treatments-xml/data/B7/57/71/B757711F3029FFDD2981FF2F2256BFB4.xml

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<mods:title id="7ED5802CF68AE55B55E0C22D3552D53C">Six new species of the genus Bellulia Fibiger, 2008 from China and Cambodia (Lepidoptera, Erebidae, Hypenodinae). Taxonomic study of Micronoctuini Contribution III</mods:title>
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<mods:namePart id="8F1E5CE5E8956CE7FD226367518098CB">Han, Hui Lin</mods:namePart>
<mods:affiliation id="E2B2B2069FEA2CF9014C0CF022913C67">School of Forestry, Northeast Forestry University, Harbin, 150040, China. E-mail: hanhuilin @ aliyun. com &amp; Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, Northeast Forestry University, Harbin, 150040, China.</mods:affiliation>
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<mods:namePart id="1C166919F8BD185BBB37CF526660F58B">Kononenko, Vladimir S.</mods:namePart>
<mods:affiliation id="671835EC7865B1C38B8239E656E5A31E">Laboratory of Entomology, Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok- 22, 690022, Russia.</mods:affiliation>
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<emphasis id="0D8A1C1B3029FFDA2981FF2F27A6BE8F" bold="true" box="[151,444,151,178]" pageId="11" pageNumber="128">
<taxonomicName id="F8FEBB8A3029FFDA2981FF2F2769BE8C" ID-CoL="B7FKB" authority="Han &amp; Kononenko, 2021" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[151,371,151,178]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="diannana" status="sp. nov.">
<emphasis id="0D8A1C1B3029FFDA2981FF2F2769BE8C" bold="true" box="[151,371,151,178]" italics="true" pageId="11" pageNumber="128">Bellulia diannana</emphasis>
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<taxonomicNameLabel id="16B9A1603029FFDA286CFF2F27A6BE8F" box="[378,444,151,177]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="77E493823029FFDA2981FF03273DBEEB" box="[151,295,187,214]" pageId="11" pageNumber="128" type="description">
<paragraph id="3F41C0093029FFDA2981FF03273DBEEB" blockId="11.[151,444,151,214]" box="[151,295,187,214]" pageId="11" pageNumber="128">
(
<figureCitation id="A7C5DC8C3029FFDA2989FF0326ECBEE8" box="[159,246,187,214]" captionStart="FIGURES 9-16" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[180,1435,338,1884]" captionTargetId="figure-89@4.[151,1435,309,1896]" captionTargetPageId="4" captionText="FIGURES 9-16. Bellulia spp., adults. 9. B. jiayiae sp. n., holotype, China; 10. B. bella (after Fibiger 2008); 11. B. kendricki (after Fibiger 2010); 12. B. suffusa (after Fibiger 2008); 13. B. basalia (after Fibiger 2010); 14. B. fuscis sp. n., holotype, Cambodia; 15. B. diannana sp. n., holotype, Cambodia; 16. B. wui (after Fibiger 2008)." figureDoi="http://doi.org/10.5281/zenodo.4561451" httpUri="https://zenodo.org/record/4561451/files/figure.png" pageId="11" pageNumber="128">Figs 15</figureCitation>
, 
<figureCitation id="A7C5DC8C3029FFDA2814FF032705BEEB" box="[258,287,187,213]" captionStart="FIGURES 29-30" captionStartId="11.[151,264,1726,1750]" captionTargetBox="[171,1405,1183,1685]" captionTargetId="figure-484@11.[151,1435,1173,1702]" captionTargetPageId="11" captionText="FIGURES 29-30. Bellulia spp., male genitalia. 29. B. fuscis sp. n., holotype, Cambodia, genit. prep. hhl-2348-1; 30. B. diannana sp. n., holotype, Cambodia, genit. prep. hhl-3538-1." figureDoi="http://doi.org/10.5281/zenodo.4561461" httpUri="https://zenodo.org/record/4561461/files/figure.png" pageId="11" pageNumber="128">30</figureCitation>
)
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<paragraph id="3F41C0093029FFDA2981FEBB2758BF93" blockId="11.[151,1437,259,1150]" pageId="11" pageNumber="128">
<emphasis id="0D8A1C1B3029FFDA2981FEBB2748BF23" bold="true" box="[151,338,259,285]" pageId="11" pageNumber="128">Type materials.</emphasis>
<materialsCitation id="8F96CA543029FFDA284CFEBB2387BF20" ID-GBIF-Occurrence="3047785313" box="[346,1437,259,286]" collectingDate="2011-07-01" collectingDateMax="2015-02-11" collectingDateMin="2011-07-01" collectorName="Y. S. Bae et al" country="Cambodia" location="Pratypes" pageId="11" pageNumber="128" specimenCount="10" specimenCount-male="10" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="0D8A1C1B3029FFDA284CFEBB27CABF23" bold="true" box="[346,464,259,285]" pageId="11" pageNumber="128">
<typeStatus id="E0457EAB3029FFDA284CFEBB27D1BF23" box="[346,459,259,285]" pageId="11" pageNumber="128" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, 
<collectingCountry id="47E980993029FFDA2B09FEBB248FBF23" box="[543,661,259,285]" name="Cambodia" pageId="11" pageNumber="128">Cambodia</collectingCountry>
, Kep, 
<date id="4B40E6C93029FFDA2BF6FEBB2549BF20" box="[736,851,259,286]" pageId="11" pageNumber="128" value="2015-02-11">
<collectingDate id="5B041F213029FFDA2BF6FEBB2549BF20" box="[736,851,259,286]" pageId="11" pageNumber="128" value="2015-02-11">11.ii.2015</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2A76FEBB25D9BF20" box="[864,963,259,286]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA2ADDFEBC25E4BF23" box="[971,1022,259,285]" italics="true" pageId="11" pageNumber="128">et al</emphasis>
., genit. prep. hhl-3538-1, coll. INU.
</materialsCitation>
<materialsCitation id="8F96CA543029FFDA2981FE9F233DBF5B" ID-GBIF-Occurrence="3047785317" collectingDate="2011-07-01" collectingDateMax="2015-02-11" collectingDateMin="2011-07-01" collectorName="Y. S. Bae et al" country="Cambodia" location="Pratypes" pageId="11" pageNumber="128" specimenCount="10" specimenCount-male="10" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="0D8A1C1B3029FFDA2981FE9F2710BF7F" bold="true" box="[151,266,295,321]" pageId="11" pageNumber="128">
<location id="3A2196D23029FFDA2981FE9F271FBF7F" LSID="urn:lsid:plazi:treatment:B757711F3029FFDD2981FF2F2256BFB4:3A2196D23029FFDA2981FE9F271FBF7F" box="[151,261,295,321]" country="Cambodia" name="Pratypes" pageId="11" pageNumber="128" stateProvince="Yunnan">Pratypes</location>
:
</emphasis>
<collectingCountry id="47E980993029FFDA2819FE9F279DBF7F" box="[271,391,295,321]" name="Cambodia" pageId="11" pageNumber="128">Cambodia</collectingCountry>
: 
<specimenCount id="29F80B803029FFDA2899FE9F27C1BF7F" box="[399,475,295,321]" pageId="11" pageNumber="128" type="male">1 male</specimenCount>
, Keo Seima Wildlife Sanctuary, 
<date id="4B40E6C93029FFDA2A28FE9F25ABBF7F" box="[830,945,295,321]" pageId="11" pageNumber="128" value="2011-07-01">
<collectingDate id="5B041F213029FFDA2A28FE9F25ABBF7F" box="[830,945,295,321]" pageId="11" pageNumber="128" value="2011-07-01">1.vii.2011</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2AACFE9F2203BF7C" box="[954,1049,295,322]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA2D0BFE902257BF7F" box="[1053,1101,295,321]" italics="true" pageId="11" pageNumber="128">et al</emphasis>
.; 
<specimenCount id="29F80B803029FFDA2D76FE9F22B6BF7F" box="[1120,1196,295,321]" pageId="11" pageNumber="128" type="male">1 male</specimenCount>
, Kep, 
<date id="4B40E6C93029FFDA2DE6FE9F2382BF7F" box="[1264,1432,295,321]" pageId="11" pageNumber="128" value="2012-07-16" valueMax="2012-07-17" valueMin="2012-07-16">
<collectingDate id="5B041F213029FFDA2DE6FE9F2382BF7F" box="[1264,1432,295,321]" pageId="11" pageNumber="128" value="2012-07-16" valueMax="2012-07-17" valueMin="2012-07-16">16-17.vii.2012</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2981FEF326EDBF58" box="[151,247,331,358]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA29EBFEF4272EBF5B" box="[253,308,331,357]" italics="true" pageId="11" pageNumber="128">et al.</emphasis>
; 
<specimenCount id="29F80B803029FFDA2854FEF32780BF5B" box="[322,410,331,357]" pageId="11" pageNumber="128" type="male">3 males</specimenCount>
, Kep, 
<date id="4B40E6C93029FFDA28F6FEF32470BF5B" box="[480,618,331,357]" pageId="11" pageNumber="128" value="2013-08-19">
<collectingDate id="5B041F213029FFDA28F6FEF32470BF5B" box="[480,618,331,357]" pageId="11" pageNumber="128" value="2013-08-19">19.viii.2013</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2B65FEF324C9BF58" box="[627,723,331,358]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA2BCFFEF42513BF5B" box="[729,777,331,357]" italics="true" pageId="11" pageNumber="128">et al</emphasis>
.; 
<specimenCount id="29F80B803029FFDA2A08FEF32571BF5B" box="[798,875,331,357]" pageId="11" pageNumber="128" type="male">1 male</specimenCount>
, Kep, 
<date id="4B40E6C93029FFDA2AA7FEF3220CBF58" box="[945,1046,331,358]" pageId="11" pageNumber="128" value="2014-02-05">
<collectingDate id="5B041F213029FFDA2AA7FEF3220CBF58" box="[945,1046,331,358]" pageId="11" pageNumber="128" value="2014-02-05">5.ii.2014</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2D36FEF3229ABF58" box="[1056,1152,331,358]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA2D90FEF422ACBF5B" box="[1158,1206,331,357]" italics="true" pageId="11" pageNumber="128">et al</emphasis>
.; 
<specimenCount id="29F80B803029FFDA2DDDFEF32339BF5B" box="[1227,1315,331,357]" pageId="11" pageNumber="128" type="male">3 males</specimenCount>
,
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Kohkong, 
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<date id="4B40E6C93029FFDA2981FED726E7BFB7" box="[151,253,367,393]" pageId="11" pageNumber="128" value="2014-02-07">7.ii.2014</date>
, 
<date id="4B40E6C93029FFDA281CFED7276ABFB4" box="[266,368,367,394]" pageId="11" pageNumber="128" value="2015-02-09">9.ii.2015</date>
</collectingDate>
, 
<collectorName id="920BA5DF3029FFDA286AFED727C5BFB4" box="[380,479,367,394]" pageId="11" pageNumber="128">Y.S. Bae</collectorName>
<emphasis id="0D8A1C1B3029FFDA28F1FEC82403BFB7" box="[487,537,367,393]" italics="true" pageId="11" pageNumber="128">et al</emphasis>
.;
</materialsCitation>
<materialsCitation id="8F96CA543029FFDA2B26FED72758BF93" ID-GBIF-Occurrence="3047785304" collectingDate="2011-07-01" collectingDateMax="2015-02-11" collectingDateMin="2011-07-01" collectorName="H. L. Han &amp; X. X. Jin &amp; H. Geng" country="Cambodia" location="Pratypes" pageId="11" pageNumber="128" specimenCount="10" specimenCount-male="10" stateProvince="Yunnan" typeStatus="holotype">
<collectingCountry id="47E980993029FFDA2B26FED7246DBFB7" box="[560,631,367,393]" name="China" pageId="11" pageNumber="128">China</collectingCountry>
: 
<specimenCount id="29F80B803029FFDA2B92FED724C9BFB7" box="[644,723,367,393]" pageId="11" pageNumber="128" type="male">1 male</specimenCount>
, 
<collectorName id="920BA5DF3029FFDA2BF6FEC82501BFB4" box="[736,795,368,394]" pageId="11" pageNumber="128">Prov.</collectorName>
<collectingRegion id="FD3A0EEB3029FFDA2A37FEC82560BFB4" box="[801,890,368,394]" country="China" name="Yunnan" pageId="11" pageNumber="128">Yunnan</collectingRegion>
, 
<collectorName id="920BA5DF3029FFDA2A90FED725CABFB7" box="[902,976,367,393]" pageId="11" pageNumber="128">Simao</collectorName>
, 
<collectorName id="920BA5DF3029FFDA2ACBFED7224ABFB7" box="[989,1104,367,393]" pageId="11" pageNumber="128">Manxieba</collectorName>
, 
<date id="4B40E6C93029FFDA2D4BFED722C5BFB7" box="[1117,1247,367,393]" pageId="11" pageNumber="128" value="2012-07-22">
<collectingDate id="5B041F213029FFDA2D4BFED722C5BFB7" box="[1117,1247,367,393]" pageId="11" pageNumber="128" value="2012-07-22">22.vii.2012</collectingDate>
</date>
, 
<collectorName id="920BA5DF3029FFDA2DFAFEC82343BFB4" box="[1260,1369,368,394]" pageId="11" pageNumber="128">H.L. Han</collectorName>
, 
<collectorName id="920BA5DF3029FFDA2C70FEC826A2BF93" pageId="11" pageNumber="128">X.X. Jin</collectorName>
&amp; 
<collectorName id="920BA5DF3029FFDA29CAFE2C2758BF93" box="[220,322,403,430]" pageId="11" pageNumber="128">H. Geng.</collectorName>
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</paragraph>
</subSubSection>
<subSubSection id="77E493823029FFDD29D1FE0F2479BF23" lastPageId="12" lastPageNumber="129" pageId="11" pageNumber="128" type="diagnosis">
<paragraph id="3F41C0093029FFDA29D1FE0F24ABBD04" blockId="11.[151,1437,259,1150]" pageId="11" pageNumber="128">
<emphasis id="0D8A1C1B3029FFDA29D1FE0F2758BFEF" bold="true" box="[199,322,439,465]" pageId="11" pageNumber="128">Diagnosis.</emphasis>
Externally this species is similar to 
<taxonomicName id="F8FEBB8A3029FFDA2BCEFE00255DBFEF" authorityName="Fibiger" authorityYear="2008" box="[728,839,439,465]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2BCEFE00255DBFEF" box="[728,839,439,465]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
(
<figureCitation id="A7C5DC8C3029FFDA2A41FE0F25B3BFEF" box="[855,937,439,465]" captionStart="FIGURES 9-16" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[180,1435,338,1884]" captionTargetId="figure-89@4.[151,1435,309,1896]" captionTargetPageId="4" captionText="FIGURES 9-16. Bellulia spp., adults. 9. B. jiayiae sp. n., holotype, China; 10. B. bella (after Fibiger 2008); 11. B. kendricki (after Fibiger 2010); 12. B. suffusa (after Fibiger 2008); 13. B. basalia (after Fibiger 2010); 14. B. fuscis sp. n., holotype, Cambodia; 15. B. diannana sp. n., holotype, Cambodia; 16. B. wui (after Fibiger 2008)." figureDoi="http://doi.org/10.5281/zenodo.4561451" httpUri="https://zenodo.org/record/4561451/files/figure.png" pageId="11" pageNumber="128">Fig. 12</figureCitation>
), but differs from in the light greyish-brown ground colour of the forewing (in 
<taxonomicName id="F8FEBB8A3029FFDA2B01FE64249FBFCB" authorityName="Fibiger" authorityYear="2008" box="[535,645,475,501]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2B01FE64249FBFCB" box="[535,645,475,501]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
it is dark); the basal line is distinct (in 
<taxonomicName id="F8FEBB8A3029FFDA2D2BFE6422B1BFCB" authorityName="Fibiger" authorityYear="2008" box="[1085,1195,475,501]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2D2BFE6422B1BFCB" box="[1085,1195,475,501]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
it is indistinct); antemedial line distinct (in 
<taxonomicName id="F8FEBB8A3029FFDA28B7FDB8240BBC27" authorityName="Fibiger" authorityYear="2008" box="[417,529,511,537]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA28B7FDB8240BBC27" box="[417,529,511,537]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
- distinct); the space between inner and outer bordering lines of the postmedial line is only little darker than the ground colour (in 
<taxonomicName id="F8FEBB8A3029FFDA2BC6FD9C2525BC03" authorityName="Fibiger" authorityYear="2008" box="[720,831,547,573]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2BC6FD9C2525BC03" box="[720,831,547,573]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
it is greyish); the reniform spot is small (in 
<taxonomicName id="F8FEBB8A3029FFDA2C38FD9C2386BC03" authorityName="Fibiger" authorityYear="2008" box="[1326,1436,547,573]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2C38FD9C2386BC03" box="[1326,1436,547,573]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
it is much larger and outlined with black). The male genitalia structure 
<taxonomicName id="F8FEBB8A3029FFDA2AACFDF02259BC5F" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[954,1091,583,609]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="diannana">
<emphasis id="0D8A1C1B3029FFDA2AACFDF02259BC5F" box="[954,1091,583,609]" italics="true" pageId="11" pageNumber="128">B. diannana</emphasis>
</taxonomicName>
(
<figureCitation id="A7C5DC8C3029FFDA2D45FDFF22BFBC5F" box="[1107,1189,583,609]" captionStart="FIGURES 29-30" captionStartId="11.[151,264,1726,1750]" captionTargetBox="[171,1405,1183,1685]" captionTargetId="figure-484@11.[151,1435,1173,1702]" captionTargetPageId="11" captionText="FIGURES 29-30. Bellulia spp., male genitalia. 29. B. fuscis sp. n., holotype, Cambodia, genit. prep. hhl-2348-1; 30. B. diannana sp. n., holotype, Cambodia, genit. prep. hhl-3538-1." figureDoi="http://doi.org/10.5281/zenodo.4561461" httpUri="https://zenodo.org/record/4561461/files/figure.png" pageId="11" pageNumber="128">Fig. 30</figureCitation>
) is close to 
<taxonomicName id="F8FEBB8A3029FFDA2C3BFDF02386BC5F" authorityName="Fibiger" authorityYear="2008" box="[1325,1436,583,609]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2C3BFDF02386BC5F" box="[1325,1436,583,609]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8FEBB8A3029FFDA29DEFDD42733BCBB" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[200,297,619,645]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA29DEFDD42733BCBB" box="[200,297,619,645]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
(
<figureCitation id="A7C5DC8C3029FFDA282CFDD32795BCBB" box="[314,399,619,645]" captionStart="FIGURES 29-30" captionStartId="11.[151,264,1726,1750]" captionTargetBox="[171,1405,1183,1685]" captionTargetId="figure-484@11.[151,1435,1173,1702]" captionTargetPageId="11" captionText="FIGURES 29-30. Bellulia spp., male genitalia. 29. B. fuscis sp. n., holotype, Cambodia, genit. prep. hhl-2348-1; 30. B. diannana sp. n., holotype, Cambodia, genit. prep. hhl-3538-1." figureDoi="http://doi.org/10.5281/zenodo.4561461" httpUri="https://zenodo.org/record/4561461/files/figure.png" pageId="11" pageNumber="128">Fig. 29</figureCitation>
), but differs by in its broader tegumen (in 
<taxonomicName id="F8FEBB8A3029FFDA2A6FFDD425F3BCBB" authorityName="Fibiger" authorityYear="2008" box="[889,1001,619,645]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2A6FFDD425F3BCBB" box="[889,1001,619,645]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8FEBB8A3029FFDA2D35FDD4229EBCBB" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[1059,1156,619,645]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2D35FDD4229EBCBB" box="[1059,1156,619,645]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
the tegumen is narrow); fused withto the fultura superior with prominent processes that are not furcate (in 
<taxonomicName id="F8FEBB8A3029FFDA2D27FD2822BABC97" authorityName="Fibiger" authorityYear="2008" box="[1073,1184,655,681]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2D27FD2822BABC97" box="[1073,1184,655,681]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8FEBB8A3029FFDA2DC1FD28232DBC97" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[1239,1335,655,681]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2DC1FD28232DBC97" box="[1239,1335,655,681]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
the right process is furcate); the juxta is ovoid, with a walnut-shaped circle opening in the center (in 
<taxonomicName id="F8FEBB8A3029FFDA2D8EFD0C231CBCF3" authorityName="Fibiger" authorityYear="2008" box="[1176,1286,691,717]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2D8EFD0C231CBCF3" box="[1176,1286,691,717]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8FEBB8A3029FFDA2C2BFD0C2386BCF3" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[1341,1436,691,717]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2C2BFD0C2386BCF3" box="[1341,1436,691,717]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
the opening is circular); the ampullae are larger than in 
<taxonomicName id="F8FEBB8A3029FFDA2A1FFD602572BCCF" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[777,872,727,754]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2A1FFD602500BCCF" box="[777,794,728,753]" italics="true" pageId="11" pageNumber="128">B</emphasis>
. 
<emphasis id="0D8A1C1B3029FFDA2A3EFD6F2572BCCF" box="[808,872,727,753]" italics="true" pageId="11" pageNumber="128">fuscis</emphasis>
</taxonomicName>
; aedeagus bears a small and triangular horn at the apex (not present in either 
<taxonomicName id="F8FEBB8A3029FFDA28A9FD442437BD2B" authorityName="Fibiger" authorityYear="2008" box="[447,557,763,789]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA28A9FD442437BD2B" box="[447,557,763,789]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
or 
<taxonomicName id="F8FEBB8A3029FFDA2B47FD4424AABD2B" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[593,688,763,789]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2B47FD4424AABD2B" box="[593,688,763,789]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
); the vesica bears seven tiny cornuti of varying size (
<taxonomicName id="F8FEBB8A3029FFDA2C16FD442377BD2B" authorityName="Fibiger" authorityYear="2008" box="[1280,1389,763,789]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="0D8A1C1B3029FFDA2C16FD442377BD2B" box="[1280,1389,763,789]" italics="true" pageId="11" pageNumber="128">B. suffusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8FEBB8A3029FFDA2981FC9826ECBD07" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[151,246,799,825]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis">
<emphasis id="0D8A1C1B3029FFDA2981FC9826ECBD07" box="[151,246,799,825]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
each have five similarly sized cornuti).
</paragraph>
<paragraph id="3F41C0093029FFDA29D1FCFB22E4BA43" blockId="11.[151,1437,259,1150]" pageId="11" pageNumber="128">
<emphasis id="0D8A1C1B3029FFDA29D1FCFB2787BD63" bold="true" box="[199,413,835,861]" pageId="11" pageNumber="128">Discription. Adult</emphasis>
(
<figureCitation id="A7C5DC8C3029FFDA28BBFCFB241ABD60" box="[429,512,835,862]" captionStart="FIGURES 9-16" captionStartId="4.[151,264,1919,1943]" captionTargetBox="[180,1435,338,1884]" captionTargetId="figure-89@4.[151,1435,309,1896]" captionTargetPageId="4" captionText="FIGURES 9-16. Bellulia spp., adults. 9. B. jiayiae sp. n., holotype, China; 10. B. bella (after Fibiger 2008); 11. B. kendricki (after Fibiger 2010); 12. B. suffusa (after Fibiger 2008); 13. B. basalia (after Fibiger 2010); 14. B. fuscis sp. n., holotype, Cambodia; 15. B. diannana sp. n., holotype, Cambodia; 16. B. wui (after Fibiger 2008)." figureDoi="http://doi.org/10.5281/zenodo.4561451" httpUri="https://zenodo.org/record/4561451/files/figure.png" pageId="11" pageNumber="128">Fig. 15</figureCitation>
). Wingspan 11.5-13.2 mm. Head, thorax and abdomen yellowish brown. Forewing pale yellowish brown, with black and dark-brown scaling; basal line black, arched, distinct, broader near costal margin; antemedial line double, arched, its inner bordering line ochreous brown, slender, outer bordering line black to blackish-brown, broader between costal and inner veins of discal cell, otherwise slender; postmedial line black, double, waved, distinct, fused to broad patch in costal area; subterminal line double, slightly waved, its inner line thin, present as only as small dot at costal margin; otherwise somewhat darker than ground colour, outer line broad and black between costa and M2, otherwise indistinct; terminal line black; fringe same as ground colour; reniform spot small, whitish yellow. Hindwing greyish brown, darker than forewing; discal spot tinged with dark, indistinct; outer wing margin weakly incurved at M2; fringe lighter than ground colour, greyish-yellow at base.
</paragraph>
<caption id="6B8190813029FFDA2981F90624FBB8C4" ID-DOI="http://doi.org/10.5281/zenodo.4561461" ID-Zenodo-Dep="4561461" httpUri="https://zenodo.org/record/4561461/files/figure.png" pageId="11" pageNumber="128" startId="11.[151,264,1726,1750]" targetBox="[171,1405,1183,1685]" targetPageId="11">
<paragraph id="3F41C0093029FFDA2981F90624FBB8C4" blockId="11.[151,1436,1726,1786]" pageId="11" pageNumber="128">
<emphasis id="0D8A1C1B3029FFDA2981F906274BB8E8" bold="true" box="[151,337,1726,1750]" pageId="11" pageNumber="128">FIGURES 29-30.</emphasis>
<taxonomicName id="F8FEBB8A3029FFDA2841F90627BDB8E8" authorityName="Fibiger" authorityYear="2008" box="[343,423,1726,1750]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="0D8A1C1B3029FFDA2841F90627BDB8E8" box="[343,423,1726,1750]" italics="true" pageId="11" pageNumber="128">Bellulia</emphasis>
</taxonomicName>
spp., male genitalia. 29. 
<taxonomicName id="F8FEBB8A3029FFDA2BBFF907251AB8E8" authority="Han &amp; Kononenko, 2021" authorityName="Han &amp; Kononenko" authorityYear="2021" box="[681,768,1726,1750]" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="fuscis" status="sp. nov.">
<emphasis id="0D8A1C1B3029FFDA2BBFF907251AB8E8" box="[681,768,1726,1750]" italics="true" pageId="11" pageNumber="128">B. fuscis</emphasis>
</taxonomicName>
<emphasis id="0D8A1C1B3029FFDA2A11F906255BB8E8" bold="true" box="[775,833,1726,1750]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="16B9A1603029FFDA2A11F906255BB8E8" box="[775,833,1726,1750]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, Cambodia, genit. prep. hhl-2348-1; 30. 
<taxonomicName id="F8FEBB8A3029FFDA2C5CF90726D0B8C4" authority="Han &amp; Kononenko, 2021" authorityName="Han &amp; Kononenko" authorityYear="2021" class="Insecta" family="Erebidae" genus="Bellulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="diannana" status="sp. nov.">
<emphasis id="0D8A1C1B3029FFDA2C5CF90726D0B8C4" italics="true" pageId="11" pageNumber="128">B. diannana</emphasis>
</taxonomicName>
<emphasis id="0D8A1C1B3029FFDA29C6F95A2713B8C4" bold="true" box="[208,265,1762,1786]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="16B9A1603029FFDA29C6F95A2713B8C4" box="[208,265,1762,1786]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, Cambodia, genit. prep. hhl-3538-1.
</paragraph>
</caption>
<paragraph id="3F41C0093029FFDD29D1F8902353BEC7" blockId="11.[151,1437,1832,2039]" lastBlockId="12.[151,1437,151,394]" lastPageId="12" lastPageNumber="129" pageId="11" pageNumber="128">
<emphasis id="0D8A1C1B3029FFDA29D1F8902769B97C" bold="true" box="[199,371,1832,1858]" pageId="11" pageNumber="128">Male genitalia</emphasis>
(
<figureCitation id="A7C5DC8C3029FFDA2892F89027C0B97C" box="[388,474,1832,1858]" captionStart="FIGURES 29-30" captionStartId="11.[151,264,1726,1750]" captionTargetBox="[171,1405,1183,1685]" captionTargetId="figure-484@11.[151,1435,1173,1702]" captionTargetPageId="11" captionText="FIGURES 29-30. Bellulia spp., male genitalia. 29. B. fuscis sp. n., holotype, Cambodia, genit. prep. hhl-2348-1; 30. B. diannana sp. n., holotype, Cambodia, genit. prep. hhl-3538-1." figureDoi="http://doi.org/10.5281/zenodo.4561461" httpUri="https://zenodo.org/record/4561461/files/figure.png" pageId="11" pageNumber="128">Fig. 30</figureCitation>
). Tegumen narrow, 5/4 as long as vinculum; its ventral part fused to fultura superior with prominent, log-shaped, curved processes on each side, left process longer than righttapered apically, right process scraping knife shape, apically sharp, triangular prism-shape basally. Vinculum thick, broad, V shaped. Saccus broad, V-shaped. Juxta ovoid, sclerotized, with walnut-shape rounded opening in center. Valva broad, slightly asymmetrical; sacculus weakly asymmetrical, right sacculus somewhat shorter and broader than left, 2/3 as long as valva, broader than base of valva, left sacculus same width as basae of valva; ampullae slightly asymmetrical, rounded, covered with heavy setae, right ampullae slightly larger than left; costa thick and arched, thinner in right valva than in left. Aedeagus with coecum 1/4 as long as aedeagus, sclerotized, strongly curved in posteriorly, with a small and triangular horn at apex; ductus ejaculatorius swollen; vesica with seven tiny variously sized medial cornuti.
</paragraph>
<paragraph id="3F41C009302EFFDD29D1FEBB2479BF23" blockId="12.[151,1437,151,394]" box="[199,611,258,285]" pageId="12" pageNumber="129">
<emphasis id="0D8A1C1B302EFFDD29D1FEBB2796BF22" bold="true" box="[199,396,258,285]" pageId="12" pageNumber="129">Female genitalia</emphasis>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="77E49382302EFFDD29D1FE9F2343BF5B" pageId="12" pageNumber="129" type="etymology">
<paragraph id="3F41C009302EFFDD29D1FE9F2343BF5B" blockId="12.[151,1437,151,394]" pageId="12" pageNumber="129">
<emphasis id="0D8A1C1B302EFFDD29D1FE9F2754BF7F" bold="true" box="[199,334,295,321]" pageId="12" pageNumber="129">Etymology.</emphasis>
The species name is the combination of two Chinese words, it refers to 
<collectingRegion id="FD3A0EEB302EFFDD2D61FE9F232DBF7F" box="[1143,1335,295,321]" country="China" name="Yunnan" pageId="12" pageNumber="129">Yunnan Province</collectingRegion>
which is called “Dian” for shortness in Chinese, and South which called “ 
<collectingRegion id="FD3A0EEB302EFFDD2A7DFEF325BBBF5B" box="[875,929,331,357]" country="Thailand" name="Nan" pageId="12" pageNumber="129">Nan</collectingRegion>
”. The name indicates South of 
<collectingRegion id="FD3A0EEB302EFFDD2DEAFEF3234FBF5B" box="[1276,1365,331,357]" country="China" name="Yunnan" pageId="12" pageNumber="129">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="77E49382302EFFDD29D1FED62256BFB4" box="[199,1100,366,394]" pageId="12" pageNumber="129" type="distribution">
<paragraph id="3F41C009302EFFDD29D1FED62256BFB4" blockId="12.[151,1437,151,394]" box="[199,1100,366,394]" pageId="12" pageNumber="129">
<emphasis id="0D8A1C1B302EFFDD29D1FED6277BBFB6" bold="true" box="[199,353,366,392]" pageId="12" pageNumber="129">Distribution.</emphasis>
<collectingCountry id="47E98099302EFFDD2871FED727B0BFB7" box="[359,426,367,393]" name="China" pageId="12" pageNumber="129">China</collectingCountry>
(Prov. 
<collectingRegion id="FD3A0EEB302EFFDD28EDFED7244CBFB7" box="[507,598,367,393]" country="China" name="Yunnan" pageId="12" pageNumber="129">Yunnan</collectingRegion>
); 
<collectingCountry id="47E98099302EFFDD2B7EFED724C1BFB7" box="[616,731,367,393]" name="Cambodia" pageId="12" pageNumber="129">Cambodia</collectingCountry>
(Prov. 
<collectingRegion id="FD3A0EEB302EFFDD2A3BFED72546BFB7" box="[813,860,367,393]" country="Cambodia" name="Kep" pageId="12" pageNumber="129">Kep</collectingRegion>
and Prov. Kohkong).
</paragraph>
</subSubSection>
</treatment>
</document>