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<mods:title id="5859956291C8D95F762444E1FE5A0973">Taxonomic review of the Thysanoptyx Hampson generic complex (Lepidoptera: Erebidae: Arctiinae: Lithosiini). II. The genus Thysanoplisna Volynkin with descriptions of seven new species from Vietnam and mainland China</mods:title>
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<subSubSection id="C37B6553FFFEFFE9FF7CBEA2FDB8FA4E" box="[151,610,1446,1473]" pageId="1" pageNumber="429" type="nomenclature">
<paragraph id="8BDE36D8FFFEFFE9FF7CBEA2FDB8FA4E" blockId="1.[151,610,1446,1473]" box="[151,610,1446,1473]" pageId="1" pageNumber="429">
<heading id="D09681B4FFFEFFE9FF7CBEA2FDB8FA4E" bold="true" box="[151,610,1446,1473]" fontSize="11" level="1" pageId="1" pageNumber="429" reason="1">
<emphasis id="B915EACAFFFEFFE9FF7CBEA2FDB8FA4E" bold="true" box="[151,610,1446,1473]" pageId="1" pageNumber="429">
Genus
<taxonomicName id="4C614D5BFFFEFFE9FF05BEA2FDB8FA4E" ID-CoL="BXN3N" authority="Volynkin, 2023" authorityName="Volynkin" authorityYear="2023" box="[238,610,1446,1473]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFEFFE9FF05BEA2FE7AFA4F" bold="true" box="[238,416,1446,1473]" italics="true" pageId="1" pageNumber="429">Thysanoplisna</emphasis>
<bibRefCitation id="EFF04B29FFFEFFE9FE4CBEA2FDB8FA4E" author="Volynkin, A. V." box="[423,610,1446,1472]" pageId="1" pageNumber="429" pagination="54 - 63" refId="ref10816" refString="Volynkin, A. V. (2023) On the taxonomy of the genus Thysanoptyx Hampson with descriptions of two new genera and a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 69, 54 - 63. https: // doi. org / 10.37828 / em. 2023.69.6" type="journal article" year="2023">Volynkin, 2023</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37B6553FFFEFFE9FF7CBEEFFCF3F98D" box="[151,809,1515,1539]" pageId="1" pageNumber="429" type="reference_group">
<paragraph id="8BDE36D8FFFEFFE9FF7CBEEFFCF3F98D" blockId="1.[151,809,1515,1539]" box="[151,809,1515,1539]" pageId="1" pageNumber="429">
<taxonomicName id="4C614D5BFFFEFFE9FF7CBEEFFE15F98D" ID-CoL="BXN3N" authority="Volynkin, 2023" authorityName="Volynkin" authorityYear="2023" box="[151,463,1515,1539]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFEFFE9FF7CBEEFFEF1F98D" box="[151,299,1515,1539]" italics="true" pageId="1" pageNumber="429">Thysanoplisna</emphasis>
<bibRefCitation id="EFF04B29FFFEFFE9FEDABEEFFE15F98D" author="Volynkin, A. V." box="[305,463,1515,1539]" pageId="1" pageNumber="429" pagination="54 - 63" refId="ref10816" refString="Volynkin, A. V. (2023) On the taxonomy of the genus Thysanoptyx Hampson with descriptions of two new genera and a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 69, 54 - 63. https: // doi. org / 10.37828 / em. 2023.69.6" type="journal article" year="2023">Volynkin, 2023</bibRefCitation>
</taxonomicName>
,
<emphasis id="B915EACAFFFEFFE9FE32BEEFFD0CF98D" box="[473,726,1515,1539]" italics="true" pageId="1" pageNumber="429">
Ecologica
<taxonomicName id="4C614D5BFFFEFFE9FDADBEE8FD0CF98D" ID-CoL="7P98C" authorityName="O. Boettger" authorityYear="1877" box="[582,726,1516,1539]" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="429" phylum="Mollusca" rank="genus">Montenegrina</taxonomicName>
</emphasis>
, 69: 55.
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFEFFE9FF7CBD36FA49F9E1" pageId="1" pageNumber="429" type="materials_examined">
<paragraph id="8BDE36D8FFFEFFE9FF7CBD36FC21F9C5" blockId="1.[151,1437,1585,2008]" box="[151,1019,1585,1612]" pageId="1" pageNumber="429">
<typeStatus id="54DA887AFFFEFFE9FF7CBD36FF15F9C2" box="[151,207,1586,1612]" pageId="1" pageNumber="429">Type</typeStatus>
species:
<taxonomicName id="4C614D5BFFFEFFE9FEDEBD35FD3CF9C5" authority="Alpheraky, 1897" authorityName="Alpheraky" authorityYear="1897" box="[309,742,1585,1611]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFEFFE9FEDEBD35FDF9F9C5" box="[309,547,1585,1611]" italics="true" pageId="1" pageNumber="429">Lithosia brevimacula</emphasis>
<bibRefCitation id="EFF04B29FFFEFFE9FDC2BD35FD3CF9C5" author="Alpheraky, S." box="[553,742,1585,1611]" pageId="1" pageNumber="429" pagination="83 - 149" refId="ref10180" refString="Alpheraky, S. (1897) Lepidopteres des provinces chinoises Se-Tchouen et Kam recueillis, en 1893, par M-r G. N. Potanine. In: Romanoff, N. M. (Ed.), Memoires sur les lepidopteres. 9. P. P. Soikine, St. Petersburg, pp. 83 - 149. [in French]" type="book chapter" year="1897">Alphéraky, 1897</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BDE36D8FFFEFFE9FF2CBD51FA49F9E1" blockId="1.[151,1437,1585,2008]" box="[199,1427,1621,1648]" pageId="1" pageNumber="429">
<emphasis id="B915EACAFFFEFFE9FF2CBD51FF25F9E1" bold="true" box="[199,255,1621,1647]" pageId="1" pageNumber="429">Note</emphasis>
. The genus is herein re-characterised following the genitalia terminology suggested by Volynkin (2024).
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFEFFEAFF2CBD7DFE15FC9B" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429" type="diagnosis">
<paragraph id="8BDE36D8FFFEFFE9FF2CBD7DFD44F8E2" blockId="1.[151,1437,1585,2008]" pageId="1" pageNumber="429">
<emphasis id="B915EACAFFFEFFE9FF2CBD7DFEE0F91D" bold="true" box="[199,314,1657,1683]" pageId="1" pageNumber="429">Diagnosis</emphasis>
.
<taxonomicName id="4C614D5BFFFEFFE9FEA2BD7DFE36F91D" authorityName="Volynkin" authorityYear="2023" box="[329,492,1657,1683]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFEFFE9FEA2BD7DFE36F91D" box="[329,492,1657,1683]" italics="true" pageId="1" pageNumber="429">Thysanoplisna</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFEFFE9FE17BD7DFDB5F91D" box="[508,623,1657,1683]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 916" captionStart-2="FIGURES 1724" captionStart-3="FIGURES 2532" captionStart-4="FIGURES 3340" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="6.[151,264,1943,1967]" captionStartId-2="10.[151,264,1943,1967]" captionStartId-3="11.[151,264,1943,1967]" captionStartId-4="14.[151,264,1943,1967]" captionTargetBox-0="[190,1397,184,1911]" captionTargetBox-1="[199,1388,182,1909]" captionTargetBox-2="[204,1383,188,1909]" captionTargetBox-3="[197,1391,182,1909]" captionTargetBox-4="[184,1420,185,1909]" captionTargetId-0="figure-15@4.[177,1410,181,1919]" captionTargetId-1="figure-15@6.[186,1401,181,1919]" captionTargetId-2="figure-15@10.[191,1395,181,1919]" captionTargetId-3="figure-26@11.[184,1404,181,1919]" captionTargetId-4="figure-15@14.[158,1430,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="14" captionText-0="FIGURES 18. Adults of Thysanoplisna spp. Depositories of the specimens: 1, 7 and 8 in MWM/ZSM; 2, 5 and 6 in NHMUK (©The Trustees of NHMUK); 3 and 4 in AFM." captionText-1="FIGURES 916. Adults of Thysanoplisna spp. The specimens are deposited in MWM/ZSM." captionText-2="FIGURES 1724. Adults of Thysanoplisna spp. The specimens are deposited in MWM/ZSM." captionText-3="FIGURES 2532. Adults of Thysanoplisna spp. Depositories of the specimens: 25, 26, and 2931 in MWM/ZSM; 27, 28 and 32 in NHMUK (©The Trustees of NHMUK)." captionText-4="FIGURES 3340. Adults of Thysanoplisna spp. (3336), Caliginoptyx vientiana (37, 38), and Medioptyx kishidai (39, 40). Depositories of the specimens: 3337, 39 and 40 in MWM/ZSM (37: ex CKC); 38 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.13747213" figureDoi-1="http://doi.org/10.5281/zenodo.13747215" figureDoi-2="http://doi.org/10.5281/zenodo.13747217" figureDoi-3="http://doi.org/10.5281/zenodo.13747219" figureDoi-4="http://doi.org/10.5281/zenodo.13747223" httpUri-0="https://zenodo.org/record/13747213/files/figure.png" httpUri-1="https://zenodo.org/record/13747215/files/figure.png" httpUri-2="https://zenodo.org/record/13747217/files/figure.png" httpUri-3="https://zenodo.org/record/13747219/files/figure.png" httpUri-4="https://zenodo.org/record/13747223/files/figure.png" pageId="1" pageNumber="429">Figs 136</figureCitation>
) is morphologically most similar to the recently described
<emphasis id="B915EACAFFFEFFE9FAE7BD7DFA47F91D" box="[1292,1437,1657,1683]" italics="true" pageId="1" pageNumber="429">Caliginoptyx</emphasis>
(
<figureCitation id="135A2A5DFFFEFFE9FF74BD99FEFCF939" box="[159,294,1693,1719]" captionStart="FIGURES 3340" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1420,185,1909]" captionTargetId="figure-15@14.[158,1430,181,1919]" captionTargetPageId="14" captionText="FIGURES 3340. Adults of Thysanoplisna spp. (3336), Caliginoptyx vientiana (37, 38), and Medioptyx kishidai (39, 40). Depositories of the specimens: 3337, 39 and 40 in MWM/ZSM (37: ex CKC); 38 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13747223" httpUri="https://zenodo.org/record/13747223/files/figure.png" pageId="1" pageNumber="429">Figs 37, 38</figureCitation>
) and
<taxonomicName id="4C614D5BFFFEFFE9FE87BD99FE38F939" box="[364,482,1693,1719]" genus="Medioptyx" pageId="1" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFEFFE9FE87BD99FE38F939" box="[364,482,1693,1719]" italics="true" pageId="1" pageNumber="429">Medioptyx</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFEFFE9FE1EBD99FDA7F939" box="[501,637,1693,1719]" captionStart="FIGURES 3340" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1420,185,1909]" captionTargetId="figure-15@14.[158,1430,181,1919]" captionTargetPageId="14" captionText="FIGURES 3340. Adults of Thysanoplisna spp. (3336), Caliginoptyx vientiana (37, 38), and Medioptyx kishidai (39, 40). Depositories of the specimens: 3337, 39 and 40 in MWM/ZSM (37: ex CKC); 38 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13747223" httpUri="https://zenodo.org/record/13747223/files/figure.png" pageId="1" pageNumber="429">Figs 39, 40</figureCitation>
) but superficially differs in the longer distal section of the forewing (between the cell and the outer margin), the ochreous grey or ochreous brown forewing ground colour (it is pale brownish-grey in
<emphasis id="B915EACAFFFEFFE9FEB7BDE1FE37F971" box="[348,493,1765,1791]" italics="true" pageId="1" pageNumber="429">Caliginoptyx</emphasis>
and bright ochreous yellow in
<taxonomicName id="4C614D5BFFFEFFE9FCA9BDE1FC62F971" box="[834,952,1765,1791]" genus="Medioptyx" pageId="1" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFEFFE9FCA9BDE1FC62F971" box="[834,952,1765,1791]" italics="true" pageId="1" pageNumber="429">Medioptyx</emphasis>
</taxonomicName>
), and the short trapezoid or rounded medial black spot, which is more similar to
<taxonomicName id="4C614D5BFFFEFFE9FDD4BC0DFCE4F8AD" authority="Walker, 1862" authorityName="Walker" authorityYear="1862" box="[575,830,1801,1827]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B915EACAFFFEFFE9FDD4BC0DFD47F8AD" box="[575,669,1801,1827]" italics="true" pageId="1" pageNumber="429">Teulisna</emphasis>
Walker, 1862
</taxonomicName>
<emphasis id="B915EACAFFFEFFE9FCACBC0EFC5EF8AD" box="[839,900,1802,1827]" italics="true" pageId="1" pageNumber="429">
<taxonomicNameLabel id="A22657B1FFFEFFE9FCACBC0EFC5EF8AD" box="[839,900,1802,1827]" pageId="1" pageNumber="429" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C614D5BFFFEFFE9FC54BC0DFB2EF8AD" authority="Volynkin, 2020" authorityName="Volynkin" authorityYear="2020" box="[959,1268,1801,1827]" class="Insecta" family="Erebidae" genus="Witteulisna" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFEFFE9FC54BC0DFBE7F8AD" box="[959,1085,1801,1827]" italics="true" pageId="1" pageNumber="429">Witteulisna</emphasis>
<bibRefCitation id="EFF04B29FFFEFFE9FBADBC0DFB2EF8AD" author="Volynkin, A. V." box="[1094,1268,1801,1827]" pageId="1" pageNumber="429" pagination="127 - 130" refId="ref10715" refString="Volynkin, A. V. (2020) Witteulisna, a new genus for a new species from North Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 38, 127 - 130. https: // doi. org / 10.37828 / em. 2020.38.16" type="journal article" year="2020">Volynkin, 2020</bibRefCitation>
</taxonomicName>
(illustrated by
<bibRefCitation id="EFF04B29FFFEFFE9FF7CBC29FE9FF8C9" author="Volynkin, A. V." box="[151,325,1837,1863]" pageId="1" pageNumber="429" pagination="127 - 130" refId="ref10715" refString="Volynkin, A. V. (2020) Witteulisna, a new genus for a new species from North Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 38, 127 - 130. https: // doi. org / 10.37828 / em. 2020.38.16" type="journal article" year="2020">Volynkin 2020</bibRefCitation>
), whereas it is shorter, diffuse and greyish-brown in
<emphasis id="B915EACAFFFEFFE9FC5CBC29FB92F8C9" box="[951,1096,1837,1863]" italics="true" pageId="1" pageNumber="429">Caliginoptyx</emphasis>
and considerably longer and occupying most of the forewing in
<taxonomicName id="4C614D5BFFFEFFE9FDCABC55FD4DF8E5" box="[545,663,1873,1899]" genus="Medioptyx" pageId="1" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFEFFE9FDCABC55FD4DF8E5" box="[545,663,1873,1899]" italics="true" pageId="1" pageNumber="429">Medioptyx</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE36D8FFFEFFEAFF2CBC71FF35FE23" blockId="1.[151,1437,1585,2008]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429">
The male genital capsule structure of
<taxonomicName id="4C614D5BFFFEFFE9FD9CBC71FCC0F801" authorityName="Volynkin" authorityYear="2023" box="[631,794,1909,1935]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFEFFE9FD9CBC71FCC0F801" box="[631,794,1909,1935]" italics="true" pageId="1" pageNumber="429">Thysanoplisna</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFEFFE9FCC0BC71FC77F81E" box="[811,941,1909,1936]" captionStart-0="FIGURES 4144" captionStart-1="FIGURES 4548" captionStart-2="FIGURES 4952" captionStart-3="FIGURES 5355" captionStartId-0="15.[151,264,1943,1967]" captionStartId-1="16.[151,264,1943,1967]" captionStartId-2="17.[151,264,1943,1967]" captionStartId-3="18.[151,264,1943,1967]" captionTargetBox-0="[226,1371,185,1912]" captionTargetBox-1="[280,1308,181,1916]" captionTargetBox-2="[206,1382,181,1912]" captionTargetBox-3="[174,1413,186,1912]" captionTargetId-0="figure-26@15.[213,1375,181,1919]" captionTargetId-1="figure-15@16.[268,1320,181,1919]" captionTargetId-2="figure-26@17.[193,1394,181,1919]" captionTargetId-3="figure-15@18.[161,1426,181,1919]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURES 4144. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." captionText-1="FIGURES 4548. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." captionText-2="FIGURES 4952. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." captionText-3="FIGURES 5355. Male genitalia of Thysanoplisna anomala (53), Caliginoptyx vientiana (54), and Medioptyx kishidai (55). The specimens dissected are deposited in MWM/ZSM." figureDoi-0="http://doi.org/10.5281/zenodo.13747225" figureDoi-1="http://doi.org/10.5281/zenodo.13747229" figureDoi-2="http://doi.org/10.5281/zenodo.13747231" figureDoi-3="http://doi.org/10.5281/zenodo.13747233" httpUri-0="https://zenodo.org/record/13747225/files/figure.png" httpUri-1="https://zenodo.org/record/13747229/files/figure.png" httpUri-2="https://zenodo.org/record/13747231/files/figure.png" httpUri-3="https://zenodo.org/record/13747233/files/figure.png" pageId="1" pageNumber="429">Figs 4153</figureCitation>
) is characterised by the combination of the following features. (1) In the conjuga, the processus momenti is weakly sclerotised while the tendon is thick and heavily sclerotised and performs a skeletal function (vs. in
<emphasis id="B915EACAFFFEFFE9FCACBCB9FC02F859" box="[839,984,1981,2007]" italics="true" pageId="1" pageNumber="429">Caliginoptyx</emphasis>
(
<figureCitation id="135A2A5DFFFEFFE9FC01BCB9FBE5F856" box="[1002,1087,1981,2008]" captionStart="FIGURES 5355" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[174,1413,186,1912]" captionTargetId="figure-15@18.[161,1426,181,1919]" captionTargetPageId="18" captionText="FIGURES 5355. Male genitalia of Thysanoplisna anomala (53), Caliginoptyx vientiana (54), and Medioptyx kishidai (55). The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747233" httpUri="https://zenodo.org/record/13747233/files/figure.png" pageId="1" pageNumber="429">Fig. 54</figureCitation>
) and
<taxonomicName id="4C614D5BFFFEFFE9FB6FBCB9FB20F859" box="[1156,1274,1981,2007]" genus="Medioptyx" pageId="1" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFEFFE9FB6FBCB9FB20F859" box="[1156,1274,1981,2007]" italics="true" pageId="1" pageNumber="429">Medioptyx</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFEFFE9FAE7BCB9FAB9F856" box="[1292,1379,1981,2008]" captionStart="FIGURES 5355" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[174,1413,186,1912]" captionTargetId="figure-15@18.[161,1426,181,1919]" captionTargetPageId="18" captionText="FIGURES 5355. Male genitalia of Thysanoplisna anomala (53), Caliginoptyx vientiana (54), and Medioptyx kishidai (55). The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747233" httpUri="https://zenodo.org/record/13747233/files/figure.png" pageId="1" pageNumber="429">Fig. 55</figureCitation>
), the conjuga is formed by the equally heavily sclerotised processus momenti and tendon). (2) The juxta consisting of two thin rod-like lateral plates, which are dorsally connected by the thin bridge and fused with two rod-like lobes of the caulis stretching laterally along the anterior (cranial) wall of the juxtal fold. (3) The membranous anellus. The juxta and anellus structures of
<taxonomicName id="4C614D5BFFFDFFEAFE44BA07FD88FE93" authorityName="Volynkin" authorityYear="2023" box="[431,594,259,285]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFE44BA07FD88FE93" box="[431,594,259,285]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
are similar to
<emphasis id="B915EACAFFFDFFEAFD1EBA07FC5CFE93" box="[757,902,259,285]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
whereas in
<taxonomicName id="4C614D5BFFFDFFEAFBE7BA07FB58FE93" box="[1036,1154,259,285]" genus="Medioptyx" pageId="2" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFDFFEAFBE7BA07FB58FE93" box="[1036,1154,259,285]" italics="true" pageId="2" pageNumber="430">Medioptyx</emphasis>
</taxonomicName>
, the dorsal section of the juxta is conical and well-sclerotised, and the anellus bears two broad ventro-lateral sclerotised plates. The vesica structure of
<taxonomicName id="4C614D5BFFFDFFEAFECEBA4FFE12FEEB" authorityName="Volynkin" authorityYear="2023" box="[293,456,331,357]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFECEBA4FFE12FEEB" box="[293,456,331,357]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
is similar to
<emphasis id="B915EACAFFFDFFEAFD82BA4FFD20FEEB" box="[617,762,331,357]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
with its sub-ejaculatorial section situated basally, but that of
<taxonomicName id="4C614D5BFFFDFFEAFF5CBA6BFE80FE07" authorityName="Volynkin" authorityYear="2023" box="[183,346,367,393]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFF5CBA6BFE80FE07" box="[183,346,367,393]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
lacks the diverticula. Unlike in
<taxonomicName id="4C614D5BFFFDFFEAFD20BA6BFCB4FE07" authorityName="Volynkin" authorityYear="2023" box="[715,878,367,393]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFD20BA6BFCB4FE07" box="[715,878,367,393]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
, the vesica ejaculatorius of
<taxonomicName id="4C614D5BFFFDFFEAFB44BA6BFAFFFE07" box="[1199,1317,367,393]" genus="Medioptyx" pageId="2" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFDFFEAFB44BA6BFAFFFE07" box="[1199,1317,367,393]" italics="true" pageId="2" pageNumber="430">Medioptyx</emphasis>
</taxonomicName>
originates distally.
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBAB3FE9BFD43" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="430">
The female genitalia of
<taxonomicName id="4C614D5BFFFDFFEAFE38BAB3FDACFE5F" authorityName="Volynkin" authorityYear="2023" box="[467,630,439,465]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFE38BAB3FDACFE5F" box="[467,630,439,465]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFDFFEAFD6EBAB3FCDCFE5F" box="[645,774,438,465]" captionStart-0="FIGURES 5661" captionStart-1="FIGURES 6267" captionStart-2="FIGURES 6870" captionStartId-0="19.[151,264,1932,1956]" captionStartId-1="20.[151,264,1932,1956]" captionStartId-2="21.[151,264,1066,1090]" captionTargetBox-0="[195,1392,184,1895]" captionTargetBox-1="[174,1413,181,1895]" captionTargetBox-2="[162,1428,188,1035]" captionTargetId-0="figure-26@19.[182,1405,181,1908]" captionTargetId-1="figure-15@20.[161,1426,181,1908]" captionTargetId-2="figure-26@21.[155,1431,181,1041]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURES 5661. Female genitalia of Thysanoplisna spp. Depositories of the specimens dissected: 56 and 57 in NHMUK (©The Trustees of NHMUK); 5861 in MWM/ZSM." captionText-1="FIGURES 6267. Female genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." captionText-2="FIGURES 6870. Female genitalia of Thysanoplisna anomala (68), Caliginoptyx vientiana (69), and Medioptyx kishidai (70). Depositories of the specimens dissected: 68 and 70 in MWM/ZSM; 69 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.13747235" figureDoi-1="http://doi.org/10.5281/zenodo.13747237" figureDoi-2="http://doi.org/10.5281/zenodo.13747239" httpUri-0="https://zenodo.org/record/13747235/files/figure.png" httpUri-1="https://zenodo.org/record/13747237/files/figure.png" httpUri-2="https://zenodo.org/record/13747239/files/figure.png" pageId="2" pageNumber="430">Figs 5668</figureCitation>
) are most similar to
<emphasis id="B915EACAFFFDFFEAFC06BAB3FBA4FE5F" box="[1005,1150,439,465]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
(
<figureCitation id="135A2A5DFFFDFFEAFB66BAB3FB04FE5F" box="[1165,1246,439,465]" captionStart="FIGURES 6870" captionStartId="21.[151,264,1066,1090]" captionTargetBox="[162,1428,188,1035]" captionTargetId="figure-26@21.[155,1431,181,1041]" captionTargetPageId="21" captionText="FIGURES 6870. Female genitalia of Thysanoplisna anomala (68), Caliginoptyx vientiana (69), and Medioptyx kishidai (70). Depositories of the specimens dissected: 68 and 70 in MWM/ZSM; 69 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13747239" httpUri="https://zenodo.org/record/13747239/files/figure.png" pageId="2" pageNumber="430">Fig. 69</figureCitation>
) but differ in the flat and evenly sclerotised ductus bursae (it is less dorso-ventrally flattened and has postero-lateral membranous areas in
<emphasis id="B915EACAFFFDFFEAFF17BAFBFE57FD97" box="[252,397,511,537]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
), the compact, elliptical or rounded signum bursae (it is markedly larger and ribbon-like in
<emphasis id="B915EACAFFFDFFEAFF5CB927FE92FDB3" box="[183,328,547,573]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
), and the ventrally situated appendix bursae (vs. dorsal position in
<emphasis id="B915EACAFFFDFFEAFBB7B927FB37FDB3" box="[1116,1261,547,573]" italics="true" pageId="2" pageNumber="430">Caliginoptyx</emphasis>
). Compared to
<taxonomicName id="4C614D5BFFFDFFEAFF7CB943FED7FDEF" box="[151,269,583,609]" genus="Medioptyx" pageId="2" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFDFFEAFF7CB943FED7FDEF" box="[151,269,583,609]" italics="true" pageId="2" pageNumber="430">Medioptyx</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A5DFFFDFFEAFEC9B943FEA0FDEF" box="[290,378,583,609]" captionStart="FIGURES 6870" captionStartId="21.[151,264,1066,1090]" captionTargetBox="[162,1428,188,1035]" captionTargetId="figure-26@21.[155,1431,181,1041]" captionTargetPageId="21" captionText="FIGURES 6870. Female genitalia of Thysanoplisna anomala (68), Caliginoptyx vientiana (69), and Medioptyx kishidai (70). Depositories of the specimens dissected: 68 and 70 in MWM/ZSM; 69 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13747239" httpUri="https://zenodo.org/record/13747239/files/figure.png" pageId="2" pageNumber="430">Fig. 70</figureCitation>
), the female genitalia of
<taxonomicName id="4C614D5BFFFDFFEAFD42B943FC96FDEF" authorityName="Volynkin" authorityYear="2023" box="[681,844,583,609]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFD42B943FC96FDEF" box="[681,844,583,609]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
have a medially constricted corpus bursae with a posterior section having a gelatinous enlargement protruding ventrally or laterally, and an anterior section bearing a signum, whereas in
<taxonomicName id="4C614D5BFFFDFFEAFE79B98BFDD2FD27" box="[402,520,655,681]" genus="Medioptyx" pageId="2" pageNumber="428" rank="genus">
<emphasis id="B915EACAFFFDFFEAFE79B98BFDD2FD27" box="[402,520,655,681]" italics="true" pageId="2" pageNumber="430">Medioptyx</emphasis>
</taxonomicName>
, the corpus bursae is sack-like and lacks both the posterior enlargement and the signum bursae.
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CB9D3FE15FC9B" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="430">
<emphasis id="B915EACAFFFDFFEAFF2CB9D3FF25FD7F" bold="true" box="[199,255,727,753]" pageId="2" pageNumber="430">Note</emphasis>
. Species of the genus significantly vary in their size and the size and shape of the forewing medial spot within the same population.
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFDFFEAFF2CB81AFDEAF9A3" pageId="2" pageNumber="430" type="distribution">
<paragraph id="8BDE36D8FFFDFFEAFF2CB81AFC78FCD3" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="430">
<emphasis id="B915EACAFFFDFFEAFF2CB81AFE83FCB6" bold="true" box="[199,345,798,824]" pageId="2" pageNumber="430">Distribution</emphasis>
. Species of the genus are known from northern Mainland Southeast Asia (
<collectingCountry id="F3767648FFFDFFEAFB51B81BFAC5FCB7" box="[1210,1311,799,825]" name="Vietnam" pageId="2" pageNumber="430">Vietnam</collectingCountry>
), mainland
<collectingCountry id="F3767648FFFDFFEAFF7CB847FF00FCD3" box="[151,218,835,861]" name="China" pageId="2" pageNumber="430">China</collectingCountry>
and the island of
<collectingRegion id="49A5F83AFFFDFFEAFE49B847FE2CFCD3" box="[418,502,835,861]" country="China" name="Hainan" pageId="2" pageNumber="430">Hainan</collectingRegion>
, but absent from the island of
<collectingCountry id="F3767648FFFDFFEAFCA3B847FC44FCD3" box="[840,926,835,861]" name="Taiwan" pageId="2" pageNumber="430">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF7CB8AAFDEAFC47" blockId="2.[151,560,942,969]" box="[151,560,942,969]" pageId="2" pageNumber="430">
<emphasis id="B915EACAFFFDFFEAFF7CB8AAFDEAFC47" bold="true" box="[151,560,942,969]" pageId="2" pageNumber="430">
Species content of
<taxonomicName id="4C614D5BFFFDFFEAFE95B8AAFDEAFC47" authorityName="Volynkin" authorityYear="2023" box="[382,560,942,969]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B915EACAFFFDFFEAFE95B8AAFDEAFC47" bold="true" box="[382,560,942,969]" italics="true" pageId="2" pageNumber="430">Thysanoplisna</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF7CB8F3FE31FB9F" blockId="2.[151,615,1015,1293]" box="[151,491,1015,1041]" pageId="2" pageNumber="430">
The
<taxonomicName id="4C614D5BFFFDFFEAFF21B8F3FE9DFB9E" baseAuthorityName="Walker" baseAuthorityYear="1854" box="[202,327,1015,1041]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFDFFEAFF21B8F3FE9DFB9E" box="[202,327,1015,1041]" italics="true" pageId="2" pageNumber="430">Th. signata</emphasis>
</taxonomicName>
species-group
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBF1FFDD3FBD7" blockId="2.[151,615,1015,1293]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BF1FFDBDFBBB" authority="(Alpheraky, 1897)" baseAuthorityName="Alpheraky" baseAuthorityYear="1897" box="[220,615,1051,1077]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFDFFEAFF37BF1FFE49FBBB" box="[220,403,1051,1077]" italics="true" pageId="2" pageNumber="430">Th. brevimacula</emphasis>
(
<bibRefCitation id="EFF04B29FFFDFFEAFE4ABF1FFD84FBBB" author="Alpheraky, S." box="[417,606,1051,1077]" pageId="2" pageNumber="430" pagination="83 - 149" refId="ref10180" refString="Alpheraky, S. (1897) Lepidopteres des provinces chinoises Se-Tchouen et Kam recueillis, en 1893, par M-r G. N. Potanine. In: Romanoff, N. M. (Ed.), Memoires sur les lepidopteres. 9. P. P. Soikine, St. Petersburg, pp. 83 - 149. [in French]" type="book chapter" year="1897">Alphéraky, 1897</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="4C614D5BFFFDFFEAFF37BF3BFDD3FBD7" authority="(Walker, 1854)" baseAuthorityName="Walker" baseAuthorityYear="1854" box="[220,521,1086,1113]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFDFFEAFF37BF3BFE80FBD6" box="[220,346,1087,1113]" italics="true" pageId="2" pageNumber="430">Th. signata</emphasis>
(
<bibRefCitation id="EFF04B29FFFDFFEAFE82BF3BFDDBFBD7" author="Walker, F." box="[361,513,1086,1113]" pageId="2" pageNumber="430" refId="ref11071" refString="Walker, F. (1854) Part I. - Lepidoptera Heterocera. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Vol. 2. Edward Newman, London, pp. 1 - 775." type="book" year="1854">Walker, 1854</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBF67FDDFFBF3" blockId="2.[151,615,1015,1293]" box="[199,517,1123,1149]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BF67FE7FFBF3" authorityName="Volynkin" authorityYear="2024" box="[220,421,1123,1149]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BF67FE7FFBF3" box="[220,421,1123,1149]" italics="true" pageId="2" pageNumber="430">Th. grandimacula</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFE47BF67FDDFFBF3" bold="true" box="[428,517,1123,1149]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFE47BF67FDDFFBF3" box="[428,517,1123,1149]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBF83FE6CFB2F" blockId="2.[151,615,1015,1293]" box="[199,438,1159,1185]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BF83FE8CFB2E" authorityName="Volynkin" authorityYear="2024" box="[220,342,1159,1185]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="minuta" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BF83FE8CFB2E" box="[220,342,1159,1185]" italics="true" pageId="2" pageNumber="430">Th. minuta</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFEB6BF83FE6CFB2F" bold="true" box="[349,438,1159,1185]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFEB6BF83FE6CFB2F" box="[349,438,1159,1185]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBFAFFE02FB4B" blockId="2.[151,615,1015,1293]" box="[199,472,1195,1221]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BFAFFEA2FB4A" authorityName="Volynkin" authorityYear="2024" box="[220,376,1195,1221]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="monotona" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BFAFFEA2FB4A" box="[220,376,1195,1221]" italics="true" pageId="2" pageNumber="430">Th. monotona</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFE94BFAFFE02FB4B" bold="true" box="[383,472,1195,1221]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFE94BFAFFE02FB4B" box="[383,472,1195,1221]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBFCBFE1BFB67" blockId="2.[151,615,1015,1293]" box="[199,449,1231,1257]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BFCBFEBBFB67" authorityName="Volynkin" authorityYear="2024" box="[220,353,1231,1257]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BFCBFEBBFB67" box="[220,353,1231,1257]" italics="true" pageId="2" pageNumber="430">Th. bachma</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFE8CBFCBFE1BFB67" bold="true" box="[359,449,1231,1257]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFE8CBFCBFE1BFB67" box="[359,449,1231,1257]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBFF7FE62FA83" blockId="2.[151,615,1015,1293]" box="[199,440,1267,1293]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BFF7FE8DFA83" authorityName="Volynkin" authorityYear="2024" box="[220,343,1267,1293]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="suffusa" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BFF7FE8DFA83" box="[220,343,1267,1293]" italics="true" pageId="2" pageNumber="430">Th. suffusa</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFEB5BFF7FE62FA83" bold="true" box="[350,440,1267,1293]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFEB5BFF7FE62FA83" box="[350,440,1267,1293]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF7CBE3FFE3DFADB" blockId="2.[151,621,1339,1473]" box="[151,487,1339,1365]" pageId="2" pageNumber="430">
The
<taxonomicName id="4C614D5BFFFDFFEAFF21BE3FFE99FADB" authorityName="Leech" authorityYear="1899" box="[202,323,1339,1365]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="directa">
<emphasis id="B915EACAFFFDFFEAFF21BE3FFE99FADB" box="[202,323,1339,1365]" italics="true" pageId="2" pageNumber="430">Th. directa</emphasis>
</taxonomicName>
species-group
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBE5BFDB7FAF7" blockId="2.[151,621,1339,1473]" box="[199,621,1375,1401]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BE5BFE26FAF7" authority="(Leech, 1899)" authorityName="Volynkin" authorityYear="2024" baseAuthorityName="Leech" baseAuthorityYear="1899" box="[220,508,1375,1401]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="directa" status="stat. rev.">
<emphasis id="B915EACAFFFDFFEAFF37BE5BFE8CFAF7" box="[220,342,1375,1401]" italics="true" pageId="2" pageNumber="430">Th. directa</emphasis>
(
<bibRefCitation id="EFF04B29FFFDFFEAFE8EBE5BFE2FFAF7" author="Leech, J. H." box="[357,501,1375,1401]" pageId="2" pageNumber="430" pagination="99 - 219" refId="ref10596" refString="Leech, J. H. (1899) Lepidoptera Heterocera from Northern China, Japan and Corea. Part II. Transactions of the Entomological Society of London, 1899 (1), 99 - 219. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 03304. x" type="journal article" year="1899">Leech, 1899</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B915EACAFFFDFFEAFDE3BE5BFDB7FAF7" bold="true" box="[520,621,1375,1401]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFDE3BE5BFDB7FAF7" box="[520,621,1375,1401]" pageId="2" pageNumber="430">stat. rev.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBE87FE0CFA13" blockId="2.[151,621,1339,1473]" box="[199,470,1411,1437]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BE87FEAFFA13" authorityName="Volynkin" authorityYear="2024" box="[220,373,1411,1437]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BE87FEAFFA13" box="[220,373,1411,1437]" italics="true" pageId="2" pageNumber="430">Th. tenebrosa</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFE97BE87FE0CFA13" bold="true" box="[380,470,1411,1437]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFE97BE87FE0CFA13" box="[380,470,1411,1437]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF2CBEA3FE69FA4F" blockId="2.[151,621,1339,1473]" box="[199,435,1447,1473]" pageId="2" pageNumber="430">
<taxonomicName id="4C614D5BFFFDFFEAFF37BEA3FE89FA4F" authorityName="Volynkin" authorityYear="2024" box="[220,339,1447,1473]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="hoenei" status="sp. nov.">
<emphasis id="B915EACAFFFDFFEAFF37BEA3FE89FA4F" box="[220,339,1447,1473]" italics="true" pageId="2" pageNumber="430">Th. hoenei</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFDFFEAFEB2BEA3FE69FA4F" bold="true" box="[345,435,1447,1473]" pageId="2" pageNumber="430">
<taxonomicNameLabel id="A22657B1FFFDFFEAFEB2BEA3FE69FA4F" box="[345,435,1447,1473]" pageId="2" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36D8FFFDFFEAFF7CBEEBFDEAF9A3" blockId="2.[151,560,1519,1581]" pageId="2" pageNumber="430">
The
<taxonomicName id="4C614D5BFFFDFFEAFF21BEEBFE8DF987" baseAuthorityName="Volynkin" baseAuthorityYear="2021" box="[202,343,1519,1545]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B915EACAFFFDFFEAFF21BEEBFE8DF987" box="[202,343,1519,1545]" italics="true" pageId="2" pageNumber="430">Th. anomala</emphasis>
</taxonomicName>
species-group
<taxonomicName id="4C614D5BFFFDFFEAFF37BD17FDEAF9A3" authority="(Volynkin, 2021)" baseAuthorityName="Volynkin" baseAuthorityYear="2021" box="[220,560,1555,1581]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B915EACAFFFDFFEAFF37BD17FEB0F9A3" box="[220,362,1555,1581]" italics="true" pageId="2" pageNumber="430">Th. anomala</emphasis>
(
<bibRefCitation id="EFF04B29FFFDFFEAFE93BD17FDFDF9A3" author="Volynkin, A. V." box="[376,551,1555,1581]" pageId="2" pageNumber="430" pagination="34 - 38" refId="ref10767" refString="Volynkin, A. V. (2021) Thysanoptyx anomala, a new species from south-western China (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 48, 34 - 38. https: // doi. org / 10.37828 / em. 2021.48.6" type="journal article" year="2021">Volynkin, 2021</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>