="species" species="carpathica" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A61DFF96AC8C6ADDFD308C91" box="[466,658,1670,1694]" italics="true" pageId="36" pageNumber="37">Platicrista carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A61DFF96AFC96ADDFD408C91" bold="true" box="[663,738,1670,1694]" pageId="36" pageNumber="37">
|
||||
<taxonomicNameLabel id="A24157BBA61DFF96AFC96ADDFD408C91" box="[663,738,1670,1694]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(paratypes, PCM): A, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A61DFF96A99D6ADCFB5E8C91" box="[1219,1276,1671,1694]" italics="true" pageId="36" pageNumber="37">in toto</emphasis>
|
||||
; B, claws I; C, claws II; D, claws III. Empty white arrowheads indicate pseudolunulae; white asterisk indicates internal pulvinus; and black asterisk indicates external pulvinus. Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C31C6559A61DFF97ADDF6B5DFE118D36" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
|
||||
<paragraph id="8BB936D2A61DFF96ADDF6B5DFC478DB5" blockId="36.[129,779,1798,1978]" lastBlockId="36.[825,1474,1798,1978]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B972EAC0A61DFF96ADDF6B5DFF558D12" box="[129,247,1798,1821]" italics="true" pageId="36" pageNumber="37">Description:</emphasis>
|
||||
Body small (
|
||||
<tableCitation id="C6840369A61DFF96ACD46B5DFE448D11" box="[394,486,1798,1822]" captionStart="Table 11" captionStartId="41.[113,168,143,167]" captionTargetPageId="41" captionText="Table 11. Measurements (in micrometres) of selected morphological structures of type series of Raribius minutissimus sp. nov. from Scotland (GB.043 + GB.059) mounted in Hoyer’s medium." pageId="36" pageNumber="37">Table 11</tableCitation>
|
||||
), elongate and slightly narrowed in its anterior part (
|
||||
<figureCitation id="133D2A57A61DFF96ACCC6B7EFE4B8D32" box="[402,489,1829,1853]" captionStart="Figure 31" captionStartId="42.[129,194,976,1000]" captionTargetBox="[129,1473,144,949]" captionTargetId="figure-5@42.[129,1473,144,949]" captionTargetPageId="42" captionText="Figure 31. Detailed morphology of Raribius minutissimus sp. nov. (PCM): A, holotype in ventral view; B, buccopharyngeal apparatus in toto; C, claws IV.Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243816" httpUri="https://zenodo.org/record/14243816/files/figure.png" pageId="36" pageNumber="37">Fig. 31A</figureCitation>
|
||||
). Head blunt. Body whitish to almost translucent. Cuticle smooth, without pores, wrinkling, or granulation. Cribriform areas not visible under PCM. Legs very short and weakly delimited from the trunk. Eyes absent in live animals. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A61DFF96AFE96BF9FCA98DB5" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[695,779,1954,1978]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="36" pageNumber="37" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A61DFF96AFE96BF9FCA98DB5" box="[695,779,1954,1978]" italics="true" pageId="36" pageNumber="37">Raribius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A61DFF96AE676B5DFCC58D11" box="[825,871,1798,1822]" pageId="36" pageNumber="37">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A61DFF96AE256B5DFC738D11" box="[891,977,1798,1822]" captionStart="Figure 31" captionStartId="42.[129,194,976,1000]" captionTargetBox="[129,1473,144,949]" captionTargetId="figure-5@42.[129,1473,144,949]" captionTargetPageId="42" captionText="Figure 31. Detailed morphology of Raribius minutissimus sp. nov. (PCM): A, holotype in ventral view; B, buccopharyngeal apparatus in toto; C, claws IV.Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243816" httpUri="https://zenodo.org/record/14243816/files/figure.png" pageId="36" pageNumber="37">Fig. 31B</figureCitation>
|
||||
). The OCA not visible under PCM. Furcae of the
|
||||
<taxonomicName id="4C064D51A61DFF96AE3E6B7DFC688D32" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[864,970,1830,1853]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="36" pageNumber="37" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A61DFF96AE3E6B7DFC688D32" box="[864,970,1830,1853]" italics="true" pageId="36" pageNumber="37">Itaquascon</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A61DFF96AE916B7DFC5C8D31" box="[975,1022,1830,1854]" pageId="36" pageNumber="37">type</typeStatus>
|
||||
, with triangular, large apices. Stylet supports S-shaped. Pharyngeal apophyses absent; instead, the pharyngeal tube is slightly enlarged at its posterior end. Pharynx almost perfectly round, with a single rectangular macroplacoid with smooth margins.
|
||||
</paragraph>
|
||||
<caption id="DF79665AA61CFF97AD2F6858FF528E5B" ID-DOI="http://doi.org/10.5281/zenodo.14243808" ID-Zenodo-Dep="14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="37" pageNumber="38" startId="37.[113,178,1027,1051]" targetBox="[116,1455,146,996]" targetPageId="37" targetType="figure">
|
||||
<paragraph id="8BB936D2A61CFF97AD2F6858FF528E5B" blockId="37.[113,1450,1027,1108]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B972EAC0A61CFF97AD2F6858FF748E14" bold="true" box="[113,214,1027,1051]" pageId="37" pageNumber="38">Figure 27.</emphasis>
|
||||
Detailed morphology of
|
||||
<taxonomicName id="4C064D51A61CFF97AC9D685FFC058E13" authority="Kaczmarek & Michalczyk, 2003" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" box="[451,935,1028,1052]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A61CFF97AC9D685FFDD48E13" box="[451,630,1028,1052]" italics="true" pageId="37" pageNumber="38">Platicrista horribilis</emphasis>
|
||||
<bibRefCitation id="EF974B23A61CFF97AF24685FFC058E13" author="Kaczmarek L & Michalczyk L" box="[634,935,1028,1052]" pageId="37" pageNumber="38" pagination="1 - 8" refId="ref36912" refString="Kaczmarek L, Michalczyk L. Platicrista horribilis, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Mongolia. Zootaxa 2003; 132: 1 - 8. https: // doi. org / 10.11646 / zootaxa. 132.1.1" type="journal article" year="2003">Kaczmarek & Michalczyk, 2003</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(PCM): A, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A61CFF97A87C685FFAFE8E14" box="[1314,1372,1028,1051]" italics="true" pageId="37" pageNumber="38">in toto</emphasis>
|
||||
; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA61CFF97AD2F6AD1FB468CAD" ID-DOI="http://doi.org/10.5281/zenodo.14243810" ID-Zenodo-Dep="14243810" box="[113,1252,1674,1698]" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="37" pageNumber="38" startId="37.[113,178,1674,1698]" targetBox="[117,1455,1148,1642]" targetPageId="37" targetType="figure">
|
||||
<paragraph id="8BB936D2A61CFF97AD2F6AD1FB468CAD" blockId="37.[113,1252,1674,1698]" box="[113,1252,1674,1698]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B972EAC0A61CFF97AD2F6AD1FF748CAD" bold="true" box="[113,214,1674,1698]" pageId="37" pageNumber="38">Figure 28.</emphasis>
|
||||
General morphology of
|
||||
<taxonomicName id="4C064D51A61CFF97ACE06AD1FDEF8CAD" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[446,589,1674,1698]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="nivea" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A61CFF97ACE06AD1FDEF8CAD" box="[446,589,1674,1698]" italics="true" pageId="37" pageNumber="38">Platicrista nivea</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A61CFF97AF0C6AD1FD3F8CAD" bold="true" box="[594,669,1674,1698]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A24157BBA61CFF97AF0C6AD1FD3F8CAD" box="[594,669,1674,1698]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype in ventral view, PCM). Scale bar is in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A61CFF97ADD36AB8FE118D36" blockId="37.[112,763,1763,1849]" pageId="37" pageNumber="38">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A61CFF97AC486AB8FED48CF4" authorityName="Ehrenberg" authorityYear="1848" box="[278,374,1763,1787]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A61CFF97AC486AB8FED48CF4" box="[278,374,1763,1787]" italics="true" pageId="37" pageNumber="38">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
type, minute and slender, with accessory points not visible under PCM (
|
||||
<figureCitation id="133D2A57A61CFF97AC846B59FDE98D15" box="[474,587,1794,1818]" captionStart="Figure 31" captionStartId="42.[129,194,976,1000]" captionTargetBox="[129,1473,144,949]" captionTargetId="figure-5@42.[129,1473,144,949]" captionTargetPageId="42" captionText="Figure 31. Detailed morphology of Raribius minutissimus sp. nov. (PCM): A, holotype in ventral view; B, buccopharyngeal apparatus in toto; C, claws IV.Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243816" httpUri="https://zenodo.org/record/14243816/files/figure.png" pageId="37" pageNumber="38">Fig. 31A, C</figureCitation>
|
||||
). Pseudolunulae, cuticular bars, and septa absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A61CFF97AD2F6B00FF508DBE" pageId="37" pageNumber="38" type="etymology">
|
||||
<paragraph id="8BB936D2A61CFF97AD2F6B00FF508DBE" blockId="37.[113,762,1883,1969]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B972EAC0A61CFF97AD2F6B00FF438D7C" box="[113,225,1883,1907]" italics="true" pageId="37" pageNumber="38">Etymology:</emphasis>
|
||||
From the Latin
|
||||
<taxonomicName id="4C064D51A61CFF97ACDB6B07FDA68D7C" authorityName="Gasiorek & Michalczyk" authorityYear="2024" box="[389,516,1884,1907]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="minutissimus">
|
||||
<emphasis id="B972EAC0A61CFF97ACDB6B07FDA68D7C" box="[389,516,1884,1907]" italics="true" pageId="37" pageNumber="38">minutissimus</emphasis>
|
||||
</taxonomicName>
|
||||
= the smallest. The species is one of the smallest eutardigrades. An adjective in nominative singular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A61CFF97AE746AB8FBC28DB9" pageId="37" pageNumber="38" type="diagnosis">
|
||||
<paragraph id="8BB936D2A61CFF97AE746AB8FBC28DB9" blockId="37.[810,1459,1763,1975]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B972EAC0A61CFF97AE746AB8FAA68CF4" box="[810,1284,1763,1787]" italics="true" pageId="37" pageNumber="38">
|
||||
Differential diagnosis:
|
||||
<taxonomicName id="4C064D51A61CFF97A9456AB8FAA68CF4" authorityName="Gasiorek & Michalczyk" authorityYear="2024" box="[1051,1284,1763,1787]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="minutissimus">Raribius minutissimus</taxonomicName>
|
||||
</emphasis>
|
||||
has short rectangular macroplacoids, whereas the pharynx is devoid of placoids in
|
||||
<taxonomicName id="4C064D51A61CFF97AEF86B79FB918D36" authorityName="NEC" baseAuthorityName="Weglarska" baseAuthorityYear="1973" box="[934,1075,1825,1849]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="pawlowskii">
|
||||
<emphasis id="B972EAC0A61CFF97AEF86B79FB918D36" box="[934,1075,1825,1849]" italics="true" pageId="37" pageNumber="38">R. pawlowskii</emphasis>
|
||||
</taxonomicName>
|
||||
. No difference was detected in claw morphology (
|
||||
<figureCitation id="133D2A57A61CFF97AEE16B1AFBA58D56" box="[959,1031,1857,1881]" captionStart="Figure 32" captionStartId="42.[129,194,1944,1968]" captionTargetBox="[129,1473,1072,1916]" captionTargetId="figure-35@42.[129,1473,1072,1916]" captionTargetPageId="42" captionText="Figure 32. Raribius pawlowskii (Węglarska, 1973) (PCM): A, claws II (an individual from Rugalo, Italy); B, claws III (an individual from Rugalo, Italy); C, claws IV (paratype). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243818" httpUri="https://zenodo.org/record/14243818/files/figure.png" pageId="37" pageNumber="38">Fig. 32</figureCitation>
|
||||
).
|
||||
<taxonomicName id="4C064D51A61CFF97A9406B1AFA118D56" authority="(Abe & Ito, 1994)" baseAuthorityName="Abe & Ito" baseAuthorityYear="1994" box="[1054,1459,1857,1881]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="globuliferus">
|
||||
<emphasis id="B972EAC0A61CFF97A9406B1AFB4D8D56" box="[1054,1263,1857,1881]" italics="true" pageId="37" pageNumber="38">Raribius globuliferus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A61CFF97A85E6B1AFA0A8D56" author="Abe W & Ito M" box="[1280,1448,1857,1881]" pageId="37" pageNumber="38" pagination="8 - 11" refId="ref34583" refString="Abe W, Ito M. Itaquascon globuliferum, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Japan. Proceedings of the Japanese Society of Systematic Zoology 1994; 51: 8 - 11." type="journal article" year="1994">Abe & Ito, 1994</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
has internal cuticular bars at the claw I–III bases and posterior bars (
|
||||
<bibRefCitation id="EF974B23A61CFF97AEF56B24FBD98D97" author="Guidetti R & Bertolani R & Nelson DR" box="[939,1147,1919,1944]" pageId="37" pageNumber="38" pagination="215 - 23" refId="ref36352" refString="Guidetti R, Bertolani R, Nelson DR. Ecological and faunistic studies on tardigrades in leaf litter of beech forests. Zoologischer Anzeiger 1999; 238: 215 - 23." type="journal article" year="1999">
|
||||
Guidetti
|
||||
<emphasis id="B972EAC0A61CFF97A9526BDBFB9C8D98" box="[1036,1086,1919,1943]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
|
||||
1999
|
||||
</bibRefCitation>
|
||||
), whereas both
|
||||
<typeStatus id="54BD8870A61CFF97A8746BDBFAC18D97" box="[1322,1379,1920,1944]" pageId="37" pageNumber="38">types</typeStatus>
|
||||
of bars are absent in
|
||||
<taxonomicName id="4C064D51A61CFF97AEE66BFBFBF88DB8" authorityName="Gasiorek & Michalczyk" authorityYear="2024" box="[952,1114,1952,1975]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="37" pageNumber="38" phylum="Tardigrada" rank="species" species="minutissimus">
|
||||
<emphasis id="B972EAC0A61CFF97AEE66BFBFBF88DB8" box="[952,1114,1952,1975]" italics="true" pageId="37" pageNumber="38">R. minutissimus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
94
data/03/AF/87/03AF87C4A61EFF95ADB06B59FD098D44.xml
Normal file
94
data/03/AF/87/03AF87C4A61EFF95ADB06B59FD098D44.xml
Normal file
|
|
@ -0,0 +1,94 @@
|
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<mods:titleInfo id="18A3C67350E99B57FA46D1068FA730D0">
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<mods:nameIdentifier id="50DDB6ED8F101F3537BB3884821BBDA1" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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,
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</taxonomicName>
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<typeStatus id="54BD8870A61EFF95AD9E6B69FF548D43" box="[192,246,1842,1868]" pageId="39" pageNumber="40">Type</typeStatus>
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species
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<taxonomicName id="4C064D51A61EFF95AC156B6AFD098D44" ID-CoL="7T84R" authority="Murray, 1905" authorityName="Murray" authorityYear="1905" box="[331,683,1841,1868]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="39" pageNumber="40" phylum="Tardigrada" rank="species" species="bullatum">
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<emphasis id="B972EAC0A61EFF95AC156B6AFDBB8D43" bold="true" box="[331,537,1841,1868]" italics="true" pageId="39" pageNumber="40">Diphascon bullatum</emphasis>
|
||||
Murray, 1905
|
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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112
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<mods:nameIdentifier id="F1BA3CBEE6F5C7B098DA0F66BEF164B1" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6641A69DA6EDF38F4074A6677B0F1491">Blagden, Brian</mods:namePart>
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<mods:affiliation id="3AF312EE5C1E0B4A273D7DB72167A55A">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:date id="D16283ADE33F6590098B52FC31CA6AD3">2024</mods:date>
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Taxonomic key to the genus
|
||||
<taxonomicName id="4C064D51A61FFF94AED06AB7FC448D09" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[910,998,1772,1798]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="38" pageNumber="39" phylum="Tardigrada" rank="genus">
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|
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|
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|
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|
||||
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||||
<paragraph id="8BB936D2A61FFF94ADC56B76FA058D4A" blockId="38.[155,1448,1772,1955]" box="[155,1447,1837,1861]" pageId="38" pageNumber="39">
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||||
<keyLead id="30F795E7A61FFF94ADC56B76FA058D4A" box="[155,1447,1837,1861]" pageId="38" pageNumber="39">
|
||||
1. Pharynx devoid of placoids ........................................................................................................................................
|
||||
<taxonomicName id="4C064D51A61FFF94A9BD6B76FA058D4A" authorityName="NEC" baseAuthorityName="Weglarska" baseAuthorityYear="1973" box="[1251,1447,1837,1861]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="38" pageNumber="39" phylum="Tardigrada" rank="species" species="pawlowskii">
|
||||
<emphasis id="B972EAC0A61FFF94A9BD6B76FA058D4A" box="[1251,1447,1837,1861]" italics="true" pageId="38" pageNumber="39">Raribius pawlowskii</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A61FFF94ADC56B16FA058D6A" blockId="38.[155,1448,1772,1955]" box="[155,1447,1869,1893]" pageId="38" pageNumber="39">
|
||||
<keyLead id="30F795E7A61FFF94ADC56B16FA058D6A" box="[155,1447,1869,1893]" pageId="38" pageNumber="39">– Pharynx with short rectangular placoids .................................................................................................................................................. 2</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A61FFF94ADC56B37FA0A8DAC" pageId="38" pageNumber="39">
|
||||
<paragraph id="8BB936D2A61FFF94ADC56B37FA058D8B" blockId="38.[155,1448,1772,1955]" box="[155,1447,1900,1924]" pageId="38" pageNumber="39">
|
||||
<keyLead id="30F795E7A61FFF94ADC56B37FA058D8B" box="[155,1447,1900,1924]" pageId="38" pageNumber="39">
|
||||
2. Internal bars I–III and posterior bars present .......................................................................................................
|
||||
<taxonomicName id="4C064D51A61FFF94A9816B37FA058D8B" baseAuthorityName="Abe & Ito" baseAuthorityYear="1994" box="[1247,1447,1900,1924]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="38" pageNumber="39" phylum="Tardigrada" rank="species" species="globuliferus">
|
||||
<emphasis id="B972EAC0A61FFF94A9816B37FA058D8B" box="[1247,1447,1900,1924]" italics="true" pageId="38" pageNumber="39">Raribius globuliferus</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A61FFF94ADC56BD7FA0A8DAC" blockId="38.[155,1448,1772,1955]" box="[155,1448,1931,1955]" pageId="38" pageNumber="39">
|
||||
<keyLead id="30F795E7A61FFF94ADC56BD7FA0A8DAC" box="[155,1448,1931,1955]" pageId="38" pageNumber="39">
|
||||
– Internal bars I–III and posterior bars absent .......................................................................................
|
||||
<taxonomicName id="4C064D51A61FFF94A9276BD0FAF38DAC" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1145,1361,1931,1955]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="38" pageNumber="39" phylum="Tardigrada" rank="species" species="minutissimus" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A61FFF94A9276BD0FAF38DAC" box="[1145,1361,1931,1955]" italics="true" pageId="38" pageNumber="39">Raribius minutissimus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA61FFF94A8086BD0FA0A8DAC" box="[1366,1448,1931,1955]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
545
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Normal file
545
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Normal file
|
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@ -0,0 +1,545 @@
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<mods:title id="11EF9D71797329FB5DF305FE033E7D03">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:affiliation id="ED6379159F6E6CAB96AD7BAACE5E0484">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="DF10D4B5DBC7A459A8534D804BB4F271">Blagden, Brian</mods:namePart>
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<mods:affiliation id="4ACB168E3BCFE7BC67D5D10440F1C250">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="DE19A28A3EEE07686AE7CB91431A1418">Morek, Witold</mods:namePart>
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<mods:affiliation id="4DC495AE1DCCC63A29A8E186A80878F9">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="DDB827523F9F4567B70CEE09B9DD4C8B">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="D8104CEE9207BBA2B3C4955E309B8D72">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<subSubSection id="C31C6559A620FFABADA46B08FDD08D62" box="[250,626,1875,1902]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BB936D2A620FFABADA46B08FDD08D62" blockId="25.[250,626,1875,1902]" box="[250,626,1875,1902]" pageId="25" pageNumber="26">
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<heading id="D0F181BEA620FFABADA46B08FDD08D62" bold="true" box="[250,626,1875,1902]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
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<taxonomicName id="4C064D51A620FFABADA46B08FDD08D62" ID-CoL="4JNBW" authority="(Murray, 1905)" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[250,626,1875,1902]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="angustata">
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<emphasis id="B972EAC0A620FFABADA46B08FDD08D62" bold="true" box="[250,626,1875,1902]" pageId="25" pageNumber="26">
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<emphasis id="B972EAC0A620FFABADA46B08FE688D62" bold="true" box="[250,458,1875,1902]" italics="true" pageId="25" pageNumber="26">Platicrista angustata</emphasis>
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(Murray, 1905)
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<paragraph id="8BB936D2A620FFABADD36B24FDAC8DB6" blockId="25.[141,734,1919,1977]" pageId="25" pageNumber="26">
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<emphasis id="B972EAC0A620FFABADD36B24FDAC8DB6" bold="true" pageId="25" pageNumber="26">
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<taxonomicName id="4C064D51A620FFABADD36B24FED78D95" authorityName="Murray" authorityYear="1905" box="[141,373,1919,1946]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="angustatum">
|
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<emphasis id="B972EAC0A620FFABADD36B24FED78D95" bold="true" box="[141,373,1919,1946]" italics="true" pageId="25" pageNumber="26">Diphascon angustatum</emphasis>
|
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</taxonomicName>
|
||||
;
|
||||
<emphasis id="B972EAC0A620FFABACDC6BDBFE578D95" bold="true" box="[386,501,1920,1946]" italics="true" pageId="25" pageNumber="26">terra typica</emphasis>
|
||||
: Loch Ness,
|
||||
<collectingCountry id="F3117642A620FFABAF276B24FD7B8D96" box="[633,729,1919,1945]" name="United Kingdom" pageId="25" pageNumber="26">Scotland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A620FFABAC036BC4FDAB8DB6" author="Murray J & The Tardigrada of the Scottish Lochs & Transactions of the Royal Society of Edinburgh & Part III" box="[349,521,1951,1977]" pageId="25" pageNumber="26" pagination="677 - 98" refId="ref38253" refString="Murray J. The Tardigrada of the Scottish Lochs. Transactions of the Royal Society of Edinburgh, Part III 1905 b; 41: 677 - 98. https: // doi. org / 10.5962 / bhl. title. 12517" type="book chapter" year="1905">Murray (1905b)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABAE126AAEFA338D00" blockId="25.[844,1425,1780,1807]" box="[844,1425,1780,1807]" pageId="25" pageNumber="26">
|
||||
<heading id="D0F181BEA620FFABAE126AAEFA338D00" bold="true" box="[844,1425,1780,1807]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
|
||||
<emphasis id="B972EAC0A620FFABAE126AAEFA338D00" bold="true" box="[844,1425,1780,1807]" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C064D51A620FFABAE126AAEFC428D00" authorityName="Murray" authorityYear="1905" box="[844,992,1781,1807]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="angustatum">
|
||||
<emphasis id="B972EAC0A620FFABAE126AAEFCFD8D00" bold="true" box="[844,863,1781,1807]" italics="true" pageId="25" pageNumber="26">D</emphasis>
|
||||
.
|
||||
<emphasis id="B972EAC0A620FFABAE376AAEFC428D00" bold="true" box="[873,992,1781,1807]" italics="true" pageId="25" pageNumber="26">angustatum</emphasis>
|
||||
</taxonomicName>
|
||||
; Forth Valley,
|
||||
<collectingCountry id="F3117642A620FFABA92B6AAFFB778D01" box="[1141,1237,1780,1806]" name="United Kingdom" pageId="25" pageNumber="26">Scotland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A620FFABA9816AAEFA2E8D01" author="Murray J" box="[1247,1420,1780,1807]" pageId="25" pageNumber="26" pagination="214 - 7" refId="ref38317" refString="Murray J. The Tardigrada of the Forth Valley (second paper). Annals of Scottish Natural History 1906 b; 60: 214 - 7." type="journal article" year="1906">Murray (1906b)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABAE6B6B64FA0A8D56" blockId="25.[821,1448,1854,1881]" box="[821,1448,1854,1881]" pageId="25" pageNumber="26">
|
||||
<heading id="D0F181BEA620FFABAE6B6B64FA0A8D56" bold="true" box="[821,1448,1854,1881]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
|
||||
<emphasis id="B972EAC0A620FFABAE6B6B64FA0A8D56" bold="true" box="[821,1448,1854,1881]" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C064D51A620FFABAE6B6B64FC6B8D56" ID-CoL="36FH4" authorityName="Murray" authorityYear="1905" box="[821,969,1855,1881]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="angustatum">
|
||||
<emphasis id="B972EAC0A620FFABAE6B6B64FCEA8D56" bold="true" box="[821,840,1855,1881]" italics="true" pageId="25" pageNumber="26">D</emphasis>
|
||||
.
|
||||
<emphasis id="B972EAC0A620FFABAE0C6B64FC6B8D56" bold="true" box="[850,969,1855,1881]" italics="true" pageId="25" pageNumber="26">angustatum</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA620FFABAE886B65FA008D57" box="[982,1442,1854,1881]" country="United Kingdom" location="Edinburgh" pageId="25" pageNumber="26" specimenCount="1" stateProvince="City of Edinburgh">
|
||||
<collectingRegion id="49C2F830A620FFABAE886B65FBE88D57" box="[982,1098,1854,1880]" country="United Kingdom" name="City of Edinburgh" pageId="25" pageNumber="26">Edinburgh</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49C2F830A620FFABA90C6B65FB2D8D57" box="[1106,1167,1854,1880]" country="United Kingdom" name="Killin" pageId="25" pageNumber="26">Killin</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F3117642A620FFABA9C76B65FB5B8D57" box="[1177,1273,1854,1880]" name="United Kingdom" pageId="25" pageNumber="26">Scotland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A620FFABA85C6B64FA008D57" author="Murray J" box="[1282,1442,1854,1881]" pageId="25" pageNumber="26" pagination="641 - 68" refId="ref38345" refString="Murray J. Scottish Tardigrada, collected by the lake survey. Transactions of the Royal Society of Edinburgh 1907; 45: 641 - 68. https: // doi. org / 10.1017 / s 0080456800011777" type="journal article" year="1907">Murray (1907)</bibRefCitation>
|
||||
</materialsCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFA8AE0D6B22FDEE8AC6" blockId="25.[851,1418,1912,1970]" lastBlockId="26.[146,760,143,201]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||||
<emphasis id="B972EAC0A620FFA8AE0D6B22FDEE8AC6" bold="true" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C064D51A620FFABAE0D6B22FC458D9C" ID-CoL="36FH4" authorityName="Murray" authorityYear="1905" box="[851,999,1913,1939]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="angustatum">
|
||||
<emphasis id="B972EAC0A620FFABAE0D6B22FCC48D9C" bold="true" box="[851,870,1913,1939]" italics="true" pageId="25" pageNumber="26">D</emphasis>
|
||||
.
|
||||
<emphasis id="B972EAC0A620FFABAE2E6B22FC458D9C" bold="true" box="[880,999,1913,1939]" italics="true" pageId="25" pageNumber="26">angustatum</emphasis>
|
||||
</taxonomicName>
|
||||
; Achill Island, Belclare,
|
||||
<collectingCountry id="F3117642A620FFABA9BC6B22FA908D9C" box="[1250,1330,1913,1939]" name="Ireland" pageId="25" pageNumber="26">Ireland</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A620FFABA8656B22FB308DBE" author="Murray J" pageId="25" pageNumber="26" pagination="1 - 16" refId="ref38408" refString="Murray J. Clare Island Survey. Arctiscoida. Proceedings of the Royal Irish Academy 1911; 31: 1 - 16." type="journal article" year="1911">Murray (1911)</bibRefCitation>
|
||||
.
|
||||
<taxonomicName id="4C064D51A623FFA8ADCC6CD4FE3A8AA5" ID-CoL="3P3GW" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[146,408,143,170]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustatus" subGenus="Diphascon">
|
||||
<emphasis id="B972EAC0A623FFA8ADCC6CD4FF548AA5" bold="true" box="[146,246,143,170]" italics="true" pageId="26" pageNumber="27">Hypsibius</emphasis>
|
||||
(
|
||||
<emphasis id="B972EAC0A623FFA8AC5B6CCBFEBE8AA5" bold="true" box="[261,284,144,170]" italics="true" pageId="26" pageNumber="27">D.</emphasis>
|
||||
)
|
||||
<emphasis id="B972EAC0A623FFA8AC756CCBFE3A8AA5" bold="true" box="[299,408,144,170]" italics="true" pageId="26" pageNumber="27">angustatus</emphasis>
|
||||
</taxonomicName>
|
||||
; Dartmoor,
|
||||
<collectingCountry id="F3117642A623FFA8AF446CCBFDD18AA5" box="[538,627,144,170]" name="United Kingdom" pageId="26" pageNumber="27">England</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F3117642A623FFA8AF226CCBFD518AA5" box="[636,755,144,170]" name="Isle of Man" pageId="26" pageNumber="27">Isle of Man</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A623FFA8AC636CF4FDE58AC6" author="Morgan CI & King PE" box="[317,583,175,201]" pageId="26" pageNumber="27" pagination="1 - 133" refId="ref38203" refString="Morgan CI, King PE. British Tardigrades. Synopses of the British Fauna (New Series) 1976; 9: 1 - 133." type="journal article" year="1976">Morgan and King (1976)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8ADD86CB6FE4F8B28" blockId="26.[134,772,237,295]" pageId="26" pageNumber="27">
|
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<emphasis id="B972EAC0A623FFA8ADD86CB6FE4F8B28" bold="true" pageId="26" pageNumber="27">
|
||||
<taxonomicName id="4C064D51A623FFA8ADD86CB6FED88B07" ID-CoL="4JNBW" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[134,378,237,264]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8ADD86CB6FF4F8B07" bold="true" box="[134,237,237,264]" italics="true" pageId="26" pageNumber="27">Platicrista</emphasis>
|
||||
aff.
|
||||
<emphasis id="B972EAC0A623FFA8AC486CB5FED88B07" bold="true" box="[278,378,238,264]" italics="true" pageId="26" pageNumber="27">angustata</emphasis>
|
||||
</taxonomicName>
|
||||
4;
|
||||
<collectingCountry id="F3117642A623FFA8ACC66CB5FE4D8B07" box="[408,495,238,264]" name="Norway" pageId="26" pageNumber="27">Norway</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A623FFA8ACA66CB5FE4A8B28" author="Gasiorek P & Michalczyk L" pageId="26" pageNumber="27" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A623FFA8ADDF6D15FF5E8E44" pageId="26" pageNumber="27" type="materials_examined">
|
||||
<paragraph id="8BB936D2A623FFA8ADDF6D15FE4B8B8A" blockId="26.[129,777,334,389]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF6D15FEEB8B69" box="[129,329,334,358]" italics="true" pageId="26" pageNumber="27">Material examined:</emphasis>
|
||||
49 individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A623FFA8ACDF6D36FE7B8B8A" box="[385,473,365,389]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="26" pageNumber="27">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8ADDF6DFCFE6488B6" blockId="26.[129,778,422,697]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF6DFCFE8F8BB1" box="[129,301,422,446]" italics="true" pageId="26" pageNumber="27">
|
||||
<typeStatus id="54BD8870A623FFA8ADDF6DFCFF738BB1" box="[129,209,423,446]" pageId="26" pageNumber="27" type="neotype">Neotype</typeStatus>
|
||||
material:
|
||||
</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA623FFA8AC6E6DFCFE6088B6" collectingDate="2014-11-01" collectingDateMax="2016-03-24" collectingDateMin="2014-11-01" collectionCode="GB" collectorName="Blagden" country="United Kingdom" elevation="451" latitude="56.959167" location="Hill of Foudland" longLatPrecision="17" longitude="-4.5677776" municipality="Institute of Zoology" pageId="26" pageNumber="27" specimenCode="GB.004.01- 06, GB.012.19-25, GB.058.01-10" specimenCount="28" stateProvince="Creag Meagaidh" typeStatus="neotype">
|
||||
<typeStatus id="54BD8870A623FFA8AC6E6DFCFE288BB0" box="[304,394,423,447]" pageId="26" pageNumber="27" type="neotype">Neotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="9D00FD5BA623FFA8ACE66DFDFDE38BB1" box="[440,577,422,446]" count="28" pageId="26" pageNumber="27" type="generic">28 specimens</specimenCount>
|
||||
(slides
|
||||
<specimenCode id="DBA09EA9A623FFA8AFD26DFDFF3F8BD1" pageId="26" pageNumber="27">GB.004.01– 06</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A623FFA8ADF36D9EFEE78BD2" box="[173,325,453,477]" pageId="26" pageNumber="27">GB.012.19–25</specimenCode>
|
||||
, and
|
||||
<specimenCode id="DBA09EA9A623FFA8ACD66D9EFD808BD2" box="[392,546,453,477]" pageId="26" pageNumber="27">GB.058.01–10</specimenCode>
|
||||
) from
|
||||
<collectingCountry id="F3117642A623FFA8AF2B6D9DFD738BD1" box="[629,721,454,478]" name="United Kingdom" pageId="26" pageNumber="27">Scotland</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A623FFA8AFBF6D9DFEA78BF2" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8AFBF6D9DFEA78BF2" country="United Kingdom" latitude="56.959167" longLatPrecision="17" longitude="-4.5677776" municipality="Institute of Zoology" name="Hill of Foudland" pageId="26" pageNumber="27" stateProvince="Creag Meagaidh">Hill of Foudland</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A623FFA8AC476DBEFE348BF2" box="[281,406,483,509]" degrees="57" direction="north" minutes="23" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="15" value="57.3875">
|
||||
57°23
|
||||
<emphasis id="B972EAC0A623FFA8AC076DB8FEFD8BF3" box="[345,351,483,508]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
15″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8ACF96DBEFDB98BF2" box="[423,539,483,509]" degrees="2" direction="west" minutes="39" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="39" value="-2.6608334">
|
||||
2°39
|
||||
<emphasis id="B972EAC0A623FFA8AC886DB8FE7E8BF3" box="[470,476,483,508]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
39″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8AF726DBEFD128BF2" box="[556,688,485,509]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="26" pageNumber="27" unit="m" value="451.0">
|
||||
<quantity id="4CFE9B37A623FFA8AF726DBEFDD78BF3" box="[556,629,485,509]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="26" pageNumber="27" unit="m" value="451.0">451 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), lichen (
|
||||
<taxonomicName id="4C064D51A623FFA8ADD26E5FFF448813" box="[140,230,516,540]" class="Lecanoromycetes" family="Cladoniaceae" genus="Cladonia" kingdom="Fungi" order="Lecanorales" pageId="26" pageNumber="27" phylum="Ascomycota" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8ADD26E5FFF448813" box="[140,230,516,540]" italics="true" pageId="26" pageNumber="27">Cladonia</emphasis>
|
||||
</taxonomicName>
|
||||
) from soil in abandoned slate quarry,
|
||||
<date id="FFB81012A623FFA8AFDB6E5FFF158833" pageId="26" pageNumber="27" value="2014-11-01">
|
||||
<collectingDate id="EFFCE9FAA623FFA8AFDB6E5FFF158833" pageId="26" pageNumber="27" value="2014-11-01">1 November 2014</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8AD986E7FFEBB8833" box="[198,281,548,572]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll., and
|
||||
<collectingCountry id="F3117642A623FFA8ACCC6E7FFE4C8833" box="[402,494,548,572]" name="United Kingdom" pageId="26" pageNumber="27">Scotland</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49C2F830A623FFA8ACA36E7FFD0E8833" box="[509,684,548,572]" pageId="26" pageNumber="27">Creag Meagaidh</collectingRegion>
|
||||
,
|
||||
<location id="8ED96009A623FFA8AFE56E7FFF408854" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8AFE56E7FFF408854" country="United Kingdom" latitude="56.959167" longLatPrecision="17" longitude="-4.5677776" municipality="Institute of Zoology" name="Lochan a' Choire" pageId="26" pageNumber="27" stateProvince="Creag Meagaidh">Lochan a’ Choire</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A623FFA8ADAA6E18FED28854" box="[244,368,577,603]" degrees="56" direction="north" minutes="57" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="33" value="56.959167">
|
||||
56°57
|
||||
<emphasis id="B972EAC0A623FFA8AC6A6E1AFE988855" box="[308,314,577,602]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
33″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8AC216E18FE518854" box="[383,499,577,603]" degrees="4" direction="west" minutes="34" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="04" value="-4.5677776">
|
||||
4°34
|
||||
<emphasis id="B972EAC0A623FFA8ACF06E1AFE168855" box="[430,436,577,602]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
04″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8AF5F6E18FD228854" box="[513,640,579,603]" metricMagnitude="2" metricUnit="m" metricValue="6.24" pageId="26" pageNumber="27" unit="m" value="624.0">
|
||||
<quantity id="4CFE9B37A623FFA8AF5F6E18FDEA8855" box="[513,584,579,603]" metricMagnitude="2" metricUnit="m" metricValue="6.24" pageId="26" pageNumber="27" unit="m" value="624.0">624 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), moss from rock in alpine grassland,
|
||||
<date id="FFB81012A623FFA8ACDD6E39FD878875" box="[387,549,610,634]" pageId="26" pageNumber="27" value="2016-03-24">
|
||||
<collectingDate id="EFFCE9FAA623FFA8ACDD6E39FD878875" box="[387,549,610,634]" pageId="26" pageNumber="27" value="2016-03-24">24 March 2016</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8AF6E6E39FD218875" box="[560,643,610,634]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll., are deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University, Kraków
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8ADDF6E80FF5E8E44" blockId="26.[128,779,730,1099]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF6E80FECC88FD" box="[129,366,730,754]" italics="true" pageId="26" pageNumber="27">Comparative material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA623FFA8ACDA6E81FF5B895F" collectingDate="2015-04-25" collectorName="Blagden" country="United Kingdom" elevation="390" latitude="57.383892" location="Gartly Forest" longLatPrecision="17" longitude="-2.7144444" pageId="26" pageNumber="27" specimenCount="5">
|
||||
<collectingCountry id="F3117642A623FFA8ACDA6E81FE4288FD" box="[388,480,730,754]" name="United Kingdom" pageId="26" pageNumber="27">Scotland</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A623FFA8ACA46E81FD3188FD" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8ACA46E81FD3188FD" box="[506,659,730,754]" country="United Kingdom" latitude="57.383892" longLatPrecision="17" longitude="-2.7144444" name="Gartly Forest" pageId="26" pageNumber="27">Gartly Forest</location>
|
||||
, southern boundary (
|
||||
<geoCoordinate id="EE325015A623FFA8ADAB6EA2FEC6891E" box="[245,356,760,785]" degrees="57" direction="north" minutes="23" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="2" value="57.383892">
|
||||
57°23
|
||||
<emphasis id="B972EAC0A623FFA8AC6A6EA3FE98891E" box="[308,314,760,785]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
2″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8AC2E6EA1FE47891E" box="[368,485,760,786]" degrees="2" direction="west" minutes="42" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="52" value="-2.7144444">
|
||||
2°42
|
||||
<emphasis id="B972EAC0A623FFA8ACC16EA3FE07891E" box="[415,421,760,785]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
52″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8ACAF6EA1FDCE891D" box="[497,620,762,786]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="27" unit="m" value="390.0">
|
||||
<quantity id="4CFE9B37A623FFA8ACAF6EA1FD94891E" box="[497,566,762,786]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="27" unit="m" value="390.0">390 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), forest, lichen (
|
||||
<taxonomicName id="4C064D51A623FFA8ADD26F42FF46893E" authorityName="Acharius" authorityYear="1803" box="[140,228,793,817]" class="Arthoniomycetes" family="Arthoniaceae" genus="Parmelia" kingdom="Fungi" order="Arthoniales" pageId="26" pageNumber="27" phylum="Ascomycota" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8ADD26F42FF46893E" box="[140,228,793,817]" italics="true" pageId="26" pageNumber="27">Parmelia</emphasis>
|
||||
</taxonomicName>
|
||||
) from stone wall,
|
||||
<date id="FFB81012A623FFA8ACF46F42FDE2893E" box="[426,576,793,817]" pageId="26" pageNumber="27" value="2015-04-25">
|
||||
<collectingDate id="EFFCE9FAA623FFA8ACF46F42FDE2893E" box="[426,576,793,817]" pageId="26" pageNumber="27" value="2015-04-25">25 April 2015</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8AF126F42FD3D893E" box="[588,671,793,817]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA623FFA8AFBC6F42FF50895F" count="5" pageId="26" pageNumber="27" type="generic">five specimens</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA623FFA8AC536F63FF6E89A1" collectingDate="2015-09-20" collectorName="Blagden" country="Scotland" county="Oxen Craig" elevation="520" latitude="57.29333" location="Oyne" longLatPrecision="17" longitude="-2.5602777" municipality="Bennachie" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Aberdeenshire">
|
||||
<collectingCounty id="62D84E5EA623FFA8AC536F63FE32895F" box="[269,400,824,848]" pageId="26" pageNumber="27">Oxen Craig</collectingCounty>
|
||||
,
|
||||
<collectingMunicipality id="6BDDACA8A623FFA8ACFA6F63FDB3895F" box="[420,529,824,848]" pageId="26" pageNumber="27">Bennachie</collectingMunicipality>
|
||||
,
|
||||
<location id="8ED96009A623FFA8AF7B6F63FDC2895F" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8AF7B6F63FDC2895F" box="[549,608,824,848]" country="United Kingdom" county="Oxen Craig" latitude="57.29333" longLatPrecision="17" longitude="-2.5602777" municipality="Bennachie" name="Oyne" pageId="26" pageNumber="27" stateProvince="Aberdeenshire">Oyne</location>
|
||||
,
|
||||
<collectingRegion id="49C2F830A623FFA8AF2A6F63FCAB895F" box="[628,777,824,848]" country="United Kingdom" name="Aberdeenshire" pageId="26" pageNumber="27">Aberdeenshire</collectingRegion>
|
||||
(
|
||||
<geoCoordinate id="EE325015A623FFA8ADD66F0CFEA78960" box="[136,261,854,879]" degrees="57" direction="north" minutes="17" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="36" value="57.29333">
|
||||
57°17
|
||||
<emphasis id="B972EAC0A623FFA8AD966F0DFF6C8960" box="[200,206,854,879]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
36″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8AC496F03FE288960" box="[279,394,854,880]" degrees="2" direction="west" minutes="33" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="37" value="-2.5602777">
|
||||
2°33
|
||||
<emphasis id="B972EAC0A623FFA8AC186F0DFEEE8960" box="[326,332,854,879]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
37″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8ACC36F0CFD81897F" box="[413,547,855,880]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="26" pageNumber="27" unit="m" value="520.0">
|
||||
<quantity id="4CFE9B37A623FFA8ACC36F0CFE458960" box="[413,487,855,879]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="26" pageNumber="27" unit="m" value="520.0">520 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), heather moor, soil from rock crevice,
|
||||
<date id="FFB81012A623FFA8AC1D6F2CFDB28980" box="[323,528,887,911]" pageId="26" pageNumber="27" value="2015-09-20">
|
||||
<collectingDate id="EFFCE9FAA623FFA8AC1D6F2CFDB28980" box="[323,528,887,911]" pageId="26" pageNumber="27" value="2015-09-20">20 September 2015</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8AF456F2CFDCC8980" box="[539,622,887,911]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA623FFA8AFF16F2CFF6489A1" count="1" pageId="26" pageNumber="27" type="generic">one specimen</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA623FFA8AD866FCCFD7389E2" collectingDate="2016-09-05" collectorName="Blagden" country="Scotland" elevation="114" latitude="57.094997" location="Drumoak" longLatPrecision="17" longitude="-2.3377776" municipality="Drum Castle" pageId="26" pageNumber="27" specimenCount="3">
|
||||
<collectingMunicipality id="6BDDACA8A623FFA8AD866FCCFEFD89A1" box="[216,351,918,942]" pageId="26" pageNumber="27">Drum Castle</collectingMunicipality>
|
||||
,
|
||||
<location id="8ED96009A623FFA8AC346FCDFE6F89A1" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8AC346FCDFE6F89A1" box="[362,461,918,942]" country="United Kingdom" latitude="57.094997" longLatPrecision="17" longitude="-2.3377776" municipality="Drum Castle" name="Drumoak" pageId="26" pageNumber="27">Drumoak</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A623FFA8AC856FCDFDFA89A1" box="[475,600,917,942]" degrees="57" direction="north" minutes="05" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="42" value="57.094997">
|
||||
57°05
|
||||
<emphasis id="B972EAC0A623FFA8AF456FCEFD8389A1" box="[539,545,917,942]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
42″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8AF3D6FCDFD7589A1" box="[611,727,917,942]" degrees="2" direction="west" minutes="20" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="16" value="-2.3377776">
|
||||
2°20
|
||||
<emphasis id="B972EAC0A623FFA8AFCC6FCEFD3A89A1" box="[658,664,917,942]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
16″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8AFBC6FCDFF6989C1" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="26" pageNumber="27" unit="m" value="114.0">
|
||||
<quantity id="4CFE9B37A623FFA8AFBC6FCDFF3589C2" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="26" pageNumber="27" unit="m" value="114.0">114 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), rural vicinity, leaf litter (
|
||||
<taxonomicName id="4C064D51A623FFA8AC866FEDFDC689C1" box="[472,612,950,974]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="robur">
|
||||
<emphasis id="B972EAC0A623FFA8AC866FEDFDC689C1" box="[472,612,950,974]" italics="true" pageId="26" pageNumber="27">Quercus robur</emphasis>
|
||||
</taxonomicName>
|
||||
) collected from soil,
|
||||
<date id="FFB81012A623FFA8ADEA6F8EFED589E2" box="[180,375,981,1005]" pageId="26" pageNumber="27" value="2016-09-05">
|
||||
<collectingDate id="EFFCE9FAA623FFA8ADEA6F8EFED589E2" box="[180,375,981,1005]" pageId="26" pageNumber="27" value="2016-09-05">5 September 2016</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8ACDD6F8EFE7489E2" box="[387,470,981,1005]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA623FFA8AF476F8EFD6889E2" box="[537,714,981,1005]" count="3" pageId="26" pageNumber="27" type="generic">three specimens</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA623FFA8AF806F8EFF5A8E44" collectingDate="2015-11-05" collectorName="Blagden" country="Scotland" elevation="20" latitude="57.146946" location="Loch Ness" longLatPrecision="17" longitude="-4.6783333" municipality="Fort Augustus" pageId="26" pageNumber="27" specimenCount="2">
|
||||
<collectingMunicipality id="6BDDACA8A623FFA8AF806F8EFF408E02" pageId="26" pageNumber="27">Fort Augustus</collectingMunicipality>
|
||||
,
|
||||
<location id="8ED96009A623FFA8ADB56FAFFEF48E03" LSID="urn:lsid:plazi:treatment:03AF87C4A620FFA8ADA46B08FBCB8830:8ED96009A623FFA8ADB56FAFFEF48E03" box="[235,342,1012,1036]" country="United Kingdom" latitude="57.146946" longLatPrecision="17" longitude="-4.6783333" municipality="Fort Augustus" name="Loch Ness" pageId="26" pageNumber="27">Loch Ness</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A623FFA8AC3D6FAFFE428E03" box="[355,480,1011,1036]" degrees="57" direction="north" minutes="08" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="49" value="57.146946">
|
||||
57°08
|
||||
<emphasis id="B972EAC0A623FFA8ACFD6FA8FE0B8E03" box="[419,425,1011,1036]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
49″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A623FFA8ACB76FAFFDFF8E03" box="[489,605,1011,1036]" degrees="4" direction="west" minutes="40" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="42" value="-4.6783333">
|
||||
4°40
|
||||
<emphasis id="B972EAC0A623FFA8AF466FA8FDBC8E03" box="[536,542,1011,1036]" italics="true" pageId="26" pageNumber="27">ʹ</emphasis>
|
||||
42″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A623FFA8AF386FAFFD6F8E03" box="[614,717,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="20.0">
|
||||
<quantity id="4CFE9B37A623FFA8AF386FAFFD388E03" box="[614,666,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="20.0">20 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), lake shore, moss from rock,
|
||||
<date id="FFB81012A623FFA8AC256848FD9E8E24" box="[379,572,1043,1068]" pageId="26" pageNumber="27" value="2015-11-05">
|
||||
<collectingDate id="EFFCE9FAA623FFA8AC256848FD9E8E24" box="[379,572,1043,1068]" pageId="26" pageNumber="27" value="2015-11-05">5 November 2015</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A623FFA8AF17684FFD3E8E23" box="[585,668,1044,1068]" pageId="26" pageNumber="27">Blagden</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA623FFA8AF81684FFF538E44" count="2" pageId="26" pageNumber="27" type="generic">two specimens</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A623FFA8ADDF683BFE4D8D26" pageId="26" pageNumber="27" type="description">
|
||||
<paragraph id="8BB936D2A623FFA8ADDF683BFF4D8C41" blockId="26.[128,780,1120,1833]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF683BFEA98E77" box="[129,267,1120,1144]" italics="true" pageId="26" pageNumber="27">Redescription:</emphasis>
|
||||
Body medium-sized to very large (
|
||||
<tableCitation id="C6840369A623FFA8AFFB683BFD5B8E77" box="[677,761,1120,1144]" captionStart="Table 6" captionStartId="27.[113,168,143,167]" captionTargetPageId="27" captionText="Table 6. Measurements (in micrometres) of selected morphological structures of neotype series (GB.004 + GB.012 + GB.058) of Platicrista angustata (Murray, 1905) in Murray (1905b) from Scotland mounted in Hoyer’s medium." pageId="26" pageNumber="27">Table 6</tableCitation>
|
||||
), elongate and slightly narrowed in its anterior part (
|
||||
<figureCitation id="133D2A57A623FFA8AFC16824FD5A8E98" box="[671,760,1151,1175]" captionStart="Figure 18" captionStartId="28.[128,193,1100,1124]" captionTargetBox="[129,1473,144,1073]" captionTargetId="figure-424@28.[129,1473,144,1073]" captionTargetPageId="28" captionText="Figure 18. Platicrista angustata (Murray, 1905) in Murray (1905b) (PCM): A, neotype habitus in ventral view; B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243784" httpUri="https://zenodo.org/record/14243784/files/figure.png" pageId="26" pageNumber="27">Fig. 18A</figureCitation>
|
||||
). Head blunt. Body white in smaller individuals, but brownish in the caudal part in large ones. Cuticle smooth, without pores or undulations, but with numerous irregular wrinkles intermingled with granulation in some individuals (
|
||||
<figureCitation id="133D2A57A623FFA8AF6568A7FD238F1B" box="[571,641,1276,1300]" captionStart="Figure 19" captionStartId="29.[114,179,1837,1861]" captionTargetBox="[332,1241,144,1809]" captionTargetId="figure-63@29.[332,1241,144,1809]" captionTargetPageId="29" captionText="Figure 19. Caudal sculpturing of Platicrista angustata (PCM): A, juvenile; B, C, adults. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243790" httpUri="https://zenodo.org/record/14243790/files/figure.png" pageId="26" pageNumber="27">Fig. 19</figureCitation>
|
||||
). Cribriform areas not visible under PCM. Legs very short and plump; legs IV barely distinguishable from the posterior body end (
|
||||
<figureCitation id="133D2A57A623FFA8AFBD6960FF098F7D" captionStart="Figure 18" captionStartId="28.[128,193,1100,1124]" captionTargetBox="[129,1473,144,1073]" captionTargetId="figure-424@28.[129,1473,144,1073]" captionTargetPageId="28" captionText="Figure 18. Platicrista angustata (Murray, 1905) in Murray (1905b) (PCM): A, neotype habitus in ventral view; B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243784" httpUri="https://zenodo.org/record/14243784/files/figure.png" pageId="26" pageNumber="27">Fig. 18A</figureCitation>
|
||||
). Eyes absent in live animals. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A623FFA8ADF76921FEAE8F9D" authorityName="Pilato" authorityYear="1987" box="[169,268,1402,1426]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8ADF76921FEAE8F9D" box="[169,268,1402,1426]" italics="true" pageId="26" pageNumber="27">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A623FFA8AC4A6921FEE68F9D" box="[276,324,1402,1426]" pageId="26" pageNumber="27">type</typeStatus>
|
||||
. The OCA absent (
|
||||
<figureCitation id="133D2A57A623FFA8AF4F6921FDC58F9D" box="[529,615,1402,1426]" captionStart="Figure 18" captionStartId="28.[128,193,1100,1124]" captionTargetBox="[129,1473,144,1073]" captionTargetId="figure-424@28.[129,1473,144,1073]" captionTargetPageId="28" captionText="Figure 18. Platicrista angustata (Murray, 1905) in Murray (1905b) (PCM): A, neotype habitus in ventral view; B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243784" httpUri="https://zenodo.org/record/14243784/files/figure.png" pageId="26" pageNumber="27">Fig. 18B</figureCitation>
|
||||
). Furcae of the
|
||||
<taxonomicName id="4C064D51A623FFA8ADDF69C2FF468FBE" authorityName="Pilato" authorityYear="1987" box="[129,228,1433,1457]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF69C2FF468FBE" box="[129,228,1433,1457]" italics="true" pageId="26" pageNumber="27">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A623FFA8ADB569C2FEB88FBE" box="[235,282,1433,1457]" pageId="26" pageNumber="27">type</typeStatus>
|
||||
, with branches pointing backwards. Pharyngeal apophyses absent; the pharyngeal tube is only slightly enlarged at its posterior end (
|
||||
<figureCitation id="133D2A57A623FFA8AC136983FE3D8FFF" box="[333,415,1496,1520]" captionStart="Figure 18" captionStartId="28.[128,193,1100,1124]" captionTargetBox="[129,1473,144,1073]" captionTargetId="figure-424@28.[129,1473,144,1073]" captionTargetPageId="28" captionText="Figure 18. Platicrista angustata (Murray, 1905) in Murray (1905b) (PCM): A, neotype habitus in ventral view; B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243784" httpUri="https://zenodo.org/record/14243784/files/figure.png" pageId="26" pageNumber="27">Fig. 18B</figureCitation>
|
||||
). Pharynx oval and broad, with two thin macroplacoids, the second being at least two times longer than the first (1 <2). The macroplacoid margins are serrated (
|
||||
<figureCitation id="133D2A57A623FFA8ADD56A6DFF7C8C41" box="[139,222,1590,1614]" captionStart="Figure 18" captionStartId="28.[128,193,1100,1124]" captionTargetBox="[129,1473,144,1073]" captionTargetId="figure-424@28.[129,1473,144,1073]" captionTargetPageId="28" captionText="Figure 18. Platicrista angustata (Murray, 1905) in Murray (1905b) (PCM): A, neotype habitus in ventral view; B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243784" httpUri="https://zenodo.org/record/14243784/files/figure.png" pageId="26" pageNumber="27">Fig. 18B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8ADC26A0EFE4D8D26" blockId="26.[128,780,1120,1833]" pageId="26" pageNumber="27">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A623FFA8AC7B6A0EFE278C62" authorityName="Ehrenberg" authorityYear="1848" box="[293,389,1621,1645]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8AC7B6A0EFE278C62" box="[293,389,1621,1645]" italics="true" pageId="26" pageNumber="27">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A623FFA8ACD56A0EFE188C62" box="[395,442,1621,1645]" pageId="26" pageNumber="27">type</typeStatus>
|
||||
, large and robust, with thick but barely divergent accessory points, acute and slender at their tips. Primary branches of internal and anterior claws particularly massive (
|
||||
<figureCitation id="133D2A57A623FFA8ADE76AE8FEA38CC4" box="[185,257,1715,1739]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Fig. 20</figureCitation>
|
||||
). Anterior and posterior claw bases with irregular margins under PCM (
|
||||
<figureCitation id="133D2A57A623FFA8AC2A6A89FE6F8CE5" box="[372,461,1746,1770]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Fig. 20B</figureCitation>
|
||||
); delicate indentation visible in SEM (
|
||||
<figureCitation id="133D2A57A623FFA8ADBC6AA9FE9A8D05" box="[226,312,1778,1802]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Fig. 20E</figureCitation>
|
||||
). Pseudolunulae present and well developed (
|
||||
<figureCitation id="133D2A57A623FFA8ADD56B4AFF598D27" box="[139,251,1809,1833]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Fig. 20A, B</figureCitation>
|
||||
). Cuticular bars absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A623FFA8ADDF6B11FD888DCF" pageId="26" pageNumber="27" type="etymology">
|
||||
<paragraph id="8BB936D2A623FFA8ADDF6B11FD888DCF" blockId="26.[129,778,1866,1985]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8ADDF6B11FF528D6D" box="[129,240,1866,1890]" italics="true" pageId="26" pageNumber="27">Etymology:</emphasis>
|
||||
From the Latin
|
||||
<taxonomicName id="4C064D51A623FFA8ACFF6B10FDAB8D6D" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[417,521,1867,1890]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustatus">
|
||||
<emphasis id="B972EAC0A623FFA8ACFF6B10FDAB8D6D" box="[417,521,1867,1890]" italics="true" pageId="26" pageNumber="27">angustatus</emphasis>
|
||||
</taxonomicName>
|
||||
= narrowed, most probably referring to the tendency, as described by Murray, of the animal to taper from legs III towards the anterior end of the body. An adjective in nominative singular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A623FFA8AE676CCBFBCB8830" pageId="26" pageNumber="27" type="diagnosis">
|
||||
<paragraph id="8BB936D2A623FFA8AE676CCBFC298AE8" blockId="26.[825,1474,144,231]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B972EAC0A623FFA8AE676CCBFBAC8AA7" box="[825,1038,144,168]" italics="true" pageId="26" pageNumber="27">Differential diagnosis:</emphasis>
|
||||
All species of
|
||||
<taxonomicName id="4C064D51A623FFA8A9F46CCBFAAF8AA7" authorityName="Pilato" authorityYear="1987" box="[1194,1293,144,168]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A623FFA8A9F46CCBFAAF8AA7" box="[1194,1293,144,168]" italics="true" pageId="26" pageNumber="27">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
that also lack cuticular bars on their legs are compared with
|
||||
<taxonomicName id="4C064D51A623FFA8A87C6CEBFA3F8AC8" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[1314,1437,176,199]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8A87C6CEBFA3F8AC8" box="[1314,1437,176,199]" italics="true" pageId="26" pageNumber="27">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
, as follows:
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8AE0A6D55FAAE8B4A" blockId="26.[852,1474,269,575]" pageId="26" pageNumber="27">
|
||||
•
|
||||
<taxonomicName id="4C064D51A623FFA8AEDA6D55FC5B8B2A" ID-CoL="4JNBW" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[900,1017,270,293]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8AEDA6D55FC5B8B2A" box="[900,1017,270,293]" italics="true" pageId="26" pageNumber="27">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C064D51A623FFA8A9226D55FADF8B2A" authority="(Lisi et al., 2019)" baseAuthorityName="Lisi" baseAuthorityYear="2019" box="[1148,1405,269,293]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="aluna">
|
||||
<emphasis id="B972EAC0A623FFA8A9226D55FB6B8B2A" box="[1148,1225,269,293]" italics="true" pageId="26" pageNumber="27">P. aluna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A623FFA8A9876D56FAD08B2A" author="Lisi O & Daza A & Londono R" box="[1241,1394,269,293]" pageId="26" pageNumber="27" pagination="1 - 20" refId="ref37545" refString="Lisi O, Daza A, Londono R et al. Meplitumen aluna gen. nov., sp. nov. an interesting eutardigrade (Hypsibiidae, Itaquasconinae) from the Sierra Nevada de Santa Marta, Colombia. ZooKeys 2019; 865: 1 - 20. https: // doi. org / 10.3897 / zookeys. 865.30705" type="journal article" year="2019">
|
||||
Lisi
|
||||
<emphasis id="B972EAC0A623FFA8A85B6D55FA968B2A" box="[1285,1332,269,293]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
|
||||
, 2019
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
by the absence of striation of the buccal tube;
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8AE0A6D17FABA8BAC" blockId="26.[852,1474,269,575]" pageId="26" pageNumber="27">
|
||||
•
|
||||
<taxonomicName id="4C064D51A623FFA8AEDA6D16FC588B6B" ID-CoL="4JNBW" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[900,1018,333,356]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8AEDA6D16FC588B6B" box="[900,1018,333,356]" italics="true" pageId="26" pageNumber="27">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C064D51A623FFA8A9DE6D16FB5C8B6B" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1152,1278,332,356]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="carpathica" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A623FFA8A9DE6D16FB5C8B6B" box="[1152,1278,332,356]" italics="true" pageId="26" pageNumber="27">P. carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A623FFA8A85D6D17FAF78B6B" bold="true" box="[1283,1365,332,356]" pageId="26" pageNumber="27">
|
||||
<taxonomicNameLabel id="A24157BBA623FFA8A85D6D17FAF78B6B" box="[1283,1365,332,356]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
by the absence of poorly developed indentation on all claw bases (compare
|
||||
<figureCitation id="133D2A57A623FFA8AEB26DD0FBFF8BAD" box="[1004,1117,395,419]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Fig. 20A, B</figureCitation>
|
||||
with
|
||||
<figureCitation id="133D2A57A623FFA8A9C86DD0FAA58BAC" box="[1174,1287,394,419]" captionStart="Figure 25" captionStartId="36.[129,194,711,735]" captionTargetBox="[129,1473,144,684]" captionTargetId="figure-167@36.[129,1473,144,684]" captionTargetPageId="36" captionText="Figure 25. Caudal wrinkling of Platicrista carpathica sp. nov. (paratype, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243804" httpUri="https://zenodo.org/record/14243804/files/figure.png" pageId="26" pageNumber="27">Fig. 25B, C</figureCitation>
|
||||
);
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8AE0A6DF1FA008BEE" blockId="26.[852,1474,269,575]" pageId="26" pageNumber="27">
|
||||
•
|
||||
<taxonomicName id="4C064D51A623FFA8AEDA6DF0FC5E8BCD" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[900,1020,427,450]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8AEDA6DF0FC5E8BCD" box="[900,1020,427,450]" italics="true" pageId="26" pageNumber="27">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C064D51A623FFA8A9D96DF0FB5A8BCD" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" box="[1159,1272,426,450]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A623FFA8A9D96DF0FB5A8BCD" box="[1159,1272,426,450]" italics="true" pageId="26" pageNumber="27">P. horribilis</emphasis>
|
||||
</taxonomicName>
|
||||
by the presence of smooth pseudolunulae (compare
|
||||
<figureCitation id="133D2A57A623FFA8A9846D92FA918BEE" box="[1242,1331,457,481]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Figs 20B</figureCitation>
|
||||
and
|
||||
<figureCitation id="133D2A57A623FFA8A83D6D92FA328BEE" box="[1379,1424,457,481]" captionStart="Figure 27" captionStartId="37.[113,178,1027,1051]" captionTargetBox="[116,1455,146,996]" captionTargetId="figure-131@37.[114,1459,144,1000]" captionTargetPageId="37" captionText="Figure 27. Detailed morphology of Platicrista horribilis Kaczmarek & Michalczyk, 2003 (PCM): A, buccopharyngeal apparatus in toto; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="26" pageNumber="27">27D</figureCitation>
|
||||
);
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A623FFA8AE0A6DB2FBCB8830" blockId="26.[852,1474,269,575]" pageId="26" pageNumber="27">
|
||||
•
|
||||
<taxonomicName id="4C064D51A623FFA8AEDA6DB2FC5B880F" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[900,1017,489,512]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A623FFA8AEDA6DB2FC5B880F" box="[900,1017,489,512]" italics="true" pageId="26" pageNumber="27">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C064D51A623FFA8A9236DB2FB64880F" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1149,1222,489,512]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="27" phylum="Tardigrada" rank="species" species="nivea" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A623FFA8A9236DB2FB64880F" box="[1149,1222,489,512]" italics="true" pageId="26" pageNumber="27">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A623FFA8A9956DB2FABF880E" bold="true" box="[1227,1309,489,513]" pageId="26" pageNumber="27">
|
||||
<taxonomicNameLabel id="A24157BBA623FFA8A9956DB2FABF880E" box="[1227,1309,489,513]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
by the presence of poorly developed indentation of claw IV bases (compare
|
||||
<figureCitation id="133D2A57A623FFA8AEEB6E7CFBAC8830" box="[949,1038,551,575]" captionStart="Figure 20" captionStartId="30.[129,194,1054,1078]" captionTargetBox="[161,1441,144,1026]" captionTargetId="figure-441@30.[161,1441,144,1026]" captionTargetPageId="30" captionText="Figure 20. Claws of Platicrista angustata: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="26" pageNumber="27">Figs 20B</figureCitation>
|
||||
and
|
||||
<figureCitation id="133D2A57A623FFA8A9606E7CFBFA8830" box="[1086,1112,551,575]" captionStart="Figure 28" captionStartId="37.[113,178,1674,1698]" captionTargetBox="[117,1455,1148,1642]" captionTargetId="figure-174@37.[114,1459,1146,1646]" captionTargetPageId="37" captionText="Figure 28. General morphology of Platicrista nivea sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243810" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="26" pageNumber="27">28</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1094
data/03/AF/87/03AF87C4A623FFA9AE006E25FB4A8DBA.xml
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1094
data/03/AF/87/03AF87C4A623FFA9AE006E25FB4A8DBA.xml
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274
data/03/AF/87/03AF87C4A625FFACAEF76AEFFBDE8C07.xml
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274
data/03/AF/87/03AF87C4A625FFACAEF76AEFFBDE8C07.xml
Normal file
|
|
@ -0,0 +1,274 @@
|
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<mods:mods id="E5B64A5C6CF8D1B247F3E5A3F2C29261" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="22ACC16EF0EB1685A183A08B04C97388">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="446C133592A8ACD6989EB3D5386AB468">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="2D3C35515ECD0CA60FEFCD8EC1D79BF3">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="5AF94475E79FBB8E8F59A0E9B3CFFD0B" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="56BE94F476BE40090ACF39086B090D8F">Blagden, Brian</mods:namePart>
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<mods:affiliation id="B068190399A9F68F23F65A08C4B35B47">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="B8B8F62F97A620B2130F251E0AA2FA38">Morek, Witold</mods:namePart>
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<mods:affiliation id="E9BA18703B9B4FEE9E6C433C40539008">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="CF5328D1D997257B4E4109311D8C834B">Michalczyk, Łukasz</mods:namePart>
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<mods:part id="BEAD551F5E8733CC9AA6B24056864670">
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<mods:date id="945DC1D3024E699D6A57FB22F1FFD592">2024</mods:date>
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|
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<treatment id="03AF87C4A625FFACAEF76AEFFBDE8C07" LSID="urn:lsid:plazi:treatment:03AF87C4A625FFACAEF76AEFFBDE8C07" httpUri="http://treatment.plazi.org/id/03AF87C4A625FFACAEF76AEFFBDE8C07" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
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<subSubSection id="C31C6559A625FFAEAEF76AEFFAF08CC1" box="[937,1362,1716,1743]" pageId="28" pageNumber="29" type="nomenclature">
|
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<paragraph id="8BB936D2A625FFAEAEF76AEFFAF08CC1" blockId="28.[825,1474,1716,1810]" box="[937,1362,1716,1743]" pageId="28" pageNumber="29">
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<taxonomicName id="4C064D51A625FFAEAEF76AEFFAA38CC1" authority="Gasiorek" authorityName="Gasiorek" authorityYear="2024" box="[937,1281,1716,1743]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="species" species="carpathica" status="sp. nov.">
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<emphasis id="B972EAC0A625FFAEAEF76AEFFB3F8CC0" bold="true" box="[937,1181,1716,1743]" italics="true" pageId="28" pageNumber="29">Platicrista carpathica</emphasis>
|
||||
Gąsiorek
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA625FFAEA8586AEFFAF08CC1" box="[1286,1362,1716,1742]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A625FFAEAE676A80FB3C8D1D" pageId="28" pageNumber="29" type="description">
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||||
ZooBank registration:
|
||||
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urn:lsid:zoobank.org:act:
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|
||||
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|
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|
||||
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|
||||
<paragraph id="8BB936D2A625FFAEAE676B72FB008D6E" blockId="28.[825,1474,1833,1889]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEAE676B72FBA08D4E" box="[825,1026,1833,1857]" italics="true" pageId="28" pageNumber="29">Material examined:</emphasis>
|
||||
24 individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A625FFAEA9646B12FB338D6E" box="[1082,1169,1865,1889]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="28" pageNumber="29">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A625FFAFAE676B25FD3A8DB9" blockId="28.[825,1474,1917,2004]" lastBlockId="29.[113,762,1887,1974]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEAE676B25FC6F8D9A" box="[825,973,1917,1941]" italics="true" pageId="28" pageNumber="29">Type material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA625FFAFAE866B26FD3A8DB9" collectionCode="PL" country="Poland" lastPageId="29" lastPageNumber="30" location="Tatra Mountains" pageId="28" pageNumber="29" specimenCode="PL.188.01, PL.088.01, PL.188.02, PL.201.01, PL.245.01-3, PL.249.01, PL.287.03, PL.296.01, PL.373.01" specimenCount="19" typeStatus="holotype">
|
||||
<typeStatus id="54BD8870A625FFAEAE866B26FB998D9A" box="[984,1083,1917,1941]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||||
(slide
|
||||
<specimenCode id="DBA09EA9A625FFAEA9D76B26FB558D9A" box="[1161,1271,1917,1941]" pageId="28" pageNumber="29">PL.188.01</specimenCode>
|
||||
) and
|
||||
<specimenCount id="9D00FD5BA625FFAEA8656B26FAF78D9A" box="[1339,1365,1917,1941]" pageId="28" pageNumber="29">18</specimenCount>
|
||||
<typeStatus id="54BD8870A625FFAEA8016B25FA608D99" box="[1375,1474,1918,1942]" pageId="28" pageNumber="29">paratypes</typeStatus>
|
||||
(slides
|
||||
<specimenCode id="DBA09EA9A625FFAEAEC96BC6FBA18DBA" box="[919,1027,1949,1973]" pageId="28" pageNumber="29">PL.088.01</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEA97C6BC6FB2C8DBA" box="[1058,1166,1949,1973]" pageId="28" pageNumber="29">PL.188.02</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEA9F36BC6FABB8DBA" box="[1197,1305,1949,1973]" pageId="28" pageNumber="29">PL.201.01</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEA8696BC7FA1C8DBB" box="[1335,1470,1948,1972]" pageId="28" pageNumber="29">PL.245.01–3</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEAE676BE7FC048DDB" box="[825,934,1980,2004]" pageId="28" pageNumber="29">PL.249.01</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEAEF06BE7FBB88DDB" box="[942,1050,1980,2004]" pageId="28" pageNumber="29">PL.287.03</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBA09EA9A625FFAEA97D6BE7FB328DDB" box="[1059,1168,1980,2004]" pageId="28" pageNumber="29">PL.296.01</specimenCode>
|
||||
, and
|
||||
<specimenCode id="DBA09EA9A625FFAEA99D6BE7FA908DDB" box="[1219,1330,1980,2004]" pageId="28" pageNumber="29">PL.373.01</specimenCode>
|
||||
) from
|
||||
<collectingCountry id="F3117642A625FFAEA82B6BE7FA1D8DDB" box="[1397,1471,1980,2004]" name="Poland" pageId="28" pageNumber="29">Poland</collectingCountry>
|
||||
, various locales in the
|
||||
<location id="8ED96009A624FFAFAC166B3BFE538D78" LSID="urn:lsid:plazi:treatment:03AF87C4A625FFACAEF76AEFFBDE8C07:8ED96009A624FFAFAC166B3BFE538D78" box="[328,497,1887,1911]" country="Poland" name="Tatra Mountains" pageId="29" pageNumber="30">Tatra Mountains</location>
|
||||
(Supporting Information,
|
||||
<tableCitation id="C6840369A624FFAFAD2F6B24FF708D98" box="[113,210,1919,1943]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="29" pageNumber="30">Table S1</tableCitation>
|
||||
), are deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University, Kraków.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF79665AA624FFAFAD2C6B76FBEF8D4A" ID-DOI="http://doi.org/10.5281/zenodo.14243790" ID-Zenodo-Dep="14243790" box="[114,1101,1837,1861]" httpUri="https://zenodo.org/record/14243790/files/figure.png" pageId="29" pageNumber="30" startId="29.[114,179,1837,1861]" targetBox="[332,1241,144,1809]" targetPageId="29" targetType="figure">
|
||||
<paragraph id="8BB936D2A624FFAFAD2C6B76FBEF8D4A" blockId="29.[114,1101,1837,1861]" box="[114,1101,1837,1861]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B972EAC0A624FFAFAD2C6B76FF748D4A" bold="true" box="[114,214,1837,1861]" pageId="29" pageNumber="30">Figure 19.</emphasis>
|
||||
Caudal sculpturing of
|
||||
<taxonomicName id="4C064D51A624FFAFACF56B76FD168D4A" authority="(PCM)" baseAuthorityName="PCM" box="[427,692,1837,1861]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="30" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A624FFAFACF56B76FDC18D4A" box="[427,611,1837,1861]" italics="true" pageId="29" pageNumber="30">Platicrista angustata</emphasis>
|
||||
(PCM)
|
||||
</taxonomicName>
|
||||
: A, juvenile; B, C, adults. Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C31C6559A624FFACAE746B3BFD938C8A" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||||
<paragraph id="8BB936D2A624FFACAE746B3BFD5F8FA5" blockId="29.[810,1461,1887,1974]" lastBlockId="30.[128,779,1175,1669]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||||
<emphasis id="B972EAC0A624FFAFAE746B3BFC038D78" box="[810,929,1888,1911]" italics="true" pageId="29" pageNumber="30">Description:</emphasis>
|
||||
Body massive (
|
||||
<tableCitation id="C6840369A624FFAFA9156B04FB3B8D77" box="[1099,1177,1887,1912]" captionStart="Table 8" captionStartId="34.[129,184,143,167]" captionTargetBox="[129,1466,217,1687]" captionTargetPageId="34" captionText="Table 8. Measurements (in micrometres) of selected morphological structures of type series (PL.088 + PL.188 + PL.201 + PL.245 + PL.249 + PL.287 + PL.296 + PL.373) of Platicrista carpathicasp. nov. from Poland mounted in Hoyer’s medium." pageId="29" pageNumber="30">Table 8</tableCitation>
|
||||
), white and elongate, with a blunt cephalic part (
|
||||
<figureCitation id="133D2A57A624FFAFA9516B24FBF48D98" box="[1039,1110,1919,1943]" captionStart="Figure 24" captionStartId="35.[113,178,1954,1978]" captionTargetBox="[114,1458,1456,1927]" captionTargetId="figure-181@35.[114,1458,1456,1927]" captionTargetPageId="35" captionText="Figure 24. General morphology of Platicrista carpathica sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243801" httpUri="https://zenodo.org/record/14243801/files/figure.png" pageId="29" pageNumber="30">Fig. 24</figureCitation>
|
||||
). Cuticle without pores, but with irregular wrinkling caudally and rudimentary granulation in the lateralmost portions of the dorsocaudal body part (
|
||||
<figureCitation id="133D2A57A627FFACAFF068CCFD5B8EA0" box="[686,761,1175,1199]" captionStart="Figure 25" captionStartId="36.[129,194,711,735]" captionTargetBox="[129,1473,144,684]" captionTargetId="figure-167@36.[129,1473,144,684]" captionTargetPageId="36" captionText="Figure 25. Caudal wrinkling of Platicrista carpathica sp. nov. (paratype, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243804" httpUri="https://zenodo.org/record/14243804/files/figure.png" pageId="30" pageNumber="31">Fig. 25</figureCitation>
|
||||
). Cribriform areas not visible under PCM. Legs short and plump. Eyes absent in unmounted animals. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A627FFACAD9D68AEFE848F02" authorityName="Pilato" authorityYear="1987" box="[195,294,1269,1293]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A627FFACAD9D68AEFE848F02" box="[195,294,1269,1293]" italics="true" pageId="30" pageNumber="31">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A627FFACAC7568AEFEFB8F02" box="[299,345,1269,1293]" pageId="30" pageNumber="31">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A627FFACAC3768AEFE1D8F02" box="[361,447,1269,1293]" captionStart="Figure 26" captionStartId="36.[129,194,1670,1694]" captionTargetBox="[131,1470,789,1638]" captionTargetId="figure-119@36.[129,1473,786,1641]" captionTargetPageId="36" captionText="Figure 26. Detailed morphology of Platicrista carpathica sp. nov. (paratypes, PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II; D, claws III. Empty white arrowheads indicate pseudolunulae; white asterisk indicates internal pulvinus; and black asterisk indicates external pulvinus. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243806" httpUri="https://zenodo.org/record/14243806/files/figure.png" pageId="30" pageNumber="31">Fig. 26A</figureCitation>
|
||||
): OCA not visible under PCM; annulation of the pharyngeal tube simple and comprising tightly merged annuli; stylet supports inserted at the junction between the buccal and pharyngeal tubes; furcae of the
|
||||
<taxonomicName id="4C064D51A627FFACAF326908FD6D8F64" authorityName="Pilato" authorityYear="1987" box="[620,719,1363,1387]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A627FFACAF326908FD6D8F64" box="[620,719,1363,1387]" italics="true" pageId="30" pageNumber="31">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A627FFACAF8A6908FCA78F64" box="[724,773,1363,1387]" pageId="30" pageNumber="31">type</typeStatus>
|
||||
; pharyngeal apophyses absent; pharynx oval, with two bar-like macroplacoids: the first much shorter than the second (1 <2).
|
||||
</paragraph>
|
||||
<caption id="DF79665AA627FFACADDF6845FBB38E5D" ID-DOI="http://doi.org/10.5281/zenodo.14243792" ID-Zenodo-Dep="14243792" httpUri="https://zenodo.org/record/14243792/files/figure.png" pageId="30" pageNumber="31" startId="30.[129,194,1054,1078]" targetBox="[161,1441,144,1026]" targetPageId="30" targetType="figure">
|
||||
<paragraph id="8BB936D2A627FFACADDF6845FBB38E5D" blockId="30.[129,1463,1054,1107]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACADDF6845FF478E39" bold="true" box="[129,229,1054,1078]" pageId="30" pageNumber="31">Figure 20.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A627FFACAC616845FE5A8E39" box="[319,504,1054,1078]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A627FFACAC616845FE5A8E39" box="[319,504,1054,1078]" italics="true" pageId="30" pageNumber="31">Platicrista angustata</emphasis>
|
||||
</taxonomicName>
|
||||
: A, claws III (neotype, PCM); B, claws IV (neotype, PCM); C, claws I (SEM); D, claws III (SEM); E, claws IV (SEM). Empty white arrowheads indicate pseudolunulae. Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A627FFACADC269EAFD938C8A" blockId="30.[128,779,1175,1669]" pageId="30" pageNumber="31">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A627FFACAC7A69EAFE268FC6" authorityName="Ehrenberg" authorityYear="1848" box="[292,388,1457,1481]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A627FFACAC7A69EAFE268FC6" box="[292,388,1457,1481]" italics="true" pageId="30" pageNumber="31">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A627FFACACD769EAFE1A8FC6" box="[393,440,1457,1481]" pageId="30" pageNumber="31">type</typeStatus>
|
||||
, large and slender (
|
||||
<figureCitation id="133D2A57A627FFACAFDC69EAFD5A8FC6" box="[642,760,1457,1481]" captionStart="Figure 26" captionStartId="36.[129,194,1670,1694]" captionTargetBox="[131,1470,789,1638]" captionTargetId="figure-119@36.[129,1473,786,1641]" captionTargetPageId="36" captionText="Figure 26. Detailed morphology of Platicrista carpathica sp. nov. (paratypes, PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II; D, claws III. Empty white arrowheads indicate pseudolunulae; white asterisk indicates internal pulvinus; and black asterisk indicates external pulvinus. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243806" httpUri="https://zenodo.org/record/14243806/files/figure.png" pageId="30" pageNumber="31">Fig. 26B–D</figureCitation>
|
||||
). Accessory points tightly adjacent to primary branches of the external/posterior claws, but with tips divergent from the primary branches of the internal/anterior claws. A weakly developed indentation on all claw bases. Septa between claw parts absent. Pseudolunulae smooth. Cuticular bars absent. Pulvini visible when legs are oriented suitably (
|
||||
<figureCitation id="133D2A57A627FFACAC976A36FD828C8A" box="[457,544,1645,1669]" captionStart="Figure 26" captionStartId="36.[129,194,1670,1694]" captionTargetBox="[131,1470,789,1638]" captionTargetId="figure-119@36.[129,1473,786,1641]" captionTargetPageId="36" captionText="Figure 26. Detailed morphology of Platicrista carpathica sp. nov. (paratypes, PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II; D, claws III. Empty white arrowheads indicate pseudolunulae; white asterisk indicates internal pulvinus; and black asterisk indicates external pulvinus. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243806" httpUri="https://zenodo.org/record/14243806/files/figure.png" pageId="30" pageNumber="31">Fig. 26C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A627FFACADDF6AF7FE828CEB" pageId="30" pageNumber="31" type="etymology">
|
||||
<paragraph id="8BB936D2A627FFACADDF6AF7FE828CEB" blockId="30.[129,778,1708,1764]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACADDF6AF7FF528CCB" box="[129,240,1708,1732]" italics="true" pageId="30" pageNumber="31">Etymology:</emphasis>
|
||||
The name signifies
|
||||
<emphasis id="B972EAC0A627FFACACE76AF6FD8A8CCB" box="[441,552,1709,1732]" italics="true" pageId="30" pageNumber="31">terra typica</emphasis>
|
||||
. An adjective in nominative singular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A627FFACADDF6B5EFBDE8C07" pageId="30" pageNumber="31" type="diagnosis">
|
||||
<paragraph id="8BB936D2A627FFACADDF6B5EFA288EA0" blockId="30.[129,778,1797,1978]" lastBlockId="30.[852,1474,1175,1544]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACADDF6B5EFECE8D12" box="[129,364,1797,1821]" italics="true" pageId="30" pageNumber="31">Differential diagnosis:</emphasis>
|
||||
There are four congeneric species characterized by an absence of cuticular bars on their legs and the presence of a poorly developed or entirely absent dorsal granulation:
|
||||
<taxonomicName id="4C064D51A627FFACAC096B3FFE0B8D74" box="[343,425,1891,1915]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="aluna">
|
||||
<emphasis id="B972EAC0A627FFACAC096B3FFE0B8D74" box="[343,425,1891,1915]" italics="true" pageId="30" pageNumber="31">P. aluna</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A627FFACACE86B3FFD968D74" box="[438,564,1892,1915]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A627FFACACE86B3FFD968D74" box="[438,564,1892,1915]" italics="true" pageId="30" pageNumber="31">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A627FFACAF1F6B3FFD328D74" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[577,656,1892,1915]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A627FFACAF1F6B3FFD328D74" box="[577,656,1892,1915]" italics="true" pageId="30" pageNumber="31">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A627FFACAF966B3FFF1E8D95" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A627FFACAF966B3FFF1E8D95" italics="true" pageId="30" pageNumber="31">P. horribilis</emphasis>
|
||||
</taxonomicName>
|
||||
(granulation absent).
|
||||
<taxonomicName id="4C064D51A627FFACACEC6BD9FD338D95" authorityName="Gasiorek" authorityYear="2024" box="[434,657,1922,1946]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="carpathica">
|
||||
<emphasis id="B972EAC0A627FFACACEC6BD9FD338D95" box="[434,657,1922,1946]" italics="true" pageId="30" pageNumber="31">Platicrista carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
can be differentiated from them based on: •
|
||||
<taxonomicName id="4C064D51A627FFACAEDA68C3FC738EA0" box="[900,977,1175,1199]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="aluna">
|
||||
<emphasis id="B972EAC0A627FFACAEDA68C3FC738EA0" box="[900,977,1175,1199]" italics="true" pageId="30" pageNumber="31">P. aluna</emphasis>
|
||||
</taxonomicName>
|
||||
, the absence of striation of the buccal tube;
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A627FFACAE0A68ECFBCF8EC1" blockId="30.[852,1474,1175,1544]" box="[852,1133,1206,1230]" pageId="30" pageNumber="31">
|
||||
•
|
||||
<taxonomicName id="4C064D51A627FFACAEDA68ECFC5B8EC1" box="[900,1017,1207,1230]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A627FFACAEDA68ECFC5B8EC1" box="[900,1017,1207,1230]" italics="true" pageId="30" pageNumber="31">P. angustata</emphasis>
|
||||
</taxonomicName>
|
||||
, see above;
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A627FFACAE0A688DFABE8F85" blockId="30.[852,1474,1175,1544]" pageId="30" pageNumber="31">
|
||||
•
|
||||
<taxonomicName id="4C064D51A627FFACAEDA688DFC518EE1" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" box="[900,1011,1238,1262]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A627FFACAEDA688DFC518EE1" box="[900,1011,1238,1262]" italics="true" pageId="30" pageNumber="31">P. horribilis</emphasis>
|
||||
</taxonomicName>
|
||||
, the indentation of the pseudolunulae under claws IV (absent in
|
||||
<taxonomicName id="4C064D51A627FFACA90868ADFB748F02" authorityName="Gasiorek" authorityYear="2024" box="[1110,1238,1269,1293]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="carpathica">
|
||||
<emphasis id="B972EAC0A627FFACA90868ADFB748F02" box="[1110,1238,1269,1293]" italics="true" pageId="30" pageNumber="31">P. carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
vs. present in
|
||||
<taxonomicName id="4C064D51A627FFACA82F68ADFC098F23" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A627FFACA82F68ADFC098F23" italics="true" pageId="30" pageNumber="31">P. horribilis</emphasis>
|
||||
</taxonomicName>
|
||||
) and the divergence of accessory points from the primary branches of the external and posterior claws (not divergent in
|
||||
<taxonomicName id="4C064D51A627FFACA958690FFB268F64" authorityName="Gasiorek" authorityYear="2024" box="[1030,1156,1363,1387]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="carpathica">
|
||||
<emphasis id="B972EAC0A627FFACA958690FFB268F64" box="[1030,1156,1363,1387]" italics="true" pageId="30" pageNumber="31">P. carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
vs. clearly divergent in
|
||||
<taxonomicName id="4C064D51A627FFACA82A690FFC098F85" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A627FFACA82A690FFC098F85" italics="true" pageId="30" pageNumber="31">P.horribilis</emphasis>
|
||||
</taxonomicName>
|
||||
; compare
|
||||
<figureCitation id="133D2A57A627FFACA9486929FB328F85" box="[1046,1168,1394,1418]" captionStart="Figure 26" captionStartId="36.[129,194,1670,1694]" captionTargetBox="[131,1470,789,1638]" captionTargetId="figure-119@36.[129,1473,786,1641]" captionTargetPageId="36" captionText="Figure 26. Detailed morphology of Platicrista carpathica sp. nov. (paratypes, PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II; D, claws III. Empty white arrowheads indicate pseudolunulae; white asterisk indicates internal pulvinus; and black asterisk indicates external pulvinus. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243806" httpUri="https://zenodo.org/record/14243806/files/figure.png" pageId="30" pageNumber="31">Figs 26B–D</figureCitation>
|
||||
and
|
||||
<figureCitation id="133D2A57A627FFACA99E6929FAA98F85" box="[1216,1291,1394,1418]" captionStart="Figure 27" captionStartId="37.[113,178,1027,1051]" captionTargetBox="[116,1455,146,996]" captionTargetId="figure-131@37.[114,1459,144,1000]" captionTargetPageId="37" captionText="Figure 27. Detailed morphology of Platicrista horribilis Kaczmarek & Michalczyk, 2003 (PCM): A, buccopharyngeal apparatus in toto; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="30" pageNumber="31">27B–D</figureCitation>
|
||||
);
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A627FFACAE0A69C9FBDE8C07" blockId="30.[852,1474,1175,1544]" pageId="30" pageNumber="31">
|
||||
•
|
||||
<taxonomicName id="4C064D51A627FFACAEDA69C9FC6F8FA6" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[900,973,1426,1449]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A627FFACAEDA69C9FC6F8FA6" box="[900,973,1426,1449]" italics="true" pageId="30" pageNumber="31">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
, the presence and form of the caudal granulation (weakly developed only in the lateralmost part of caudal trunk in
|
||||
<taxonomicName id="4C064D51A627FFACAE84698AFBF48FE7" authorityName="Gasiorek" authorityYear="2024" box="[986,1110,1488,1512]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="carpathica">
|
||||
<emphasis id="B972EAC0A627FFACAE84698AFBF48FE7" box="[986,1110,1488,1512]" italics="true" pageId="30" pageNumber="31">P. carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
vs. strongly developed on the entire caudal trunk in
|
||||
<taxonomicName id="4C064D51A627FFACA97C69ABFBC98C08" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1058,1131,1520,1543]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A627FFACA97C69ABFBC98C08" box="[1058,1131,1520,1543]" italics="true" pageId="30" pageNumber="31">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
178
data/03/AF/87/03AF87C4A625FFAEAD8568EBFA928C96.xml
Normal file
178
data/03/AF/87/03AF87C4A625FFAEAD8568EBFA928C96.xml
Normal file
|
|
@ -0,0 +1,178 @@
|
|||
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<mods:mods id="14DF89CE9D39F28F77299B578961D560" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0B21E3B2AF1A4FB06F7C4D2FD26B0BB2">
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<mods:title id="1697DBF413D4068592CCF7B7CACAECD1">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="E402CEED5AD7C64ECF1FF9903335848B">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="7F4C87AC94C94537CDCA85AD690336A1">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
<mods:nameIdentifier id="0D3A47C140A822B80F03EBFD459470B5" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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</mods:role>
|
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<mods:namePart id="00A92CBCDECD07E2392C9C99BA415AE8">Blagden, Brian</mods:namePart>
|
||||
<mods:affiliation id="F8612C3B3F796FFBCC45DF83A99D8DF0">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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</mods:role>
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<mods:namePart id="C5F186945AD75DB07887B8843AE80FB0">Morek, Witold</mods:namePart>
|
||||
<mods:affiliation id="A7EBAAD2FC79FE8986E2CA6DFA21127E">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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</mods:role>
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<mods:namePart id="DA8CD81016B6CAE4BB56BEDC9869B7A5">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="61695579242EE3CFA180A08D541CCD73">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:title id="A415EF660040DE08B9159A40AAC4F044">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="2AD66A0DDF2B455C7F9097012E665B5B">
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<mods:date id="12D784E3F1B631F4B6E4FA40D1A6192E">2024</mods:date>
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<mods:number id="35F9E2DFE63348DE3A1BAA5CA63E235C">2023-11-14</mods:number>
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|
||||
</mods:location>
|
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<mods:classification id="64E9474B82BBFCCC8F266918DD02ACCD">journal article</mods:classification>
|
||||
<mods:identifier id="27AB9B1030E44D592AF630FD1B0619BD" type="DOI">10.1093/zoolinnean/zlad151</mods:identifier>
|
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|
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|
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<mods:identifier id="2940C73025AF9380461B7762BB8FFC79" type="ZooBank">044A402-2A0F-4135-9410-7DE081CB11C4Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03AF87C4A625FFAEAD8568EBFA928C96" LSID="urn:lsid:plazi:treatment:03AF87C4A625FFAEAD8568EBFA928C96" httpUri="http://treatment.plazi.org/id/03AF87C4A625FFAEAD8568EBFA928C96" lastPageNumber="29" pageId="28" pageNumber="29">
|
||||
<subSubSection id="C31C6559A625FFAEAD8568EBFD0D8EC5" box="[219,687,1200,1227]" pageId="28" pageNumber="29" type="nomenclature">
|
||||
<paragraph id="8BB936D2A625FFAEAD8568EBFD0D8EC5" blockId="28.[219,687,1200,1227]" box="[219,687,1200,1227]" pageId="28" pageNumber="29">
|
||||
<heading id="D0F181BEA625FFAEAD8568EBFD0D8EC5" bold="true" box="[219,687,1200,1227]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
|
||||
<taxonomicName id="4C064D51A625FFAEAD8568EBFD0D8EC5" ID-CoL="9X739" authority="Miller & J. D. Miller, 2021" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[219,687,1200,1227]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A625FFAEAD8568EBFD0D8EC5" bold="true" box="[219,687,1200,1227]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEAD8568EBFE028EC4" bold="true" box="[219,416,1200,1227]" italics="true" pageId="28" pageNumber="29">Platicrista brunsoni</emphasis>
|
||||
Miller & J.D.
|
||||
<bibRefCitation id="EF974B23A625FFAEAF7368EBFD0D8EC5" author="Miller WR & Miller JD" box="[557,687,1200,1226]" pageId="28" pageNumber="29" pagination="98 - 105" refId="ref38051" refString="Miller WR, Miller JD. Tardigrades of North America: Platicrista brunsoni nov. sp. (Parachela, Hypsibiidae, Itaquasconinae) from the Bob Marshall Wilderness Area of Montana. Northwest Science 2021; 95: 98 - 105. https: // doi. org / 10.3955 / 046.095.0106" type="journal article" year="2021">Miller, 2021</bibRefCitation>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A625FFAEADDF68AAFD6B8F27" pageId="28" pageNumber="29" type="materials_examined">
|
||||
<paragraph id="8BB936D2A625FFAEADDF68AAFD6B8F27" blockId="28.[129,777,1265,1321]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEADDF68AAFEE68F06" box="[129,324,1265,1289]" italics="true" pageId="28" pageNumber="29">Material examined:</emphasis>
|
||||
Two individuals in total (originating from northern Wyoming; loaned by courtesy of Harry Meyer).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A625FFAEADDF6912FBA18F08" pageId="28" pageNumber="29" type="description">
|
||||
<paragraph id="8BB936D2A625FFAEADDF6912FF718C32" blockId="28.[127,779,1353,2004]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEADDF6912FEC78F6E" box="[129,357,1353,1377]" italics="true" pageId="28" pageNumber="29">Amended description:</emphasis>
|
||||
Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A625FFAEADDF6932FF468F8E" authorityName="Pilato" authorityYear="1987" box="[129,228,1385,1409]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEADDF6932FF468F8E" box="[129,228,1385,1409]" italics="true" pageId="28" pageNumber="29">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A625FFAEADB66932FEB48F8E" box="[232,278,1385,1409]" pageId="28" pageNumber="29">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A625FFAEAC786932FED98F8E" box="[294,379,1385,1409]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22A</figureCitation>
|
||||
): OCA not visible under PCM; simple pharyngeal annulation composed of tightly arranged single annuli; stylet supports inserted at the junction between the buccal and pharyngeal tubes; furcae of the
|
||||
<taxonomicName id="4C064D51A625FFAEACA4699CFDFF8FD0" authorityName="Pilato" authorityYear="1987" box="[506,605,1479,1503]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEACA4699CFDFF8FD0" box="[506,605,1479,1503]" italics="true" pageId="28" pageNumber="29">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A625FFAEAF3C699CFD328FD0" box="[610,656,1479,1503]" pageId="28" pageNumber="29">type</typeStatus>
|
||||
(cochlear); pharyngeal apophyses not visible; pharynx oval, with two bar-like macroplacoids: the first much shorter than the second (1 <2).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A625FFAEADC26A1FFD2C8D37" blockId="28.[127,779,1353,2004]" pageId="28" pageNumber="29">
|
||||
Claws massive, of the
|
||||
<taxonomicName id="4C064D51A625FFAEAC286A1FFE748C53" authorityName="Ehrenberg" authorityYear="1848" box="[374,470,1604,1628]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEAC286A1FFE748C53" box="[374,470,1604,1628]" italics="true" pageId="28" pageNumber="29">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A625FFAEAC856A1EFDAB8C52" box="[475,521,1605,1629]" pageId="28" pageNumber="29">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A625FFAEAF466A1FFD2E8C53" box="[536,652,1604,1628]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22B–D</figureCitation>
|
||||
). Accessory points divergent only on the primary branches of the posterior claws (
|
||||
<figureCitation id="133D2A57A625FFAEAC466AD8FEDB8C94" box="[280,377,1667,1691]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22D</figureCitation>
|
||||
). Septa between claw parts absent. Pseudolunulae present at the internal and anterior claw bases. Irregular granulation on legs IV. Internal cuticular bars II–III with irregular margins (
|
||||
<figureCitation id="133D2A57A625FFAEAC226ABAFE758CF6" box="[380,471,1761,1785]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22C</figureCitation>
|
||||
). Internal, anterior, and posterior claw bases delicately indented (
|
||||
<figureCitation id="133D2A57A625FFAEACA76B5BFDCE8D17" box="[505,620,1792,1816]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22B–D</figureCitation>
|
||||
). Basal portion of the posterior claws greatly broadened (
|
||||
<figureCitation id="133D2A57A625FFAEAF786B44FDDC8D37" box="[550,638,1823,1848]" captionStart="Figure 22" captionStartId="33.[113,178,1025,1049]" captionTargetBox="[114,1458,144,997]" captionTargetId="figure-150@33.[114,1458,144,997]" captionTargetPageId="33" captionText="Figure 22. Detailed morphology of Platicrista brunsoni Miller & J.D. Miller, 2021 (PCM): A, buccopharyngeal apparatus in toto; B, claws I; C, claws II (black arrowhead indicates internal bar); D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243796" httpUri="https://zenodo.org/record/14243796/files/figure.png" pageId="28" pageNumber="29">Fig. 22D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A625FFAEADC26B64FBA18F08" blockId="28.[127,779,1353,2004]" lastBlockId="28.[825,1474,1201,1287]" pageId="28" pageNumber="29">
|
||||
Dorsal cuticle with large polygonal granulation that extends to the lateral body parts (including legs). Granulation is most distinct in the cephalic (
|
||||
<figureCitation id="133D2A57A625FFAEACDE6B26FE7B8D99" box="[384,473,1917,1942]" captionStart="Figure 23" captionStartId="33.[113,178,1672,1696]" captionTargetBox="[114,1458,1102,1644]" captionTargetId="figure-194@33.[114,1458,1102,1644]" captionTargetPageId="33" captionText="Figure 23. Dorsal sculpturing of Platicrista brunsoni (PCM): A, rostral part; B, C, central part; D, caudal part. Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243799" httpUri="https://zenodo.org/record/14243799/files/figure.png" pageId="28" pageNumber="29">Fig. 23A</figureCitation>
|
||||
) to median and more caudal portions of the cuticle (
|
||||
<figureCitation id="133D2A57A625FFAEAC336BC6FE1D8DBA" box="[365,447,1949,1973]" captionStart="Figure 23" captionStartId="33.[113,178,1672,1696]" captionTargetBox="[114,1458,1102,1644]" captionTargetId="figure-194@33.[114,1458,1102,1644]" captionTargetPageId="33" captionText="Figure 23. Dorsal sculpturing of Platicrista brunsoni (PCM): A, rostral part; B, C, central part; D, caudal part. Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243799" httpUri="https://zenodo.org/record/14243799/files/figure.png" pageId="28" pageNumber="29">Fig. 23B</figureCitation>
|
||||
). However, polygons may merge at the level of legs II–III, and form
|
||||
<emphasis id="B972EAC0A625FFAEAC876BE6FDAC8DDB" box="[473,526,1981,2004]" italics="true" pageId="28" pageNumber="29">striae</emphasis>
|
||||
(
|
||||
<figureCitation id="133D2A57A625FFAEAF436BE7FDD68DDB" box="[541,628,1980,2004]" captionStart="Figure 23" captionStartId="33.[113,178,1672,1696]" captionTargetBox="[114,1458,1102,1644]" captionTargetId="figure-194@33.[114,1458,1102,1644]" captionTargetPageId="33" captionText="Figure 23. Dorsal sculpturing of Platicrista brunsoni (PCM): A, rostral part; B, C, central part; D, caudal part. Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243799" httpUri="https://zenodo.org/record/14243799/files/figure.png" pageId="28" pageNumber="29">Fig. 23C</figureCitation>
|
||||
). Polygons are different in shape in the caudalmost body portion, being more convex and irregular in shape (
|
||||
<figureCitation id="133D2A57A625FFAEA925688BFB768EE7" box="[1147,1236,1232,1256]" captionStart="Figure 23" captionStartId="33.[113,178,1672,1696]" captionTargetBox="[114,1458,1102,1644]" captionTargetId="figure-194@33.[114,1458,1102,1644]" captionTargetPageId="33" captionText="Figure 23. Dorsal sculpturing of Platicrista brunsoni (PCM): A, rostral part; B, C, central part; D, caudal part. Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243799" httpUri="https://zenodo.org/record/14243799/files/figure.png" pageId="28" pageNumber="29">Fig. 23D</figureCitation>
|
||||
). Cribriform areas not visible under PCM.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A625FFAEAE676972FA928C96" pageId="28" pageNumber="29" type="discussion">
|
||||
<paragraph id="8BB936D2A625FFAEAE676972FA928C96" blockId="28.[825,1475,1321,1689]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B972EAC0A625FFAEAE676972FC358F4E" box="[825,919,1321,1345]" italics="true" pageId="28" pageNumber="29">Remarks:</emphasis>
|
||||
The original description contained several errors. The buccal tube width should not have been measured, at least not in the
|
||||
<typeStatus id="54BD8870A625FFAEAE01693CFC1F8F70" box="[863,957,1383,1407]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||||
, because it is evident from the photograph that the buccopharyngeal tube had collapsed (
|
||||
<bibRefCitation id="EF974B23A625FFAEA99769DCFA198F90" author="Miller WR & Miller JD" box="[1225,1467,1415,1439]" pageId="28" pageNumber="29" pagination="98 - 105" refId="ref38051" refString="Miller WR, Miller JD. Tardigrades of North America: Platicrista brunsoni nov. sp. (Parachela, Hypsibiidae, Itaquasconinae) from the Bob Marshall Wilderness Area of Montana. Northwest Science 2021; 95: 98 - 105. https: // doi. org / 10.3955 / 046.095.0106" type="journal article" year="2021">Miller and Miller 2021</bibRefCitation>
|
||||
: fig. 2B). This occurs frequently during mounting of
|
||||
<taxonomicName id="4C064D51A625FFAEA80769FDFA1E8FB1" authorityName="Pilato" authorityYear="1987" box="[1369,1468,1446,1470]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEA80769FDFA1E8FB1" box="[1369,1468,1446,1470]" italics="true" pageId="28" pageNumber="29">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A625FFAEAE67699DFC018FD2" authorityName="Pilato" authorityYear="1997" box="[825,931,1478,1501]" class="Eutardigrada" family="Hypsibiidae" genus="Astatumen" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEAE67699DFC018FD2" box="[825,931,1478,1501]" italics="true" pageId="28" pageNumber="29">Astatumen</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A625FFAEAEB3699DFBF58FD2" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1005,1111,1478,1501]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEAEB3699DFBF58FD2" box="[1005,1111,1478,1501]" italics="true" pageId="28" pageNumber="29">Itaquascon</emphasis>
|
||||
</taxonomicName>
|
||||
specimens on permanent slides (P. Gąsiorek, pers. obs.) and might be related to their shared similarities in the ultrastructure of the apparatus. The claws of
|
||||
<taxonomicName id="4C064D51A625FFAEAE676A7FFC058C34" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[825,935,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A625FFAEAE676A7FFC058C34" box="[825,935,1571,1595]" italics="true" pageId="28" pageNumber="29">P. brunsoni</emphasis>
|
||||
</taxonomicName>
|
||||
are of the
|
||||
<taxonomicName id="4C064D51A625FFAEA9456A78FBD98C34" authorityName="Ehrenberg" authorityYear="1848" box="[1051,1147,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEA9456A78FBD98C34" box="[1051,1147,1571,1595]" italics="true" pageId="28" pageNumber="29">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
type, not the
|
||||
<taxonomicName id="4C064D51A625FFAEA84C6A78FA258C34" authorityName="Thulin" authorityYear="1928" box="[1298,1415,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A625FFAEA84C6A78FA258C34" box="[1298,1415,1571,1595]" italics="true" pageId="28" pageNumber="29">Isohypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
type. Finally, the dorsal sculpturing is not uniform across the dorsum, because the caudalmost polygonal tubercles are disparate in form. This makes
|
||||
<taxonomicName id="4C064D51A625FFAEAEB36AD9FBFA8C96" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[1005,1112,1665,1689]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A625FFAEAEB36AD9FBFA8C96" box="[1005,1112,1665,1689]" italics="true" pageId="28" pageNumber="29">P. brunsoni</emphasis>
|
||||
</taxonomicName>
|
||||
similar to
|
||||
<taxonomicName id="4C064D51A625FFAEA99C6AD9FA888C96" authorityName="Kathman" authorityYear="1990" box="[1218,1322,1665,1689]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="28" pageNumber="29" phylum="Tardigrada" rank="species" species="cheleusis">
|
||||
<emphasis id="B972EAC0A625FFAEA99C6AD9FA888C96" box="[1218,1322,1665,1689]" italics="true" pageId="28" pageNumber="29">P. cheleusis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
303
data/03/AF/87/03AF87C4A626FF92ADA16B38FA8C8DB6.xml
Normal file
303
data/03/AF/87/03AF87C4A626FF92ADA16B38FA8C8DB6.xml
Normal file
|
|
@ -0,0 +1,303 @@
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<mods:nameIdentifier id="63E4B768FCD0B012FA7D30799C8BDEEE" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:location id="D071E32DBB4C01300A6F93378E76EBC7">
|
||||
<mods:url id="E8DE410E6567E05BE68B61F8852BD083">http://dx.doi.org/10.1093/zoolinnean/zlad151</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="C736EF1FABE5A19855927EB882885C7C">journal article</mods:classification>
|
||||
<mods:identifier id="38757352E5A2EBE018015233D379B119" type="DOI">10.1093/zoolinnean/zlad151</mods:identifier>
|
||||
<mods:identifier id="9CE070A4207F936D8580F7212410D6F8" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="23C108299CAB003A193CA991F6E724D9" type="Zenodo-Dep">14243740</mods:identifier>
|
||||
<mods:identifier id="6505DA9EB31733009C4A31AAC5B32BB8" type="ZooBank">044A402-2A0F-4135-9410-7DE081CB11C4Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03AF87C4A626FF92ADA16B38FA8C8DB6" LSID="urn:lsid:plazi:treatment:03AF87C4A626FF92ADA16B38FA8C8DB6" httpUri="http://treatment.plazi.org/id/03AF87C4A626FF92ADA16B38FA8C8DB6" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||||
<subSubSection id="C31C6559A626FFADADA16B38FDCE8D71" box="[255,620,1891,1918]" pageId="31" pageNumber="32" type="nomenclature">
|
||||
<paragraph id="8BB936D2A626FFADADA16B38FDCE8D71" blockId="31.[113,763,1891,1985]" box="[255,620,1891,1918]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADADA16B38FDCE8D71" bold="true" box="[255,620,1891,1918]" pageId="31" pageNumber="32">
|
||||
<taxonomicName id="4C064D51A626FFADADA16B38FDB98D72" authority="Gasiorek" authorityName="Gasiorek" authorityYear="2024" box="[255,539,1891,1918]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="31" pageNumber="32" phylum="Tardigrada" rank="species" species="nivea" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A626FFADADA16B38FE158D71" bold="true" box="[255,439,1891,1918]" italics="true" pageId="31" pageNumber="32">Platicrista nivea</emphasis>
|
||||
Gąsiorek
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA626FFADAF7E6B3FFDCE8D71" box="[544,620,1892,1918]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A626FFADAD2F6BD1FE408DCE" pageId="31" pageNumber="32" type="description">
|
||||
<paragraph id="8BB936D2A626FFADAD2F6BD1FE408DCE" blockId="31.[113,763,1891,1985]" pageId="31" pageNumber="32">
|
||||
ZooBank registration:
|
||||
<uri id="FF973AD0A626FFADACDB6BD1FE408DCE" pageId="31" pageNumber="32">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFA00C07A626FFADAFDD6BD1FE408DCE" pageId="31" pageNumber="32">3988853F- DDB5-4217-8293-701E05D226BD</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A626FF92AE746A95FE648FA4" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
|
||||
<paragraph id="8BB936D2A626FFADAE746A95FB318D09" blockId="31.[810,1459,1742,1798]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE746A95FC4D8CE9" box="[810,1007,1742,1766]" italics="true" pageId="31" pageNumber="32">Material examined:</emphasis>
|
||||
Four individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A626FFADA9756AB6FB208D09" box="[1067,1154,1773,1798]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="31" pageNumber="32">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A626FF92AE746B75FE648FA4" blockId="31.[810,1460,1837,1986]" lastBlockId="32.[129,778,1332,1451]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE746B75FC1B8D4A" box="[810,953,1837,1861]" italics="true" pageId="31" pageNumber="32">Type material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA626FF92AE9E6B76FE648FA4" collectingDate="2020-02-10" collectionCode="TW" collectorName="Dreyer" country="Taiwan" county="Institute of Zoology" elevation="3200" lastPageId="32" lastPageNumber="33" latitude="24.3975" location="Snow Mountain" longLatPrecision="21" longitude="121.23444" municipality="North Peak" pageId="31" pageNumber="32" specimenCode="TW.008.14, TW.008.15" specimenCount="2" typeStatus="holotype">
|
||||
<typeStatus id="54BD8870A626FFADAE9E6B76FB818D4A" box="[960,1059,1837,1861]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
|
||||
(slide
|
||||
<specimenCount id="9D00FD5BA626FFADA9366B76FB428D4A" box="[1128,1248,1837,1861]" pageId="31" pageNumber="32">TW.006.07</specimenCount>
|
||||
) from
|
||||
<collectingCountry id="F3117642A626FFADA87B6B76FAD38D4A" box="[1317,1393,1837,1861]" name="Taiwan" pageId="31" pageNumber="32">Taiwan</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A626FFADA8246B76FC338D6B" LSID="urn:lsid:plazi:treatment:03AF87C4A626FF92ADA16B38FA8C8DB6:8ED96009A626FFADA8246B76FC338D6B" country="Taiwan" county="Institute of Zoology" latitude="24.3975" longLatPrecision="21" longitude="121.23444" municipality="North Peak" name="Snow Mountain" pageId="31" pageNumber="32">Snow Mountain</location>
|
||||
(Xueshan),
|
||||
<location id="8ED96009A626FFADA9416B16FB2A8D6B" LSID="urn:lsid:plazi:treatment:03AF87C4A626FF92ADA16B38FA8C8DB6:8ED96009A626FFADA9416B16FB2A8D6B" box="[1055,1160,1868,1892]" country="Taiwan" county="Institute of Zoology" latitude="24.3975" longLatPrecision="21" longitude="121.23444" municipality="North Peak" name="East Peak" pageId="31" pageNumber="32">East Peak</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A626FFADA9C36B17FABB8D6B" box="[1181,1305,1867,1892]" degrees="24" direction="north" minutes="23" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="18" value="24.388332">
|
||||
24°23
|
||||
<emphasis id="B972EAC0A626FFADA9826B10FB408D6B" box="[1244,1250,1867,1892]" italics="true" pageId="31" pageNumber="32">ʹ</emphasis>
|
||||
18″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A626FFADA8746B17FA0D8D6B" box="[1322,1455,1867,1892]" degrees="121" direction="east" minutes="15" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="39" value="121.26083">
|
||||
121°15
|
||||
<emphasis id="B972EAC0A626FFADA82A6B10FAD88D6B" box="[1396,1402,1867,1892]" italics="true" pageId="31" pageNumber="32">ʹ</emphasis>
|
||||
39″E
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A626FFADAE746B37FC0C8D8C" box="[810,942,1899,1924]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="31" pageNumber="32" unit="m" value="3200.0">
|
||||
<quantity id="4CFE9B37A626FFADAE746B37FCD88D8C" box="[810,890,1900,1924]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="31" pageNumber="32" unit="m" value="3200.0">3200 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), moss from rock in alpine grassland,
|
||||
<date id="FFB81012A626FFADA86B6B37FCC38DAC" pageId="31" pageNumber="32" value="2020-02-10">
|
||||
<collectingDate id="EFFCE9FAA626FFADA86B6B37FCC38DAC" pageId="31" pageNumber="32" value="2020-02-10">10 February 2020</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A626FFADAE306BD0FC178DAC" box="[878,949,1931,1955]" pageId="31" pageNumber="32">Dreyer</collectorName>
|
||||
coll., and
|
||||
<specimenCount id="9D00FD5BA626FFADA9766BD0FBED8DAD" box="[1064,1103,1931,1954]" pageId="31" pageNumber="32">two</specimenCount>
|
||||
<typeStatus id="54BD8870A626FFADA9066BD0FB198DAC" box="[1112,1211,1931,1955]" pageId="31" pageNumber="32">paratypes</typeStatus>
|
||||
(slides
|
||||
<specimenCode id="DBA09EA9A626FFADA84E6BD0FA268DAC" box="[1296,1412,1931,1955]" pageId="31" pageNumber="32">TW.008.14</specimenCode>
|
||||
and
|
||||
<specimenCode id="DBA09EA9A626FFADAE746BF1FC018DCD" box="[810,931,1962,1986]" pageId="31" pageNumber="32">TW.008.15</specimenCode>
|
||||
) from
|
||||
<collectingCountry id="F3117642A626FFADAEB06BF1FB988DCD" box="[1006,1082,1962,1986]" name="Taiwan" pageId="31" pageNumber="32">Taiwan</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A626FFADA9186BF1FB4C8DCD" LSID="urn:lsid:plazi:treatment:03AF87C4A626FF92ADA16B38FA8C8DB6:8ED96009A626FFADA9186BF1FB4C8DCD" box="[1094,1262,1962,1986]" country="Taiwan" county="Institute of Zoology" latitude="24.3975" longLatPrecision="21" longitude="121.23444" municipality="North Peak" name="Snow Mountain" pageId="31" pageNumber="32">Snow Mountain</location>
|
||||
(Xueshan),
|
||||
<collectingMunicipality id="6BDDACA8A626FF92A82D6BF1FF138F42" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">North Peak</collectingMunicipality>
|
||||
(
|
||||
<geoCoordinate id="EE325015A619FF92AD98696EFEE08F42" box="[198,322,1332,1357]" degrees="24" direction="north" minutes="23" orientation="latitude" pageId="32" pageNumber="33" precision="15" seconds="51" value="24.3975">
|
||||
24°23
|
||||
<emphasis id="B972EAC0A619FF92AC5B696FFEA98F42" box="[261,267,1332,1357]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
|
||||
51″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A619FF92AC0D696EFE758F42" box="[339,471,1332,1357]" degrees="121" direction="east" minutes="14" orientation="longitude" pageId="32" pageNumber="33" precision="15" seconds="04" value="121.23444">
|
||||
121°14
|
||||
<emphasis id="B972EAC0A619FF92ACC2696FFE008F42" box="[412,418,1332,1357]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
|
||||
04″E
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A619FF92ACB7696EFDD88F42" box="[489,634,1333,1357]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="32" pageNumber="33" unit="m" value="3700.0">
|
||||
<quantity id="4CFE9B37A619FF92ACB7696EFD9D8F42" box="[489,575,1333,1357]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="32" pageNumber="33" unit="m" value="3700.0">3700 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), moss from rock in alpine grassland,
|
||||
<date id="FFB81012A619FF92AC23690EFD948F62" box="[381,566,1364,1389]" pageId="32" pageNumber="33" value="2020-02-10">
|
||||
<collectingDate id="EFFCE9FAA619FF92AC23690EFD948F62" box="[381,566,1364,1389]" pageId="32" pageNumber="33" value="2020-02-10">10 February 2020</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A619FF92AF1E690EFD258F63" box="[576,647,1365,1388]" pageId="32" pageNumber="33">Dreyer</collectorName>
|
||||
coll., are deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University, Kraków.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF79665AA619FF92ADDF688AFB788F0A" ID-DOI="http://doi.org/10.5281/zenodo.14243794" ID-Zenodo-Dep="14243794" httpUri="https://zenodo.org/record/14243794/files/figure.png" pageId="32" pageNumber="33" startId="32.[129,194,1233,1257]" targetBox="[131,1470,146,1201]" targetPageId="32" targetType="figure">
|
||||
<paragraph id="8BB936D2A619FF92ADDF688AFB788F0A" blockId="32.[129,1457,1233,1285]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B972EAC0A619FF92ADDF688AFF478EE6" bold="true" box="[129,229,1233,1257]" pageId="32" pageNumber="33">Figure 21.</emphasis>
|
||||
Detailed morphology of
|
||||
<taxonomicName id="4C064D51A619FF92AC8C688AFD2D8EE6" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[466,655,1233,1257]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="borneensis" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A619FF92AC8C688AFD2D8EE6" box="[466,655,1233,1257]" italics="true" pageId="32" pageNumber="33">Platicrista borneensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A619FF92AFCA688AFD7D8EE6" bold="true" box="[660,735,1233,1257]" pageId="32" pageNumber="33">
|
||||
<taxonomicNameLabel id="A24157BBA619FF92AFCA688AFD7D8EE6" box="[660,735,1233,1257]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(PCM): A, holotype in ventral view (white arrowheads indicate median and posterior bars); B, buccopharyngeal apparatus
|
||||
<emphasis id="B972EAC0A619FF92AF6968B6FDD38F0A" box="[567,625,1261,1285]" italics="true" pageId="32" pageNumber="33">in toto</emphasis>
|
||||
(paratype); C, claws II (paratype). Scale bars are in micrometres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C31C6559A619FF92ADDF6996FB558F63" pageId="32" pageNumber="33" type="description">
|
||||
<paragraph id="8BB936D2A619FF92ADDF6996FDD88D52" blockId="32.[128,780,1484,1979]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B972EAC0A619FF92ADDF6996FF568FEB" box="[129,244,1485,1508]" italics="true" pageId="32" pageNumber="33">Description:</emphasis>
|
||||
Body massive (
|
||||
<tableCitation id="C6840369A619FF92ACC86997FE4F8FEA" box="[406,493,1484,1509]" captionStart="Table 10" captionStartId="39.[113,168,143,167]" captionTargetPageId="39" captionText="Table 10. Measurements (in micrometres) of selected morphological structures of type series of Platicrista nivea sp. nov. from Taiwan (TW.006 + TW.008) mounted in Hoyer’s medium." pageId="32" pageNumber="33">Table 10</tableCitation>
|
||||
), white and elongated, with blunt cephalic part (
|
||||
<figureCitation id="133D2A57A619FF92AC0769B7FE008C0B" box="[345,418,1516,1540]" captionStart="Figure 28" captionStartId="37.[113,178,1674,1698]" captionTargetBox="[117,1455,1148,1642]" captionTargetId="figure-174@37.[114,1459,1146,1646]" captionTargetPageId="37" captionText="Figure 28. General morphology of Platicrista nivea sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243810" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="32" pageNumber="33">Fig. 28</figureCitation>
|
||||
). Cuticle without pores, but with both irregular wrinkling and distinct, minute granulation in the caudal body part (
|
||||
<figureCitation id="133D2A57A619FF92AC1C6A71FE2B8C4C" box="[322,393,1578,1603]" captionStart="Figure 29" captionStartId="40.[129,194,1261,1285]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-146@40.[129,1473,144,1233]" captionTargetPageId="40" captionText="Figure 29. Caudal sculpturing of Platicrista nivea sp. nov. (paratypes, PCM). Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243812" httpUri="https://zenodo.org/record/14243812/files/figure.png" pageId="32" pageNumber="33">Fig. 29</figureCitation>
|
||||
). Cribriform areas identifiable under PCM (
|
||||
<figureCitation id="133D2A57A619FF92AD956A11FE808C6D" box="[203,290,1610,1634]" captionStart="Figure 29" captionStartId="40.[129,194,1261,1285]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-146@40.[129,1473,144,1233]" captionTargetPageId="40" captionText="Figure 29. Caudal sculpturing of Platicrista nivea sp. nov. (paratypes, PCM). Scale bars: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.14243812" httpUri="https://zenodo.org/record/14243812/files/figure.png" pageId="32" pageNumber="33">Fig. 29A</figureCitation>
|
||||
). Legs short and plump; legs IV poorly delimited from the trunk (
|
||||
<figureCitation id="133D2A57A619FF92AC116A32FE318C8E" box="[335,403,1641,1665]" captionStart="Figure 28" captionStartId="37.[113,178,1674,1698]" captionTargetBox="[117,1455,1148,1642]" captionTargetId="figure-174@37.[114,1459,1146,1646]" captionTargetPageId="37" captionText="Figure 28. General morphology of Platicrista nivea sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243810" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="32" pageNumber="33">Fig. 28</figureCitation>
|
||||
). Eyes absent in unmounted animals. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A619FF92ACB16AD3FDED8CAF" authorityName="Pilato" authorityYear="1987" box="[495,591,1672,1696]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A619FF92ACB16AD3FDED8CAF" box="[495,591,1672,1696]" italics="true" pageId="32" pageNumber="33">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A619FF92AF056AD2FD2A8CAE" box="[603,648,1673,1697]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A619FF92AFC36AD3FD558CAE" box="[669,759,1672,1697]" captionStart="Figure 30" captionStartId="40.[129,194,1670,1694]" captionTargetBox="[131,1470,1334,1637]" captionTargetId="figure-162@40.[129,1474,1332,1641]" captionTargetPageId="40" captionText="Figure 30. Detailed morphology of Platicrista nivea sp. nov. (PCM): A, buccopharyngeal apparatus in toto (paratype); B, claws II (holotype). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243814" httpUri="https://zenodo.org/record/14243814/files/figure.png" pageId="32" pageNumber="33">Fig. 30A</figureCitation>
|
||||
): OCA not visible under PCM; annulation of the pharyngeal tube simple (single bifurcations visible) and comprising tightly merged annuli; stylet supports embedded at the border between buccal and pharyngeal tube; furcae of the
|
||||
<taxonomicName id="4C064D51A619FF92ACAD6B5DFDF18D11" authorityName="Pilato" authorityYear="1987" box="[499,595,1798,1822]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A619FF92ACAD6B5DFDF18D11" box="[499,595,1798,1822]" italics="true" pageId="32" pageNumber="33">Platicrista</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A619FF92AF036B5DFD2F8D11" box="[605,653,1798,1822]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
; pharyngeal apophyses absent; pharynx oval, with two bar-like macroplacoids: the first much shorter than the second one (1 <2).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A619FF92ADC26B3FFB558F63" blockId="32.[128,780,1484,1979]" lastBlockId="32.[825,1474,1333,1388]" pageId="32" pageNumber="33">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A619FF92AC7F6B3FFE238D73" authorityName="Ehrenberg" authorityYear="1848" box="[289,385,1892,1916]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A619FF92AC7F6B3FFE238D73" box="[289,385,1892,1916]" italics="true" pageId="32" pageNumber="33">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A619FF92ACD86B3FFE178D73" box="[390,437,1892,1916]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
, large and slender (
|
||||
<figureCitation id="133D2A57A619FF92AF226B3FFD678D73" box="[636,709,1892,1916]" captionStart="Figure 28" captionStartId="37.[113,178,1674,1698]" captionTargetBox="[117,1455,1148,1642]" captionTargetId="figure-174@37.[114,1459,1146,1646]" captionTargetPageId="37" captionText="Figure 28. General morphology of Platicrista nivea sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243810" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="32" pageNumber="33">Figs 28</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A619FF92AF8E6B3FFD5A8D73" box="[720,760,1892,1916]" captionStart="Figure 30" captionStartId="40.[129,194,1670,1694]" captionTargetBox="[131,1470,1334,1637]" captionTargetId="figure-162@40.[129,1474,1332,1641]" captionTargetPageId="40" captionText="Figure 30. Detailed morphology of Platicrista nivea sp. nov. (PCM): A, buccopharyngeal apparatus in toto (paratype); B, claws II (holotype). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243814" httpUri="https://zenodo.org/record/14243814/files/figure.png" pageId="32" pageNumber="33">30B</figureCitation>
|
||||
). Accessory points divergent from all claw branches. Indentation of claw bases absent. Septa between claw parts absent. Pseudolunulae present, but clearly discernible only under anterior claws (
|
||||
<figureCitation id="133D2A57A619FF92AEE0690FFBA18F63" box="[958,1027,1364,1388]" captionStart="Figure 28" captionStartId="37.[113,178,1674,1698]" captionTargetBox="[117,1455,1148,1642]" captionTargetId="figure-174@37.[114,1459,1146,1646]" captionTargetPageId="37" captionText="Figure 28. General morphology of Platicrista nivea sp. nov. (holotype in ventral view, PCM). Scale bar is in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243810" httpUri="https://zenodo.org/record/14243810/files/figure.png" pageId="32" pageNumber="33">Fig. 28</figureCitation>
|
||||
). Cuticular bars absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A619FF92AE6769D2FC2D8FD0" pageId="32" pageNumber="33" type="etymology">
|
||||
<paragraph id="8BB936D2A619FF92AE6769D2FC2D8FD0" blockId="32.[825,1475,1417,1503]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B972EAC0A619FF92AE6769D2FC0A8FAE" box="[825,936,1417,1441]" italics="true" pageId="32" pageNumber="33">Etymology:</emphasis>
|
||||
From the Latin
|
||||
<emphasis id="B972EAC0A619FF92A90F69D2FB2F8FAF" box="[1105,1165,1417,1440]" italics="true" pageId="32" pageNumber="33">niveus</emphasis>
|
||||
= snowy, referring to the
|
||||
<typeStatus id="54BD8870A619FF92A8CA69D2FA608FAE" box="[1428,1474,1417,1441]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
locality in Snow Mountain (
|
||||
<collectingRegion id="49C2F830A619FF92A90C69F3FB098FCF" box="[1106,1195,1448,1472]" country="China" name="Taiwan)" pageId="32" pageNumber="33">Taiwan)</collectingRegion>
|
||||
. An adjective in nominative singular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A619FF92AE6769A7FA8C8DB6" pageId="32" pageNumber="33" type="diagnosis">
|
||||
<paragraph id="8BB936D2A619FF92AE6769A7FC548CBE" blockId="32.[824,1474,1532,1713]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B972EAC0A619FF92AE6769A7FBB18C1B" box="[825,1043,1532,1556]" italics="true" pageId="32" pageNumber="33">Differential diagnosis:</emphasis>
|
||||
There are two congeneric species with strong dorsal sculpturing:
|
||||
<taxonomicName id="4C064D51A619FF92A9146A47FB158C3C" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[1098,1207,1563,1587]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A619FF92A9146A47FB158C3C" box="[1098,1207,1563,1587]" italics="true" pageId="32" pageNumber="33">P. brunsoni</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A619FF92A9B56A47FAF18C3C" authorityName="Kathman" authorityYear="1990" box="[1259,1363,1563,1587]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="cheleusis">
|
||||
<emphasis id="B972EAC0A619FF92A9B56A47FAF18C3C" box="[1259,1363,1563,1587]" italics="true" pageId="32" pageNumber="33">P. cheleusis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="4C064D51A619FF92A8016A40FCCE8C5D" authorityName="Gasiorek" authorityYear="2024" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A619FF92A8016A40FCCE8C5D" italics="true" pageId="32" pageNumber="33">Platicrista nivea</emphasis>
|
||||
</taxonomicName>
|
||||
can be differentiated from them based on differences in the
|
||||
<typeStatus id="54BD8870A619FF92AE676A01FCC58C7D" box="[825,871,1626,1650]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
of sculpturing (minute granules restricted to the caudal part in
|
||||
<taxonomicName id="4C064D51A619FF92AE0A6A21FC3F8C9E" authorityName="Gasiorek" authorityYear="2024" box="[852,925,1658,1681]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A619FF92AE0A6A21FC3F8C9E" box="[852,925,1658,1681]" italics="true" pageId="32" pageNumber="33">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
vs. polygonal tubercles covering the entire dorsum in both species) and:
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A619FF92AE0A6A95FB508D0A" blockId="32.[852,1474,1741,1891]" pageId="32" pageNumber="33">
|
||||
• for
|
||||
<taxonomicName id="4C064D51A619FF92AEF86A95FBB28CEA" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[934,1040,1741,1765]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A619FF92AEF86A95FBB28CEA" box="[934,1040,1741,1765]" italics="true" pageId="32" pageNumber="33">P. brunsoni</emphasis>
|
||||
</taxonomicName>
|
||||
, the presence of internal bars II–III (absent in
|
||||
<taxonomicName id="4C064D51A619FF92AEC36AB6FC448D0B" authorityName="Gasiorek" authorityYear="2024" box="[925,998,1773,1796]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A619FF92AEC36AB6FC448D0B" box="[925,998,1773,1796]" italics="true" pageId="32" pageNumber="33">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
vs. present in
|
||||
<taxonomicName id="4C064D51A619FF92A92B6AB6FB428D0A" authorityName="Miller & J. D. Miller" authorityYear="2021" box="[1141,1248,1773,1797]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="brunsoni">
|
||||
<emphasis id="B972EAC0A619FF92A92B6AB6FB428D0A" box="[1141,1248,1773,1797]" italics="true" pageId="32" pageNumber="33">P. brunsoni</emphasis>
|
||||
</taxonomicName>
|
||||
);
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A619FF92AE0A6B57FC5B8D6C" blockId="32.[852,1474,1741,1891]" pageId="32" pageNumber="33">
|
||||
• for
|
||||
<taxonomicName id="4C064D51A619FF92AEF76B56FBB38D2B" authorityName="Kathman" authorityYear="1990" box="[937,1041,1804,1828]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="cheleusis">
|
||||
<emphasis id="B972EAC0A619FF92AEF76B56FBB38D2B" box="[937,1041,1804,1828]" italics="true" pageId="32" pageNumber="33">P. cheleusis</emphasis>
|
||||
</taxonomicName>
|
||||
, the presence of internal and median bars I–III and posterior bars (absent in
|
||||
<taxonomicName id="4C064D51A619FF92A9B46B77FA968D4C" authorityName="Gasiorek" authorityYear="2024" box="[1258,1332,1836,1859]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="nivea">
|
||||
<emphasis id="B972EAC0A619FF92A9B46B77FA968D4C" box="[1258,1332,1836,1859]" italics="true" pageId="32" pageNumber="33">P. nivea</emphasis>
|
||||
</taxonomicName>
|
||||
vs. present in
|
||||
<taxonomicName id="4C064D51A619FF92AEDA6B10FC4B8D6C" authorityName="Kathman" authorityYear="1990" box="[900,1001,1867,1891]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="cheleusis">
|
||||
<emphasis id="B972EAC0A619FF92AEDA6B10FC4B8D6C" box="[900,1001,1867,1891]" italics="true" pageId="32" pageNumber="33">P. cheleusis</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A619FF92AE0B6BD9FA8C8DB6" blockId="32.[825,1474,1922,1977]" pageId="32" pageNumber="33">
|
||||
<taxonomicName id="4C064D51A619FF92AE0B6BD9FBBD8D95" box="[853,1055,1922,1946]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="angustata">
|
||||
<emphasis id="B972EAC0A619FF92AE0B6BD9FBBD8D95" box="[853,1055,1922,1946]" italics="true" pageId="32" pageNumber="33">Platicrista angustata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A619FF92A9086BD8FB748D95" authorityName="Gasiorek" authorityYear="2024" box="[1110,1238,1922,1946]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="32" pageNumber="33" phylum="Tardigrada" rank="species" species="carpathica">
|
||||
<emphasis id="B972EAC0A619FF92A9086BD8FB748D95" box="[1110,1238,1922,1946]" italics="true" pageId="32" pageNumber="33">P. carpathica</emphasis>
|
||||
</taxonomicName>
|
||||
have a similar-looking dorsocaudal cuticle; see above for differentiation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
878
data/03/AF/87/03AF87C4A627FFADAE316A77FD9C8D4B.xml
Normal file
878
data/03/AF/87/03AF87C4A627FFADAE316A77FD9C8D4B.xml
Normal file
|
|
@ -0,0 +1,878 @@
|
|||
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<mods:nameIdentifier id="571CFC8AA9B90BD7C0C909E14D908249" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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|
||||
<treatment id="03AF87C4A627FFADAE316A77FD9C8D4B" LSID="urn:lsid:plazi:treatment:03AF87C4A627FFADAE316A77FD9C8D4B" httpUri="http://treatment.plazi.org/id/03AF87C4A627FFADAE316A77FD9C8D4B" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
|
||||
<subSubSection id="C31C6559A627FFACAE316A77FA2E8C49" box="[879,1420,1580,1607]" pageId="30" pageNumber="31" type="nomenclature">
|
||||
<paragraph id="8BB936D2A627FFACAE316A77FA2E8C49" blockId="30.[879,1420,1580,1607]" box="[879,1420,1580,1607]" pageId="30" pageNumber="31">
|
||||
<heading id="D0F181BEA627FFACAE316A77FA2E8C49" bold="true" box="[879,1420,1580,1607]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
|
||||
<taxonomicName id="4C064D51A627FFACAE316A77FA2E8C49" ID-CoL="4JNBY" authority="Kaczmarek & Michalczyk, 2003" authorityName="Kaczmarek & Michalczyk" authorityYear="2003" box="[879,1420,1580,1607]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="species" species="horribilis">
|
||||
<emphasis id="B972EAC0A627FFACAE316A77FA2E8C49" bold="true" box="[879,1420,1580,1607]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACAE316A77FB948C48" bold="true" box="[879,1078,1580,1607]" italics="true" pageId="30" pageNumber="31">Platicrista horribilis</emphasis>
|
||||
<bibRefCitation id="EF974B23A627FFACA9656A76FA2E8C49" author="Kaczmarek L & Michalczyk L" box="[1083,1420,1580,1607]" pageId="30" pageNumber="31" pagination="1 - 8" refId="ref36912" refString="Kaczmarek L, Michalczyk L. Platicrista horribilis, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Mongolia. Zootaxa 2003; 132: 1 - 8. https: // doi. org / 10.11646 / zootaxa. 132.1.1" type="journal article" year="2003">Kaczmarek & Michalczyk, 2003</bibRefCitation>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A627FFACAE676A36FB008CAB" pageId="30" pageNumber="31" type="materials_examined">
|
||||
<paragraph id="8BB936D2A627FFACAE676A36FB008CAB" blockId="30.[825,1474,1645,1700]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACAE676A36FBA08C8A" box="[825,1026,1645,1669]" italics="true" pageId="30" pageNumber="31">Material examined:</emphasis>
|
||||
25 individuals in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A627FFACA9646AD7FB338CAB" box="[1082,1169,1676,1700]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="30" pageNumber="31">Table S1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A627FFACAE676A9DFBD58DB6" pageId="30" pageNumber="31" type="description">
|
||||
<paragraph id="8BB936D2A627FFACAE676A9DFBD58DB6" blockId="30.[825,1476,1734,1977]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B972EAC0A627FFACAE676A9DFBAF8CD1" box="[825,1037,1734,1758]" italics="true" pageId="30" pageNumber="31">Amended description:</emphasis>
|
||||
The OCA not visible under PCM; simple pharyngeal annulation composed of tightly arranged single annuli; pharyngeal apophyses absent (
|
||||
<figureCitation id="133D2A57A627FFACA9996B5FFA878D13" box="[1223,1317,1796,1820]" captionStart="Figure 27" captionStartId="37.[113,178,1027,1051]" captionTargetBox="[116,1455,146,996]" captionTargetId="figure-131@37.[114,1459,144,1000]" captionTargetPageId="37" captionText="Figure 27. Detailed morphology of Platicrista horribilis Kaczmarek & Michalczyk, 2003 (PCM): A, buccopharyngeal apparatus in toto; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="30" pageNumber="31">Fig. 27A</figureCitation>
|
||||
). Large claws of the
|
||||
<taxonomicName id="4C064D51A627FFACAEDB6B7FFC478D33" authorityName="Ehrenberg" authorityYear="1848" box="[901,997,1828,1852]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="30" pageNumber="31" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A627FFACAEDB6B7FFC478D33" box="[901,997,1828,1852]" italics="true" pageId="30" pageNumber="31">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A627FFACAEAE6B7FFBBC8D33" box="[1008,1054,1828,1852]" pageId="30" pageNumber="31">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A627FFACA96D6B7FFB0C8D33" box="[1075,1198,1828,1852]" captionStart="Figure 27" captionStartId="37.[113,178,1027,1051]" captionTargetBox="[116,1455,146,996]" captionTargetId="figure-131@37.[114,1459,144,1000]" captionTargetPageId="37" captionText="Figure 27. Detailed morphology of Platicrista horribilis Kaczmarek & Michalczyk, 2003 (PCM): A, buccopharyngeal apparatus in toto; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="30" pageNumber="31">Fig. 27B–D</figureCitation>
|
||||
). Accessory points divergent from both primary and secondary branches of all claws. Pseudolunulae present, indented only on legs IV. Internal, anterior, and posterior claw bases with delicate indentations (
|
||||
<figureCitation id="133D2A57A627FFACA8C26BD9FC268DB6" captionStart="Figure 27" captionStartId="37.[113,178,1027,1051]" captionTargetBox="[116,1455,146,996]" captionTargetId="figure-131@37.[114,1459,144,1000]" captionTargetPageId="37" captionText="Figure 27. Detailed morphology of Platicrista horribilis Kaczmarek & Michalczyk, 2003 (PCM): A, buccopharyngeal apparatus in toto; B, claws I (white asterisk indicates internal pulvinus); C, claws III; D, claws IV (empty white arrowhead indicates pseudolunule). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243808" httpUri="https://zenodo.org/record/14243808/files/figure.png" pageId="30" pageNumber="31">Fig. 27B–D</figureCitation>
|
||||
). Cuticular bars absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF79665AA626FFADAD2F6CD4FC878ACC" pageId="31" pageNumber="32" startId="31.[113,168,143,167]" targetBox="[113,1451,217,1687]" targetIsTable="true" targetPageId="31" targetType="table">
|
||||
<paragraph id="8BB936D2A626FFADAD2F6CD4FC878ACC" blockId="31.[113,1409,143,195]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAD2F6CD4FF1D8AA8" bold="true" box="[113,191,143,167]" pageId="31" pageNumber="32">Table 7.</emphasis>
|
||||
Measurements (in micrometres) of selected morphological structures of type series (MY.090 + MY.091 + MY.100 + MY.826) of
|
||||
<taxonomicName id="4C064D51A626FFADAD2F6CF0FE8C8ACC" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[113,302,171,195]" class="Eutardigrada" family="Hypsibiidae" genus="Platicrista" kingdom="Animalia" order="Parachela" pageId="31" pageNumber="32" phylum="Tardigrada" rank="species" species="borneensis" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A626FFADAD2F6CF0FE8C8ACC" box="[113,302,171,195]" italics="true" pageId="31" pageNumber="32">Platicrista borneensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A626FFADAC6D6CF0FEDC8ACC" bold="true" box="[307,382,171,195]" pageId="31" pageNumber="32">
|
||||
<taxonomicNameLabel id="A24157BBA626FFADAC6D6CF0FEDC8ACC" box="[307,382,171,195]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
from Malaysia mounted in Hoyer’s medium.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A626FFADAD2F6C82FADD8C99" pageId="31" pageNumber="32">
|
||||
<table id="F906C472A626004DAD2F6C82FA098C98" box="[113,1451,217,1687]" gridcols="10" gridrows="43" pageId="31" pageNumber="32">
|
||||
<tr id="35363490A626004DAD2F6C82FA098AFE" box="[113,1451,217,241]" gridrow="0" pageId="31" pageNumber="32" rowspan-3="1" rowspan-5="1" rowspan-7="1">
|
||||
<th id="76E75DECA626004DAD2F6C82FE428AFE" box="[113,480,217,241]" gridcol="0" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAD2F6C82FF768AFE" bold="true" box="[113,212,217,241]" pageId="31" pageNumber="32">Character</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA626004DAF596C82FDBB8AFE" box="[519,537,217,241]" gridcol="1" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAF596C82FDBB8AFF" bold="true" box="[519,537,217,240]" italics="true" pageId="31" pageNumber="32">N</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA626004DAF146C82FD618AFE" box="[586,707,217,241]" gridcol="2" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAF146C82FD2B8AFE" bold="true" box="[586,649,217,241]" pageId="31" pageNumber="32">Range</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA626004DAEC96C82FC6D8AFE" box="[919,975,217,241]" gridcol="4" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEC96C82FC6D8AFE" bold="true" box="[919,975,217,241]" pageId="31" pageNumber="32">Mean</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA626004DA93E6C82FB2B8AFE" box="[1120,1161,217,241]" gridcol="6" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA93E6C82FBDD8AFE" bold="true" box="[1120,1151,217,241]" pageId="31" pageNumber="32">SD</emphasis>
|
||||
</th>
|
||||
<th id="76E75DECA626004DA84F6C82FA098AFE" box="[1297,1451,217,241]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA84F6C82FAD28AFE" bold="true" box="[1297,1392,217,241]" pageId="31" pageNumber="32">Holotype</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6D5EFA098B12" box="[113,1451,261,285]" gridrow="1" pageId="31" pageNumber="32" rowspan-0="1" rowspan-1="1">
|
||||
<td id="76E75DECA626004DAF146D5EFD618B12" box="[586,707,261,285]" gridcol="2" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAF146D5EFDC88B12" bold="true" box="[586,618,261,285]" pageId="31" pageNumber="32">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAFAF6D5EFCCB8B12" box="[753,873,261,285]" gridcol="3" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFAF6D5EFCA78B13" bold="true" box="[753,773,261,284]" italics="true" pageId="31" pageNumber="32">pt</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96D5EFC6D8B12" box="[919,975,261,285]" gridcol="4" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEC96D5EFC158B12" bold="true" box="[919,951,261,285]" pageId="31" pageNumber="32">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEA26D5EFB938B12" box="[1020,1073,261,285]" gridcol="5" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26D5EFBB28B13" bold="true" box="[1020,1040,261,284]" italics="true" pageId="31" pageNumber="32">pt</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6D5EFB2B8B12" box="[1120,1161,261,285]" gridcol="6" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA93E6D5EFB228B12" bold="true" box="[1120,1152,261,285]" pageId="31" pageNumber="32">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA9E76D5EFB408B12" box="[1209,1250,261,285]" gridcol="7" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76D5EFB6F8B13" bold="true" box="[1209,1229,261,284]" italics="true" pageId="31" pageNumber="32">pt</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6D5EFAE58B12" box="[1297,1351,261,285]" gridcol="8" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA84F6D5EFA938B12" bold="true" box="[1297,1329,261,285]" pageId="31" pageNumber="32">µm</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA8286D5EFA098B12" box="[1398,1451,261,285]" gridcol="9" gridrow="1" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286D5EFA288B13" bold="true" box="[1398,1418,261,284]" italics="true" pageId="31" pageNumber="32">pt</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6D6EFA098B42" box="[113,1451,309,333]" gridrow="2" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6D6EFE428B42" box="[113,480,309,333]" gridcol="0" gridrow="2" pageId="31" pageNumber="32">Body length</th>
|
||||
<td id="76E75DECA626004DAF596D6EFDBB8B42" box="[519,537,309,333]" gridcol="1" gridrow="2" pageId="31" pageNumber="32">8</td>
|
||||
<td id="76E75DECA626004DAF146D6EFD618B42" box="[586,707,309,333]" gridcol="2" gridrow="2" pageId="31" pageNumber="32">271–346</td>
|
||||
<td id="76E75DECA626004DAFAF6D6EFCCB8B42" box="[753,873,309,333]" gridcol="3" gridrow="2" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE5D6D6EFCC18B42" box="[771,867,309,333]" italics="true" pageId="31" pageNumber="32">925–1139</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96D6EFC6D8B42" box="[919,975,309,333]" gridcol="4" gridrow="2" pageId="31" pageNumber="32">318</td>
|
||||
<td id="76E75DECA626004DAEA26D6EFB938B42" box="[1020,1073,309,333]" gridcol="5" gridrow="2" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26D6EFB8E8B42" box="[1020,1068,309,333]" italics="true" pageId="31" pageNumber="32">1028</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6D6EFB2B8B42" box="[1120,1161,309,333]" gridcol="6" gridrow="2" pageId="31" pageNumber="32">27</td>
|
||||
<td id="76E75DECA626004DA9E76D6EFB408B42" box="[1209,1250,309,333]" gridcol="7" gridrow="2" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76D6EFB738B42" box="[1209,1233,309,333]" italics="true" pageId="31" pageNumber="32">80</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6D6EFAE58B42" box="[1297,1351,309,333]" gridcol="8" gridrow="2" pageId="31" pageNumber="32">336</td>
|
||||
<td id="76E75DECA626004DA8286D6EFA098B42" box="[1398,1451,309,333]" gridcol="9" gridrow="2" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286D6EFA048B42" box="[1398,1446,309,333]" italics="true" pageId="31" pageNumber="32">1139</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6D0CFA098B60" box="[113,1451,343,367]" gridrow="3" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F6D0CFE428B60" box="[113,480,343,367]" gridcol="0" gridrow="3" pageId="31" pageNumber="32">Buccopharyngeal tube</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6D23FA098B9E" box="[113,1451,376,401]" gridrow="4" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6D23FE428B9E" box="[113,480,376,401]" gridcol="0" gridrow="4" pageId="31" pageNumber="32">Buccal tube length</th>
|
||||
<td id="76E75DECA626004DAF596D23FDBB8B9E" box="[519,537,376,401]" gridcol="1" gridrow="4" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146D23FD618B9E" box="[586,707,376,401]" gridcol="2" gridrow="4" pageId="31" pageNumber="32">29.3–33.5</td>
|
||||
<td id="76E75DECA626004DAFAF6D23FCCB8B9E" box="[753,873,376,401]" gridcol="3" gridrow="4" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DAEC96D23FC6D8B9E" box="[919,975,376,401]" gridcol="4" gridrow="4" pageId="31" pageNumber="32">30.8</td>
|
||||
<td id="76E75DECA626004DAEA26D23FB938B9E" box="[1020,1073,376,401]" gridcol="5" gridrow="4" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DA93E6D23FB2B8B9E" box="[1120,1161,376,401]" gridcol="6" gridrow="4" pageId="31" pageNumber="32">1.5</td>
|
||||
<td id="76E75DECA626004DA9E76D23FB408B9E" box="[1209,1250,376,401]" gridcol="7" gridrow="4" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DA84F6D23FAE58B9E" box="[1297,1351,376,401]" gridcol="8" gridrow="4" pageId="31" pageNumber="32">29.5</td>
|
||||
<td id="76E75DECA626004DA8286D23FA098B9E" box="[1398,1451,376,401]" gridcol="9" gridrow="4" pageId="31" pageNumber="32">–</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6DC1FA098BBD" box="[113,1451,410,434]" gridrow="5" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6DC1FE428BBD" box="[113,480,410,434]" gridcol="0" gridrow="5" pageId="31" pageNumber="32">Pharyngeal tube length</th>
|
||||
<td id="76E75DECA626004DAF596DC1FDBB8BBD" box="[519,537,410,434]" gridcol="1" gridrow="5" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146DC1FD618BBD" box="[586,707,410,434]" gridcol="2" gridrow="5" pageId="31" pageNumber="32">15.6–26.1</td>
|
||||
<td id="76E75DECA626004DAFAF6DC1FCCB8BBD" box="[753,873,410,434]" gridcol="3" gridrow="5" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06DC1FCFE8BBD" box="[766,860,410,434]" italics="true" pageId="31" pageNumber="32">53.2–83.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96DC1FC6D8BBD" box="[919,975,410,434]" gridcol="4" gridrow="5" pageId="31" pageNumber="32">21.8</td>
|
||||
<td id="76E75DECA626004DAEA26DC1FB938BBD" box="[1020,1073,410,434]" gridcol="5" gridrow="5" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26DC1FB878BBD" box="[1020,1061,410,434]" italics="true" pageId="31" pageNumber="32">70.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6DC1FB2B8BBD" box="[1120,1161,410,434]" gridcol="6" gridrow="5" pageId="31" pageNumber="32">3.6</td>
|
||||
<td id="76E75DECA626004DA9E76DC1FB408BBD" box="[1209,1250,410,434]" gridcol="7" gridrow="5" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76DC1FB408BBD" box="[1209,1250,410,434]" italics="true" pageId="31" pageNumber="32">10.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6DC1FAE58BBD" box="[1297,1351,410,434]" gridcol="8" gridrow="5" pageId="31" pageNumber="32">23.5</td>
|
||||
<td id="76E75DECA626004DA8286DC1FA098BBD" box="[1398,1451,410,434]" gridcol="9" gridrow="5" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286DC1FA3D8BBD" box="[1398,1439,410,434]" italics="true" pageId="31" pageNumber="32">79.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6DE7FA098BDB" box="[113,1451,444,468]" gridrow="6" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6DE7FE428BDB" box="[113,480,444,468]" gridcol="0" gridrow="6" pageId="31" pageNumber="32">Buccopharyngeal tube length</th>
|
||||
<td id="76E75DECA626004DAF596DE7FDBB8BDB" box="[519,537,444,468]" gridcol="1" gridrow="6" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146DE7FD618BDB" box="[586,707,444,468]" gridcol="2" gridrow="6" pageId="31" pageNumber="32">44.9–58.8</td>
|
||||
<td id="76E75DECA626004DAFAF6DE7FCCB8BDB" box="[753,873,444,468]" gridcol="3" gridrow="6" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFAC6DE7FCCB8BDB" box="[754,873,444,468]" italics="true" pageId="31" pageNumber="32">153.2–183.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96DE7FC6D8BDB" box="[919,975,444,468]" gridcol="4" gridrow="6" pageId="31" pageNumber="32">52.7</td>
|
||||
<td id="76E75DECA626004DAEA26DE7FB938BDB" box="[1020,1073,444,468]" gridcol="5" gridrow="6" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26DE7FB938BDB" box="[1020,1073,444,468]" italics="true" pageId="31" pageNumber="32">170.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6DE7FB2B8BDB" box="[1120,1161,444,468]" gridcol="6" gridrow="6" pageId="31" pageNumber="32">4.7</td>
|
||||
<td id="76E75DECA626004DA9E76DE7FB408BDB" box="[1209,1250,444,468]" gridcol="7" gridrow="6" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76DE7FB408BDB" box="[1209,1250,444,468]" italics="true" pageId="31" pageNumber="32">10.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6DE7FAE58BDB" box="[1297,1351,444,468]" gridcol="8" gridrow="6" pageId="31" pageNumber="32">53.0</td>
|
||||
<td id="76E75DECA626004DA8286DE7FA098BDB" box="[1398,1451,444,468]" gridcol="9" gridrow="6" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286DE7FA098BDB" box="[1398,1451,444,468]" italics="true" pageId="31" pageNumber="32">179.7</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6D85FA098BF9" box="[113,1451,478,502]" gridrow="7" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6D85FE428BF9" box="[113,480,478,502]" gridcol="0" gridrow="7" pageId="31" pageNumber="32">Buccal/pharyngeal tube length ratio</th>
|
||||
<td id="76E75DECA626004DAF596D85FDBB8BF9" box="[519,537,478,502]" gridcol="1" gridrow="7" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146D85FD618BF9" box="[586,707,478,502]" gridcol="2" gridrow="7" pageId="31" pageNumber="32">119%–188%</td>
|
||||
<td id="76E75DECA626004DAFAF6D85FCCB8BF9" box="[753,873,478,502]" gridcol="3" gridrow="7" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DAEC96D85FC6D8BF9" box="[919,975,478,502]" gridcol="4" gridrow="7" pageId="31" pageNumber="32">144%</td>
|
||||
<td id="76E75DECA626004DAEA26D85FB938BF9" box="[1020,1073,478,502]" gridcol="5" gridrow="7" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DA93E6D85FB2B8BF9" box="[1120,1161,478,502]" gridcol="6" gridrow="7" pageId="31" pageNumber="32">23%</td>
|
||||
<td id="76E75DECA626004DA9E76D85FB408BF9" box="[1209,1250,478,502]" gridcol="7" gridrow="7" pageId="31" pageNumber="32">–</td>
|
||||
<td id="76E75DECA626004DA84F6D85FAE58BF9" box="[1297,1351,478,502]" gridcol="8" gridrow="7" pageId="31" pageNumber="32">126%</td>
|
||||
<td id="76E75DECA626004DA8286D85FA098BF9" box="[1398,1451,478,502]" gridcol="9" gridrow="7" pageId="31" pageNumber="32">–</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6E5BFA098817" box="[113,1451,512,536]" gridrow="8" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6E5BFE428817" box="[113,480,512,536]" gridcol="0" gridrow="8" pageId="31" pageNumber="32">Stylet support insertion point</th>
|
||||
<td id="76E75DECA626004DAF596E5BFDBB8817" box="[519,537,512,536]" gridcol="1" gridrow="8" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146E5BFD618817" box="[586,707,512,536]" gridcol="2" gridrow="8" pageId="31" pageNumber="32">23.4–28.0</td>
|
||||
<td id="76E75DECA626004DAFAF6E5BFCCB8817" box="[753,873,512,536]" gridcol="3" gridrow="8" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06E5BFCFE8817" box="[766,860,512,536]" italics="true" pageId="31" pageNumber="32">78.7–85.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96E5BFC6D8817" box="[919,975,512,536]" gridcol="4" gridrow="8" pageId="31" pageNumber="32">25.4</td>
|
||||
<td id="76E75DECA626004DAEA26E5BFB938817" box="[1020,1073,512,536]" gridcol="5" gridrow="8" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26E5BFB878817" box="[1020,1061,512,536]" italics="true" pageId="31" pageNumber="32">82.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6E5BFB2B8817" box="[1120,1161,512,536]" gridcol="6" gridrow="8" pageId="31" pageNumber="32">1.8</td>
|
||||
<td id="76E75DECA626004DA9E76E5BFB408817" box="[1209,1250,512,536]" gridcol="7" gridrow="8" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76E5BFB748817" box="[1209,1238,512,536]" italics="true" pageId="31" pageNumber="32">2.6</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6E5BFAE58817" box="[1297,1351,512,536]" gridcol="8" gridrow="8" pageId="31" pageNumber="32">25.1</td>
|
||||
<td id="76E75DECA626004DA8286E5BFA098817" box="[1398,1451,512,536]" gridcol="9" gridrow="8" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286E5BFA3D8817" box="[1398,1439,512,536]" italics="true" pageId="31" pageNumber="32">85.1</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6E79FA098835" box="[113,1451,546,570]" gridrow="9" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6E79FE428835" box="[113,480,546,570]" gridcol="0" gridrow="9" pageId="31" pageNumber="32">Buccal tube external width</th>
|
||||
<td id="76E75DECA626004DAF596E79FDBB8835" box="[519,537,546,570]" gridcol="1" gridrow="9" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146E79FD618835" box="[586,707,546,570]" gridcol="2" gridrow="9" pageId="31" pageNumber="32">3.1–4.1</td>
|
||||
<td id="76E75DECA626004DAFAF6E79FCCB8835" box="[753,873,546,570]" gridcol="3" gridrow="9" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06E79FCFE8835" box="[766,860,546,570]" italics="true" pageId="31" pageNumber="32">10.2–13.6</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96E79FC6D8835" box="[919,975,546,570]" gridcol="4" gridrow="9" pageId="31" pageNumber="32">3.7</td>
|
||||
<td id="76E75DECA626004DAEA26E79FB938835" box="[1020,1073,546,570]" gridcol="5" gridrow="9" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26E79FB878835" box="[1020,1061,546,570]" italics="true" pageId="31" pageNumber="32">12.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6E79FB2B8835" box="[1120,1161,546,570]" gridcol="6" gridrow="9" pageId="31" pageNumber="32">0.4</td>
|
||||
<td id="76E75DECA626004DA9E76E79FB408835" box="[1209,1250,546,570]" gridcol="7" gridrow="9" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76E79FB748835" box="[1209,1238,546,570]" italics="true" pageId="31" pageNumber="32">1.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6E79FAE58835" box="[1297,1351,546,570]" gridcol="8" gridrow="9" pageId="31" pageNumber="32">4.0</td>
|
||||
<td id="76E75DECA626004DA8286E79FA098835" box="[1398,1451,546,570]" gridcol="9" gridrow="9" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286E79FA3D8835" box="[1398,1439,546,570]" italics="true" pageId="31" pageNumber="32">13.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6E18FA098853" box="[113,1451,579,604]" gridrow="10" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6E18FE428853" box="[113,480,579,604]" gridcol="0" gridrow="10" pageId="31" pageNumber="32">Buccal tube internal width</th>
|
||||
<td id="76E75DECA626004DAF596E18FDBB8853" box="[519,537,579,604]" gridcol="1" gridrow="10" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146E18FD618853" box="[586,707,579,604]" gridcol="2" gridrow="10" pageId="31" pageNumber="32">2.0–2.8</td>
|
||||
<td id="76E75DECA626004DAFAF6E18FCCB8853" box="[753,873,579,604]" gridcol="3" gridrow="10" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE546E18FCF28854" box="[778,848,579,603]" italics="true" pageId="31" pageNumber="32">6.6–9.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96E18FC6D8853" box="[919,975,579,604]" gridcol="4" gridrow="10" pageId="31" pageNumber="32">2.5</td>
|
||||
<td id="76E75DECA626004DAEA26E18FB938853" box="[1020,1073,579,604]" gridcol="5" gridrow="10" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26E1FFBBB8853" box="[1020,1049,580,604]" italics="true" pageId="31" pageNumber="32">8.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6E18FB2B8853" box="[1120,1161,579,604]" gridcol="6" gridrow="10" pageId="31" pageNumber="32">0.3</td>
|
||||
<td id="76E75DECA626004DA9E76E18FB408853" box="[1209,1250,579,604]" gridcol="7" gridrow="10" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76E1FFB748853" box="[1209,1238,580,604]" italics="true" pageId="31" pageNumber="32">0.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6E18FAE58853" box="[1297,1351,579,604]" gridcol="8" gridrow="10" pageId="31" pageNumber="32">2.7</td>
|
||||
<td id="76E75DECA626004DA8286E18FA098853" box="[1398,1451,579,604]" gridcol="9" gridrow="10" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286E1FFA318853" box="[1398,1427,580,604]" italics="true" pageId="31" pageNumber="32">9.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6E3EFA098872" box="[113,1451,613,637]" gridrow="11" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F6E3EFE428872" box="[113,480,613,637]" gridcol="0" gridrow="11" pageId="31" pageNumber="32">Placoid lengths</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6EDCFA098890" box="[113,1451,647,671]" gridrow="12" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6EDCFE428890" box="[113,480,647,671]" gridcol="0" gridrow="12" pageId="31" pageNumber="32">Macroplacoid 1</th>
|
||||
<td id="76E75DECA626004DAF596EDCFDBB8890" box="[519,537,647,671]" gridcol="1" gridrow="12" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146EDCFD618890" box="[586,707,647,671]" gridcol="2" gridrow="12" pageId="31" pageNumber="32">2.5–4.1</td>
|
||||
<td id="76E75DECA626004DAFAF6EDCFCCB8890" box="[753,873,647,671]" gridcol="3" gridrow="12" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE546EDCFCFE8890" box="[778,860,647,671]" italics="true" pageId="31" pageNumber="32">8.5–12.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96EDCFC6D8890" box="[919,975,647,671]" gridcol="4" gridrow="12" pageId="31" pageNumber="32">3.6</td>
|
||||
<td id="76E75DECA626004DAEA26EDCFB938890" box="[1020,1073,647,671]" gridcol="5" gridrow="12" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26EDCFB878890" box="[1020,1061,647,671]" italics="true" pageId="31" pageNumber="32">11.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6EDCFB2B8890" box="[1120,1161,647,671]" gridcol="6" gridrow="12" pageId="31" pageNumber="32">0.5</td>
|
||||
<td id="76E75DECA626004DA9E76EDCFB408890" box="[1209,1250,647,671]" gridcol="7" gridrow="12" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76EDCFB748890" box="[1209,1238,647,671]" italics="true" pageId="31" pageNumber="32">1.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6EDCFAE58890" box="[1297,1351,647,671]" gridcol="8" gridrow="12" pageId="31" pageNumber="32">3.7</td>
|
||||
<td id="76E75DECA626004DA8286EDCFA098890" box="[1398,1451,647,671]" gridcol="9" gridrow="12" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286EDCFA3D8890" box="[1398,1439,647,671]" italics="true" pageId="31" pageNumber="32">12.5</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6EF2FA0988CE" box="[113,1451,681,705]" gridrow="13" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6EF2FE4288CE" box="[113,480,681,705]" gridcol="0" gridrow="13" pageId="31" pageNumber="32">Macroplacoid 2</th>
|
||||
<td id="76E75DECA626004DAF596EF2FDBB88CE" box="[519,537,681,705]" gridcol="1" gridrow="13" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146EF2FD6188CE" box="[586,707,681,705]" gridcol="2" gridrow="13" pageId="31" pageNumber="32">8.5–12.1</td>
|
||||
<td id="76E75DECA626004DAFAF6EF2FCCB88CE" box="[753,873,681,705]" gridcol="3" gridrow="13" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06EF2FCFE88CE" box="[766,860,681,705]" italics="true" pageId="31" pageNumber="32">29.0–36.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96EF2FC6D88CE" box="[919,975,681,705]" gridcol="4" gridrow="13" pageId="31" pageNumber="32">10.2</td>
|
||||
<td id="76E75DECA626004DAEA26EF2FB9388CE" box="[1020,1073,681,705]" gridcol="5" gridrow="13" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26EF2FB8788CE" box="[1020,1061,681,705]" italics="true" pageId="31" pageNumber="32">32.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6EF2FB2B88CE" box="[1120,1161,681,705]" gridcol="6" gridrow="13" pageId="31" pageNumber="32">1.4</td>
|
||||
<td id="76E75DECA626004DA9E76EF2FB4088CE" box="[1209,1250,681,705]" gridcol="7" gridrow="13" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76EF2FB7488CE" box="[1209,1238,681,705]" italics="true" pageId="31" pageNumber="32">3.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6EF2FAE588CE" box="[1297,1351,681,705]" gridcol="8" gridrow="13" pageId="31" pageNumber="32">10.0</td>
|
||||
<td id="76E75DECA626004DA8286EF2FA0988CE" box="[1398,1451,681,705]" gridcol="9" gridrow="13" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286EF2FA3D88CE" box="[1398,1439,681,705]" italics="true" pageId="31" pageNumber="32">33.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6E90FA0988EC" box="[113,1451,715,739]" gridrow="14" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6E90FE4288EC" box="[113,480,715,739]" gridcol="0" gridrow="14" pageId="31" pageNumber="32">Macroplacoid row</th>
|
||||
<td id="76E75DECA626004DAF596E90FDBB88EC" box="[519,537,715,739]" gridcol="1" gridrow="14" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146E90FD6188EC" box="[586,707,715,739]" gridcol="2" gridrow="14" pageId="31" pageNumber="32">11.7–16.8</td>
|
||||
<td id="76E75DECA626004DAFAF6E90FCCB88EC" box="[753,873,715,739]" gridcol="3" gridrow="14" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06E90FCFE88EC" box="[766,860,715,739]" italics="true" pageId="31" pageNumber="32">39.9–50.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96E90FC6D88EC" box="[919,975,715,739]" gridcol="4" gridrow="14" pageId="31" pageNumber="32">14.6</td>
|
||||
<td id="76E75DECA626004DAEA26E90FB9388EC" box="[1020,1073,715,739]" gridcol="5" gridrow="14" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26E90FB8788EC" box="[1020,1061,715,739]" italics="true" pageId="31" pageNumber="32">47.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6E90FB2B88EC" box="[1120,1161,715,739]" gridcol="6" gridrow="14" pageId="31" pageNumber="32">1.8</td>
|
||||
<td id="76E75DECA626004DA9E76E90FB4088EC" box="[1209,1250,715,739]" gridcol="7" gridrow="14" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76E90FB7488EC" box="[1209,1238,715,739]" italics="true" pageId="31" pageNumber="32">4.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6E90FAE588EC" box="[1297,1351,715,739]" gridcol="8" gridrow="14" pageId="31" pageNumber="32">14.6</td>
|
||||
<td id="76E75DECA626004DA8286E90FA0988EC" box="[1398,1451,715,739]" gridcol="9" gridrow="14" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286E90FA3D88EC" box="[1398,1439,715,739]" italics="true" pageId="31" pageNumber="32">49.5</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6EB6FA09890A" box="[113,1451,749,773]" gridrow="15" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F6EB6FE42890A" box="[113,480,749,773]" gridcol="0" gridrow="15" pageId="31" pageNumber="32">Claw 1 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6F55FA098928" box="[113,1451,782,807]" gridrow="16" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6F55FE428928" box="[113,480,782,807]" gridcol="0" gridrow="16" pageId="31" pageNumber="32">External base</th>
|
||||
<td id="76E75DECA626004DAF596F55FDBB8928" box="[519,537,782,807]" gridcol="1" gridrow="16" pageId="31" pageNumber="32">4</td>
|
||||
<td id="76E75DECA626004DAF146F55FD618928" box="[586,707,782,807]" gridcol="2" gridrow="16" pageId="31" pageNumber="32">2.6–3.5</td>
|
||||
<td id="76E75DECA626004DAFAF6F55FCCB8928" box="[753,873,782,807]" gridcol="3" gridrow="16" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE546F54FCFE8928" box="[778,860,783,807]" italics="true" pageId="31" pageNumber="32">8.8–11.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96F55FC6D8928" box="[919,975,782,807]" gridcol="4" gridrow="16" pageId="31" pageNumber="32">3.0</td>
|
||||
<td id="76E75DECA626004DAEA26F55FB938928" box="[1020,1073,782,807]" gridcol="5" gridrow="16" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26F54FBBB8928" box="[1020,1049,783,807]" italics="true" pageId="31" pageNumber="32">9.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6F55FB2B8928" box="[1120,1161,782,807]" gridcol="6" gridrow="16" pageId="31" pageNumber="32">0.4</td>
|
||||
<td id="76E75DECA626004DA9E76F55FB408928" box="[1209,1250,782,807]" gridcol="7" gridrow="16" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76F54FB748928" box="[1209,1238,783,807]" italics="true" pageId="31" pageNumber="32">1.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6F55FAE58928" box="[1297,1351,782,807]" gridcol="8" gridrow="16" pageId="31" pageNumber="32">3.5</td>
|
||||
<td id="76E75DECA626004DA8286F55FA098928" box="[1398,1451,782,807]" gridcol="9" gridrow="16" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286F54FA3D8928" box="[1398,1439,783,807]" italics="true" pageId="31" pageNumber="32">11.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6F6BFA098946" box="[113,1451,816,841]" gridrow="17" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6F6BFE428946" box="[113,480,816,841]" gridcol="0" gridrow="17" pageId="31" pageNumber="32">External primary branch</th>
|
||||
<td id="76E75DECA626004DAF596F6BFDBB8946" box="[519,537,816,841]" gridcol="1" gridrow="17" pageId="31" pageNumber="32">4</td>
|
||||
<td id="76E75DECA626004DAF146F6BFD618946" box="[586,707,816,841]" gridcol="2" gridrow="17" pageId="31" pageNumber="32">6.7–7.5</td>
|
||||
<td id="76E75DECA626004DAFAF6F6BFCCB8946" box="[753,873,816,841]" gridcol="3" gridrow="17" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06F6BFCFF8947" box="[766,861,816,840]" italics="true" pageId="31" pageNumber="32">22.6–25.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96F6BFC6D8946" box="[919,975,816,841]" gridcol="4" gridrow="17" pageId="31" pageNumber="32">7.1</td>
|
||||
<td id="76E75DECA626004DAEA26F6BFB938946" box="[1020,1073,816,841]" gridcol="5" gridrow="17" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26F6AFB878946" box="[1020,1061,817,841]" italics="true" pageId="31" pageNumber="32">24.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6F6BFB2B8946" box="[1120,1161,816,841]" gridcol="6" gridrow="17" pageId="31" pageNumber="32">0.4</td>
|
||||
<td id="76E75DECA626004DA9E76F6BFB408946" box="[1209,1250,816,841]" gridcol="7" gridrow="17" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76F6AFB748946" box="[1209,1238,817,841]" italics="true" pageId="31" pageNumber="32">1.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6F6BFAE58946" box="[1297,1351,816,841]" gridcol="8" gridrow="17" pageId="31" pageNumber="32">7.5</td>
|
||||
<td id="76E75DECA626004DA8286F6BFA098946" box="[1398,1451,816,841]" gridcol="9" gridrow="17" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286F6AFA3D8946" box="[1398,1439,817,841]" italics="true" pageId="31" pageNumber="32">25.4</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6F09FA098965" box="[113,1451,850,874]" gridrow="18" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6F09FE428965" box="[113,480,850,874]" gridcol="0" gridrow="18" pageId="31" pageNumber="32">External secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596F09FDBB8965" box="[519,537,850,874]" gridcol="1" gridrow="18" pageId="31" pageNumber="32">4</td>
|
||||
<td id="76E75DECA626004DAF146F09FD618965" box="[586,707,850,874]" gridcol="2" gridrow="18" pageId="31" pageNumber="32">3.9–4.6</td>
|
||||
<td id="76E75DECA626004DAFAF6F09FCCB8965" box="[753,873,850,874]" gridcol="3" gridrow="18" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06F09FCFE8965" box="[766,860,850,874]" italics="true" pageId="31" pageNumber="32">12.8–14.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96F09FC6D8965" box="[919,975,850,874]" gridcol="4" gridrow="18" pageId="31" pageNumber="32">4.2</td>
|
||||
<td id="76E75DECA626004DAEA26F09FB938965" box="[1020,1073,850,874]" gridcol="5" gridrow="18" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26F09FB878965" box="[1020,1061,850,874]" italics="true" pageId="31" pageNumber="32">13.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6F09FB2B8965" box="[1120,1161,850,874]" gridcol="6" gridrow="18" pageId="31" pageNumber="32">0.3</td>
|
||||
<td id="76E75DECA626004DA9E76F09FB408965" box="[1209,1250,850,874]" gridcol="7" gridrow="18" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76F09FB748965" box="[1209,1238,850,874]" italics="true" pageId="31" pageNumber="32">0.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6F09FAE58965" box="[1297,1351,850,874]" gridcol="8" gridrow="18" pageId="31" pageNumber="32">4.1</td>
|
||||
<td id="76E75DECA626004DA8286F09FA098965" box="[1398,1451,850,874]" gridcol="9" gridrow="18" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286F09FA3D8965" box="[1398,1439,850,874]" italics="true" pageId="31" pageNumber="32">13.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6F2FFA098983" box="[113,1451,884,908]" gridrow="19" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6F2FFE428983" box="[113,480,884,908]" gridcol="0" gridrow="19" pageId="31" pageNumber="32">Internal base</th>
|
||||
<td id="76E75DECA626004DAF596F2FFDBB8983" box="[519,537,884,908]" gridcol="1" gridrow="19" pageId="31" pageNumber="32">1</td>
|
||||
<td id="76E75DECA626004DAF146F2FFD618983" box="[586,707,884,908]" gridcol="2" gridrow="19" pageId="31" pageNumber="32">3.7–3.7</td>
|
||||
<td id="76E75DECA626004DAFAF6F2FFCCB8983" box="[753,873,884,908]" gridcol="3" gridrow="19" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06F2FFCFE8983" box="[766,860,884,908]" italics="true" pageId="31" pageNumber="32">11.0–11.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96F2FFC6D8983" box="[919,975,884,908]" gridcol="4" gridrow="19" pageId="31" pageNumber="32">3.7</td>
|
||||
<td id="76E75DECA626004DAEA26F2FFB938983" box="[1020,1073,884,908]" gridcol="5" gridrow="19" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26F2FFB878983" box="[1020,1061,884,908]" italics="true" pageId="31" pageNumber="32">11.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6F2FFB2B8983" box="[1120,1161,884,908]" gridcol="6" gridrow="19" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76F2FFB408983" box="[1209,1250,884,908]" gridcol="7" gridrow="19" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76F2EFB608983" box="[1209,1218,885,908]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6F2FFAE58983" box="[1297,1351,884,908]" gridcol="8" gridrow="19" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286F2FFA098983" box="[1398,1451,884,908]" gridcol="9" gridrow="19" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286F2EFADD8983" box="[1398,1407,885,908]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6FCDFA0989A1" box="[113,1451,918,942]" gridrow="20" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6FCDFE4289A1" box="[113,480,918,942]" gridcol="0" gridrow="20" pageId="31" pageNumber="32">Internal primary branch</th>
|
||||
<td id="76E75DECA626004DAF596FCDFDBB89A1" box="[519,537,918,942]" gridcol="1" gridrow="20" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF146FCDFD6189A1" box="[586,707,918,942]" gridcol="2" gridrow="20" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF6FCDFCCB89A1" box="[753,873,918,942]" gridcol="3" gridrow="20" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE766FCDFC9389A2" box="[808,817,918,941]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96FCDFC6D89A1" box="[919,975,918,942]" gridcol="4" gridrow="20" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA26FCDFB9389A1" box="[1020,1073,918,942]" gridcol="5" gridrow="20" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26FCDFBA789A2" box="[1020,1029,918,941]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6FCDFB2B89A1" box="[1120,1161,918,942]" gridcol="6" gridrow="20" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76FCDFB4089A1" box="[1209,1250,918,942]" gridcol="7" gridrow="20" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76FCDFB6089A2" box="[1209,1218,918,941]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6FCDFAE589A1" box="[1297,1351,918,942]" gridcol="8" gridrow="20" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286FCDFA0989A1" box="[1398,1451,918,942]" gridcol="9" gridrow="20" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286FCDFADD89A2" box="[1398,1407,918,941]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6FE3FA0989DF" box="[113,1451,952,976]" gridrow="21" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6FE3FE4289DF" box="[113,480,952,976]" gridcol="0" gridrow="21" pageId="31" pageNumber="32">Internal secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596FE3FDBB89DF" box="[519,537,952,976]" gridcol="1" gridrow="21" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF146FE3FD6189DF" box="[586,707,952,976]" gridcol="2" gridrow="21" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF6FE3FCCB89DF" box="[753,873,952,976]" gridcol="3" gridrow="21" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE766FE3FC9389C0" box="[808,817,952,975]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96FE3FC6D89DF" box="[919,975,952,976]" gridcol="4" gridrow="21" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA26FE3FB9389DF" box="[1020,1073,952,976]" gridcol="5" gridrow="21" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26FE3FBA789C0" box="[1020,1029,952,975]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6FE3FB2B89DF" box="[1120,1161,952,976]" gridcol="6" gridrow="21" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76FE3FB4089DF" box="[1209,1250,952,976]" gridcol="7" gridrow="21" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76FE3FB6089C0" box="[1209,1218,952,975]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6FE3FAE589DF" box="[1297,1351,952,976]" gridcol="8" gridrow="21" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286FE3FA0989DF" box="[1398,1451,952,976]" gridcol="9" gridrow="21" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286FE3FADD89C0" box="[1398,1407,952,975]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6F81FA0989FD" box="[113,1451,986,1010]" gridrow="22" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F6F81FE4289FD" box="[113,480,986,1010]" gridcol="0" gridrow="22" pageId="31" pageNumber="32">Claw 2 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6FA0FA098E1B" box="[113,1451,1019,1044]" gridrow="23" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6FA0FE428E1B" box="[113,480,1019,1044]" gridcol="0" gridrow="23" pageId="31" pageNumber="32">External base</th>
|
||||
<td id="76E75DECA626004DAF596FA0FDBB8E1B" box="[519,537,1019,1044]" gridcol="1" gridrow="23" pageId="31" pageNumber="32">8</td>
|
||||
<td id="76E75DECA626004DAF146FA0FD618E1B" box="[586,707,1019,1044]" gridcol="2" gridrow="23" pageId="31" pageNumber="32">2.6–4.5</td>
|
||||
<td id="76E75DECA626004DAFAF6FA0FCCB8E1B" box="[753,873,1019,1044]" gridcol="3" gridrow="23" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE546FA7FCFE8E1B" box="[778,860,1020,1044]" italics="true" pageId="31" pageNumber="32">8.9–14.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96FA0FC6D8E1B" box="[919,975,1019,1044]" gridcol="4" gridrow="23" pageId="31" pageNumber="32">3.4</td>
|
||||
<td id="76E75DECA626004DAEA26FA0FB938E1B" box="[1020,1073,1019,1044]" gridcol="5" gridrow="23" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26FA7FB878E1B" box="[1020,1061,1020,1044]" italics="true" pageId="31" pageNumber="32">11.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6FA0FB2B8E1B" box="[1120,1161,1019,1044]" gridcol="6" gridrow="23" pageId="31" pageNumber="32">0.7</td>
|
||||
<td id="76E75DECA626004DA9E76FA0FB408E1B" box="[1209,1250,1019,1044]" gridcol="7" gridrow="23" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76FA7FB748E1B" box="[1209,1238,1020,1044]" italics="true" pageId="31" pageNumber="32">2.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6FA0FAE58E1B" box="[1297,1351,1019,1044]" gridcol="8" gridrow="23" pageId="31" pageNumber="32">3.3</td>
|
||||
<td id="76E75DECA626004DA8286FA0FA098E1B" box="[1398,1451,1019,1044]" gridcol="9" gridrow="23" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286FA7FA3D8E1B" box="[1398,1439,1020,1044]" italics="true" pageId="31" pageNumber="32">11.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6846FA098E3A" box="[113,1451,1053,1077]" gridrow="24" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6846FE428E3A" box="[113,480,1053,1077]" gridcol="0" gridrow="24" pageId="31" pageNumber="32">External primary branch</th>
|
||||
<td id="76E75DECA626004DAF596846FDBB8E3A" box="[519,537,1053,1077]" gridcol="1" gridrow="24" pageId="31" pageNumber="32">8</td>
|
||||
<td id="76E75DECA626004DAF146846FD618E3A" box="[586,707,1053,1077]" gridcol="2" gridrow="24" pageId="31" pageNumber="32">6.5–9.4</td>
|
||||
<td id="76E75DECA626004DAFAF6846FCCB8E3A" box="[753,873,1053,1077]" gridcol="3" gridrow="24" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06846FCFF8E3A" box="[766,861,1053,1077]" italics="true" pageId="31" pageNumber="32">22.2–29.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96846FC6D8E3A" box="[919,975,1053,1077]" gridcol="4" gridrow="24" pageId="31" pageNumber="32">7.9</td>
|
||||
<td id="76E75DECA626004DAEA26846FB938E3A" box="[1020,1073,1053,1077]" gridcol="5" gridrow="24" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26846FB878E3A" box="[1020,1061,1053,1077]" italics="true" pageId="31" pageNumber="32">25.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6846FB2B8E3A" box="[1120,1161,1053,1077]" gridcol="6" gridrow="24" pageId="31" pageNumber="32">1.0</td>
|
||||
<td id="76E75DECA626004DA9E76846FB408E3A" box="[1209,1250,1053,1077]" gridcol="7" gridrow="24" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76846FB748E3A" box="[1209,1238,1053,1077]" italics="true" pageId="31" pageNumber="32">2.9</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6846FAE58E3A" box="[1297,1351,1053,1077]" gridcol="8" gridrow="24" pageId="31" pageNumber="32">8.7</td>
|
||||
<td id="76E75DECA626004DA8286846FA098E3A" box="[1398,1451,1053,1077]" gridcol="9" gridrow="24" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286846FA3D8E3A" box="[1398,1439,1053,1077]" italics="true" pageId="31" pageNumber="32">29.5</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6864FA098E58" box="[113,1451,1087,1111]" gridrow="25" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6864FE428E58" box="[113,480,1087,1111]" gridcol="0" gridrow="25" pageId="31" pageNumber="32">External secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596864FDBB8E58" box="[519,537,1087,1111]" gridcol="1" gridrow="25" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146864FD618E58" box="[586,707,1087,1111]" gridcol="2" gridrow="25" pageId="31" pageNumber="32">3.9–5.2</td>
|
||||
<td id="76E75DECA626004DAFAF6864FCCB8E58" box="[753,873,1087,1111]" gridcol="3" gridrow="25" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06864FCFE8E58" box="[766,860,1087,1111]" italics="true" pageId="31" pageNumber="32">13.3–16.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96864FC6D8E58" box="[919,975,1087,1111]" gridcol="4" gridrow="25" pageId="31" pageNumber="32">4.6</td>
|
||||
<td id="76E75DECA626004DAEA26864FB938E58" box="[1020,1073,1087,1111]" gridcol="5" gridrow="25" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26864FB878E58" box="[1020,1061,1087,1111]" italics="true" pageId="31" pageNumber="32">14.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6864FB2B8E58" box="[1120,1161,1087,1111]" gridcol="6" gridrow="25" pageId="31" pageNumber="32">0.5</td>
|
||||
<td id="76E75DECA626004DA9E76864FB408E58" box="[1209,1250,1087,1111]" gridcol="7" gridrow="25" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76864FB748E58" box="[1209,1238,1087,1111]" italics="true" pageId="31" pageNumber="32">1.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6864FAE58E58" box="[1297,1351,1087,1111]" gridcol="8" gridrow="25" pageId="31" pageNumber="32">4.6</td>
|
||||
<td id="76E75DECA626004DA8286864FA098E58" box="[1398,1451,1087,1111]" gridcol="9" gridrow="25" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286864FA3D8E58" box="[1398,1439,1087,1111]" italics="true" pageId="31" pageNumber="32">15.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F683AFA098E76" box="[113,1451,1121,1145]" gridrow="26" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F683AFE428E76" box="[113,480,1121,1145]" gridcol="0" gridrow="26" pageId="31" pageNumber="32">Internal base</th>
|
||||
<td id="76E75DECA626004DAF59683AFDBB8E76" box="[519,537,1121,1145]" gridcol="1" gridrow="26" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF14683AFD618E76" box="[586,707,1121,1145]" gridcol="2" gridrow="26" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF683AFCCB8E76" box="[753,873,1121,1145]" gridcol="3" gridrow="26" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE766839FC938E76" box="[808,817,1122,1145]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC9683AFC6D8E76" box="[919,975,1121,1145]" gridcol="4" gridrow="26" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA2683AFB938E76" box="[1020,1073,1121,1145]" gridcol="5" gridrow="26" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26839FBA78E76" box="[1020,1029,1122,1145]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E683AFB2B8E76" box="[1120,1161,1121,1145]" gridcol="6" gridrow="26" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E7683AFB408E76" box="[1209,1250,1121,1145]" gridcol="7" gridrow="26" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76839FB608E76" box="[1209,1218,1122,1145]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F683AFAE58E76" box="[1297,1351,1121,1145]" gridcol="8" gridrow="26" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA828683AFA098E76" box="[1398,1451,1121,1145]" gridcol="9" gridrow="26" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286839FADD8E76" box="[1398,1407,1122,1145]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F68D8FA098E94" box="[113,1451,1155,1179]" gridrow="27" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F68D8FE428E94" box="[113,480,1155,1179]" gridcol="0" gridrow="27" pageId="31" pageNumber="32">Internal primary branch</th>
|
||||
<td id="76E75DECA626004DAF5968D8FDBB8E94" box="[519,537,1155,1179]" gridcol="1" gridrow="27" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF1468D8FD618E94" box="[586,707,1155,1179]" gridcol="2" gridrow="27" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF68D8FCCB8E94" box="[753,873,1155,1179]" gridcol="3" gridrow="27" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE7668D8FC938E95" box="[808,817,1155,1178]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC968D8FC6D8E94" box="[919,975,1155,1179]" gridcol="4" gridrow="27" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA268D8FB938E94" box="[1020,1073,1155,1179]" gridcol="5" gridrow="27" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA268D8FBA78E95" box="[1020,1029,1155,1178]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E68D8FB2B8E94" box="[1120,1161,1155,1179]" gridcol="6" gridrow="27" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E768D8FB408E94" box="[1209,1250,1155,1179]" gridcol="7" gridrow="27" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E768D8FB608E95" box="[1209,1218,1155,1178]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F68D8FAE58E94" box="[1297,1351,1155,1179]" gridcol="8" gridrow="27" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA82868D8FA098E94" box="[1398,1451,1155,1179]" gridcol="9" gridrow="27" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA82868D8FADD8E95" box="[1398,1407,1155,1178]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F68FEFA098EB2" box="[113,1451,1189,1213]" gridrow="28" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F68FEFE428EB2" box="[113,480,1189,1213]" gridcol="0" gridrow="28" pageId="31" pageNumber="32">Internal secondary branch</th>
|
||||
<td id="76E75DECA626004DAF5968FEFDBB8EB2" box="[519,537,1189,1213]" gridcol="1" gridrow="28" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF1468FEFD618EB2" box="[586,707,1189,1213]" gridcol="2" gridrow="28" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF68FEFCCB8EB2" box="[753,873,1189,1213]" gridcol="3" gridrow="28" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE7668FEFC938EB3" box="[808,817,1189,1212]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC968FEFC6D8EB2" box="[919,975,1189,1213]" gridcol="4" gridrow="28" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA268FEFB938EB2" box="[1020,1073,1189,1213]" gridcol="5" gridrow="28" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA268FEFBA78EB3" box="[1020,1029,1189,1212]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E68FEFB2B8EB2" box="[1120,1161,1189,1213]" gridcol="6" gridrow="28" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E768FEFB408EB2" box="[1209,1250,1189,1213]" gridcol="7" gridrow="28" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E768FEFB608EB3" box="[1209,1218,1189,1212]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F68FEFAE58EB2" box="[1297,1351,1189,1213]" gridcol="8" gridrow="28" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA82868FEFA098EB2" box="[1398,1451,1189,1213]" gridcol="9" gridrow="28" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA82868FEFADD8EB3" box="[1398,1407,1189,1212]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F689CFA098ED0" box="[113,1451,1223,1247]" gridrow="29" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F689CFE428ED0" box="[113,480,1223,1247]" gridcol="0" gridrow="29" pageId="31" pageNumber="32">Claw 3 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F68B3FA098F0E" box="[113,1451,1256,1281]" gridrow="30" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F68B3FE428F0E" box="[113,480,1256,1281]" gridcol="0" gridrow="30" pageId="31" pageNumber="32">External base</th>
|
||||
<td id="76E75DECA626004DAF5968B3FDBB8F0E" box="[519,537,1256,1281]" gridcol="1" gridrow="30" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF1468B3FD618F0E" box="[586,707,1256,1281]" gridcol="2" gridrow="30" pageId="31" pageNumber="32">2.6–5.6</td>
|
||||
<td id="76E75DECA626004DAFAF68B3FCCB8F0E" box="[753,873,1256,1281]" gridcol="3" gridrow="30" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE5468B2FCFF8F0E" box="[778,861,1257,1281]" italics="true" pageId="31" pageNumber="32">8.9–17.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC968B3FC6D8F0E" box="[919,975,1256,1281]" gridcol="4" gridrow="30" pageId="31" pageNumber="32">3.6</td>
|
||||
<td id="76E75DECA626004DAEA268B3FB938F0E" box="[1020,1073,1256,1281]" gridcol="5" gridrow="30" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA268B2FB878F0E" box="[1020,1061,1257,1281]" italics="true" pageId="31" pageNumber="32">12.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E68B3FB2B8F0E" box="[1120,1161,1256,1281]" gridcol="6" gridrow="30" pageId="31" pageNumber="32">1.0</td>
|
||||
<td id="76E75DECA626004DA9E768B3FB408F0E" box="[1209,1250,1256,1281]" gridcol="7" gridrow="30" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E768B2FB748F0E" box="[1209,1238,1257,1281]" italics="true" pageId="31" pageNumber="32">3.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F68B3FAE58F0E" box="[1297,1351,1256,1281]" gridcol="8" gridrow="30" pageId="31" pageNumber="32">3.7</td>
|
||||
<td id="76E75DECA626004DA82868B3FA098F0E" box="[1398,1451,1256,1281]" gridcol="9" gridrow="30" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA82868B2FA3D8F0E" box="[1398,1439,1257,1281]" italics="true" pageId="31" pageNumber="32">12.5</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6951FA098F2D" box="[113,1451,1290,1314]" gridrow="31" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6951FE428F2D" box="[113,480,1290,1314]" gridcol="0" gridrow="31" pageId="31" pageNumber="32">External primary branch</th>
|
||||
<td id="76E75DECA626004DAF596951FDBB8F2D" box="[519,537,1290,1314]" gridcol="1" gridrow="31" pageId="31" pageNumber="32">7</td>
|
||||
<td id="76E75DECA626004DAF146951FD618F2D" box="[586,707,1290,1314]" gridcol="2" gridrow="31" pageId="31" pageNumber="32">6.7–9.4</td>
|
||||
<td id="76E75DECA626004DAFAF6951FCCB8F2D" box="[753,873,1290,1314]" gridcol="3" gridrow="31" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06951FCFE8F2D" box="[766,860,1290,1314]" italics="true" pageId="31" pageNumber="32">22.9–29.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96951FC6D8F2D" box="[919,975,1290,1314]" gridcol="4" gridrow="31" pageId="31" pageNumber="32">8.0</td>
|
||||
<td id="76E75DECA626004DAEA26951FB938F2D" box="[1020,1073,1290,1314]" gridcol="5" gridrow="31" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26951FB878F2D" box="[1020,1061,1290,1314]" italics="true" pageId="31" pageNumber="32">26.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6951FB2B8F2D" box="[1120,1161,1290,1314]" gridcol="6" gridrow="31" pageId="31" pageNumber="32">0.9</td>
|
||||
<td id="76E75DECA626004DA9E76951FB408F2D" box="[1209,1250,1290,1314]" gridcol="7" gridrow="31" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76951FB748F2D" box="[1209,1238,1290,1314]" italics="true" pageId="31" pageNumber="32">2.8</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6951FAE58F2D" box="[1297,1351,1290,1314]" gridcol="8" gridrow="31" pageId="31" pageNumber="32">8.6</td>
|
||||
<td id="76E75DECA626004DA8286951FA098F2D" box="[1398,1451,1290,1314]" gridcol="9" gridrow="31" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286951FA3D8F2D" box="[1398,1439,1290,1314]" italics="true" pageId="31" pageNumber="32">29.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6977FA098F4B" box="[113,1451,1324,1348]" gridrow="32" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6977FE428F4B" box="[113,480,1324,1348]" gridcol="0" gridrow="32" pageId="31" pageNumber="32">External secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596977FDBB8F4B" box="[519,537,1324,1348]" gridcol="1" gridrow="32" pageId="31" pageNumber="32">6</td>
|
||||
<td id="76E75DECA626004DAF146977FD618F4B" box="[586,707,1324,1348]" gridcol="2" gridrow="32" pageId="31" pageNumber="32">3.9–5.2</td>
|
||||
<td id="76E75DECA626004DAFAF6977FCCB8F4B" box="[753,873,1324,1348]" gridcol="3" gridrow="32" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06977FCFF8F4B" box="[766,861,1324,1348]" italics="true" pageId="31" pageNumber="32">13.3–17.6</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96977FC6D8F4B" box="[919,975,1324,1348]" gridcol="4" gridrow="32" pageId="31" pageNumber="32">4.7</td>
|
||||
<td id="76E75DECA626004DAEA26977FB938F4B" box="[1020,1073,1324,1348]" gridcol="5" gridrow="32" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26977FB878F4B" box="[1020,1061,1324,1348]" italics="true" pageId="31" pageNumber="32">15.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6977FB2B8F4B" box="[1120,1161,1324,1348]" gridcol="6" gridrow="32" pageId="31" pageNumber="32">0.5</td>
|
||||
<td id="76E75DECA626004DA9E76977FB408F4B" box="[1209,1250,1324,1348]" gridcol="7" gridrow="32" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76977FB748F4B" box="[1209,1238,1324,1348]" italics="true" pageId="31" pageNumber="32">1.7</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6977FAE58F4B" box="[1297,1351,1324,1348]" gridcol="8" gridrow="32" pageId="31" pageNumber="32">5.2</td>
|
||||
<td id="76E75DECA626004DA8286977FA098F4B" box="[1398,1451,1324,1348]" gridcol="9" gridrow="32" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286977FA3D8F4B" box="[1398,1439,1324,1348]" italics="true" pageId="31" pageNumber="32">17.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6915FA098F69" box="[113,1451,1358,1382]" gridrow="33" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6915FE428F69" box="[113,480,1358,1382]" gridcol="0" gridrow="33" pageId="31" pageNumber="32">Internal base</th>
|
||||
<td id="76E75DECA626004DAF596915FDBB8F69" box="[519,537,1358,1382]" gridcol="1" gridrow="33" pageId="31" pageNumber="32">1</td>
|
||||
<td id="76E75DECA626004DAF146915FD618F69" box="[586,707,1358,1382]" gridcol="2" gridrow="33" pageId="31" pageNumber="32">3.3–3.3</td>
|
||||
<td id="76E75DECA626004DAFAF6915FCCB8F69" box="[753,873,1358,1382]" gridcol="3" gridrow="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06915FCFE8F69" box="[766,860,1358,1382]" italics="true" pageId="31" pageNumber="32">11.1–11.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96915FC6D8F69" box="[919,975,1358,1382]" gridcol="4" gridrow="33" pageId="31" pageNumber="32">3.3</td>
|
||||
<td id="76E75DECA626004DAEA26915FB938F69" box="[1020,1073,1358,1382]" gridcol="5" gridrow="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26915FB878F69" box="[1020,1061,1358,1382]" italics="true" pageId="31" pageNumber="32">11.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6915FB2B8F69" box="[1120,1161,1358,1382]" gridcol="6" gridrow="33" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76915FB408F69" box="[1209,1250,1358,1382]" gridcol="7" gridrow="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76915FB608F6A" box="[1209,1218,1358,1381]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6915FAE58F69" box="[1297,1351,1358,1382]" gridcol="8" gridrow="33" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286915FA098F69" box="[1398,1451,1358,1382]" gridcol="9" gridrow="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286915FADD8F6A" box="[1398,1407,1358,1381]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F692BFA098F87" box="[113,1451,1392,1416]" gridrow="34" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F692BFE428F87" box="[113,480,1392,1416]" gridcol="0" gridrow="34" pageId="31" pageNumber="32">Internal primary branch</th>
|
||||
<td id="76E75DECA626004DAF59692BFDBB8F87" box="[519,537,1392,1416]" gridcol="1" gridrow="34" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF14692BFD618F87" box="[586,707,1392,1416]" gridcol="2" gridrow="34" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF692BFCCB8F87" box="[753,873,1392,1416]" gridcol="3" gridrow="34" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE76692BFC938F88" box="[808,817,1392,1415]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC9692BFC6D8F87" box="[919,975,1392,1416]" gridcol="4" gridrow="34" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA2692BFB938F87" box="[1020,1073,1392,1416]" gridcol="5" gridrow="34" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA2692BFBA78F88" box="[1020,1029,1392,1415]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E692BFB2B8F87" box="[1120,1161,1392,1416]" gridcol="6" gridrow="34" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E7692BFB408F87" box="[1209,1250,1392,1416]" gridcol="7" gridrow="34" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E7692BFB608F88" box="[1209,1218,1392,1415]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F692BFAE58F87" box="[1297,1351,1392,1416]" gridcol="8" gridrow="34" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA828692BFA098F87" box="[1398,1451,1392,1416]" gridcol="9" gridrow="34" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA828692BFADD8F88" box="[1398,1407,1392,1415]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F69C9FA098FA5" box="[113,1451,1426,1450]" gridrow="35" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F69C9FE428FA5" box="[113,480,1426,1450]" gridcol="0" gridrow="35" pageId="31" pageNumber="32">Internal secondary branch</th>
|
||||
<td id="76E75DECA626004DAF5969C9FDBB8FA5" box="[519,537,1426,1450]" gridcol="1" gridrow="35" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF1469C9FD618FA5" box="[586,707,1426,1450]" gridcol="2" gridrow="35" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF69C9FCCB8FA5" box="[753,873,1426,1450]" gridcol="3" gridrow="35" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE7669C9FC938FA6" box="[808,817,1426,1449]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC969C9FC6D8FA5" box="[919,975,1426,1450]" gridcol="4" gridrow="35" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA269C9FB938FA5" box="[1020,1073,1426,1450]" gridcol="5" gridrow="35" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA269C9FBA78FA6" box="[1020,1029,1426,1449]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E69C9FB2B8FA5" box="[1120,1161,1426,1450]" gridcol="6" gridrow="35" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E769C9FB408FA5" box="[1209,1250,1426,1450]" gridcol="7" gridrow="35" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E769C9FB608FA6" box="[1209,1218,1426,1449]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F69C9FAE58FA5" box="[1297,1351,1426,1450]" gridcol="8" gridrow="35" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA82869C9FA098FA5" box="[1398,1451,1426,1450]" gridcol="9" gridrow="35" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA82869C9FADD8FA6" box="[1398,1407,1426,1449]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F69E8FA098FC3" box="[113,1451,1459,1484]" gridrow="36" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||||
<th id="76E75DECA626004DAD2F69E8FE428FC3" box="[113,480,1459,1484]" gridcol="0" gridrow="36" pageId="31" pageNumber="32">Claw 4 heights</th>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F698EFA098FE2" box="[113,1451,1493,1517]" gridrow="37" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F698EFE428FE2" box="[113,480,1493,1517]" gridcol="0" gridrow="37" pageId="31" pageNumber="32">Anterior base</th>
|
||||
<td id="76E75DECA626004DAF59698EFDBB8FE2" box="[519,537,1493,1517]" gridcol="1" gridrow="37" pageId="31" pageNumber="32">5</td>
|
||||
<td id="76E75DECA626004DAF14698EFD618FE2" box="[586,707,1493,1517]" gridcol="2" gridrow="37" pageId="31" pageNumber="32">3.6–4.3</td>
|
||||
<td id="76E75DECA626004DAFAF698EFCCB8FE2" box="[753,873,1493,1517]" gridcol="3" gridrow="37" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA0698EFCFE8FE2" box="[766,860,1493,1517]" italics="true" pageId="31" pageNumber="32">10.7–13.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC9698EFC6D8FE2" box="[919,975,1493,1517]" gridcol="4" gridrow="37" pageId="31" pageNumber="32">3.9</td>
|
||||
<td id="76E75DECA626004DAEA2698EFB938FE2" box="[1020,1073,1493,1517]" gridcol="5" gridrow="37" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA2698EFB878FE2" box="[1020,1061,1493,1517]" italics="true" pageId="31" pageNumber="32">12.5</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E698EFB2B8FE2" box="[1120,1161,1493,1517]" gridcol="6" gridrow="37" pageId="31" pageNumber="32">0.3</td>
|
||||
<td id="76E75DECA626004DA9E7698EFB408FE2" box="[1209,1250,1493,1517]" gridcol="7" gridrow="37" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E7698EFB748FE2" box="[1209,1238,1493,1517]" italics="true" pageId="31" pageNumber="32">1.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F698EFAE58FE2" box="[1297,1351,1493,1517]" gridcol="8" gridrow="37" pageId="31" pageNumber="32">3.8</td>
|
||||
<td id="76E75DECA626004DA828698EFA098FE2" box="[1398,1451,1493,1517]" gridcol="9" gridrow="37" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA828698EFA3D8FE2" box="[1398,1439,1493,1517]" italics="true" pageId="31" pageNumber="32">12.9</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F69ACFA098C00" box="[113,1451,1527,1551]" gridrow="38" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F69ACFE428C00" box="[113,480,1527,1551]" gridcol="0" gridrow="38" pageId="31" pageNumber="32">Anterior primary branch</th>
|
||||
<td id="76E75DECA626004DAF5969ACFDBB8C00" box="[519,537,1527,1551]" gridcol="1" gridrow="38" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF1469ACFD618C00" box="[586,707,1527,1551]" gridcol="2" gridrow="38" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF69ACFCCB8C00" box="[753,873,1527,1551]" gridcol="3" gridrow="38" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE7669A3FC938C00" box="[808,817,1528,1551]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC969ACFC6D8C00" box="[919,975,1527,1551]" gridcol="4" gridrow="38" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA269ACFB938C00" box="[1020,1073,1527,1551]" gridcol="5" gridrow="38" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA269A3FBA78C00" box="[1020,1029,1528,1551]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E69ACFB2B8C00" box="[1120,1161,1527,1551]" gridcol="6" gridrow="38" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E769ACFB408C00" box="[1209,1250,1527,1551]" gridcol="7" gridrow="38" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E769A3FB608C00" box="[1209,1218,1528,1551]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F69ACFAE58C00" box="[1297,1351,1527,1551]" gridcol="8" gridrow="38" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA82869ACFA098C00" box="[1398,1451,1527,1551]" gridcol="9" gridrow="38" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA82869A3FADD8C00" box="[1398,1407,1528,1551]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6A42FA098C3E" box="[113,1451,1561,1585]" gridrow="39" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6A42FE428C3E" box="[113,480,1561,1585]" gridcol="0" gridrow="39" pageId="31" pageNumber="32">Anterior secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596A42FDBB8C3E" box="[519,537,1561,1585]" gridcol="1" gridrow="39" pageId="31" pageNumber="32">0</td>
|
||||
<td id="76E75DECA626004DAF146A42FD618C3E" box="[586,707,1561,1585]" gridcol="2" gridrow="39" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAFAF6A42FCCB8C3E" box="[753,873,1561,1585]" gridcol="3" gridrow="39" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAE766A42FC938C3F" box="[808,817,1561,1584]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96A42FC6D8C3E" box="[919,975,1561,1585]" gridcol="4" gridrow="39" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DAEA26A42FB938C3E" box="[1020,1073,1561,1585]" gridcol="5" gridrow="39" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26A42FBA78C3F" box="[1020,1029,1561,1584]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6A42FB2B8C3E" box="[1120,1161,1561,1585]" gridcol="6" gridrow="39" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76A42FB408C3E" box="[1209,1250,1561,1585]" gridcol="7" gridrow="39" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76A42FB608C3F" box="[1209,1218,1561,1584]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6A42FAE58C3E" box="[1297,1351,1561,1585]" gridcol="8" gridrow="39" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286A42FA098C3E" box="[1398,1451,1561,1585]" gridcol="9" gridrow="39" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286A42FADD8C3F" box="[1398,1407,1561,1584]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6A60FA098C5C" box="[113,1451,1595,1619]" gridrow="40" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6A60FE428C5C" box="[113,480,1595,1619]" gridcol="0" gridrow="40" pageId="31" pageNumber="32">Posterior base</th>
|
||||
<td id="76E75DECA626004DAF596A60FDBB8C5C" box="[519,537,1595,1619]" gridcol="1" gridrow="40" pageId="31" pageNumber="32">3</td>
|
||||
<td id="76E75DECA626004DAF146A60FD618C5C" box="[586,707,1595,1619]" gridcol="2" gridrow="40" pageId="31" pageNumber="32">3.6–4.6</td>
|
||||
<td id="76E75DECA626004DAFAF6A60FCCB8C5C" box="[753,873,1595,1619]" gridcol="3" gridrow="40" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06A60FCFE8C5C" box="[766,860,1595,1619]" italics="true" pageId="31" pageNumber="32">12.3–14.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96A60FC6D8C5C" box="[919,975,1595,1619]" gridcol="4" gridrow="40" pageId="31" pageNumber="32">4.0</td>
|
||||
<td id="76E75DECA626004DAEA26A60FB938C5C" box="[1020,1073,1595,1619]" gridcol="5" gridrow="40" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26A60FB878C5C" box="[1020,1061,1595,1619]" italics="true" pageId="31" pageNumber="32">13.3</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6A60FB2B8C5C" box="[1120,1161,1595,1619]" gridcol="6" gridrow="40" pageId="31" pageNumber="32">0.5</td>
|
||||
<td id="76E75DECA626004DA9E76A60FB408C5C" box="[1209,1250,1595,1619]" gridcol="7" gridrow="40" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76A60FB748C5C" box="[1209,1238,1595,1619]" italics="true" pageId="31" pageNumber="32">1.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6A60FAE58C5C" box="[1297,1351,1595,1619]" gridcol="8" gridrow="40" pageId="31" pageNumber="32">3.9</td>
|
||||
<td id="76E75DECA626004DA8286A60FA098C5C" box="[1398,1451,1595,1619]" gridcol="9" gridrow="40" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286A60FA3D8C5C" box="[1398,1439,1595,1619]" italics="true" pageId="31" pageNumber="32">13.2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6A06FA098C7A" box="[113,1451,1629,1653]" gridrow="41" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6A06FE428C7A" box="[113,480,1629,1653]" gridcol="0" gridrow="41" pageId="31" pageNumber="32">Posterior primary branch</th>
|
||||
<td id="76E75DECA626004DAF596A06FDBB8C7A" box="[519,537,1629,1653]" gridcol="1" gridrow="41" pageId="31" pageNumber="32">3</td>
|
||||
<td id="76E75DECA626004DAF146A06FD618C7A" box="[586,707,1629,1653]" gridcol="2" gridrow="41" pageId="31" pageNumber="32">8.1–11.3</td>
|
||||
<td id="76E75DECA626004DAFAF6A06FCCB8C7A" box="[753,873,1629,1653]" gridcol="3" gridrow="41" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06A06FCFE8C7A" box="[766,860,1629,1653]" italics="true" pageId="31" pageNumber="32">27.6–35.2</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96A06FC6D8C7A" box="[919,975,1629,1653]" gridcol="4" gridrow="41" pageId="31" pageNumber="32">9.8</td>
|
||||
<td id="76E75DECA626004DAEA26A06FB938C7A" box="[1020,1073,1629,1653]" gridcol="5" gridrow="41" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26A06FB878C7A" box="[1020,1061,1629,1653]" italics="true" pageId="31" pageNumber="32">32.1</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6A06FB2B8C7A" box="[1120,1161,1629,1653]" gridcol="6" gridrow="41" pageId="31" pageNumber="32">1.6</td>
|
||||
<td id="76E75DECA626004DA9E76A06FB408C7A" box="[1209,1250,1629,1653]" gridcol="7" gridrow="41" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76A06FB748C7A" box="[1209,1238,1629,1653]" italics="true" pageId="31" pageNumber="32">4.0</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6A06FAE58C7A" box="[1297,1351,1629,1653]" gridcol="8" gridrow="41" pageId="31" pageNumber="32">9.9</td>
|
||||
<td id="76E75DECA626004DA8286A06FA098C7A" box="[1398,1451,1629,1653]" gridcol="9" gridrow="41" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA8286A06FA3D8C7A" box="[1398,1439,1629,1653]" italics="true" pageId="31" pageNumber="32">33.6</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35363490A626004DAD2F6A25FA098C98" box="[113,1451,1662,1687]" gridrow="42" pageId="31" pageNumber="32">
|
||||
<th id="76E75DECA626004DAD2F6A25FE428C98" box="[113,480,1662,1687]" gridcol="0" gridrow="42" pageId="31" pageNumber="32">Posterior secondary branch</th>
|
||||
<td id="76E75DECA626004DAF596A25FDBB8C98" box="[519,537,1662,1687]" gridcol="1" gridrow="42" pageId="31" pageNumber="32">1</td>
|
||||
<td id="76E75DECA626004DAF146A25FD618C98" box="[586,707,1662,1687]" gridcol="2" gridrow="42" pageId="31" pageNumber="32">4.8–4.8</td>
|
||||
<td id="76E75DECA626004DAFAF6A25FCCB8C98" box="[753,873,1662,1687]" gridcol="3" gridrow="42" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAFA06A25FCFE8C99" box="[766,860,1662,1686]" italics="true" pageId="31" pageNumber="32">16.4–16.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DAEC96A25FC6D8C98" box="[919,975,1662,1687]" gridcol="4" gridrow="42" pageId="31" pageNumber="32">4.8</td>
|
||||
<td id="76E75DECA626004DAEA26A25FB938C98" box="[1020,1073,1662,1687]" gridcol="5" gridrow="42" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAEA26A25FB878C99" box="[1020,1061,1662,1686]" italics="true" pageId="31" pageNumber="32">16.4</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA93E6A25FB2B8C98" box="[1120,1161,1662,1687]" gridcol="6" gridrow="42" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA9E76A25FB408C98" box="[1209,1250,1662,1687]" gridcol="7" gridrow="42" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADA9E76A24FB608C99" box="[1209,1218,1663,1686]" italics="true" pageId="31" pageNumber="32">?</emphasis>
|
||||
</td>
|
||||
<td id="76E75DECA626004DA84F6A25FAE58C98" box="[1297,1351,1662,1687]" gridcol="8" gridrow="42" pageId="31" pageNumber="32">?</td>
|
||||
<td id="76E75DECA626004DA8286A25FA098C98" box="[1398,1451,1662,1687]" gridcol="9" gridrow="42" pageId="31" pageNumber="32">?</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<subSubSection id="C31C6559A626FFADAD2F6A95FD9C8D4B" pageId="31" pageNumber="32" type="discussion">
|
||||
<paragraph id="8BB936D2A626FFADAD2F6A95FD9C8D4B" blockId="31.[113,763,1742,1860]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B972EAC0A626FFADAD2F6A95FF6D8CE9" box="[113,207,1742,1766]" italics="true" pageId="31" pageNumber="32">Remarks:</emphasis>
|
||||
The indentation of claw IV bases is not always as well developed as depicted in the original description (
|
||||
<bibRefCitation id="EF974B23A626FFADAFD66AB6FEF38D2A" author="Kaczmarek L & Michalczyk L" pageId="31" pageNumber="32" pagination="1 - 8" refId="ref36912" refString="Kaczmarek L, Michalczyk L. Platicrista horribilis, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Mongolia. Zootaxa 2003; 132: 1 - 8. https: // doi. org / 10.11646 / zootaxa. 132.1.1" type="journal article" year="2003">Kaczmarek and Michalczyk 2003</bibRefCitation>
|
||||
). A more comprehensive morphometric dataset for the species is presented in
|
||||
<tableCitation id="C6840369A626FFADACB36B77FD9A8D4B" box="[493,568,1836,1860]" captionStart="Table 9" captionStartId="38.[129,184,143,167]" captionTargetBox="[129,1466,217,1687]" captionTargetPageId="38" captionText="Table 9. Measurements (in micrometres) of selected morphological structures of Platicrista horribilis from Montenegro (ME.008) mounted in Hoyer’s medium." pageId="31" pageNumber="32">Table 9</tableCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1018
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1018
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157
data/03/AF/87/03AF87C4A62CFFA7AF1D693DFA3B8CD7.xml
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157
data/03/AF/87/03AF87C4A62CFFA7AF1D693DFA3B8CD7.xml
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|
|
@ -0,0 +1,157 @@
|
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<mods:title id="9E26919AF59858CA3F4D24F57A02A00A">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="62DA48501D4A32E7053F1FF1627BD431">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="1122845EE66339EFA5DEA97D2323966A">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="C42B628EAF098633EEFDD50A546E0238" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F0002A3C631E3FD62ED86A0A4FB45F40">Blagden, Brian</mods:namePart>
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<mods:affiliation id="4D9D952B2650428A70B68DA8543FC4BE">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="2F1FC8AD20CDC051CACB3B8EB123D088">Morek, Witold</mods:namePart>
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<mods:affiliation id="39D93C5D88845556FCEE41DA50B5F169">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="B7D0D67E36C142559FAD57ABCC591F53">Michalczyk, Łukasz</mods:namePart>
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<mods:part id="7FE3DC9B35A40FEC39A700F318312326">
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<mods:date id="3FE0195270D5F16A7CA28DE2CB57AF34">2024</mods:date>
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<mods:number id="F3351E7CECFCB021F7D554AE0C81B27E">2023-11-14</mods:number>
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<mods:classification id="FC6324469EC4EDAEA668C9C8F2AB59BF">journal article</mods:classification>
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<mods:identifier id="6CC7818BEDFCFF40F52C1D672B90EC26" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="8A9858941951DDCC906E89245CF4395E" type="Zenodo-Dep">14243740</mods:identifier>
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<mods:identifier id="2F3A632FA1BA647DF0F9908262B8111A" type="ZooBank">044A402-2A0F-4135-9410-7DE081CB11C4Corresponding</mods:identifier>
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</mods:mods>
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<treatment id="03AF87C4A62CFFA7AF1D693DFA3B8CD7" LSID="urn:lsid:plazi:treatment:03AF87C4A62CFFA7AF1D693DFA3B8CD7" httpUri="http://treatment.plazi.org/id/03AF87C4A62CFFA7AF1D693DFA3B8CD7" lastPageNumber="22" pageId="21" pageNumber="22">
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<subSubSection id="C31C6559A62CFFA7AF1D693DFC438F8F" box="[579,993,1382,1408]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BB936D2A62CFFA7AF1D693DFC438F8F" blockId="21.[137,1437,1382,1753]" box="[579,993,1382,1408]" pageId="21" pageNumber="22">
|
||||
Taxonomic key to the genus
|
||||
<taxonomicName id="4C064D51A62CFFA7AE2A693DFC438F8F" authorityName="Gasiorek & Michalczyk" authorityYear="2020" box="[884,993,1382,1408]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="genus">
|
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<emphasis id="B972EAC0A62CFFA7AE2A693DFC438F8F" box="[884,993,1382,1408]" italics="true" pageId="21" pageNumber="22">Guidettion</emphasis>
|
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</taxonomicName>
|
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A62CFFA7ADD769FCFF5F8FD1" pageId="21" pageNumber="22" type="discussion">
|
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<paragraph id="8BB936D2A62CFFA7ADD769FCFF5F8FD1" blockId="21.[137,1437,1382,1753]" pageId="21" pageNumber="22">
|
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<emphasis id="B972EAC0A62CFFA7ADD769FCFE158FB0" box="[137,439,1447,1471]" italics="true" pageId="21" pageNumber="22">
|
||||
Remark:
|
||||
<taxonomicName id="4C064D51A62CFFA7ADBB69FCFE158FB0" baseAuthorityName="Thulin" baseAuthorityYear="1928" box="[229,439,1447,1471]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="arduifrons">Guidettion arduifrons</taxonomicName>
|
||||
</emphasis>
|
||||
<emphasis id="B972EAC0A62CFFA7ACE369FCFDB58FB0" bold="true" box="[445,535,1447,1471]" pageId="21" pageNumber="22">nom. inq</emphasis>
|
||||
. is not included in the key, owing to the unavailability of traits differentiating it from
|
||||
<taxonomicName id="4C064D51A62CFFA7A8DB69FCFF558FD1" baseAuthorityName="Thulin" baseAuthorityYear="1928" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="prorsirostre">
|
||||
<emphasis id="B972EAC0A62CFFA7A8DB69FCFF558FD1" italics="true" pageId="21" pageNumber="22">G. prorsirostre</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A62CFFA7ADD269BDFA3B8CD7" pageId="21" pageNumber="22" type="key">
|
||||
<key id="FF9761E3A62CFFA7ADD269BDFA3B8CD7" pageId="21" pageNumber="22">
|
||||
<keyStep id="30F22E77A62CFFA7ADD269BDFA3A8C12" pageId="21" pageNumber="22">
|
||||
<paragraph id="8BB936D2A62CFFA7ADD269BDFA3B8FF2" blockId="21.[137,1437,1382,1753]" box="[140,1433,1510,1534]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD269BDFA3B8FF2" box="[140,1433,1510,1534]" pageId="21" pageNumber="22">
|
||||
1. Macroplacoids granular ........................................................................................................................................
|
||||
<taxonomicName id="4C064D51A62CFFA7A9EC69BDFA3B8FF2" ID-CoL="9X7JD" baseAuthorityName="Pilato & Dastych" baseAuthorityYear="1974" box="[1202,1433,1510,1534]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="montigenum">
|
||||
<emphasis id="B972EAC0A62CFFA7A9EC69BDFA3B8FF2" box="[1202,1433,1510,1534]" italics="true" pageId="21" pageNumber="22">Guidettion montigenum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26A5EFA3A8C12" blockId="21.[137,1437,1382,1753]" box="[140,1432,1541,1565]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26A5EFA3A8C12" box="[140,1432,1541,1565]" pageId="21" pageNumber="22">– Macroplacoid bar-like .................................................................................................................................................................................. 2</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A62CFFA7ADD26A7FFA3A8C53" pageId="21" pageNumber="22">
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26A7FFA3A8C33" blockId="21.[137,1437,1382,1753]" box="[140,1432,1572,1596]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26A7FFA3A8C33" box="[140,1432,1572,1596]" pageId="21" pageNumber="22">
|
||||
2. Caudal cuticular granulation present .......................................................................................................................
|
||||
<taxonomicName id="4C064D51A62CFFA7A98C6A7FFA3A8C33" ID-CoL="9X964" baseAuthorityName="Bartos" baseAuthorityYear="1935" box="[1234,1432,1572,1596]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="clavatum">
|
||||
<emphasis id="B972EAC0A62CFFA7A98C6A7FFA3A8C33" box="[1234,1432,1572,1596]" italics="true" pageId="21" pageNumber="22">Guidettion clavatum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26A1FFA3A8C53" blockId="21.[137,1437,1382,1753]" box="[140,1432,1604,1628]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26A1FFA3A8C53" box="[140,1432,1604,1628]" pageId="21" pageNumber="22">– Caudal cuticular granulation absent ......................................................................................................................................................... 3</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A62CFFA7ADD26A38FA3A8C95" pageId="21" pageNumber="22">
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26A38FA3A8C74" blockId="21.[137,1437,1382,1753]" box="[140,1432,1635,1659]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26A38FA3A8C74" box="[140,1432,1635,1659]" pageId="21" pageNumber="22">
|
||||
3. Pharyngeal apophyses absent ..................................................................................................................................
|
||||
<taxonomicName id="4C064D51A62CFFA7A9996A38FA3A8C74" ID-CoL="9X9C7" box="[1223,1432,1635,1659]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="modestum">
|
||||
<emphasis id="B972EAC0A62CFFA7A9996A38FA3A8C74" box="[1223,1432,1635,1659]" italics="true" pageId="21" pageNumber="22">Guidettion modestum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26AD8FA3A8C95" blockId="21.[137,1437,1382,1753]" box="[140,1432,1666,1690]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26AD8FA3A8C95" box="[140,1432,1666,1690]" pageId="21" pageNumber="22">– Pharyngeal apophyses present ................................................................................................................................................................... 4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A62CFFA7ADD26AF9FA3B8CD7" pageId="21" pageNumber="22">
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26AF9FA3B8CB6" blockId="21.[137,1437,1382,1753]" box="[140,1433,1698,1722]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26AF9FA3B8CB6" box="[140,1433,1698,1722]" pageId="21" pageNumber="22">
|
||||
4.
|
||||
<emphasis id="B972EAC0A62CFFA7ADE26AF9FF728CB6" box="[188,208,1698,1721]" italics="true" pageId="21" pageNumber="22">pt</emphasis>
|
||||
of SSIP> 66% ......................................................................................................................................................
|
||||
<taxonomicName id="4C064D51A62CFFA7A9E06AF9FA3B8CB6" ID-CoL="9WV5P" baseAuthorityName="Thulin" baseAuthorityYear="1928" box="[1214,1433,1698,1722]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="prorsirostre">
|
||||
<emphasis id="B972EAC0A62CFFA7A9E06AF9FA3B8CB6" box="[1214,1433,1698,1722]" italics="true" pageId="21" pageNumber="22">Guidettion prorsirostre</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A62CFFA7ADD26A9AFA3B8CD7" blockId="21.[137,1437,1382,1753]" box="[140,1433,1729,1753]" pageId="21" pageNumber="22">
|
||||
<keyLead id="30F795E7A62CFFA7ADD26A9AFA3B8CD7" box="[140,1433,1729,1753]" pageId="21" pageNumber="22">
|
||||
<emphasis id="B972EAC0A62CFFA7ADD26A9AFF728CD6" box="[140,208,1729,1753]" italics="true" pageId="21" pageNumber="22">– pt</emphasis>
|
||||
of SSIP <66% .........................................................................................................................................
|
||||
<emphasis id="B972EAC0A62CFFA7A92A6A9AFA978CD6" box="[1140,1333,1729,1753]" italics="true" pageId="21" pageNumber="22">
|
||||
<taxonomicName id="4C064D51A62CFFA7A92A6A9AFB7E8CD6" ID-CoL="9WSYR" authorityName="Gasiorek & Michalczyk" authorityYear="2020" box="[1140,1244,1729,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="genus">Guidettion</taxonomicName>
|
||||
<taxonomicName id="4C064D51A62CFFA7A9BF6A99FA978CD6" ID-CoL="9X9BY" baseAuthorityName="Pilato, Sabella, D'Urso & Lisi" baseAuthorityYear="2017" box="[1249,1333,1730,1753]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="22" phylum="Tardigrada" rank="species" species="vexatum">vexatum</taxonomicName>
|
||||
</emphasis>
|
||||
<emphasis id="B972EAC0A62CFFA7A8646A9AFA318CD6" bold="true" box="[1338,1427,1729,1753]" pageId="21" pageNumber="22">nom. inq</emphasis>
|
||||
.
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
442
data/03/AF/87/03AF87C4A62EFFABAE1169DEFA3A8CB6.xml
Normal file
442
data/03/AF/87/03AF87C4A62EFFABAE1169DEFA3A8CB6.xml
Normal file
|
|
@ -0,0 +1,442 @@
|
|||
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<mods:title id="CBE692A499942491D8B6CAA758089449">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="F8316510DBC2EAB4ABF97B9F09856E67">Gąsiorek, Piotr</mods:namePart>
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<mods:affiliation id="81EAF4D256F8B0A63403C694C97B740F">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="01757666F18992B1E21615240FF6D620" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BA5D5CCEEFC89DB0632C89F0711AFDB3">Blagden, Brian</mods:namePart>
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<mods:affiliation id="23BE0FF12EB98EFBCCBA2AFD9E2536C5">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:affiliation id="3A5D5AE3C0D69B32E0CC63FF71263868">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="CCB530C16D442505FA2CCFC41EAA6F4C">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="7D181AA2B4E948530342FEEB0ED1004D">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:date id="921032B13E5147F5D9D4AB7FB560C5F0">2024</mods:date>
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<treatment id="03AF87C4A62EFFABAE1169DEFA3A8CB6" LSID="urn:lsid:plazi:treatment:03AF87C4A62EFFABAE1169DEFA3A8CB6" httpUri="http://treatment.plazi.org/id/03AF87C4A62EFFABAE1169DEFA3A8CB6" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
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<subSubSection id="C31C6559A62EFFA5AE1169DEFB028FB0" pageId="23" pageNumber="24" type="nomenclature">
|
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<paragraph id="8BB936D2A62EFFA5AE1169DEFB028FB0" blockId="23.[847,1422,1413,1471]" pageId="23" pageNumber="24">
|
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<emphasis id="B972EAC0A62EFFA5AE1169DEFB028FB0" bold="true" pageId="23" pageNumber="24">
|
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Composition of
|
||||
<taxonomicName id="4C064D51A62EFFA5AEA269DDFBC98FAF" ID-CoL="55TB" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1020,1131,1414,1440]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="genus">
|
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<emphasis id="B972EAC0A62EFFA5AEA269DDFBC98FAF" bold="true" box="[1020,1131,1414,1440]" italics="true" pageId="23" pageNumber="24">Itaquascon</emphasis>
|
||||
</taxonomicName>
|
||||
and its consensus amended diagnosis
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="C31C6559A62EFFABAE746997FEF38E08" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="discussion">
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<paragraph id="8BB936D2A62EFFA5AE746997FA988D53" blockId="23.[809,1461,1484,1978]" pageId="23" pageNumber="24">
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Taxonomy of
|
||||
<taxonomicName id="4C064D51A62EFFA5AEEB6997FBBD8FEC" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[949,1055,1484,1507]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A62EFFA5AEEB6997FBBD8FEC" box="[949,1055,1484,1507]" italics="true" pageId="23" pageNumber="24">Itaquascon</emphasis>
|
||||
</taxonomicName>
|
||||
appears very challenging, mainly owing to the fact that, like
|
||||
<taxonomicName id="4C064D51A62EFFA5A95D69B7FBF98C0C" authorityName="Pilato & Binda" authorityYear="1987" box="[1027,1115,1516,1539]" class="Eutardigrada" family="Hypsibiidae" genus="Parascon" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A62EFFA5A95D69B7FBF98C0C" box="[1027,1115,1516,1539]" italics="true" pageId="23" pageNumber="24">Parascon</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A62EFFA5A9CC69B0FB448C0C" authorityName="Gasiorek & Michalczyk" authorityYear="2020" box="[1170,1254,1515,1539]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A62EFFA5A9CC69B0FB448C0C" box="[1170,1254,1515,1539]" italics="true" pageId="23" pageNumber="24">Raribius</emphasis>
|
||||
</taxonomicName>
|
||||
, the genus is rarely found and, if encountered, usually only in small numbers (i.e. fewer than
|
||||
<specimenCount id="9D00FD5BA62EFFA5AEFD6A71FBE38C4D" box="[931,1089,1578,1602]" count="5" pageId="23" pageNumber="24" type="generic">five specimens</specimenCount>
|
||||
). This rarity means that opportunities for extensive morphometric analysis and the drawing of robust conclusions based on intra- and interspecific variability of phenotypic traits are also limited and infrequent. Luckily, we obtained an enormously high number of individuals of
|
||||
<taxonomicName id="4C064D51A62EFFA5AE746A9CFC0A8CD1" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[810,936,1734,1758]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="species" species="serratulum">
|
||||
<emphasis id="B972EAC0A62EFFA5AE746A9CFC0A8CD1" box="[810,936,1734,1758]" italics="true" pageId="23" pageNumber="24">I. serratulum</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<quantity id="4CFE9B37A62EFFA5AEE96A9DFC4D8CD1" box="[951,1007,1734,1758]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="23" pageNumber="24" unit="in" value="50.0">50 in</quantity>
|
||||
total), which represents the largest number of specimens recorded in the literature so far (Węglarska isolated tens of individuals of
|
||||
<taxonomicName id="4C064D51A62EFFA5A9506B5DFB358D12" authorityName="NEC" baseAuthorityName="Weglarska" baseAuthorityYear="1973" box="[1038,1175,1797,1821]" class="Eutardigrada" family="Hypsibiidae" genus="Raribius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="species" species="pawlowskii">
|
||||
<emphasis id="B972EAC0A62EFFA5A9506B5DFB358D12" box="[1038,1175,1797,1821]" italics="true" pageId="23" pageNumber="24">R. pawlowskii</emphasis>
|
||||
</taxonomicName>
|
||||
from a single moss sample collected in the Polish Tatras, but unfortunately did not publish that interesting finding; B. Węglarska, pers. comm.).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A62EFFABAE1B6B38FE70880E" blockId="23.[809,1461,1484,1978]" lastBlockId="25.[112,764,144,826]" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
|
||||
According to its original description,
|
||||
<taxonomicName id="4C064D51A62EFFA5A9966B3FFAFF8D74" authorityName="Maucci" authorityYear="1973" box="[1224,1373,1891,1915]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="24" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A62EFFA5A9966B3FFAFF8D74" box="[1224,1373,1891,1915]" italics="true" pageId="23" pageNumber="24">I. placophorum</emphasis>
|
||||
</taxonomicName>
|
||||
exhibits a dorsal papilla (
|
||||
<bibRefCitation id="EF974B23A62EFFA5AE836BD9FBC48D95" author="Maucci W" box="[989,1126,1922,1946]" pageId="23" pageNumber="24" pagination="169 - 221" refId="ref37716" refString="Maucci W. Tardigradi muscicoli della Turchia. Memorie del Museo Civico di Storia Naturale di Verona 1973; 20: 169 - 221." type="journal article" year="1973">Maucci 1973</bibRefCitation>
|
||||
), a character never recorded in any other eutardigrade, making it a suspicious taxon. Thus, we examined the
|
||||
<typeStatus id="54BD8870A620FFABADA06CCBFEF88AA7" box="[254,346,144,168]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||||
of this species and found that the ‘papilla’ is simply a conspicuous ventral cloacal fold. In a flattened individual, this ventral fold is visible through the translucent dorsal cuticle even when focusing on the dorsum (hence Maucci’s misinterpretation). It is not the first time that ventral structures have been interpreted as dorsal characters (see the case of the no longer valid ‘
|
||||
<taxonomicName id="4C064D51A620FFABAC436D17FE5D8B6B" box="[285,511,332,356]" class="Eutardigrada" family="Milnesiidae" genus="Milnesium" kingdom="Animalia" order="Apochela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="trispinosum">
|
||||
<emphasis id="B972EAC0A620FFABAC436D17FE5D8B6B" box="[285,511,332,356]" italics="true" pageId="25" pageNumber="26">Milnesium trispinosum</emphasis>
|
||||
</taxonomicName>
|
||||
’ as discussed by
|
||||
<bibRefCitation id="EF974B23A620FFABAFEB6D17FF078B8C" author="Suzuki AC" pageId="25" pageNumber="26" pagination="431 - 3" refId="ref39941" refString="Suzuki AC. Specimens with an artifact appearing as ' three spines' in Milnesium tardigradum var. trispinosa Rahm, 1931 (Tardigrada). Zoological Science 2016; 33: 431 - 3. https: // doi. org / 10.2108 / zs 150184" type="journal article" year="2016">Suzuki 2016</bibRefCitation>
|
||||
). Internal bars on legs II–III of the
|
||||
<taxonomicName id="4C064D51A620FFABAF7A6D37FD198B8C" authorityName="Maucci" authorityYear="1973" box="[548,699,363,387]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A620FFABAF7A6D37FD198B8C" box="[548,699,363,387]" italics="true" pageId="25" pageNumber="26">I. placophorum</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A620FFABAF9D6D30FF3D8BAC" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||||
are thin but clearly visible (
|
||||
<figureCitation id="133D2A57A620FFABACEA6DD0FDA48BAC" box="[436,518,395,419]" captionStart="Figure 17" captionStartId="24.[129,194,1926,1950]" captionTargetBox="[129,1473,1360,1899]" captionTargetId="figure-4@24.[129,1473,1360,1899]" captionTargetPageId="24" captionText="Figure 17. Claws of Itaquascon spp. (PCM, paratypes): A, claws IV of Itaquascon mongolicus Kaczmarek et al., 2002 (note serrated margin of the anterior claw basis); B, claws II of Itaquascon placophorum Maucci, 1973 (black arrowhead indicates internal bar). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243782" httpUri="https://zenodo.org/record/14243782/files/figure.png" pageId="25" pageNumber="26">Fig. 17B</figureCitation>
|
||||
), like those in
|
||||
<taxonomicName id="4C064D51A620FFABAFC66DD0FD568BAC" box="[664,756,395,419]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="biserovi">
|
||||
<emphasis id="B972EAC0A620FFABAFC66DD0FD568BAC" box="[664,756,395,419]" italics="true" pageId="25" pageNumber="26">I. biserovi</emphasis>
|
||||
</taxonomicName>
|
||||
. Furthermore, we independently confirm the findings of
|
||||
<bibRefCitation id="EF974B23A620FFABAFFA6DF1FF568BEE" author="Guidetti R & Bertolani R & Nelson DR" pageId="25" pageNumber="26" pagination="215 - 23" refId="ref36352" refString="Guidetti R, Bertolani R, Nelson DR. Ecological and faunistic studies on tardigrades in leaf litter of beech forests. Zoologischer Anzeiger 1999; 238: 215 - 23." type="journal article" year="1999">
|
||||
Guidetti
|
||||
<emphasis id="B972EAC0A620FFABAD2F6D91FF018BEE" box="[113,163,457,481]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
(1999)
|
||||
</bibRefCitation>
|
||||
, who validated the presence of stylet supports in
|
||||
<taxonomicName id="4C064D51A620FFABAD2F6DB2FE6F880E" authority="(Massa et al. 2021)" baseAuthorityName="Massa" baseAuthorityYear="2021" box="[113,461,489,513]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A620FFABAD2F6DB2FEA7880E" box="[113,261,489,513]" italics="true" pageId="25" pageNumber="26">I. placophorum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A620FFABAC4B6DB2FE63880E" author="Massa E & Guidetti R & Cesari M" box="[277,449,489,513]" pageId="25" pageNumber="26" refId="ref37665" refString="Massa E, Guidetti R, Cesari M et al. Tardigrades of Kristianstads Vattenrike Biosphere Reserve with description of four new species from Sweden. Scientific Reports 2021; 11: 4861. https: // doi. org / 10.1038 / s 41598 - 021 - 83627 - w" type="journal volume" year="2021">
|
||||
Massa
|
||||
<emphasis id="B972EAC0A620FFABAC046DB2FE2B880E" box="[346,393,489,513]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF79665AA621FFAAADDF68BEFA058F15" ID-DOI="http://doi.org/10.5281/zenodo.14243780" ID-Zenodo-Dep="14243780" httpUri="https://zenodo.org/record/14243780/files/figure.png" pageId="24" pageNumber="25" startId="24.[129,194,1253,1277]" targetBox="[129,1473,144,1225]" targetPageId="24" targetType="figure">
|
||||
<paragraph id="8BB936D2A621FFAAADDF68BEFA058F15" blockId="24.[129,1447,1253,1306]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B972EAC0A621FFAAADDF68BEFF478EF2" bold="true" box="[129,229,1253,1277]" pageId="24" pageNumber="25">Figure 16.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A621FFAAAC6168BDFDAB8EF2" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[319,521,1253,1277]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="24" pageNumber="25" phylum="Tardigrada" rank="species" species="serratulum" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A621FFAAAC6168BDFDAB8EF2" box="[319,521,1253,1277]" italics="true" pageId="24" pageNumber="25">Itaquascon serratulum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A621FFAAAF5368BEFDFA8EF2" bold="true" box="[525,600,1253,1277]" pageId="24" pageNumber="25">
|
||||
<taxonomicNameLabel id="A24157BBA621FFAAAF5368BEFDFA8EF2" box="[525,600,1253,1277]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(paratypes): A, claws II (PCM; black arrowhead indicates an internal bar); B, claws IV (PCM); C, claws II (SEM); D, claws IV (SEM). Note aberrant spurs on the basal portions of external and posterior claws. Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF79665AA621FFAAADDF6BDDFAF58DB4" ID-DOI="http://doi.org/10.5281/zenodo.14243782" ID-Zenodo-Dep="14243782" httpUri="https://zenodo.org/record/14243782/files/figure.png" pageId="24" pageNumber="25" startId="24.[129,194,1926,1950]" targetBox="[129,1473,1360,1899]" targetPageId="24" targetType="figure">
|
||||
<paragraph id="8BB936D2A621FFAAADDF6BDDFAF58DB4" blockId="24.[129,1472,1926,1979]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B972EAC0A621FFAAADDF6BDDFF478D90" bold="true" box="[129,229,1926,1951]" pageId="24" pageNumber="25">Figure 17.</emphasis>
|
||||
Claws of
|
||||
<taxonomicName id="4C064D51A621FFAAAC616BDCFE6F8D91" box="[319,461,1927,1950]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="24" pageNumber="25" phylum="Tardigrada" rank="species" species="undetermined">
|
||||
<emphasis id="B972EAC0A621FFAAAC616BDCFE038D91" box="[319,417,1927,1950]" italics="true" pageId="24" pageNumber="25">Itaquascon</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
(PCM, paratypes): A, claws IV of
|
||||
<taxonomicName id="4C064D51A621FFAAAE4F6BDCFB118D90" authority="Kaczmarek et al., 2002" authorityName="Kaczmarek" authorityYear="2002" box="[785,1203,1927,1951]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="24" pageNumber="25" phylum="Tardigrada" rank="species" species="mongolicus">
|
||||
<emphasis id="B972EAC0A621FFAAAE4F6BDCFC798D90" box="[785,987,1927,1951]" italics="true" pageId="24" pageNumber="25">Itaquascon mongolicus</emphasis>
|
||||
<bibRefCitation id="EF974B23A621FFAAAEBE6BDCFB118D90" author="Kaczmarek L & Michalczyk L & Weglarska B" box="[992,1203,1927,1951]" pageId="24" pageNumber="25" pagination="1 - 3" refId="ref36996" refString="Kaczmarek L, Michalczyk L, Weglarska B. Itaquascon mongolicus, a new species of Eutardigrada from Mongolia. Genus 2002; 13: 1 - 3." type="journal article" year="2002">
|
||||
Kaczmarek
|
||||
<emphasis id="B972EAC0A621FFAAA9116BDCFBD88D90" box="[1103,1146,1927,1951]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||||
, 2002
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(note serrated margin of the anterior claw basis); B, claws II of
|
||||
<taxonomicName id="4C064D51A621FFAAAC9E6BF8FC818DB4" authority="Maucci, 1973" authorityName="Maucci" authorityYear="1973" box="[448,803,1955,1979]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="24" pageNumber="25" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A621FFAAAC9E6BF8FD3C8DB4" box="[448,670,1955,1979]" italics="true" pageId="24" pageNumber="25">Itaquascon placophorum</emphasis>
|
||||
<bibRefCitation id="EF974B23A621FFAAAFFD6BF8FC818DB4" author="Maucci W" box="[675,803,1955,1979]" pageId="24" pageNumber="25" pagination="169 - 221" refId="ref37716" refString="Maucci W. Tardigradi muscicoli della Turchia. Memorie del Museo Civico di Storia Naturale di Verona 1973; 20: 169 - 221." type="journal article" year="1973">Maucci, 1973</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(black arrowhead indicates internal bar). Scale bars: 10 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A620FFABADD36E53FB5F8B8C" blockId="25.[112,764,144,826]" lastBlockId="25.[809,1461,144,826]" pageId="25" pageNumber="26">
|
||||
There also seems to be a misunderstanding about the pattern of the pharyngeal tube annulation in the genus
|
||||
<taxonomicName id="4C064D51A620FFABAFD26E73FD578830" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[652,757,552,575]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A620FFABAFD26E73FD578830" box="[652,757,552,575]" italics="true" pageId="25" pageNumber="26">Itaquascon</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation id="EF974B23A620FFABAD2F6E1CFE6F8850" author="Gasiorek P & Michalczyk L" box="[113,461,583,607]" pageId="25" pageNumber="26" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
|
||||
, based on high-quality SEM images, described the annulation in
|
||||
<taxonomicName id="4C064D51A620FFABAF526E3CFD308871" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[524,658,614,638]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="serratulum" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A620FFABAF526E3CFD308871" box="[524,658,614,638]" italics="true" pageId="25" pageNumber="26">I. serratulum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A620FFABAFFE6E3DFD598871" bold="true" box="[672,763,614,638]" pageId="25" pageNumber="26">
|
||||
<taxonomicNameLabel id="A24157BBA620FFABAFFE6E3DFD598871" box="[672,763,614,638]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<taxonomicName id="4C064D51A620FFABAD226EDDFEB28892" box="[124,272,646,669]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="undetermined">
|
||||
<emphasis id="B972EAC0A620FFABAD226EDDFF448892" box="[124,230,646,669]" italics="true" pageId="25" pageNumber="26">Itaquascon</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
then) as ‘complex, with annuli forking and merging irregularly’ (figs 4i–j therein). In contrast,
|
||||
<bibRefCitation id="EF974B23A620FFABAFDB6EFEFF1E88D3" author="Massa E & Guidetti R & Cesari M" pageId="25" pageNumber="26" refId="ref37665" refString="Massa E, Guidetti R, Cesari M et al. Tardigrades of Kristianstads Vattenrike Biosphere Reserve with description of four new species from Sweden. Scientific Reports 2021; 11: 4861. https: // doi. org / 10.1038 / s 41598 - 021 - 83627 - w" type="journal volume" year="2021">
|
||||
Massa
|
||||
<emphasis id="B972EAC0A620FFABAF946EFEFD5988B2" box="[714,763,677,701]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
(2021)
|
||||
</bibRefCitation>
|
||||
, when interpreting the somewhat blurred SEM images of the buccal apparatuses of
|
||||
<taxonomicName id="4C064D51A620FFABACC96EBFFF078914" authority="Massa et al., 2021" authorityName="Massa" authorityYear="2021" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="magnussoni">
|
||||
<emphasis id="B972EAC0A620FFABACC96EBFFDD888F4" box="[407,634,740,763]" italics="true" pageId="25" pageNumber="26">Itaquascon magnussoni</emphasis>
|
||||
<bibRefCitation id="EF974B23A620FFABAF216EBFFF078914" author="Massa E & Guidetti R & Cesari M" pageId="25" pageNumber="26" refId="ref37665" refString="Massa E, Guidetti R, Cesari M et al. Tardigrades of Kristianstads Vattenrike Biosphere Reserve with description of four new species from Sweden. Scientific Reports 2021; 11: 4861. https: // doi. org / 10.1038 / s 41598 - 021 - 83627 - w" type="journal volume" year="2021">
|
||||
Massa
|
||||
<emphasis id="B972EAC0A620FFABAF9B6EBFFD5488F4" box="[709,758,739,763]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
, 2021
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A620FFABAD896F58FECE8914" authorityName="Maucci" authorityYear="1973" box="[215,364,771,795]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A620FFABAD896F58FECE8914" box="[215,364,771,795]" italics="true" pageId="25" pageNumber="26">I. placophorum</emphasis>
|
||||
</taxonomicName>
|
||||
(figs 5e–f and SM6 therein), described the annulation in these species as an ‘alternating hexagonal wire mesh’ and concluded that it is a different morphotype from that shown by
|
||||
<bibRefCitation id="EF974B23A620FFABAECE6CF4FB428AC8" author="Gasiorek P & Michalczyk L" box="[912,1248,175,199]" pageId="25" pageNumber="26" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
|
||||
, thus suggesting that there is intrageneric variability. However, a careful comparison of the 2020 and 2021 microphotographs shows that the pattern in all analysed species is similar. In other words, the differences seem to be semantic, rather than morphological. Nevertheless, better-quality images of additional
|
||||
<taxonomicName id="4C064D51A620FFABA9D66D16FAE38B6B" box="[1160,1345,332,356]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B972EAC0A620FFABA9D66D16FB508B6B" box="[1160,1266,333,356]" italics="true" pageId="25" pageNumber="26">Itaquascon</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
are needed to allow more sound conclusions to be drawn.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABAE1B6DD0FC478935" blockId="25.[809,1461,144,826]" pageId="25" pageNumber="26">
|
||||
The other difference in our interpretations pertains to the pharyngeal structures. Unlike
|
||||
<bibRefCitation id="EF974B23A620FFABA9C76DF1FAC78BCD" author="Massa E & Guidetti R & Cesari M" box="[1177,1381,426,450]" pageId="25" pageNumber="26" refId="ref37665" refString="Massa E, Guidetti R, Cesari M et al. Tardigrades of Kristianstads Vattenrike Biosphere Reserve with description of four new species from Sweden. Scientific Reports 2021; 11: 4861. https: // doi. org / 10.1038 / s 41598 - 021 - 83627 - w" type="journal volume" year="2021">
|
||||
Massa
|
||||
<emphasis id="B972EAC0A620FFABA9BE6DF0FAB08BCD" box="[1248,1298,426,450]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
(2021)
|
||||
</bibRefCitation>
|
||||
, we do not regard the extremely thin cuticular ridges present in the pharynx of most
|
||||
<taxonomicName id="4C064D51A620FFABAEBE6DB2FB03880E" box="[992,1185,489,513]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B972EAC0A620FFABAEBE6DB2FBED880F" box="[992,1103,489,512]" italics="true" pageId="25" pageNumber="26">Itaquascon</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
as typical macroplacoids. Consequently, we merge the amended diagnoses of the genus (
|
||||
<bibRefCitation id="EF974B23A620FFABAE6B6E7CFB298830" author="Gasiorek P & Michalczyk L" box="[821,1163,551,575]" pageId="25" pageNumber="26" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk 2020</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A620FFABA9C56E73FAFF8830" author="Massa E & Guidetti R & Cesari M" box="[1179,1373,551,575]" pageId="25" pageNumber="26" refId="ref37665" refString="Massa E, Guidetti R, Cesari M et al. Tardigrades of Kristianstads Vattenrike Biosphere Reserve with description of four new species from Sweden. Scientific Reports 2021; 11: 4861. https: // doi. org / 10.1038 / s 41598 - 021 - 83627 - w" type="journal volume" year="2021">
|
||||
Massa
|
||||
<emphasis id="B972EAC0A620FFABA9B66E73FABF8830" box="[1256,1309,551,575]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
) into a single, unified diagnosis that addresses these disparate points of view: ‘Dorsoposterior apodeme (drop-like thickening) between the buccal tube and the pharyngeal tube absent. S-shaped (sinusoidal) stylet supports. Pharynx elongated, oval or pyriform in shape, with placoids reduced to thin cuticular ridges or absent. Claws of the
|
||||
<taxonomicName id="4C064D51A620FFABA9EB6EB8FAB888F4" authorityName="Ehrenberg" authorityYear="1848" box="[1205,1306,739,763]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A620FFABA9EB6EB8FAB888F4" box="[1205,1306,739,763]" italics="true" pageId="25" pageNumber="26">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A620FFABA87E6EBFFAF188F3" box="[1312,1363,740,764]" pageId="25" pageNumber="26">type</typeStatus>
|
||||
; external claws with primary branches markedly longer than the secondary branches’.
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABAF1C6F34FEF38E08" blockId="25.[137,1435,879,1721]" pageId="25" pageNumber="26">
|
||||
Taxonomic key to the genus
|
||||
<emphasis id="B972EAC0A620FFABAE2D6F2BFE0289C6" italics="true" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C064D51A620FFABAE2D6F2BFC408986" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[883,994,880,905]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="genus">Itaquascon</taxonomicName>
|
||||
Remark:
|
||||
<taxonomicName id="4C064D51A620FFABADB96FEAFE0289C6" baseAuthorityName="Mihelcic" baseAuthorityYear="1971" box="[231,416,945,969]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="enckelli">Itaquascon enckelli</taxonomicName>
|
||||
</emphasis>
|
||||
(Mihelčič, 1971/72) and
|
||||
<taxonomicName id="4C064D51A620FFABAFEB6FEAFB8B89C6" authority="(Mihelcic, 1971)" baseAuthorityName="Mihelcic" baseAuthorityYear="1971" box="[693,1065,945,969]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="simplex">
|
||||
<emphasis id="B972EAC0A620FFABAFEB6FEAFCD389C6" box="[693,881,945,969]" italics="true" pageId="25" pageNumber="26">Itaquascon simplex</emphasis>
|
||||
(Mihelčič, 1971)
|
||||
</taxonomicName>
|
||||
have outdated descriptions that do not permit comparison with, or exclusion of other species. Thus, both are designated as
|
||||
<emphasis id="B972EAC0A620FFABA94E6F8BFB0C89E7" bold="true" box="[1040,1198,976,1000]" italics="true" pageId="25" pageNumber="26">nomen dubium</emphasis>
|
||||
and, consequently, not included in the key.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A620FFABADD26854FA3A8CB6" pageId="25" pageNumber="26" type="key">
|
||||
<key id="FF9761E3A620FFABADD26854FA3A8CB6" pageId="25" pageNumber="26">
|
||||
<keyStep id="30F22E77A620FFABADD26854FA3A8E49" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26854FA3A8E28" blockId="25.[137,1435,879,1721]" box="[140,1432,1038,1063]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26854FA3A8E28" box="[140,1432,1038,1063]" pageId="25" pageNumber="26">
|
||||
1. The buccal tube longer than half of the total length of the buccopharyngeal tube (
|
||||
<emphasis id="B972EAC0A620FFABAEAB6854FBB68E28" box="[1013,1044,1039,1063]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
> 50%) .................................................. 2
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD26875FA3A8E49" blockId="25.[137,1435,879,1721]" box="[140,1432,1070,1094]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26875FA3A8E49" box="[140,1432,1070,1094]" pageId="25" pageNumber="26">
|
||||
– The buccal tube shorter than half of the total length of the buccopharyngeal tube (
|
||||
<emphasis id="B972EAC0A620FFABAEA36875FBBE8E49" box="[1021,1052,1070,1094]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
<50%) ................................................ 3
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26816FA3A8E8B" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26816FA3A8E6A" blockId="25.[137,1435,879,1721]" box="[140,1432,1101,1125]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26816FA3A8E6A" box="[140,1432,1101,1125]" pageId="25" pageNumber="26">
|
||||
2. Claws with distinct pseudolunulae and indented bases, cuticular bars on legs I–III absent ....................
|
||||
<taxonomicName id="4C064D51A620FFABA9E06815FA3A8E6A" ID-CoL="3QDS5" box="[1214,1432,1101,1125]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="mongolicus">
|
||||
<emphasis id="B972EAC0A620FFABA9E06815FA3A8E6A" box="[1214,1432,1101,1125]" italics="true" pageId="25" pageNumber="26">Itaquascon mongolicus</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD26836FA3A8E8B" blockId="25.[137,1435,879,1721]" box="[140,1432,1133,1157]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26836FA3A8E8B" box="[140,1432,1133,1157]" pageId="25" pageNumber="26">
|
||||
– Claws with indistinct pseudolunulae and smooth bases, cuticular bars on legs I–III present ......................
|
||||
<taxonomicName id="4C064D51A620FFABA9846836FA3A8E8B" ID-CoL="9WQDR" authorityName="Pilato & Lisi" authorityYear="2009" box="[1242,1432,1133,1156]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="pisoniae">
|
||||
<emphasis id="B972EAC0A620FFABA9846836FA3A8E8B" box="[1242,1432,1133,1156]" italics="true" pageId="25" pageNumber="26">Itaquascon pisoniae</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD268D7FA3A8ECC" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD268D7FA3A8EAB" blockId="25.[137,1435,879,1721]" box="[140,1432,1164,1188]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD268D7FA3A8EAB" box="[140,1432,1164,1188]" pageId="25" pageNumber="26">3. Cuticular internal bars on legs I–III present ............................................................................................................................................ 4</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD268F7FA3A8ECC" blockId="25.[137,1435,879,1721]" box="[140,1432,1195,1219]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD268F7FA3A8ECC" box="[140,1432,1195,1219]" pageId="25" pageNumber="26">– Cuticular internal bars on legs I–III absent ............................................................................................................................................. 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26890FA3B8F0D" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26890FA3B8EEC" blockId="25.[137,1435,879,1721]" box="[140,1433,1227,1251]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26890FA3B8EEC" box="[140,1433,1227,1251]" pageId="25" pageNumber="26">
|
||||
4. Claws IV with indented bases ................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9376890FAE08EEC" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" box="[1129,1346,1227,1251]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="serratulum" status="sp. nov.">
|
||||
<emphasis id="B972EAC0A620FFABA9376890FAE08EEC" box="[1129,1346,1227,1251]" italics="true" pageId="25" pageNumber="26">Itaquascon serratulum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A620FFABA8196890FA3B8EEC" bold="true" box="[1351,1433,1227,1251]" pageId="25" pageNumber="26">
|
||||
<taxonomicNameLabel id="A24157BBA620FFABA8196890FA3B8EEC" box="[1351,1433,1227,1251]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD268B1FA3B8F0D" blockId="25.[137,1435,879,1721]" box="[140,1433,1258,1282]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD268B1FA3B8F0D" box="[140,1433,1258,1282]" pageId="25" pageNumber="26">
|
||||
– Claws IV with smooth bases ........................................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA98168B0FA3B8F0D" ID-CoL="3QDRY" box="[1247,1433,1258,1282]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="biserovi">
|
||||
<emphasis id="B972EAC0A620FFABA98168B0FA3B8F0D" box="[1247,1433,1258,1282]" italics="true" pageId="25" pageNumber="26">Itaquascon biserovi</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26952FA3A8F4F" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26952FA3A8F2E" blockId="25.[137,1435,879,1721]" box="[140,1432,1289,1313]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26952FA3A8F2E" box="[140,1432,1289,1313]" pageId="25" pageNumber="26">5. Pharynx with reduced macroplacoids (cuticular ridges) ...................................................................................................................... 6</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD26972FA3A8F4F" blockId="25.[137,1435,879,1721]" box="[140,1432,1321,1345]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26972FA3A8F4F" box="[140,1432,1321,1345]" pageId="25" pageNumber="26">– Pharynx without any cuticular structures ................................................................................................................................................ 7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26913FA3A8FB1" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26913FA3A8F70" blockId="25.[137,1435,879,1721]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26913FA3A8F70" pageId="25" pageNumber="26">
|
||||
6. The ratio between the length of the buccal tube and the total length of the buccopharyngeal tube
|
||||
<emphasis id="B972EAC0A620FFABA9CA6913FB118F6F" box="[1172,1203,1352,1376]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
<20% .......................... ..................................................................................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9F66933FA3A8F70" ID-CoL="3QDS7" authorityName="Maucci" authorityYear="1973" box="[1192,1432,1383,1407]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="placophorum">
|
||||
<emphasis id="B972EAC0A620FFABA9F66933FA3A8F70" box="[1192,1432,1383,1407]" italics="true" pageId="25" pageNumber="26">Itaquascon placophorum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD269DCFA3A8FB1" blockId="25.[137,1435,879,1721]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD269DCFA3A8FB1" pageId="25" pageNumber="26">
|
||||
– The ratio between the length of the buccal tube and the total length of the buccopharyngeal tube
|
||||
<emphasis id="B972EAC0A620FFABA9CA69DCFB118F90" box="[1172,1203,1415,1439]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
> 45% .......................... .....................................................................................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9E969FCFA3A8FB1" ID-CoL="9X6R2" box="[1207,1432,1447,1470]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="magnussoni">
|
||||
<emphasis id="B972EAC0A620FFABA9E969FCFA3A8FB1" box="[1207,1432,1447,1470]" italics="true" pageId="25" pageNumber="26">Itaquascon magnussoni</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD2699DFA3A8FF2" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD2699DFA3A8FD2" blockId="25.[137,1435,879,1721]" box="[140,1432,1477,1502]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD2699DFA3A8FD2" box="[140,1432,1477,1502]" pageId="25" pageNumber="26">
|
||||
7. The percentage ratio between anterior and posterior claw
|
||||
<emphasis id="B972EAC0A620FFABAE5A699DFCBF8FD2" box="[772,797,1478,1501]" italics="true" pageId="25" pageNumber="26">ap</emphasis>
|
||||
<58% ................................................................................................ 8
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD269BEFA3A8FF2" blockId="25.[137,1435,879,1721]" box="[140,1432,1508,1533]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD269BEFA3A8FF2" box="[140,1432,1508,1533]" pageId="25" pageNumber="26">
|
||||
– The percentage ratio between anterior and posterior claw
|
||||
<emphasis id="B972EAC0A620FFABAE5A69BEFCBF8FF3" box="[772,797,1509,1532]" italics="true" pageId="25" pageNumber="26">ap</emphasis>
|
||||
> 58% ................................................................................................ 9
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26A5FFA3B8C34" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26A5FFA3B8C13" blockId="25.[137,1435,879,1721]" box="[140,1433,1540,1564]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26A5FFA3B8C13" box="[140,1433,1540,1564]" pageId="25" pageNumber="26">
|
||||
8. The percentage ratio between anterior and posterior claw
|
||||
<emphasis id="B972EAC0A620FFABAE5A6A5EFCBF8C13" box="[772,797,1541,1564]" italics="true" pageId="25" pageNumber="26">ap</emphasis>
|
||||
= 54–56% ....................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9E16A5EFA3B8C13" ID-CoL="3QDSC" authorityName="de Barros" authorityYear="1939" box="[1215,1433,1540,1564]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="umbellinae">
|
||||
<emphasis id="B972EAC0A620FFABA9E16A5EFA3B8C13" box="[1215,1433,1540,1564]" italics="true" pageId="25" pageNumber="26">Itaquascon umbellinae</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD26A7FFA3B8C34" blockId="25.[137,1435,879,1721]" box="[140,1433,1571,1596]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26A7FFA3B8C34" box="[140,1433,1571,1596]" pageId="25" pageNumber="26">
|
||||
– The percentage ratio between anterior and posterior claw
|
||||
<emphasis id="B972EAC0A620FFABAE5A6A7FFCBF8C34" box="[772,797,1572,1595]" italics="true" pageId="25" pageNumber="26">ap</emphasis>
|
||||
<51% ....................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9CB6A7FFA3B8C34" ID-CoL="3QDS2" authorityName="Pilato, Binda & Claxton" authorityYear="2002" box="[1173,1433,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="cambewarrense">
|
||||
<emphasis id="B972EAC0A620FFABA9CB6A7FFA3B8C34" box="[1173,1433,1571,1595]" italics="true" pageId="25" pageNumber="26">Itaquascon cambewarrense</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A620FFABADD26A18FA3A8CB6" pageId="25" pageNumber="26">
|
||||
<paragraph id="8BB936D2A620FFABADD26A18FA3A8C75" blockId="25.[137,1435,879,1721]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26A18FA3A8C75" pageId="25" pageNumber="26">
|
||||
9. The ratio between the length of the buccal tube and the total length of the buccopharyngeal tube
|
||||
<emphasis id="B972EAC0A620FFABA9F56A18FB688C54" box="[1195,1226,1603,1627]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
<45%, the external buccal tube width
|
||||
<emphasis id="B972EAC0A620FFABAC266A38FE2E8C75" box="[376,396,1635,1658]" italics="true" pageId="25" pageNumber="26">pt</emphasis>
|
||||
> 15% ..................................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9E96A38FA3A8C75" ID-CoL="3QDSD" authorityName="Pilato, Binda & Claxton" authorityYear="2002" box="[1207,1432,1634,1658]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="unguiculum">
|
||||
<emphasis id="B972EAC0A620FFABA9E96A38FA3A8C75" box="[1207,1432,1634,1658]" italics="true" pageId="25" pageNumber="26">Itaquascon unguiculum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A620FFABADD26AD9FA3A8CB6" blockId="25.[137,1435,879,1721]" pageId="25" pageNumber="26">
|
||||
<keyLead id="30F795E7A620FFABADD26AD9FA3A8CB6" pageId="25" pageNumber="26">
|
||||
– The ratio between the length of the buccal tube and the total length of the buccopharyngeal tube
|
||||
<emphasis id="B972EAC0A620FFABA9F56ADAFB688C96" box="[1195,1226,1665,1689]" italics="true" pageId="25" pageNumber="26">pbf</emphasis>
|
||||
> 45%, the external buccal tube width
|
||||
<emphasis id="B972EAC0A620FFABAC266AFAFE2E8CB7" box="[376,396,1697,1720]" italics="true" pageId="25" pageNumber="26">pt</emphasis>
|
||||
<15% ...........................................................................................................................................
|
||||
<taxonomicName id="4C064D51A620FFABA9B46AFAFA3A8CB6" ID-CoL="9WQDQ" authorityName="Lisi, Londoño & Quiroga" authorityYear="2014" box="[1258,1432,1697,1721]" class="Eutardigrada" family="Hypsibiidae" genus="Itaquascon" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="26" phylum="Tardigrada" rank="species" species="pilatoi">
|
||||
<emphasis id="B972EAC0A620FFABA9B46AFAFA3A8CB6" box="[1258,1432,1697,1721]" italics="true" pageId="25" pageNumber="26">Itaquascon pilatoi</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
170
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Normal file
170
data/03/AF/87/03AF87C4A632FFB9AC7168DDFB248FA7.xml
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|
|
@ -0,0 +1,170 @@
|
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<mods:title id="FFCB7A20F867AE1F68DB369FC6DEBD04">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:affiliation id="F28A23AD261E77AB68B74F8BA32A9DE6">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="8E1612D7F8947C9D07223CA9335515A9" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0C060EE0866D5ABCB857565341420B56">Blagden, Brian</mods:namePart>
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<mods:affiliation id="B5AABD4C6C64BA4C69C289587F387A80">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:affiliation id="D0B43F421A0165DEA4FE187789B7EDCB">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="E4732F773A3A5EDD8E18BD6538FD1765">Michalczyk, Łukasz</mods:namePart>
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<treatment id="03AF87C4A632FFB9AC7168DDFB248FA7" LSID="urn:lsid:plazi:treatment:03AF87C4A632FFB9AC7168DDFB248FA7" httpUri="http://treatment.plazi.org/id/03AF87C4A632FFB9AC7168DDFB248FA7" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="C31C6559A632FFB9AC7168DDFD9E8EAE" box="[303,572,1158,1185]" pageId="11" pageNumber="12" type="nomenclature">
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<emphasis id="B972EAC0A632FFB9AC7168DDFD9E8EAE" bold="true" box="[303,572,1158,1185]" pageId="11" pageNumber="12">
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Composition of
|
||||
<taxonomicName id="4C064D51A632FFB9AC8268DDFD9E8EAE" authorityName="Pilato" authorityYear="1987" box="[476,572,1158,1185]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AC8268DDFD9E8EAE" bold="true" box="[476,572,1158,1185]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A632FFB9AD2F68F6FB248FA7" pageId="11" pageNumber="12" type="discussion">
|
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<paragraph id="8BB936D2A632FFB9AD2F68F6FB248FA7" blockId="11.[112,764,1158,1441]" lastBlockId="11.[810,1460,985,1448]" pageId="11" pageNumber="12">
|
||||
The genus
|
||||
<taxonomicName id="4C064D51A632FFB9ADB868F6FEE68ECA" authorityName="Pilato" authorityYear="1987" box="[230,324,1197,1221]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9ADB868F6FEE68ECA" box="[230,324,1197,1221]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
was inferred as polyphyletic by
|
||||
<bibRefCitation id="EF974B23A632FFB9AFC568F6FF578EEA" author="Bertolani R & Guidetti R & Marchioro T" pageId="11" pageNumber="12" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A632FFB9AD2F6896FF018EEB" box="[113,163,1228,1252]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||||
(2014)
|
||||
</bibRefCitation>
|
||||
.
|
||||
<bibRefCitation id="EF974B23A632FFB9AC5F6897FDFD8EEA" author="Gasiorek P & Michalczyk L" box="[257,607,1228,1253]" pageId="11" pageNumber="12" pagination="499 - 515" refId="ref36073" refString="Gasiorek P, Michalczyk L. Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta 2020; 49: 499 - 515. https: // doi. org / 10.1111 / zsc. 12424" type="journal article" year="2020">Gąsiorek and Michalczyk (2020)</bibRefCitation>
|
||||
confirmed the non-monophyly of
|
||||
<taxonomicName id="4C064D51A632FFB9AC1D68B7FE038F0B" authorityName="Pilato" authorityYear="1987" box="[323,417,1260,1284]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AC1D68B7FE038F0B" box="[323,417,1260,1284]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
and established
|
||||
<taxonomicName id="4C064D51A632FFB9AF0468B7FD668F0B" authorityName="Gasiorek & Michalczyk" authorityYear="2020" box="[602,708,1260,1284]" class="Eutardigrada" family="Hypsibiidae" genus="Guidettion" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="22" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AF0468B7FD668F0B" box="[602,708,1260,1284]" italics="true" pageId="11" pageNumber="12">Guidettion</emphasis>
|
||||
</taxonomicName>
|
||||
(the latter having only three bar-like macroplacoids in the pharynx). However, the genus is still heterogeneous in terms of the number of macroplacoids: the majority of species exhibit three placoids, whereas there is a small group of species with only two macroplacoids, i.e.
|
||||
<taxonomicName id="4C064D51A632FFB9AC6369D2FE0B8FAF" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[317,425,1416,1440]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A632FFB9AC6369D2FE0B8FAF" box="[317,425,1416,1440]" italics="true" pageId="11" pageNumber="12">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A632FFB9ACE969D2FDD58FAF" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[439,631,1416,1440]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A632FFB9ACE969D2FDD58FAF" box="[439,631,1416,1440]" italics="true" pageId="11" pageNumber="12">A. diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A632FFB9AF2169D2FC0189FE" bold="true" box="[639,931,985,1441]" pageId="11" pageNumber="12">
|
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<taxonomicNameLabel id="A24157BBA632FFB9AF2169D2FC0189FE" box="[639,931,985,1441]" pageId="11" pageNumber="12">comb. nov. et stat. rev.</taxonomicNameLabel>
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</emphasis>
|
||||
, and
|
||||
<taxonomicName id="4C064D51A632FFB9AE856F82FB8089FE" box="[987,1058,985,1009]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gani">
|
||||
<emphasis id="B972EAC0A632FFB9AE856F82FB8089FE" box="[987,1058,985,1009]" italics="true" pageId="11" pageNumber="12">A. gani</emphasis>
|
||||
</taxonomicName>
|
||||
. The adoption of molecular phylogenetic methods in tardigrade research has shown that the number and shape of placoids in the pharynx is indicative of common ancestry (e.g.
|
||||
<bibRefCitation id="EF974B23A632FFB9AEE3686CFB2E8E40" author="Gasiorek P & Stec D & Morek W" box="[957,1164,1079,1103]" pageId="11" pageNumber="12" pagination="78 - 132" refId="ref36279" refString="Gasiorek P, Stec D, Morek W et al. Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology 2019; 88: 78 - 132. https: // doi. org / 10.1163 / 18759866 - 20191350" type="journal article" year="2019">
|
||||
Gąsiorek
|
||||
<emphasis id="B972EAC0A632FFB9A97E6863FBF38E40" box="[1056,1105,1079,1103]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||||
2019
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A632FFB9A9C6686CFAC08E40" author="Guidetti R & Giovannini I & Del Papa V" box="[1176,1378,1079,1103]" pageId="11" pageNumber="12" pagination="1099 - 117" refId="ref36442" refString="Guidetti R, Giovannini I, Del Papa V et al. Phylogeny of the asexual lineage Murrayidae (Macrobiotoidea, Eutardigrada) with the description of Paramurrayon gen. nov. and Paramurrayon meieri sp. nov. Invertebrate Systematics 2022; 36: 1099 - 117. https: // doi. org / 10.1071 / IS 22031" type="journal article" year="2022">
|
||||
Guidetti
|
||||
<emphasis id="B972EAC0A632FFB9A9A86863FA838E40" box="[1270,1313,1079,1103]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
|
||||
. 2022
|
||||
</bibRefCitation>
|
||||
). Thus, it is plausible that
|
||||
<taxonomicName id="4C064D51A632FFB9AEA6680CFBC58E61" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1016,1127,1110,1134]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A632FFB9AEA6680CFBC58E61" box="[1016,1127,1110,1134]" italics="true" pageId="11" pageNumber="12">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A632FFB9A9FC680CFAC68E61" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[1186,1380,1110,1134]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A632FFB9A9FC680CFAC68E61" box="[1186,1380,1110,1134]" italics="true" pageId="11" pageNumber="12">A. diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A632FFB9A831680CFC7D8E81" bold="true" pageId="11" pageNumber="12">
|
||||
<taxonomicNameLabel id="A24157BBA632FFB9A831680CFC7D8E81" pageId="11" pageNumber="12">comb. nov. et stat. rev.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
are closely related and represent a separate evolutionary lineage. If reciprocal monophyly is confirmed molecularly, they should be erected as a new genus. However,
|
||||
<taxonomicName id="4C064D51A632FFB9AE72688FFCD78EE4" box="[812,885,1236,1259]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gani">
|
||||
<emphasis id="B972EAC0A632FFB9AE72688FFCD78EE4" box="[812,885,1236,1259]" italics="true" pageId="11" pageNumber="12">A. gani</emphasis>
|
||||
</taxonomicName>
|
||||
is designated as
|
||||
<emphasis id="B972EAC0A632FFB9A974688FFAA78EE4" bold="true" box="[1066,1285,1235,1259]" italics="true" pageId="11" pageNumber="12">nomen inquirendum</emphasis>
|
||||
, because having two rod-like macroplacoids arranged in parentheses (i.e. as in the members of the genus
|
||||
<taxonomicName id="4C064D51A632FFB9A9156949FB0D8F25" box="[1099,1199,1298,1322]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9A9156949FB0D8F25" box="[1099,1199,1298,1322]" italics="true" pageId="11" pageNumber="12">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
) it does not fit the current diagnosis of
|
||||
<taxonomicName id="4C064D51A632FFB9AEB66969FBE48F45" authorityName="Pilato" authorityYear="1987" box="[1000,1094,1330,1354]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AEB66969FBE48F45" box="[1000,1094,1330,1354]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
. Moreover, recently three species of
|
||||
<taxonomicName id="4C064D51A632FFB9AE12690AFC088F66" authorityName="Pilato" authorityYear="1987" box="[844,938,1361,1385]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AE12690AFC088F66" box="[844,938,1361,1385]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
, each with three placoids, were transferred to
|
||||
<taxonomicName id="4C064D51A632FFB9AE74692BFB0B8F86" authority="(Zawierucha et al. 2023)" baseAuthorityName="Zawierucha" baseAuthorityYear="2023" box="[810,1193,1392,1417]" class="Eutardigrada" family="Hypsibiidae" genus="Kararehius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AE74692BFC3A8F87" box="[810,920,1392,1416]" italics="true" pageId="11" pageNumber="12">Kararehius</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A632FFB9AEF5692BFB3F8F86" author="Zawierucha K & Stec D & Dearden PK" box="[939,1181,1392,1417]" pageId="11" pageNumber="12" refId="ref40518" refString="Zawierucha K, Stec D, Dearden PK et al. Two new tardigrade genera from New Zealand's Southern Alp glaciers display morphological stasis and parallel evolution. Molecular Phylogenetics and Evolution 2023; 178: 107634. https: // doi. org / 10.1016 / j. ympev. 2022.107634" type="journal volume" year="2023">
|
||||
Zawierucha
|
||||
<emphasis id="B972EAC0A632FFB9A971692AFBC38F87" box="[1071,1121,1392,1416]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Their systematic affinity remains to be verified genetically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
251
data/03/AF/87/03AF87C4A632FFBEAC3569A7FA05880A.xml
Normal file
251
data/03/AF/87/03AF87C4A632FFBEAC3569A7FA05880A.xml
Normal file
|
|
@ -0,0 +1,251 @@
|
|||
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<mods:mods id="92BA67468E5A1451D2997275E46742C3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B3152AFF1768B2902456461AE78BDF1C">
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<mods:title id="5648DC0A19F40F0A4AEC7070482716DF">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<mods:namePart id="C30EC95E3A6AC7BB4B13D86B66E769F3">Gąsiorek, Piotr</mods:namePart>
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||||
<mods:affiliation id="34EFB048C876863A56246C6F8F351D17">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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||||
<mods:nameIdentifier id="1EBB252319BE7B215927D70C03021637" type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="588AEE8C98C5661B14D01E7E84384D2E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F473160AED6CA1184D8FC33D2C6B29D">Blagden, Brian</mods:namePart>
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<mods:affiliation id="E90AFB5FE11F345B970067E6A933203E">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:namePart id="09533F381125E2BF6F4B88629BB354E3">Morek, Witold</mods:namePart>
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<mods:affiliation id="F196E0FF72D35E6ABDE046D8235FE855">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Tree of Life Programme, Wellcome Sanger Institute, Hinxton, Cambridge CB 10 1 SA, United Kingdom</mods:affiliation>
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<mods:namePart id="B30B72ECB22EA0CC6890E6266023ADCA">Michalczyk, Łukasz</mods:namePart>
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<mods:affiliation id="B148DAB0C83941FBA2B82D08D7FF69C1">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland</mods:affiliation>
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<mods:typeOfResource id="3FEEAEB96D89EE18CF6E26BC7FF0090A">text</mods:typeOfResource>
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<mods:title id="F176CC561B1B4B32988D1844B9C5AC4F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="8B63B1A630ED9A20B9D905BFA4C7E130">
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<mods:date id="594247764873A27F75306438F3C3D2C3">2024</mods:date>
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<mods:number id="88244C56B64DECDB98F84FF0C74D5208">2023-11-14</mods:number>
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<mods:classification id="DD973D579F2BA433F27342E0E2CC4D78">journal article</mods:classification>
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</mods:mods>
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<treatment id="03AF87C4A632FFBEAC3569A7FA05880A" LSID="urn:lsid:plazi:treatment:03AF87C4A632FFBEAC3569A7FA05880A" httpUri="http://treatment.plazi.org/id/03AF87C4A632FFBEAC3569A7FA05880A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<subSubSection id="C31C6559A632FFB9AC3569A7FB1B8C19" box="[363,1209,1532,1559]" pageId="11" pageNumber="12" type="nomenclature">
|
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<paragraph id="8BB936D2A632FFB9AC3569A7FB1B8C19" blockId="11.[137,1436,1532,1985]" box="[363,1209,1532,1559]" pageId="11" pageNumber="12">
|
||||
Taxonomic key to the genus
|
||||
<taxonomicName id="4C064D51A632FFB9AFC269A7FD5F8C19" authorityName="Pilato" authorityYear="1987" box="[668,765,1532,1558]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AFC269A7FD5F8C19" box="[668,765,1532,1558]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
and species of
|
||||
<taxonomicName id="4C064D51A632FFB9AEC269A7FBAB8C19" authorityName="Plate" authorityYear="1888" box="[924,1033,1532,1558]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AEC269A7FBAB8C19" box="[924,1033,1532,1558]" italics="true" pageId="11" pageNumber="12">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
lacking a DABT
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A632FFB9ADD76A1BFB438CB9" pageId="11" pageNumber="12" type="discussion">
|
||||
<paragraph id="8BB936D2A632FFB9ADD76A1BFB438CB9" blockId="11.[137,1436,1532,1985]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B972EAC0A632FFB9ADD76A1BFED38C57" box="[137,369,1600,1624]" italics="true" pageId="11" pageNumber="12">
|
||||
Remark:
|
||||
<taxonomicName id="4C064D51A632FFB9ADBB6A1BFED38C57" box="[229,369,1600,1624]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gani">Adropion gani</taxonomicName>
|
||||
</emphasis>
|
||||
requires further investigation (see above) in order to ascribe it to the correct genus (i.e. a new genus or
|
||||
<taxonomicName id="4C064D51A632FFB9ADD56A04FF498C78" box="[139,235,1631,1655]" class="Eutardigrada" family="Hypsibiidae" genus="Pilatobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9ADD56A04FF498C78" box="[139,235,1631,1655]" italics="true" pageId="11" pageNumber="12">Pilatobius</emphasis>
|
||||
</taxonomicName>
|
||||
). Therefore, it is excluded from the key, together with
|
||||
<taxonomicName id="4C064D51A632FFB9AE496A3BFC228C78" baseAuthorityName="Rudescu" baseAuthorityYear="1964" box="[791,896,1632,1655]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="marcusi">
|
||||
<emphasis id="B972EAC0A632FFB9AE496A3BFC228C78" box="[791,896,1632,1655]" italics="true" pageId="11" pageNumber="12">A. marcusi</emphasis>
|
||||
</taxonomicName>
|
||||
sp. dub. Note that although
|
||||
<taxonomicName id="4C064D51A632FFB9A9FC6A3BFAA08C78" authorityName="Dastych" authorityYear="1984" box="[1186,1282,1632,1655]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A632FFB9A9FC6A3BFAA08C78" box="[1186,1282,1632,1655]" italics="true" pageId="11" pageNumber="12">D. greveni</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C064D51A632FFB9A86A6A3BFE308C98" authority="Dastych & McInnes, 1996" authorityName="Dastych & McInnes" authorityYear="1996" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="mauccii">
|
||||
<emphasis id="B972EAC0A632FFB9A86A6A3BFA3E8C78" box="[1332,1436,1632,1655]" italics="true" pageId="11" pageNumber="12">D. mauccii</emphasis>
|
||||
<bibRefCitation id="EF974B23A632FFB9ADD76A24FE308C98" author="Dastych H & McInnes SJ" box="[137,402,1663,1687]" pageId="11" pageNumber="12" pagination="35 - 41" refId="ref35756" refString="Dastych H, McInnes SJ. A new species of the genus Diphascon (Tardigrada) from the maritime Antarctic. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 1996; 12: 35 - 41." type="journal article" year="1996">Dastych & McInnes, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
belong within
|
||||
<taxonomicName id="4C064D51A632FFB9AF766A24FBAF8C98" authority="(Tumanov and Tsvetkova 2023)" baseAuthorityName="Tumanov and Tsvetkova" baseAuthorityYear="2023" box="[552,1037,1663,1687]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">
|
||||
Diphasconinae (
|
||||
<bibRefCitation id="EF974B23A632FFB9AF8F6A24FBA08C98" author="Tumanov DV & Tsvetkova AY" box="[721,1026,1663,1687]" pageId="11" pageNumber="12" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, they cannot be distinguished morphologically from
|
||||
<taxonomicName id="4C064D51A632FFB9AC456AC5FED58CB9" authorityName="Pilato" authorityYear="1987" box="[283,375,1694,1718]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A632FFB9AC456AC5FED58CB9" box="[283,375,1694,1718]" italics="true" pageId="11" pageNumber="12">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
based on the currently recognized taxonomic criteria and are thus included in the key.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A632FFBEADD26A96FA05880A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="key">
|
||||
<key id="FF9761E3A632FFBEADD26A96FA05880A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||||
<keyStep id="30F22E77A632FFB9ADD26A96FA3A8D0A" pageId="11" pageNumber="12">
|
||||
<paragraph id="8BB936D2A632FFB9ADD26A96FA3A8CEA" blockId="11.[137,1436,1532,1985]" box="[140,1432,1741,1765]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26A96FA3A8CEA" box="[140,1432,1741,1765]" pageId="11" pageNumber="12">1. Macroplacoids granular ............................................................................................................................................................................... 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A632FFB9ADD26AB6FA3A8D0A" blockId="11.[137,1436,1532,1985]" box="[140,1432,1773,1797]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26AB6FA3A8D0A" box="[140,1432,1773,1797]" pageId="11" pageNumber="12">– Macroplacoids bar-shaped .......................................................................................................................................................................... 4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A632FFB9ADD26B57FA3A8D4C" pageId="11" pageNumber="12">
|
||||
<paragraph id="8BB936D2A632FFB9ADD26B57FA3A8D2B" blockId="11.[137,1436,1532,1985]" box="[140,1432,1804,1828]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26B57FA3A8D2B" box="[140,1432,1804,1828]" pageId="11" pageNumber="12">
|
||||
2. Internal bars I–III present .........................................................................................................................................
|
||||
<taxonomicName id="4C064D51A632FFB9A9926B57FA3A8D2B" authorityName="Pilato, Claxton & Horning" authorityYear="1991" box="[1228,1432,1804,1828]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="gordonense">
|
||||
<emphasis id="B972EAC0A632FFB9A9926B57FA3A8D2B" box="[1228,1432,1804,1828]" italics="true" pageId="11" pageNumber="12">Adropion gordonense</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A632FFB9ADD26B77FA3A8D4C" blockId="11.[137,1436,1532,1985]" box="[140,1432,1835,1859]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26B77FA3A8D4C" box="[140,1432,1835,1859]" pageId="11" pageNumber="12">– Internal bars I–III absent ............................................................................................................................................................................. 3</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A632FFB9ADD26B10FA3B8D8D" pageId="11" pageNumber="12">
|
||||
<paragraph id="8BB936D2A632FFB9ADD26B10FA3B8D6C" blockId="11.[137,1436,1532,1985]" box="[140,1433,1867,1891]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26B10FA3B8D6C" box="[140,1433,1867,1891]" pageId="11" pageNumber="12">
|
||||
3. Macroplacoid length sequence 1 <2 <3, dorsal granulation absent ...............................................................
|
||||
<taxonomicName id="4C064D51A632FFB9A98C6B10FA3B8D6C" baseAuthorityName="X. Li" baseAuthorityYear="2007" box="[1234,1433,1867,1891]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="linzhiensis">
|
||||
<emphasis id="B972EAC0A632FFB9A98C6B10FA3B8D6C" box="[1234,1433,1867,1891]" italics="true" pageId="11" pageNumber="12">Adropion linzhiensis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A632FFB9ADD26B31FA3B8D8D" blockId="11.[137,1436,1532,1985]" box="[140,1433,1898,1922]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26B31FA3B8D8D" box="[140,1433,1898,1922]" pageId="11" pageNumber="12">
|
||||
– Macroplacoid length sequence 2 <1 <3, dorsal granulation present ..........................................................
|
||||
<taxonomicName id="4C064D51A632FFB9A99D6B31FA3B8D8D" box="[1219,1433,1898,1922]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="afroglacialis">
|
||||
<emphasis id="B972EAC0A632FFB9A99D6B31FA3B8D8D" box="[1219,1433,1898,1922]" italics="true" pageId="11" pageNumber="12">Adropion afroglacialis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A632FFB9ADD26BD2FA3A8DCE" pageId="11" pageNumber="12">
|
||||
<paragraph id="8BB936D2A632FFB9ADD26BD2FA3A8DAE" blockId="11.[137,1436,1532,1985]" box="[140,1432,1929,1953]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26BD2FA3A8DAE" box="[140,1432,1929,1953]" pageId="11" pageNumber="12">4. Three macroplacoids present ...................................................................................................................................................................... 5</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A632FFB9ADD26BF2FA3A8DCE" blockId="11.[137,1436,1532,1985]" box="[140,1432,1961,1985]" pageId="11" pageNumber="12">
|
||||
<keyLead id="30F795E7A632FFB9ADD26BF2FA3A8DCE" box="[140,1432,1961,1985]" pageId="11" pageNumber="12">– Two macroplacoids present ........................................................................................................................................................................ 9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A635FFBEADC56CEFFA058AE4" pageId="12" pageNumber="13">
|
||||
<paragraph id="8BB936D2A635FFBEADC56CEFFA058AC3" blockId="12.[155,1448,180,517]" box="[155,1447,180,204]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56CEFFA058AC3" box="[155,1447,180,204]" pageId="12" pageNumber="13">5. Both microplacoid and septulum present ................................................................................................................................................ 6</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A635FFBEADC56C8FFA058AE4" blockId="12.[155,1448,180,517]" box="[155,1447,211,235]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56C8FFA058AE4" box="[155,1447,211,235]" pageId="12" pageNumber="13">– Only microplacoid or septulum present .................................................................................................................................................. 7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A635FFBEADC56CA9FA0A8B25" pageId="12" pageNumber="13">
|
||||
<paragraph id="8BB936D2A635FFBEADC56CA9FA058B05" blockId="12.[155,1448,180,517]" box="[155,1447,242,267]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56CA9FA058B05" box="[155,1447,242,267]" pageId="12" pageNumber="13">
|
||||
6. Bases of external claws I-III with indented bases,
|
||||
<emphasis id="B972EAC0A635FFBEAFE46CA8FD6C8B05" box="[698,718,243,266]" italics="true" pageId="12" pageNumber="13">pt</emphasis>
|
||||
of SSIP = 67–75% .............................................................
|
||||
<taxonomicName id="4C064D51A635FFBEA9A86CA8FA058B05" authority="Bases" authorityName="Bases" box="[1270,1447,243,267]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A635FFBEA9A86CA8FA058B05" box="[1270,1447,243,267]" italics="true" pageId="12" pageNumber="13">Diphascon greveni</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A635FFBEADC56D49FA0A8B25" blockId="12.[155,1448,180,517]" box="[155,1448,274,298]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56D49FA0A8B25" box="[155,1448,274,298]" pageId="12" pageNumber="13">
|
||||
– Bases of external claws I-III with smooth bases,
|
||||
<emphasis id="B972EAC0A635FFBEAFF06D48FD608B25" box="[686,706,275,298]" italics="true" pageId="12" pageNumber="13">pt</emphasis>
|
||||
of SSIP = 64–65% ...................................................................
|
||||
<taxonomicName id="4C064D51A635FFBEA8546D49FA0A8B25" authority="Microplacoid" authorityName="Microplacoid" box="[1290,1448,274,298]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="onorei">
|
||||
<emphasis id="B972EAC0A635FFBEA8546D49FA0A8B25" box="[1290,1448,274,298]" italics="true" pageId="12" pageNumber="13">Adropion onorei</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A635FFBEADC56D69FA058B66" pageId="12" pageNumber="13">
|
||||
<paragraph id="8BB936D2A635FFBEADC56D69FA0A8B46" blockId="12.[155,1448,180,517]" box="[155,1448,305,329]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56D69FA0A8B46" box="[155,1448,305,329]" pageId="12" pageNumber="13">
|
||||
7. Microplacoid absent, septulum present ....................................................................................................................
|
||||
<taxonomicName id="4C064D51A635FFBEA9AE6D6AFA0A8B46" authority="Microplacoid" authorityName="Microplacoid" box="[1264,1448,305,329]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="mauccii">
|
||||
<emphasis id="B972EAC0A635FFBEA9AE6D6AFA0A8B46" box="[1264,1448,305,329]" italics="true" pageId="12" pageNumber="13">Diphascon mauccii</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A635FFBEADC56D0AFA058B66" blockId="12.[155,1448,180,517]" box="[155,1447,337,361]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56D0AFA058B66" box="[155,1447,337,361]" pageId="12" pageNumber="13">– Microplacoid present, septulum absent ................................................................................................................................................... 8</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A635FFBEADC56D2BFA058BA8" pageId="12" pageNumber="13">
|
||||
<paragraph id="8BB936D2A635FFBEADC56D2BFA0A8B87" blockId="12.[155,1448,180,517]" box="[155,1448,368,392]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56D2BFA0A8B87" box="[155,1448,368,392]" pageId="12" pageNumber="13">
|
||||
8. Eyes present .................................................................................................................................
|
||||
<taxonomicName id="4C064D51A635FFBEA9736D2BFA8F8B87" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Thulin" baseAuthorityYear="1911" box="[1069,1325,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="ommatophorum" status="comb. nov.">
|
||||
<emphasis id="B972EAC0A635FFBEA9736D2BFA8F8B87" box="[1069,1325,368,392]" italics="true" pageId="12" pageNumber="13">Adropion ommatophorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A635FFBEA86F6D2BFA0A8B87" bold="true" box="[1329,1448,368,392]" pageId="12" pageNumber="13">
|
||||
<taxonomicNameLabel id="A24157BBA635FFBEA86F6D2BFA0A8B87" box="[1329,1448,368,392]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A635FFBEADC56DCBFA058BA8" blockId="12.[155,1448,180,517]" box="[155,1447,399,424]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56DCBFA058BA8" box="[155,1447,399,424]" pageId="12" pageNumber="13">
|
||||
– Eyes absent .......................................................................................................................................................................
|
||||
<taxonomicName id="4C064D51A635FFBEA9AC6DD4FA058BA8" authority="Cuticular" authorityName="Cuticular" box="[1266,1447,399,423]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="scoticum">
|
||||
<emphasis id="B972EAC0A635FFBEA9AC6DD4FA058BA8" box="[1266,1447,399,423]" italics="true" pageId="12" pageNumber="13">Adropion scoticum</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30F22E77A635FFBEADC56DF4FA05880A" pageId="12" pageNumber="13">
|
||||
<paragraph id="8BB936D2A635FFBEADC56DF4FA058BC8" blockId="12.[155,1448,180,517]" box="[155,1447,431,455]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56DF4FA058BC8" box="[155,1447,431,455]" pageId="12" pageNumber="13">
|
||||
9. Cuticular bars absent (the edges of external pulvini I–III may be faintly visible) ..............................................
|
||||
<taxonomicName id="4C064D51A635FFBEA9A56DF4FA058BC8" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1275,1447,431,455]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A635FFBEA9A56DF4FA058BC8" box="[1275,1447,431,455]" italics="true" pageId="12" pageNumber="13">Adropion belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A635FFBEADC56D95FA05880A" blockId="12.[155,1448,180,517]" pageId="12" pageNumber="13">
|
||||
<keyLead id="30F795E7A635FFBEADC56D95FA05880A" pageId="12" pageNumber="13">
|
||||
– Median and internal cuticular bars I–III and posterior bars between bases of claw IV present ...................................................... ....................................................................................................................................
|
||||
<taxonomicName id="4C064D51A635FFBEAEEB6DB6FB11880A" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[949,1203,493,517]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A635FFBEAEEB6DB6FB11880A" box="[949,1203,493,517]" italics="true" pageId="12" pageNumber="13">Adropion diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA635FFBEA9E76DB6FA05880A" box="[1209,1447,493,517]" pageId="12" pageNumber="13">comb. nov. et stat. rev.</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
2968
data/03/AF/87/03AF87C4A635FFA1AC5D6EFBFBF38DB2.xml
Normal file
2968
data/03/AF/87/03AF87C4A635FFA1AC5D6EFBFBF38DB2.xml
Normal file
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|
|
@ -1,110 +0,0 @@
|
|||
<document id="73F8B15A7DFF4CFB298858DFC632AE96" ID-DOI="10.1093/zoolinnean/zlad151" ID-ISSN="0024-4082" ID-Zenodo-Dep="14243740" ID-ZooBank="044A402-2A0F-4135-9410-7DE081CB11C4Corresponding" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732840371232" checkinUser="plazi" docAuthor="Gąsiorek, Piotr, Blagden, Brian, Morek, Witold & Michalczyk, Łukasz" docDate="2024" docId="03AF87C4A635FFBEAC5D6EFBFA8C8879" docLanguage="en" docName="zlad151.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad151" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.6:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="6" docTitle="Guidettion prorsirostre" docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FF96FFBCA639FFB2AD5E6C5BFFA28A0F" masterDocTitle="What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="13" updateTime="1732902770667" updateUser="ExternalLinkService">
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<mods:mods id="D03DBA65977EC801A822EACFD7B74FA3" xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo id="B0E70B1AEF5B89794783028DA45022E7">
|
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<mods:title id="0C1294AEF0D4DF24A6A9FB0B6960E08B">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
|
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</mods:titleInfo>
|
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<mods:name id="4D3985F32EA5A03613D1C6D3DA523716" type="personal">
|
||||
<mods:role id="2962571CC7A02D82B8F9E6486D21EDFF">
|
||||
<mods:roleTerm id="5A18A64085634D879CAFE1FFA51CBB9B">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="1689DCB6D05B76D8632A823B32D63BE4">Gąsiorek, Piotr</mods:namePart>
|
||||
<mods:affiliation id="33E3D2717237F4EEFF5CCAFC1A12E347">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland & Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade 5 - 7, DK- 1350, Universitetsparken 15, DK- 2100, Copenhagen, Denmark</mods:affiliation>
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.
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<taxonomicName id="4C064D51A63DFFB6A9496DEAFAF68BC4" ID-CoL="JG4" authority="Dastych, 1992" authorityName="Dastych" authorityYear="1992" box="[1047,1364,433,459]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">
|
||||
Diphasconinae
|
||||
<bibRefCitation id="EF974B23A63DFFB6A9E26DEAFAF68BC4" author="Dastych H" box="[1212,1364,433,459]" pageId="4" pageNumber="5" pagination="125 - 39" refId="ref35602" refString="Dastych H. Paradiphascon manningi gen. n. sp. n., a new water-bear from South Africa, with the erecting of a new subfamily Diphasconinae (Tardigrada). Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut, Hamburg 1992; 89: 125 - 39." type="journal article" year="1992">Dastych, 1992</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="C31C6559A63DFFB6AE676D83FA6089A9" pageId="4" pageNumber="5" type="discussion">
|
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<paragraph id="8BB936D2A63DFFB6AE676D83FA6089A9" blockId="4.[825,1475,472,934]" pageId="4" pageNumber="5">
|
||||
The phylogenetic position of
|
||||
<taxonomicName id="4C064D51A63DFFB6A92E6D83FA608BFF" authority="Dastych, 1984" authorityName="Dastych" authorityYear="1984" box="[1136,1474,472,496]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A63DFFB6A92E6D83FA868BE0" box="[1136,1316,472,496]" italics="true" pageId="4" pageNumber="5">Diphascon greveni</emphasis>
|
||||
<bibRefCitation id="EF974B23A63DFFB6A8726D83FA608BFF" author="Dastych H" box="[1324,1474,472,496]" pageId="4" pageNumber="5" pagination="377 - 436" refId="ref35540" refString="Dastych H. The Tardigrada from Antarctica with descriptions of several new species. Acta Zoologica Cracoviensia 1984; 27: 377 - 436." type="journal article" year="1984">Dastych, 1984</bibRefCitation>
|
||||
</taxonomicName>
|
||||
among the
|
||||
<taxonomicName id="4C064D51A63DFFB6AEF56DACFBE18800" authorityName="Dastych" authorityYear="1992" box="[939,1091,503,527]" class="Eutardigrada" family="Hypsibiidae" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="subFamily" subFamily="Diphasconinae">Diphasconinae</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133D2A57A63DFFB6A90A6DACFB2E8800" box="[1108,1164,503,527]" captionStart="Figure 2" captionStartId="5.[113,178,1715,1739]" captionTargetBox="[115,1454,150,1684]" captionTargetId="graphics-78@5.[115,1181,156,1684]" captionTargetPageId="5" captionText="Figure 2. Phylogenetic relationships within Hypsibiidae in the Bayesian and maximum likelihood analyses (identical topologies; the scale refers to the Bayesian tree and represents substitutions per position; posterior probability values are provided above the nodes, whereas bootstrap values can be found below the nodes; *, maximal support; #, no support).Calohypsibius ornatus constitutes an outgroup. Newly added species and populations are in bold." figureDoi="http://doi.org/10.5281/zenodo.14243746" httpUri="https://zenodo.org/record/14243746/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||||
) further undermines the diagnosis of this group, which had previously been challenged by the inclusion of taxa lacking a DABT (
|
||||
<bibRefCitation id="EF974B23A63DFFB6A9986E6DFCCF8862" author="Tumanov DV & Tsvetkova AY" pageId="4" pageNumber="5" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF974B23A63DFFB6AE246E0EFBC68862" author="Zawierucha K & Stec D & Dearden PK" box="[890,1124,597,621]" pageId="4" pageNumber="5" refId="ref40518" refString="Zawierucha K, Stec D, Dearden PK et al. Two new tardigrade genera from New Zealand's Southern Alp glaciers display morphological stasis and parallel evolution. Molecular Phylogenetics and Evolution 2023; 178: 107634. https: // doi. org / 10.1016 / j. ympev. 2022.107634" type="journal volume" year="2023">
|
||||
Zawierucha
|
||||
<emphasis id="B972EAC0A63DFFB6AEA76E0DFB888862" box="[1017,1066,597,621]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="4C064D51A63DFFB6A9256E0EFB408862" authorityName="Plate" authorityYear="1888" box="[1147,1250,597,621]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63DFFB6A9256E0EFB408862" box="[1147,1250,597,621]" italics="true" pageId="4" pageNumber="5">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
should group species with an apodeme according to its last broad diagnosis (Pilato 1987,
|
||||
<bibRefCitation id="EF974B23A63DFFB6AE276ECFFBE488A3" author="Bertolani R & Guidetti R & Marchioro T" box="[889,1094,660,684]" pageId="4" pageNumber="5" pagination="110 - 26" refId="ref34993" refString="Bertolani R, Guidetti R, Marchioro T et al. Phylogeny of Eutardigrada: new molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 2014; 76: 110 - 26. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
|
||||
Bertolani
|
||||
<emphasis id="B972EAC0A63DFFB6AE836ECFFBAF88A3" box="[989,1037,660,684]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
), although its morphology is diversified among species currently classified within this genus (
|
||||
<figureCitation id="133D2A57A63DFFB6A8246EE8FA1088C4" box="[1402,1458,691,715]" captionStart="Figure 3" captionStartId="6.[129,194,1333,1357]" captionTargetBox="[131,1470,146,1304]" captionTargetId="figure-244@6.[129,1473,144,1306]" captionTargetPageId="6" captionText="Figure 3. Various Diphasconinae, morphology of the DABT (SEM): A, Diphascon higginsi Binda, 1971 (dorsal view); B, Diphascon higginsi (lateral view); C, Diphascon pingue (Marcus, 1936) (lateral view); D, Diphascon pinguiforme Pilato & Binda, 1997/8 (dorsolateral view). Scale bars: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.14243748" httpUri="https://zenodo.org/record/14243748/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
|
||||
). The
|
||||
<typeStatus id="54BD8870A63DFFB6AE396E88FC3788E4" box="[871,917,723,747]" pageId="4" pageNumber="5">type</typeStatus>
|
||||
species of
|
||||
<taxonomicName id="4C064D51A63DFFB6A9586E89FBCE88E5" authorityName="Plate" authorityYear="1888" box="[1030,1132,722,746]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63DFFB6A9586E89FBCE88E5" box="[1030,1132,722,746]" italics="true" pageId="4" pageNumber="5">Diphascon</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C064D51A63DFFB6A9296E89FA1F88E5" authority="Plate, 1888" authorityName="Plate" authorityYear="1888" box="[1143,1469,722,746]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="chilenense">
|
||||
<emphasis id="B972EAC0A63DFFB6A9296E89FAE688E5" box="[1143,1348,722,746]" italics="true" pageId="4" pageNumber="5">Diphascon chilenense</emphasis>
|
||||
<bibRefCitation id="EF974B23A63DFFB6A8156E89FA1F88E5" author="Plate L" box="[1355,1469,722,746]" pageId="4" pageNumber="5" pagination="487 - 550" refId="ref39183" refString="Plate L. Beitrage zur Naturgeschichte der Tardigraden. Zoologische Jahrbucher 1888; 3: 487 - 550." type="journal article" year="1888">Plate, 1888</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, and members of the
|
||||
<taxonomicName id="4C064D51A63DFFB6A94F6EA9FB628906" baseAuthorityName="Marcus" baseAuthorityYear="1936" box="[1041,1216,754,778]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="pingue">
|
||||
<emphasis id="B972EAC0A63DFFB6A94F6EA9FB628906" box="[1041,1216,754,778]" italics="true" pageId="4" pageNumber="5">Diphascon pingue</emphasis>
|
||||
</taxonomicName>
|
||||
group (
|
||||
<bibRefCitation id="EF974B23A63DFFB6A8486EA9FCD38926" author="Pilato G & Binda MG" pageId="4" pageNumber="5" pagination="181 - 5" refId="ref38708" refString="Pilato G, Binda MG. A comparison of Diphascon (D.) alpinum Murray, 1906, D. (D.) chilenense Plate, 1889 and D. (D.) pingue Marcus, 1936 (Tardigrada), and description of a new species. Zoologischer Anzeiger 1997 / 8; 236: 181 - 5." type="journal article" year="1997">Pilato and Binda 1997</bibRefCitation>
|
||||
/8, 1999) have a disparate pharyngeal apparatus (short, rodshaped, or granular macroplacoids;
|
||||
<figureCitation id="133D2A57A63DFFB6A9C16F6BFB748947" box="[1183,1238,816,840]" captionStart="Figure 4" captionStartId="7.[113,178,925,949]" captionTargetBox="[119,756,145,895]" captionTargetId="figure-733@7.[118,758,144,897]" captionTargetPageId="7" captionText="Figure 4. The Diphascon pingue group, morphology of pharyngeal structures (SEM): A, Diphascon pingue; B, C, Diphascon pinguiforme. Scale bars: 2 μm." figureDoi="http://doi.org/10.5281/zenodo.14243750" httpUri="https://zenodo.org/record/14243750/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||||
) and claw morphology (minute claws with smooth bases and no bars) with respect to
|
||||
<taxonomicName id="4C064D51A63DFFB6AE676F2BFC358988" authorityName="Dastych" authorityYear="1984" box="[825,919,880,903]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A63DFFB6AE676F2BFC358988" box="[825,919,880,903]" italics="true" pageId="4" pageNumber="5">D. greveni</emphasis>
|
||||
</taxonomicName>
|
||||
, thus highlighting the artificial nature of the current definition of
|
||||
<taxonomicName id="4C064D51A63DFFB6AEC76FD5FBD789A9" authority="Plate, 1888" authorityName="Plate" authorityYear="1888" box="[921,1141,910,934]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63DFFB6AEC76FD5FBA289A9" box="[921,1024,910,934]" italics="true" pageId="4" pageNumber="5">Diphascon</emphasis>
|
||||
<bibRefCitation id="EF974B23A63DFFB6A95B6FD5FBD789A9" author="Plate L" box="[1029,1141,910,934]" pageId="4" pageNumber="5" pagination="487 - 550" refId="ref39183" refString="Plate L. Beitrage zur Naturgeschichte der Tardigraden. Zoologische Jahrbucher 1888; 3: 487 - 550." type="journal article" year="1888">Plate, 1888</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A63DFFB6A9DD6FD4FA1089A9" author="Tumanov DV & Tsvetkova AY" box="[1155,1458,910,934]" pageId="4" pageNumber="5" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova 2023</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
197
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Normal file
197
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|
||||
</mods:mods>
|
||||
<treatment id="03AF87C4A63DFFB7AE8D685BFB028DB5" LSID="urn:lsid:plazi:treatment:03AF87C4A63DFFB7AE8D685BFB028DB5" httpUri="http://treatment.plazi.org/id/03AF87C4A63DFFB7AE8D685BFB028DB5" lastPageId="5" lastPageNumber="5" pageId="4" pageNumber="5">
|
||||
<subSubSection id="C31C6559A63DFFB6AE8D685BFA8A8E14" box="[979,1320,1024,1051]" pageId="4" pageNumber="5" type="nomenclature">
|
||||
<paragraph id="8BB936D2A63DFFB6AE8D685BFA8A8E14" blockId="4.[979,1320,1024,1051]" box="[979,1320,1024,1051]" pageId="4" pageNumber="5">
|
||||
<heading id="D0F181BEA63DFFB6AE8D685BFA8A8E14" bold="true" box="[979,1320,1024,1051]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
|
||||
<taxonomicName id="4C064D51A63DFFB6AE8D685BFA8A8E14" ID-CoL="7T83Y" authority="Dastych, 1984" authorityName="Dastych" authorityYear="1984" box="[979,1320,1024,1051]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A63DFFB6AE8D685BFA8A8E14" bold="true" box="[979,1320,1024,1051]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B972EAC0A63DFFB6AE8D685BFB298E14" bold="true" box="[979,1163,1024,1051]" italics="true" pageId="4" pageNumber="5">Diphascon greveni</emphasis>
|
||||
<bibRefCitation id="EF974B23A63DFFB6A9CE685AFA8A8E14" author="Dastych H" box="[1168,1320,1025,1051]" pageId="4" pageNumber="5" pagination="377 - 436" refId="ref35540" refString="Dastych H. The Tardigrada from Antarctica with descriptions of several new species. Acta Zoologica Cracoviensia 1984; 27: 377 - 436." type="journal article" year="1984">Dastych, 1984</bibRefCitation>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63DFFB6AE63686BFB7F8F0A" pageId="4" pageNumber="5" type="reference_group">
|
||||
<paragraph id="8BB936D2A63DFFB6AE63686BFB4A8E86" blockId="4.[829,1471,1072,1161]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B972EAC0A63DFFB6AE63686BFB4A8E86" bold="true" pageId="4" pageNumber="5">
|
||||
<taxonomicName id="4C064D51A63DFFB6AE63686BFB818E44" baseAuthorityName="Murray" baseAuthorityYear="1905" box="[829,1059,1072,1099]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="scoticus" subGenus="Diphascon">
|
||||
<emphasis id="B972EAC0A63DFFB6AE63686BFC038E44" bold="true" box="[829,929,1072,1099]" italics="true" pageId="4" pageNumber="5">Hypsibius</emphasis>
|
||||
(
|
||||
<emphasis id="B972EAC0A63DFFB6AEEE686AFC6A8E44" bold="true" box="[944,968,1073,1099]" italics="true" pageId="4" pageNumber="5">D.</emphasis>
|
||||
)
|
||||
<emphasis id="B972EAC0A63DFFB6AE89686AFB818E44" bold="true" box="[983,1059,1073,1099]" italics="true" pageId="4" pageNumber="5">scoticus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<collectingCountry id="F3117642A63DFFB6A970686BFB6B8E45" box="[1070,1225,1072,1098]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="5">South Georgia</collectingCountry>
|
||||
, South
|
||||
<collectingCountry id="F3117642A63DFFB6A849686BFA188E44" box="[1303,1466,1072,1099]" name="United Kingdom" pageId="4" pageNumber="5">Orkney Islands</collectingCountry>
|
||||
, South
|
||||
<collectingRegion id="49C2F830A63DFFB6AECF680BFBE28E65" box="[913,1088,1104,1130]" country="United Kingdom" name="Shetland Islands" pageId="4" pageNumber="5">Shetland Islands</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49C2F830A63DFFB6A917680BFB498E65" box="[1097,1259,1104,1130]" country="United Kingdom" name="Marguerite Bay" pageId="4" pageNumber="5">Marguerite Bay</collectingRegion>
|
||||
, Alexander Island;
|
||||
<bibRefCitation id="EF974B23A63DFFB6A94D6834FB608E86" author="Jennings PG" box="[1043,1218,1135,1161]" pageId="4" pageNumber="5" pagination="77 - 95" refId="ref36812" refString="Jennings PG. Tardigrada from the Antarctic Peninsula and Scotia Ridge region. British Antarctic Survey Bulletin 1976 a; 44: 77 - 95." type="journal article" year="1976">Jennings (1976a</bibRefCitation>
|
||||
, b).
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A63DFFB6AE0668C4FA018EB6" blockId="4.[856,1443,1183,1210]" box="[856,1443,1183,1210]" pageId="4" pageNumber="5">
|
||||
<heading id="D0F181BEA63DFFB6AE0668C4FA018EB6" bold="true" box="[856,1443,1183,1210]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
|
||||
<emphasis id="B972EAC0A63DFFB6AE0668C4FA018EB6" bold="true" box="[856,1443,1183,1210]" pageId="4" pageNumber="5">
|
||||
<taxonomicName id="4C064D51A63DFFB6AE0668C4FBB28EB6" ID-CoL="7T83Y" authorityName="Dastych" authorityYear="1984" box="[856,1040,1183,1210]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A63DFFB6AE0668C4FBB28EB6" bold="true" box="[856,1040,1183,1210]" italics="true" pageId="4" pageNumber="5">Diphascon greveni</emphasis>
|
||||
</taxonomicName>
|
||||
; King George Island;
|
||||
<bibRefCitation id="EF974B23A63DFFB6A9AB68C4FA3F8EB6" author="Dastych H" box="[1269,1437,1183,1209]" pageId="4" pageNumber="5" pagination="377 - 436" refId="ref35540" refString="Dastych H. The Tardigrada from Antarctica with descriptions of several new species. Acta Zoologica Cracoviensia 1984; 27: 377 - 436." type="journal article" year="1984">Dastych (1984)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A63DFFB6AE386890FB7F8F0A" blockId="4.[870,1429,1227,1285]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B972EAC0A63DFFB6AE386890FB7F8F0A" bold="true" pageId="4" pageNumber="5">
|
||||
<taxonomicName id="4C064D51A63DFFB6AE386890FBBC8EE9" authorityName="Dastych" authorityYear="1984" box="[870,1054,1227,1254]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="greveni">
|
||||
<emphasis id="B972EAC0A63DFFB6AE386890FBBC8EE9" bold="true" box="[870,1054,1227,1254]" italics="true" pageId="4" pageNumber="5">Diphascon greveni</emphasis>
|
||||
</taxonomicName>
|
||||
; King George Island;
|
||||
<bibRefCitation id="EF974B23A63DFFB6A85D6897FB7A8F0A" author="Tumanov DV & Tsvetkova AY" pageId="4" pageNumber="5" pagination="50 - 74" refId="ref40208" refString="Tumanov DV, Tsvetkova AY. Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica 2023; 32: 50 - 74. https: // doi. org / 10.31610 / zsr / 2023.32.1.50" type="journal article" year="2023">Tumanov and Tsvetkova (2023)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63DFFB6AE676974FB008F69" pageId="4" pageNumber="5" type="materials_examined">
|
||||
<paragraph id="8BB936D2A63DFFB6AE676974FB008F69" blockId="4.[824,1477,1327,1978]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B972EAC0A63DFFB6AE676974FBA28F48" box="[825,1024,1327,1351]" italics="true" pageId="4" pageNumber="5">Material examined:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA63DFFB6A9536974FB008F69" pageId="4" pageNumber="5" specimenCount="128">
|
||||
<specimenCount id="9D00FD5BA63DFFB6A9536974FB138F48" box="[1037,1201,1327,1351]" pageId="4" pageNumber="5">128 individuals</specimenCount>
|
||||
in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A63DFFB6A9646915FB338F69" box="[1082,1169,1358,1382]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="4" pageNumber="5">Table S1</tableCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63DFFB7AE676935FB028DB5" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||||
<paragraph id="8BB936D2A63DFFB7AE676935FB028DB5" blockId="4.[824,1477,1327,1978]" lastBlockId="5.[810,1461,1891,1978]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||||
<emphasis id="B972EAC0A63DFFB6AE676935FBAD8F89" box="[825,1039,1390,1414]" italics="true" pageId="4" pageNumber="5">Amended description:</emphasis>
|
||||
Large, body can exceed 500 μm (
|
||||
<tableCitation id="C6840369A63DFFB6A8D46935FCE48FAA" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="4" pageNumber="5">Table 1</tableCitation>
|
||||
), with a poorly demarcated, blunt cephalic region and a large interval between legs III and IV (
|
||||
<figureCitation id="133D2A57A63DFFB6A9DA69F7FB6E8FCB" box="[1156,1228,1452,1476]" captionStart="Figure 5" captionStartId="9.[113,178,836,860]" captionTargetBox="[116,1454,146,804]" captionTargetId="figure-648@9.[114,1458,144,808]" captionTargetPageId="9" captionText="Figure 5. General morphology of Diphascon greveni Dastych, 1984 (PCM, topotypes): A, specimen in toto (ventral view); B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243752" httpUri="https://zenodo.org/record/14243752/files/figure.png" pageId="4" pageNumber="5">Fig. 5A</figureCitation>
|
||||
). Lacking a DABT (
|
||||
<figureCitation id="133D2A57A63DFFB6A8C769F7FCF78FEC" captionStart="Figure 5" captionStartId="9.[113,178,836,860]" captionTargetBox="[116,1454,146,804]" captionTargetId="figure-648@9.[114,1458,144,808]" captionTargetPageId="9" captionText="Figure 5. General morphology of Diphascon greveni Dastych, 1984 (PCM, topotypes): A, specimen in toto (ventral view); B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243752" httpUri="https://zenodo.org/record/14243752/files/figure.png" pageId="4" pageNumber="5">Figs 5B</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A63DFFB6AE3C6997FC228FEB" box="[866,896,1484,1508]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">6A</figureCitation>
|
||||
). The pharyngeal annulation simple in both PCM (
|
||||
<figureCitation id="133D2A57A63DFFB6A8C36997FCF68C0C" captionStart="Figure 5" captionStartId="9.[113,178,836,860]" captionTargetBox="[116,1454,146,804]" captionTargetId="figure-648@9.[114,1458,144,808]" captionTargetPageId="9" captionText="Figure 5. General morphology of Diphascon greveni Dastych, 1984 (PCM, topotypes): A, specimen in toto (ventral view); B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243752" httpUri="https://zenodo.org/record/14243752/files/figure.png" pageId="4" pageNumber="5">Fig. 5B</figureCitation>
|
||||
) and SEM (
|
||||
<figureCitation id="133D2A57A63DFFB6AE8F69B0FBEC8C0D" box="[977,1102,1515,1539]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6A, D, E</figureCitation>
|
||||
), but occasional annular bifurcations present (
|
||||
<figureCitation id="133D2A57A63DFFB6AECD6A51FC518C2D" box="[915,1011,1546,1570]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6D, E</figureCitation>
|
||||
). One band of teeth in the OCA (
|
||||
<figureCitation id="133D2A57A63DFFB6A8186A51FA288C2D" box="[1350,1418,1546,1570]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6B</figureCitation>
|
||||
), not detectable in PCM (
|
||||
<figureCitation id="133D2A57A63DFFB6A9576A71FBED8C4E" box="[1033,1103,1577,1602]" captionStart="Figure 5" captionStartId="9.[113,178,836,860]" captionTargetBox="[116,1454,146,804]" captionTargetId="figure-648@9.[114,1458,144,808]" captionTargetPageId="9" captionText="Figure 5. General morphology of Diphascon greveni Dastych, 1984 (PCM, topotypes): A, specimen in toto (ventral view); B, buccopharyngeal apparatus in toto. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243752" httpUri="https://zenodo.org/record/14243752/files/figure.png" pageId="4" pageNumber="5">Fig. 5B</figureCitation>
|
||||
). Lateral porous areas in the walls of the buccal crown reduced and partly hidden under the apophyses for the insertion of stylet muscles (AISMs) (
|
||||
<figureCitation id="133D2A57A63DFFB6A9B16A33FA958C8F" box="[1263,1335,1640,1664]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6C</figureCitation>
|
||||
). Furca of the
|
||||
<taxonomicName id="4C064D51A63DFFB6AE666AD3FC358CAF" authorityName="Ehrenberg" authorityYear="1848" box="[824,919,1672,1696]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63DFFB6AE666AD3FC358CAF" box="[824,919,1672,1696]" italics="true" pageId="4" pageNumber="5">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A63DFFB6AEC36AD3FC688CAF" box="[925,970,1672,1696]" pageId="4" pageNumber="5">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A63DFFB6AE846AD3FBBD8CAF" box="[986,1055,1672,1696]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6F</figureCitation>
|
||||
). Large pharyngeal apophyses. Three bar-shaped macroplacoids occasionally merged, but microplacoid and septulum always separate (
|
||||
<figureCitation id="133D2A57A63DFFB6A9266A9DFB668CD1" box="[1144,1220,1734,1758]" captionStart="Figure 6" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[265,1336,146,1805]" captionTargetId="figure-4@10.[263,1339,144,1808]" captionTargetPageId="10" captionText="Figure 6. Detailed morphology of buccopharyngeal apparatus of Diphascon greveni (SEM, topotypes): A, in toto; B, OCA; C, AISM in lateral view; D, annulation of pharyngeal tube; E, annuli in close-up; F, furca; G, pharynx. Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243754" httpUri="https://zenodo.org/record/14243754/files/figure.png" pageId="4" pageNumber="5">Fig. 6G</figureCitation>
|
||||
). Claws of the
|
||||
<taxonomicName id="4C064D51A63DFFB6A83B6A9DFA668CD1" authorityName="Ehrenberg" authorityYear="1848" box="[1381,1476,1734,1758]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63DFFB6A83B6A9DFA668CD1" box="[1381,1476,1734,1758]" italics="true" pageId="4" pageNumber="5">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A63DFFB6AE676ABDFCCA8CF1" box="[825,872,1766,1790]" pageId="4" pageNumber="5">type</typeStatus>
|
||||
. Claw morphology (
|
||||
<figureCitation id="133D2A57A63DFFB6A91D6ABDFB208CF2" box="[1091,1154,1766,1790]" captionStart="Figure 7" captionStartId="11.[113,178,888,912]" captionTargetBox="[114,1458,144,860]" captionTargetId="figure-272@11.[114,1458,144,860]" captionTargetPageId="11" captionText="Figure 7. Claws of Diphascon greveni (PCM, topotypes): A, claws I; B, claws II; C, claws III; D, E, claws IV; F, claws III of a juvenile. Black arrowheads indicate internal bars. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243758" httpUri="https://zenodo.org/record/14243758/files/figure.png" pageId="4" pageNumber="5">Figs 7</figureCitation>
|
||||
,
|
||||
<figureCitation id="133D2A57A63DFFB6A9CF6ABDFB3C8CF1" box="[1169,1182,1766,1790]" captionStart="Figure 8" captionStartId="12.[129,194,1992,2016]" captionTargetBox="[129,1473,802,1964]" captionTargetId="figure-133@12.[129,1473,802,1964]" captionTargetPageId="12" captionText="Figure 8. Claws of Diphascon greveni (SEM, topotypes):A, claws I; B, claws III; C, D, claws IV. Black arrowheads indicate internal bars; white asterisks indicate internal pulvini. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243762" httpUri="https://zenodo.org/record/14243762/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
|
||||
) largely conforming to that described by
|
||||
<bibRefCitation id="EF974B23A63DFFB6AE976B5EFBD28D12" author="Dastych H" box="[969,1136,1797,1821]" pageId="4" pageNumber="5" pagination="377 - 436" refId="ref35540" refString="Dastych H. The Tardigrada from Antarctica with descriptions of several new species. Acta Zoologica Cracoviensia 1984; 27: 377 - 436." type="journal article" year="1984">Dastych (1984)</bibRefCitation>
|
||||
, but in rare cases the posterior claw bases were not broadened (almost all occurring asymmetrically on one leg,
|
||||
<figureCitation id="133D2A57A63DFFB6AEA56B1FFBE58D54" box="[1019,1095,1860,1884]" captionStart="Figure 7" captionStartId="11.[113,178,888,912]" captionTargetBox="[114,1458,144,860]" captionTargetId="figure-272@11.[114,1458,144,860]" captionTargetPageId="11" captionText="Figure 7. Claws of Diphascon greveni (PCM, topotypes): A, claws I; B, claws II; C, claws III; D, E, claws IV; F, claws III of a juvenile. Black arrowheads indicate internal bars. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243758" httpUri="https://zenodo.org/record/14243758/files/figure.png" pageId="4" pageNumber="5">Fig. 7D</figureCitation>
|
||||
) with respect to external claws I–III (
|
||||
<figureCitation id="133D2A57A63DFFB6AE1A6B38FC0D8D74" box="[836,943,1891,1915]" captionStart="Figure 7" captionStartId="11.[113,178,888,912]" captionTargetBox="[114,1458,144,860]" captionTargetId="figure-272@11.[114,1458,144,860]" captionTargetPageId="11" captionText="Figure 7. Claws of Diphascon greveni (PCM, topotypes): A, claws I; B, claws II; C, claws III; D, E, claws IV; F, claws III of a juvenile. Black arrowheads indicate internal bars. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243758" httpUri="https://zenodo.org/record/14243758/files/figure.png" pageId="4" pageNumber="5">Fig. 7A–C</figureCitation>
|
||||
). Some posterior claw bases so enlarged as to take almost a conical shape (
|
||||
<figureCitation id="133D2A57A63DFFB6A9696BD9FB228D95" box="[1079,1152,1922,1946]" captionStart="Figure 7" captionStartId="11.[113,178,888,912]" captionTargetBox="[114,1458,144,860]" captionTargetId="figure-272@11.[114,1458,144,860]" captionTargetPageId="11" captionText="Figure 7. Claws of Diphascon greveni (PCM, topotypes): A, claws I; B, claws II; C, claws III; D, E, claws IV; F, claws III of a juvenile. Black arrowheads indicate internal bars. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243758" httpUri="https://zenodo.org/record/14243758/files/figure.png" pageId="4" pageNumber="5">Fig. 7E</figureCitation>
|
||||
). Claws of juveniles (probably hatchlings) delicate and with less well-developed indentation (
|
||||
<figureCitation id="133D2A57A63CFFB7AD226B38FF608D75" box="[124,194,1891,1915]" captionStart="Figure 7" captionStartId="11.[113,178,888,912]" captionTargetBox="[114,1458,144,860]" captionTargetId="figure-272@11.[114,1458,144,860]" captionTargetPageId="11" captionText="Figure 7. Claws of Diphascon greveni (PCM, topotypes): A, claws I; B, claws II; C, claws III; D, E, claws IV; F, claws III of a juvenile. Black arrowheads indicate internal bars. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243758" httpUri="https://zenodo.org/record/14243758/files/figure.png" pageId="5" pageNumber="6">Fig. 7F</figureCitation>
|
||||
). Rarely, this indentation may also be developed on the internal claw bases (
|
||||
<figureCitation id="133D2A57A63CFFB7AC166BD9FE338D95" box="[328,401,1922,1946]" captionStart="Figure 8" captionStartId="12.[129,194,1992,2016]" captionTargetBox="[129,1473,802,1964]" captionTargetId="figure-133@12.[129,1473,802,1964]" captionTargetPageId="12" captionText="Figure 8. Claws of Diphascon greveni (SEM, topotypes):A, claws I; B, claws III; C, D, claws IV. Black arrowheads indicate internal bars; white asterisks indicate internal pulvini. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243762" httpUri="https://zenodo.org/record/14243762/files/figure.png" pageId="5" pageNumber="6">Fig. 8B</figureCitation>
|
||||
). Internal bars visible under SEM as longitudinal thickenings adjacent to the internal claw bases (
|
||||
<figureCitation id="133D2A57A63CFFB7AE6B6B38FCDD8D74" box="[821,895,1891,1915]" captionStart="Figure 8" captionStartId="12.[129,194,1992,2016]" captionTargetBox="[129,1473,802,1964]" captionTargetId="figure-133@12.[129,1473,802,1964]" captionTargetPageId="12" captionText="Figure 8. Claws of Diphascon greveni (SEM, topotypes):A, claws I; B, claws III; C, D, claws IV. Black arrowheads indicate internal bars; white asterisks indicate internal pulvini. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243762" httpUri="https://zenodo.org/record/14243762/files/figure.png" pageId="5" pageNumber="6">Fig. 8A</figureCitation>
|
||||
). Accessory points on claws IV more conspicuous and more widely divergent from the primary branches than on claws I–III (compare
|
||||
<figureCitation id="133D2A57A63CFFB7AE9B6BFAFB8B8DB6" box="[965,1065,1953,1977]" captionStart="Figure 8" captionStartId="12.[129,194,1992,2016]" captionTargetBox="[129,1473,802,1964]" captionTargetId="figure-133@12.[129,1473,802,1964]" captionTargetPageId="12" captionText="Figure 8. Claws of Diphascon greveni (SEM, topotypes):A, claws I; B, claws III; C, D, claws IV. Black arrowheads indicate internal bars; white asterisks indicate internal pulvini. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.14243762" httpUri="https://zenodo.org/record/14243762/files/figure.png" pageId="5" pageNumber="6">Fig. 8A, B</figureCitation>
|
||||
with C, D).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1337
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@ -0,0 +1,403 @@
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<mods:title id="C758A2092EEFDF406C7E840759739115">What is a ‘ strong’ synapomorphy? Redescriptions of Murray’s type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela)</mods:title>
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<subSubSection id="C31C6559A63FFFB4AC4F6A22FDDB8C9C" box="[273,633,1657,1684]" pageId="6" pageNumber="7" type="nomenclature">
|
||||
<paragraph id="8BB936D2A63FFFB4AC4F6A22FDDB8C9C" blockId="6.[273,633,1657,1684]" box="[273,633,1657,1684]" pageId="6" pageNumber="7">
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<heading id="D0F181BEA63FFFB4AC4F6A22FDDB8C9C" bold="true" box="[273,633,1657,1684]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
|
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<taxonomicName id="4C064D51A63FFFB4AC4F6A22FDDB8C9C" ID-CoL="9WNK9" authority="(Richters, 1911)" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[273,633,1657,1684]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63FFFB4AC4F6A22FDDB8C9C" bold="true" box="[273,633,1657,1684]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B972EAC0A63FFFB4AC4F6A22FE678C9B" bold="true" box="[273,453,1657,1684]" italics="true" pageId="6" pageNumber="7">Adropion belgicae</emphasis>
|
||||
(Richters, 1911)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63FFFB4ADF36AFDFCA78D1F" pageId="6" pageNumber="7" type="reference_group">
|
||||
<paragraph id="8BB936D2A63FFFB4ADF36AFDFDF28CEF" blockId="6.[173,733,1702,1760]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B972EAC0A63FFFB4ADF36AFDFDF28CEF" bold="true" pageId="6" pageNumber="7">
|
||||
<taxonomicName id="4C064D51A63FFFB4ADF36AFDFECF8CCE" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[173,365,1702,1729]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63FFFB4ADF36AFDFECF8CCE" bold="true" box="[173,365,1702,1729]" italics="true" pageId="6" pageNumber="7">Diphascon belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<emphasis id="B972EAC0A63FFFB4AC276AFCFE4E8CCF" bold="true" box="[377,492,1702,1729]" italics="true" pageId="6" pageNumber="7">terra typica</emphasis>
|
||||
: Advent Fjord, Hopen,
|
||||
<collectingCountry id="F3117642A63FFFB4AC676A9EFE3A8CD0" box="[313,408,1733,1759]" name="Svalbard and Jan Mayen" pageId="6" pageNumber="7">Svalbard</collectingCountry>
|
||||
; Richters (1911).
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A63FFFB4ADDA6AACFCA78D1F" blockId="6.[132,773,1782,1809]" box="[132,773,1782,1809]" pageId="6" pageNumber="7">
|
||||
<heading id="D0F181BEA63FFFB4ADDA6AACFCA78D1F" bold="true" box="[132,773,1782,1809]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
|
||||
<emphasis id="B972EAC0A63FFFB4ADDA6AACFCA78D1F" bold="true" box="[132,773,1782,1809]" pageId="6" pageNumber="7">
|
||||
<taxonomicName id="4C064D51A63FFFB4ADDA6AACFF488D1E" ID-CoL="7T83R" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[132,234,1782,1809]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63FFFB4ADDA6AACFF488D1E" bold="true" box="[132,234,1782,1809]" italics="true" pageId="6" pageNumber="7">D. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
; Albert I Land,Atomfjella,
|
||||
<collectingCountry id="F3117642A63FFFB4ACA56AADFDF58D1F" box="[507,599,1782,1808]" name="Svalbard and Jan Mayen" pageId="6" pageNumber="7">Svalbard</collectingCountry>
|
||||
;
|
||||
<bibRefCitation id="EF974B23A63FFFB4AF006AADFCA28D1F" author="Dastych H" box="[606,768,1782,1808]" pageId="6" pageNumber="7" pagination="169 - 214" refId="ref35565" refString="Dastych H. West Spitsbergen Tardigrada. Acta Zoologica Cracoviensia 1985; 28: 169 - 214." type="journal article" year="1985">Dastych(1985)</bibRefCitation>
|
||||
.
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63FFFB4ADDF6B69FA1D8C4D" pageId="6" pageNumber="7" type="materials_examined">
|
||||
<paragraph id="8BB936D2A63FFFB4ADDF6B69FE4B8D65" blockId="6.[129,777,1842,1898]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B972EAC0A63FFFB4ADDF6B69FEEB8D45" box="[129,329,1842,1866]" italics="true" pageId="6" pageNumber="7">Material examined:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA63FFFB4AC096B69FE4B8D65" pageId="6" pageNumber="7" specimenCount="21">
|
||||
<specimenCount id="9D00FD5BA63FFFB4AC096B69FE528D45" box="[343,496,1842,1866]" pageId="6" pageNumber="7">21 individuals</specimenCount>
|
||||
in total (for details, see Supporting Information,
|
||||
<tableCitation id="C6840369A63FFFB4ACDF6B0AFE7B8D65" box="[385,473,1873,1898]" captionStart="Table 1" captionStartId="8.[129,184,143,167]" captionTargetPageId="8" captionText="Table 1. Measurements (in micrometres) of selected morphological structures of topotypic specimens (AQ.149) of Diphascon greveni Dastych, 1984 from King George Island (Antarctica) mounted in Hoyer’s medium.In this and subsequent tables, N is the number of specimens/ structures measured; ‘Range’ refers to the smallest and the largest structure among all measured specimens; and SD is the standard deviation." pageId="6" pageNumber="7">Table S1</tableCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A63FFFB4ADDF6BDCFA1D8C4D" blockId="6.[129,777,1925,1982]" lastBlockId="6.[825,1474,1484,1602]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B972EAC0A63FFFB4ADDF6BDCFEFD8D91" box="[129,351,1926,1950]" italics="true" pageId="6" pageNumber="7">Comparative material:</emphasis>
|
||||
<materialsCitation id="3B6E3C8FA63FFFB4AC366BDDFC138C0C" collectingDate="2014-11-01" collectorName="Blagden" country="United Kingdom" elevation="451" latitude="57.3875" location="Hill of Foudland" longLatPrecision="17" longitude="-2.6608334" pageId="6" pageNumber="7" specimenCount="3">
|
||||
<collectingCountry id="F3117642A63FFFB4AC366BDDFE668D91" box="[360,452,1926,1950]" name="United Kingdom" pageId="6" pageNumber="7">Scotland</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A63FFFB4AC916BDDFDDE8D91" LSID="urn:lsid:plazi:treatment:03AF87C4A63FFFB5AC4F6A22FA91897C:8ED96009A63FFFB4AC916BDDFDDE8D91" box="[463,636,1926,1950]" country="United Kingdom" latitude="57.3875" longLatPrecision="17" longitude="-2.6608334" name="Hill of Foudland" pageId="6" pageNumber="7">Hill of Foudland</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A63FFFB4AFD56BDDFCA78D91" box="[651,773,1925,1950]" degrees="57" direction="north" minutes="23" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="15" value="57.3875">
|
||||
57°23
|
||||
<emphasis id="B972EAC0A63FFFB4AF966BDEFD6C8D91" box="[712,718,1925,1950]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||||
15″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A63FFFB4ADDF6BFDFF568DB2" box="[129,244,1956,1982]" degrees="2" direction="west" minutes="39" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="39" value="-2.6608334">
|
||||
2°39
|
||||
<emphasis id="B972EAC0A63FFFB4ADEE6BFFFF148DB2" box="[176,182,1956,1981]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||||
39″W
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A63FFFB4AC5A6BFEFE248DB2" box="[260,390,1957,1981]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="6" pageNumber="7" unit="m" value="451.0">
|
||||
<quantity id="4CFE9B37A63FFFB4AC5A6BFEFEEE8DB2" box="[260,332,1957,1981]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="6" pageNumber="7" unit="m" value="451.0">451 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), old slate quarry, lichen (
|
||||
<taxonomicName id="4C064D51A63FFFB4AFF16BFEFC058FEB" authority="P. Browne" authorityName="P. Browne" box="[687,935,1484,1981]" class="Lecanoromycetes" family="Cladoniaceae" genus="Cladonia" kingdom="Fungi" order="Lecanorales" pageId="6" pageNumber="7" phylum="Ascomycota" rank="genus">
|
||||
<emphasis id="B972EAC0A63FFFB4AFF16BFEFCAB8DB2" box="[687,777,1957,1981]" italics="true" pageId="6" pageNumber="7">Cladonia</emphasis>
|
||||
P. Browne
|
||||
</taxonomicName>
|
||||
) from soil,
|
||||
<date id="FFB81012A63FFFB4A97B6997FB458FEB" box="[1061,1255,1484,1508]" pageId="6" pageNumber="7" value="2014-11-01">
|
||||
<collectingDate id="EFFCE9FAA63FFFB4A97B6997FB458FEB" box="[1061,1255,1484,1508]" pageId="6" pageNumber="7" value="2014-11-01">1 November 2014</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A63FFFB4A9AD6997FAE48FEB" box="[1267,1350,1484,1508]" pageId="6" pageNumber="7">Blagden</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA63FFFB4A8D46997FC088C0C" count="3" pageId="6" pageNumber="7" type="generic">three specimens</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B6E3C8FA63FFFB4AE9B69B0FA198C4D" collectingDate="2014-09-03" collectorName="Morek" country="Poland" elevation="9" latitude="54.71472" location="Slowinski National Park" longLatPrecision="17" longitude="17.223333" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<collectingCountry id="F3117642A63FFFB4AE9B69B0FBAD8C0C" box="[965,1039,1515,1539]" name="Poland" pageId="6" pageNumber="7">Poland</collectingCountry>
|
||||
,
|
||||
<location id="8ED96009A63FFFB4A97C69B0FA898C0C" LSID="urn:lsid:plazi:treatment:03AF87C4A63FFFB5AC4F6A22FA91897C:8ED96009A63FFFB4A97C69B0FA898C0C" box="[1058,1323,1515,1539]" country="Poland" latitude="54.71472" longLatPrecision="17" longitude="17.223333" name="Slowinski National Park" pageId="6" pageNumber="7">Słowiński National Park</location>
|
||||
(
|
||||
<geoCoordinate id="EE325015A63FFFB4A81F69B0FA1C8C0C" box="[1345,1470,1514,1539]" degrees="54" direction="north" minutes="42" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="53" value="54.71472">
|
||||
54°42
|
||||
<emphasis id="B972EAC0A63FFFB4A8DF69B1FA258C0C" box="[1409,1415,1514,1539]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||||
53″N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE325015A63FFFB4AE676A50FC128C2D" box="[825,944,1545,1571]" degrees="17" direction="east" minutes="13" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="24" value="17.223333">
|
||||
17°13
|
||||
<emphasis id="B972EAC0A63FFFB4AE2B6A52FCD98C2D" box="[885,891,1545,1570]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||||
24″E
|
||||
</geoCoordinate>
|
||||
;
|
||||
<elevation id="002BD1E1A63FFFB4AEE46A50FBB68C2C" box="[954,1044,1547,1571]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="m" value="9.0">
|
||||
<quantity id="4CFE9B37A63FFFB4AEE46A50FC438C2D" box="[954,993,1547,1571]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="m" value="9.0">9 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
), the Scots pine (
|
||||
<taxonomicName id="4C064D51A63FFFB4A9966A57FAF58C2C" box="[1224,1367,1547,1571]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="sylvestris">
|
||||
<emphasis id="B972EAC0A63FFFB4A9966A57FAF58C2C" box="[1224,1367,1547,1571]" italics="true" pageId="6" pageNumber="7">Pinus sylvestris</emphasis>
|
||||
</taxonomicName>
|
||||
L.) forest, moss from soil,
|
||||
<date id="FFB81012A63FFFB4AE876A71FB348C4D" box="[985,1174,1578,1602]" pageId="6" pageNumber="7" value="2014-09-03">
|
||||
<collectingDate id="EFFCE9FAA63FFFB4AE876A71FB348C4D" box="[985,1174,1578,1602]" pageId="6" pageNumber="7" value="2014-09-03">3 September 2014</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26F35304A63FFFB4A9C16A71FB418C4D" box="[1183,1251,1578,1602]" pageId="6" pageNumber="7">Morek</collectorName>
|
||||
coll. (
|
||||
<specimenCount id="9D00FD5BA63FFFB4A87E6A71FA168C4D" box="[1312,1460,1578,1602]" count="1" pageId="6" pageNumber="7" type="generic">one specimen</specimenCount>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63FFFB5AE676A37FDB88DB8" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
|
||||
<paragraph id="8BB936D2A63FFFB5AE676A37FE968EC2" blockId="6.[824,1476,1644,1982]" lastBlockId="7.[113,763,1079,1448]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||||
<emphasis id="B972EAC0A63FFFB4AE676A37FBB08C8B" box="[825,1042,1644,1668]" italics="true" pageId="6" pageNumber="7">Amended description:</emphasis>
|
||||
Body medium-sized to large (
|
||||
<tableCitation id="C6840369A63FFFB4A83F6A37FA138C8B" box="[1377,1457,1644,1668]" captionStart="Table 2" captionStartId="13.[115,170,143,167]" captionTargetPageId="13" captionText="Table 2.Measurements (in micrometres) of selected morphological structures of topotypic specimens (NO.025) of Adropion belgicae (Richters, 1911) from Svalbard mounted in Hoyer’s medium." pageId="6" pageNumber="7">Table 2</tableCitation>
|
||||
), elongated and massive, of similar width over its entire length. Smaller specimens white, but larger brownish at least in the caudal part, often extending to the whole body (
|
||||
<figureCitation id="133D2A57A63FFFB4A8456A91FAC08CED" box="[1307,1378,1738,1762]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="6" pageNumber="7">Fig. 9A</figureCitation>
|
||||
). Cuticle smooth, without pores, wrinkling, or granulation. Head blunt. Cribriform areas not visible under PCM. Legs short and stout. Eyes absent in live animals. Buccopharyngeal apparatus of the
|
||||
<taxonomicName id="4C064D51A63FFFB4AE676B1CFC378D50" authorityName="Pilato" authorityYear="1987" box="[825,917,1863,1887]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="7" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63FFFB4AE676B1CFC378D50" box="[825,917,1863,1887]" italics="true" pageId="6" pageNumber="7">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A63FFFB4AEC56B13FC6B8D6F" box="[923,969,1864,1888]" pageId="6" pageNumber="7">type</typeStatus>
|
||||
(
|
||||
<figureCitation id="133D2A57A63FFFB4AE876B1CFB828D6F" box="[985,1056,1863,1888]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="6" pageNumber="7">Fig. 9B</figureCitation>
|
||||
). The OCA not visible under PCM (
|
||||
<figureCitation id="133D2A57A63FFFB4A8C26B1CFCF78D70" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="6" pageNumber="7">Fig. 9B</figureCitation>
|
||||
). Furcae of the
|
||||
<taxonomicName id="4C064D51A63FFFB4A9576B3CFBCB8D70" authorityName="Ehrenberg" authorityYear="1848" box="[1033,1129,1895,1919]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="7" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63FFFB4A9576B3CFBCB8D70" box="[1033,1129,1895,1919]" italics="true" pageId="6" pageNumber="7">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A63FFFB4A92D6B3CFB018D70" box="[1139,1187,1895,1919]" pageId="6" pageNumber="7">type</typeStatus>
|
||||
. Pharyngeal apophyses absent (
|
||||
<figureCitation id="133D2A57A63FFFB4AE266BDDFC618D91" box="[888,963,1926,1950]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="6" pageNumber="7">Fig. 9B</figureCitation>
|
||||
). Pharynx oval and broad. Macroplacoid length sequence 1 <2; all macroplacoids elongated and thin, with irregular, deeply serrated margins (
|
||||
<figureCitation id="133D2A57A63EFFB5ACAF686CFDC38E40" box="[497,609,1079,1103]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9B–D</figureCitation>
|
||||
). The second macroplacoid 2–2.5 times longer than the first, with a clearly swollen terminal part separated from the rest of the structure by a constriction (
|
||||
<figureCitation id="133D2A57A63EFFB5AC5168CEFEDB8EA2" box="[271,377,1173,1197]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9C, D</figureCitation>
|
||||
). Microplacoid small and of irregular shape (
|
||||
<figureCitation id="133D2A57A63EFFB5ADE268EEFE868EC2" box="[188,292,1205,1229]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9B–D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF79665AA63EFFB5AD2F6FC6FEB289E2" ID-DOI="http://doi.org/10.5281/zenodo.14243750" ID-Zenodo-Dep="14243750" httpUri="https://zenodo.org/record/14243750/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,925,949]" targetBox="[119,756,145,895]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BB936D2A63EFFB5AD2F6FC6FEB289E2" blockId="7.[113,755,925,1005]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B972EAC0A63EFFB5AD2F6FC6FF6B89BA" bold="true" box="[113,201,925,949]" pageId="7" pageNumber="8">Figure 4.</emphasis>
|
||||
The
|
||||
<taxonomicName id="4C064D51A63EFFB5ADA96FC6FE3589BA" baseAuthorityName="Marcus" baseAuthorityYear="1936" box="[247,407,925,949]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="pingue">
|
||||
<emphasis id="B972EAC0A63EFFB5ADA96FC6FE3589BA" box="[247,407,925,949]" italics="true" pageId="7" pageNumber="8">Diphascon pingue</emphasis>
|
||||
</taxonomicName>
|
||||
group, morphology of pharyngeal structures (SEM): A,
|
||||
<taxonomicName id="4C064D51A63EFFB5AC606FE2FE7C89DE" baseAuthorityName="Marcus" baseAuthorityYear="1936" box="[318,478,953,977]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="pingue">
|
||||
<emphasis id="B972EAC0A63EFFB5AC606FE2FE7C89DE" box="[318,478,953,977]" italics="true" pageId="7" pageNumber="8">Diphascon pingue</emphasis>
|
||||
</taxonomicName>
|
||||
; B, C,
|
||||
<taxonomicName id="4C064D51A63EFFB5AF436FE2FD4C89DE" authorityName="Pilato & Binda" authorityYear="1997" box="[541,750,953,977]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="pinguiforme">
|
||||
<emphasis id="B972EAC0A63EFFB5AF436FE2FD4C89DE" box="[541,750,953,977]" italics="true" pageId="7" pageNumber="8">Diphascon pinguiforme</emphasis>
|
||||
</taxonomicName>
|
||||
. Scale bars: 2 μm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB936D2A63EFFB5ADD3688FFEE58FA7" blockId="7.[113,763,1079,1448]" pageId="7" pageNumber="8">
|
||||
Claws of the
|
||||
<taxonomicName id="4C064D51A63EFFB5AC4B688FFED78EE3" authorityName="Ehrenberg" authorityYear="1848" box="[277,373,1236,1260]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5AC4B688FFED78EE3" box="[277,373,1236,1260]" italics="true" pageId="7" pageNumber="8">Hypsibius</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="54BD8870A63EFFB5AC24688EFE0B8EE2" box="[378,425,1237,1261]" pageId="7" pageNumber="8">type</typeStatus>
|
||||
, large and robust, with divergent accessory points. Claw septa usually clearly visible, dividing the claw into three portions (
|
||||
<figureCitation id="133D2A57A63EFFB5AC2A6948FE7E8F24" box="[372,476,1299,1323]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9E–G</figureCitation>
|
||||
). Pseudolunulae weakly delineated and only sometimes visible at the base of the internal and anterior claws (
|
||||
<figureCitation id="133D2A57A63EFFB5AC12690AFE198F66" box="[332,443,1361,1385]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9F, G</figureCitation>
|
||||
). Cuticular bars absent; only slightly thickened edges of pulvini may be visible on the internal side of legs (
|
||||
<figureCitation id="133D2A57A63EFFB5ADAE69CBFE958FA7" box="[240,311,1424,1448]" captionStart="Figure 9" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[196,1406,144,1809]" captionTargetId="figure-5@14.[196,1406,144,1809]" captionTargetPageId="14" captionText="Figure 9. Adropion belgicae (Richters, 1911) (topotypes): A, specimen in toto (PCM, ventral view); B, buccopharyngeal apparatus in toto (PCM); C, pharynx in dorsal view (SEM; note irregular margins of placoids); D, pharynx in lateral view (SEM); E, claws II (PCM; white arrowhead indicates the external pulvinus); F, claws II seen at a different angle (PCM); G, claws IV (PCM). Scale bars are in micrometres." figureDoi="http://doi.org/10.5281/zenodo.14243766" httpUri="https://zenodo.org/record/14243766/files/figure.png" pageId="7" pageNumber="8">Fig. 9E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB936D2A63EFFB5AD2F6992FDB88DB8" blockId="7.[113,763,1481,1975]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B972EAC0A63EFFB5AD2F6992FF6D8FEE" box="[113,207,1481,1505]" italics="true" pageId="7" pageNumber="8">Remarks:</emphasis>
|
||||
A more detailed description of
|
||||
<taxonomicName id="4C064D51A63EFFB5AF496991FD3F8FEE" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[535,669,1481,1505]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AF496991FD8D8FEE" box="[535,559,1482,1505]" italics="true" pageId="7" pageNumber="8">A.</emphasis>
|
||||
cf.
|
||||
<emphasis id="B972EAC0A63EFFB5AF0C6992FD3F8FEE" box="[594,669,1481,1505]" italics="true" pageId="7" pageNumber="8">belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
was provided by
|
||||
<bibRefCitation id="EF974B23A63EFFB5AD8F69B2FED18C0E" author="Bernard EC" box="[209,371,1513,1537]" pageId="7" pageNumber="8" pagination="476 - 82" refId="ref34843" refString="Bernard EC. A new species of Hexapodibius from North America, with a redescription of Diphascon belgicae (Tardigrada). Transactions of the American Microscopical Society 1977; 96: 476 - 82. https: // doi. org / 10.2307 / 3225665" type="journal article" year="1977">Bernard (1977)</bibRefCitation>
|
||||
. Based on claw (stumpy and with distinct septa) and buccopharyngeal apparatus morphology, it is obvious that the specimens collected by Bernard in Michigan represented a similar species, but not
|
||||
<taxonomicName id="4C064D51A63EFFB5AF5C6A1CFDCE8C50" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[514,620,1607,1631]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AF5C6A1CFDCE8C50" box="[514,620,1607,1631]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A24157BBA63EFFB5AF2B6A1CFD328C51" box="[629,656,1607,1630]" pageId="7" pageNumber="8" sensu="stricto">
|
||||
<emphasis id="B972EAC0A63EFFB5AF2B6A1CFD288C51" box="[629,650,1607,1630]" italics="true" pageId="7" pageNumber="8">s.s</emphasis>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
They had three well-developed
|
||||
<typeStatus id="54BD8870A63EFFB5AC0F6A3CFE2A8C70" box="[337,392,1639,1663]" pageId="7" pageNumber="8">types</typeStatus>
|
||||
of cuticular bars (internal, median, and anterior), as opposed to only pulvini present in
|
||||
<taxonomicName id="4C064D51A63EFFB5AFD36ADDFD578C92" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[653,757,1669,1693]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AFD36ADDFD578C92" box="[653,757,1669,1693]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
. Thus, the American specimens must have belonged to a new species. Likewise,
|
||||
<taxonomicName id="4C064D51A63EFFB5AC686A9EFD388CD3" authority="Ito, 1991" authorityName="Ito" authorityYear="1991" box="[310,666,1732,1756]" class="Eutardigrada" family="Hypsibiidae" genus="Fujiscon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="diphasconiellum">
|
||||
<emphasis id="B972EAC0A63EFFB5AC686A9EFD8C8CD3" box="[310,558,1732,1756]" italics="true" pageId="7" pageNumber="8">Fujiscon diphasconiellum</emphasis>
|
||||
<bibRefCitation id="EF974B23A63EFFB5AF646A9EFD388CD3" author="Ito M" box="[570,666,1732,1756]" pageId="7" pageNumber="8" pagination="30 - 43" refId="ref36735" refString="Ito M. Taxonomic study on the Eutardigrada from the northern slope of Mt. Fuji, Central Japan. Proceedings of the Japanese Society of Systematic Zoology 1991; 45: 30 - 43." type="journal article" year="1991">Ito, 1991</bibRefCitation>
|
||||
</taxonomicName>
|
||||
is a different species owing to the dissimilar arrangement of bars; its current status should be
|
||||
<taxonomicName id="4C064D51A63EFFB5AC366B58FDC48D14" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[360,614,1795,1819]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A63EFFB5AC366B58FDC48D14" box="[360,614,1795,1819]" italics="true" pageId="7" pageNumber="8">Adropion diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A63EFFB5AF356B58FF6F8D35" bold="true" pageId="7" pageNumber="8">
|
||||
<taxonomicNameLabel id="A24157BBA63EFFB5AF356B58FF6F8D35" pageId="7" pageNumber="8">comb. nov. et stat. rev.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, by withdrawing it from an incorrect synonymy with
|
||||
<taxonomicName id="4C064D51A63EFFB5AD2F6B19FEDB8D55" authority="(Maucci 1996)" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[113,377,1857,1882]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AD2F6B19FF7A8D56" box="[113,216,1857,1881]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A63EFFB5ADB76B1AFECC8D56" author="Maucci W" box="[233,366,1857,1881]" pageId="7" pageNumber="8" pagination="185 - 204" refId="ref37799" refString="Maucci W. Tardigrada of the Arctic tundra with descriptions of two new species. Zoological Journal of the Linnean Society 1996; 116: 185 - 204. https: // doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02343. x" type="journal article" year="1996">Maucci 1996</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and pending a redescription.
|
||||
<bibRefCitation id="EF974B23A63EFFB5AFF86B1AFF198D76" author="Dastych H" pageId="7" pageNumber="8" pagination="1 - 255" refId="ref35583" refString="Dastych H. The Tardigrada of Poland. Monografie Fauny Polski 1988; 16: 1 - 255." type="journal article" year="1988">Dastych (1988)</bibRefCitation>
|
||||
was the first to mention the brown colour in
|
||||
<taxonomicName id="4C064D51A63EFFB5AFCD6B3AFD598D76" baseAuthorityName="Maucci" baseAuthorityYear="1996" box="[659,763,1889,1913]" class="Eutardigrada" family="Hypsibiidae" genus="Diphascon" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AFCD6B3AFD598D76" box="[659,763,1889,1913]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
he collected in
|
||||
<collectingCountry id="F3117642A63EFFB5AC536BDBFEF58D97" box="[269,343,1920,1944]" name="Poland" pageId="7" pageNumber="8">Poland</collectingCountry>
|
||||
. He also illustrated the conspicuous claw septa that are characteristic of the species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63EFFB5AE746CCBFBE28AE8" pageId="7" pageNumber="8" type="etymology">
|
||||
<paragraph id="8BB936D2A63EFFB5AE746CCBFBE28AE8" blockId="7.[810,1460,144,231]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B972EAC0A63EFFB5AE746CCBFC3B8AA7" box="[810,921,144,168]" italics="true" pageId="7" pageNumber="8">Etymology:</emphasis>
|
||||
Named after the steamship
|
||||
<collectingCountry id="F3117642A63EFFB5A99C6CCBFAAC8AA7" box="[1218,1294,144,168]" name="Belgium" pageId="7" pageNumber="8">Belgica</collectingCountry>
|
||||
, used in expeditions to Antarctic (1897–1899) and Arctic (1905, 1907–1909). A noun in genitive singular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31C6559A63EFFB5AE746D53FA91897C" pageId="7" pageNumber="8" type="diagnosis">
|
||||
<paragraph id="8BB936D2A63EFFB5AE746D53FA91897C" blockId="7.[809,1461,264,883]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B972EAC0A63EFFB5AE746D53FC5E8B2F" box="[810,1020,264,288]" italics="true" pageId="7" pageNumber="8">Differential diagnosis:</emphasis>
|
||||
There are only two other
|
||||
<taxonomicName id="4C064D51A63EFFB5A9A96D53FA208B10" box="[1271,1410,264,288]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="undetermined">
|
||||
<emphasis id="B972EAC0A63EFFB5A9A96D53FAF18B2F" box="[1271,1363,264,288]" italics="true" pageId="7" pageNumber="8">Adropion</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
with two macroplacoids in the pharynx:
|
||||
<taxonomicName id="4C064D51A63EFFB5A9F66D73FAC48B30" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[1192,1382,295,319]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A63EFFB5A9F66D73FAC48B30" box="[1192,1382,295,319]" italics="true" pageId="7" pageNumber="8">A. diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A63EFFB5A82E6D7CFC768B50" bold="true" pageId="7" pageNumber="8">
|
||||
<taxonomicNameLabel id="A24157BBA63EFFB5A82E6D7CFC768B50" pageId="7" pageNumber="8">comb. nov. et stat. rev.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C064D51A63EFFB5A9576D1CFAEC8B50" authority="(Sun et al., 2014)" baseAuthorityName="Sun" baseAuthorityYear="2014" box="[1033,1358,327,351]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="gani">
|
||||
<emphasis id="B972EAC0A63EFFB5A9576D1CFB378B51" box="[1033,1173,327,351]" italics="true" pageId="7" pageNumber="8">Adropion gani</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A63EFFB5A9FB6D1CFAE08B50" author="Sun X & Li X & Feng W" box="[1189,1346,327,351]" pageId="7" pageNumber="8" pagination="323 - 8" refId="ref39797" refString="Sun X, Li X, Feng W. Two new species of Tardigrada (Echiniscidae, Hypsibiidae) fromChina. ProceedingsoftheBiologicalSocietyofWashington 2014; 126: 323 - 8. https: // doi. org / 10.2988 / 0006 - 324 X- 126.4.323" type="journal article" year="2014">
|
||||
Sun
|
||||
<emphasis id="B972EAC0A63EFFB5A98D6D1CFAA18B50" box="[1235,1283,327,351]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
, 2014
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(a poorly diagnosed species with macroplacoids arranged rhomboidally; see below for the composition of the genus).
|
||||
<taxonomicName id="4C064D51A63EFFB5A8586DDEFA178B92" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1286,1461,389,413]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5A8586DDEFA178B92" box="[1286,1461,389,413]" italics="true" pageId="7" pageNumber="8">Adropion belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
has no cuticular bars, which distinguishes it sufficiently from
|
||||
<taxonomicName id="4C064D51A63EFFB5AE746D9FFC438BD3" authorityName="Gąsiorek & Blagden & Morek & Michalczyk" authorityYear="2024" baseAuthorityName="Gąsiorek & Blagden & Morek & Michalczyk" baseAuthorityYear="1991" box="[810,993,452,476]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="diphasconiellum" status="comb. nov. et stat. rev.">
|
||||
<emphasis id="B972EAC0A63EFFB5AE746D9FFC438BD3" box="[810,993,452,476]" italics="true" pageId="7" pageNumber="8">A. diphasconiellum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B972EAC0A63EFFB5AEBB6D9FFB6C8BD3" bold="true" box="[997,1230,452,476]" pageId="7" pageNumber="8">
|
||||
<taxonomicNameLabel id="A24157BBA63EFFB5AEBB6D9FFB6C8BD3" box="[997,1230,452,476]" pageId="7" pageNumber="8">comb. nov. et stat. rev.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, which possesses
|
||||
<specimenCount id="9D00FD5BA63EFFB5A8206D9FFCC28BF3" count="3" pageId="7" pageNumber="8" type="generic" typeStatus="types">three types</specimenCount>
|
||||
of bars (internal, median, and anterior). Morphometric data would probably also be useful in separation of the two species; however, such data are unavailable owing to the outdated description of the former taxon. Because of general similarities in the morphology of the buccopharyngeal apparatus,
|
||||
<taxonomicName id="4C064D51A63EFFB5AE746EDBFC2C8897" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[810,910,640,664]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5AE746EDBFC2C8897" box="[810,910,640,664]" italics="true" pageId="7" pageNumber="8">A.belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
canbemisidentifiedasamemberofthegenus
|
||||
<taxonomicName id="4C064D51A63EFFB5A8166EDAFC0688B8" authority="Pilato, 1987" authorityName="Pilato" authorityYear="1987" class="Eutardigrada" family="Hypsibiidae" genus="Mesocrista" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5A8166EDAFA128897" box="[1352,1456,641,664]" italics="true" pageId="7" pageNumber="8">Mesocrista</emphasis>
|
||||
Pilato, 1987
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EF974B23A63EFFB5AEEA6EC4FBDC88B8" author="Gasiorek P & Stec D & Morek W" box="[948,1150,671,695]" pageId="7" pageNumber="8" pagination="2803 - 28" refId="ref36118" refString="Gasiorek P, Stec D, Morek W et al. An integrative revision of Mesocrista Pilato, 1987 (Tardigrada: Eutardigrada: Hypsibiidae). Journal of Natural History 2016; 50: 2803 - 28. https: // doi. org / 10.1080 / 00222 933.2016. 1234654" type="journal article" year="2016">
|
||||
Gąsiorek
|
||||
<emphasis id="B972EAC0A63EFFB5A9486EFBFBE788B8" box="[1046,1093,671,695]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
). Nevertheless,
|
||||
<taxonomicName id="4C064D51A63EFFB5A87A6EFBFA2E88B8" authorityName="Pilato" authorityYear="1987" box="[1316,1420,672,695]" class="Eutardigrada" family="Hypsibiidae" genus="Mesocrista" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5A87A6EFBFA2E88B8" box="[1316,1420,672,695]" italics="true" pageId="7" pageNumber="8">Mesocrista</emphasis>
|
||||
</taxonomicName>
|
||||
has a buccal tube that is wider than in
|
||||
<taxonomicName id="4C064D51A63EFFB5A9DB6EE4FB4388D8" authorityName="Pilato" authorityYear="1987" box="[1157,1249,703,727]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5A9DB6EE4FB4388D8" box="[1157,1249,703,727]" italics="true" pageId="7" pageNumber="8">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
(the thinnest buccal tube measured in
|
||||
<taxonomicName id="4C064D51A63EFFB5AEB76E84FBF388F9" authorityName="Pilato" authorityYear="1987" box="[1001,1105,735,758]" class="Eutardigrada" family="Hypsibiidae" genus="Mesocrista" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5AEB76E84FBF388F9" box="[1001,1105,735,758]" italics="true" pageId="7" pageNumber="8">Mesocrista</emphasis>
|
||||
</taxonomicName>
|
||||
had an external diameter of ~3.5 μm, vs. ~2.5 μm at its widest in
|
||||
<taxonomicName id="4C064D51A63EFFB5A93D6EA5FB6B891A" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1123,1225,765,789]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5A93D6EA5FB6B891A" box="[1123,1225,765,789]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
), double annulation of the pharyngeal tube (single annulation in
|
||||
<taxonomicName id="4C064D51A63EFFB5A9816F46FA9B893A" authorityName="Pilato" authorityYear="1987" box="[1247,1337,797,821]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="genus">
|
||||
<emphasis id="B972EAC0A63EFFB5A9816F46FA9B893A" box="[1247,1337,797,821]" italics="true" pageId="7" pageNumber="8">Adropion</emphasis>
|
||||
</taxonomicName>
|
||||
), and
|
||||
<specimenCount id="9D00FD5BA63EFFB5A8206F46FCC1895B" count="3" pageId="7" pageNumber="8" type="generic" typeStatus="types">three types</specimenCount>
|
||||
of cuticular bars at the claw bases (typically only the edges of external pulvini faintly observable in
|
||||
<taxonomicName id="4C064D51A63EFFB5A9E56F07FA80897C" baseAuthorityName="Richters" baseAuthorityYear="1911" box="[1211,1314,859,883]" class="Eutardigrada" family="Hypsibiidae" genus="Adropion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="8" phylum="Tardigrada" rank="species" species="belgicae">
|
||||
<emphasis id="B972EAC0A63EFFB5A9E56F07FA80897C" box="[1211,1314,859,883]" italics="true" pageId="7" pageNumber="8">A. belgicae</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue