266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document id="3DB821B472D7FD95229284AD49816CCA" ID-DOI="10.5281/zenodo.5391440" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630315127817" checkinUser="marcus" docAuthor="Monniot, Françoise & Monniot, Claude" docDate="2001" docId="F57D87A3FF4B31ADEA4FFBB7FDA41080" docLanguage="en" docName="Zoosystema.2001.23.2.201-383.pdf" docOrigin="Zoosystema 23 (2)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Microcosmus pacificus Monniot & Monniot 2001, n. sp." docType="treatment" docVersion="4" lastPageNumber="351" masterDocId="0944FFDBFFDF313BEB47FF93FFBE1602" masterDocTitle="Ascidians from the tropical western Pacific" masterLastPageNumber="383" masterPageNumber="201" pageNumber="349" updateTime="1699223418854" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="7ED2CDBE0D33C32402F31001B2C64D99" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CA7D66768259D18717EA8B79204F9AE9">
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<mods:title id="DA6B49A3BC5FFB7B80F82C1CE182D069">Ascidians from the tropical western Pacific</mods:title>
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</mods:titleInfo>
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<mods:name id="2CC10705B741C7A9460114B2119E444F" type="personal">
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<mods:role id="DF2A0551A741F04E5C5D8D5E99656C39">
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<mods:roleTerm id="FD638FA3050CB163404AC9E32D18D1BE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7F730BF97783E246A8AA187CB13241D7">Monniot, Françoise</mods:namePart>
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</mods:name>
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<mods:name id="C10BBFC20232B61BFC14FB4B93864485" type="personal">
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<mods:role id="30CD73A3F6B1647C37A61EB4F1AA744E">
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<mods:roleTerm id="53647BEFE688EA1EE925573AD8E6DF7A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBC1277A3A58878BB743D1AB86FA372D">Monniot, Claude</mods:namePart>
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<mods:affiliation id="C6DD5AC62252A21DB43368AC567832CA">UPESA 8044, Laboratoire de Biologie des Invertébrés marins et Malacologie, Muséum national d’Histoire naturelle, 55 rue Buffon, F- 75005 Paris (France) monniot @ mnhn. fr.</mods:affiliation>
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<mods:nameIdentifier id="7ECB29A74B5131C65FD8A306EC9C382E" type="email">monniot@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="060A95ED230CF460D02C362851B60BBA">text</mods:typeOfResource>
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<mods:titleInfo id="C392680B1AE7188F234F99E41D480940">
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<mods:title id="73FFEC27EB66F6F9F11F5FA9EBD15001">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="85B020CBDFF8972375222C3398E9D68C">
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<mods:date id="BA75F16BCF54B1298759E3992A3E2F88">2001</mods:date>
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<mods:detail id="DA5177DD381F79817A35D929B5424C6B" type="volume">
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<mods:number id="5D07A72EA73EAD5E4066F8D983A2C1A5">23</mods:number>
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</mods:detail>
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<mods:detail id="00BAE7FF269CEBB19AB87A96E0B3FA94" type="issue">
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<mods:number id="1DD76B126033FBC2A7D23D36EE34CC7E">2</mods:number>
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</mods:detail>
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<mods:start id="AF448285F9AD7F7A97EE1B4DDA4E418C">201</mods:start>
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<mods:end id="0FD5AFFFCBFC8AA93C656811E682282C">383</mods:end>
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</mods:part>
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<mods:classification id="48701D3A57445EDD865DD8820386C306">journal article</mods:classification>
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<mods:identifier id="16F8E41AF27B37334714657746D3AFB3" type="DOI">10.5281/zenodo.5391440</mods:identifier>
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<mods:identifier id="94CC171B35A328CD48ED56C3E4278CA2" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="072BE7EC05A26202793F1BBDCA69F986" type="Zenodo-Dep">5391440</mods:identifier>
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<treatment id="F57D87A3FF4B31ADEA4FFBB7FDA41080" ID-DOI="http://doi.org/10.5281/zenodo.5468118" ID-Zenodo-Dep="5468118" LSID="urn:lsid:plazi:treatment:F57D87A3FF4B31ADEA4FFBB7FDA41080" httpUri="http://treatment.plazi.org/id/F57D87A3FF4B31ADEA4FFBB7FDA41080" lastPageId="150" lastPageNumber="351" pageId="148" pageNumber="349">
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<subSubSection id="35CE653EFF4B31AFEA4FFBB7FD85123C" box="[264,571,1060,1087]" pageId="148" pageNumber="349" type="nomenclature">
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<paragraph id="7D6B36B5FF4B31AFEA4FFBB7FD85123C" blockId="148.[264,571,1060,1118]" box="[264,571,1060,1087]" pageId="148" pageNumber="349">
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<heading id="262381D9FF4B31AFEA4FFBB7FD85123C" box="[264,571,1060,1087]" centered="true" fontSize="11" level="2" pageId="148" pageNumber="349" reason="2">
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<taxonomicName id="BAD44D36FF4B31AFEA4FFBB7FE46123D" ID-CoL="73FRP" authority="Monniot & Monniot, 2001" authorityName="Monniot & Monniot" authorityYear="2001" box="[264,504,1060,1087]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="148" pageNumber="349" phylum="Chordata" rank="species" species="pacificus" status="sp. nov.">
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<emphasis id="4FA0EAA7FF4B31AFEA4FFBB7FE46123D" bold="true" box="[264,504,1060,1087]" italics="true" pageId="148" pageNumber="349">Microcosmus pacificus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="549357DCFF4B31AFEAB8FBB7FD85123C" box="[511,571,1060,1086]" pageId="148" pageNumber="349" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="35CE653EFF4B31AFEA04FBD7FE41125C" box="[323,511,1092,1118]" pageId="148" pageNumber="349" type="description">
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<paragraph id="7D6B36B5FF4B31AFEA04FBD7FE41125C" blockId="148.[264,571,1060,1118]" box="[323,511,1092,1118]" pageId="148" pageNumber="349">
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(
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<figureCitation id="E5EF2A30FF4B31AFEA0DFBD7FE7F125C" box="[330,449,1092,1118]" captionStart="FIG" captionStartId="146.[151,162,1628,1645]" captionTargetBox="[151,1247,215,1579]" captionTargetId="graphics-80@146.[475,1236,860,1566]" captionTargetPageId="146" captionText="FIG. 108. — Siphonal spinules; A, Pyura gangelion (Savigny, 1816); B, Pyura styeliformis n. sp.; C, Microcosmus bitunicatus n. sp.; D, Microcosmus pacificus n. sp. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5391655" httpUri="https://zenodo.org/record/5391655/files/figure.png" pageId="148" pageNumber="349">Figs 108D</figureCitation>
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;
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<figureCitation id="E5EF2A30FF4B31AFEA8AFBD7FE46125C" box="[461,504,1092,1118]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">109</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="35CE653EFF4B31AFEBD0FBEFFD8312CA" pageId="148" pageNumber="349" type="materials_examined">
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||
<paragraph id="7D6B36B5FF4B31AFEBD0FBEFFD8312CA" blockId="148.[151,684,1148,1355]" pageId="148" pageNumber="349">
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||
<collectionCode id="1BC5AE70FF4B31AFEBD0FBEFFE8D1290" box="[151,307,1148,1171]" pageId="148" pageNumber="349">
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<typeStatus id="A26F8817FF4B31AFEBD0FBEFFF771293" box="[151,201,1148,1171]" pageId="148" pageNumber="349">TYPE</typeStatus>
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MATERIAL
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||
</collectionCode>
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. —
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<emphasis id="4FA0EAA7FF4B31AFEA23FBEFFD861297" bold="true" box="[356,568,1148,1173]" pageId="148" pageNumber="349">
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||
<collectingCountry id="05C37625FF4B31AFEA23FBEFFD8A1297" box="[356,564,1148,1173]" name="Papua New Guinea" pageId="148" pageNumber="349">Papua New Guinea</collectingCountry>
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.
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</emphasis>
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<collectingRegion id="BF10F857FF4B31AFE906FBEEFF4C12AC" country="Papua New Guinea" name="Milne Bay" pageId="148" pageNumber="349">Milne Bay Province</collectingRegion>
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, Samarai Island,
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<geoCoordinate id="18E05072FF4B31AFEAE3FB05FDA912AC" box="[420,535,1174,1198]" degrees="10" direction="south" minutes="36.98" orientation="latitude" pageId="148" pageNumber="349" precision="9" value="-10.616333">10°36.98’S</geoCoordinate>
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||
,
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||
<geoCoordinate id="18E05072FF4B31AFE962FB05FD1612AC" box="[549,680,1174,1198]" degrees="150" direction="east" minutes="39.77" orientation="longitude" pageId="148" pageNumber="349" precision="9" value="150.66283">150°39.77’E</geoCoordinate>
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,
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<quantity id="BA2C9B50FF4B31AFEBD0FB23FF7312C4" box="[151,205,1199,1224]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="148" pageNumber="349" unit="m" value="27.0">27 m</quantity>
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,
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<date id="096A1075FF4B31AFEB90FB23FEF712CA" box="[215,329,1200,1224]" pageId="148" pageNumber="349" value="1998-06-10">
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10.
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<collectionCode id="1BC5AE70FF4B31AFEBB3FB23FEAF12CA" box="[244,273,1200,1224]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="148" pageNumber="349">VI</collectionCode>
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.1998
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</date>
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(
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<collectionCode id="1BC5AE70FF4B31AFEA10FB23FE1512CA" box="[343,427,1200,1224]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="148" pageNumber="349">MNHN</collectionCode>
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S2
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<collectionCode id="1BC5AE70FF4B31AFEA96FB23FDBC12CA" box="[465,514,1200,1224]" country="India" httpUri="http://grbio.org/cool/is9g-wzwv" name="Mar Ivanios College (Zoology museum)" pageId="148" pageNumber="349">MIC</collectionCode>
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159).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="35CE653EFF4B31AFEBD0FB44FE4F1349" pageId="148" pageNumber="349" type="etymology">
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<paragraph id="7D6B36B5FF4B31AFEBD0FB44FDCF12F2" blockId="148.[151,684,1148,1355]" box="[151,625,1239,1264]" pageId="148" pageNumber="349">ETYMOLOGY. — Named after the Pacific Ocean.</paragraph>
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<paragraph id="7D6B36B5FF4B31AFEBD0FA92FE4F1349" blockId="148.[151,684,1148,1355]" pageId="148" pageNumber="349">
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OTHER MATERIAL EXAMINED. —
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<emphasis id="4FA0EAA7FF4B31AFE959FA92FD12131B" bold="true" box="[542,684,1281,1305]" pageId="148" pageNumber="349">
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||
<collectingCountry id="05C37625FF4B31AFE959FA92FD19131B" box="[542,679,1281,1305]" name="Philippines" pageId="148" pageNumber="349">Philippines</collectingCountry>
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.
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</emphasis>
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Musorstom 3,
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<geoCoordinate id="18E05072FF4B31AFEA7AFA89FE211330" box="[317,415,1306,1330]" degrees="11" direction="north" minutes="37" orientation="latitude" pageId="148" pageNumber="349" precision="925" value="11.616667">11°37’N</geoCoordinate>
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,
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||
<geoCoordinate id="18E05072FF4B31AFEAE9FA89FDA41330" box="[430,538,1306,1330]" degrees="121" direction="east" minutes="43" orientation="longitude" pageId="148" pageNumber="349" precision="925" value="121.71667">121°43’E</geoCoordinate>
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,
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||
<quantity id="BA2C9B50FF4B31AFE96FFA89FD161332" box="[552,680,1305,1330]" metricMagnitude="2" metricUnit="m" metricValue="1.21" metricValueMax="1.22" metricValueMin="1.2" pageId="148" pageNumber="349" unit="m" value="121.0" valueMax="122.0" valueMin="120.0">120-122 m</quantity>
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||
,
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<date id="096A1075FF4B31AFEBD0FAA0FF431349" box="[151,253,1331,1355]" pageId="148" pageNumber="349" value="1985-06-05">5.VI.1985</date>
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(MNHN S2 MIC 160).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="35CE653EFF4B31AFEBD0FA14FB831080" pageId="148" pageNumber="349" type="description">
|
||
<paragraph id="7D6B36B5FF4B31AFEBD0FA14FE96139D" blockId="148.[151,684,1415,1666]" box="[151,296,1415,1441]" pageId="148" pageNumber="349">DESCRIPTION</paragraph>
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<paragraph id="7D6B36B5FF4B31AFEBD0FA3BFC9A1720" blockId="148.[151,684,1415,1666]" lastBlockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">
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Both specimens were collected on a sandy bottom. They are very similar,
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<quantity id="BA2C9B50FF4B31AFEA8CFA5BFDBC13E3" box="[459,514,1479,1506]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="148" pageNumber="349" unit="cm" value="3.0">3 cm</quantity>
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in length, covered with coarse sand and broken shells. The siphons are well apart. The body wall is thin and pale in formalin except for the red extremity of the siphons. Internally the siphons are long and narrow. The internal spinules are arranged in chevrons and their section is angular (
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<figureCitation id="E5EF2A30FF4B31AFEF19FF3BFB6D16C0" box="[1118,1235,168,194]" captionStart="FIG" captionStartId="146.[151,162,1628,1645]" captionTargetBox="[151,1247,215,1579]" captionTargetId="graphics-80@146.[475,1236,860,1566]" captionTargetPageId="146" captionText="FIG. 108. — Siphonal spinules; A, Pyura gangelion (Savigny, 1816); B, Pyura styeliformis n. sp.; C, Microcosmus bitunicatus n. sp.; D, Microcosmus pacificus n. sp. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5391655" httpUri="https://zenodo.org/record/5391655/files/figure.png" pageId="148" pageNumber="349">Fig. 108D</figureCitation>
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). The musculature forms a network of crossed and regularly spaced bundles issuing from both siphons.
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</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFEBBFB9A1460" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">
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The tentacles are inserted at the base of a high velum. There are 12 large, bushy tentacles in two orders of size with three orders of branchings (
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<figureCitation id="E5EF2A30FF4B31AFE995FE1BFCF617A0" box="[722,840,392,418]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109D</figureCitation>
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); smaller ones are irregularly intercalated. The prepharyngeal band is very close to the branchial tissue. It is deeply indented dorsally. The dorsal tubercle is protruding; its opening is horse-shoe shaped, anteriorly opened with horns inwardly rolled (
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<figureCitation id="E5EF2A30FF4B31AFE83BFDBBFC4F1440" box="[892,1009,552,578]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109D</figureCitation>
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). The neural ganglion is particularly long (
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<figureCitation id="E5EF2A30FF4B31AFE8E5FDDBFBA91460" box="[930,1047,584,610]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109D</figureCitation>
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).
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</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFDFBFC0114A0" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">The dorsal lamina is a high blade of equal height along its whole length.</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFD3BFB9714E0" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">The branchial sac counts seven folds on each side. The formula on the right side is:</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFD74FB6F1503" blockId="148.[714,1248,168,1666]" box="[714,1233,743,770]" pageId="148" pageNumber="349">
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R.E.?
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<emphasis id="4FA0EAA7FF4B31AFE856FD7BFC901500" bold="true" box="[785,814,744,770]" pageId="148" pageNumber="349">10</emphasis>
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2
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<emphasis id="4FA0EAA7FF4B31AFE80DFD7BFCD91500" bold="true" box="[842,871,744,770]" pageId="148" pageNumber="349">15</emphasis>
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||
2
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<emphasis id="4FA0EAA7FF4B31AFE8C4FD7BFC1E1500" bold="true" box="[899,928,744,770]" pageId="148" pageNumber="349">17</emphasis>
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||
1
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||
<emphasis id="4FA0EAA7FF4B31AFE8FCFD7BFC661500" bold="true" box="[955,984,744,770]" pageId="148" pageNumber="349">19</emphasis>
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1
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||
<emphasis id="4FA0EAA7FF4B31AFE8B3FD7BFBAF1500" bold="true" box="[1012,1041,744,770]" pageId="148" pageNumber="349">19</emphasis>
|
||
1
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||
<emphasis id="4FA0EAA7FF4B31AFEF6BFD7BFBF71500" bold="true" box="[1068,1097,744,770]" pageId="148" pageNumber="349">19</emphasis>
|
||
1
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||
<emphasis id="4FA0EAA7FF4B31AFEF22FD7BFB3C1500" bold="true" box="[1125,1154,744,770]" pageId="148" pageNumber="349">17</emphasis>
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2 D.L.
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</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFC9BFC6A15A0" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">The folds are high, overlapping each other. There are five to six stigmata per mesh between the folds and three to four on the folds. At the top of the folds one or two stigmata are curved. There are parastigmatic vessels.</paragraph>
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<paragraph id="7D6B36B5FF4B31AFE98DFC3BFC5A1260" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">
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The gut makes a very long and curved loop with both limbs very close together (
|
||
<figureCitation id="E5EF2A30FF4B31AFEF71FC5BFB6F15E0" box="[1078,1233,968,994]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109A, C</figureCitation>
|
||
). The stomach is only slightly enlarged. It is covered by a massive hepatic gland constituted of lamellae. Many filiform papillae arise from the hepatic lobes (
|
||
<figureCitation id="E5EF2A30FF4B31AFE822FBDBFC691260" box="[869,983,1096,1122]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7D6B36B5FF4B31AFE98DFBFBFCD613C0" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">
|
||
The rectum is cylindrical, its posterior part attached to the oesophagus. The anus is wide open with two lobes. There is one gonad on each side, protruding well above the body wall (
|
||
<figureCitation id="E5EF2A30FF4B31AFE995FB7BFC931300" box="[722,813,1256,1282]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109</figureCitation>
|
||
A-C). The left gonad lies in the secondary loop of the gut but does not overlap the rectum (
|
||
<figureCitation id="E5EF2A30FF4B31AFE84BFABBFC321340" box="[780,908,1320,1346]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109C</figureCitation>
|
||
). The general outline of the gonads is oval. The male lobules surround the sinuous ovaries and penetrate them in furrows (
|
||
<figureCitation id="E5EF2A30FF4B31AFE995FA1BFC8113A0" box="[722,831,1416,1442]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109E</figureCitation>
|
||
) The gonoducts are joined, opening at the same level.
|
||
</paragraph>
|
||
<paragraph id="7D6B36B5FF4B31AFE98DFA5BFCEF1040" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">
|
||
The endocarps are irregular in size, and lobulated. They lie inside the gut loop, on both sides of the body wall ventrally, and above the gonads (
|
||
<figureCitation id="E5EF2A30FF4B31AFE995F9BBFCFB1040" box="[722,837,1576,1602]" captionStart="FIG" captionStartId="149.[151,162,1442,1459]" captionTargetBox="[151,1244,166,1407]" captionTargetId="figure-73@149.[148,1249,169,1415]" captionTargetPageId="149" captionText="FIG. 109. — Microcosmus pacificus n. sp.; A, B, left and right sides of the body; C, body ventrally opened; D, dorsal tubercle; E, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="148" pageNumber="349">Fig. 109C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7D6B36B5FF4B31AFE98DF9DBFB831080" blockId="148.[714,1248,168,1666]" pageId="148" pageNumber="349">There is a large cloacal velum with an irregular ring of filiform papillae at its base.</paragraph>
|
||
</subSubSection>
|
||
<caption id="29AB663DFF4A31AEEBD0FA31FDD813CA" ID-DOI="http://doi.org/10.5281/zenodo.5391657" ID-Zenodo-Dep="5391657" httpUri="https://zenodo.org/record/5391657/files/figure.png" pageId="149" pageNumber="350" startId="149.[151,162,1442,1459]" targetBox="[151,1244,166,1407]" targetPageId="149">
|
||
<paragraph id="7D6B36B5FF4A31AEEBD0FA31FDD813CA" blockId="149.[151,1248,1442,1480]" pageId="149" pageNumber="350">
|
||
FIG. 109. —
|
||
<taxonomicName id="BAD44D36FF4A31AEEA45FA31FE7613B1" authority="Monniot & Monniot, 2001" authorityName="Monniot & Monniot" authorityYear="2001" box="[258,456,1442,1459]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="149" pageNumber="350" phylum="Chordata" rank="species" species="pacificus" status="sp. nov.">
|
||
<emphasis id="4FA0EAA7FF4A31AEEA45FA31FE7613B1" box="[258,456,1442,1459]" italics="true" pageId="149" pageNumber="350">Microcosmus pacificus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="549357DCFF4A31AEEA89FA31FDBD13B1" box="[462,515,1442,1459]" pageId="149" pageNumber="350" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="4FA0EAA7FF4A31AEE94CFA31FDA613B1" bold="true" box="[523,536,1442,1459]" pageId="149" pageNumber="350">A</emphasis>
|
||
,
|
||
<emphasis id="4FA0EAA7FF4A31AEE962FA31FD8C13B1" bold="true" box="[549,562,1442,1459]" pageId="149" pageNumber="350">B</emphasis>
|
||
, left and right sides of the body;
|
||
<emphasis id="4FA0EAA7FF4A31AEE810FA31FCDB13B1" bold="true" box="[855,869,1442,1459]" pageId="149" pageNumber="350">C</emphasis>
|
||
, body ventrally opened;
|
||
<emphasis id="4FA0EAA7FF4A31AEEF79FA31FBF213B1" bold="true" box="[1086,1100,1442,1459]" pageId="149" pageNumber="350">D</emphasis>
|
||
, dorsal tubercle;
|
||
<emphasis id="4FA0EAA7FF4A31AEEBD0FA24FF1D13CA" bold="true" box="[151,163,1463,1480]" pageId="149" pageNumber="350">E</emphasis>
|
||
, gonad. Scale bars: A, B, 1 cm; C, 5 mm; D, E, 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="35CE653EFF4A31AEEBD0F994FB081080" pageId="149" pageNumber="350" type="discussion">
|
||
<paragraph id="7D6B36B5FF4A31AEEBD0F994FF45101D" blockId="149.[151,684,1543,1666]" box="[151,251,1543,1569]" pageId="149" pageNumber="350">REMARKS</paragraph>
|
||
<paragraph id="7D6B36B5FF4A31AEEBD0F9B4FB081080" blockId="149.[151,684,1543,1666]" lastBlockId="149.[714,1247,1544,1666]" pageId="149" pageNumber="350">
|
||
By its sediment-covered tunic, the length of its well-separated siphons, its branchial sac, and its long narrow gut loop, this species resembles
|
||
<taxonomicName id="BAD44D36FF4A31AEE98DF99BFB611020" authority="Michaelsen, 1918" authorityName="Michaelsen" authorityYear="1918" box="[714,1247,1544,1570]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="149" pageNumber="350" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="4FA0EAA7FF4A31AEE98DF99BFBAE1020" box="[714,1040,1544,1570]" italics="true" pageId="149" pageNumber="350">Microcosmus madagascariensis</emphasis>
|
||
<bibRefCitation id="19454B44FF4A31AEEF5FF99BFB611020" author="MICHAELSEN W." box="[1048,1247,1544,1570]" pageId="149" pageNumber="350" pagination="1 - 68" refId="ref86283" refString="MICHAELSEN W. 1918 - Die Ptychobranchien und Diktyobranchien Ascidien des Westlichen Indischen Ozeans. Mittelungen aus dem Zoologishen Museum Hamburg 35: 1 - 68." type="journal article" year="1918">Michaelsen, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="05C37625FF4A31AEE842F9BBFC361040" box="[773,904,1576,1602]" name="Madagascar" pageId="149" pageNumber="350">Madagascar</collectingCountry>
|
||
. But the latter has the left gonad overlapping the gut loop. As well, the endocarps are differently distributed in these two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="29AB663DFF4931ADEBEAFC78FB7415FE" ID-DOI="http://doi.org/10.5281/zenodo.5391659" ID-Zenodo-Dep="5391659" box="[173,1226,1003,1020]" httpUri="https://zenodo.org/record/5391659/files/figure.png" pageId="150" pageNumber="351" startId="150.[173,184,1003,1020]" targetBox="[151,1250,163,970]" targetPageId="150">
|
||
<paragraph id="7D6B36B5FF4931ADEBEAFC78FB7415FE" blockId="150.[173,1226,1003,1020]" box="[173,1226,1003,1020]" pageId="150" pageNumber="351">
|
||
FIG. 110. —
|
||
<taxonomicName id="BAD44D36FF4931ADEA55FC78FE5915FE" authority="Monniot & Monniot, 2001" authorityName="Monniot & Monniot" authorityYear="2001" box="[274,487,1003,1020]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="vesiculosus" status="sp. nov.">
|
||
<emphasis id="4FA0EAA7FF4931ADEA55FC78FE5915FE" box="[274,487,1003,1020]" italics="true" pageId="150" pageNumber="351">Microcosmus vesiculosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="549357DCFF4931ADEAABFC78FDA015FE" box="[492,542,1003,1020]" pageId="150" pageNumber="351" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="4FA0EAA7FF4931ADE962FC78FD8C15FE" bold="true" box="[549,562,1003,1020]" pageId="150" pageNumber="351">A</emphasis>
|
||
,
|
||
<emphasis id="4FA0EAA7FF4931ADE97BFC78FDF715FE" bold="true" box="[572,585,1003,1020]" pageId="150" pageNumber="351">B</emphasis>
|
||
, left and right sides of the body;
|
||
<emphasis id="4FA0EAA7FF4931ADE81BFC78FCD415FE" bold="true" box="[860,874,1003,1020]" pageId="150" pageNumber="351">C</emphasis>
|
||
, body ventrally opened. Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="35CE653EFF4931ADEBD0FBDBFDA41080" pageId="150" pageNumber="351" type="reference_group">
|
||
<paragraph id="7D6B36B5FF4931ADEBD0FBDBFE081300" blockId="150.[151,684,1095,1666]" pageId="150" pageNumber="351">
|
||
<taxonomicName id="BAD44D36FF4931ADEBD0FBDBFDD21260" ID-CoL="6RG3W" authority="Hartmeyer, 1919" authorityName="Hartmeyer" authorityYear="1919" box="[151,620,1095,1122]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="agglutinans">
|
||
<emphasis id="4FA0EAA7FF4931ADEBD0FBDBFE1F1260" box="[151,417,1096,1122]" italics="true" pageId="150" pageNumber="351">Microcosmus agglutinans</emphasis>
|
||
<bibRefCitation id="19454B44FF4931ADEAECFBD4FDD21260" author="HARTMEYER R." box="[427,620,1095,1122]" pageId="150" pageNumber="351" pagination="1 - 150" refId="ref85067" refString="HARTMEYER R. 1919. - Ascidien, in Results of Dr E. Mjobergs Swedish scientific expedition in Australia 1910 - 13. Kunglinga Svenska vetenskapsakademiens Handlingar 60 (4): 1 - 150." type="journal article" year="1919">Hartmeyer, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="BF10F857FF4931ADEBD0FBF4FEF11280" box="[151,335,1127,1154]" country="Australia" name="Western Australia" pageId="150" pageNumber="351">western Australia</collectingRegion>
|
||
is also closely allied. It differs in having its the siphons close together, and it has a more accentuated secondary loop of the gut with the anus level with the top of the primary gut loop. It has no oral velum.
|
||
</paragraph>
|
||
<paragraph id="7D6B36B5FF4931ADEBD0FA94FDA41080" blockId="150.[151,684,1095,1666]" pageId="150" pageNumber="351">
|
||
<bibRefCitation id="19454B44FF4931ADEBD0FA94FDAE1320" author="HARTMEYER R. & MICHAELSEN W." box="[151,528,1287,1314]" pageId="150" pageNumber="351" pagination="251 - 460" refId="ref85104" refString="HARTMEYER R. & MICHAELSEN W. 1928. - Ascidiae Dictyobranchiae und Ptychobranchiae. Fauna Sudwest-Australiens 5 (6): 251 - 460." type="journal article" year="1928">Hartmeyer & Michaelsen (1928)</bibRefCitation>
|
||
synonymised
|
||
<taxonomicName id="BAD44D36FF4931ADEBD0FABBFED21340" ID-CoL="73FRR" authorityName="Michaelsen" authorityYear="1918" box="[151,364,1320,1346]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="4FA0EAA7FF4931ADEBD0FABBFED21340" box="[151,364,1320,1346]" italics="true" pageId="150" pageNumber="351">M. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="BAD44D36FF4931ADEAE3FABBFDFF1340" ID-CoL="6RG3W" authorityName="Hartmeyer" authorityYear="1919" box="[420,577,1320,1346]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="agglutinans">
|
||
<emphasis id="4FA0EAA7FF4931ADEAE3FABBFDFF1340" box="[420,577,1320,1346]" italics="true" pageId="150" pageNumber="351">M. agglutinans</emphasis>
|
||
</taxonomicName>
|
||
, an opinion that
|
||
<bibRefCitation id="19454B44FF4931ADEBBDFAD4FE3F1360" author="KOTT P." box="[250,385,1351,1378]" pageId="150" pageNumber="351" pagination="1 - 440" refId="ref85894" refString="KOTT P. 1985. - The Australian Ascidiacea. Part 1: Phlebobranchiata and Stolidobranchiata. Memoirs of the Queensland Museum 23: 1 - 440." type="journal article" year="1985">Kott (1985)</bibRefCitation>
|
||
supported, but we believe that these are different species. Kott’s description corresponds better to
|
||
<taxonomicName id="BAD44D36FF4931ADEAC9FA1BFD8E13A0" ID-CoL="6RG3W" authorityName="Hartmeyer" authorityYear="1919" box="[398,560,1416,1442]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="agglutinans">
|
||
<emphasis id="4FA0EAA7FF4931ADEAC9FA1BFD8E13A0" box="[398,560,1416,1442]" italics="true" pageId="150" pageNumber="351">M. agglutinans</emphasis>
|
||
</taxonomicName>
|
||
. Moreover the description given by
|
||
<bibRefCitation id="19454B44FF4931ADEA8FFA34FD3E13C0" author="VASSEUR P." box="[456,640,1447,1474]" pageId="150" pageNumber="351" pagination="912 - 923" refId="ref89327" refString="VASSEUR P. 1969. - Deuxieme contribution l'etude des ascidies de Madagascar region de Tulear. Bulletin du Museum national d'Histoire naturelle (2), 40 (5): 912 - 923." type="journal article" year="1969">Vasseur (1969)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="BAD44D36FF4931ADEBD0FA5BFED213E0" ID-CoL="73FRR" authorityName="Michaelsen" authorityYear="1918" box="[151,364,1480,1506]" class="Ascidiacea" family="Pyuridae" genus="Microcosmus" kingdom="Animalia" order="Stolidobranchia" pageId="150" pageNumber="351" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="4FA0EAA7FF4931ADEBD0FA5BFED213E0" box="[151,364,1480,1506]" italics="true" pageId="150" pageNumber="351">M. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
collected in
|
||
<collectingCountry id="05C37625FF4931ADEABDFA5BFDC713E0" box="[506,633,1480,1506]" name="Madagascar" pageId="150" pageNumber="351">Madagascar</collectingCountry>
|
||
corresponds well to the description of Michaelsen’s
|
||
<typeStatus id="A26F8817FF4931ADEBD0F994FF7B1023" box="[151,197,1543,1569]" pageId="150" pageNumber="351">type</typeStatus>
|
||
from the same locality. The geographic distance separating them – one in
|
||
<collectingCountry id="05C37625FF4931ADEABEF9BBFDC21040" box="[505,636,1576,1602]" name="Madagascar" pageId="150" pageNumber="351">Madagascar</collectingCountry>
|
||
, the other in western Australia – is an additional argument for separating the two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |