treatments-xml/data/3C/70/D6/3C70D6BD67545BB8BCCF987233CF5995.xml
2024-06-21 12:34:03 +02:00

280 lines
21 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.81.67785" ID-GBIF-Dataset="c2ed1993-fbdc-4b34-974c-fcc1bcdc8c80" ID-PMC="PMC8213683" ID-Pensoft-Pub="1314-4049-81-1" ID-Pensoft-UUID="D672632A0F1A525A81CADAD8623D67FF" ID-PubMed="34163305" ModsDocID="1314-4049-81-1" checkinTime="1623443205026" checkinUser="pensoft" docAuthor="Reblova, Martina, Nekvindova, Jana &amp; Miller, Andrew N." docDate="2021" docId="3C70D6BD67545BB8BCCF987233CF5995" docLanguage="en" docName="MycoKeys 81: 1-44" docOrigin="MycoKeys 81" docPubDate="2021-06-11" docSource="http://dx.doi.org/10.3897/mycokeys.81.67785" docTitle="Catenularia angulospora (Linder) E. W. Mason, Mycol. Pap. 5: 121. 1941." docType="treatment" docVersion="4" id="D672632A0F1A525A81CADAD8623D67FF" lastPageNumber="1" masterDocId="D672632A0F1A525A81CADAD8623D67FF" masterDocTitle="Phylogeny and taxonomy of Catenularia and similar fungi with catenate conidia" masterLastPageNumber="44" masterPageNumber="1" pageNumber="1" updateTime="1668137208642" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogeny and taxonomy of Catenularia and similar fungi with catenate conidia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reblova, Martina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5229-1709</mods:nameIdentifier>
<mods:affiliation>Czech Academy of Sciences, Institute of Botany, Pruhonice 252 43, Czech Republic</mods:affiliation>
<mods:nameIdentifier type="email">martina.reblova@ibot.cas.cz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nekvindova, Jana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2861-5483</mods:nameIdentifier>
<mods:affiliation>Department of Clinical Biochemistry and Diagnostics, University Hospital Hradec Kralove, Hradec Kralove 500 05, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Andrew N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7300-0069</mods:nameIdentifier>
<mods:affiliation>Illinois Natural History Survey, University of Illinois Urbana-Champaign, Champaign, Illinois 61820, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>81</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.81.67785</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.81.67785</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-81-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">D672632A0F1A525A81CADAD8623D67FF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182654388" LSID="urn:lsid:plazi:treatment:3C70D6BD67545BB8BCCF987233CF5995" httpUri="http://treatment.plazi.org/id/3C70D6BD67545BB8BCCF987233CF5995" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="3C70D6BD-6754-5BB8-BCCF-987233CF5995" authority="(Linder) E. W. Mason, Mycol. Pap. 5: 121. 1941." authorityName="(Linder) E. W. Mason, Mycol. Pap. 5: 121. 1941." class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia angulospora" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="angulospora">Catenularia angulospora (Linder) E.W. Mason, Mycol. Pap. 5: 121. 1941.</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Catenularia angulospora (holotype FH 00965375) A vertical section of the wood with colony B-D conidiophores with conidia. Scale bars: 500 μm (A); 50 μm (B-D)." figureDoi="10.3897/mycokeys.81.67785.figure3" httpUri="https://binary.pensoft.net/fig/554153" pageId="0" pageNumber="1">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Linder" authorityYear="1933" class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Haplochalara" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Haplochalara angulospora" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="angulospora">Haplochalara angulospora</taxonomicName>
Basionym.
<taxonomicName authorityName="Linder" authorityYear="1933" class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Haplochalara" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Haplochalara angulospora" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="angulospora">Haplochalara angulospora</taxonomicName>
Linder, Mycologia 25: 347. 1933.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="N.D.Sharma" authorityYear="1980" class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia hughesii" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hughesii">Catenularia hughesii</taxonomicName>
Synonym.?
<taxonomicName authorityName="N.D.Sharma" authorityYear="1980" class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia hughesii" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hughesii">Catenularia hughesii</taxonomicName>
N.D. Sharma, J. Indian bot. Soc. 59: 73. 1980.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Colonies on natural substrate effuse, hairy to velutinous, dark brown to almost black.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Anamorph</emphasis>
.
</emphasis>
Conidiophores 77-220
<normalizedToken originalValue="×">x</normalizedToken>
4.5-6(-7)
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, 7-8
<normalizedToken originalValue="μm">μm</normalizedToken>
above the base, macronematous, solitary or arise in tufts, erect, straight or slightly flexuous, unbranched, dark brown, paler towards the apex, septate. Capitate hyphae absent. Conidiogenous cells 18-25
<normalizedToken originalValue="×">x</normalizedToken>
3.5-4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
tapering to ca. 2.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, integrated, terminal, monophialidic, extending percurrently, obclavate to subcylindrical or slightly lageniform, pale brown, paler towards the apex; collarettes 3-4
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, 1.5(-2)
<normalizedToken originalValue="μm">μm</normalizedToken>
deep, funnel-shaped, subhyaline, smooth, margin entire. Conidia 6-8(-9)
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 4.5-6(-7)
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at the apical end, ca. 2
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at the basal hilum (mean
<normalizedToken originalValue="±">+/-</normalizedToken>
SD = 7.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
<normalizedToken originalValue="×">x</normalizedToken>
6.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
2.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.0
<normalizedToken originalValue="μm">μm</normalizedToken>
), rounded-obconic in side view, with three blunt corners when viewed from above, broadly rounded to flattened at the apex, truncate at the basal scar, pale brown to pale fuscous, thick-walled, smooth; formed singly, adhered in basipetal chains.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Teleomorph</emphasis>
.
</emphasis>
Unknown.
</paragraph>
<caption doi="10.3897/mycokeys.81.67785.figure3" httpUri="https://binary.pensoft.net/fig/554153" pageId="0" pageNumber="1" start="Figure 3" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia angulospora" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">Catenularia angulospora</emphasis>
</taxonomicName>
(holotype FH 00965375)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
vertical section of the wood with colony
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
conidiophores with conidia. Scale bars: 500
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
); 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="specimen examined">
<paragraph pageId="0" pageNumber="1">Specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
USA - Kentucky • near Louisville; on decaying beech log; 23 Mar. 1928; D.H. Linder (
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">holotype</emphasis>
of
<taxonomicName lsidName="C. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">C. angulospora</emphasis>
</taxonomicName>
FH herbarium 00965375, as microscopic slides).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat and geographical distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Saprobe on dead culms of
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Bambusa" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Bambusa" order="Graminales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bambusa</emphasis>
</taxonomicName>
sp., decaying wood of
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Fagus</emphasis>
</taxonomicName>
sp. and other unknown hosts in freshwater and terrestrial habitats. It is known in China, India and the USA (
<bibRefCitation DOI="https://doi.org/10.1080/00275514.1933.12020675" author="Linder, DH" journalOrPublisher="Mycologia" pageId="0" pageNumber="1" pagination="342 - 349" refId="B46" refString="Linder, DH, 1933. North American Hyphomycetes. I. Two new Helicosporeae and the new genera Haplochalara and Paspalomyces. Mycologia 25: 342 - 349, DOI: https://doi.org/10.1080/00275514.1933.12020675" title="North American Hyphomycetes. I. Two new Helicosporeae and the new genera Haplochalara and Paspalomyces." url="https://doi.org/10.1080/00275514.1933.12020675" volume="25" year="1933">Linder 1933</bibRefCitation>
;
<bibRefCitation author="Sharma, ND" journalOrPublisher="Journal of the Indian Botanical Society" pageId="0" pageNumber="1" pagination="72 - 77" refId="B78" refString="Sharma, ND, 1980. Some additions to fungi of India - VIII. Journal of the Indian Botanical Society 59: 72 - 77" title="Some additions to fungi of India - VIII." volume="59" year="1980">Sharma 1980</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/s13225-019-00438-1" author="Luo, ZL" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="1" pagination="451 - 660" refId="B50" refString="Luo, ZL, Hyde, KD, Liu, J-K, Maharachchikumbura, SSN, Rajesh, J, Bao, D-F, Bhat, DJ, Lin, C-G, Li, W-L, Yang, J, Liu, N-G, Lu, Y-Z, Jayawardena, RS, Li, J-F, Su, H-Y, 2019. Freshwater Sordariomycetes. Fungal Diversity 99: 451 - 660, DOI: https://doi.org/10.1007/s13225-019-00438-1" title="Freshwater Sordariomycetes." url="https://doi.org/10.1007/s13225-019-00438-1" volume="99" year="2019">Luo et al. 2019</bibRefCitation>
as
<taxonomicName lsidName="C. cubensis" pageId="0" pageNumber="1" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. cubensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
For additional description and illustration, see
<bibRefCitation DOI="https://doi.org/10.1007/s13225-019-00438-1" author="Luo, ZL" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="1" pagination="451 - 660" refId="B50" refString="Luo, ZL, Hyde, KD, Liu, J-K, Maharachchikumbura, SSN, Rajesh, J, Bao, D-F, Bhat, DJ, Lin, C-G, Li, W-L, Yang, J, Liu, N-G, Lu, Y-Z, Jayawardena, RS, Li, J-F, Su, H-Y, 2019. Freshwater Sordariomycetes. Fungal Diversity 99: 451 - 660, DOI: https://doi.org/10.1007/s13225-019-00438-1" title="Freshwater Sordariomycetes." url="https://doi.org/10.1007/s13225-019-00438-1" volume="99" year="2019">Luo et al. (2019</bibRefCitation>
, as
<taxonomicName lsidName="C. cubensis" pageId="0" pageNumber="1" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. cubensis</emphasis>
</taxonomicName>
).
<bibRefCitation DOI="https://doi.org/10.1080/0028825X.1965.10876990" author="Hughes, SJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="1" pagination="136 - 150" refId="B37" refString="Hughes, SJ, 1965. New Zealand fungi 3. Catenularia Grove. New Zealand Journal of Botany 3: 136 - 150, DOI: https://doi.org/10.1080/0028825X.1965.10876990" title="New Zealand fungi 3. Catenularia Grove." url="https://doi.org/10.1080/0028825X.1965.10876990" volume="3" year="1965">Hughes (1965)</bibRefCitation>
revised the type material of
<taxonomicName lsidName="H. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">H. angulospora</emphasis>
</taxonomicName>
, and despite the striking similarities to other
<taxonomicName class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Catenularia</emphasis>
</taxonomicName>
, he kept the species in
<taxonomicName authorityName="Linder" authorityYear="1933" class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Haplochalara" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Haplochalara" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haplochalara</emphasis>
</taxonomicName>
due to the absence of capitate hyphae.
<bibRefCitation author="Sharma, ND" journalOrPublisher="Journal of the Indian Botanical Society" pageId="0" pageNumber="1" pagination="72 - 77" refId="B78" refString="Sharma, ND, 1980. Some additions to fungi of India - VIII. Journal of the Indian Botanical Society 59: 72 - 77" title="Some additions to fungi of India - VIII." volume="59" year="1980">Sharma (1980)</bibRefCitation>
described
<taxonomicName lsidName="C. hughesii" pageId="0" pageNumber="1" rank="species" species="hughesii">
<emphasis italics="true" pageId="0" pageNumber="1">C. hughesii</emphasis>
</taxonomicName>
on dead bamboo culms in India with pale brown to brown conidia 6-8
<normalizedToken originalValue="×">x</normalizedToken>
4.5-5.8
<normalizedToken originalValue="μm">μm</normalizedToken>
and conidiophores up to 270
<normalizedToken originalValue="×">x</normalizedToken>
5-7
<normalizedToken originalValue="μm">μm</normalizedToken>
. Although the holotype of this species was not available for study, a detailed morphological comparison of its original description and illustration with
<taxonomicName lsidName="C. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">C. angulospora</emphasis>
</taxonomicName>
suggests that they are conspecific.
<bibRefCitation DOI="https://doi.org/10.1007/s13225-019-00438-1" author="Luo, ZL" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="1" pagination="451 - 660" refId="B50" refString="Luo, ZL, Hyde, KD, Liu, J-K, Maharachchikumbura, SSN, Rajesh, J, Bao, D-F, Bhat, DJ, Lin, C-G, Li, W-L, Yang, J, Liu, N-G, Lu, Y-Z, Jayawardena, RS, Li, J-F, Su, H-Y, 2019. Freshwater Sordariomycetes. Fungal Diversity 99: 451 - 660, DOI: https://doi.org/10.1007/s13225-019-00438-1" title="Freshwater Sordariomycetes." url="https://doi.org/10.1007/s13225-019-00438-1" volume="99" year="2019">Luo et al. (2019)</bibRefCitation>
reported this species as
<taxonomicName lsidName="C. cubensis" pageId="0" pageNumber="1" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. cubensis</emphasis>
</taxonomicName>
(strain MFLUCC 18-1331) from China, characterised by the absence of capitate hyphae and cuneiform, greyish-brown to brown conidia 6-8
<normalizedToken originalValue="×">x</normalizedToken>
4-6
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
In the phylogenetic analysis, the strain of
<taxonomicName lsidName="C. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">C. angulospora</emphasis>
</taxonomicName>
MFLUCC 18-1331 clustered as a sister to
<taxonomicName lsidName="C. cubensis" pageId="0" pageNumber="1" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. cubensis</emphasis>
</taxonomicName>
S.M.H. 3258, but their relationship is not statistically supported. Both species are, however, very similar.
<taxonomicName class="Sordariomycetes" family="Chaetosphaeriaceae" genus="Catenularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Catenularia cubensis" order="Chaetosphaeriales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">Catenularia cubensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Holubova-Jechova, V" journalOrPublisher="Mycotaxon" pageId="0" pageNumber="1" pagination="277 - 292" refId="B31" refString="Holubova-Jechova, V, 1982. New or interesting phialidic hyphomycetes from Cuba. Mycotaxon 15: 277 - 292" title="New or interesting phialidic hyphomycetes from Cuba." volume="15" year="1982">
<normalizedToken originalValue="Holubová-Jechová">Holubova-Jechova</normalizedToken>
1982
</bibRefCitation>
) differs from
<taxonomicName lsidName="C. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">C. angulospora</emphasis>
</taxonomicName>
in brown to dark brown conidia, slightly narrower at the apical end (5.5-8.5
<normalizedToken originalValue="×">x</normalizedToken>
3.5-5.5
<normalizedToken originalValue="μm">μm</normalizedToken>
), and presence of capitate hyphae scattered among the conidiophores. The ITS sequence identity between
<taxonomicName lsidName="C. cubensis" pageId="0" pageNumber="1" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. cubensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. angulospora" pageId="0" pageNumber="1" rank="species" species="angulospora">
<emphasis italics="true" pageId="0" pageNumber="1">C. angulospora</emphasis>
</taxonomicName>
is 96.5% and supports our conclusion to treat them as separate species.
</paragraph>
</subSubSection>
</treatment>
</document>