treatments-xml/data/03/E5/B5/03E5B501A008FF94FED4FDD0FCCAFA79.xml

<document id="62190A30D1613670875BC374593F0BE3" ID-DOI="10.5281/zenodo.157371" ID-GBIF-Dataset="2cfbd9b1-3637-4729-b74f-768ead0726e1" ID-ISSN="1175­5326" ID-Zenodo-Dep="157371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461015336210" checkinUser="plazi" docAuthor="Santos, Hugo Ricardo Secioso, Gomes, Ulisses Leite &amp; Charvet-Almeida, Patrícia" docDate="2004" docId="03E5B501A008FF94FED4FDD0FCCAFA79" docLanguage="en" docName="zt00492.pdf" docOrigin="Zootaxa 492" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Dasyatis colarensis Santos, Gomes &amp; Charvet-Almeida, 2004, n. sp." docType="treatment" docUuid="C633B59E-A366-45DC-A02C-3DF56D2AD36E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="FFDCCD79A00BFF9DFFDCFF9EFFE5FFC7" masterDocTitle="A new species of whiptail stingray of the genus Dasyatis Rafinesque, 1810 from the Southwestern Atlantic Ocean (Chondrichthyes: Myliobatiformes: Dasyatidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="4" updateTime="1698568624532" updateUser="plazi">
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<mods:title id="83C73E7887D361ADAAED32F277A4E508">A new species of whiptail stingray of the genus Dasyatis Rafinesque, 1810 from the Southwestern Atlantic Ocean (Chondrichthyes: Myliobatiformes: Dasyatidae)</mods:title>
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<mods:namePart id="4D8A2C3D30707FE785B0336735D607C1">Santos, Hugo Ricardo Secioso</mods:namePart>
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<mods:namePart id="A132768C31C247D61DDF1D1B13B15B1B">Gomes, Ulisses Leite</mods:namePart>
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<mods:namePart id="4B57F8285F4DCEB965561D07CA970489">Charvet-Almeida, Patrícia</mods:namePart>
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<mods:date id="AD2D717AF7533AC17052F5F86855D998">2004</mods:date>
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<treatment id="03E5B501A008FF94FED4FDD0FCCAFA79" ID-DOI="http://doi.org/10.5281/zenodo.6272913" ID-GBIF-Taxon="119601482" ID-Zenodo-Dep="6272913" LSID="urn:lsid:zoobank.org:act:C633B59E-A366-45DC-A02C-3DF56D2AD36E" httpUri="http://treatment.plazi.org/id/03E5B501A008FF94FED4FDD0FCCAFA79" lastPageId="9" lastPageNumber="10" pageId="3" pageNumber="4">
<subSubSection id="C356579CA008FF9EFED4FDD0FC49FDAF" box="[264,940,590,616]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BF30417A008FF9EFED4FDD0FC49FDAF" blockId="3.[264,940,590,616]" box="[264,940,590,616]" pageId="3" pageNumber="4">
<heading id="D0BBB37BA008FF9EFED4FDD0FC49FDAF" bold="true" box="[264,940,590,616]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B938D805A008FF9EFED4FDD0FC49FDAF" bold="true" box="[264,940,590,616]" pageId="3" pageNumber="4">
<taxonomicName id="4C4C7F94A008FF9EFED4FDD0FE15FDAF" ID-CoL="6CBT4" box="[264,496,590,616]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A008FF9EFED4FDD0FE15FDAF" bold="true" box="[264,496,590,616]" italics="true" pageId="3" pageNumber="4">Dasyatis colarensis</emphasis>
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(
<figureCitation id="13771892A008FF9EFD95FDD0FD38FDAF" box="[585,733,590,616]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="5.[264,383,921,945]" captionStartId-1="5.[264,367,1861,1885]" captionStartId-2="6.[264,367,840,864]" captionStartId-3="6.[264,367,1488,1512]" captionStartId-4="7.[264,367,1801,1825]" captionTargetBox-0="[295,1297,313,893]" captionTargetBox-1="[430,1156,1103,1836]" captionTargetBox-2="[479,1096,310,790]" captionTargetBox-3="[428,1159,943,1461]" captionTargetBox-4="[571,1030,1035,1788]" captionTargetId-1="figure@5.[430,1157,1103,1837]" captionTargetId-2="figure@6.[442,1122,293,835]" captionTargetId-3="figure@6.[428,1160,941,1461]" captionTargetId-4="figure@7.[564,1055,1023,1794]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 2. Dasyatis colarensis n. sp, MNRJ 25179, holotype, male, 630 DW. 1, dorsal view; 2, ventral view." captionText-1="FIGURE 3: Dasyatis colarensis n. sp., UERJ 2006, paratype, male, 530 DW. Dorsal view." captionText-2="FIGURE 4: Dasyatis colarensis n. sp. Schematic drawing of oral papillae from the holotype." captionText-3="FIGURE 5: Dasyatis colarensis n. sp., UERJ 2006, paratype. Detail of mouth showing the coloration of the lower lip margin." captionText-4="FIGURE 6: Dasyatis colarensis n. sp. Schematic drawing of the right clasper of holotype in dorsal view. Abbreviations: mts, medial terminal margin serrae; hyp, hypopyle; pe, pent." httpUri-0="https://zenodo.org/record/168967/files/figure.png" httpUri-1="https://zenodo.org/record/168968/files/figure.png" httpUri-2="https://zenodo.org/record/157373/files/figure.png" httpUri-3="https://zenodo.org/record/168969/files/figure.png" httpUri-4="https://zenodo.org/record/157375/files/figure.png" pageId="3" pageNumber="4">Figures 1–6</figureCitation>
, Tables I and II)
</emphasis>
</heading>
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<subSubSection id="C356579CA008FF9EFED4FD0AFC51FC01" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BF30417A008FF9EFED4FD0AFDB4FCE1" blockId="3.[264,1325,660,966]" pageId="3" pageNumber="4">
<emphasis id="B938D805A008FF9EFED4FD0AFE96FD6A" box="[264,371,660,685]" italics="true" pageId="3" pageNumber="4">
<typeStatus id="54F7BAB5A008FF9EFED4FD0AFE96FD6A" box="[264,371,660,685]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
— 
<collectionCode id="ED5D9CD2A008FF9EFE43FD0AFE08FD69" box="[415,493,660,686]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="3" pageNumber="4">MNRJ</collectionCode>
25179, mature male, 
<quantity id="4CB4A9F2A008FF9EFD30FD0AFCBEFD69" box="[748,859,660,686]" metricMagnitude="0" metricUnit="m" metricValue="2.07" pageId="3" pageNumber="4" unit="mm" value="2070.0">2070 mm</quantity>
TL, 
<quantity id="4CB4A9F2A008FF9EFC48FD0AFC10FD69" box="[916,1013,660,686]" metricMagnitude="-1" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="4" unit="mm" value="630.0">630 mm</quantity>
DW, Marajó Bay, close to Colares Island, district of Colares, Pará State, 
<collectingCountry id="F35B4487A008FF9EFC95FD22FC71FD11" box="[841,916,700,726]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
, 
<geoCoordinate id="EE7862D0A008FF9EFC79FD22FBBBFD11" box="[933,1118,700,726]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-0.9108333">00° 54’ 39’’ S</geoCoordinate>
, 
<geoCoordinate id="EE7862D0A008FF9EFBACFD22FAC2FD11" box="[1136,1319,700,726]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="15" value="-48.289165">48° 17’21’’ W</geoCoordinate>
, approximately 6 meters deep, water temperature 28.6°C, salinity 4 ppt, collector, P. Charvet­Almeida, 
<date id="FFF222D7A008FF9EFE75FC92FDA9FCE1" box="[425,588,780,806]" pageId="3" pageNumber="4" value="2001-12-05">05/ XII/ 2001</date>
.
</paragraph>
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<typeStatus id="54F7BAB5A008FF9EFEE4FCAAFE4AFC8A" box="[312,431,820,845]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
</emphasis>
— 
<collectionCode id="ED5D9CD2A008FF9EFE3DFCAAFDC2FC89" box="[481,551,820,846]" pageId="3" pageNumber="4">UERJ</collectionCode>
2006, subadult male, 
<quantity id="4CB4A9F2A008FF9EFCEAFCAAFC4CFC89" box="[822,937,820,846]" metricMagnitude="0" metricUnit="m" metricValue="1.81" pageId="3" pageNumber="4" unit="mm" value="1810.0">1810 mm</quantity>
TL, 
<quantity id="4CB4A9F2A008FF9EFC34FCAAFBA8FC89" box="[1000,1101,820,846]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="4" unit="mm" value="530.0">530 mm</quantity>
DW, Marajó Bay, close to Colares Islands, district of Colares, Pará State, 
<collectingCountry id="F35B4487A008FF9EFC13FCC2FBFFFCB1" box="[975,1050,860,886]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
, collector P. Charvet­ Almeida, 
<date id="FFF222D7A008FF9EFE5CFC1AFDC5FC59" box="[384,544,900,926]" pageId="3" pageNumber="4" value="2003-11-05">05/ XI/ 2003</date>
; 
<collectionCode id="ED5D9CD2A008FF9EFDEDFC1AFD8AFC59" box="[561,623,900,926]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="4">MCP</collectionCode>
34811, juvenile male, 
<quantity id="4CB4A9F2A008FF9EFC56FC1AFC1BFC59" box="[906,1022,900,926]" metricMagnitude="0" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="4" unit="mm" value="1260.0">1260 mm</quantity>
TL, 
<quantity id="4CB4A9F2A008FF9EFB9CFC1AFB40FC59" box="[1088,1189,900,926]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="4" unit="mm" value="330.0">330 mm</quantity>
DW, same locality and collector as the other 
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, 0 
<date id="FFF222D7A008FF9EFCCBFC32FC55FC01" box="[791,944,940,966]" pageId="3" pageNumber="4" value="2003-11-04">4 / XI / 2003</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C356579CA008FF9EFED4FC69FD03FA96" pageId="3" pageNumber="4" type="diagnosis">
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<emphasis id="B938D805A008FF9EFED4FC69FE9AFBD7" box="[264,383,1015,1040]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF30417A008FF9EFEE4FB81FD03FA96" blockId="3.[264,1324,1015,1361]" pageId="3" pageNumber="4">
<taxonomicName id="4C4C7F94A008FF9EFEE4FB81FDF9FBFF" box="[312,540,1055,1080]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A008FF9EFEE4FB81FDF9FBFF" box="[312,540,1055,1080]" italics="true" pageId="3" pageNumber="4">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA008FF9EFDF8FB81FD87FBFE" box="[548,610,1055,1081]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
is distinguished by the combination of the following characters: a diamond­shaped (rhomboid) disc and an elongate snout (preorbital distance ranging from 35.7 to 37.7 % in DW); presence of a dark and well defined band outlining the lower lips; row of small tubercles along midline of disc from shoulder region to base of tail, tubercles somewhat randomly distributed; posterior margins of pectoral fins uniformly rounded; triangular­shaped pelvic fins with posterior margin almost straight or slightly sinuous, pelvic fin tips exceeding posterior margins of disc; dorsal caudal keel absent in the 
<typeStatus id="54F7BAB5A008FF9EFED4FAA9FE94FA96" box="[264,369,1335,1361]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, but vestigial in both 
<typeStatus id="54F7BAB5A008FF9EFDB2FAA9FD07FA96" box="[622,738,1335,1361]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C356579CA008FF95FED4FA19FDA0FD29" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
<paragraph id="8BF30417A008FF9EFED4FA19FE77FA67" blockId="3.[264,1324,1415,1881]" box="[264,402,1415,1440]" pageId="3" pageNumber="4">
<emphasis id="B938D805A008FF9EFED4FA19FE77FA67" box="[264,402,1415,1440]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="8BF30417A008FF9EFEE4FA31FDFCFA36" blockId="3.[264,1324,1415,1881]" pageId="3" pageNumber="4">Proportional dimensions as percentages of DW and meristic data are given respectively in tables I and II.</paragraph>
<paragraph id="8BF30417A008FF9EFEE4FA61FC35F926" blockId="3.[264,1324,1415,1881]" pageId="3" pageNumber="4">Snout long (preorbital length 35.7 to 37.7 % of DW) and acutely pointed. Maximum anterior angle in front of spiracles (snout angle) about 95° to 100°. Anterior margins of disc slightly concave with posterior margins convex. Disc width about equal to disc length (from 100.7 to 103.3 %). Eyes small, horizontal eye diameter 18.9 to 24.2 % in interorbital width. Spiracles well developed (horizontal eye diameter 35.5 to 47.0% of spiracle length). Posterior margins of pectoral fins perfectly rounded.</paragraph>
<paragraph id="8BF30417A008FF99FEE4F971FD57FE11" blockId="3.[264,1324,1415,1881]" lastBlockId="4.[264,1325,284,470]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Corners of nasal curtain rounded; nasal curtain with fringed posterior margin and with a median hiatus. Mouth arched, with a straight and dark band outlining the lower lips. Median portion of lower jaw (symphysial region) slightly concave. Teeth in a pavement pattern; cusps pointed and curved backwards in the 
<typeStatus id="54F7BAB5A00FFF99FC51FE82FC12FEF1" box="[909,1015,284,310]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
. Subadult specimen with teeth near symphysis almost pointed. Dental formulae: 43–45/45–60. Three to four papillae in transverse series on mouth floor. In the 
<typeStatus id="54F7BAB5A00FFF99FCEEFEF2FC79FE41" box="[818,924,364,390]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, the central papilla is curved forward, the two lateral ones show a singular bifurcated tip (
<typeStatus id="54F7BAB5A00FFF99FC10FE0AFBDBFE69" box="[972,1086,404,430]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
have no bifurcated papillae). Gill slit apertures sinuous.
</paragraph>
<paragraph id="8BF30417A00FFF99FED4FD95FD3DFD8C" blockId="4.[264,1322,523,587]" pageId="4" pageNumber="5">
<emphasis id="B938D805A00FFF99FED4FD95FE99FDE4" bold="true" box="[264,380,523,547]" pageId="4" pageNumber="5">TABLE I:</emphasis>
Proportional morphometrics of 
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and 
<typeStatus id="54F7BAB5A00FFF99FCADFD92FC32FDE4" box="[881,983,524,547]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C4C7F94A00FFF99FC23FD92FB6AFDE4" box="[1023,1167,524,547]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00FFF99FC23FD92FB6AFDE4" box="[1023,1167,524,547]" italics="true" pageId="4" pageNumber="5">D. colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00FFF99FB4BFD92FB37FDE4" box="[1175,1234,524,547]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
(in mm and as % of DW). SD = standard deviation.
</paragraph>
<paragraph id="8BF30417A00FFF99FECAFDF0FAFAF891" pageId="4" pageNumber="5">
<table id="F94CF6B7A00F0062FECCFDF3FAC0F891" box="[272,1317,621,1878]" gridcols="4" gridrows="34" pageId="4" pageNumber="5">
<tr id="357C0655A00F0062FECCFDF3FAC0FD07" box="[272,1317,621,704]" gridrow="0" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFDF3FDCFFD07" box="[272,554,621,704]" gridcol="0" gridrow="0" pageId="4" pageNumber="5">Parameters</th>
<th id="76AD6F29A00F0062FD93FDF3FD19FD07" box="[591,764,621,704]" gridcol="1" gridrow="0" pageId="4" pageNumber="5">HOLOTYPE MNRJ 25179 mm % of DW</th>
<th id="76AD6F29A00F0062FCC3FDF3FC26FD07" box="[799,963,621,704]" gridcol="2" gridrow="0" pageId="4" pageNumber="5">PARATYPE UERJ 2006 mm % of DW</th>
<th id="76AD6F29A00F0062FC31FDF3FAC0FD07" box="[1005,1317,621,704]" gridcol="3" gridrow="0" pageId="4" pageNumber="5">PARATYPE AV E ­ SD M C P 3 4 8 11 RAGE mm % of DW</th>
</tr>
<tr id="357C0655A00F0062FECCFD4AFAC0FD2E" box="[272,1317,724,745]" gridrow="1" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFD4AFDCFFD2E" box="[272,554,724,745]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">total length</th>
<td id="76AD6F29A00F0062FD93FD4AFD19FD2E" box="[591,764,724,745]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">2070 328.5</td>
<td id="76AD6F29A00F0062FCC3FD4AFC26FD2E" box="[799,963,724,745]" gridcol="2" gridrow="1" pageId="4" pageNumber="5">1810 341.5</td>
<td id="76AD6F29A00F0062FC31FD4AFAC0FD2E" box="[1005,1317,724,745]" gridcol="3" gridrow="1" pageId="4" pageNumber="5">1260 381.8 350.6 27.7</td>
</tr>
<tr id="357C0655A00F0062FECCFD69FAC0FCCB" box="[272,1317,759,780]" gridrow="2" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFD69FDCFFCCB" box="[272,554,759,780]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">disc width</th>
<td id="76AD6F29A00F0062FD93FD69FD19FCCB" box="[591,764,759,780]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">630 100</td>
<td id="76AD6F29A00F0062FCC3FD69FC26FCCB" box="[799,963,759,780]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">530 100</td>
<td id="76AD6F29A00F0062FC31FD69FAC0FCCB" box="[1005,1317,759,780]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">330 100 100 0</td>
</tr>
<tr id="357C0655A00F0062FECCFC84FAC0FCE8" box="[272,1317,794,815]" gridrow="3" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFC84FDCFFCE8" box="[272,554,794,815]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">disc length</th>
<td id="76AD6F29A00F0062FD93FC84FD19FCE8" box="[591,764,794,815]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">635 100.7</td>
<td id="76AD6F29A00F0062FCC3FC84FC26FCE8" box="[799,963,794,815]" gridcol="2" gridrow="3" pageId="4" pageNumber="5">548 103.3</td>
<td id="76AD6F29A00F0062FC31FC84FAC0FCE8" box="[1005,1317,794,815]" gridcol="3" gridrow="3" pageId="4" pageNumber="5">335 101.5 101.9 1.3</td>
</tr>
<tr id="357C0655A00F0062FECCFCA3FAC0FC95" box="[272,1317,829,850]" gridrow="4" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFCA3FDCFFC95" box="[272,554,829,850]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">internal disc length</th>
<td id="76AD6F29A00F0062FD93FCA3FD19FC95" box="[591,764,829,850]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">561 89.4</td>
<td id="76AD6F29A00F0062FCC3FCA3FC26FC95" box="[799,963,829,850]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">460 86.7</td>
<td id="76AD6F29A00F0062FC31FCA3FAC0FC95" box="[1005,1317,829,850]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">290 87.8 87.9 1.1</td>
</tr>
<tr id="357C0655A00F0062FECCFCFEFAC0FCB2" box="[272,1317,864,885]" gridrow="5" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFCFEFDCFFCB2" box="[272,554,864,885]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">precloacal length</th>
<td id="76AD6F29A00F0062FD93FCFEFD19FCB2" box="[591,764,864,885]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">530 84.1</td>
<td id="76AD6F29A00F0062FCC3FCFEFC26FCB2" box="[799,963,864,885]" gridcol="2" gridrow="5" pageId="4" pageNumber="5">470 88.6</td>
<td id="76AD6F29A00F0062FC31FCFEFAC0FCB2" box="[1005,1317,864,885]" gridcol="3" gridrow="5" pageId="4" pageNumber="5">285 86.3 86.3 2.2</td>
</tr>
<tr id="357C0655A00F0062FECCFC1DFAC0FC5F" box="[272,1317,899,920]" gridrow="6" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFC1DFDCFFC5F" box="[272,554,899,920]" gridcol="0" gridrow="6" pageId="4" pageNumber="5">cloaca­ sting length</th>
<td id="76AD6F29A00F0062FD93FC1DFD19FC5F" box="[591,764,899,920]" gridcol="1" gridrow="6" pageId="4" pageNumber="5">235 37.3</td>
<td id="76AD6F29A00F0062FCC3FC1DFC26FC5F" box="[799,963,899,920]" gridcol="2" gridrow="6" pageId="4" pageNumber="5">220 41.5</td>
<td id="76AD6F29A00F0062FC31FC1DFAC0FC5F" box="[1005,1317,899,920]" gridcol="3" gridrow="6" pageId="4" pageNumber="5">135 40.9 39.9 2.2</td>
</tr>
<tr id="357C0655A00F0062FECCFC38FAC0FC7C" box="[272,1317,934,955]" gridrow="7" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFC38FDCFFC7C" box="[272,554,934,955]" gridcol="0" gridrow="7" pageId="4" pageNumber="5">caudal length</th>
<td id="76AD6F29A00F0062FD93FC38FD19FC7C" box="[591,764,934,955]" gridcol="1" gridrow="7" pageId="4" pageNumber="5">1494 237</td>
<td id="76AD6F29A00F0062FCC3FC38FC26FC7C" box="[799,963,934,955]" gridcol="2" gridrow="7" pageId="4" pageNumber="5">1111 209.6</td>
<td id="76AD6F29A00F0062FC31FC38FAC0FC7C" box="[1005,1317,934,955]" gridcol="3" gridrow="7" pageId="4" pageNumber="5">970 296.3 247.7 44.3</td>
</tr>
<tr id="357C0655A00F0062FECCFC57FAC0FC19" box="[272,1317,969,990]" gridrow="8" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFC57FDCFFC19" box="[272,554,969,990]" gridcol="0" gridrow="8" pageId="4" pageNumber="5">caudal diameter</th>
<td id="76AD6F29A00F0062FD93FC57FD19FC19" box="[591,764,969,990]" gridcol="1" gridrow="8" pageId="4" pageNumber="5">14 2.2</td>
<td id="76AD6F29A00F0062FCC3FC57FC26FC19" box="[799,963,969,990]" gridcol="2" gridrow="8" pageId="4" pageNumber="5">11 2.0</td>
<td id="76AD6F29A00F0062FC31FC57FAC0FC19" box="[1005,1317,969,990]" gridcol="3" gridrow="8" pageId="4" pageNumber="5">9 2.7 2.3 0.3</td>
</tr>
<tr id="357C0655A00F0062FECCFC73FAC0FBC5" box="[272,1317,1005,1026]" gridrow="9" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFC73FDCFFBC5" box="[272,554,1005,1026]" gridcol="0" gridrow="9" pageId="4" pageNumber="5">height of ventral finfold</th>
<td id="76AD6F29A00F0062FD93FC73FD19FBC5" box="[591,764,1005,1026]" gridcol="1" gridrow="9" pageId="4" pageNumber="5">4 0.6</td>
<td id="76AD6F29A00F0062FCC3FC73FC26FBC5" box="[799,963,1005,1026]" gridcol="2" gridrow="9" pageId="4" pageNumber="5">3 0.5</td>
<td id="76AD6F29A00F0062FC31FC73FAC0FBC5" box="[1005,1317,1005,1026]" gridcol="3" gridrow="9" pageId="4" pageNumber="5">2 2.1 1.1 0.8</td>
</tr>
<tr id="357C0655A00F0062FECCFB91FAC0FBE3" box="[272,1317,1039,1060]" gridrow="10" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFB91FDCFFBE3" box="[272,554,1039,1060]" gridcol="0" gridrow="10" pageId="4" pageNumber="5">length of ventral finfold</th>
<td id="76AD6F29A00F0062FD93FB91FD19FBE3" box="[591,764,1039,1060]" gridcol="1" gridrow="10" pageId="4" pageNumber="5">235 37.3</td>
<td id="76AD6F29A00F0062FCC3FB91FC26FBE3" box="[799,963,1039,1060]" gridcol="2" gridrow="10" pageId="4" pageNumber="5">240 45.2</td>
<td id="76AD6F29A00F0062FC31FB91FAC0FBE3" box="[1005,1317,1039,1060]" gridcol="3" gridrow="10" pageId="4" pageNumber="5">170 51.5 44.6 7.1</td>
</tr>
<tr id="357C0655A00F0062FECCFBADFAC0FB8F" box="[272,1317,1075,1096]" gridrow="11" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFBADFDCFFB8F" box="[272,554,1075,1096]" gridcol="0" gridrow="11" pageId="4" pageNumber="5">horizontal eye diameter</th>
<td id="76AD6F29A00F0062FD93FBADFD19FB8F" box="[591,764,1075,1096]" gridcol="1" gridrow="11" pageId="4" pageNumber="5">12.8 2.0</td>
<td id="76AD6F29A00F0062FCC3FBADFC26FB8F" box="[799,963,1075,1096]" gridcol="2" gridrow="11" pageId="4" pageNumber="5">11 2.0</td>
<td id="76AD6F29A00F0062FC31FBADFAC0FB8F" box="[1005,1317,1075,1096]" gridcol="3" gridrow="11" pageId="4" pageNumber="5">8 2.4 2.1 0.2</td>
</tr>
<tr id="357C0655A00F0062FECCFBC8FAC0FBAC" box="[272,1317,1110,1131]" gridrow="12" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFBC8FDCFFBAC" box="[272,554,1110,1131]" gridcol="0" gridrow="12" pageId="4" pageNumber="5">orbit length</th>
<td id="76AD6F29A00F0062FD93FBC8FD19FBAC" box="[591,764,1110,1131]" gridcol="1" gridrow="12" pageId="4" pageNumber="5">16 2.53</td>
<td id="76AD6F29A00F0062FCC3FBC8FC26FBAC" box="[799,963,1110,1131]" gridcol="2" gridrow="12" pageId="4" pageNumber="5">17 3.2</td>
<td id="76AD6F29A00F0062FC31FBC8FAC0FBAC" box="[1005,1317,1110,1131]" gridcol="3" gridrow="12" pageId="4" pageNumber="5">11 3.33 2.8 0.4</td>
</tr>
<tr id="357C0655A00F0062FECCFBE7FAC0FB49" box="[272,1317,1145,1166]" gridrow="13" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFBE7FDCFFB49" box="[272,554,1145,1166]" gridcol="0" gridrow="13" pageId="4" pageNumber="5">interorbital width</th>
<td id="76AD6F29A00F0062FD93FBE7FD19FB49" box="[591,764,1145,1166]" gridcol="1" gridrow="13" pageId="4" pageNumber="5">61 9.6</td>
<td id="76AD6F29A00F0062FCC3FBE7FC26FB49" box="[799,963,1145,1166]" gridcol="2" gridrow="13" pageId="4" pageNumber="5">58 10.9</td>
<td id="76AD6F29A00F0062FC31FBE7FAC0FB49" box="[1005,1317,1145,1166]" gridcol="3" gridrow="13" pageId="4" pageNumber="5">33 11.3 10.2 0.6</td>
</tr>
<tr id="357C0655A00F0062FECCFB02FAC0FB76" box="[272,1317,1180,1201]" gridrow="14" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFB02FDCFFB76" box="[272,554,1180,1201]" gridcol="0" gridrow="14" pageId="4" pageNumber="5">spiracle length</th>
<td id="76AD6F29A00F0062FD93FB02FD19FB76" box="[591,764,1180,1201]" gridcol="1" gridrow="14" pageId="4" pageNumber="5">36 5.7</td>
<td id="76AD6F29A00F0062FCC3FB02FC26FB76" box="[799,963,1180,1201]" gridcol="2" gridrow="14" pageId="4" pageNumber="5">28 5.2</td>
<td id="76AD6F29A00F0062FC31FB02FAC0FB76" box="[1005,1317,1180,1201]" gridcol="3" gridrow="14" pageId="4" pageNumber="5">17 5.1 5.3 5.3</td>
</tr>
<tr id="357C0655A00F0062FECCFB21FAC0FB13" box="[272,1317,1215,1236]" gridrow="15" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFB21FDCFFB13" box="[272,554,1215,1236]" gridcol="0" gridrow="15" pageId="4" pageNumber="5">interspiracular length</th>
<td id="76AD6F29A00F0062FD93FB21FD19FB13" box="[591,764,1215,1236]" gridcol="1" gridrow="15" pageId="4" pageNumber="5">90 14.2</td>
<td id="76AD6F29A00F0062FCC3FB21FC26FB13" box="[799,963,1215,1236]" gridcol="2" gridrow="15" pageId="4" pageNumber="5">73 13.7</td>
<td id="76AD6F29A00F0062FC31FB21FAC0FB13" box="[1005,1317,1215,1236]" gridcol="3" gridrow="15" pageId="4" pageNumber="5">50 15.1 14.4 0.6</td>
</tr>
<tr id="357C0655A00F0062FECCFB7CFAC0FB30" box="[272,1317,1250,1271]" gridrow="16" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFB7CFDCFFB30" box="[272,554,1250,1271]" gridcol="0" gridrow="16" pageId="4" pageNumber="5">preoral length</th>
<td id="76AD6F29A00F0062FD93FB7CFD19FB30" box="[591,764,1250,1271]" gridcol="1" gridrow="16" pageId="4" pageNumber="5">220 34.9</td>
<td id="76AD6F29A00F0062FCC3FB7CFC26FB30" box="[799,963,1250,1271]" gridcol="2" gridrow="16" pageId="4" pageNumber="5">195 36.7</td>
<td id="76AD6F29A00F0062FC31FB7CFAC0FB30" box="[1005,1317,1250,1271]" gridcol="3" gridrow="16" pageId="4" pageNumber="5">115 34.8 35.5 1.1</td>
</tr>
<tr id="357C0655A00F0062FECCFA9BFAC0FADD" box="[272,1317,1285,1306]" gridrow="17" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFA9BFDCFFADD" box="[272,554,1285,1306]" gridcol="0" gridrow="17" pageId="4" pageNumber="5">preorbital length</th>
<td id="76AD6F29A00F0062FD93FA9BFD19FADD" box="[591,764,1285,1306]" gridcol="1" gridrow="17" pageId="4" pageNumber="5">225 35.7</td>
<td id="76AD6F29A00F0062FCC3FA9BFC26FADD" box="[799,963,1285,1306]" gridcol="2" gridrow="17" pageId="4" pageNumber="5">200 37.7</td>
<td id="76AD6F29A00F0062FC31FA9BFAC0FADD" box="[1005,1317,1285,1306]" gridcol="3" gridrow="17" pageId="4" pageNumber="5">120 36.6 36.6 1.0</td>
</tr>
<tr id="357C0655A00F0062FECCFAB9FAC0FA85" box="[272,1317,1319,1346]" gridrow="18" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFAB9FDCFFA85" box="[272,554,1319,1346]" gridcol="0" gridrow="18" pageId="4" pageNumber="5">1st and 5st gill slit distance</th>
<td id="76AD6F29A00F0062FD93FAB9FD19FA85" box="[591,764,1319,1346]" gridcol="1" gridrow="18" pageId="4" pageNumber="5">70 11.1</td>
<td id="76AD6F29A00F0062FCC3FAB9FC26FA85" box="[799,963,1319,1346]" gridcol="2" gridrow="18" pageId="4" pageNumber="5">55 10.3</td>
<td id="76AD6F29A00F0062FC31FAB9FAC0FA85" box="[1005,1317,1319,1346]" gridcol="3" gridrow="18" pageId="4" pageNumber="5">36 10.9 10.7 0.3</td>
</tr>
<tr id="357C0655A00F0062FECCFAD1FAC0FAAC" box="[272,1317,1359,1387]" gridrow="19" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFAD1FDCFFAAC" box="[272,554,1359,1387]" gridcol="0" gridrow="19" pageId="4" pageNumber="5">1st gill slits distance</th>
<td id="76AD6F29A00F0062FD93FAD1FD19FAAC" box="[591,764,1359,1387]" gridcol="1" gridrow="19" pageId="4" pageNumber="5">105 16.6</td>
<td id="76AD6F29A00F0062FCC3FAD1FC26FAAC" box="[799,963,1359,1387]" gridcol="2" gridrow="19" pageId="4" pageNumber="5">90 16.9</td>
<td id="76AD6F29A00F0062FC31FAD1FAC0FAAC" box="[1005,1317,1359,1387]" gridcol="3" gridrow="19" pageId="4" pageNumber="5">56 16.9 16.8 0.1</td>
</tr>
<tr id="357C0655A00F0062FECCFAE7FAC0FA49" box="[272,1317,1401,1422]" gridrow="20" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFAE7FDCFFA49" box="[272,554,1401,1422]" gridcol="0" gridrow="20" pageId="4" pageNumber="5">mouth width</th>
<td id="76AD6F29A00F0062FD93FAE7FD19FA49" box="[591,764,1401,1422]" gridcol="1" gridrow="20" pageId="4" pageNumber="5">48 7.6</td>
<td id="76AD6F29A00F0062FCC3FAE7FC26FA49" box="[799,963,1401,1422]" gridcol="2" gridrow="20" pageId="4" pageNumber="5">44 8.3</td>
<td id="76AD6F29A00F0062FC31FAE7FAC0FA49" box="[1005,1317,1401,1422]" gridcol="3" gridrow="20" pageId="4" pageNumber="5">31 9.3 8.4 8.4</td>
</tr>
<tr id="357C0655A00F0062FECCFA02FAC0FA76" box="[272,1317,1436,1457]" gridrow="21" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFA02FDCFFA76" box="[272,554,1436,1457]" gridcol="0" gridrow="21" pageId="4" pageNumber="5">internasal width</th>
<td id="76AD6F29A00F0062FD93FA02FD19FA76" box="[591,764,1436,1457]" gridcol="1" gridrow="21" pageId="4" pageNumber="5">64 10.1</td>
<td id="76AD6F29A00F0062FCC3FA02FC26FA76" box="[799,963,1436,1457]" gridcol="2" gridrow="21" pageId="4" pageNumber="5">57 10.7</td>
<td id="76AD6F29A00F0062FC31FA02FAC0FA76" box="[1005,1317,1436,1457]" gridcol="3" gridrow="21" pageId="4" pageNumber="5">36 10.9 10.6 0.3</td>
</tr>
<tr id="357C0655A00F0062FECCFA21FAC0FA13" box="[272,1317,1471,1492]" gridrow="22" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFA21FDCFFA13" box="[272,554,1471,1492]" gridcol="0" gridrow="22" pageId="4" pageNumber="5">pelvic fin width</th>
<td id="76AD6F29A00F0062FD93FA21FD19FA13" box="[591,764,1471,1492]" gridcol="1" gridrow="22" pageId="4" pageNumber="5">124 19.6</td>
<td id="76AD6F29A00F0062FCC3FA21FC26FA13" box="[799,963,1471,1492]" gridcol="2" gridrow="22" pageId="4" pageNumber="5">120 22.6</td>
<td id="76AD6F29A00F0062FC31FA21FAC0FA13" box="[1005,1317,1471,1492]" gridcol="3" gridrow="22" pageId="4" pageNumber="5">77 23.3 21.8 1.9</td>
</tr>
<tr id="357C0655A00F0062FECCFA7CFAC0FA30" box="[272,1317,1506,1527]" gridrow="23" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCFA7CFDCFFA30" box="[272,554,1506,1527]" gridcol="0" gridrow="23" pageId="4" pageNumber="5">pelvic fin length</th>
<td id="76AD6F29A00F0062FD93FA7CFD19FA30" box="[591,764,1506,1527]" gridcol="1" gridrow="23" pageId="4" pageNumber="5">135 21.4</td>
<td id="76AD6F29A00F0062FCC3FA7CFC26FA30" box="[799,963,1506,1527]" gridcol="2" gridrow="23" pageId="4" pageNumber="5">110 20.7</td>
<td id="76AD6F29A00F0062FC31FA7CFAC0FA30" box="[1005,1317,1506,1527]" gridcol="3" gridrow="23" pageId="4" pageNumber="5">66 20 20.7 0.7</td>
</tr>
<tr id="357C0655A00F0062FECCF99BFAC0F9DD" box="[272,1317,1541,1562]" gridrow="24" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF99BFDCFF9DD" box="[272,554,1541,1562]" gridcol="0" gridrow="24" pageId="4" pageNumber="5">base of pelvic fin</th>
<td id="76AD6F29A00F0062FD93F99BFD19F9DD" box="[591,764,1541,1562]" gridcol="1" gridrow="24" pageId="4" pageNumber="5">96 15.2</td>
<td id="76AD6F29A00F0062FCC3F99BFC26F9DD" box="[799,963,1541,1562]" gridcol="2" gridrow="24" pageId="4" pageNumber="5">80 15.0</td>
<td id="76AD6F29A00F0062FC31F99BFAC0F9DD" box="[1005,1317,1541,1562]" gridcol="3" gridrow="24" pageId="4" pageNumber="5">41 12.4 14.2 1.5</td>
</tr>
<tr id="357C0655A00F0062FECCF9B6FAC0F9FA" box="[272,1317,1576,1597]" gridrow="25" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF9B6FDCFF9FA" box="[272,554,1576,1597]" gridcol="0" gridrow="25" pageId="4" pageNumber="5">anterior margin of pelvic fin</th>
<td id="76AD6F29A00F0062FD93F9B6FD19F9FA" box="[591,764,1576,1597]" gridcol="1" gridrow="25" pageId="4" pageNumber="5">14 2.2</td>
<td id="76AD6F29A00F0062FCC3F9B6FC26F9FA" box="[799,963,1576,1597]" gridcol="2" gridrow="25" pageId="4" pageNumber="5">125 26.0</td>
<td id="76AD6F29A00F0062FC31F9B6FAC0F9FA" box="[1005,1317,1576,1597]" gridcol="3" gridrow="25" pageId="4" pageNumber="5">86 26.6 23.9 1.9</td>
</tr>
<tr id="357C0655A00F0062FECCF9D5FAC0F9A7" box="[272,1317,1611,1632]" gridrow="26" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF9D5FDCFF9A7" box="[272,554,1611,1632]" gridcol="0" gridrow="26" pageId="4" pageNumber="5">posterior margin of pelvic fin</th>
<td id="76AD6F29A00F0062FD93F9D5FD19F9A7" box="[591,764,1611,1632]" gridcol="1" gridrow="26" pageId="4" pageNumber="5">25 3.9</td>
<td id="76AD6F29A00F0062FCC3F9D5FC26F9A7" box="[799,963,1611,1632]" gridcol="2" gridrow="26" pageId="4" pageNumber="5">100 23.3</td>
<td id="76AD6F29A00F0062FC31F9D5FAC0F9A7" box="[1005,1317,1611,1632]" gridcol="3" gridrow="26" pageId="4" pageNumber="5">77 23.3 27.9 10.9</td>
</tr>
<tr id="357C0655A00F0062FECCF9F0FAC0F944" box="[272,1317,1646,1667]" gridrow="27" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF9F0FDCFF944" box="[272,554,1646,1667]" gridcol="0" gridrow="27" pageId="4" pageNumber="5">width of pelvic fin girdle</th>
<td id="76AD6F29A00F0062FD93F9F0FD19F944" box="[591,764,1646,1667]" gridcol="1" gridrow="27" pageId="4" pageNumber="5">82 13.0</td>
<td id="76AD6F29A00F0062FCC3F9F0FC26F944" box="[799,963,1646,1667]" gridcol="2" gridrow="27" pageId="4" pageNumber="5">68 13.3</td>
<td id="76AD6F29A00F0062FC31F9F0FAC0F944" box="[1005,1317,1646,1667]" gridcol="3" gridrow="27" pageId="4" pageNumber="5">41 13.3 12.9 0.1</td>
</tr>
<tr id="357C0655A00F0062FECCF90FFAC0F961" box="[272,1317,1681,1702]" gridrow="28" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF90FFDCFF961" box="[272,554,1681,1702]" gridcol="0" gridrow="28" pageId="4" pageNumber="5">clasper inner length</th>
<td id="76AD6F29A00F0062FD93F90FFD19F961" box="[591,764,1681,1702]" gridcol="1" gridrow="28" pageId="4" pageNumber="5">140 22.2</td>
<td id="76AD6F29A00F0062FCC3F90FFC26F961" box="[799,963,1681,1702]" gridcol="2" gridrow="28" pageId="4" pageNumber="5">100 18.8</td>
<td id="76AD6F29A00F0062FC31F90FFAC0F961" box="[1005,1317,1681,1702]" gridcol="3" gridrow="28" pageId="4" pageNumber="5">20 6.0 15.7 8.5</td>
</tr>
<tr id="357C0655A00F0062FECCF92AFAC0F90E" box="[272,1317,1716,1737]" gridrow="29" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF92AFDCFF90E" box="[272,554,1716,1737]" gridcol="0" gridrow="29" pageId="4" pageNumber="5">clasper outer length</th>
<td id="76AD6F29A00F0062FD93F92AFD19F90E" box="[591,764,1716,1737]" gridcol="1" gridrow="29" pageId="4" pageNumber="5">85 13.4</td>
<td id="76AD6F29A00F0062FCC3F92AFC26F90E" box="[799,963,1716,1737]" gridcol="2" gridrow="29" pageId="4" pageNumber="5">75 13.7</td>
<td id="76AD6F29A00F0062FC31F92AFAC0F90E" box="[1005,1317,1716,1737]" gridcol="3" gridrow="29" pageId="4" pageNumber="5">12 3.6 10.3 5.7</td>
</tr>
<tr id="357C0655A00F0062FECCF949FAC0F92B" box="[272,1317,1751,1772]" gridrow="30" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF949FDCFF92B" box="[272,554,1751,1772]" gridcol="0" gridrow="30" pageId="4" pageNumber="5">clasper glans</th>
<td id="76AD6F29A00F0062FD93F949FD19F92B" box="[591,764,1751,1772]" gridcol="1" gridrow="30" pageId="4" pageNumber="5">44 31.4</td>
<td id="76AD6F29A00F0062FCC3F949FC26F92B" box="[799,963,1751,1772]" gridcol="2" gridrow="30" pageId="4" pageNumber="5">38 35.0</td>
<td id="76AD6F29A00F0062FC31F949FAC0F92B" box="[1005,1317,1751,1772]" gridcol="3" gridrow="30" pageId="4" pageNumber="5">7 38 7.0 2.8</td>
</tr>
<tr id="357C0655A00F0062FECCF965FAC0F8D7" box="[272,1317,1787,1808]" gridrow="31" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF965FDCFF8D7" box="[272,554,1787,1808]" gridcol="0" gridrow="31" pageId="4" pageNumber="5">snout­ sting length</th>
<td id="76AD6F29A00F0062FD93F965FD19F8D7" box="[591,764,1787,1808]" gridcol="1" gridrow="31" pageId="4" pageNumber="5">797 126.5</td>
<td id="76AD6F29A00F0062FCC3F965FC26F8D7" box="[799,963,1787,1808]" gridcol="2" gridrow="31" pageId="4" pageNumber="5">200 132.0</td>
<td id="76AD6F29A00F0062FC31F965FAC0F8D7" box="[1005,1317,1787,1808]" gridcol="3" gridrow="31" pageId="4" pageNumber="5">420 127.2 128.6 3.0</td>
</tr>
<tr id="357C0655A00F0062FECCF880FAC0F8F4" box="[272,1317,1822,1843]" gridrow="32" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF880FDCFF8F4" box="[272,554,1822,1843]" gridcol="0" gridrow="32" pageId="4" pageNumber="5">prenasal length</th>
<td id="76AD6F29A00F0062FD93F880FD19F8F4" box="[591,764,1822,1843]" gridcol="1" gridrow="32" pageId="4" pageNumber="5">186 29.5</td>
<td id="76AD6F29A00F0062FCC3F880FC26F8F4" box="[799,963,1822,1843]" gridcol="2" gridrow="32" pageId="4" pageNumber="5">170 32.0</td>
<td id="76AD6F29A00F0062FC31F880FAC0F8F4" box="[1005,1317,1822,1843]" gridcol="3" gridrow="32" pageId="4" pageNumber="5">100 30.3 30.6 30.6</td>
</tr>
<tr id="357C0655A00F0062FECCF8DFFAC0F891" box="[272,1317,1857,1878]" gridrow="33" pageId="4" pageNumber="5">
<th id="76AD6F29A00F0062FECCF8DFFDCFF891" box="[272,554,1857,1878]" gridcol="0" gridrow="33" pageId="4" pageNumber="5">caudal base­sting</th>
<td id="76AD6F29A00F0062FD93F8DFFD19F891" box="[591,764,1857,1878]" gridcol="1" gridrow="33" pageId="4" pageNumber="5">220 34.9</td>
<td id="76AD6F29A00F0062FCC3F8DFFC26F891" box="[799,963,1857,1878]" gridcol="2" gridrow="33" pageId="4" pageNumber="5">200 37.7</td>
<td id="76AD6F29A00F0062FC31F8DFFAC0F891" box="[1005,1317,1857,1878]" gridcol="3" gridrow="33" pageId="4" pageNumber="5">125 37.8 36.8 1.6</td>
</tr>
</table>
</paragraph>
<caption id="DF33549FA00EFF98FED4FC07FE48FC14" httpUri="https://zenodo.org/record/168967/files/figure.png" pageId="5" pageNumber="6" targetBox="[295,1297,313,893]" targetPageId="5">
<paragraph id="8BF30417A00EFF98FED4FC07FE48FC14" blockId="5.[264,1322,921,979]" pageId="5" pageNumber="6">
<emphasis id="B938D805A00EFF98FED4FC07FE51FC76" bold="true" box="[264,436,921,945]" pageId="5" pageNumber="6">FIGURES 1–2.</emphasis>
<taxonomicName id="4C4C7F94A00EFF98FE60FC04FD68FC76" box="[444,653,922,945]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A00EFF98FE60FC04FD68FC76" box="[444,653,922,945]" italics="true" pageId="5" pageNumber="6">Dasyatis colarensis</emphasis>
</taxonomicName>
n. sp, MNRJ 25179, holotype, male, 630 DW. 1, dorsal view; 2, ventral view.
</paragraph>
</caption>
<caption id="DF33549FA00EFF98FED4F8DBFB35F89B" box="[264,1232,1861,1885]" httpUri="https://zenodo.org/record/168968/files/figure.png" pageId="5" pageNumber="6" targetBox="[430,1156,1103,1836]" targetPageId="5">
<paragraph id="8BF30417A00EFF98FED4F8DBFB35F89B" blockId="5.[264,1232,1861,1885]" box="[264,1232,1861,1885]" pageId="5" pageNumber="6">
<emphasis id="B938D805A00EFF98FED4F8DBFE66F89A" bold="true" box="[264,387,1861,1885]" pageId="5" pageNumber="6">FIGURE 3</emphasis>
: 
<taxonomicName id="4C4C7F94A00EFF98FE4EF8DBFD86F89B" box="[402,611,1861,1884]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00EFF98FE4EF8DBFD86F89B" box="[402,611,1861,1884]" italics="true" pageId="5" pageNumber="6">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00EFF98FDB5F8DBFD41F89B" box="[617,676,1861,1884]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, UERJ 2006, paratype, male, 530 DW. Dorsal view.
</paragraph>
</caption>
<caption id="DF33549FA00DFF9BFED4FCD6FB0EFC98" box="[264,1259,840,864]" httpUri="https://zenodo.org/record/157373/files/figure.png" pageId="6" pageNumber="7" targetBox="[479,1096,310,790]" targetPageId="6">
<paragraph id="8BF30417A00DFF9BFED4FCD6FB0EFC98" blockId="6.[264,1259,840,864]" box="[264,1259,840,864]" pageId="6" pageNumber="7">
<emphasis id="B938D805A00DFF9BFED4FCD6FE66FCA7" bold="true" box="[264,387,840,864]" pageId="6" pageNumber="7">FIGURE 4</emphasis>
: 
<taxonomicName id="4C4C7F94A00DFF9BFE4EFCD6FD86FC98" box="[402,611,840,863]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00DFF9BFE4EFCD6FD86FC98" box="[402,611,840,863]" italics="true" pageId="6" pageNumber="7">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00DFF9BFDB5FCD6FD47FC98" box="[617,674,840,863]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
Schematic drawing of oral papillae from the holotype.
</paragraph>
</caption>
<caption id="DF33549FA00DFF9BFED4FA4EFDD0F9CD" httpUri="https://zenodo.org/record/168969/files/figure.png" pageId="6" pageNumber="7" targetBox="[428,1159,943,1461]" targetPageId="6">
<paragraph id="8BF30417A00DFF9BFED4FA4EFDD0F9CD" blockId="6.[264,1322,1488,1546]" pageId="6" pageNumber="7">
<emphasis id="B938D805A00DFF9BFED4FA4EFE61FA2F" bold="true" box="[264,388,1488,1512]" pageId="6" pageNumber="7">FIGURE 5</emphasis>
: 
<taxonomicName id="4C4C7F94A00DFF9BFE4FFA4EFD86FA20" box="[403,611,1488,1511]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00DFF9BFE4FFA4EFD86FA20" box="[403,611,1488,1511]" italics="true" pageId="6" pageNumber="7">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00DFF9BFDB7FA4EFD43FA20" box="[619,678,1488,1511]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
, UERJ 2006, paratype. Detail of mouth showing the coloration of the lower lip margin.
</paragraph>
</caption>
<paragraph id="8BF30417A00DFF9BFEE4F9C4FBCAF8A3" blockId="6.[264,1323,1626,1892]" pageId="6" pageNumber="7">
Pelvic fins triangular, with posterior margin almost straight or slightly sinuous. Tip of pelvic fin sharp, exceeding the limits of the posterior margin of disc. Well developed claspers in the 
<typeStatus id="54F7BAB5A00DFF9BFE67F934FDC7F903" box="[443,546,1706,1732]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
(with residual traces of sperm at the moment of capture). Medial margin of clasper groove slightly serrated at the glans. Tail (which seems complete) more than twice disc width (209.6 to 296.3%). Dorsal caudal keel absent in 
<typeStatus id="54F7BAB5A00DFF9BFBE0F964FB47F8D3" box="[1084,1186,1786,1812]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
but present in both 
<typeStatus id="54F7BAB5A00DFF9BFEBDF8BCFE34F8FB" box="[353,465,1826,1852]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
as a low vestigial keel. Ventral finfold not high, its maximum height less than one third of caudal diameter at sting level (from 22.2 to 28.5%).
</paragraph>
<paragraph id="8BF30417A00CFF9AFED4FE83FB80FEF1" blockId="7.[264,1125,284,310]" box="[264,1125,284,310]" pageId="7" pageNumber="8">
<emphasis id="B938D805A00CFF9AFED4FE83FE99FEF2" bold="true" box="[264,380,285,309]" pageId="7" pageNumber="8">TABLE II</emphasis>
: Meristic data of 
<typeStatus id="54F7BAB5A00CFF9AFDE5FE83FD73FEF3" box="[569,662,285,308]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54F7BAB5A00CFF9AFD16FE83FCD5FEF3" box="[714,816,285,308]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C4C7F94A00CFF9AFC88FE80FBC1FEF2" box="[852,1060,286,309]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00CFF9AFC88FE80FBC1FEF2" box="[852,1060,286,309]" italics="true" pageId="7" pageNumber="8">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00CFF9AFBF0FE83FB80FEF1" box="[1068,1125,284,310]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF30417A00CFF95FD3AFEC7FDBCFD89" blockId="7.[417,1277,345,398]" lastBlockId="8.[264,1323,284,750]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<typeStatus id="54F7BAB5A00CFF9AFD3AFEC7FCDAFEA9" box="[742,831,345,366]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
<typeStatus id="54F7BAB5A00CFF9AFC15FEC7FB08FEA9" box="[969,1261,345,366]" pageId="7" pageNumber="8" type="paratype">Paratype Paratype</typeStatus>
Countings MNRJ 25179 UERJ 2006 MCP 34811 Central upper surface of disc covered with small, widespread, and flattened tubercles in a band along midline from level of orbits to base of tail. Tubercles also on rostral, preorbital and interorbital regions. Margin of disc naked. Dorsal thorn counts: 6 to 9 nuchals, 1 to 2 scapulars, 2 to 3 suprascapulars on the left side and 1 to 3 on the right, 12 to 17 median dorsals, and 6 to 9 median caudals. Ventral finfold, pelvic fins, claspers and ventral surface of the disc smooth. Sting missing in 
<typeStatus id="54F7BAB5A003FF95FD6CFE7AFCF2FE39" box="[688,791,484,510]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(probably removed, original groove present). Counts of lateral sting barbs: 42 (left) and 48 (right) in juvenile 
<typeStatus id="54F7BAB5A003FF95FC22FD92FB83FDE1" box="[1022,1126,524,550]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
, 69 (left) and 70 (right) in subadult specimen.
</paragraph>
<paragraph id="8BF30417A00CFF9AFECCFE00FB3FFC5B" pageId="7" pageNumber="8">
<table id="F94CF6B7A00C0062FECCFE00FB18FC5B" box="[272,1277,414,924]" gridcols="4" gridrows="14" pageId="7" pageNumber="8">
<tr id="357C0655A00C0062FECCFE00FB18FE74" box="[272,1277,414,435]" gridrow="0" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFE00FDAEFE74" box="[272,587,414,435]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">teeth (superior jaw)</th>
<th id="76AD6F29A00C0062FD2EFE00FCD1FE74" box="[754,820,414,435]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">43</th>
<th id="76AD6F29A00C0062FC14FE00FBFAFE74" box="[968,1055,414,435]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">43</th>
<th id="76AD6F29A00C0062FB70FE00FB18FE74" box="[1196,1277,414,435]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">45</th>
</tr>
<tr id="357C0655A00C0062FECCFE5DFB18FE1F" box="[272,1277,451,472]" gridrow="1" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFE5DFDAEFE1F" box="[272,587,451,472]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">teeth (inferior jaw)</th>
<td id="76AD6F29A00C0062FD2EFE5DFCD1FE1F" box="[754,820,451,472]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">60</td>
<td id="76AD6F29A00C0062FC14FE5DFBFAFE1F" box="[968,1055,451,472]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">51</td>
<td id="76AD6F29A00C0062FB70FE5DFB18FE1F" box="[1196,1277,451,472]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">45</td>
</tr>
<tr id="357C0655A00C0062FECCFE76FB18FE3A" box="[272,1277,488,509]" gridrow="2" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFE76FDAEFE3A" box="[272,587,488,509]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">vellum papillae</th>
<td id="76AD6F29A00C0062FD2EFE76FCD1FE3A" box="[754,820,488,509]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">41</td>
<td id="76AD6F29A00C0062FC14FE76FBFAFE3A" box="[968,1055,488,509]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">damaged</td>
<td id="76AD6F29A00C0062FB70FE76FB18FE3A" box="[1196,1277,488,509]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">41</td>
</tr>
<tr id="357C0655A00C0062FECCFD90FB18FDE4" box="[272,1277,526,547]" gridrow="3" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFD90FDAEFDE4" box="[272,587,526,547]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">pelvic radials (left / right)</th>
<td id="76AD6F29A00C0062FD2EFD90FCD1FDE4" box="[754,820,526,547]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">26 / 26</td>
<td id="76AD6F29A00C0062FC14FD90FBFAFDE4" box="[968,1055,526,547]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">­</td>
<td id="76AD6F29A00C0062FB70FD90FB18FDE4" box="[1196,1277,526,547]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">­</td>
</tr>
<tr id="357C0655A00C0062FECCFDADFB18FD8F" box="[272,1277,563,584]" gridrow="4" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFDADFDAEFD8F" box="[272,587,563,584]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">pre­sting vertebral centra</th>
<td id="76AD6F29A00C0062FD2EFDADFCD1FD8F" box="[754,820,563,584]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">127</td>
<td id="76AD6F29A00C0062FC14FDADFBFAFD8F" box="[968,1055,563,584]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">­</td>
<td id="76AD6F29A00C0062FB70FDADFB18FD8F" box="[1196,1277,563,584]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">­</td>
</tr>
<tr id="357C0655A00C0062FECCFDC6FB18FDAA" box="[272,1277,600,621]" gridrow="5" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFDC6FDAEFDAA" box="[272,587,600,621]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">monospondylic vertebral centra</th>
<td id="76AD6F29A00C0062FD2EFDC6FCD1FDAA" box="[754,820,600,621]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">46</td>
<td id="76AD6F29A00C0062FC14FDC6FBFAFDAA" box="[968,1055,600,621]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">­</td>
<td id="76AD6F29A00C0062FB70FDC6FB18FDAA" box="[1196,1277,600,621]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">­</td>
</tr>
<tr id="357C0655A00C0062FECCFDE3FB18FD55" box="[272,1277,637,658]" gridrow="6" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFDE3FDAEFD55" box="[272,587,637,658]" gridcol="0" gridrow="6" pageId="7" pageNumber="8">nuchalspines</th>
<td id="76AD6F29A00C0062FD2EFDE3FCD1FD55" box="[754,820,637,658]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">9</td>
<td id="76AD6F29A00C0062FC14FDE3FBFAFD55" box="[968,1055,637,658]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">9</td>
<td id="76AD6F29A00C0062FB70FDE3FB18FD55" box="[1196,1277,637,658]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">6</td>
</tr>
<tr id="357C0655A00C0062FECCFD3DFB18FD7F" box="[272,1277,675,696]" gridrow="7" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFD3DFDAEFD7F" box="[272,587,675,696]" gridcol="0" gridrow="7" pageId="7" pageNumber="8">scapulars thorns</th>
<td id="76AD6F29A00C0062FD2EFD3DFCD1FD7F" box="[754,820,675,696]" gridcol="1" gridrow="7" pageId="7" pageNumber="8">1</td>
<td id="76AD6F29A00C0062FC14FD3DFBFAFD7F" box="[968,1055,675,696]" gridcol="2" gridrow="7" pageId="7" pageNumber="8">2</td>
<td id="76AD6F29A00C0062FB70FD3DFB18FD7F" box="[1196,1277,675,696]" gridcol="3" gridrow="7" pageId="7" pageNumber="8">1</td>
</tr>
<tr id="357C0655A00C0062FECCFD56FB18FD1A" box="[272,1277,712,733]" gridrow="8" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFD56FDAEFD1A" box="[272,587,712,733]" gridcol="0" gridrow="8" pageId="7" pageNumber="8">suprascapular thorns (left / right)</th>
<td id="76AD6F29A00C0062FD2EFD56FCD1FD1A" box="[754,820,712,733]" gridcol="1" gridrow="8" pageId="7" pageNumber="8">3 / 3</td>
<td id="76AD6F29A00C0062FC14FD56FBFAFD1A" box="[968,1055,712,733]" gridcol="2" gridrow="8" pageId="7" pageNumber="8">2 / 2</td>
<td id="76AD6F29A00C0062FB70FD56FB18FD1A" box="[1196,1277,712,733]" gridcol="3" gridrow="8" pageId="7" pageNumber="8">3 / 1</td>
</tr>
<tr id="357C0655A00C0062FECCFD73FB18FCC5" box="[272,1277,749,770]" gridrow="9" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFD73FDAEFCC5" box="[272,587,749,770]" gridcol="0" gridrow="9" pageId="7" pageNumber="8">mediodorsal thorns</th>
<td id="76AD6F29A00C0062FD2EFD73FCD1FCC5" box="[754,820,749,770]" gridcol="1" gridrow="9" pageId="7" pageNumber="8">12</td>
<td id="76AD6F29A00C0062FC14FD73FBFAFCC5" box="[968,1055,749,770]" gridcol="2" gridrow="9" pageId="7" pageNumber="8">17</td>
<td id="76AD6F29A00C0062FB70FD73FB18FCC5" box="[1196,1277,749,770]" gridcol="3" gridrow="9" pageId="7" pageNumber="8">16</td>
</tr>
<tr id="357C0655A00C0062FECCFC8CFB18FCE0" box="[272,1277,786,807]" gridrow="10" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFC8CFDAEFCE0" box="[272,587,786,807]" gridcol="0" gridrow="10" pageId="7" pageNumber="8">mediocaudal thorns</th>
<td id="76AD6F29A00C0062FD2EFC8CFCD1FCE0" box="[754,820,786,807]" gridcol="1" gridrow="10" pageId="7" pageNumber="8">9</td>
<td id="76AD6F29A00C0062FC14FC8CFBFAFCE0" box="[968,1055,786,807]" gridcol="2" gridrow="10" pageId="7" pageNumber="8">8</td>
<td id="76AD6F29A00C0062FB70FC8CFB18FCE0" box="[1196,1277,786,807]" gridcol="3" gridrow="10" pageId="7" pageNumber="8">6</td>
</tr>
<tr id="357C0655A00C0062FECCFCA9FB18FC8B" box="[272,1277,823,844]" gridrow="11" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFCA9FDAEFC8B" box="[272,587,823,844]" gridcol="0" gridrow="11" pageId="7" pageNumber="8">oral papillae</th>
<td id="76AD6F29A00C0062FD2EFCA9FCD1FC8B" box="[754,820,823,844]" gridcol="1" gridrow="11" pageId="7" pageNumber="8">3</td>
<td id="76AD6F29A00C0062FC14FCA9FBFAFC8B" box="[968,1055,823,844]" gridcol="2" gridrow="11" pageId="7" pageNumber="8">4</td>
<td id="76AD6F29A00C0062FB70FCA9FB18FC8B" box="[1196,1277,823,844]" gridcol="3" gridrow="11" pageId="7" pageNumber="8">3</td>
</tr>
<tr id="357C0655A00C0062FECCFCC3FB18FCB5" box="[272,1277,861,882]" gridrow="12" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFCC3FDAEFCB5" box="[272,587,861,882]" gridcol="0" gridrow="12" pageId="7" pageNumber="8">pelvic fin fringes</th>
<td id="76AD6F29A00C0062FD2EFCC3FCD1FCB5" box="[754,820,861,882]" gridcol="1" gridrow="12" pageId="7" pageNumber="8">18</td>
<td id="76AD6F29A00C0062FC14FCC3FBFAFCB5" box="[968,1055,861,882]" gridcol="2" gridrow="12" pageId="7" pageNumber="8">17</td>
<td id="76AD6F29A00C0062FB70FCC3FB18FCB5" box="[1196,1277,861,882]" gridcol="3" gridrow="12" pageId="7" pageNumber="8">17</td>
</tr>
<tr id="357C0655A00C0062FECCFC1CFB18FC5B" box="[272,1277,898,924]" gridrow="13" pageId="7" pageNumber="8">
<th id="76AD6F29A00C0062FECCFC1CFDAEFC5B" box="[272,587,898,924]" gridcol="0" gridrow="13" pageId="7" pageNumber="8">snout angle</th>
<td id="76AD6F29A00C0062FD2EFC1CFCD1FC5B" box="[754,820,898,924]" gridcol="1" gridrow="13" pageId="7" pageNumber="8">96°</td>
<td id="76AD6F29A00C0062FC14FC1CFBFAFC5B" box="[968,1055,898,924]" gridcol="2" gridrow="13" pageId="7" pageNumber="8">95°</td>
<td id="76AD6F29A00C0062FB70FC1CFB18FC5B" box="[1196,1277,898,924]" gridcol="3" gridrow="13" pageId="7" pageNumber="8">100°</td>
</tr>
</table>
</paragraph>
<caption id="DF33549FA00CFF9AFED4F897FB94F884" httpUri="https://zenodo.org/record/157375/files/figure.png" pageId="7" pageNumber="8" targetBox="[571,1030,1035,1788]" targetPageId="7">
<paragraph id="8BF30417A00CFF9AFED4F897FB94F884" blockId="7.[264,1323,1801,1859]" pageId="7" pageNumber="8">
<emphasis id="B938D805A00CFF9AFED4F897FE66F8E6" bold="true" box="[264,387,1801,1825]" pageId="7" pageNumber="8">FIGURE 6</emphasis>
: 
<taxonomicName id="4C4C7F94A00CFF9AFE4EF897FD87F8E7" box="[402,610,1801,1824]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="colarensis" status="sp. nov.">
<emphasis id="B938D805A00CFF9AFE4EF897FD87F8E7" box="[402,610,1801,1824]" italics="true" pageId="7" pageNumber="8">Dasyatis colarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B657EA00CFF9AFDB6F897FD46F8E7" box="[618,675,1801,1824]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
Schematic drawing of the right clasper of holotype in dorsal view. Abbreviations: mts, medial terminal margin serrae; hyp, hypopyle; pe, pent.
</paragraph>
</caption>
<paragraph id="8BF30417A003FF95FEE4FDC2FDA0FD29" blockId="8.[264,1323,284,750]" pageId="8" pageNumber="9">Coloration in life very similar to that in preservative. Dorsal surface of the disc light brown; ventral surface whitish­gray with margins of pectoral fins and posterior margins of pelvic fins darker. Anterior margin of pelvic fin whitish to yellowish. Claspers and caudal ventral finfold dark brown.</paragraph>
</subSubSection>
<subSubSection id="C356579CA003FF95FED4FCBAFC12FC4C" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BF30417A003FF95FED4FCBAFE60FCFA" blockId="8.[264,1323,804,907]" box="[264,389,804,829]" pageId="8" pageNumber="9">
<emphasis id="B938D805A003FF95FED4FCBAFE60FCFA" box="[264,389,804,829]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
</paragraph>
<paragraph id="8BF30417A003FF95FEE4FCD2FC12FC4C" blockId="8.[264,1323,804,907]" pageId="8" pageNumber="9">
The specific name was selected for the geographic location of 
<typeStatus id="54F7BAB5A003FF95FBD1FCD2FB92FCA1" box="[1037,1143,844,870]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, from the proximities of Colares Island, district of Colares (Pará State, 
<collectingCountry id="F35B4487A003FF95FC41FCEFFC0FFC4C" box="[925,1002,881,907]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C356579CA003FF95FED4FC27FE6CFC15" box="[264,393,953,978]" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BF30417A003FF95FED4FC27FE6CFC15" blockId="8.[264,1324,953,1894]" box="[264,393,953,978]" pageId="8" pageNumber="9">
<emphasis id="B938D805A003FF95FED4FC27FE6CFC15" box="[264,393,953,978]" italics="true" pageId="8" pageNumber="9">Discussion</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C356579CA003FF95FEE4FC42FDACF9B1" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BF30417A003FF95FEE4FC42FC33FB21" blockId="8.[264,1324,953,1894]" pageId="8" pageNumber="9">
<taxonomicName id="4C4C7F94A003FF95FEE4FC42FDC7FC32" ID-CoL="6CBT4" box="[312,546,988,1013]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FEE4FC42FDC7FC32" box="[312,546,988,1013]" italics="true" pageId="8" pageNumber="9">Dasyatis colarensis</emphasis>
</taxonomicName>
resembles the diamond shaped, long snouted 
<taxonomicName id="4C4C7F94A003FF95FBBFFC42FB2FFC32" ID-CoL="3ZY5" box="[1123,1226,988,1013]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938D805A003FF95FBBFFC42FB2FFC32" box="[1123,1226,988,1013]" italics="true" pageId="8" pageNumber="9">Dasyatis</emphasis>
</taxonomicName>
species group of 
<bibRefCitation id="EFDD79E6A003FF95FEA6FB9AFD02FBD9" author="Compagno" box="[378,743,1028,1054]" pageId="8" pageNumber="11" refString="Compagno, L. J. V. &amp; Roberts, T. (1982) Freshwater stingrays (Dasyatidae) of Southeast Asia and New Guinea, with description of a new species of Himantura and reports of unidentified species. Environmental Biology of Fishes, 7 (4), 321 - 339." type="journal article" year="1982">Compagno and Roberts (1982</bibRefCitation>
, 
<bibRefCitation id="EFDD79E6A003FF95FD29FB9AFCD1FBD9" author="Compagno" box="[757,820,1028,1054]" pageId="8" pageNumber="11" refString="Compagno, L. J. V. &amp; Roberts, T. R. (1984) Marine and freshwater stingrays (Dasyatidae) of West Africa, with description of a new species. Proceedings of California Academy of Sciences, 43 (18), 283 - 300." type="journal article" year="1984">1984</bibRefCitation>
), 
<bibRefCitation id="EFDD79E6A003FF95FC90FB9AFB7DFBD9" author="Nishida" box="[844,1176,1028,1054]" pageId="8" pageNumber="12" refString="Nishida, K. &amp; Nakaya, K. (1990) Taxonomy of the genus Dasyatis (Elasmobranchii, Dasyatididae) from the North Pacific. In. H. L. Pratts, Taniuchi, K and S. H. Gruber (Ed) Elasmobranchs as living resources; advances in the biology, ecology, systematics and the status of the fisheries. United States Department of Commerce. National Oceanic and Atmospheric Administration, Report 90, National Marine Fisheries Service, 327 - 346" type="book chapter" year="1990">Nishida and Nakaya (1990)</bibRefCitation>
and 
<bibRefCitation id="EFDD79E6A003FF95FB0BFB9AFE4FFB81" author="Rosenberger" pageId="8" pageNumber="12" refString="Rosenberger, L. J. (2001) Phylogenetic relationships within the stingray genus Dasyatis (Chondrichthyes: Dasyatidae). Copeia, 2001 (3), 615 - 627." type="journal article" year="2001">Rosenberger (2001)</bibRefCitation>
. According to 
<bibRefCitation id="EFDD79E6A003FF95FD88FBB2FCDAFB81" author="Rosenberger" box="[596,831,1068,1094]" pageId="8" pageNumber="12" refString="Rosenberger, L. J. (2001) Phylogenetic relationships within the stingray genus Dasyatis (Chondrichthyes: Dasyatidae). Copeia, 2001 (3), 615 - 627." type="journal article" year="2001">Rosenberger (2001)</bibRefCitation>
, in the western Atlantic this species group is represented by 
<taxonomicName id="4C4C7F94A003FF95FE39FBCAFD58FBAA" ID-CoL="347BB" box="[485,701,1108,1133]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="geijskesi">
<emphasis id="B938D805A003FF95FE39FBCAFD58FBAA" box="[485,701,1108,1133]" italics="true" pageId="8" pageNumber="9">Dasyatis geijskesi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C7F94A003FF95FD0CFBCAFCA9FBAA" ID-CoL="347BM" box="[720,844,1108,1134]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A003FF95FD0CFBCAFD00FBAA" box="[720,741,1108,1133]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FD25FBCAFCA9FBAA" box="[761,844,1108,1133]" italics="true" pageId="8" pageNumber="9">guttata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C4C7F94A003FF95FC4AFBCAFBE8FBAA" ID-CoL="347D8" box="[918,1037,1108,1134]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FC4AFBCAFC4EFBAA" box="[918,939,1108,1133]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FC63FBCAFBE8FBAA" box="[959,1037,1108,1133]" italics="true" pageId="8" pageNumber="9">sabina</emphasis>
</taxonomicName>
, which have preorbital snout lengths exceeding 25% of DW. The other western Atlantic species of 
<taxonomicName id="4C4C7F94A003FF95FB4BFBE2FB1BFB52" ID-CoL="3ZY5" box="[1175,1278,1148,1173]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938D805A003FF95FB4BFBE2FB1BFB52" box="[1175,1278,1148,1173]" italics="true" pageId="8" pageNumber="9">Dasyatis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C7F94A003FF95FAD2FBE2FE60FB7A" ID-CoL="6CC4W" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="americana">
<emphasis id="B938D805A003FF95FAD2FBE2FAC6FB52" box="[1294,1315,1148,1173]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FED4FB3AFE60FB7A" box="[264,389,1188,1213]" italics="true" pageId="8" pageNumber="9">americana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C7F94A003FF95FE4BFB3AFDD4FB7A" ID-CoL="6CBT3" box="[407,561,1188,1214]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="centroura">
<emphasis id="B938D805A003FF95FE4BFB3AFE49FB7A" box="[407,428,1188,1213]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FE63FB3AFDD4FB7A" box="[447,561,1188,1213]" italics="true" pageId="8" pageNumber="9">centroura</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C7F94A003FF95FD9FFB3AFD3FFB7A" ID-CoL="347CN" box="[579,730,1188,1214]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="marianae">
<emphasis id="B938D805A003FF95FD9FFB3AFDBDFB7A" box="[579,600,1188,1213]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FDB6FB3AFD3FFB7A" box="[618,730,1188,1213]" italics="true" pageId="8" pageNumber="9">marianae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C4C7F94A003FF95FCFDFB3AFC8AFB7A" ID-CoL="6CC56" box="[801,879,1188,1214]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="say">
<emphasis id="B938D805A003FF95FCFDFB3AFCD3FB7A" box="[801,822,1188,1213]" italics="true" pageId="8" pageNumber="9">D</emphasis>
. 
<emphasis id="B938D805A003FF95FC94FB3AFC8AFB7A" box="[840,879,1188,1213]" italics="true" pageId="8" pageNumber="9">say</emphasis>
</taxonomicName>
either have or lack diamond shaped discs but all have snouts that are less than 25% of disc width.
</paragraph>
<paragraph id="8BF30417A003FF95FEE4FB6AFDACF9B1" blockId="8.[264,1324,953,1894]" pageId="8" pageNumber="9">
The shape of the disc, which is laterally rounded, promptly differentiates 
<taxonomicName id="4C4C7F94A003FF95FB69FB6AFACEFACA" ID-CoL="347D8" box="[1205,1323,1268,1293]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FB69FB6AFACEFACA" box="[1205,1323,1268,1293]" italics="true" pageId="8" pageNumber="9">D. sabina</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C4C7F94A003FF95FE96FA82FE03FAF2" ID-CoL="6CBT4" box="[330,486,1308,1333]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FE96FA82FE03FAF2" box="[330,486,1308,1333]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
, which has angular pectoral fin edges. In 
<taxonomicName id="4C4C7F94A003FF95FC06FA82FBA8FAF2" ID-CoL="347D8" box="[986,1101,1308,1333]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FC06FA82FBA8FAF2" box="[986,1101,1308,1333]" italics="true" pageId="8" pageNumber="9">D. sabina</emphasis>
</taxonomicName>
the horizontal eye diameter is more developed (53% to 88% of spiracle length), whereas in 
<taxonomicName id="4C4C7F94A003FF95FB82FADAFB1FFA9A" ID-CoL="6CBT4" box="[1118,1274,1348,1373]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FB82FADAFB1FFA9A" box="[1118,1274,1348,1373]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
this proportion is 35.5 to 47.0 %. In 
<taxonomicName id="4C4C7F94A003FF95FD58FAF2FD12FA42" ID-CoL="347D8" box="[644,759,1388,1413]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FD58FAF2FD12FA42" box="[644,759,1388,1413]" italics="true" pageId="8" pageNumber="9">D. sabina</emphasis>
</taxonomicName>
the upper surface of tail has a definite and high finfold, and the ventral finfold is larger than the caudal height. In 
<taxonomicName id="4C4C7F94A003FF95FBCFFA0AFB4AFA6A" ID-CoL="6CBT4" box="[1043,1199,1428,1453]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FBCFFA0AFB4AFA6A" box="[1043,1199,1428,1453]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
the dorsal finfold is absent (
<typeStatus id="54F7BAB5A003FF95FE01FA22FDA2FA11" box="[477,583,1468,1494]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
) or vestigial (
<typeStatus id="54F7BAB5A003FF95FD2EFA22FC83FA11" box="[754,870,1468,1494]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
) and the height of the ventral finfold does not reach one­half of tail. The snout of 
<taxonomicName id="4C4C7F94A003FF95FCD1FA7AFC9AFA3A" ID-CoL="347D8" box="[781,895,1508,1533]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FCD1FA7AFC9AFA3A" box="[781,895,1508,1533]" italics="true" pageId="8" pageNumber="9">D. sabina</emphasis>
</taxonomicName>
is shorter than in 
<taxonomicName id="4C4C7F94A003FF95FB8CFA7AFB0FFA3A" ID-CoL="6CBT4" box="[1104,1258,1508,1533]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FB8CFA7AFB0FFA3A" box="[1104,1258,1508,1533]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
; horizontal eye diameter is 5.6 to 6.9% of preoral length in 
<taxonomicName id="4C4C7F94A003FF95FC63F992FBBAF9E2" ID-CoL="6CBT4" box="[959,1119,1548,1573]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FC63F992FBBAF9E2" box="[959,1119,1548,1573]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
, and the preoral length is 34.8 to 36.7 % of DW. In 
<taxonomicName id="4C4C7F94A003FF95FD6DF9AAFCC1F98A" ID-CoL="347D8" box="[689,804,1588,1613]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sabina">
<emphasis id="B938D805A003FF95FD6DF9AAFCC1F98A" box="[689,804,1588,1613]" italics="true" pageId="8" pageNumber="9">D. sabina</emphasis>
</taxonomicName>
these percentages are, respectively, 14% to 22% and 27. 7 % to 26.4%.
</paragraph>
</subSubSection>
<subSubSection id="C356579CA003FF94FEE4F91AFCCAFA79" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF30417A003FF94FEE4F91AFBEFFE41" blockId="8.[264,1324,953,1894]" lastBlockId="9.[264,1325,284,1470]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<taxonomicName id="4C4C7F94A003FF95FEE4F91AFE37F95A" box="[312,466,1668,1693]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FEE4F91AFE37F95A" box="[312,466,1668,1693]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C4C7F94A003FF95FD89F91AFD20F95A" box="[597,709,1668,1693]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A003FF95FD89F91AFD20F95A" box="[597,709,1668,1693]" italics="true" pageId="8" pageNumber="9">D guttata</emphasis>
</taxonomicName>
in possessing a rough patch of small tubercles at the midline dorsal area and on the elongated snout. The dorsal patch of tubercles of the disc in mature 
<taxonomicName id="4C4C7F94A003FF95FEB8F94AFE3AF92A" box="[356,479,1748,1773]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A003FF95FEB8F94AFE3AF92A" box="[356,479,1748,1773]" italics="true" pageId="8" pageNumber="9">D. guttata</emphasis>
</taxonomicName>
forms a more or less defined band, while in 
<taxonomicName id="4C4C7F94A003FF95FBD7F94AFB4CF92A" box="[1035,1193,1748,1773]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FBD7F94AFB4CF92A" box="[1035,1193,1748,1773]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
the dorsal tubercles present a random arrangement. The snout of 
<taxonomicName id="4C4C7F94A003FF95FC58F962FC1FF8D2" box="[900,1018,1788,1813]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A003FF95FC58F962FC1FF8D2" box="[900,1018,1788,1813]" italics="true" pageId="8" pageNumber="9">D. guttata</emphasis>
</taxonomicName>
is shorter than in 
<taxonomicName id="4C4C7F94A003FF95FB17F962FEA9F8FA" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FB17F962FEA9F8FA" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
. The horizontal eye diameter is 10% to 39% in preoral length in 
<taxonomicName id="4C4C7F94A003FF95FB90F8BAFB26F8FA" box="[1100,1219,1828,1853]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A003FF95FB90F8BAFB26F8FA" box="[1100,1219,1828,1853]" italics="true" pageId="8" pageNumber="9">D. guttata</emphasis>
</taxonomicName>
whereas in 
<taxonomicName id="4C4C7F94A003FF95FEFAF8D2FE24F8A2" box="[294,449,1868,1893]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A003FF95FEFAF8D2FE24F8A2" box="[294,449,1868,1893]" italics="true" pageId="8" pageNumber="9">D. colarensis</emphasis>
</taxonomicName>
it is 5.6 to 6.9 %. In addition, the preoral length is 21.5 to 25.5% of DW in 
<taxonomicName id="4C4C7F94A002FF94FED4FE82FE9BFEF2" box="[264,382,284,309]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A002FF94FED4FE82FE9BFEF2" box="[264,382,284,309]" italics="true" pageId="9" pageNumber="10">D. guttata</emphasis>
</taxonomicName>
, and 34.9% to 36.7% in 
<taxonomicName id="4C4C7F94A002FF94FD42FE82FCDCFEF2" box="[670,825,284,309]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FD42FE82FCDCFEF2" box="[670,825,284,309]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
. The posterior margins of pectoral fins are perfectly rounded in 
<taxonomicName id="4C4C7F94A002FF94FE23FEDAFD79FE9A" box="[511,668,324,349]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FE23FEDAFD79FE9A" box="[511,668,324,349]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
and angular in 
<taxonomicName id="4C4C7F94A002FF94FC88FEDAFC2EFE9A" box="[852,971,324,349]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A002FF94FC88FEDAFC2EFE9A" box="[852,971,324,349]" italics="true" pageId="9" pageNumber="10">D. guttata</emphasis>
</taxonomicName>
. The posterior margins of the pelvic fins are rounded in 
<taxonomicName id="4C4C7F94A002FF94FDE7FEF2FD57FE42" box="[571,690,364,389]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B938D805A002FF94FDE7FEF2FD57FE42" box="[571,690,364,389]" italics="true" pageId="9" pageNumber="10">D. guttata</emphasis>
</taxonomicName>
and pointed in 
<taxonomicName id="4C4C7F94A002FF94FCB4FEF2FBE6FE42" box="[872,1027,364,389]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FCB4FEF2FBE6FE42" box="[872,1027,364,389]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FE0AFDE6FD89" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
<taxonomicName id="4C4C7F94A002FF94FEE4FE0AFDEFFE6A" box="[312,522,404,429]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geijskesi">
<emphasis id="B938D805A002FF94FEE4FE0AFDEFFE6A" box="[312,522,404,429]" italics="true" pageId="9" pageNumber="10">Dasyatis geijskesi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C4C7F94A002FF94FD78FE0AFCA5FE6A" box="[676,832,404,429]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FD78FE0AFCA5FE6A" box="[676,832,404,429]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
in possessing a very long preorbital snout length (40 to 50% of DW vs. 35.7 to 37.7% in 
<taxonomicName id="4C4C7F94A002FF94FC9CFE22FC09FE12" box="[832,1004,444,469]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="coloarensis">
<emphasis id="B938D805A002FF94FC9CFE22FC09FE12" box="[832,1004,444,469]" italics="true" pageId="9" pageNumber="10">D. coloarensis</emphasis>
</taxonomicName>
), very long prenasal snout length (37 to 45% of DW vs. 29.5 to 32.7%), very broad pelvic fins (38 to 42.9% of DW vs. 19.6 to 23.3%), and narrow pelvic fin base length (79.3 to 91.1% of internasal length vs. 119.2 to 124.2%).
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FDC2FC3CFCF9" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
With respect to 
<taxonomicName id="4C4C7F94A002FF94FE2DFDC2FDBDFDB2" box="[497,600,604,629]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938D805A002FF94FE2DFDC2FDBDFDB2" box="[497,600,604,629]" italics="true" pageId="9" pageNumber="10">Dasyatis</emphasis>
</taxonomicName>
from other regions, 
<taxonomicName id="4C4C7F94A002FF94FC94FDC2FC01FDB2" box="[840,996,604,629]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FC94FDC2FC01FDB2" box="[840,996,604,629]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C4C7F94A002FF94FBB4FDC2FE25FD59" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zugei">
<emphasis id="B938D805A002FF94FBB4FDC2FB2FFDB2" box="[1128,1226,604,629]" italics="true" pageId="9" pageNumber="10">D. zugei</emphasis>
(Müller &amp; Henle, 1841)
</taxonomicName>
from 
<collectingCountry id="F35B4487A002FF94FDDBFD1AFDB5FD59" box="[519,592,644,670]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
, 
<collectingCountry id="F35B4487A002FF94FD80FD1AFD26FD59" box="[604,707,644,670]" name="Pakistan" pageId="9" pageNumber="10">Pakistan</collectingCountry>
, 
<collectingCountry id="F35B4487A002FF94FD12FD1AFCA5FD59" box="[718,832,644,670]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
and the Sea of 
<collectingCountry id="F35B4487A002FF94FC29FD1AFBDEFD59" box="[1013,1083,644,670]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
, 
<taxonomicName id="4C4C7F94A002FF94FB99FD1AFE59FD01" authority="Steindachner, 1892" authorityName="Steindachner" authorityYear="1892" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="navarrae">
<emphasis id="B938D805A002FF94FB99FD1AFB30FD5A" box="[1093,1237,644,669]" italics="true" pageId="9" pageNumber="10">D. navarrae</emphasis>
(Steindachner, 1892)
</taxonomicName>
and 
<taxonomicName id="4C4C7F94A002FF94FDDDFD32FC25FD01" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[513,960,684,710]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bennetti">
<emphasis id="B938D805A002FF94FDDDFD32FD6CFD02" box="[513,649,684,709]" italics="true" pageId="9" pageNumber="10">D. bennetti</emphasis>
(Müller &amp; Henle, 1841)
</taxonomicName>
from southeast Asia, and 
<taxonomicName id="4C4C7F94A002FF94FAD2FD32FD3BFD29" authority="Nishida and Nakaya, 1988" authorityName="Nishida and Nakaya" authorityYear="1988" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="acutirostra">
<emphasis id="B938D805A002FF94FAD2FD32FE6CFD2A" italics="true" pageId="9" pageNumber="10">D. acutirostra</emphasis>
(Nishida and Nakaya, 1988)
</taxonomicName>
from the Sea of 
<collectingCountry id="F35B4487A002FF94FC74FD4AFC0EFD29" box="[936,1003,724,750]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
and the East 
<collectingCountry id="F35B4487A002FF94FB50FD4AFB37FD29" box="[1164,1234,724,750]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
Sea, in having an elongated snout which reaches more than 25% of disc length (
<bibRefCitation id="EFDD79E6A002FF94FBBDFD62FB08FCD1" author="Chen" box="[1121,1261,764,790]" pageId="9" pageNumber="11" refString="Chen, J. T. F. (1948) Notes on the fish-fauna of Taiwan in the collections of the Taiwan Musem. I. Some records of Platosomae from Taiwan, with description of a new species of Dasyatis. Quarterly Journal of the Taiwan Museum, 1 (3), 1 - 12." type="journal article" year="1948">Chen, 1948</bibRefCitation>
; 
<bibRefCitation id="EFDD79E6A002FF94FB26FD62FE4DFCF9" author="Chu" pageId="9" pageNumber="11" refString="Chu, Y. T &amp; Wen, M. C. (1982) A Study of the lateral-line canals system and that of Lorenzini ampullae and tubules of elasmobranchiate fishes of China. In: Monography of Fishes of China. Vo l 2., Science and Technology Press, Shangai, pp. 117 - 126." type="book chapter" year="1982">Chu &amp; Wen, 1982</bibRefCitation>
; 
<bibRefCitation id="EFDD79E6A002FF94FE68FCBAFD7AFCF9" author="Monkolprasit" box="[436,671,804,830]" pageId="9" pageNumber="12" refString="Monkolprasit, S. (1984) The Cartilaginous Fishes (Class Elasmobranchii) Found in Thai Waters and Adjacent Areas. Kasetsart University, Bangkok, 175 pp." type="book" year="1984">Monkolprasit, 1984</bibRefCitation>
; 
<bibRefCitation id="EFDD79E6A002FF94FD77FCBAFC28FCF9" author="Nishida" box="[683,973,804,830]" pageId="9" pageNumber="12" refString="Nishida, K. &amp; Nakaya, K. (1990) Taxonomy of the genus Dasyatis (Elasmobranchii, Dasyatididae) from the North Pacific. In. H. L. Pratts, Taniuchi, K and S. H. Gruber (Ed) Elasmobranchs as living resources; advances in the biology, ecology, systematics and the status of the fisheries. United States Department of Commerce. National Oceanic and Atmospheric Administration, Report 90, National Marine Fisheries Service, 327 - 346" type="book chapter" year="1990">Nishida &amp; Nakaya,1990</bibRefCitation>
).
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FCD2FDD8FC71" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
The pale­edged stingray 
<taxonomicName id="4C4C7F94A002FF94FD86FCD2FD59FCA2" box="[602,700,844,869]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zugei">
<emphasis id="B938D805A002FF94FD86FCD2FD59FCA2" box="[602,700,844,869]" italics="true" pageId="9" pageNumber="10">D. zugei</emphasis>
</taxonomicName>
and the blackish stingray 
<taxonomicName id="4C4C7F94A002FF94FC33FCD2FB9AFCA2" box="[1007,1151,844,869]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="navarrae">
<emphasis id="B938D805A002FF94FC33FCD2FB9AFCA2" box="[1007,1151,844,869]" italics="true" pageId="9" pageNumber="10">D. navarrae</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="4C4C7F94A002FF94FAD2FCD2FE9AFC4A" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FAD2FCD2FE9AFC4A" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
by presenting a well developed upper finfold in the former, and vestigial or absent finfold in the latter.
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FC5AFCD8FBB9" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
The frilltail stingray 
<taxonomicName id="4C4C7F94A002FF94FDEBFC5AFD58FC1A" box="[567,701,964,989]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bennetti">
<emphasis id="B938D805A002FF94FDEBFC5AFD58FC1A" box="[567,701,964,989]" italics="true" pageId="9" pageNumber="10">D. bennetti</emphasis>
</taxonomicName>
lacks a patch of tubercles on the back of the disc (present in 
<taxonomicName id="4C4C7F94A002FF94FE52FC72FDCFFBC2" box="[398,554,1004,1029]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FE52FC72FDCFFBC2" box="[398,554,1004,1029]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), its ventral finfold length is 60% to 67% of DW (37.3 to 51.5% in 
<taxonomicName id="4C4C7F94A002FF94FEFBFB8AFE27FBEA" box="[295,450,1044,1069]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FEFBFB8AFE27FBEA" box="[295,450,1044,1069]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), its preoral length ranges from 25.7% to 27.4% of DW (34.8 to 36.7% in 
<taxonomicName id="4C4C7F94A002FF94FED4FBA2FE40FB92" box="[264,421,1084,1109]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FED4FBA2FE40FB92" box="[264,421,1084,1109]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), its pelvic radials range from 21 to 24 (
<quantity id="4CB4A9F2A002FF94FC5BFBA2FC20FB91" box="[903,965,1084,1110]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="9" pageNumber="10" unit="in" value="26.0">26 in</quantity>
<taxonomicName id="4C4C7F94A002FF94FC12FBA2FB8FFB92" box="[974,1130,1084,1109]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FC12FBA2FB8FFB92" box="[974,1130,1084,1109]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), and it has 119 pre­sting vertebral centra (
<quantity id="4CB4A9F2A002FF94FDE2FBFAFD6CFBB9" box="[574,649,1124,1150]" metricMagnitude="0" metricUnit="m" metricValue="3.2257999999999996" pageId="9" pageNumber="10" unit="in" value="127.0">127 in</quantity>
<taxonomicName id="4C4C7F94A002FF94FD4DFBFAFCC8FBBA" box="[657,813,1124,1149]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FD4DFBFAFCC8FBBA" box="[657,813,1124,1149]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FB12FC2DFB31" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
The smalltooth stingray 
<taxonomicName id="4C4C7F94A002FF94FD89FB12FD50FB62" box="[597,693,1164,1190]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rudis">
<emphasis id="B938D805A002FF94FD89FB12FD8FFB62" box="[597,618,1164,1189]" italics="true" pageId="9" pageNumber="10">D</emphasis>
. 
<emphasis id="B938D805A002FF94FDA5FB12FD50FB62" box="[633,693,1164,1189]" italics="true" pageId="9" pageNumber="10">rudis</emphasis>
</taxonomicName>
possesses the entire dorsal surface of disc rough with minute and dense asperities and no middorsal row of spines (
<taxonomicName id="4C4C7F94A002FF94FC02FB2AFB9EFB0A" box="[990,1147,1204,1229]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FC02FB2AFB9EFB0A" box="[990,1147,1204,1229]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
presents small tubercles only at disc centre and a middorsal row of spines).
</paragraph>
<paragraph id="8BF30417A002FF94FEE4FA9AFCCAFA79" blockId="9.[264,1325,284,1470]" pageId="9" pageNumber="10">
The sharpnose stingray 
<taxonomicName id="4C4C7F94A002FF94FD8AFA9AFD18FADA" box="[598,765,1284,1309]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="acutirostra">
<emphasis id="B938D805A002FF94FD8AFA9AFD18FADA" box="[598,765,1284,1309]" italics="true" pageId="9" pageNumber="10">D. acutirostra</emphasis>
</taxonomicName>
presents a ventral finfold length ranging from 15.0% to 22.9% of DW (37.3 to 51.5% in 
<taxonomicName id="4C4C7F94A002FF94FCD3FAB2FC49FA82" box="[783,940,1324,1349]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FCD3FAB2FC49FA82" box="[783,940,1324,1349]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), a preoral length ranging from 37.9% to 44.2% of DW (34.8 to 36.7% in 
<taxonomicName id="4C4C7F94A002FF94FD20FACAFC7DFAAA" box="[764,920,1364,1389]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FD20FACAFC7DFAAA" box="[764,920,1364,1389]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), lacks a dorsal patch of tubercles and suprascapular thorns (both present in 
<taxonomicName id="4C4C7F94A002FF94FD20FAE2FC7DFA52" box="[764,920,1404,1429]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FD20FAE2FC7DFA52" box="[764,920,1404,1429]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
), and lacks oral papillae (three to four well developed papillae in 
<taxonomicName id="4C4C7F94A002FF94FDA0FA3AFCFBFA7A" box="[636,798,1444,1469]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="colarensis">
<emphasis id="B938D805A002FF94FDA0FA3AFCFBFA7A" box="[636,798,1444,1469]" italics="true" pageId="9" pageNumber="10">D. colarensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>