treatments-xml/data/B7/11/B2/B711B23FFECE8709DA3FC284FC237E3C.xml

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<mods:title id="3E0D99050032AD72BB51A5644041206F">Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph</mods:title>
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<mods:namePart id="33E0CEEF549A9052EC007E1A7F140370">Nakayama, Naohide</mods:namePart>
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<mods:affiliation id="713E76954CABA550EF638A0797E66DF4">Department of Marine Biology, School of Marine Science and Technology, Tokai University, 3 - 20 - 1 Orido, Shimizu, Shizuoka 424 - 8610, Japan ® naonakayama @ tsc. u-tokai. ac. jp or ®</mods:affiliation>
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<treatment id="B711B23FFECE8709DA3FC284FC237E3C" ID-DOI="http://doi.org/10.5281/zenodo.6422850" ID-GBIF-Taxon="194588107" ID-Zenodo-Dep="6422850" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C" httpUri="http://treatment.plazi.org/id/B711B23FFECE8709DA3FC284FC237E3C" lastPageId="283" lastPageNumber="284" pageId="280" pageNumber="281">
<subSubSection id="77A250A2FECE870ADA3FC284FB837DDC" pageId="280" pageNumber="281" type="nomenclature">
<paragraph id="3F070329FECE870ADA3FC284FB1F7DF3" blockId="280.[810,1272,172,268]" box="[810,1272,172,199]" pageId="280" pageNumber="281">
<heading id="644FB445FECE870ADA3FC284FB1F7DF3" box="[810,1272,172,199]" fontSize="11" level="2" pageId="280" pageNumber="281" reason="2">
<taxonomicName id="F8B878AAFECE870ADA3FC284FB1F7DF3" ID-CoL="57PJN" authority="(Gunther, 1877)" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[810,1272,172,199]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECE870ADA3FC284FB1F7DF3" bold="true" box="[810,1272,172,199]" pageId="280" pageNumber="281">
<emphasis id="0DCCDF3BFECE870ADA3FC284FBCC7DF2" bold="true" box="[810,1067,172,199]" italics="true" pageId="280" pageNumber="281">Trachonurus villosus</emphasis>
(Günther, 1877)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3F070329FECE870ADA3FC2E7FB837DDC" blockId="280.[810,1272,172,268]" box="[810,1124,206,233]" pageId="280" pageNumber="281">[Japanese name: Wata-hige]</paragraph>
</subSubSection>
<subSubSection id="77A250A2FECE870ADA3FC2D9FB4D7F2E" pageId="280" pageNumber="281" type="description">
<paragraph id="3F070329FECE870ADA3FC2D9FB417C3E" blockId="280.[810,1272,172,268]" box="[810,1190,241,268]" pageId="280" pageNumber="281">
(
<figureCitation id="A7831FACFECE870ADA27C2D9FC307C39" box="[818,983,241,268]" captionStart-0="FIGURE 193" captionStart-1="FIGURE 194" captionStart-2="FIGURE 195" captionStartId-0="281.[136,235,1467,1491]" captionStartId-1="282.[136,235,685,709]" captionStartId-2="282.[136,235,1399,1423]" captionTargetBox-0="[151,1436,158,1428]" captionTargetBox-1="[151,1436,159,652]" captionTargetBox-2="[151,1436,850,1360]" captionTargetId-0="figure-243@281.[151,1436,158,1428]" captionTargetId-1="figure-256@282.[151,1436,159,652]" captionTargetId-2="figure-301@282.[151,1436,850,1360]" captionTargetPageId-0="281" captionTargetPageId-1="282" captionTargetPageId-2="282" captionText-0="FIGURE 193. Fresh specimens of Trachonurus villosus. (A) BSKU 23083, 45.2 mm HL, 290+ mm TL, off Kochi, Kochi Pref., Pacific, 1043 m depth; (B) BSKU 43458, 79.7 mm HL, 463+ mm TL, south-southwest of Okino-shima Island, Kochi Pref., Pacific, 1075–1092 m depth; (C) HUMZ 71883, 110 mm HL, 675+ mm TL, off Minami-torishima Island, Pacific. Lateral views. [Photos: (A–B) BSKU; (C) HUMZ]" captionText-1="FIGURE 194. Holotype of Coryphaenoides villosus (= Trachonurus villosus). BMNH 1887.12.7.105, 26.0 mm HL, 193+ mm TL, Sagami Bay, Pacific, 631 m depth. (A) Lateral view; (B) dorsal and (C) ventral views of the head and body. [Photos: N. Nakayama]" captionText-2="FIGURE 195. Scanning electron micrographs showing a body scale (from the dorsum below the interdorsal space) of Trachonurus villosus. BSKU 43458, 79.7 mm HL. (A) View from above; (B) oblique view. [Photos: N. Nakayama]" figureDoi-0="http://doi.org/10.5281/zenodo.6423202" figureDoi-1="http://doi.org/10.5281/zenodo.6423204" figureDoi-2="http://doi.org/10.5281/zenodo.6423206" httpUri-0="https://zenodo.org/record/6423202/files/figure.png" httpUri-1="https://zenodo.org/record/6423204/files/figure.png" httpUri-2="https://zenodo.org/record/6423206/files/figure.png" pageId="280" pageNumber="281">Figs. 193–195</figureCitation>
; Appendix 3-12B)
</paragraph>
<paragraph id="3F070329FECE870ADA3FC310FA4D7CA6" blockId="280.[810,1452,312,1559]" pageId="280" pageNumber="281">
<taxonomicName id="F8B878AAFECE870ADA3FC310FBFF7C78" authorityName="Gunther" authorityYear="1877" box="[810,1048,312,333]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECE870ADA3FC310FBFF7C78" box="[810,1048,312,333]" italics="true" pageId="280" pageNumber="281">Coryphaenoides villosus</emphasis>
</taxonomicName>
<materialsCitation id="8FD00974FECE870ADD34C311FA427CA6" ID-GBIF-Occurrence="3750954874" collectionCode="BMNH" county="Challenger" elevation="631" location="Yeddo" municipality="Eno-shima Island" pageId="280" pageNumber="281" specimenCode="BMNH 1887.12" specimenCount="1" typeStatus="holotype">
G̹nther, 1877:441 [original description; 
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: 
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.7.105, from “south of 
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” (off 
<collectingMunicipality id="DF639953FECE870ADA4AC354FBEC7CA7" box="[863,1035,380,402]" pageId="280" pageNumber="281">Eno-shima Island</collectingMunicipality>
), 
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<collectingCounty id="D6667BA5FECE870ADD09C354FB6F7CA4" box="[1052,1160,380,401]" pageId="280" pageNumber="281">Challenger</collectingCounty>
</emphasis>
sta. 232, in 345 ftm (
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<elevation id="B495E41AFECE870ADC40C354FA707CA6" box="[1365,1431,380,403]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="280" pageNumber="281" unit="m" value="631.0">631 m</elevation>
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)]
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.
</paragraph>
<paragraph id="3F070329FECE870ADA3FC3B7FB4D7F2E" blockId="280.[810,1452,312,1559]" pageId="280" pageNumber="281">
<taxonomicName id="F8B878AAFECE870ADA3FC3B7FC047C86" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[810,995,414,436]" family="Macrouridae" genus="Macrurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECE870ADA3FC3B7FC047C86" box="[810,995,414,436]" italics="true" pageId="280" pageNumber="281">Macrurus villosus</emphasis>
</taxonomicName>
: 
<materialsCitation id="8FD00974FECE870ADAEEC3B7FB417F2E" ID-GBIF-Occurrence="3750954646" collectionCode="BMNH" pageId="280" pageNumber="281" specimenCode="BMNH 1887.12" specimenCount="2" typeStatus="holotype">
G̹nther 1887:142–143, pl. XXXVI, fig. B [description; in part, 
<specimenCount id="29BEC8A0FECE870ADD72C3E9FB517CE3" box="[1127,1206,449,470]" pageId="280" pageNumber="281" type="generic">1 spec.</specimenCount>
, 
<specimenCode id="6F1EAB52FECE870ADDDCC3E9FA967CE3" box="[1225,1393,448,470]" collectionCode="BMNH" pageId="280" pageNumber="281">BMNH 1887.12</specimenCode>
.7.105 (
<typeStatus id="E003BD8BFECE870ADA73C3CBFC277CCC" box="[870,960,483,505]" pageId="280" pageNumber="281" type="holotype">holotype</typeStatus>
); remaining 
<specimenCount id="29BEC8A0FECE870ADD2AC3CBFB617CCD" box="[1087,1158,483,504]" pageId="280" pageNumber="281" type="generic">1 spec.</specimenCount>
probably represents different species (fide Iwamoto 1997:945)]
</materialsCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="3F070329FECE870ADA3FC00FFA857B23" blockId="280.[810,1452,312,1559]" pageId="280" pageNumber="281">
<taxonomicName id="F8B878AAFECE870ADA3FC00FFC117F09" ID-CoL="57PJN" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[810,1014,551,572]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECE870ADA3FC00FFC117F09" box="[810,1014,551,572]" italics="true" pageId="280" pageNumber="281">Trachonurus villosus</emphasis>
</taxonomicName>
: 
<bibRefCitation id="5B297ED8FECE870ADD13C00FFAF57F09" author="Jordan, D. S. &amp; Snyder, J. O." box="[1030,1298,551,573]" pageId="280" pageNumber="281" pagination="31 - 159" refId="ref290097" refString="Jordan, D. S. &amp; Snyder, J. O. (1901) A preliminary check list of the fishes of Japan. Annotationes Zoologicae Japonenses, 3, 31 - 159." type="journal article" year="1901">
<collectingCountry id="47AF43B9FECE870ADD13C00FFBA17F08" box="[1030,1094,551,573]" name="Jordan" pageId="280" pageNumber="281">Jordan</collectingCountry>
&amp; Snyder 1901:120
</bibRefCitation>
(listed; 
<collectingCountry id="47AF43B9FECE870ADC73C00FFA467F09" box="[1382,1441,551,572]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<collectingCountry id="47AF43B9FECE870ADA4AC061FC787F6A" box="[863,927,585,607]" name="Jordan" pageId="280" pageNumber="281">Jordan</collectingCountry>
&amp; Gilbert 
<emphasis id="0DCCDF3BFECE870ADD1DC061FBFC7F6B" box="[1032,1051,585,606]" italics="true" pageId="280" pageNumber="281">in</emphasis>
<bibRefCitation id="5B297ED8FECE870ADD34C061FAFA7F6A" author="Jordan, D. S. &amp; Starks, E. C." box="[1057,1309,585,607]" pageId="280" pageNumber="281" pagination="577 - 630" refId="ref290133" refString="Jordan, D. S. &amp; Starks, E. C. (1904) List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the U. S. Fish Commission, 22, 577 - 630, pls. 1 - 8." type="journal article" year="1904">
<collectingCountry id="47AF43B9FECE870ADD34C061FB867F6A" box="[1057,1121,585,607]" name="Jordan" pageId="280" pageNumber="281">Jordan</collectingCountry>
&amp; Starks 1904:621
</bibRefCitation>
[after G̹nther (1877, 1887)]; 
<bibRefCitation id="5B297ED8FECE870ADAEFC043FB067FB5" author="Jordan, D. S. &amp; Tanaka, S. &amp; Snyder, J. O." box="[1018,1249,619,641]" pageId="280" pageNumber="281" pagination="1 - 497" refId="ref290203" refString="Jordan, D. S., Tanaka, S. &amp; Snyder, J. O. (1913) A catalogue of the fishes of Japan. Journal of the College of Science, Imperial University of Tokyo, 33, 1 - 497." type="journal article" year="1913">
<collectingCountry id="47AF43B9FECE870ADAEFC043FBDD7FB4" box="[1018,1082,619,641]" name="Jordan" pageId="280" pageNumber="281">Jordan</collectingCountry>
<emphasis id="0DCCDF3BFECE870ADD53C043FB9B7FB5" box="[1094,1148,619,640]" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
1913:420
</bibRefCitation>
(listed; 
<collectingCountry id="47AF43B9FECE870ADC2EC043FA927FB5" box="[1339,1397,619,640]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
; new Japanese name: “Wata-hige”); 
<bibRefCitation id="5B297ED8FECE870ADD8DC0A5FA4B7F97" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[1176,1452,652,675]" pageId="280" pageNumber="281" pagination="135 - 214" refId="ref285931" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1916) Report on the Japanese macrourid fishes collected by the United States fisheries steamer &quot; Albatross &quot; in 1906, with a synopsis of the genera. Proceedings of the United States National Museum, 51, 135 - 214, pls. 8 - 11. http: // dx. doi. org / 10.5479 / si. 00963801.51 - 2149.135" type="journal article" year="1916">Gilbert &amp; Hubbs 1916:205</bibRefCitation>
(description; 
<specimenCount id="29BEC8A0FECE870ADACAC087FBC37FF1" box="[991,1060,687,708]" pageId="280" pageNumber="281" type="generic">4 spec.</specimenCount>
from off Cape Shionomisaki and southsouthwest of Izu Island); 
<bibRefCitation id="5B297ED8FECE870ADD5AC0F9FAF97FD3" author="Kamohara, T." box="[1103,1310,720,743]" pageId="280" pageNumber="281" pagination="533 - 544" refId="ref290351" refString="Kamohara, T. (1931 b) Kochi-shi fukin no gyorui tsuiki (1) [Additions to the fishes collected in the vicinity of Kochi City (1)]. Dobutsugaku Zasshi [Zoological Magazine, Japan], 43, 533 - 544. [In Japanese.]" type="journal article" year="1931">Kamohara 1931b:544</bibRefCitation>
(questionable; listed; 
<collectingRegion id="FD7CCDCBFECE870ADABDC0DBFC007E3C" box="[936,999,755,777]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADD15C0DBFB2C7E3C" author="Kamohara, T." box="[1024,1227,755,777]" pageId="280" pageNumber="281" pagination="1196 - 1202" refId="ref290404" refString="Kamohara, T. (1934 a) Tosaoki no shinkaigyo ni tsuite [Deepsea fishes of the Province of Tosa]. &quot; Shokubutsu oyobi Dobutsu &quot; [Plants and Animals], 2, 1196 - 1202. [In Japanese.]" type="journal article" year="1934">Kamohara 1934a:54</bibRefCitation>
(questionable; listed; 
<collectingRegion id="FD7CCDCBFECE870ADA4AC13DFC797E1E" box="[863,926,789,811]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADAA2C13DFB127E1E" author="Okada, Y. &amp; Matsubara, K." box="[951,1269,789,811]" pageId="280" pageNumber="281" refId="ref297013" refString="Okada, Y. &amp; Matsubara, K. (1938) Keys to the Fishes and Fish- Like Animals of Japan Including Kuril Islands, Southern Sakhalin, Bonin Islands, Ryukyu Islands, Korea and Formosa. Sanseido Co., Ltd., Tokyo, xl + 584 pp., 113 pls. [In Japanese.]" type="book" year="1938">Okada &amp; Matsubara 1938:453</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADC73C13DFA467E1F" box="[1382,1441,789,810]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADA4AC11FFBFC7E79" author="Kamohara, T." box="[863,1051,823,845]" pageId="280" pageNumber="281" refId="ref290644" refString="Kamohara, T. (1938) On the Offshore Bottom-Fishes of Prov. Tosa, Shikoku, Japan. Maruzen Co., Ltd., Tokyo, 86 pp." type="book" year="1938">Kamohara 1938:73</bibRefCitation>
(questionable; spec. from 
<collectingRegion id="FD7CCDCBFECE870ADC3FC11FFA457E78" box="[1322,1442,823,845]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi Pref.</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADA4AC171FBC97E5A" author="Kamohara, T." box="[863,1070,857,879]" pageId="280" pageNumber="281" refId="ref290676" refString="Kamohara, T. (1950) Description of the Fishes from the Provinces of Tosa and Kishu, Japan. Kochi-Ken Bunkyo Kyokai [Kochi Literature Education Association], Kochi, 288 + 5 + 48 + 26 pp. [In Japanese.]" type="book" year="1950">Kamohara 1950:279</bibRefCitation>
(questionable; listed; 
<collectingRegion id="FD7CCDCBFECE870ADC36C171FA457E5A" box="[1315,1442,857,879]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi Pref.</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADA4AC153FBFC7EA4" author="Kamohara, T." box="[863,1051,891,913]" pageId="280" pageNumber="281" pagination="1 - 122" refId="ref290723" refString="Kamohara, T. (1952) Revised descriptions of the offshore bottom-fishes of Prov. Tosa, Shikoku, Japan. Reports of the Kochi University, Natural Science, (3), 1 - 122." type="journal article" year="1952">Kamohara 1952:99</bibRefCitation>
(questionable; spec. from 
<collectingRegion id="FD7CCDCBFECE870ADC3FC153FA457EA4" box="[1322,1442,891,913]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi Pref.</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADA4AC1B5FBD17E86" author="Matsubara, K." box="[863,1078,925,947]" pageId="280" pageNumber="281" refId="ref293415" refString="Matsubara, K. (1955) Fish Morphology and Hierarchy. Ishizaki Shoten, Tokyo, xi + v + 1950 + viii pp., 135 pls. [In Japanese.]" type="book" year="1955">Matsubara 1955:1317</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADD88C1B5FB3F7E87" box="[1181,1240,925,946]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADDF8C1B5FA4C7E86" author="Kamohara, T." box="[1261,1451,925,947]" pageId="280" pageNumber="281" pagination="1 - 76" refId="ref290804" refString="Kamohara, T. (1958) A catalogue of fishes of Kochi Prefecture (Province Tosa), Japan. Reports of the Usa Marine Biological Station, 5, 1 - 76." type="journal article" year="1958">Kamohara 1958:74</bibRefCitation>
(questionable; listed; 
<collectingRegion id="FD7CCDCBFECE870ADD5DC197FB2E7EE0" box="[1096,1225,959,981]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi Pref.</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADDF3C197FA4C7EE0" author="Kamohara, T." box="[1254,1451,959,981]" pageId="280" pageNumber="281" pagination="1 - 99" refId="ref290959" refString="Kamohara, T. (1964) Revised catalogue of fishes of Kochi Prefecture, Japan. Reports of the Usa Marine Biological Station, 11, 1 - 99." type="journal article" year="1964">Kamohara 1964:96</bibRefCitation>
(questionable; listed; 
<collectingRegion id="FD7CCDCBFECE870ADD26C1C9FB4D7EC2" box="[1075,1194,993,1015]" country="Japan" name="Kochi" pageId="280" pageNumber="281">Kochi Pref.</collectingRegion>
); 
<bibRefCitation id="5B297ED8FECE870ADDA9C1C9FA617EC2" author="Okamura, O." box="[1212,1414,993,1015]" pageId="280" pageNumber="281" refId="ref297275" refString="Okamura, O. (1970 a) Fauna Japonica, Macrourina (Pisces). Academic Press of Japan, Tokyo, 216 pp., 64 pls." type="book" year="1970">Okamura 1970a:106</bibRefCitation>
, pl. XXIII, text-fig. 44 (description; 
<specimenCount id="29BEC8A0FECE870ADD87C62BFB32792D" box="[1170,1237,1027,1048]" pageId="280" pageNumber="281" type="generic">1 spec.</specimenCount>
from East 
<collectingCountry id="47AF43B9FECE870ADC29C62BFA92792C" box="[1340,1397,1027,1049]" name="China" pageId="280" pageNumber="281">China</collectingCountry>
Sea); 
<bibRefCitation id="5B297ED8FECE870ADA4AC60DFBE7790E" author="Okamura, O." box="[863,1024,1061,1083]" pageId="280" pageNumber="281" pagination="1 - 179" refId="ref297305" refString="Okamura, O. (1970 b) Studies on the macrouroid fishes of Japan: morphology, ecology and phylogeny. Reports of the Usa Marine Biological Station, 17, 1 - 179, pls. I-V." type="journal article" year="1970">Okamura 1970b</bibRefCitation>
: table 1 (listed; 
<collectingCountry id="47AF43B9FECE870ADD8BC60DFB3E790F" box="[1182,1241,1061,1082]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADDFFC60DFC5E7969" author="Tominaga, Y. &amp; Uyeno, T." pageId="280" pageNumber="281" pagination="437 - 574" refId="ref301688" refString="Tominaga, Y. &amp; Uyeno, T. (1981) List of Japanese fishes. In: Yasuda, F., Takagi, K., Tominaga, Y., Uyeno, T., Abe, T., Ishiyama, R., Iwai, T., Ochiai, A., Kuronuma, K. &amp; Nakamura, M. (Eds.), Dictionary of Japanese Fish Names and Their Foreign Equivalents. Sanseido Co., Ltd., Tokyo, pp. 437 - 574." type="book chapter" year="1981">Tominaga &amp; Uyeno 1981:489</bibRefCitation>
(listed; 
<collectingCountry id="47AF43B9FECE870ADD1AC66FFBAD7969" box="[1039,1098,1095,1116]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADD4BC66FFAFA7968" author="Okamura, O." box="[1118,1309,1095,1117]" pageId="280" pageNumber="281" pagination="93 - 99" refId="ref297499" refString="Okamura, O. (1984 b) Macrouroidei. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (Eds.), The Fishes of the Japanese Archipelago. 1 st Edition. Tokai University Press, Tokyo, pp. 93 - 99, pls. 79 - 83. [In Japanese.]" type="book chapter" year="1984">Okamura 1984b:94</bibRefCitation>
, pl. 81, fig. L (compiled); 
<bibRefCitation id="5B297ED8FECE870ADAC9C641FB75794B" author="Okamura, O." box="[988,1170,1129,1151]" pageId="280" pageNumber="281" pagination="344 - 373" refId="ref297606" refString="Okamura, O. (1988) Macrouroidei. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (Eds.), The Fishes of the Japanese Archipelago. 2 nd Edition. Tokai University Press, Tokyo, pp. 93 - 99, 453, pls. 79 - 83, 344 - 373." type="journal article" year="1988">Okamura 1988:94</bibRefCitation>
, pl. 81, fig. L (compiled); 
<bibRefCitation id="5B297ED8FECE870ADA4AC6A3FB4D7995" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[863,1194,1163,1184]" pageId="280" pageNumber="281" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto 1992:77</bibRefCitation>
(unnecessary 
<typeStatus id="E003BD8BFECE870ADC46C6A3FA4C7994" box="[1363,1451,1163,1185]" pageId="280" pageNumber="281" type="lectotype">lectotype</typeStatus>
designation; synonymy; comments on 
<emphasis id="0DCCDF3BFECE870ADDF3C685FAB579F7" box="[1254,1362,1197,1218]" italics="true" pageId="280" pageNumber="281">Challenger</emphasis>
spec.; 2 additional spec. from Sala y Gomez Ridge); 
<bibRefCitation id="5B297ED8FECE870ADC14C6E7FA4C79D0" author="Nakabo, T." box="[1281,1451,1231,1253]" pageId="280" pageNumber="281" pagination="1276 - 1277" refId="ref295357" refString="Nakabo, T. (1993) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 1 st Edition. Tokai University Press, Tokyo, pp. 353 - 371, 1276 - 1277. [In Japanese.]" type="journal article" year="1993">Nakabo 1993:356</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADAD7C6D9FC1A7833" box="[962,1021,1265,1286]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADD02C6D9FB3A7833" author="Okamura, O." box="[1047,1245,1265,1287]" pageId="280" pageNumber="281" pagination="124 - 129" refId="ref297681" refString="Okamura, O. (1997) Macrouridae. In: Okamura, O. &amp; Amaoka, K. (Eds.), Sea Fishes of Japan. Yama-Kei Publishers Co., Ltd., Tokyo, pp. 124 - 129. [In Japanese.]" type="book chapter" year="1997">Okamura 1997:128</bibRefCitation>
, fig. 7 (compiled); 
<bibRefCitation id="5B297ED8FECE870ADA4AC73BFBEB781D" author="Nakabo, T." box="[863,1036,1299,1321]" pageId="280" pageNumber="281" pagination="417 - 435" refId="ref295412" refString="Nakabo, T. (2000) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 2 nd Edition. Tokai University Press, Tokyo, pp. 417 - 435, 1494. [In Japanese.]" type="journal article" year="2000">Nakabo 2000:420</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADD71C73BFB78781D" box="[1124,1183,1299,1320]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADDA5C73BFA4C781C" author="Shinohara, G. &amp; Endo, H. &amp; Matsuura, K. &amp; Machida, Y. &amp; Honda, H." box="[1200,1451,1299,1321]" pageId="280" pageNumber="281" pagination="283 - 343" refId="ref299992" refString="Shinohara, G., Endo, H., Matsuura, K., Machida, Y. &amp; Honda, H. (2001) Annotated checklist of the deepwater fishes from Tosa Bay, Japan. National Science Museum Tokyo Monographs, (20), 283 - 343." type="journal article" year="2001">
Shinohara 
<emphasis id="0DCCDF3BFECE870ADC0CC73CFAAC781D" box="[1305,1355,1299,1321]" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
2001:306
</bibRefCitation>
(listed; Tosa Bay); 
<bibRefCitation id="5B297ED8FECE870ADD03C71DFB25787F" author="Nakabo, T." box="[1046,1218,1333,1355]" pageId="280" pageNumber="281" pagination="1491 - 1492" refId="ref295465" refString="Nakabo, T. (2002) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. English Edition. Tokai University Press, Tokyo, pp. 417 - 435, 1491 - 1492. [In Japanese.]" type="journal article" year="2002">Nakabo 2002:420</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADC0DC71DFAB4787F" box="[1304,1363,1333,1354]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); 
<bibRefCitation id="5B297ED8FECE870ADC77C71DFC2A7859" author="Yoda, M. &amp; Tokimura, M. &amp; Horikawa, H. &amp; Yamada, U." pageId="280" pageNumber="281" refId="ref303555" refString="Yoda, M., Tokimura, M., Horikawa, H. &amp; Yamada, U. (2002) A Catalogue of Fishes from the East China and Yellow Seas with Their Local Names. Seikai National Fisheries Research Institute, Fisheries Research Agency, Nagasaki, 41 pp. [In Japanese.]" type="book" year="2002">
Yoda 
<emphasis id="0DCCDF3BFECE870ADC8FC71EFC9E7859" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
2002:11
</bibRefCitation>
(listed; East 
<collectingCountry id="47AF43B9FECE870ADD44C77FFB6D7858" box="[1105,1162,1367,1389]" name="China" pageId="280" pageNumber="281">China</collectingCountry>
and Yellow Seas); 
<bibRefCitation id="5B297ED8FECE870ADC5FC77FFC1278BA" author="Shinohara, G. &amp; Sato, T. &amp; Aonuma, Y. &amp; Horikawa, H. &amp; Matsuura, K. &amp; Nakabo, T. &amp; Sato, K." pageId="280" pageNumber="281" pagination="385 - 452" refId="ref300310" refString="Shinohara, G., Sato, T., Aonuma, Y., Horikawa, H., Matsuura, K., Nakabo, T. &amp; Sato, K. (2005) Annotated checklist of deep-sea fishes from the waters around the Ryukyu Islands, Japan. National Science Museum Tokyo Monographs, (29), 385 - 452." type="journal article" year="2005">
Shinohara 
<emphasis id="0DCCDF3BFECE870ADA4AC752FC7578BB" box="[863,914,1401,1423]" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
2005:418
</bibRefCitation>
(listed; Ryukyu Islands); 
<bibRefCitation id="5B297ED8FECE870ADDE3C751FA4F78BA" author="Shao, K. - T. &amp; Iwamoto, T. &amp; Ho, H. - C. &amp; Cheng, T. - Y. &amp; Chen, C. - Y." box="[1270,1448,1401,1423]" pageId="280" pageNumber="281" pagination="17 - 29" refId="ref299606" refString="Shao, K. - T., Iwamoto, T., Ho, H. - C., Cheng, T. - Y. &amp; Chen, C. - Y. (2008 b) Species composition and distribution pattern of grenadiers (family [sic] Bathygadidae, Macrouridae, and Macrourididae [sic]) from Taiwan. In: Orlov, A. M. &amp; Iwamoto, T. (Eds.), Grenadiers of the World Ocean: Biology, Stock Assessment, and Fisheries. American Fisheries Society Symposium 63. American Fisheries Society, Maryland, pp. 17 - 29." type="book chapter" year="2008">
Shao 
<emphasis id="0DCCDF3BFECE870ADC3BC752FA8678BB" box="[1326,1377,1401,1423]" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
2008b
</bibRefCitation>
: table 2 (
<specimenCount id="29BEC8A0FECE870ADABBC7B3FC147885" box="[942,1011,1435,1457]" pageId="280" pageNumber="281" type="generic">5 spec.</specimenCount>
listed from South 
<collectingCountry id="47AF43B9FECE870ADDB2C7B3FB077884" box="[1191,1248,1435,1457]" name="China" pageId="280" pageNumber="281">China</collectingCountry>
Sea); 
<bibRefCitation id="5B297ED8FECE870ADC0BC7B3FC5D78E6" author="Nakabo, T. &amp; Kai, Y." pageId="280" pageNumber="281" pagination="1872 - 1876" refId="ref295555" refString="Nakabo, T. &amp; Kai, Y. (2013) Macrouridae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species. 3 rd Edition. Tokai University Press, Hadano, pp. 493 - 512, 1872 - 1876. [In Japanese.]" type="journal article" year="2013">Nakabo &amp; Kai 2013:496</bibRefCitation>
(in key; 
<collectingCountry id="47AF43B9FECE870ADD0DC795FBB478E7" box="[1048,1107,1469,1490]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
); Iwamoto 
<emphasis id="0DCCDF3BFECE870ADDD0C796FB1F78E7" box="[1221,1272,1469,1491]" italics="true" pageId="280" pageNumber="281">et al.</emphasis>
2015:104, fig. 26 (brief description; 
<specimenCount id="29BEC8A0FECE870ADD08C7F7FB8078C1" box="[1053,1127,1503,1524]" pageId="280" pageNumber="281" type="generic">4 spec.</specimenCount>
from southwestern 
<collectingCountry id="47AF43B9FECE870ADC23C7F7FA9B78C0" box="[1334,1404,1503,1525]" name="Taiwan" pageId="280" pageNumber="281">Taiwan</collectingCountry>
and South 
<collectingCountry id="47AF43B9FECE870ADA88C429FC317B22" box="[925,982,1537,1559]" name="China" pageId="280" pageNumber="281">China</collectingCountry>
Sea); 
<bibRefCitation id="5B297ED8FECE870ADD01C429FB2A7B23" author="Motomura, H." box="[1044,1229,1537,1558]" pageId="280" pageNumber="281" refId="ref295068" refString="Motomura, H. (2020) List of Japan's All Fish Species. Current Standard Japanese and Scientific Names of All Fish Species Recorded from Japanese Waters. The Kagoshima University Museum, Kagoshima, 560 pp. [In Japanese.]" type="book" year="2020">Motomura 2020:39</bibRefCitation>
(listed; 
<collectingCountry id="47AF43B9FECE870ADC09C429FAB07B23" box="[1308,1367,1537,1558]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FECE870ADA3FC469FBFE7A7F" pageId="280" pageNumber="281" type="diagnosis">
<paragraph id="3F070329FECE870ADA3FC469FBFE7A7F" blockId="280.[810,1452,1601,2070]" pageId="280" pageNumber="281">
<emphasis id="0DCCDF3BFECE870ADA3FC469FC437B6E" bold="true" box="[810,932,1601,1627]" pageId="280" pageNumber="281">Diagnosis.</emphasis>
A species of 
<taxonomicName id="F8B878AAFECE870ADD7CC469FB1F7B6E" authorityName="Gunther" authorityYear="1887" box="[1129,1272,1601,1627]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFECE870ADD7CC469FB1F7B6E" box="[1129,1272,1601,1627]" italics="true" pageId="280" pageNumber="281">Trachonurus</emphasis>
</taxonomicName>
with grooved lateral line; pelvic-fin rays 7; gular and lower portions of branchiostegal membranes heavily scaled; outer series of premaxillary tooth band distinctly enlarged; pectoralfin rays i13–i17 (usually i15); inner gill rakers on first arch 10–12; body scales small, longitudinal scales 32–46; scales below first dorsal-fin midbase 4.5–8, below second dorsal-fin origin 5–9.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FECE8709DA4AC579FD607C18" lastPageId="283" lastPageNumber="284" pageId="280" pageNumber="281" type="materials_examined">
<paragraph id="3F070329FECE870BDA4AC579FBB47523" blockId="280.[810,1452,1601,2070]" lastBlockId="281.[810,1452,1669,2070]" lastPageId="281" lastPageNumber="282" pageId="280" pageNumber="281">
<emphasis id="0DCCDF3BFECE870ADA4AC579FBB67A5E" bold="true" box="[863,1105,1873,1899]" pageId="280" pageNumber="281">Material examined.</emphasis>
<materialsCitation id="8FD00974FECE870ADD74C57AFA4B7523" ID-GBIF-Occurrence="3750955487" collectingDate="1875-03-12" collectionCode="BMNH" country="Japan" county="Challenger" elevation="631" latitude="35.1833" location="Eno-shima Island" longLatPrecision="6" longitude="139.4667" municipality="Sagami Bay" pageId="280" pageNumber="281" specimenCode="BMNH 1887.12" specimenCount="10" stateProvince="Kanagawa" typeStatus="holotype">
<specimenCount id="29BEC8A0FECE870ADD74C57AFAE37A5E" box="[1121,1284,1873,1900]" pageId="280" pageNumber="281" type="generic">10 specimens</specimenCount>
. 
<emphasis id="0DCCDF3BFECE870ADC02C57AFBA87ABB" bold="true" pageId="280" pageNumber="281">
<typeStatus id="E003BD8BFECE870ADC02C57AFA637A59" box="[1303,1412,1874,1900]" pageId="280" pageNumber="281" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="F8B878AAFECE870ADA3FC55BFBA17ABB" authorityName="Gunther" authorityYear="1877" box="[810,1094,1907,1934]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="280" pageNumber="281" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECE870ADA3FC55BFBA17ABB" bold="true" box="[810,1094,1907,1934]" italics="true" pageId="280" pageNumber="281">Coryphaenoides villosus</emphasis>
</taxonomicName>
:
</emphasis>
* 
<specimenCode id="6F1EAB52FECE870ADD71C55CFAC47ABB" box="[1124,1315,1907,1934]" collectionCode="BMNH" pageId="280" pageNumber="281">BMNH 1887.12</specimenCode>
.7.105 (26.0 mm HL, 193+ mm TL), off 
<location id="3A6755F2FECE870ADD76C5BEFACD7A85" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECE870ADD76C5BEFACD7A85" box="[1123,1322,1942,1968]" country="Japan" county="Challenger" latitude="35.1833" longLatPrecision="6" longitude="139.4667" municipality="Sagami Bay" name="Eno-shima Island" pageId="280" pageNumber="281" stateProvince="Kanagawa">Eno-shima Island</location>
, 
<collectingRegion id="FD7CCDCBFECE870ADC20C5BEFC847AE7" country="Japan" name="Kanagawa" pageId="280" pageNumber="281">Kanagawa Pref.</collectingRegion>
, 
<collectingMunicipality id="DF639953FECE870ADA67C590FC187AE7" box="[882,1023,1976,2002]" pageId="280" pageNumber="281">Sagami Bay</collectingMunicipality>
, 
<collectingCountry id="47AF43B9FECE870ADD1AC590FBB57AE7" box="[1039,1106,1976,2002]" name="Japan" pageId="280" pageNumber="281">Japan</collectingCountry>
, 
<geoCoordinate id="5A8C65EEFECE870ADD77C59FFB3B7AE7" box="[1122,1244,1975,2002]" degrees="35.1833" direction="north" orientation="latitude" pageId="280" pageNumber="281" precision="5" value="35.1833">35.1833ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECE870ADDF9C59FFA957AE4" box="[1260,1394,1975,2001]" degrees="139.4667" direction="east" orientation="longitude" pageId="280" pageNumber="281" precision="5" value="139.4667">139.4667ºE</geoCoordinate>
, 345 ftm (
<quantity id="F840AECCFECE870ADA76C5F2FC567AC1" box="[867,945,2010,2036]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="280" pageNumber="281" unit="m" value="631.0">
<elevation id="B495E41AFECE870ADA76C5F2FC567AC1" box="[867,945,2010,2036]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="280" pageNumber="281" unit="m" value="631.0">631 m</elevation>
</quantity>
), 
<emphasis id="0DCCDF3BFECE870ADADDC5F2FBA17AC1" box="[968,1094,2010,2036]" italics="true" pageId="280" pageNumber="281">
<collectingCounty id="D6667BA5FECE870ADADDC5F2FBA17AC1" box="[968,1094,2010,2036]" pageId="280" pageNumber="281">Challenger</collectingCounty>
</emphasis>
sta. 232, trawl and dredge, 
<date id="4B0625E9FECE870ADC9AC5F2FC467523" pageId="280" pageNumber="281" value="1875-03-12">
<collectingDate id="5B42DC01FECE870ADC9AC5F2FC467523" pageId="280" pageNumber="281" value="1875-03-12">12 Mar. 1875</collectingDate>
</date>
(only photographed and briefly examined in
</materialsCitation>
this study). 
<emphasis id="0DCCDF3BFECF870BD803C4AEFE1F7B95" bold="true" box="[278,504,1670,1696]" pageId="281" pageNumber="282">
Non-types: 
<collectingCountry id="47AF43B9FECF870BD8B0C4AEFE157B95" box="[421,498,1670,1696]" name="Japan" pageId="281" pageNumber="282">Japan</collectingCountry>
:
</emphasis>
<materialsCitation id="8FD00974FECF870BDB11C4AEFE437A1D" ID-GBIF-Occurrence="3750954956" collectingDate="1978-01-23" collectionCode="BSKU" collectorName="S. Kishida &amp; Y. Kinoshita" elevation="725" latitude="26.4633" location="Trough" longLatPrecision="6" longitude="124.8483" municipality="Miyako-jima Island" pageId="281" pageNumber="282" specimenCode="BSKU 26555" specimenCount="1" stateProvince="Okinawa">
<specimenCode id="6F1EAB52FECF870BDB11C4AEFD467B95" box="[516,673,1669,1696]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="281" pageNumber="282">BSKU 26555</specimenCode>
(1, 
<quantity id="F840AECCFECF870BDBCCC4ADFF537BF7" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="281" pageNumber="282" unit="mm" value="25.8">25.8 mm</quantity>
HL, 114+ mm TL), off 
<collectingMunicipality id="DF639953FECF870BD8AEC48FFD7D7BF4" box="[443,666,1703,1729]" pageId="281" pageNumber="282">Miyako-jima Island</collectingMunicipality>
, 
<collectingRegion id="FD7CCDCBFECF870BDBB0C48FFCED7BF4" box="[677,778,1703,1729]" country="Japan" name="Okinawa" pageId="281" pageNumber="282">Okinawa</collectingRegion>
<location id="3A6755F2FECF870BD99DC4E1FF3B7BD6" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BD99DC4E1FF3B7BD6" box="[136,220,1737,1763]" latitude="26.4633" longLatPrecision="6" longitude="124.8483" municipality="Miyako-jima Island" name="Trough" pageId="281" pageNumber="282" stateProvince="Okinawa">Trough</location>
, 
<geoCoordinate id="5A8C65EEFECF870BD9FDC4E1FE847BD6" box="[232,355,1737,1763]" degrees="26.4633" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="26.4633">26.4633ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BD87BC4E2FE137BD1" box="[366,500,1738,1764]" degrees="124.8483" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="124.8483">124.8483ºE</geoCoordinate>
, 
<quantity id="F840AECCFECF870BDB15C4E1FD667BD1" box="[512,641,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="281" pageNumber="282" unit="m" value="725.0" valueMax="770.0" valueMin="680.0">
<elevation id="B495E41AFECF870BDB15C4E1FD667BD1" box="[512,641,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="281" pageNumber="282" unit="m" value="725.0" valueMax="770.0" valueMin="680.0">680–770 m</elevation>
</quantity>
, F/ 
<collectorName id="924D66FFFECF870BDBB0C4E2FF1C7A30" pageId="281" pageNumber="282">
V 
<emphasis id="0DCCDF3BFECF870BDBD5C4E2FF1C7A30" italics="true" pageId="281" pageNumber="282">Yuryomaru, No.</emphasis>
</collectorName>
8, tr. 10, bottom trawl, coll. 
<collectorName id="924D66FFFECF870BDB51C4C4FD5D7A30" box="[580,698,1771,1798]" pageId="281" pageNumber="282">S. Kishida</collectorName>
and 
<collectorName id="924D66FFFECF870BDBE7C4C4FF1F7A12" pageId="281" pageNumber="282">Y. Kinoshita</collectorName>
, 
<date id="4B0625E9FECF870BD81DC526FE437A1D" box="[264,420,1806,1832]" pageId="281" pageNumber="282" value="1978-01-23">
<collectingDate id="5B42DC01FECF870BD81DC526FE437A1D" box="[264,420,1806,1832]" pageId="281" pageNumber="282" value="1978-01-23">23 Jan. 1978</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BD8A0C526FF1F7ABB" ID-GBIF-Occurrence="3750955348" collectingDate="1877-06-18" collectorName="V Tansei-maru" elevation="918" location="Trough" pageId="281" pageNumber="282" specimenCount="1" stateProvince="Okinawa">
NSMT-P 93826 (1, 
<quantity id="F840AECCFECF870BDBB4C525FCED7A1D" box="[673,778,1805,1832]" metricMagnitude="-2" metricUnit="m" metricValue="5.77" pageId="281" pageNumber="282" unit="mm" value="57.7">57.7 mm</quantity>
HL, 362+ mm TL), northern 
<collectingRegion id="FD7CCDCBFECF870BD8CDC507FDDA7A7C" box="[472,573,1839,1865]" country="Japan" name="Okinawa" pageId="281" pageNumber="282">Okinawa</collectingRegion>
<location id="3A6755F2FECF870BDB53C507FD7D7A7C" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDB53C507FD7D7A7C" box="[582,666,1839,1865]" name="Trough" pageId="281" pageNumber="282" stateProvince="Okinawa">Trough</location>
, 
<quantity id="F840AECCFECF870BDBBDC518FF477A59" metricMagnitude="2" metricUnit="m" metricValue="9.18" metricValueMax="9.19" metricValueMin="9.17" pageId="281" pageNumber="282" unit="m" value="918.0" valueMax="919.0" valueMin="917.0">
<elevation id="B495E41AFECF870BDBBDC518FF477A59" metricMagnitude="2" metricUnit="m" metricValue="9.18" metricValueMax="9.19" metricValueMin="9.17" pageId="281" pageNumber="282" unit="m" value="918.0" valueMax="919.0" valueMin="917.0">917–919 m</elevation>
</quantity>
, R/ 
<collectorName id="924D66FFFECF870BD9DCC57AFE937A5E" box="[201,372,1874,1900]" pageId="281" pageNumber="282">
V 
<emphasis id="0DCCDF3BFECF870BD9F2C57AFE937A5E" box="[231,372,1874,1899]" italics="true" pageId="281" pageNumber="282">Tansei-maru</emphasis>
</collectorName>
, cr. KT-93-09, 3-m beam trawl, 
<date id="4B0625E9FECF870BDBFBC57AFF1F7ABB" pageId="281" pageNumber="282" value="1877-06-18">
<collectingDate id="5B42DC01FECF870BDBFBC57AFF1F7ABB" pageId="281" pageNumber="282" value="1877-06-18">18 Jun. 1877</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BD816C55CFCE27AE7" ID-GBIF-Occurrence="3750955313" collectingDate="1975-01-26" collectionCode="BSKU" collectorName="Kaiyomaru" elevation="1043" latitude="32.98" location="Kochi" longLatPrecision="6" longitude="133.855" pageId="281" pageNumber="282" specimenCode="BSKU 23083" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BD816C55CFE7C7ABB" box="[259,411,1908,1934]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="281" pageNumber="282">BSKU 23083</specimenCode>
(1, 
<quantity id="F840AECCFECF870BD8D2C55BFDCC7ABB" box="[455,555,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="281" pageNumber="282" unit="mm" value="45.2">45.2 mm</quantity>
HL, 290+ mm TL), off 
<collectingRegion id="FD7CCDCBFECF870BD9A5C5BEFF107A85" box="[176,247,1942,1968]" country="Japan" name="Kochi" pageId="281" pageNumber="282">Kochi</collectingRegion>
, 
<geoCoordinate id="5A8C65EEFECF870BD811C5BEFE987A85" box="[260,383,1942,1968]" degrees="32.9800" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="32.98">32.9800ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BD89EC5BDFDF67A85" box="[395,529,1941,1968]" degrees="133.8550" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="133.855">133.8550ºE</geoCoordinate>
, 
<quantity id="F840AECCFECF870BDB09C5BEFD937A85" box="[540,628,1942,1968]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="281" pageNumber="282" unit="m" value="1043.0">
<elevation id="B495E41AFECF870BDB09C5BEFD937A85" box="[540,628,1942,1968]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="281" pageNumber="282" unit="m" value="1043.0">1043 m</elevation>
</quantity>
, FRV 
<emphasis id="0DCCDF3BFECF870BDBABC5BEFF247AE4" italics="true" pageId="281" pageNumber="282">
<collectorName id="924D66FFFECF870BDBABC5BEFF247AE4" pageId="281" pageNumber="282">Kaiyomaru</collectorName>
</emphasis>
, cr. SK-75-2, sta. 9, tr. 2, bottom trawl, 
<date id="4B0625E9FECF870BDB63C59FFCE27AE7" box="[630,773,1975,2002]" pageId="281" pageNumber="282" value="1975-01-26">
<collectingDate id="5B42DC01FECF870BDB63C59FFCE27AE7" box="[630,773,1975,2002]" pageId="281" pageNumber="282" value="1975-01-26">26 Jan. 1975</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BD99DC5F2FAF37B95" ID-GBIF-Occurrence="3750954301" collectingDate="1975-01-26" collectionCode="BSKU" collectorName="Kaiyo-maru" elevation="945" latitude="33.035" location="Tosa Bay" longLatPrecision="6" longitude="133.75" pageId="281" pageNumber="282" specimenCode="BSKU 23055" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BD99DC5F2FEC77AC1" box="[136,288,2009,2036]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="281" pageNumber="282">BSKU 23055</specimenCode>
(1, 
<quantity id="F840AECCFECF870BD85BC5F2FE547AC1" box="[334,435,2010,2036]" metricMagnitude="-2" metricUnit="m" metricValue="6.01" pageId="281" pageNumber="282" unit="mm" value="60.1">60.1 mm</quantity>
HL, 383+ mm TL), 
<location id="3A6755F2FECF870BDB8EC5F2FCE27AC1" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDB8EC5F2FCE27AC1" box="[667,773,2010,2036]" latitude="33.035" longLatPrecision="6" longitude="133.75" name="Tosa Bay" pageId="281" pageNumber="282" stateProvince="Kochi">Tosa Bay</location>
, 
<geoCoordinate id="5A8C65EEFECF870BD99DC5D3FEE47523" box="[136,259,2043,2070]" degrees="33.0350" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="33.035">33.0350ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BD818C5D3FE747523" box="[269,403,2043,2070]" degrees="133.7500" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="133.75">133.7500ºE</geoCoordinate>
, 
<quantity id="F840AECCFECF870BD889C5D3FE037523" box="[412,484,2043,2070]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="281" pageNumber="282" unit="m" value="945.0">
<elevation id="B495E41AFECF870BD889C5D3FE037523" box="[412,484,2043,2070]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="281" pageNumber="282" unit="m" value="945.0">945 m</elevation>
</quantity>
, FRV 
<emphasis id="0DCCDF3BFECF870BDB3DC5D4FD487520" box="[552,687,2044,2069]" italics="true" pageId="281" pageNumber="282">
<collectorName id="924D66FFFECF870BDB3DC5D4FD487520" box="[552,687,2044,2069]" pageId="281" pageNumber="282">Kaiyo-maru</collectorName>
</emphasis>
, cr. SK-75-2, sta. 18, tr. 3, bottom trawl, 
<date id="4B0625E9FECF870BDD9DC4ADFAF37B95" box="[1160,1300,1669,1696]" pageId="281" pageNumber="282" value="1975-01-26">
<collectingDate id="5B42DC01FECF870BDD9DC4ADFAF37B95" box="[1160,1300,1669,1696]" pageId="281" pageNumber="282" value="1975-01-26">26 Jan. 1975</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BDC08C4AEFB017BD1" ID-GBIF-Occurrence="3750955123" collectingDate="1988-09-13" collectionCode="SNFR" collectorName="Kotaka-maru" elevation="750" location="Tosa Bay" pageId="281" pageNumber="282" specimenCount="1" stateProvince="Kochi">
<collectionCode id="59A99BECFECF870BDC08C4AEFA847B95" box="[1309,1379,1670,1696]" pageId="281" pageNumber="282">SNFR</collectionCode>
uncat. (1, 
<quantity id="F840AECCFECF870BDA44C48FFC507BF7" box="[849,951,1703,1730]" metricMagnitude="-2" metricUnit="m" metricValue="5.63" pageId="281" pageNumber="282" unit="mm" value="56.3">56.3 mm</quantity>
HL, 328+ mm TL), 
<location id="3A6755F2FECF870BDDB3C480FAF67BF7" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDDB3C480FAF67BF7" box="[1190,1297,1704,1730]" name="Tosa Bay" pageId="281" pageNumber="282" stateProvince="Kochi">Tosa Bay</location>
, 
<quantity id="F840AECCFECF870BDC0AC48FFA8E7BF7" box="[1311,1385,1703,1730]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="281" pageNumber="282" unit="m" value="750.0">
<elevation id="B495E41AFECF870BDC0AC48FFA8E7BF7" box="[1311,1385,1703,1730]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="281" pageNumber="282" unit="m" value="750.0">750 m</elevation>
</quantity>
, FRV 
<emphasis id="0DCCDF3BFECF870BDA3FC4E1FC587BD6" box="[810,959,1737,1763]" italics="true" pageId="281" pageNumber="282">
<collectorName id="924D66FFFECF870BDA3FC4E1FC587BD6" box="[810,959,1737,1763]" pageId="281" pageNumber="282">Kotaka-maru</collectorName>
</emphasis>
, otter trawl, 
<date id="4B0625E9FECF870BDD5EC4E2FB017BD1" box="[1099,1254,1738,1764]" pageId="281" pageNumber="282" value="1988-09-13">
<collectingDate id="5B42DC01FECF870BDD5EC4E2FB017BD1" box="[1099,1254,1738,1764]" pageId="281" pageNumber="282" value="1988-09-13">13 Sept. 1988</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BDDE4C4E2FC477A5E" ID-GBIF-Occurrence="3750955535" collectingDate="1986-11-02" collectionCode="BSKU" collectorName="V Tansei-maru" elevation="1084" latitude="32.545" location="Okinoshima Island" longLatPrecision="6" longitude="132.475" pageId="281" pageNumber="282" specimenCode="BSKU 43458" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BDDE4C4E2FA607BD1" box="[1265,1415,1737,1764]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="281" pageNumber="282">BSKU 43458</specimenCode>
(1, 
<quantity id="F840AECCFECF870BDA3FC4C4FC687A33" box="[810,911,1772,1798]" metricMagnitude="-2" metricUnit="m" metricValue="7.97" pageId="281" pageNumber="282" unit="mm" value="79.7">79.7 mm</quantity>
HL, 463+ mm TL), south-southwest of 
<location id="3A6755F2FECF870BDC49C4C3FC5B7A12" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDC49C4C3FC5B7A12" latitude="32.545" longLatPrecision="6" longitude="132.475" name="Okinoshima Island" pageId="281" pageNumber="282" stateProvince="Kochi">Okinoshima Island</location>
, 
<geoCoordinate id="5A8C65EEFECF870BDADFC525FBA27A1D" box="[970,1093,1805,1832]" degrees="32.5450" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="32.545">32.5450ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BDD47C525FB3F7A1D" box="[1106,1240,1805,1832]" degrees="132.4750" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="132.475">132.4750ºE</geoCoordinate>
, 
<quantity id="F840AECCFECF870BDDF1C525FA647A1D" box="[1252,1411,1805,1832]" metricMagnitude="3" metricUnit="m" metricValue="1.0835" metricValueMax="1.092" metricValueMin="1.075" pageId="281" pageNumber="282" unit="m" value="1083.5" valueMax="1092.0" valueMin="1075.0">
<elevation id="B495E41AFECF870BDDF1C525FA647A1D" box="[1252,1411,1805,1832]" metricMagnitude="3" metricUnit="m" metricValue="1.0835" metricValueMax="1.092" metricValueMin="1.075" pageId="281" pageNumber="282" unit="m" value="1083.5" valueMax="1092.0" valueMin="1075.0">1075–1092 m</elevation>
</quantity>
, R/ 
<collectorName id="924D66FFFECF870BDA3FC518FC337A7C" box="[810,980,1840,1866]" pageId="281" pageNumber="282">
V 
<emphasis id="0DCCDF3BFECF870BDA52C518FC337A7C" box="[839,980,1840,1865]" italics="true" pageId="281" pageNumber="282">Tansei-maru</emphasis>
</collectorName>
, cr. KT-86-16, sta. C, 3-m beam trawl, 
<date id="4B0625E9FECF870BDC88C518FC477A5E" pageId="281" pageNumber="282" value="1986-11-02">
<collectingDate id="5B42DC01FECF870BDC88C518FC477A5E" pageId="281" pageNumber="282" value="1986-11-02">2 Nov. 1986</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BDABEC57AFB797A85" ID-GBIF-Occurrence="3750954548" collectingDate="1965-10-05" collectionCode="BSKU" collectorName="Soyo-maru" latitude="34.3333" location="Izu Peninsula" longLatPrecision="6" longitude="138.8167" pageId="281" pageNumber="282" specimenCode="BSKU 20536" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BDABEC57AFBA57A59" box="[939,1090,1873,1900]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="281" pageNumber="282">BSKU 20536</specimenCode>
(1, 
<quantity id="F840AECCFECF870BDD79C57AFB377A59" box="[1132,1232,1874,1900]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="281" pageNumber="282" unit="mm" value="37.2">37.2 mm</quantity>
HL, 229+ mm TL), south of 
<location id="3A6755F2FECF870BDA8EC55CFBD97AB8" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDA8EC55CFBD97AB8" box="[923,1086,1907,1934]" latitude="34.3333" longLatPrecision="6" longitude="138.8167" name="Izu Peninsula" pageId="281" pageNumber="282" stateProvince="Kochi">Izu Peninsula</location>
, 
<geoCoordinate id="5A8C65EEFECF870BDD44C55CFB2C7ABB" box="[1105,1227,1908,1934]" degrees="34.3333" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="34.3333">34.3333ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BDDCBC55BFA837AB8" box="[1246,1380,1907,1933]" degrees="138.8167" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="138.8167">138.8167ºE</geoCoordinate>
, FRV 
<emphasis id="0DCCDF3BFECF870BDA3FC5BEFC427A85" box="[810,933,1942,1968]" italics="true" pageId="281" pageNumber="282">
<collectorName id="924D66FFFECF870BDA3FC5BEFC427A85" box="[810,933,1942,1968]" pageId="281" pageNumber="282">Soyo-maru</collectorName>
</emphasis>
, sta. 30, 
<date id="4B0625E9FECF870BDD07C5BDFB797A85" box="[1042,1182,1941,1968]" pageId="281" pageNumber="282" value="1965-10-05">
<collectingDate id="5B42DC01FECF870BDD07C5BDFB797A85" box="[1042,1182,1941,1968]" pageId="281" pageNumber="282" value="1965-10-05">5 Oct. 1965</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="8FD00974FECF870BDDB8C5BEFBDE7AE7" ID-GBIF-Occurrence="3750955472" collectionCode="HUMZ" pageId="281" pageNumber="282" specimenCode="HUMZ 71882" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BDDB8C5BEFAA87A85" box="[1197,1359,1942,1968]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="281" pageNumber="282">HUMZ 71882</specimenCode>
(
<quantity id="F840AECCFECF870BDC70C5BEFCB17AE7" metricMagnitude="0" metricUnit="m" metricValue="1.124" pageId="281" pageNumber="282" unit="mm" value="1124.0">1, 124 mm</quantity>
HL, 495+ mm TL)
</materialsCitation>
, 
<materialsCitation id="8FD00974FECF870BDD53C590FBA87523" ID-GBIF-Occurrence="3750954400" collectingDate="1977-10-10" collectionCode="HUMZ" latitude="23.8667" location="Minami-torishima Island" longLatPrecision="7" longitude="150.6667" pageId="281" pageNumber="282" specimenCode="HUMZ 71883" specimenCount="1" stateProvince="Kochi">
<specimenCode id="6F1EAB52FECF870BDD53C590FB0F7AE7" box="[1094,1256,1976,2002]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="281" pageNumber="282">HUMZ 71883</specimenCode>
(
<quantity id="F840AECCFECF870BDDE9C590FA917AE7" box="[1276,1398,1976,2002]" metricMagnitude="0" metricUnit="m" metricValue="1.1099999999999999" pageId="281" pageNumber="282" unit="mm" value="1110.0">1, 110 mm</quantity>
HL, 675+ mm TL), off 
<location id="3A6755F2FECF870BDD11C5F2FAF87AC1" LSID="urn:lsid:plazi:treatment:B711B23FFECE8709DA3FC284FC237E3C:3A6755F2FECF870BDD11C5F2FAF87AC1" box="[1028,1311,2010,2036]" latitude="23.8667" longLatPrecision="7" longitude="150.6667" name="Minami-torishima Island" pageId="281" pageNumber="282" stateProvince="Kochi">Minami-torishima Island</location>
, 
<geoCoordinate id="5A8C65EEFECF870BDC39C5F2FA407AC1" box="[1324,1447,2010,2036]" degrees="23.8667" direction="north" orientation="latitude" pageId="281" pageNumber="282" precision="5" value="23.8667">23.8667ºN</geoCoordinate>
, 
<geoCoordinate id="5A8C65EEFECF870BDA3FC5D3FC487523" box="[810,943,2043,2070]" degrees="150.6667" direction="east" orientation="longitude" pageId="281" pageNumber="282" precision="5" value="150.6667">150.6667ºE</geoCoordinate>
, 
<date id="4B0625E9FECF870BDAAFC5D4FBA87523" box="[954,1103,2044,2070]" pageId="281" pageNumber="282" value="1977-10-10">
<collectingDate id="5B42DC01FECF870BDAAFC5D4FBA87523" box="[954,1103,2044,2070]" pageId="281" pageNumber="282" value="1977-10-10">10 Oct. 1977</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="6BC753A1FECF870BD99DC793FE7B7B0C" ID-DOI="http://doi.org/10.5281/zenodo.6423202" ID-Zenodo-Dep="6423202" httpUri="https://zenodo.org/record/6423202/files/figure.png" pageId="281" pageNumber="282" startId="281.[136,235,1467,1491]" targetBox="[151,1436,158,1428]" targetPageId="281">
<paragraph id="3F070329FECF870BD99DC793FE7B7B0C" blockId="281.[136,1452,1467,1593]" pageId="281" pageNumber="282">
<emphasis id="0DCCDF3BFECF870BD99DC793FEF978E6" bold="true" box="[136,286,1467,1491]" pageId="281" pageNumber="282">FIGURE 193.</emphasis>
Fresh specimens of 
<taxonomicName id="F8B878AAFECF870BD8E7C793FD2F78E6" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[498,712,1467,1491]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="281" pageNumber="282" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECF870BD8E7C793FD2F78E6" box="[498,712,1467,1491]" italics="true" pageId="281" pageNumber="282">Trachonurus villosus</emphasis>
</taxonomicName>
. (A) BSKU 23083, 45.2 mm HL, 290+ mm TL, off Kochi, Kochi Pref., Pacific, 1043 m depth; (B) BSKU 43458, 79.7 mm HL, 463+ mm TL, south-southwest of Okino-shima Island, Kochi Pref., Pacific, 1075–1092 m depth; (C) HUMZ 71883, 110 mm HL, 675+ mm TL, off Minami-torishima Island, Pacific. Lateral views. [Photos: (A–B) BSKU; (C) HUMZ]
</paragraph>
</caption>
<caption id="6BC753A1FECC8708D99DC085FF1A7E3C" ID-DOI="http://doi.org/10.5281/zenodo.6423204" ID-Zenodo-Dep="6423204" httpUri="https://zenodo.org/record/6423204/files/figure.png" pageId="282" pageNumber="283" startId="282.[136,235,685,709]" targetBox="[151,1436,159,652]" targetPageId="282">
<paragraph id="3F070329FECC8708D99DC085FF1A7E3C" blockId="282.[136,1452,685,777]" pageId="282" pageNumber="283">
<emphasis id="0DCCDF3BFECC8708D99DC085FEF87FF0" bold="true" box="[136,287,685,709]" pageId="282" pageNumber="283">FIGURE 194.</emphasis>
Holotype of 
<taxonomicName id="F8B878AAFECC8708D8B9C085FD407FF0" authorityName="Gunther" authorityYear="1877" box="[428,679,685,709]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="282" pageNumber="283" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECC8708D8B9C085FD407FF0" box="[428,679,685,709]" italics="true" pageId="282" pageNumber="283">Coryphaenoides villosus</emphasis>
</taxonomicName>
(= 
<taxonomicName id="F8B878AAFECC8708DBDAC085FC417FF0" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[719,934,685,709]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="282" pageNumber="283" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECC8708DBDAC085FC417FF0" box="[719,934,685,709]" italics="true" pageId="282" pageNumber="283">Trachonurus villosus</emphasis>
</taxonomicName>
). BMNH 1887.12.7.105, 26.0 mm HL, 193+ mm TL, Sagami Bay, Pacific, 631 m depth. (A) Lateral view; (B) dorsal and (C) ventral views of the head and body. [Photos: N. Nakayama]
</paragraph>
</caption>
<caption id="6BC753A1FECC8708D99DC75FFB747884" ID-DOI="http://doi.org/10.5281/zenodo.6423206" ID-Zenodo-Dep="6423206" httpUri="https://zenodo.org/record/6423206/files/figure.png" pageId="282" pageNumber="283" startId="282.[136,235,1399,1423]" targetBox="[151,1436,850,1360]" targetPageId="282">
<paragraph id="3F070329FECC8708D99DC75FFB747884" blockId="282.[136,1452,1399,1457]" pageId="282" pageNumber="283">
<emphasis id="0DCCDF3BFECC8708D99DC75FFEFB78BA" bold="true" box="[136,284,1399,1423]" pageId="282" pageNumber="283">FIGURE 195.</emphasis>
Scanning electron micrographs showing a body scale (from the dorsum below the interdorsal space) of 
<taxonomicName id="F8B878AAFECC8708DC3FC75FFF327884" baseAuthorityName="Gunther" baseAuthorityYear="1877" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="282" pageNumber="283" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECC8708DC3FC75FFF327884" italics="true" pageId="282" pageNumber="283">Trachonurus villosus</emphasis>
</taxonomicName>
. BSKU 43458, 79.7 mm HL. (A) View from above; (B) oblique view. [Photos: N. Nakayama]
</paragraph>
</caption>
<paragraph id="3F070329FECC8708D9A8C7D5FD357A33" blockId="282.[136,778,1533,2071]" pageId="282" pageNumber="283">
<emphasis id="0DCCDF3BFECC8708D9A8C7D5FDE27B2D" bold="true" box="[189,517,1533,1560]" pageId="282" pageNumber="283">Counts and measurements.</emphasis>
Based on 
<specimenCount id="29BEC8A0FECC8708DB6AC7D6FCED7B22" box="[639,778,1533,1560]" pageId="282" pageNumber="283" type="generic">9 specimens</specimenCount>
(
<quantity id="F840AECCFECC8708D985C437FED07B0F" box="[144,311,1567,1594]" metricMagnitude="-2" metricUnit="m" metricValue="7.49" metricValueMax="12.4" metricValueMin="2.58" pageId="282" pageNumber="283" unit="mm" value="74.9" valueMax="124.0" valueMin="25.8">25.8–124 mm</quantity>
HL, 
<quantity id="F840AECCFECC8708D891C437FDC57B0F" box="[388,546,1567,1594]" metricMagnitude="-1" metricUnit="m" metricValue="3.945" metricValueMax="6.75" metricValueMin="1.14" pageId="282" pageNumber="283" unit="mm" value="394.5" valueMax="675.0" valueMin="114.0">114–675 mm</quantity>
TL). Counts: first dorsal-fin rays II,7–9; pectoral-fin rays i13–i17; pelvic-fin rays 7; gill rakers on first arch (outer/inner) 4–9/10–12, on second arch 9–14/10–12; longitudinal scales 32–46; transverse scale rows below first dorsal-fin origin 6.5– 8.5, below first dorsal-fin midbase 4.5–8, below second dorsal-fin origin 5–9, above anal-fin origin 22–33.5.
</paragraph>
<paragraph id="3F070329FECC8709D9A8C525FE927DDF" blockId="282.[136,778,1533,2071]" lastBlockId="283.[136,778,173,2070]" lastPageId="283" lastPageNumber="284" pageId="282" pageNumber="283">The following measurements are in % of HL, followed by those in % of PRL in parentheses: snout length 24–29 (31–40); orbit diameter 24–33 (32–42); postorbital length 46–55 (60–71); postrostral length 73–79 (–); orbit–preopercle distance 30–39 (39–51); suborbital width 10–14 (14–18); upper-jaw length 31– 38 (41–49); length of rictus 26–33 (35–42); length of premaxillary tooth band 20–26 (26–34); preoral length 12–16 (16–21); distance between tip and lateral angle of snout 10–16 (13–21); snout width 18–24 (23–31); internasal width 19–23 (25–32); interorbital width 32– 38 (43–51); body width over pectoral-fin bases 38–66 (50–90); body depth at first dorsal-fin origin 66–104 (89–135); body depth at anal-fin origin 52–93 (68–119); prepelvic length 114–134 (149–181); preanus length 137–165 (180–214); preanal length 144–172 (189–230); isthmus–pelvic distance 55–65 (72–88); isthmus–anus distance 74–95 (98–123); isthmus–anal distance 82–108 (108–140); pelvic–anal distance 26–46 (34–58); anusanal distance 8–15 (10–20); pelvic-fin length 29–45 (38–60); pectoral-fin length 42–54 (56–71); predorsal length 109–121 (143–164); height of first dorsal fin 51– 65 (67–87); length of first dorsal-fin base 16–23 (21–32); interdorsal length 27–45 (36–61); length of gill slit 14–20 (19–25); length of posterior nostril 5–10 (7–14); barbel length 9–12 (11–16).</paragraph>
<paragraph id="3F070329FECD8709D9A8C2D8FD607C18" blockId="283.[136,778,173,2070]" pageId="283" pageNumber="284">
<emphasis id="0DCCDF3BFECD8709D9A8C2D8FF127C3F" bold="true" box="[189,245,240,266]" pageId="283" pageNumber="284">Size.</emphasis>
Attains about 
<quantity id="F840AECCFECD8709D8B6C2D9FE0B7C3E" box="[419,492,241,267]" metricMagnitude="-1" metricUnit="m" metricValue="6.8" pageId="283" pageNumber="284" unit="cm" value="68.0">68 cm</quantity>
TL (
<collectionCode id="59A99BECFECD8709DB3CC2D9FD9A7C3E" box="[553,637,241,267]" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="283" pageNumber="284">HUMZ</collectionCode>
<quantity id="F840AECCFECD8709DB9DC2D9FF537C18" metricMagnitude="4" metricUnit="m" metricValue="7.1883675" pageId="283" pageNumber="284" unit="mm" value="7.1883675E7">71883, 675 mm</quantity>
TL, off Minami-torishima Island, 
<collectingCountry id="47AF43B9FECD8709DB22C33BFD9B7C18" box="[567,636,275,301]" name="Japan" pageId="283" pageNumber="284">Japan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FECD8709D9A8C31DFDEA7F6A" pageId="283" pageNumber="284" type="distribution">
<paragraph id="3F070329FECD8709D9A8C31DFDEA7F6A" blockId="283.[136,778,173,2070]" pageId="283" pageNumber="284">
<emphasis id="0DCCDF3BFECD8709D9A8C31DFEB07C7A" bold="true" box="[189,343,309,335]" pageId="283" pageNumber="284">Distribution.</emphasis>
RecordedfromthenorthwesternPacific off 
<collectingCountry id="47AF43B9FECD8709D9A6C37FFF147C44" box="[179,243,343,369]" name="Japan" pageId="283" pageNumber="284">Japan</collectingCountry>
and 
<collectingCountry id="47AF43B9FECD8709D824C37FFE647C44" box="[305,387,343,369]" name="Taiwan" pageId="283" pageNumber="284">Taiwan</collectingCountry>
as well as from the southeastern Pacific at the Sala y Gomez Ridge; depth range 
<quantity id="F840AECCFECD8709DB8FC351FF797C80" metricMagnitude="2" metricUnit="m" metricValue="8.77" metricValueMax="12.4" metricValueMin="5.14" pageId="283" pageNumber="284" unit="m" value="877.0" valueMax="1240.0" valueMin="514.0">514–1240 m</quantity>
(
<bibRefCitation id="5B297ED8FECD8709D9B8C3B3FE3C7C80" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[173,475,411,437]" pageId="283" pageNumber="284" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto 1992</bibRefCitation>
; 
<bibRefCitation id="5B297ED8FECD8709D8FDC3B3FD487C80" author="Shao, K. - T. &amp; Ho, H. - C. &amp; Lin, P. - L. &amp; Lee, P. - F. &amp; Lee, M. - Y. &amp; Tsai, C. - Y. &amp; Liao, Y. - C. &amp; Lin, Y. - C. &amp; Chen, J. - P. &amp; Yeh, H. - M." box="[488,687,411,437]" pageId="283" pageNumber="284" pagination="233 - 271" refId="ref299493" refString="Shao, K. - T., Ho, H. - C., Lin, P. - L., Lee, P. - F., Lee, M. - Y., Tsai, C. - Y., Liao, Y. - C., Lin, Y. - C., Chen, J. - P. &amp; Yeh, H. - M. (2008 a) A checklist of the fishes of southern Taiwan, northern South China Sea. The Raffles Bulletin of Zoology, (suppl. 19), 233 - 271." type="journal article" year="2008">
Shao 
<emphasis id="0DCCDF3BFECD8709DB3DC3B4FD867C80" box="[552,609,411,437]" italics="true" pageId="283" pageNumber="284">et al.</emphasis>
2008a
</bibRefCitation>
, 
<bibRefCitation id="5B297ED8FECD8709DBA9C3B3FCE27C80" author="Shao, K. - T. &amp; Iwamoto, T. &amp; Ho, H. - C. &amp; Cheng, T. - Y. &amp; Chen, C. - Y." box="[700,773,411,437]" pageId="283" pageNumber="284" pagination="17 - 29" refId="ref299606" refString="Shao, K. - T., Iwamoto, T., Ho, H. - C., Cheng, T. - Y. &amp; Chen, C. - Y. (2008 b) Species composition and distribution pattern of grenadiers (family [sic] Bathygadidae, Macrouridae, and Macrourididae [sic]) from Taiwan. In: Orlov, A. M. &amp; Iwamoto, T. (Eds.), Grenadiers of the World Ocean: Biology, Stock Assessment, and Fisheries. American Fisheries Society Symposium 63. American Fisheries Society, Maryland, pp. 17 - 29." type="book chapter" year="2008">2008b</bibRefCitation>
; this study). In Japanese waters, known from the Pacific off southern 
<collectingCountry id="47AF43B9FECD8709D830C3F7FE827CCC" box="[293,357,479,505]" name="Japan" pageId="283" pageNumber="284">Japan</collectingCountry>
northward to the Miura Peninsula (
<geoCoordinate id="5A8C65EEFECD8709D985C029FF1B7F29" box="[144,252,513,540]" degrees="139.62" direction="east" orientation="longitude" pageId="283" pageNumber="284" precision="555" value="139.62">139.62ºE</geoCoordinate>
), 
<collectingRegion id="FD7CCDCBFECD8709D807C029FE907F2E" box="[274,375,513,539]" country="Japan" name="Okinawa" pageId="283" pageNumber="284">Okinawa</collectingRegion>
Trough, and off Minami-torishima Island, at depths of 631‾ 
<quantity id="F840AECCFECD8709D8BAC00BFDEF7F08" box="[431,520,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.187" pageId="283" pageNumber="284" unit="m" value="1187.0">1187 m</quantity>
(Appendix 3-12B). It appears to be rare wherever found.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FECD8709D9A8C04FFC237E3C" pageId="283" pageNumber="284" type="discussion">
<paragraph id="3F070329FECD8709D9A8C04FFE0B781F" blockId="283.[136,778,173,2070]" pageId="283" pageNumber="284">
<emphasis id="0DCCDF3BFECD8709D9A8C04FFDD97FB4" bold="true" box="[189,574,615,641]" pageId="283" pageNumber="284">Comments on type specimen.</emphasis>
G̹nther (1877) originally described the species (as 
<taxonomicName id="F8B878AAFECD8709DB47C0A1FF397FF0" authorityName="Gunther" authorityYear="1877" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DB47C0A1FF397FF0" italics="true" pageId="283" pageNumber="284">Coryphaenoides villosus</emphasis>
</taxonomicName>
) based on a single small specimen (BMNH 1887.12.7.105, 26 mm HL, 193+ mm TL; 
<figureCitation id="A7831FACFECD8709DB8EC0E5FCE67FD2" box="[667,769,717,743]" captionStart="FIGURE 194" captionStartId="282.[136,235,685,709]" captionTargetBox="[151,1436,159,652]" captionTargetId="figure-256@282.[151,1436,159,652]" captionTargetPageId="282" captionText="FIGURE 194. Holotype of Coryphaenoides villosus (= Trachonurus villosus). BMNH 1887.12.7.105, 26.0 mm HL, 193+ mm TL, Sagami Bay, Pacific, 631 m depth. (A) Lateral view; (B) dorsal and (C) ventral views of the head and body. [Photos: N. Nakayama]" figureDoi="http://doi.org/10.5281/zenodo.6423204" httpUri="https://zenodo.org/record/6423204/files/figure.png" pageId="283" pageNumber="284">Fig. 194</figureCitation>
) collected from “south of Yeddo”, 
<collectingCountry id="47AF43B9FECD8709DB1AC0D8FDB57E3F" box="[527,594,752,778]" name="Japan" pageId="283" pageNumber="284">Japan</collectingCountry>
. Ten years after the original description, he (1887) reported an additional specimen from the south of the 
<collectingCountry id="47AF43B9FECD8709DB0AC11BFD7A7E78" box="[543,669,819,845]" name="Philippines" pageId="283" pageNumber="284">Philippines</collectingCountry>
(BMNH 1887.12.7.106, 45 mm HL, 246+ mm TL). 
<bibRefCitation id="5B297ED8FECD8709DB9DC17DFEBA7EA4" author="Sazonov, Y. I. &amp; Iwamoto, T." pageId="283" pageNumber="284" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto(1992:78)</bibRefCitation>
erroneously considered the 
<collectingCountry id="47AF43B9FECD8709DB99C15FFCED7EA4" box="[652,778,887,913]" name="Philippines" pageId="283" pageNumber="284">Philippines</collectingCountry>
specimen as a 
<typeStatus id="E003BD8BFECD8709D855C1B2FE707E81" box="[320,407,922,948]" pageId="283" pageNumber="284" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="F8B878AAFECD8709D8DEC1B2FDA37E86" authorityName="Gunther" authorityYear="1877" box="[459,580,921,947]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709D8DEC1B2FDA37E86" box="[459,580,921,947]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
, designating the Japanese specimen as the 
<typeStatus id="E003BD8BFECD8709D8B7C193FDEE7EE0" box="[418,521,955,981]" pageId="283" pageNumber="284" type="lectotype">lectotype</typeStatus>
of this species.This act was probably due to their recognition that the 
<collectingCountry id="47AF43B9FECD8709DB99C1F5FCED7EC2" box="[652,778,989,1015]" name="Philippines" pageId="283" pageNumber="284">Philippines</collectingCountry>
specimen “appears to be a different species with a longer upper jaw, longer barbel, longer first gill slit, wider 1D.– 2D. interspace, and larger scales”. According to Iwamoto (1997:945), this specimen is unable to be allocated to any known species, partly owing to its poor condition. The above 
<collectingCountry id="47AF43B9FECD8709D9C3C681FEB379F6" box="[214,340,1193,1219]" name="Philippines" pageId="283" pageNumber="284">Philippines</collectingCountry>
specimen is not included in the type series of 
<taxonomicName id="F8B878AAFECD8709D9E4C6E4FE8279D0" authorityName="Gunther" authorityYear="1877" box="[241,357,1227,1253]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709D9E4C6E4FE8279D0" box="[241,357,1227,1253]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
, being designated as a 
<typeStatus id="E003BD8BFECD8709DB66C6E3FCED79D0" box="[627,778,1227,1253]" pageId="283" pageNumber="284" type="paralectotype">paralectotype</typeStatus>
of 
<taxonomicName id="F8B878AAFECD8709D9B3C6C5FE467832" authorityName=": Okamura" authorityYear="1970" box="[166,417,1261,1287]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="asper">
<emphasis id="0DCCDF3BFECD8709D9B3C6C5FE467832" box="[166,417,1261,1287]" italics="true" pageId="283" pageNumber="284">Coryphaenoides asper</emphasis>
</taxonomicName>
in this study (see the Comments on type specimens of 
<taxonomicName id="F8B878AAFECD8709D869C738FE3C781C" authorityName=": Okamura" authorityYear="1970" box="[380,475,1296,1322]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="asper">
<emphasis id="0DCCDF3BFECD8709D869C738FE3C781C" box="[380,475,1296,1322]" italics="true" pageId="283" pageNumber="284">C. asper</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3F070329FECD8709D9A8C719FE637B0C" blockId="283.[136,778,173,2070]" pageId="283" pageNumber="284">
<emphasis id="0DCCDF3BFECD8709D9A8C719FED6787E" bold="true" box="[189,305,1329,1355]" pageId="283" pageNumber="284">Remarks.</emphasis>
<bibRefCitation id="5B297ED8FECD8709D829C719FD65787E" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[316,642,1329,1356]" pageId="283" pageNumber="284" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto (1992)</bibRefCitation>
reported 
<taxonomicName id="F8B878AAFECD8709DBE0C71AFF397858" authorityName="Gunther" authorityYear="1877" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DBE0C71AFF397858" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
from the Sala y Gomez Ridge in the southeastern Pacific, but noted that their specimens differ from the 
<typeStatus id="E003BD8BFECD8709D99DC7BFFF0D7884" box="[136,234,1431,1457]" pageId="283" pageNumber="284" type="holotype">holotype</typeStatus>
from 
<collectingCountry id="47AF43B9FECD8709D825C7B0FE977887" box="[304,368,1432,1458]" name="Japan" pageId="283" pageNumber="284">Japan</collectingCountry>
in many aspects, including the orbit diameter, suborbital width, orbit–preopercle distance, upper-jaw length, and the number of pelvic-fin rays. A further study is required to confirm the conspecificity of these two populations.
</paragraph>
<paragraph id="3F070329FECD8709D9A8C469FC237E3C" blockId="283.[136,778,173,2070]" lastBlockId="283.[810,1452,173,777]" pageId="283" pageNumber="284">
<emphasis id="0DCCDF3BFECD8709D9A8C469FE847B6E" bold="true" box="[189,355,1601,1627]" pageId="283" pageNumber="284">Comparisons.</emphasis>
The species of 
<taxonomicName id="F8B878AAFECD8709DB02C469FD417B6E" authorityName="Gunther" authorityYear="1887" box="[535,678,1601,1627]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFECD8709DB02C469FD417B6E" box="[535,678,1601,1627]" italics="true" pageId="283" pageNumber="284">Trachonurus</emphasis>
</taxonomicName>
are very similar to each other in general appearance, with most species having few diagnostic features that allow ready identification (Iwamoto 1997:942). Among the other five species of the genus (see the Remarks of the genus), 
<taxonomicName id="F8B878AAFECD8709DBE0C4E2FDF87A33" authority="Iwamoto &amp; McMillan, 1997" authorityName="Iwamoto &amp; McMillan" authorityYear="1997" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="gagates">
<emphasis id="0DCCDF3BFECD8709DBE0C4E2FF387A30" italics="true" pageId="283" pageNumber="284">T. gagates</emphasis>
<bibRefCitation id="5B297ED8FECD8709D9F0C4C4FDF87A33" author="Iwamoto, T. &amp; McMillan, P." box="[229,543,1771,1798]" pageId="283" pageNumber="284" pagination="255 - 259" refId="ref288623" refString="Iwamoto, T. &amp; McMillan, P. (1997) A new grenadier, genus Trachonurus, from New Zealand and Australia (Pisces: Gadiformes: Macrouridae). Memoirs of the Museum of Victoria, 56, 255 - 259." type="journal article" year="1997">Iwamoto &amp; McMillan, 1997</bibRefCitation>
</taxonomicName>
may not be confused with any other congeners in the absence of a grooved lateral line on the body (vs. present).However, the remaining four species are quite similar to 
<taxonomicName id="F8B878AAFECD8709D8AAC57AFDD57A5E" authorityName="Gunther" authorityYear="1877" box="[447,562,1873,1899]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709D8AAC57AFDD57A5E" box="[447,562,1873,1899]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
, and they are most likely to be misidentified without a detailed examination of morphology. Based on published information, 
<taxonomicName id="F8B878AAFECD8709DB82C5BEFCED7A85" authorityName="Gunther" authorityYear="1877" box="[663,778,1942,1968]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DB82C5BEFCED7A85" box="[663,778,1942,1968]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="F8B878AAFECD8709D802C590FD5D7AE7" authority="Gilbert &amp; Cramer, 1897" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[279,698,1976,2002]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="sentipellis">
<emphasis id="0DCCDF3BFECD8709D802C590FE427AE7" box="[279,421,1976,2002]" italics="true" pageId="283" pageNumber="284">T. sentipellis</emphasis>
<bibRefCitation id="5B297ED8FECD8709D8B9C590FD5D7AE7" author="Gilbert, C. H. &amp; Cramer, F." box="[428,698,1976,2002]" pageId="283" pageNumber="284" pagination="403 - 435" refId="ref285862" refString="Gilbert, C. H. &amp; Cramer, F. (1897) Report on the fishes dredged in deep water near the Hawaiian Islands, with descriptions and figures of twenty-three new species. Proceedings of the United States National Museum 19, 403 - 435, pls. XXXVI-XLVIII. http: // dx. doi. org / 10.5479 / si. 00963801.19 - 1114.403" type="journal article" year="1897">
Gilbert &amp; Cramer, 
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</bibRefCitation>
</taxonomicName>
in that the outer premaxillary teeth are distinctly enlarged (vs. scarcely or not at all enlarged, fide 
<bibRefCitation id="5B297ED8FECD8709DB0EC5D4FC707DF2" author="Iwamoto, T. &amp; Williams, A." box="[539,919,173,2070]" pageId="283" pageNumber="284" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams 1999:213</bibRefCitation>
), and from 
<taxonomicName id="F8B878AAFECD8709DD0EC286FADE7DF2" authority="Iwamoto, 1997" authorityName="Iwamoto" authorityYear="1997" box="[1051,1337,173,199]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="robinsi">
<emphasis id="0DCCDF3BFECD8709DD0EC286FB617DF2" box="[1051,1158,173,199]" italics="true" pageId="283" pageNumber="284">T. robinsi</emphasis>
Iwamoto, 
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</taxonomicName>
in having fewer spinules on the body scales (see Iwamoto 1997: fig. 2). According to Iwamoto (1997:948), 
<taxonomicName id="F8B878AAFECD8709DC19C2DAFA997C3E" authorityName="Gunther" authorityYear="1877" box="[1292,1406,241,267]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DC19C2DAFA997C3E" box="[1292,1406,241,267]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="F8B878AAFECD8709DD3DC33CFA4C7C1B" authority="(Goode &amp; Bean, 1885)" baseAuthorityName="Goode &amp; Bean" baseAuthorityYear="1885" box="[1064,1451,275,302]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="sulcatus">
<emphasis id="0DCCDF3BFECD8709DD3DC33CFB477C18" box="[1064,1184,275,301]" italics="true" pageId="283" pageNumber="284">T. sulcatus</emphasis>
(
<bibRefCitation id="5B297ED8FECD8709DDBAC33BFA447C1B" author="Goode, G. B. &amp; Bean, T. H." box="[1199,1443,275,302]" pageId="283" pageNumber="284" pagination="589 - 605" refId="ref286338" refString="Goode, G. B. &amp; Bean, T. H. (1885) Descriptions of new fishes obtainedbytheUnitedStatesFishCommissionmainlyfrom deep water off the Atlantic and Gulf coasts. Proceedings of the United States National Museum, 8, 589 - 605. http: // dx. doi. org / 10.5479 / si. 00963801.8 - 543.589" type="journal article" year="1885">Goode &amp; Bean, 1885</bibRefCitation>
)
</taxonomicName>
in its slightly lower dimensions of the upper-jaw length, suborbital width, and snout length. However, these differences become negligible when considering variation of the 
<taxonomicName id="F8B878AAFECD8709DA60C3B4FC0E7C80" authorityName="Gunther" authorityYear="1877" box="[885,1001,411,437]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DA60C3B4FC0E7C80" box="[885,1001,411,437]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
specimens examined in this study. The differences between 
<taxonomicName id="F8B878AAFECD8709DD1AC396FB987CE2" authorityName="Gunther" authorityYear="1877" box="[1039,1151,445,471]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="villosus">
<emphasis id="0DCCDF3BFECD8709DD1AC396FB987CE2" box="[1039,1151,445,471]" italics="true" pageId="283" pageNumber="284">T. villosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8B878AAFECD8709DDA1C396FBE77CCC" authority="Iwamoto &amp; Williams, 1999" authorityName="Iwamoto &amp; Williams" authorityYear="1999" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="species" species="yiwardaus">
<emphasis id="0DCCDF3BFECD8709DDA1C396FAA57CE2" box="[1204,1346,445,471]" italics="true" pageId="283" pageNumber="284">T. yiwardaus</emphasis>
<bibRefCitation id="5B297ED8FECD8709DC5DC395FBE77CCC" author="Iwamoto, T. &amp; Williams, A." pageId="283" pageNumber="284" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams, 1999</bibRefCitation>
</taxonomicName>
were not discussed in the original description of the latter (
<bibRefCitation id="5B297ED8FECD8709DD2BC029FA887F2E" author="Iwamoto, T. &amp; Williams, A." box="[1086,1391,513,539]" pageId="283" pageNumber="284" pagination="105 - 243" refId="ref289322" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of western and northwestern Australia. Proceedings of the California Academy of Sciences, 51, 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams 1999</bibRefCitation>
), and should be confirmed by a direct comparison of the two species in the future. As noted by Iwamoto (1997:948), there remains much work to be done on 
<taxonomicName id="F8B878AAFECD8709DDF2C04FFA917FB4" authorityName="Gunther" authorityYear="1887" box="[1255,1398,615,641]" class="Actinopterygii" family="Macrouridae" genus="Trachonurus" kingdom="Animalia" order="Gadiformes" pageId="283" pageNumber="284" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFECD8709DDF2C04FFA917FB4" box="[1255,1398,615,641]" italics="true" pageId="283" pageNumber="284">Trachonurus</emphasis>
</taxonomicName>
, and additional characters are obviously necessary to further characterize each species. A molecular approach may be helpful to clarify the problems within this difficult group of grenadiers.
</paragraph>
</subSubSection>
</treatment>
</document>