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<document id="64F12C9515CEFD58BF3CE56C74B1A3F3" ID-DOI="10.11646/phytotaxa.446.2.4" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727828871079" checkinUser="felipe" docAuthor="Chen, Junhao &amp; Wong, Khoon Meng" docDate="2020" docId="03AF8798FFC4FF9BFF6EFEE0FD3E11E1" docLanguage="en" docName="phytotaxa.446.2.4.pdf" docOrigin="Phytotaxa 446 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.446.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Timonius hirsutus Merrill 1917" docType="treatment" docVersion="1" lastPageNumber="117" masterDocId="FF96FFE0FFC7FF98FFE6FFB2FFA91269" masterDocTitle="Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae)" masterLastPageNumber="120" masterPageNumber="114" pageNumber="117" updateTime="1727834409287" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7E1B5476490F664E5F2D9DEA4DDED2A2">Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae)</mods:title>
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<mods:title id="667D8C9E5DE93CD1799C22AB86A5081A">Phytotaxa</mods:title>
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<mods:date id="8C32D4A0FB41E0463D408545F0447CDA">2020</mods:date>
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<treatment id="03AF8798FFC4FF9BFF6EFEE0FD3E11E1" LSID="urn:lsid:plazi:treatment:03AF8798FFC4FF9BFF6EFEE0FD3E11E1" httpUri="http://treatment.plazi.org/id/03AF8798FFC4FF9BFF6EFEE0FD3E11E1" lastPageNumber="117" pageId="3" pageNumber="117">
<subSubSection id="C31C6505FFC4FF9BFF6EFEE0FDE21305" box="[136,587,338,365]" pageId="3" pageNumber="117" type="nomenclature">
<paragraph id="8BB9368EFFC4FF9BFF6EFEE0FDE21305" blockId="3.[136,587,338,365]" box="[136,587,338,365]" pageId="3" pageNumber="117">
<heading id="D0F181E2FFC4FF9BFF6EFEE0FDE21305" box="[136,587,338,365]" fontSize="11" level="2" pageId="3" pageNumber="117" reason="3">
<treatmentCitationGroup id="AB1611A0FFC4FF9BFF6EFEE0FDE21305" box="[136,587,338,365]" pageId="3" pageNumber="117">
<treatmentCitation id="0AA7109FFFC4FF9BFF6EFEE0FDE21305" author="Merrill, E. D." box="[136,587,338,365]" page=")" pageId="3" pageNumber="117" year="1917">
<taxonomicName id="4C064D0DFFC4FF9BFF6EFEE0FDE21305" ID-CoL="56ZHS" authority="Merrill (1917 b: 242)" authorityName="Merrill" authorityPageNumber="242" authorityYear="1917" baseAuthorityName="Elmer" box="[136,587,338,365]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B972EA9CFFC4FF9BFF6EFEE0FEF51304" bold="true" box="[136,348,338,365]" italics="true" pageId="3" pageNumber="117">Timonius hirsutus</emphasis>
<bibRefCitation id="EF974B7FFFC4FF9BFE84FEE0FDE21305" author="Merrill, E. D." box="[354,587,338,364]" pageId="3" pageNumber="117" pagination="189 - 247" refId="ref5492" refString="Merrill, E. D. (1917 b) Alabastra Borneensia. Journal of the Straits Branch of the Royal Asiatic Society 77: 189 - 247." type="journal article" year="1917">Merrill (1917b: 242)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C6505FFC4FF9BFF6EFE2CFD3E11E1" pageId="3" pageNumber="117" type="materials_examined">
<paragraph id="8BB9368EFFC4FF9BFF6EFE2CFEFA13B1" blockId="3.[136,1451,414,472]" pageId="3" pageNumber="117">
<emphasis id="B972EA9CFFC4FF9BFF6EFE2CFF6B13DA" bold="true" box="[136,194,414,435]" pageId="3" pageNumber="117">Type:</emphasis>
<materialsCitation id="3B6E3CD3FFC4FF9BFF3CFE2CFEE613B1" collectingDate="1895-04-20" country="Malaysia" location="Baram" municipality="Miri" pageId="3" pageNumber="117" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
BORNEO.
<collectingCountry id="F311761EFFC4FF9BFEAEFE2CFE0B13DD" box="[328,418,414,436]" name="Malaysia" pageId="3" pageNumber="117">Malaysia</collectingCountry>
, Sarawak,
<location id="8ED96055FFC4FF9BFDEFFE2CFDE213DA" LSID="urn:lsid:plazi:treatment:03AF8798FFC4FF9BFF6EFEE0FD3E11E1:8ED96055FFC4FF9BFDEFFE2CFDE213DA" box="[521,587,414,435]" country="Malaysia" municipality="Miri" name="Baram" pageId="3" pageNumber="117">Baram</location>
,
<collectingMunicipality id="6BDDACF4FFC4FF9BFDB2FE2CFD2813DD" box="[596,641,414,436]" pageId="3" pageNumber="117">Miri</collectingMunicipality>
,
<date id="FFB8104EFFC4FF9BFD6CFE2CFCA913DD" box="[650,768,414,436]" pageId="3" pageNumber="117" value="1895-04-20">
<collectingDate id="EFFCE9A6FFC4FF9BFD6CFE2CFCA913DD" box="[650,768,414,436]" pageId="3" pageNumber="117" value="1895-04-20">20 Apr 1895</collectingDate>
</date>
(
<specimenCount id="9D00FD07FFC4FF9BFCEBFE2CFC8813DA" box="[781,801,414,435]" count="1" pageId="3" pageNumber="117" type="male"></specimenCount>
),
<emphasis id="B972EA9CFFC4FF9BFCD4FE2DFC2513DA" box="[818,908,414,436]" italics="true" pageId="3" pageNumber="117">Hose 651</emphasis>
(
<typeStatus id="54BD882CFFC4FF9BFC7EFE2CFC5B13DD" box="[920,1010,414,436]" pageId="3" pageNumber="117" type="lectotype">lectotype</typeStatus>
A [A00312908], designated here;
<typeStatus id="54BD882CFFC4FF9BFAD1FE2CFA0213DD" box="[1335,1451,414,436]" pageId="3" pageNumber="117" type="isolectotype">isolectotype</typeStatus>
L [L0001476])
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9368EFFC4FF9BFF6EFDB4FE4810D9" blockId="3.[136,1452,518,905]" pageId="3" pageNumber="117">
<emphasis id="B972EA9CFFC4FF9BFF6EFDB4FF7D1049" bold="true" box="[136,212,518,544]" pageId="3" pageNumber="117">Notes:</emphasis>
—Although Merrill mentioned a single collection (
<emphasis id="B972EA9CFFC4FF9BFCE5FDB5FCC21049" box="[771,875,518,544]" italics="true" pageId="3" pageNumber="117">Hose 651</emphasis>
) in his protologue, a few duplicates of this collection exist, and apparently there was also a PNH specimen that must have been destroyed during the war. The
<typeStatus id="54BD882CFFC4FF9BFACCFD98FA25102D" box="[1322,1420,554,580]" pageId="3" pageNumber="117" type="holotype">holotype</typeStatus>
of Merrills Bornean species was deposited in PNH (
<bibRefCitation id="EF974B7FFFC4FF9BFD5AFDFCFCF71001" author="Merrill, E. D." box="[700,862,590,616]" pageId="3" pageNumber="117" pagination="75 - 117" refId="ref5456" refString="Merrill, E. D. (1917 a) Contributions to our knowledge of the flora of Borneo. Journal of the Straits Branch of the Royal Asiatic Society 76: 75 - 117." type="journal article" year="1917">Merrill 1917a</bibRefCitation>
).
<bibRefCitation id="EF974B7FFFC4FF9BFC94FDFCFBE91000" author="Chen, J. &amp; Tan, H. T. W. &amp; Wong, K. M." box="[882,1088,590,617]" pageId="3" pageNumber="117" pagination="420 - 430" refId="ref4802" refString="Chen, J., Tan, H. T. W. &amp; Wong, K. M. (2015) A revision of Timonius (Rubiaceae) in Kinabalu Park, Borneo, with notes on typification and species distinction. Plant Ecology and Evolution 148: 420 - 430. https: // doi. org / 10.5091 / plecevo. 2015.1073" type="journal article" year="2015">
Chen
<emphasis id="B972EA9CFFC4FF9BFC52FDFDFC4F1001" box="[948,998,590,616]" italics="true" pageId="3" pageNumber="117">et al</emphasis>
. (2015)
</bibRefCitation>
stated that the
<typeStatus id="54BD882CFFC4FF9BFB0AFDFCFAE71001" box="[1260,1358,590,616]" pageId="3" pageNumber="117" type="holotype">holotype</typeStatus>
in PNH was destroyed, but did not select a
<typeStatus id="54BD882CFFC4FF9BFDC7FDC0FD2110E5" box="[545,648,626,652]" pageId="3" pageNumber="117" type="lectotype">lectotype</typeStatus>
among the extant type material. Therefore, we here designate a well-preserved A sheet as
<typeStatus id="54BD882CFFC4FF9BFE95FD24FE7410D9" box="[371,477,662,688]" pageId="3" pageNumber="117" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BB9368EFFC4FF9BFF5BFD08FD3E11E1" blockId="3.[136,1452,518,905]" pageId="3" pageNumber="117">
The name “
<taxonomicName id="4C064D0DFFC4FF9BFEA3FD09FDA610BD" authorityName="Merrill" authorityYear="1917" baseAuthorityName="Elmer" box="[325,527,698,724]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B972EA9CFFC4FF9BFEA3FD09FDA610BD" box="[325,527,698,724]" italics="true" pageId="3" pageNumber="117">Timonius hirsutus</emphasis>
</taxonomicName>
” has been used three times. The name was first used by
<bibRefCitation id="EF974B7FFFC4FF9BFB42FD08FAFF10BD" author="Merrill, E. D." box="[1188,1366,698,724]" pageId="3" pageNumber="117" pagination="189 - 247" refId="ref5492" refString="Merrill, E. D. (1917 b) Alabastra Borneensia. Journal of the Straits Branch of the Royal Asiatic Society 77: 189 - 247." type="journal article" year="1917">Merrill (1917b)</bibRefCitation>
for the Bornean species above. The second name
<taxonomicName id="4C064D0DFFC4FF9BFD8FFD6DFBAD1091" authority="Ridley (1918: 81)" authorityName="Ridley" authorityPageNumber="81" authorityYear="1918" box="[617,1028,734,760]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B972EA9CFFC4FF9BFD8FFD6DFC9B1091" box="[617,818,734,760]" italics="true" pageId="3" pageNumber="117">Timonius hirsutus</emphasis>
<bibRefCitation id="EF974B7FFFC4FF9BFCDCFD6CFBAD1091" author="Ridley, H. N." box="[826,1028,734,760]" pageId="3" pageNumber="117" pagination="63 - 100" refId="ref5679" refString="Ridley, H. N. (1918) New and rare Malayan plants, Series X. Journal of the Straits Branch of the Royal Asiatic Society 79: 63 - 100." type="journal article" year="1918">Ridley (1918: 81)</bibRefCitation>
</taxonomicName>
postdates the Bornean name and is a synonym of the Malayan species
<taxonomicName id="4C064D0DFFC4FF9BFE1BFCB0FBBB1175" authority="(Craib 1932: 132) Chaw" authorityName="Chaw" baseAuthorityName="Craib" baseAuthorityPageNumber="132" baseAuthorityYear="1932" box="[509,1042,770,796]" class="Magnoliopsida" family="Rubiaceae" genus="Antirhea" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="atropurpurea">
<emphasis id="B972EA9CFFC4FF9BFE1BFCB0FD521175" box="[509,763,770,796]" italics="true" pageId="3" pageNumber="117">Antirhea atropurpurea</emphasis>
(
<bibRefCitation id="EF974B7FFFC4FF9BFCEFFCB0FC6B1175" author="Craib, W. G." box="[777,962,770,796]" pageId="3" pageNumber="117" refId="ref4993" refString="Craib, W. G. (1932) Florae Siamensis Enumeratio Vol. 2. W. H. Mundie, The Bangkok Times Press, Bangkok, 476 pp." type="book" year="1932">Craib 1932: 132</bibRefCitation>
) Chaw
</taxonomicName>
in
<bibRefCitation id="EF974B7FFFC4FF9BFBD3FCB1FAAD1175" author="Wong, K. M." box="[1077,1284,770,797]" pageId="3" pageNumber="117" pagination="491 - 518" refId="ref6127" refString="Wong, K. M. (1988) The Antirheoideae (Rubiaceae) of the Malay Peninsula. Kew Bulletin 43: 491 - 518. https: // doi. org / 10.2307 / 4118980" type="journal article" year="1988">Wong (1988: 493)</bibRefCitation>
. Subsequently,
<bibRefCitation id="EF974B7FFFC4FF9BFF6EFC94FED21129" author="Merrill, E. D." box="[136,379,806,832]" pageId="3" pageNumber="117" pagination="425 - 485" refId="ref5569" refString="Merrill, E. D. (1921 b) Studies on Philippine Rubiaceae, IV. Philippine Journal of Science 17: 425 - 485." type="journal article" year="1921">Merrill (1921b: 480)</bibRefCitation>
transferred
<taxonomicName id="4C064D0DFFC4FF9BFDEBFC95FC151128" authority="Elmer (1913: 1857)" authorityName="Elmer" authorityPageNumber="1857" authorityYear="1913" box="[525,956,806,833]" class="Magnoliopsida" family="Rubiaceae" genus="Greenea" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis id="B972EA9CFFC4FF9BFDEBFC95FD611129" box="[525,712,806,833]" italics="true" pageId="3" pageNumber="117">Greenea hirsuta</emphasis>
<bibRefCitation id="EF974B7FFFC4FF9BFD32FC94FC151128" author="Elmer, A. D. E." box="[724,956,806,833]" pageId="3" pageNumber="117" pagination="1855 - 1905" refId="ref5249" refString="Elmer, A. D. E. (1913) Rubiaceae from Mount Urdaneta. Leaflets of Philippine Botany 5: 1855 - 1905." type="journal article" year="1913">Elmer (1913: 1857)</bibRefCitation>
</taxonomicName>
into
<taxonomicName id="4C064D0DFFC4FF9BFBE6FC95FBCE1129" baseAuthorityName="Valeton" baseAuthorityYear="1909" box="[1024,1127,807,832]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis id="B972EA9CFFC4FF9BFBE6FC95FBCE1129" box="[1024,1127,807,832]" italics="true" pageId="3" pageNumber="117">Timonius</emphasis>
</taxonomicName>
, generating the illegitimate combination
<taxonomicName id="4C064D0DFFC4FF9BFEFCFCF9FD2A110C" authority="(Elmer) Merr." authorityName="Merrill" authorityYear="1917" baseAuthorityName="Elmer" box="[282,643,842,869]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B972EA9CFFC4FF9BFEFCFCF9FE48110D" box="[282,481,842,868]" italics="true" pageId="3" pageNumber="117">Timonius hirsutus</emphasis>
(Elmer) Merr.
</taxonomicName>
for a species from the
<collectingCountry id="F311761EFFC4FF9BFC9BFCF8FC56110D" box="[893,1023,842,868]" name="Philippines" pageId="3" pageNumber="117">Philippines</collectingCountry>
. Therefore,
<taxonomicName id="4C064D0DFFC4FF9BFB65FCF9FA26110C" authority="Merr." authorityName="Merrill" authorityYear="1917" baseAuthorityName="Elmer" box="[1155,1423,842,869]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="117" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B972EA9CFFC4FF9BFB65FCF9FAE3110D" box="[1155,1354,842,868]" italics="true" pageId="3" pageNumber="117">Timonius hirsutus</emphasis>
Merr.
</taxonomicName>
as used for the Bornean species takes precedence.
</paragraph>
</subSubSection>
</treatment>
</document>