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<mods:title id="1E682A65ABC81F1B66457C3FB82FD09C">Review of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) from Papua New Guinea, with Descriptions of Five New Species</mods:title>
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<mods:title id="E865FBFBC76389C825C74C8E657A315E">The Coleopterists Bulletin</mods:title>
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<paragraph id="8BE136A9FF88FF924BAAFC917D9AFC30" blockId="8.[713,1144,856,903]" pageId="8" pageNumber="686">
<emphasis id="B92AEABBFF88FF924BAAFC917A3EFCDA" bold="true" box="[713,1144,856,877]" pageId="8" pageNumber="686">
3.
<taxonomicName id="4C5E4D2AFF88FF924B80FC917A3EFCDA" ID-CoL="8MN59" authority="Gueorguiev, 2013" authorityName="Gueorguiev" authorityYear="2013" box="[739,1144,856,877]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="baehri">
<emphasis id="B92AEABBFF88FF924B80FC917DF8FCDA" bold="true" box="[739,958,856,877]" italics="true" pageId="8" pageNumber="686">Mecyclothorax baehri</emphasis>
<bibRefCitation id="EFCF4B58FF88FF924AA6FC917A3EFCDA" author="Gueorguiev, B." box="[965,1144,856,877]" pageId="8" pageNumber="686" pagination="52 - 70" refId="ref18062" refString="Gueorguiev, B. 2013. Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae). Zootaxa 3709: 52 - 70." type="journal article" year="2013">Guéorguiev, 2013</bibRefCitation>
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(
<figureCitation id="13652A2CFF88FF924A0FFCBB7DF7FC30" box="[876,945,882,903]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="8" pageNumber="686">Figs. 2</figureCitation>
,
<figureCitation id="13652A2CFF88FF924ADEFCBB7D90FC30" box="[957,982,882,903]" captionStart="Fig" captionStartId="17.[197,228,896,915]" captionTargetBox="[175,1191,199,878]" captionTargetId="figure-399@17.[173,1195,194,881]" captionTargetPageId="17" captionText="Fig. 32. Localities in Papua New Guinea from which Mecyclothorax spp. were collected and the particular species collected at each locality. 1 = Mt. Capella — M. baehri; 2 = Mt. Fugilil — M. brispex and M. medioconstrictus; 3 = Kubor Range — M. kubor; 4 = Mt. Wilhelm — M. sedlaceki; 5 = Finisterre Range — M. kavanaughi; 6 = Mt. Missim — M. gressitti; 7 = Mt. Kaindi, Edie Creek and Bulldog Road — M. andersoni, M. cuccodoroi, and M. loebli; 8 = Mt. Amungwiwa—M. amingwiwae." figureDoi="http://doi.org/10.5281/zenodo.5377933" httpUri="https://zenodo.org/record/5377933/files/figure.png" pageId="8" pageNumber="686">32</figureCitation>
)
</paragraph>
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<subSubSection id="C3446522FF88FF924BCEFC547D77FC7A" pageId="8" pageNumber="686" type="reference_group">
<paragraph id="8BE136A9FF88FF924BCEFC547D77FC7A" blockId="8.[685,1171,925,973]" pageId="8" pageNumber="686">
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<treatmentCitation id="0AFF10B8FF88FF924BCEFC547A17FC05" author="Gueorguiev, B." box="[685,1105,925,946]" page="53" pageId="8" pageNumber="686" year="2013">
<taxonomicName id="4C5E4D2AFF88FF924BCEFC547A17FC05" ID-CoL="8MN59" authority="Gueorguiev 2013: 53" authorityName="Gueorguiev" authorityPageNumber="53" authorityYear="2013" box="[685,1105,925,946]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="baehri">
<emphasis id="B92AEABBFF88FF924BCEFC547D39FC05" box="[685,895,925,946]" italics="true" pageId="8" pageNumber="686">Mecyclothorax baehri</emphasis>
<bibRefCitation id="EFCF4B58FF88FF924AE6FC547A17FC05" author="Gueorguiev, B." box="[901,1105,925,946]" pageId="8" pageNumber="686" pagination="52 - 70" refId="ref18062" refString="Gueorguiev, B. 2013. Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae). Zootaxa 3709: 52 - 70." type="journal article" year="2013">Guéorguiev 2013: 53</bibRefCitation>
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;
<treatmentCitation id="0AFF10B8FF88FF924D38FC547D6BFC7A" author="Baehr, M." page="126" pageId="8" pageNumber="686" year="2014">
<bibRefCitation id="EFCF4B58FF88FF924D38FC547D6BFC7A" author="Baehr, M." pageId="8" pageNumber="686" pagination="123 - 129" refId="ref17601" refString="Baehr, M. 2014. A new species of the genus Mecyclothorax Sharp from New Guinea (Coleoptera, Carabidae, Psydrini, Mecyclothoracina). Spixiana 37: 123 - 129." type="journal article" year="2014">Baehr 2014: 126</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C3446522FF88FF934BA6FC167F22FF46" lastPageId="9" lastPageNumber="687" pageId="8" pageNumber="686" type="diagnosis">
<paragraph id="8BE136A9FF88FF934BA6FC167F22FF46" blockId="8.[685,1173,991,1730]" lastBlockId="9.[173,660,194,401]" lastPageId="9" lastPageNumber="687" pageId="8" pageNumber="686">
<emphasis id="B92AEABBFF88FF924BA6FC167D6BFC43" bold="true" box="[709,813,991,1012]" pageId="8" pageNumber="686">Diagnosis.</emphasis>
Among
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species, only this and
<taxonomicName id="4C5E4D2AFF88FF924A54FC337DBAFBB8" authorityName="Liebherr" authorityYear="2017" box="[823,1020,1018,1039]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="medioconstrictus">
<emphasis id="B92AEABBFF88FF924A54FC337DBAFBB8" box="[823,1020,1018,1039]" italics="true" pageId="8" pageNumber="686">M. medioconstrictus</emphasis>
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can be characterized by the presence of a quadrisetose pronotum, three dorsal elytral setae, and both subapical and apical elytral setae. The subapical seta is positioned within the seventh elytral stria apicad the apical juncture of striae 3 and 4, and the apical seta occurs just laterad the apex of stria 2 as it curves toward the sutural stria. This species can be diagnosed by the distinctly punctate pronotal median base, with about 20 round punctures on each side surrounded with distinct microsculpture consisting of isodiametric and slightly transversely stretched sculpticells. Conversely, the pronotal median base in
<taxonomicName id="4C5E4D2AFF88FF924D5FFAF07D67FADF" authorityName="Liebherr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="medioconstrictus">
<emphasis id="B92AEABBFF88FF924D5FFAF07D67FADF" italics="true" pageId="8" pageNumber="686">M. medioconstrictus</emphasis>
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is less distinctly punctate, with only about 14 smaller, isolated punctures each side that are surrounded by shallowly margined, transverse sculpticells. The anterior transverse pronotal impression is also very deep and well defined medially in this species (
<figureCitation id="13652A2CFF88FF924A2CFA117DC8FA5A" box="[847,910,1496,1517]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="8" pageNumber="686">Fig. 2</figureCitation>
), whereas it is much less developed in
<taxonomicName id="4C5E4D2AFF88FF924A4FFA3A7DABF9BF" authorityName="Liebherr" authorityYear="2017" box="[812,1005,1523,1544]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="medioconstrictus">
<emphasis id="B92AEABBFF88FF924A4FFA3A7DABF9BF" box="[812,1005,1523,1544]" italics="true" pageId="8" pageNumber="686">M. medioconstrictus</emphasis>
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(
<figureCitation id="13652A2CFF88FF924A99FA3A7A77F9BF" box="[1018,1073,1523,1544]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="8" pageNumber="686">Fig. 3</figureCitation>
) where its position is defined by the raised anterior callosity, not by any depression of the disc posterad the callosity. The eyes of
<taxonomicName id="4C5E4D2AFF88FF924AECF98B7DB4F9E0" box="[911,1010,1602,1623]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="baehri">
<emphasis id="B92AEABBFF88FF924AECF98B7DB4F9E0" box="[911,1010,1602,1623]" italics="true" pageId="8" pageNumber="686">M. baehri</emphasis>
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are much more protruding, with the ocular lobe behind the eye nearly perpendicular to the gena, whereas the posterior surface of the ocular lobe in
<taxonomicName id="4C5E4D2AFF88FF924D5AF95B7D67F975" authorityName="Liebherr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="686" phylum="Arthropoda" rank="species" species="medioconstrictus">
<emphasis id="B92AEABBFF88FF924D5AF95B7D67F975" italics="true" pageId="8" pageNumber="686">M. medioconstrictus</emphasis>
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is broader behind the eye and meets the gena at a distinctly obtuse angle. Standardized body length is
<quantity id="4CA69B4CFF89FF934864FF157F26FF46" box="[263,352,220,241]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="9" pageNumber="687" unit="mm" value="5.05">5.05 mm</quantity>
.
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<subSubSection id="C3446522FF89FF9349A6FF3E7F51FE26" pageId="9" pageNumber="687" type="distribution">
<paragraph id="8BE136A9FF89FF9349A6FF3E7F51FE26" blockId="9.[173,660,194,401]" pageId="9" pageNumber="687">
<emphasis id="B92AEABBFF89FF9349A6FF3E7F81FEBB" bold="true" box="[197,455,247,268]" pageId="9" pageNumber="687">Distribution and Habitat.</emphasis>
The elevation of Mt. Capella, the
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locality, is consistent with that reported in
<bibRefCitation id="EFCF4B58FF89FF934841FEE57F8BFEF6" author="Wikipedia" box="[290,461,300,321]" pageId="9" pageNumber="687" refId="ref19640" refString="Wikipedia. 2017. List of highest mountains of New Guinea. en. wikipedia. org / wiki / List _ of _ highest _ mountains _ of _ New _ Guinea (accessed 21 May 2017)." type="book" year="2017">Wikipedia (2017)</bibRefCitation>
with the associated geographic coordinates used in
<figureCitation id="13652A2CFF89FF9348B5FE8F7C5AFEEC" box="[470,540,326,347]" captionStart="Fig" captionStartId="17.[197,228,896,915]" captionTargetBox="[175,1191,199,878]" captionTargetId="figure-399@17.[173,1195,194,881]" captionTargetPageId="17" captionText="Fig. 32. Localities in Papua New Guinea from which Mecyclothorax spp. were collected and the particular species collected at each locality. 1 = Mt. Capella — M. baehri; 2 = Mt. Fugilil — M. brispex and M. medioconstrictus; 3 = Kubor Range — M. kubor; 4 = Mt. Wilhelm — M. sedlaceki; 5 = Finisterre Range — M. kavanaughi; 6 = Mt. Missim — M. gressitti; 7 = Mt. Kaindi, Edie Creek and Bulldog Road — M. andersoni, M. cuccodoroi, and M. loebli; 8 = Mt. Amungwiwa—M. amingwiwae." figureDoi="http://doi.org/10.5281/zenodo.5377933" httpUri="https://zenodo.org/record/5377933/files/figure.png" pageId="9" pageNumber="687">Fig. 32</figureCitation>
. Based upon
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, the mountain at this elevation is forested.
</paragraph>
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