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<mods:title id="060BCF5B03E4BB6CDB16BAF526A59C37">Review of the genus Paradonus Stibick, 1971 (Elateridae: Negastriinae), with three new North American species</mods:title>
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5. 
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<emphasis id="D887EACD4B798928E3F243B32DBF92BC" bold="true" box="[181,472,153,180]" italics="true" pageId="10" pageNumber="534">Paradonus obliquatulus</emphasis>
(
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.
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<paragraph id="EA4C36DF4B798928E3D043972D7392DF" blockId="10.[151,724,153,215]" box="[151,276,189,215]" pageId="10" pageNumber="534">Figs. 4, 14.</paragraph>
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<emphasis id="D887EACD4B798928E3D042282DCC9312" box="[151,427,258,282]" italics="true" pageId="10" pageNumber="534">Cryptohypnus obliquatulus</emphasis>
<bibRefCitation id="8E624B2E4B798928E2F6422B2EF99312" author="Melsheimer, F. E." box="[433,670,257,282]" pageId="10" pageNumber="534" pagination="213 - 223" refId="ref15708" refString="Melsheimer, F. E. (1845) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2 (1844 - 1845), 213 - 223." type="journal article" year="1845">Melsheimer, 1845: 214</bibRefCitation>
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; 
<treatmentCitation id="6B5210CE4B798928E1EE42282F089312" author="LeConte, J. L." box="[681,879,257,282]" page="488" pageId="10" pageNumber="534" year="1853">
<bibRefCitation id="8E624B2E4B798928E1EE42282F089312" author="LeConte, J. L." box="[681,879,257,282]" pageId="10" pageNumber="534" pagination="405 - 508" refId="ref15451" refString="LeConte, J. L. (1853) Revision of the Elateridae of the United States. Transactions of the American Philosophical Society, 10 (2), 405 - 508. https: // doi. org / 10.2307 / 1005288" type="journal article" year="1853">LeConte 1853: 488</bibRefCitation>
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; Candèze 1860: 90; 
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<bibRefCitation id="8E624B2E4B798928E779422828B39312" author="Horn, G. H." box="[1086,1236,258,282]" pageId="10" pageNumber="534" pagination="1 - 29" refId="ref15233" refString="Horn, G. H. (1891) A monograph of the species of Cryptohypnus of Boreal America. Transactions of the American Entomological Society, 18, 1 - 29." type="journal article" year="1891">Horn 1891: 27</bibRefCitation>
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<taxonomicName id="2DF34D5C4B798928E3D0420A2E3B9331" authority="(Melsheimer, 1845)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[151,604,288,313]" class="Insecta" family="Elateridae" genus="Hypnoidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="obliquatulus">
<emphasis id="D887EACD4B798928E3D0420A2DED9330" box="[151,394,288,312]" italics="true" pageId="10" pageNumber="534">Hypnoidus obliquatulus</emphasis>
(
<bibRefCitation id="8E624B2E4B798928E2D0420A2E349331" author="Melsheimer, F. E." box="[407,595,288,313]" pageId="10" pageNumber="534" pagination="213 - 223" refId="ref15708" refString="Melsheimer, F. E. (1845) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2 (1844 - 1845), 213 - 223." type="journal article" year="1845">Melsheimer, 1845</bibRefCitation>
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; various authors.
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<emphasis id="D887EACD4B798928E3D0426A2DED935F" box="[151,394,319,343]" italics="true" pageId="10" pageNumber="534">Negastrius obliquatulus</emphasis>
(
<bibRefCitation id="8E624B2E4B798928E2D042142E34935F" author="Melsheimer, F. E." box="[407,595,318,343]" pageId="10" pageNumber="534" pagination="213 - 223" refId="ref15708" refString="Melsheimer, F. E. (1845) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2 (1844 - 1845), 213 - 223." type="journal article" year="1845">Melsheimer, 1845</bibRefCitation>
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; various authors, noted in 
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<bibRefCitation id="8E624B2E4B798928E020421528C6935F" author="van Herk, W. G. &amp; Vernon, R. S." box="[871,1185,318,343]" pageId="10" pageNumber="534" pagination="87 - 117" refId="ref16106" refString="van Herk, W. G. &amp; Vernon, R. S. (2014) Click Beetles and Wireworms (Coleoptera: Elateridae) of Alberta, Saskatchewan, and Manitoba. In: Giberson, D. J. &amp; Carcamo, H. A. (Eds.), Arthropods of Canadian Grasslands. Vol. 4. Biodiversity and Systematics Part 2. Biological Survey of Canada, Ottawa, Ontario, pp. 87 - 117." type="book chapter" year="2014">van Herk and Vernon 2014: 93</bibRefCitation>
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<emphasis id="D887EACD4B798928E3D042742DEA937E" box="[151,397,350,374]" italics="true" pageId="10" pageNumber="534">Paradonus obliquatulus</emphasis>
(
<bibRefCitation id="8E624B2E4B798928E2DC42772E3E937E" author="Melsheimer, F. E." box="[411,601,349,374]" pageId="10" pageNumber="534" pagination="213 - 223" refId="ref15708" refString="Melsheimer, F. E. (1845) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2 (1844 - 1845), 213 - 223." type="journal article" year="1845">Melsheimer, 1845</bibRefCitation>
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; 
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<bibRefCitation id="8E624B2E4B798928E12A42772F40937E" author="Stibick, J. N. L." box="[621,807,349,374]" pageId="10" pageNumber="534" pagination="99 - 131" refId="ref16033" refString="Stibick, J. N. L. (1991) North American Negastriinae (Coleoptera, Elateridae): The Negastriinae of the Eastern United States and Adjacent Canada. Insecta Mundi, 4 (1990), 99 - 131." type="journal article" year="1991">Stibick 1991: 122</bibRefCitation>
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; 
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Majka 
<emphasis id="D887EACD4B798928E03C42742FCE937E" box="[891,937,350,374]" italics="true" pageId="10" pageNumber="534">et al</emphasis>
. 2011:142
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; 
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Bousquet 
<emphasis id="D887EACD4B798928E7CC427428DE937E" box="[1163,1209,350,374]" italics="true" pageId="10" pageNumber="534">et al</emphasis>
. 2013: 189
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; 
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<bibRefCitation id="8E624B2E4B798928E67A42742DFA939C" author="van Herk, W. G. &amp; Vernon, R. S." pageId="10" pageNumber="534" pagination="87 - 117" refId="ref16106" refString="van Herk, W. G. &amp; Vernon, R. S. (2014) Click Beetles and Wireworms (Coleoptera: Elateridae) of Alberta, Saskatchewan, and Manitoba. In: Giberson, D. J. &amp; Carcamo, H. A. (Eds.), Arthropods of Canadian Grasslands. Vol. 4. Biodiversity and Systematics Part 2. Biological Survey of Canada, Ottawa, Ontario, pp. 87 - 117." type="book chapter" year="2014">van Herk and Vernon 2014: 93</bibRefCitation>
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<treatmentCitation id="6B5210CE4B798928E3D042B12E6793BB" author="Notman, H." box="[151,512,410,435]" page="159" pageId="10" pageNumber="534" year="1921">
<taxonomicName id="2DF34D5C4B798928E3D042B12E6793BB" authority="Notman, 1921: 159" authorityName="Notman" authorityPageNumber="159" authorityYear="1921" box="[151,512,410,435]" class="Insecta" family="Elateridae" genus="Hypnoidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="felti">
<emphasis id="D887EACD4B798928E3D042B12D5593BB" box="[151,306,411,435]" italics="true" pageId="10" pageNumber="534">Hypnoidus felti</emphasis>
<bibRefCitation id="8E624B2E4B798928E27E42B12E6793BB" author="Notman, H." box="[313,512,410,435]" pageId="10" pageNumber="534" pagination="145 - 160" refId="ref15749" refString="Notman, H. (1921) Some new genera and species of Coleoptera collected at Westfield, Chautauqua Co, N. Y. Journal of the New York Entomological Society, 29, 145 - 160." type="journal article" year="1921">Notman, 1921: 159</bibRefCitation>
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<paragraph id="EA4C36DF4B798928E3D042CB2EBD93F3" blockId="10.[151,1436,481,940]" box="[151,730,481,507]" pageId="10" pageNumber="534">Diagnosis: See key above for diagnostic characters.</paragraph>
<paragraph id="EA4C36DF4B798928E380412F2FEF9017" blockId="10.[151,1436,481,940]" box="[199,904,517,543]" pageId="10" pageNumber="534">
Over 
<specimenCount id="FCF5FD564B798928E241412F2DF99017" box="[262,414,517,543]" pageId="10" pageNumber="534" type="generic">20 specimens</specimenCount>
examined from INHS, MCZC, and LSAM.
</paragraph>
<paragraph id="EA4C36DF4B798928E38041032E5C9148" blockId="10.[151,1436,481,940]" pageId="10" pageNumber="534">
This species is the only one that historically has consistently been considered distinct from 
<taxonomicName id="2DF34D5C4B798928E7F341002CBB906F" authority="(Stibick 1991)" baseAuthorityName="Stibick" baseAuthorityYear="1991" class="Insecta" family="Elateridae" genus="Paradonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="D887EACD4B798928E7F341002959904B" box="[1204,1342,553,579]" italics="true" pageId="10" pageNumber="534">P. pectoralis</emphasis>
(
<bibRefCitation id="8E624B2E4B798928E60C41032CB4906F" author="Stibick, J. N. L." pageId="10" pageNumber="534" pagination="99 - 131" refId="ref16033" refString="Stibick, J. N. L. (1991) North American Negastriinae (Coleoptera, Elateridae): The Negastriinae of the Eastern United States and Adjacent Canada. Insecta Mundi, 4 (1990), 99 - 131." type="journal article" year="1991">Stibick 1991</bibRefCitation>
)
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. Stibick stresses a piceous pronotum in his key to species, but the pronotum is often lighter in specimens I have seen. I have found many specimens of 
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<emphasis id="D887EACD4B798928E13941582EE89083" box="[638,655,626,651]" italics="true" pageId="10" pageNumber="534">P</emphasis>
. 
<emphasis id="D887EACD4B798928E1DB415B2E899083" box="[668,750,625,651]" italics="true" pageId="10" pageNumber="534">beckeri</emphasis>
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, 
<taxonomicName id="2DF34D5C4B798928E1BC41582FE89083" authorityName="Stibick" authorityYear="1991" box="[763,911,626,652]" class="Insecta" family="Elateridae" genus="Paradonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="jerseiensis">
<emphasis id="D887EACD4B798928E1BC41582F6B9083" box="[763,780,626,651]" italics="true" pageId="10" pageNumber="534">P</emphasis>
. 
<emphasis id="D887EACD4B798928E05E41582FE89083" box="[793,911,626,651]" italics="true" pageId="10" pageNumber="534">jerseiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="2DF34D5C4B798928E08C415828329083" baseAuthorityName="Stibick" baseAuthorityYear="1991" box="[971,1109,625,651]" class="Insecta" family="Elateridae" genus="Paradonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="D887EACD4B798928E08C415828329083" box="[971,1109,625,651]" italics="true" pageId="10" pageNumber="534">P. pectoralis</emphasis>
</taxonomicName>
(see below) mixed under this name. 
<bibRefCitation id="8E624B2E4B798928E3A541BC2E4190A7" author="van Herk, W. G. &amp; Vernon, R. S." box="[226,550,661,688]" pageId="10" pageNumber="534" pagination="87 - 117" refId="ref16106" refString="van Herk, W. G. &amp; Vernon, R. S. (2014) Click Beetles and Wireworms (Coleoptera: Elateridae) of Alberta, Saskatchewan, and Manitoba. In: Giberson, D. J. &amp; Carcamo, H. A. (Eds.), Arthropods of Canadian Grasslands. Vol. 4. Biodiversity and Systematics Part 2. Biological Survey of Canada, Ottawa, Ontario, pp. 87 - 117." type="book chapter" year="2014">Van Herk and Vernon (2014)</bibRefCitation>
report this species from 
<collectingRegion id="2837F83D4B798928E07B41BF2FF390A7" box="[828,916,661,687]" country="Canada" name="Alberta" pageId="10" pageNumber="534">Alberta</collectingRegion>
. This record may represent a misidentification of the wide-spread 
<taxonomicName id="2DF34D5C4B798928E23441902E6690DB" baseAuthorityName="Stibick" baseAuthorityYear="1991" box="[371,513,697,723]" class="Insecta" family="Elateridae" genus="Paradonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="D887EACD4B798928E23441902E6690DB" box="[371,513,697,723]" italics="true" pageId="10" pageNumber="534">P. pectoralis</emphasis>
</taxonomicName>
, which is found throughout 
<collectingRegion id="2837F83D4B798928E00141902FCD90DC" box="[838,938,698,724]" country="United States of America" name="Montana" pageId="10" pageNumber="534">Montana</collectingRegion>
south of 
<collectingRegion id="2837F83D4B798928E7514193280A90DB" box="[1046,1133,697,723]" country="Canada" name="Alberta" pageId="10" pageNumber="534">Alberta</collectingRegion>
, as 
<taxonomicName id="2DF34D5C4B798928E7DD4190292190DB" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[1178,1350,697,724]" class="Insecta" family="Elateridae" genus="Paradonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="obliquatulus">
<emphasis id="D887EACD4B798928E7DD419028CC90DB" box="[1178,1195,698,723]" italics="true" pageId="10" pageNumber="534">P</emphasis>
. 
<emphasis id="D887EACD4B798928E7FD4193292190DB" box="[1210,1350,697,723]" italics="true" pageId="10" pageNumber="534">obliquatulus</emphasis>
</taxonomicName>
is only known as far west as Southeast 
<collectingRegion id="2837F83D4B798928E2B041F72EF390FF" box="[503,660,733,759]" country="United States of America" name="South Dakota" pageId="10" pageNumber="534">South Dakota</collectingRegion>
. In addition to the localities given in 
<bibRefCitation id="8E624B2E4B798928E76A41F728DA90FF" author="Stibick, J. N. L." box="[1069,1213,733,759]" pageId="10" pageNumber="534" pagination="99 - 131" refId="ref16033" refString="Stibick, J. N. L. (1991) North American Negastriinae (Coleoptera, Elateridae): The Negastriinae of the Eastern United States and Adjacent Canada. Insecta Mundi, 4 (1990), 99 - 131." type="journal article" year="1991">Stibick 1991</bibRefCitation>
, I add the following localities: 
<emphasis id="D887EACD4B798928E24B402B2DF29114" bold="true" box="[268,405,769,796]" pageId="10" pageNumber="534">
<collectingRegion id="2837F83D4B798928E24B402B2DF79114" box="[268,400,769,796]" country="United States of America" name="Illinois" pageId="10" pageNumber="534">ILLINOIS</collectingRegion>
:
</emphasis>
Elgin; Havana; and Grand Tower (INHS 830141—INHS 830153). 
<emphasis id="D887EACD4B798928E7DF402B29279114" bold="true" box="[1176,1344,769,796]" pageId="10" pageNumber="534">
<collectingRegion id="2837F83D4B798928E7DF402B295C9114" box="[1176,1339,769,796]" country="United States of America" name="Louisiana" pageId="10" pageNumber="534">LOUISIANA</collectingRegion>
:
</emphasis>
Bossier Parish (4, LSAM), 
<emphasis id="D887EACD4B798928E228400C2E539148" bold="true" box="[367,564,806,832]" pageId="10" pageNumber="534">new state record</emphasis>
.
</paragraph>
<paragraph id="EA4C36DF4B798928E3804063285791A4" blockId="10.[151,1436,481,940]" pageId="10" pageNumber="534">
Distribution (State and Provence only): 
<emphasis id="D887EACD4B798928E1C640632F63916C" bold="true" box="[641,772,841,868]" pageId="10" pageNumber="534">
<collectingCountry id="92E4764F4B798928E1C640632E98916C" box="[641,767,841,868]" name="Canada" pageId="10" pageNumber="534">CANADA</collectingCountry>
:
</emphasis>
<collectingRegion id="2837F83D4B798928E04D40632F07916B" box="[778,864,841,867]" country="Canada" name="Ontario" pageId="10" pageNumber="534">Ontario</collectingRegion>
(
<bibRefCitation id="8E624B2E4B798928E02A40632830916B" author="Bousquet, Y. &amp; Bouchard, P. &amp; Davies, A. E. &amp; Sikes, D. S." box="[877,1111,841,868]" pageId="10" pageNumber="534" refId="ref15082" refString="Bousquet, Y., Bouchard, P., Davies, A. E. &amp; Sikes, D. S. (2013) Checklist of beetles (Coleoptera) of Canada and Alaska, second edition. Pensoft, Sofia, 44 pp. https: // doi. org / 10.3897 / zookeys. 360.4742" type="book" year="2013">Bousquet et al. 2013</bibRefCitation>
). 
<emphasis id="D887EACD4B798928E72F406328CE916C" bold="true" box="[1128,1193,841,868]" pageId="10" pageNumber="534">
<collectingCountry id="92E4764F4B798928E72F406328C3916C" box="[1128,1188,841,868]" name="United States of America" pageId="10" pageNumber="534">USA</collectingCountry>
:
</emphasis>
<collectingRegion id="2837F83D4B798928E7E8406329FF916B" box="[1199,1432,841,867]" country="United States of America" name="District of Columbia" pageId="10" pageNumber="534">District of Columbia</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E3D040472C8E918F" box="[151,233,877,903]" country="United States of America" name="Illinois" pageId="10" pageNumber="534">Illinois</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E3B140472D2A918F" box="[246,333,877,903]" country="United States of America" name="Indiana" pageId="10" pageNumber="534">Indiana</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E21D40442DF29180" box="[346,405,878,904]" country="United States of America" name="Iowa" pageId="10" pageNumber="534">Iowa</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E2E640442D929180" box="[417,501,878,904]" country="United States of America" name="Kansas" pageId="10" pageNumber="534">Kansas</collectingRegion>
, 
<emphasis id="D887EACD4B798928E14540472E1F9180" bold="true" box="[514,632,877,904]" pageId="10" pageNumber="534">
<collectingRegion id="2837F83D4B798928E14540472E1F9180" box="[514,632,877,904]" country="United States of America" name="Louisiana" pageId="10" pageNumber="534">Louisiana</collectingRegion>
</emphasis>
, 
<collectingRegion id="2837F83D4B798928E1CF40472EB5918F" box="[648,722,877,903]" country="United States of America" name="Maine" pageId="10" pageNumber="534">Maine</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E19840472F2A918F" box="[735,845,877,903]" country="United States of America" name="Michigan" pageId="10" pageNumber="534">Michigan</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E01D40442FB89180" box="[858,991,878,904]" country="United States of America" name="New Jersey" pageId="10" pageNumber="534">New Jersey</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E0AC40442805918F" box="[1003,1122,877,904]" country="United States of America" name="New York" pageId="10" pageNumber="534">New York</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E729404728CE918F" box="[1134,1193,877,903]" country="United States of America" name="Ohio" pageId="10" pageNumber="534">Ohio</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E7F140472929918F" box="[1206,1358,877,903]" country="United States of America" name="Pennsylvania" pageId="10" pageNumber="534">Pennsylvania</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E61C40472C8C91A3" country="United States of America" name="South Dakota" pageId="10" pageNumber="534">South Dakota</collectingRegion>
, 
<collectingRegion id="2837F83D4B798928E3B140BB2D0891A3" box="[246,367,913,939]" country="United States of America" name="Wisconsin" pageId="10" pageNumber="534">Wisconsin</collectingRegion>
. (
<bibRefCitation id="8E624B2E4B798928E2C740BB2E7591A3" author="Stibick, J. N. L." box="[384,530,913,939]" pageId="10" pageNumber="534" pagination="99 - 131" refId="ref16033" refString="Stibick, J. N. L. (1991) North American Negastriinae (Coleoptera, Elateridae): The Negastriinae of the Eastern United States and Adjacent Canada. Insecta Mundi, 4 (1990), 99 - 131." type="journal article" year="1991">Stibick 1991</bibRefCitation>
, bolded states are new state records noted here).
</paragraph>
</subSubSection>
</treatment>
</document>