818 lines
97 KiB
XML
818 lines
97 KiB
XML
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<mods:title id="A041C2183DA4BAD6918A4843D2459EFC">Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history</mods:title>
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<mods:namePart id="7E08B86F0D16DE42A97D4E761689AFC3">Lobón-Rovira, Javier</mods:namePart>
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<mods:namePart id="05F4A7208A430BFE8FDE58A7B112020D">Conradie, Werner</mods:namePart>
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<mods:namePart id="35A6D0D7E7C3FB482F9923FB980DE767">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="42556DF0173A853C7F025A38706B6F7C">Júlio, Timóteo</mods:namePart>
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<mods:namePart id="EAE38E89E08A4F389927FAB2E03852DD">Stanley, Edward L.</mods:namePart>
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<mods:namePart id="EBA4B1CDA55FBCA485B630210094DA2F">Behangana, Mathias</mods:namePart>
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<mods:namePart id="C6FEE7819F4B67F5FF4EF5BDC6EF0D1A">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="3C60CFD576A8ACCAD778A21FBF1DBE60">Greenbaum, Eli</mods:namePart>
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<mods:date id="CE8D2E37E5B4241AA7C06FF8278E7FD2">2024</mods:date>
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<treatment id="03B30955FFF3FFD9FF3444C6FC56F8A3" ID-DOI="http://doi.org/10.5281/zenodo.13302903" ID-Zenodo-Dep="13302903" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3" httpUri="http://treatment.plazi.org/id/03B30955FFF3FFD9FF3444C6FC56F8A3" lastPageId="38" lastPageNumber="485" pageId="36" pageNumber="483">
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<taxonomicName id="4C1AC3C0FFF3FFDBFF3444C6FDBCFB62" ID-CoL="C7JCT" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[246,560,1212,1238]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="485" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
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<emphasis id="B96E6451FFF3FFDBFF3444C6FDBCFB62" bold="true" box="[246,560,1212,1238]" italics="true" pageId="36" pageNumber="483">LYGODACTYLUS GAMBLEI</emphasis>
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<taxonomicNameLabel id="A25DD92AFFF3FFDBFDF644C6FD18FB62" box="[564,660,1212,1238]" pageId="36" pageNumber="483" rank="species">SP.NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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<subSubSection id="C300EBC8FFF3FFDBFF5C4499FD66FA39" pageId="36" pageNumber="483" type="description">
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<paragraph id="8BA5B843FFF3FFDBFF5C4499FD60FB4F" blockId="36.[158,748,1251,1275]" box="[158,748,1251,1275]" pageId="36" pageNumber="483">
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(
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<figureCitation id="1321A4C6FFF3FFDBFF6B4499FE95FB4F" box="[169,281,1251,1275]" captionStart-0="Figure 23" captionStart-1="Figure 24" captionStartId-0="37.[113,178,1204,1228]" captionStartId-1="38.[129,194,1232,1256]" captionTargetBox-0="[114,1458,144,1176]" captionTargetBox-1="[129,1473,144,1205]" captionTargetId-0="figure-366@37.[114,1458,144,1176]" captionTargetId-1="figure-368@38.[129,1473,144,1205]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." captionText-1="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi-0="http://doi.org/10.5281/zenodo.13302917" figureDoi-1="http://doi.org/10.5281/zenodo.13302919" httpUri-0="https://zenodo.org/record/13302917/files/figure.png" httpUri-1="https://zenodo.org/record/13302919/files/figure.png" pageId="36" pageNumber="483">Figs 23–24</figureCitation>
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,
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<tableCitation id="C6988DF8FFF3FFDBFEE64499FEE3FB4F" box="[292,367,1251,1275]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="36" pageNumber="483" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
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; Supporting Information, Table S10)
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</paragraph>
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<paragraph id="8BA5B843FFF3FFDBFF434566FE38FAE7" blockId="36.[129,777,1308,1363]" pageId="36" pageNumber="483">
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<emphasis id="B96E6451FFF3FFDBFF434566FED2FA80" box="[129,350,1308,1332]" italics="true" pageId="36" pageNumber="483">Zoobank registration:</emphasis>
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https://zoobank.org/
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<uuid id="FFBC8296FFF3FFDBFD8E4566FE38FAE7" pageId="36" pageNumber="483">411D833D-2017- 4930-BD11-B783AAD6A9B7</uuid>
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</paragraph>
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<paragraph id="8BA5B843FFF3FFDBFF634509FD66FA39" blockId="36.[161,746,1395,1421]" box="[161,746,1395,1421]" pageId="36" pageNumber="483">
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<emphasis id="B96E6451FFF3FFDBFF634509FD66FA39" box="[161,746,1395,1421]" italics="true" pageId="36" pageNumber="483">
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<taxonomicName id="4C1AC3C0FFF3FFDBFF634509FE73FA39" authorityName="' (Schmidt" authorityYear="1919" box="[161,511,1395,1421]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">Lygodactylus picturatus gutturalis</taxonomicName>
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: de Witte (1953) [part].
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</emphasis>
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</paragraph>
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</subSubSection>
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<paragraph id="8BA5B843FFF3FFDBFF5B45CBFD1AFA5F" blockId="36.[129,778,1457,1677]" pageId="36" pageNumber="483">
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<emphasis id="B96E6451FFF3FFDBFF5B45CBFD1AFA5F" italics="true" pageId="36" pageNumber="483">
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<taxonomicName id="4C1AC3C0FFF3FFDBFF5B45CBFE05FA7F" ID-CoL="3WQ3T" authority="(Bocage, 1873)" box="[153,393,1457,1483]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">Lygodactylus gutturalis</taxonomicName>
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:
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<bibRefCitation id="EF8BC5B2FFF3FFDBFE5745C8FDABFA7F" author="Pasteur G" box="[405,551,1457,1483]" pageId="36" pageNumber="483" pagination="1 - 132" refId="ref36328" refString="Pasteur G. Recherches sur l'evolution des lygodactyles, lezards Afro- Malgaches actuels. Travaux de l'Institut Scientifique Cherifien, Zoologie 1965 (1964); 29: 1 - 132." type="journal article" year="1964">Pasteur (1964)</bibRefCitation>
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[part]; Broadley and Cotterill (2004) [part];
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<bibRefCitation id="EF8BC5B2FFF3FFDBFE1E45ABFDDDFA5F" author="Roll B" box="[476,593,1489,1515]" pageId="36" pageNumber="483" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
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[part].
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</emphasis>
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</paragraph>
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|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFF43458DFEE6F9BB" ID-CoL="C7JCT" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[129,362,1527,1551]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFF43458DFEE6F9BB" bold="true" box="[129,362,1527,1551]" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFEB7458DFE41F9BB" box="[373,461,1527,1551]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a miombo-savannah form within subgroup C, described above.It also differs by a minimum of
|
||
<emphasis id="B96E6451FFF3FFDBFF5F464DFF2AF9FA" box="[157,166,1591,1614]" italics="true" pageId="36" pageNumber="483">c</emphasis>
|
||
. 9.96% for the
|
||
<emphasis id="B96E6451FFF3FFDBFE88464CFEFDF9FA" box="[330,369,1590,1614]" italics="true" pageId="36" pageNumber="483">16S</emphasis>
|
||
mitochondrial gene (Table 2) from its sister taxon
|
||
<emphasis id="B96E6451FFF3FFDBFF3E462CFE61F9D9" bold="true" box="[252,493,1621,1645]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFF3E462CFE19F9D9" ID-CoL="C7JCY" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[252,405,1621,1645]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFF3E462CFE19F9D9" bold="true" box="[252,405,1621,1645]" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFE58462FFE61F9D9" box="[410,493,1621,1645]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF3FFDBFE3F462FFDC7F9D9" box="[509,587,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="36" pageNumber="483">Fig. 2A</figureCitation>
|
||
) and lacks nuclear haplotype sharing in
|
||
<emphasis id="B96E6451FFF3FFDBFE94460FFE1FF939" box="[342,403,1653,1677]" italics="true" pageId="36" pageNumber="483">RAG1</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF3FFDBFE61460FFD87F939" box="[419,523,1653,1677]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="36" pageNumber="483">Fig. 2B–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300EBC8FFF3FFD9FF4346D4FC56F8A3" lastPageId="38" lastPageNumber="485" pageId="36" pageNumber="483" type="materials_examined">
|
||
<paragraph id="8BA5B843FFF3FFDBFF4346D4FE72F890" blockId="36.[129,778,1710,1828]" pageId="36" pageNumber="483">
|
||
<materialsCitation id="3B72B21EFFF3FFDBFF4346D4FE72F890" collectingDate="2010-01-19" collectionCode="UTEP, ELI" country="Democratic Republic of the Congo" elevation="634" latitude="-7.29363" location="Manono" longLatPrecision="1" longitude="27.39472" municipality="Tanganyika Province" pageId="36" pageNumber="483" specimenCode="UTEP 22587, ELI 340" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B96E6451FFF3FFDBFF4346D4FF6DF972" box="[129,225,1710,1734]" italics="true" pageId="36" pageNumber="483">
|
||
<typeStatus id="54A106E1FFF3FFDBFF4346D4FF50F972" box="[129,220,1710,1734]" pageId="36" pageNumber="483" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DBBC1038FFF3FFDBFF3246D4FEF0F972" box="[240,380,1710,1734]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="36" pageNumber="483" type="Museum">UTEP 22587</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFF3FFDBFE4846D4FE64F972" box="[394,488,1710,1734]" collectionCode="ELI" pageId="36" pageNumber="483">ELI 340</specimenCode>
|
||
), male with regenerated tail and ventral incision, collected at
|
||
<location id="8EC5EE98FFF3FFDBFE0F46B4FDAAF951" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFE0F46B4FDAAF951" box="[461,550,1742,1765]" country="Democratic Republic of the Congo" latitude="-7.29363" longLatPrecision="1" longitude="27.39472" municipality="Tanganyika Province" name="Manono" pageId="36" pageNumber="483">Manono</location>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFDEC46B7FC89F951" box="[558,773,1741,1765]" pageId="36" pageNumber="483">Tanganyika Province</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFF434697FF35F8B1" box="[129,185,1773,1797]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFF3FFDBFF024697FEBCF8B1" box="[192,304,1773,1797]" degrees="07.29363" direction="south" orientation="latitude" pageId="36" pageNumber="483" precision="1" value="-7.29363">S07.29363</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFF3FFDBFEFA4697FE26F8B1" box="[312,426,1773,1797]" degrees="27.39472" direction="east" orientation="longitude" pageId="36" pageNumber="483" precision="1" value="27.39472">E27.39472</geoCoordinate>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFE704697FDAFF8B1" box="[434,547,1773,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.34" pageId="36" pageNumber="483" unit="m" value="634.0">
|
||
<quantity id="4CE215A6FFF3FFDBFE704697FE7FF8B0" box="[434,499,1773,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.34" pageId="36" pageNumber="483" unit="m" value="634.0">634 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFD854697FD60F8B1" box="[583,748,1773,1797]" pageId="36" pageNumber="483" value="2010-01-19">
|
||
<collectingDate id="EFE0676BFFF3FFDBFD854697FD60F8B1" box="[583,748,1773,1797]" pageId="36" pageNumber="483" value="2010-01-19">19 January 2010</collectingDate>
|
||
</date>
|
||
by locals and brought to Eli Greenbaum.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF3FFDBFF43473CFB58FCED" blockId="36.[129,778,1861,1980]" lastBlockId="36.[825,1475,144,858]" pageId="36" pageNumber="483">
|
||
<emphasis id="B96E6451FFF3FFDBFF43473CFE1BF8E9" box="[129,407,1862,1885]" italics="true" pageId="36" pageNumber="483">
|
||
<typeStatus id="54A106E1FFF3FFDBFF43473CFF6CF8E9" box="[129,224,1862,1885]" pageId="36" pageNumber="483" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D1C73CAFFF3FFDBFF30473CFE07F8E9" box="[242,395,1862,1885]" count="9" pageId="36" pageNumber="483" type="generic">nine specimens</specimenCount>
|
||
):
|
||
</emphasis>
|
||
<materialsCitation id="3B72B21EFFF3FFDBFE6F473CFAE1FF1C" collectingDate="2010-01-18" collectionCode="UTEP, ELI" collectorName="Mulongo" country="Democratic Republic of the Congo" elevation="875" latitude="-7.65509" location="Haut-Lomami Province" longLatPrecision="1" longitude="27.34027" municipality="Aristote" pageId="36" pageNumber="483" specimenCode="UTEP 22584, ELI 293" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFE6F473CFDB0F8E9" box="[429,572,1861,1885]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="36" pageNumber="483" type="Museum">UTEP 22584</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFF3FFDBFD8C473FFD22F8E9" box="[590,686,1861,1885]" collectionCode="ELI" pageId="36" pageNumber="483">ELI 293</specimenCode>
|
||
), female with original tail and ventral incision, collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFD65471FFC8AF8C9" box="[679,774,1893,1917]" pageId="36" pageNumber="483">Mulongo</collectorName>
|
||
,
|
||
<location id="8EC5EE98FFF3FFDBFF4347FEFEF4F828" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFF4347FEFEF4F828" box="[129,376,1924,1948]" country="Democratic Republic of the Congo" latitude="-7.65509" longLatPrecision="1" longitude="27.34027" name="Haut-Lomami Province" pageId="36" pageNumber="483">Haut-Lomami Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFE4747FEFE31F828" box="[389,445,1924,1948]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFF3FFDBFE0847FEFDB6F828" box="[458,570,1924,1948]" degrees="07.65509" direction="south" orientation="latitude" pageId="36" pageNumber="483" precision="1" value="-7.65509">S07.65509</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFF3FFDBFD8447FEFD34F828" box="[582,696,1924,1948]" degrees="27.34027" direction="east" orientation="longitude" pageId="36" pageNumber="483" precision="1" value="27.34027">E27.34027</geoCoordinate>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFD0747FEFF21F80F" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="36" pageNumber="483" unit="m" value="875.0">
|
||
<quantity id="4CE215A6FFF3FFDBFD0747FEFC86F82F" box="[709,778,1924,1948]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="36" pageNumber="483" unit="m" value="875.0">875 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFF1147D9FEF7F80F" box="[211,379,1955,1980]" pageId="36" pageNumber="483" value="2010-01-18">
|
||
<collectingDate id="EFE0676BFFF3FFDBFF1147D9FEF7F80F" box="[211,379,1955,1980]" pageId="36" pageNumber="483" value="2010-01-18">18 January 2010</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFF3FFDBFE6247D9FDF2F80F" box="[416,638,1955,1979]" pageId="36" pageNumber="483">Chifundera Kusamba</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFD4A47D9FD6BF80F" box="[648,743,1955,1979]" pageId="36" pageNumber="483">Wandege</collectorName>
|
||
M.
|
||
<collectorName id="26EFDD95FFF3FFDBFCFB40EAFC1BFF1C" box="[825,919,144,168]" pageId="36" pageNumber="483">Muninga</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFC6040EAFB97FF1C" box="[930,1051,144,168]" pageId="36" pageNumber="483">Mwenebatu</collectorName>
|
||
M.
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFB8640EAFB14FF1C" box="[1092,1176,144,168]" pageId="36" pageNumber="483">Aristote</collectingMunicipality>
|
||
, and
|
||
<location id="8EC5EE98FFF3FFDBFB0C40EAFAE1FF1C" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB0C40EAFAE1FF1C" box="[1230,1389,144,168]" country="Democratic Republic of the Congo" municipality="Aristote" name="Eli Greenbaum" pageId="36" pageNumber="483">Eli Greenbaum</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFABA40EAFCFCFEF1" collectingDate="1947-10-04" collectingDateMax="1947-10-08" collectingDateMin="1947-10-04" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="715" location="Haut-Katanga Province" municipality="Upemba National Park" pageId="36" pageNumber="483" specimenCode="RBINS 2721, RBINS 6157" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFABA40EAFCE1FF73" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2721</specimenCode>
|
||
(formerly under
|
||
<specimenCode id="DBBC1038FFF3FFDBFBEB40D5FB3BFF73" box="[1065,1207,175,199]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6157</specimenCode>
|
||
), adult male, collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFCFB40B5FC43FF53" box="[825,975,207,231]" pageId="36" pageNumber="483">Rivière Kande</collectorName>
|
||
, affluent de la rive gauche
|
||
<collectorName id="26EFDD95FFF3FFDBFB2740B5FAE7FF53" box="[1253,1387,207,231]" pageId="36" pageNumber="483">de la Lupiala</collectorName>
|
||
et sousaffluent de la rive droite de la
|
||
<collectorName id="26EFDD95FFF3FFDBFBBD4094FB33FEB2" box="[1151,1215,238,262]" pageId="36" pageNumber="483">Lufira</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFB094094FA33FEB2" box="[1227,1471,238,262]" pageId="36" pageNumber="483">Upemba National Park</collectingMunicipality>
|
||
,
|
||
<location id="8EC5EE98FFF3FFDBFCFB4174FBA5FE91" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFCFB4174FBA5FE91" box="[825,1065,269,294]" country="Democratic Republic of the Congo" municipality="Upemba National Park" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFBF34177FBE5FE91" box="[1073,1129,269,293]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFBB34177FA9BFE91" box="[1137,1303,269,293]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="36" pageNumber="483" unit="m" value="715.0" valueMax="730.0" valueMin="700.0">
|
||
<quantity id="4CE215A6FFF3FFDBFBB34177FB6BFE91" box="[1137,1255,269,293]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="36" pageNumber="483" unit="m" value="715.0" valueMax="730.0" valueMin="700.0">700–730 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="FFA49E83FFF3FFDBFA824177FCFCFEF1" pageId="36" pageNumber="483" value="1947-10-04" valueMax="1947-10-08" valueMin="1947-10-04">
|
||
<collectingDate id="EFE0676BFFF3FFDBFA824177FCFCFEF1" pageId="36" pageNumber="483" value="1947-10-04" valueMax="1947-10-08" valueMin="1947-10-04">4–8 October 1947</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFCBE4157FC41FE17" collectingDate="1947-10-28" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="700" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 2722, RBINS 6160" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFCBE4157FB8EFEF1" box="[892,1026,301,325]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2722</specimenCode>
|
||
(formerly under
|
||
<specimenCode id="DBBC1038FFF3FFDBFB754157FACEFEF0" box="[1207,1346,300,325]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6160</specimenCode>
|
||
), adult male collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFC734136FBDCFED0" box="[945,1104,332,356]" pageId="36" pageNumber="483">Rivière Lukawe</collectorName>
|
||
, affluent de la rive droite
|
||
<collectorName id="26EFDD95FFF3FFDBFA894136FA33FED0" box="[1355,1471,332,356]" pageId="36" pageNumber="483">de la Lufira</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFCFB4111FC37FE37" box="[825,955,363,387]" pageId="36" pageNumber="483">Upemba NP</collectorName>
|
||
,
|
||
<location id="8EC5EE98FFF3FFDBFC044116FB35FE37" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFC044116FB35FE37" box="[966,1209,363,388]" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFB064111FB70FE37" box="[1220,1276,363,387]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFAC54111FAF2FE37" box="[1287,1406,363,387]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">
|
||
<quantity id="4CE215A6FFF3FFDBFAC54111FAC7FE37" box="[1287,1355,363,387]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFA6A4111FC41FE17" pageId="36" pageNumber="483" value="1947-10-28">
|
||
<collectingDate id="EFE0676BFFF3FFDBFA6A4111FC41FE17" pageId="36" pageNumber="483" value="1947-10-28">28 October 1947</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFC1A41F1FB06FE16" box="[984,1162,394,419]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 2723" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFC1A41F1FBD1FE17" box="[984,1117,395,419]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2723</specimenCode>
|
||
and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B72B21EFFF3FFDBFB5241F1FC52FD94" collectingDate="1948-06-03" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="890" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 6124" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFB5241F1FA9AFE16" box="[1168,1302,394,419]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6124</specimenCode>
|
||
(formerly under
|
||
<specimenCode id="DBBC1038FFF3FFDBFCFB41D0FC4DFE76" box="[825,961,426,450]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6124</specimenCode>
|
||
), male and female, respectively, collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFAB541D0FA33FE76" box="[1399,1471,426,450]" pageId="36" pageNumber="483">Munoi</collectorName>
|
||
, bifurcation de la rivière
|
||
<collectorName id="26EFDD95FFF3FFDBFBF741B3FB08FE55" box="[1077,1156,457,481]" pageId="36" pageNumber="483">Lupiala</collectorName>
|
||
, affluent de la rive droite
|
||
<collectorName id="26EFDD95FFF3FFDBFA5241B3FCF5FDB4" pageId="36" pageNumber="483">de la Lufira</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFC414192FB88FDB4" box="[899,1028,488,512]" pageId="36" pageNumber="483">Upemba NP</collectorName>
|
||
,
|
||
<location id="8EC5EE98FFF3FFDBFBCD4193FA8CFDB4" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFBCD4193FA8CFDB4" box="[1039,1280,488,513]" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFAC94193FACFFDB5" box="[1291,1347,489,513]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFA8F4193FA4EFDB4" box="[1357,1474,488,513]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">
|
||
<quantity id="4CE215A6FFF3FFDBFA8F4193FA1CFDB4" box="[1357,1424,489,513]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">890 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFC9E4272FC52FD94" box="[860,990,520,544]" pageId="36" pageNumber="483" value="1948-06-03">
|
||
<collectingDate id="EFE0676BFFF3FFDBFC9E4272FC52FD94" box="[860,990,520,544]" pageId="36" pageNumber="483" value="1948-06-03">3 June 1948</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFC284272FABEFD29" collectingDate="1947-05-28" collectingDateMax="1947-05-31" collectingDateMin="1947-05-28" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="890" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 2725-27, RBINS 6122" specimenCount="12" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFC284272FB1BFD94" box="[1002,1175,520,544]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2725–27</specimenCode>
|
||
(formerly in a series of
|
||
<specimenCount id="9D1C73CAFFF3FFDBFA574272FC28FD8B" count="9" pageId="36" pageNumber="483" type="generic">nine specimens</specimenCount>
|
||
under
|
||
<specimenCode id="DBBC1038FFF3FFDBFC2D425DFBF5FD8B" box="[1007,1145,551,575]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6122</specimenCode>
|
||
),
|
||
<specimenCount id="9D1C73CAFFF3FFDBFB4E4252FB79FD8B" box="[1164,1269,551,575]" count="2" pageId="36" pageNumber="483" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="9D1C73CAFFF3FFDBFAEB4252FA17FD8B" box="[1321,1435,551,575]" count="1" pageId="36" pageNumber="483" type="female">one female</specimenCount>
|
||
, respectively, collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFBE7423CFBE1FDEA" box="[1061,1133,582,606]" pageId="36" pageNumber="483">Munoi</collectorName>
|
||
, bifurcation de la rivière
|
||
<collectorName id="26EFDD95FFF3FFDBFAB2423CFA33FDEA" box="[1392,1471,582,606]" pageId="36" pageNumber="483">Lupiala</collectorName>
|
||
, affluent de la rive droite
|
||
<collectorName id="26EFDD95FFF3FFDBFBEC421CFB2FFDCA" box="[1070,1187,614,638]" pageId="36" pageNumber="483">de la Lufira</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFB69421CFAA6FDC9" box="[1195,1322,614,638]" pageId="36" pageNumber="483">Upemba NP</collectorName>
|
||
,
|
||
<location id="8EC5EE98FFF3FFDBFAF1421CFC19FD29" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAF1421CFC19FD29" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFC6242FFFC54FD29" box="[928,984,645,669]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFC2042FFFBDBFD29" box="[994,1111,645,669]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">
|
||
<quantity id="4CE215A6FFF3FFDBFC2042FFFBA9FD29" box="[994,1061,645,669]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">890 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFBBD42FFFABEFD29" box="[1151,1330,645,670]" pageId="36" pageNumber="483" value="1947-05-28" valueMax="1947-05-31" valueMin="1947-05-28">
|
||
<collectingDate id="EFE0676BFFF3FFDBFBBD42FFFABEFD29" box="[1151,1330,645,670]" pageId="36" pageNumber="483" value="1947-05-28" valueMax="1947-05-31" valueMin="1947-05-28">28–31 May 1947</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFAFF42FFFBBFFCAF" collectingDate="1949-01-31" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="700" location="Haut-Katanga Province" municipality="Upemba" pageId="36" pageNumber="483" specimenCode="RBINS 2728, RBINS 6156" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFF3FFDBFAFF42FFFA4EFD29" box="[1341,1474,645,669]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2728</specimenCode>
|
||
(formerly under
|
||
<specimenCode id="DBBC1038FFF3FFDBFC3442DFFB04FD08" box="[1014,1160,676,701]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6156</specimenCode>
|
||
), collected at
|
||
<collectorName id="26EFDD95FFF3FFDBFAED42DEFA32FD08" box="[1327,1470,676,700]" pageId="36" pageNumber="483">Kaswabilenga</collectorName>
|
||
, région du cours inférieur
|
||
<collectorName id="26EFDD95FFF3FFDBFBFC42BEFB48FD68" box="[1086,1220,708,732]" pageId="36" pageNumber="483">de la Lupiala</collectorName>
|
||
, affluent de la rive droite
|
||
<collectorName id="26EFDD95FFF3FFDBFCFB4299FC3CFD4F" box="[825,944,739,763]" pageId="36" pageNumber="483">de la Lufira</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFC7B4299FB9DFD4F" box="[953,1041,739,763]" pageId="36" pageNumber="483">Upemba</collectingMunicipality>
|
||
NP,
|
||
<location id="8EC5EE98FFF3FFDBFB81429EFAB9FD4F" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB81429EFAB9FD4F" box="[1091,1333,739,764]" country="Democratic Republic of the Congo" municipality="Upemba" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFAFD4299FAFBFD4F" box="[1343,1399,739,763]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
,
|
||
<elevation id="00375F70FFF3FFDBFA424299FCE9FCAE" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">
|
||
<quantity id="4CE215A6FFF3FFDBFA424299FA4FFD4F" box="[1408,1475,739,763]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFF3FFDBFC484379FBBFFCAF" box="[906,1075,771,795]" pageId="36" pageNumber="483" value="1949-01-31">
|
||
<collectingDate id="EFE0676BFFF3FFDBFC484379FBBFFCAF" box="[906,1075,771,795]" pageId="36" pageNumber="483" value="1949-01-31">31 January 1949</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
. All
|
||
<typeStatus id="54A106E1FFF3FFDBFB9D4379FB34FCAF" box="[1119,1208,771,795]" pageId="36" pageNumber="483" type="paratype">paratype</typeStatus>
|
||
material from
|
||
<materialsCitation id="3B72B21EFFF3FFDBFA8E4379FB5CFCED" collectionCode="RBINS" collectorName="G. - F. de Witte" country="Democratic Republic of the Congo" location="Katanga region" pageId="36" pageNumber="483" specimenCount="1" stateProvince="Katanga" typeStatus="paratype">
|
||
<collectionCode id="ED0B2086FFF3FFDBFA8E4379FA1BFCAF" box="[1356,1431,771,795]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS</collectionCode>
|
||
was collected by
|
||
<collectorName id="26EFDD95FFF3FFDBFC784358FBC6FC8E" box="[954,1098,802,826]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
|
||
during his expeditions in the former
|
||
<collectingRegion id="49DE76A1FFF3FFDBFCFB4338FC58FCED" box="[825,980,833,858]" country="Democratic Republic of the Congo" name="Katanga" pageId="36" pageNumber="483">Katanga region</collectingRegion>
|
||
between 1946 and 1949
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF3FFDBFCFB4300FA03FB39" blockId="36.[825,1475,890,1165]" pageId="36" pageNumber="483">
|
||
<emphasis id="B96E6451FFF3FFDBFCFB4300FC73FC26" box="[825,1023,890,914]" italics="true" pageId="36" pageNumber="483">Additional material:</emphasis>
|
||
<materialsCitation id="3B72B21EFFF3FFDBFBCB4301FC1AFC65" collectingDate="1932-09" collectionCode="RMCA, R" collectorName="G. F. Overlaet" country="Democratic Republic of the Congo" location="Kapanga" municipality="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCount="1">
|
||
<collectionCode id="ED0B2086FFF3FFDBFBCB4301FBD9FC27" box="[1033,1109,891,915]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="36" pageNumber="483" type="Museum">RMCA</collectionCode>
|
||
<collectionCode id="ED0B2086FFF3FFDBFB9B4301FBE0FC27" box="[1113,1132,891,915]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="483">R</collectionCode>
|
||
.8933, male, collected at
|
||
<location id="8EC5EE98FFF3FFDBFAA14301FA33FC27" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAA14301FA33FC27" box="[1379,1471,891,915]" country="Democratic Republic of the Congo" municipality="Haut-Katanga Province" name="Kapanga" pageId="36" pageNumber="483">Kapanga</location>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFCFB43E0FBBFFC06" box="[825,1075,922,946]" pageId="36" pageNumber="483">Haut-Katanga Province</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFB8743E0FBF1FC06" box="[1093,1149,922,946]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
, in
|
||
<date id="FFA49E83FFF3FFDBFB7343E0FAEEFC06" box="[1201,1378,922,946]" pageId="36" pageNumber="483" value="1932-09">
|
||
<collectingDate id="EFE0676BFFF3FFDBFB7343E0FAEEFC06" box="[1201,1378,922,946]" pageId="36" pageNumber="483" value="1932-09">September 1932</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFF3FFDBFA5543E0FC1AFC65" pageId="36" pageNumber="483">G.F. Overlaet</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFC6343C3FBDFFC65" box="[929,1107,953,977]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 6121" specimenCount="1">
|
||
<specimenCode id="DBBC1038FFF3FFDBFC6343C3FBAAFC65" box="[929,1062,953,977]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6121</specimenCode>
|
||
and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B72B21EFFF3FFDBFB9843C3FC35FC44" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 20749, RBINS 6121" specimenCount="1">
|
||
<specimenCode id="DBBC1038FFF3FFDBFB9843C3FB60FC65" box="[1114,1260,953,977]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 20749</specimenCode>
|
||
(formerly in
|
||
<specimenCode id="DBBC1038FFF3FFDBFAB543C3FCE1FC44" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6121</specimenCode>
|
||
series)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B72B21EFFF3FFDBFC0143A2FA32FB9B" collectingDate="1945-07-25" collectingDateMax="1945-07-27" collectingDateMin="1945-07-25" collectorName="de la Grande Kafwe & G. - F. de Witte" country="Democratic Republic of the Congo" county="Haut-Katanga Province" location="Kambi" municipality="Upemba" pageId="36" pageNumber="483" specimenCount="1">
|
||
male and female, respectively, collected at
|
||
<location id="8EC5EE98FFF3FFDBFAB543A2FA33FC44" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAB543A2FA33FC44" box="[1399,1471,984,1008]" country="Democratic Republic of the Congo" county="Haut-Katanga Province" municipality="Upemba" name="Kambi" pageId="36" pageNumber="483">Kambi</location>
|
||
, affluent
|
||
<collectorName id="26EFDD95FFF3FFDBFC4C4382FBD0FBA4" box="[910,1116,1016,1040]" pageId="36" pageNumber="483">de la Grande Kafwe</collectorName>
|
||
(
|
||
<location id="8EC5EE98FFF3FFDBFBA84382FB66FBA4" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFBA84382FB66FBA4" box="[1130,1258,1016,1040]" country="Democratic Republic of the Congo" county="Haut-Katanga Province" municipality="Upemba" name="Masombwe" pageId="36" pageNumber="483">Masombwe</location>
|
||
),
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFB394382FADFFBA4" box="[1275,1363,1016,1040]" pageId="36" pageNumber="483">Upemba</collectingMunicipality>
|
||
NP,
|
||
<collectingCounty id="62C4C0CFFFF3FFDBFA444382FC63FB9B" pageId="36" pageNumber="483">Haut-Katanga Province</collectingCounty>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFC3B446DFBBDFB9B" box="[1017,1073,1047,1071]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
, on
|
||
<date id="FFA49E83FFF3FFDBFB9E446DFA8AFB9B" box="[1116,1286,1047,1071]" pageId="36" pageNumber="483" value="1945-07-25" valueMax="1945-07-27" valueMin="1945-07-25">
|
||
<collectingDate id="EFE0676BFFF3FFDBFB9E446DFA8AFB9B" box="[1116,1286,1047,1071]" pageId="36" pageNumber="483" value="1945-07-25" valueMax="1945-07-27" valueMin="1945-07-25">25–27 July 1945</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFF3FFDBFAE9446DFA32FB9B" box="[1323,1470,1047,1071]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFF3FFDBFCFB444CFB82FBFA" box="[825,1038,1078,1102]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 6139" specimenCount="1">
|
||
<specimenCode id="DBBC1038FFF3FFDBFCFB444CFC4CFBFA" box="[825,960,1078,1102]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6139</specimenCode>
|
||
(male)
|
||
</materialsCitation>
|
||
and
|
||
<materialsCitation id="3B72B21EFFF3FFDBFB84444CFB82FBDA" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 20748, RBINS 6139" specimenCount="1">
|
||
<specimenCode id="DBBC1038FFF3FFDBFB84444CFB56FBFA" box="[1094,1242,1078,1102]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 20748</specimenCode>
|
||
(female) (formerly in
|
||
<specimenCode id="DBBC1038FFF3FFDBFCFB442CFC4DFBDA" box="[825,961,1110,1134]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6139</specimenCode>
|
||
series)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B72B21EFFF3FFDBFBDE442CFA07FB39" collectingDate="1945-08" collectorName="Upemba NP & G. - F. de Witte" country="Democratic Republic of the Congo" location="Mabwe" municipality="Haut-Lomami Province" pageId="36" pageNumber="483" specimenCount="1">
|
||
collected at
|
||
<location id="8EC5EE98FFF3FFDBFB5F442CFB66FBDA" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB5F442CFB66FBDA" box="[1181,1258,1110,1134]" country="Democratic Republic of the Congo" municipality="Haut-Lomami Province" name="Mabwe" pageId="36" pageNumber="483">Mabwe</location>
|
||
,
|
||
<collectorName id="26EFDD95FFF3FFDBFB34442CFAF6FBD9" box="[1270,1402,1110,1134]" pageId="36" pageNumber="483">Upemba NP</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFF3FFDBFA44442CFC7EFB39" pageId="36" pageNumber="483">Haut-Lomami Province</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFF3FFDBFC39440FFBBFFB39" box="[1019,1075,1141,1165]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
|
||
, in
|
||
<date id="FFA49E83FFF3FFDBFB97440FFB5BFB39" box="[1109,1239,1141,1165]" pageId="36" pageNumber="483" value="1945-08">
|
||
<collectingDate id="EFE0676BFFF3FFDBFB97440FFB5BFB39" box="[1109,1239,1141,1165]" pageId="36" pageNumber="483" value="1945-08">August 1945</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFF3FFDBFB38440FFA07FB39" box="[1274,1419,1141,1165]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF3FFDBFCFB44D5FBC9FA16" blockId="36.[825,1475,1198,1974]" pageId="36" pageNumber="483">
|
||
<emphasis id="B96E6451FFF3FFDBFCFB44D5FC13FB72" box="[825,927,1199,1222]" italics="true" pageId="36" pageNumber="483">Diagnosis:</emphasis>
|
||
A large
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFBC044D4FB0DFB72" box="[1026,1153,1198,1222]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96E6451FFF3FFDBFBC044D4FB0DFB72" box="[1026,1153,1198,1222]" italics="true" pageId="36" pageNumber="483">Lygodactylus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<collectionCode id="ED0B2086FFF3FFDBFB4D44D4FB4DFB72" box="[1167,1217,1198,1222]" pageId="36" pageNumber="483">SVL</collectionCode>
|
||
to
|
||
<quantity id="4CE215A6FFF3FFDBFB2744D4FACFFB72" box="[1253,1347,1198,1222]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="36" pageNumber="483" unit="mm" value="39.8">39.8 mm</quantity>
|
||
(mean 36.1 ±
|
||
<quantity id="4CE215A6FFF3FFDBFC8D44B4FC28FB52" box="[847,932,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="36" pageNumber="483" unit="mm" value="2.1">2.1 mm</quantity>
|
||
)] with 6–9 supralabials and 6–7 infralabials. As with other
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFCB84497FC75FAB1" box="[890,1017,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96E6451FFF3FFDBFCB84497FC75FAB1" box="[890,1017,1261,1285]" italics="true" pageId="36" pageNumber="483">Lygodactylus</emphasis>
|
||
</taxonomicName>
|
||
within the
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFBB94494FB78FAB1" box="[1147,1268,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFF3FFDBFBB94494FB78FAB1" box="[1147,1268,1261,1285]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
group, this species can be easily differentiated from the
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFB774577FAA0FA90" authorityName="Gunther" authorityYear="1893" box="[1205,1324,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="angularis">
|
||
<emphasis id="B96E6451FFF3FFDBFB774577FAA0FA90" box="[1205,1324,1292,1316]" italics="true" pageId="36" pageNumber="483">L. angularis</emphasis>
|
||
</taxonomicName>
|
||
group species by the gular ornamentation, dorsal pattern, and lack of terminal scansors on the tail tip, and from other members of the
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFCFB4511FC39FA36" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[825,949,1387,1410]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="picturatus">
|
||
<emphasis id="B96E6451FFF3FFDBFCFB4511FC39FA36" box="[825,949,1387,1410]" italics="true" pageId="36" pageNumber="483">L. picturatus</emphasis>
|
||
</taxonomicName>
|
||
group based on dorsal coloration and gular pattern (see
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFCAB45F0FC51FA16" box="[873,989,1418,1442]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFF3FFDBFCAB45F0FC51FA16" box="[873,989,1418,1442]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
account).
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF3FFDAFC9745D2FF54FA5C" blockId="36.[825,1475,1198,1974]" lastBlockId="37.[113,763,1331,1512]" lastPageId="37" lastPageNumber="484" pageId="36" pageNumber="483">
|
||
<emphasis id="B96E6451FFF3FFDBFC9745D2FB13FA75" bold="true" box="[853,1183,1448,1473]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFC9745D2FBB1FA74" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[853,1085,1448,1472]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFC9745D2FBB1FA74" bold="true" box="[853,1085,1448,1472]" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB8A45D3FB13FA75" box="[1096,1183,1449,1473]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from other members within the
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFB8145B3FB3AFA54" box="[1091,1206,1480,1504]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFF3FFDBFB8145B3FB3AFA54" box="[1091,1206,1480,1504]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
subgroup by the following combination of characters: dorsal coloration lacking dorsolateral series of cream ocelli on flanks (present in
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFAEA4672FA12F9AB" box="[1320,1438,1543,1567]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFF3FFDBFAEA4672FA12F9AB" box="[1320,1438,1543,1567]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96E6451FFF3FFDBFA6E467DFC71F98A" bold="true" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFA6E467DFC13F98A" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFA6E467DFC13F98A" bold="true" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC6A465CFC71F98A" box="[936,1021,1574,1598]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B96E6451FFF3FFDBFBC9465DFB46F98A" bold="true" box="[1035,1226,1574,1598]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFBC9465DFBE1F98A" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1035,1133,1574,1598]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFBC9465DFBE1F98A" bold="true" box="[1035,1133,1574,1598]" italics="true" pageId="36" pageNumber="483">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFBB7465CFB46F98A" box="[1141,1226,1574,1598]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFAC5465DFAF7F98A" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1287,1403,1574,1598]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="dysmicus">
|
||
<emphasis id="B96E6451FFF3FFDBFAC5465DFAF7F98A" box="[1287,1403,1574,1598]" italics="true" pageId="36" pageNumber="483">L. dysmicus</emphasis>
|
||
</taxonomicName>
|
||
), with two parallel dorsolateral lines from head to forelimbs, the lower line extending on to the anterior side of the forelimb, reaching the cubital fossa (
|
||
<figureCitation id="1321A4C6FFF3FFDBFC3046FEFBC0F928" box="[1010,1100,1668,1692]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="36" pageNumber="483">Fig. 25B</figureCitation>
|
||
); males usually with three
|
||
<emphasis id="B96E6451FFF3FFDBFAA046FEFAFFF928" bold="true" box="[1378,1395,1668,1692]" pageId="36" pageNumber="483">∩</emphasis>
|
||
<emphasis id="B96E6451FFF3FFDBFAB146FFFAF6F928" box="[1395,1402,1669,1692]" italics="true" pageId="36" pageNumber="483">-</emphasis>
|
||
shaped chevrons (vs. one or two in
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFB9546DEFB47F90F" authorityName=", Schmidt" authorityYear="1919" box="[1111,1227,1699,1723]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="depressus">
|
||
<emphasis id="B96E6451FFF3FFDBFB9546DEFB47F90F" box="[1111,1227,1699,1723]" italics="true" pageId="36" pageNumber="483">L. depressus</emphasis>
|
||
</taxonomicName>
|
||
; a broken chevron in
|
||
<emphasis id="B96E6451FFF3FFDBFA6E46DEFB86F96F" bold="true" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFA6E46DEFC39F96E" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFA6E46DEFC39F96E" bold="true" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC7B46B9FB86F96F" box="[953,1034,1731,1755]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; and the exceptional inverted
|
||
<emphasis id="B96E6451FFF3FFDBFAFA46BEFACBF96F" box="[1336,1351,1732,1755]" italics="true" pageId="36" pageNumber="483">Y</emphasis>
|
||
-shaped pattern in
|
||
<emphasis id="B96E6451FFF3FFDBFC474699FBF7F94E" bold="true" box="[901,1147,1762,1786]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFC474699FBADF94E" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[901,1057,1762,1786]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFC474699FBADF94E" bold="true" box="[901,1057,1762,1786]" italics="true" pageId="36" pageNumber="483">L. mirabundus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFBEA4698FBF7F94E" box="[1064,1147,1762,1786]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) and thicker gular lines than
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFA6C4699FC18F8AD" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFF3FFDBFA6C4699FC18F8AD" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFC634778FB98F8AD" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[929,1044,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="dysmicus">
|
||
<emphasis id="B96E6451FFF3FFDBFC634778FB98F8AD" box="[929,1044,1793,1817]" italics="true" pageId="36" pageNumber="483">L. dysmicus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B96E6451FFF3FFDBFB8F4778FC65F88D" bold="true" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFB8F4778FB5CF8AD" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1101,1232,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov">
|
||
<emphasis id="B96E6451FFF3FFDBFB8F4778FB5CF8AD" bold="true" box="[1101,1232,1793,1817]" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB154778FAA9F8AE" box="[1239,1317,1794,1818]" pageId="36" pageNumber="483" rank="species">sp. nov</taxonomicNameLabel>
|
||
..
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFAF5477BFC00F88C" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFAF5477BFC00F88C" bold="true" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC56475BFC65F88D" box="[916,1001,1825,1849]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
always has three postmental scales, with the lateral pair separated by a posterior extension of the mental (
|
||
<figureCitation id="1321A4C6FFF3FFDBFC864725FC2DF8CC" box="[836,929,1887,1912]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="36" pageNumber="483">Fig. 25A</figureCitation>
|
||
). It can be differentiated from
|
||
<emphasis id="B96E6451FFF3FFDBFB1D471AFA31F8CC" bold="true" box="[1247,1469,1887,1912]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFB1D471AFAEEF8C3" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1247,1378,1887,1911]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFB1D471AFAEEF8C3" bold="true" box="[1247,1378,1887,1911]" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFAAB471AFA31F8CC" box="[1385,1469,1888,1912]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B96E6451FFF3FFDBFCFB4705FC7CF823" bold="true" box="[825,1008,1918,1943]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFCFB4705FC14F822" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[825,920,1918,1942]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFCFB4705FC14F822" bold="true" box="[825,920,1918,1942]" italics="true" pageId="36" pageNumber="483">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC5C4705FC7CF823" box="[926,1008,1919,1943]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<emphasis id="B96E6451FFF3FFDBFBE54705FA9BF823" bold="true" box="[1063,1303,1918,1943]" pageId="36" pageNumber="483">
|
||
<taxonomicName id="4C1AC3C0FFF3FFDBFBE54705FB33F822" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1063,1215,1918,1942]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF3FFDBFBE54705FB33F822" bold="true" box="[1063,1215,1918,1942]" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB074705FA9BF823" box="[1221,1303,1919,1943]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having a proportionally larger tibia (CL/
|
||
<collectionCode id="ED0B2086FFF3FFDBFBA147E4FB1AF802" box="[1123,1174,1950,1974]" pageId="36" pageNumber="483">SVL</collectionCode>
|
||
≥ 0.19 vs. CL/
|
||
<collectionCode id="ED0B2086FFF3FFDBFAF547E4FAE4F802" box="[1335,1384,1950,1974]" pageId="36" pageNumber="483">SVL</collectionCode>
|
||
≤ 0.17). As in its sister taxon
|
||
<emphasis id="B96E6451FFF2FFDAFE8A454EFDB4FAF8" bold="true" box="[328,568,1331,1356]" pageId="37" pageNumber="484">
|
||
<taxonomicName id="4C1AC3C0FFF2FFDAFE8A454EFE6CFAFF" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[328,480,1331,1355]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF2FFDAFE8A454EFE6CFAFF" bold="true" box="[328,480,1331,1355]" italics="true" pageId="37" pageNumber="484">L. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFE24454EFDB4FAF8" box="[486,568,1332,1356]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, females lack gular patterning (present in all the other members of the group,
|
||
<figureCitation id="1321A4C6FFF2FFDAFD164529FF2DFA3E" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="37" pageNumber="484">Fig. 25A</figureCitation>
|
||
).
|
||
<emphasis id="B96E6451FFF2FFDAFF704508FE61FA3E" bold="true" box="[178,493,1394,1418]" pageId="37" pageNumber="484">
|
||
<taxonomicName id="4C1AC3C0FFF2FFDAFF704508FE1AFA3E" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[178,406,1394,1418]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF2FFDAFF704508FE1AFA3E" bold="true" box="[178,406,1394,1418]" italics="true" pageId="37" pageNumber="484">Lygodactylus gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFE594508FE61FA3E" box="[411,493,1394,1418]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from
|
||
<emphasis id="B96E6451FFF2FFDAFFB345E8FEE8FA1E" bold="true" box="[113,356,1425,1450]" pageId="37" pageNumber="484">
|
||
<taxonomicName id="4C1AC3C0FFF2FFDAFFB345E8FE80FA1D" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[113,268,1425,1449]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF2FFDAFFB345E8FE80FA1D" bold="true" box="[113,268,1425,1449]" italics="true" pageId="37" pageNumber="484">L. mirabundus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFED045E8FEE8FA1E" box="[274,356,1426,1450]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having a lower number of precloacal pores (7–9 vs. 10) and by subtle morphometric differences (
|
||
<tableCitation id="C6988DF8FFF2FFDAFFBE45AAFF4BFA5C" box="[124,199,1488,1512]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="37" pageNumber="484" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF65E8CBFFF2FFDAFFB344CEFADFFAB0" ID-DOI="http://doi.org/10.5281/zenodo.13302917" ID-Zenodo-Dep="13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="37" pageNumber="484" startId="37.[113,178,1204,1228]" targetBox="[114,1458,144,1176]" targetPageId="37" targetType="figure">
|
||
<paragraph id="8BA5B843FFF2FFDAFFB344CEFADFFAB0" blockId="37.[113,1458,1203,1284]" pageId="37" pageNumber="484">
|
||
<emphasis id="B96E6451FFF2FFDAFFB344CEFF5AFB78" bold="true" box="[113,214,1204,1228]" pageId="37" pageNumber="484">Figure 23.</emphasis>
|
||
A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of
|
||
<emphasis id="B96E6451FFF2FFDAFCBD44CEFBD6FB78" bold="true" box="[895,1114,1203,1228]" pageId="37" pageNumber="484">
|
||
<taxonomicName id="4C1AC3C0FFF2FFDAFCBD44CEFB87FB7F" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[895,1035,1203,1227]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF2FFDAFCBD44CEFB87FB7F" bold="true" box="[895,1035,1203,1227]" italics="true" pageId="37" pageNumber="484">L. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFBCD44CEFBD6FB78" box="[1039,1114,1204,1228]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of
|
||
<emphasis id="B96E6451FFF2FFDAFF4B4496FEB3FAB0" bold="true" box="[137,319,1259,1284]" pageId="37" pageNumber="484">
|
||
<taxonomicName id="4C1AC3C0FFF2FFDAFF4B4496FF7CFAB7" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[137,240,1259,1283]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF2FFDAFF4B4496FF7CFAB7" bold="true" box="[137,240,1259,1283]" italics="true" pageId="37" pageNumber="484">L. gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFF364496FEB3FAB0" box="[244,319,1260,1284]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA5B843FFF2FFDAFFB34672FEB4F928" blockId="37.[113,762,1544,1693]" pageId="37" pageNumber="484">
|
||
<emphasis id="B96E6451FFF2FFDAFFB34672FF6DF994" box="[113,225,1544,1568]" italics="true" pageId="37" pageNumber="484">Etymology:</emphasis>
|
||
We name this new species after the American evolutionary biologist and herpetologist Tony Gamble of Marquette University, in recognition of his substantial contributions to the evolutionary biology of geckos. The name is a patronym formed in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF2FFD9FFB346D7FC86FAD8" blockId="37.[113,762,1708,1952]" lastBlockId="38.[129,778,1333,1388]" lastPageId="38" lastPageNumber="485" pageId="37" pageNumber="484">
|
||
<emphasis id="B96E6451FFF2FFDAFFB346D7FE1EF970" box="[113,402,1708,1732]" italics="true" pageId="37" pageNumber="484">
|
||
Description of the
|
||
<typeStatus id="54A106E1FFF2FFDAFE8246D6FE1EF970" box="[320,402,1708,1732]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF2FFDAFE7F46D6FD81F970" box="[445,525,1708,1732]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="37" pageNumber="484">Fig. 24</figureCitation>
|
||
): Measurements and meristic characters of the
|
||
<typeStatus id="54A106E1FFF2FFDAFEB846B1FE59F957" box="[378,469,1739,1763]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
|
||
are presented in Supporting Information, Table S10. Adult male, with a snout–vent length (
|
||
<collectionCode id="ED0B2086FFF2FFDAFFBB4770FF20F896" box="[121,172,1802,1826]" pageId="37" pageNumber="484">SVL</collectionCode>
|
||
) of 35.0 mm and a regenerated tail of 31.0 mm.Body slender and nape slightly distinct. Head slightly broader than body, and moderate head length (
|
||
<collectionCode id="ED0B2086FFF2FFDAFEA94733FE17F8D5" box="[363,411,1865,1889]" pageId="37" pageNumber="484">HW</collectionCode>
|
||
/HL 0.66). Canthus rostralis not prominent. Eye diameter
|
||
<quantity id="4CE215A6FFF2FFDAFEB04712FE40F834" box="[370,460,1896,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.51" pageId="37" pageNumber="484" unit="mm" value="2.51">2.51 mm</quantity>
|
||
with circular pupil. Ear to eye distance slightly larger than the orbit diameter (
|
||
<quantity id="4CE215A6FFF2FFDAFD8E47FDFD22F814" box="[588,686,1927,1952]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="37" pageNumber="484" unit="mm" value="3.26">3.26 mm</quantity>
|
||
). Snout rounded and moderately pointed. Granular scale of frontal larger than occipital scales, with 25 small interorbital granular scales. Dorsal scales granular from rostral to tail. Rostral undivided, in narrow contact with nostril. Two small symmetrical internasals. Seven supralabials and six infralabials. Prenasal contacts the 1st supralabial. Nostril circular and surrounded by 1st supralabial, prenasal, one supranasal, and one postnasal. Four or five rows of scales between supralabials and the orbit. Mental large, triangular, and posteriorly in contact with three rounded symmetrical postmental scales. Mental scales extending between the lateral postmentals reaching ~50% of the medial surface suture of the postmentals. Five smaller post-postmental scales. Gular scales granular, rounded, and smaller than ventral scales. Ventrals imbricate, denticulate, and larger than dorsal scales, with 19 scale rows across the venter. Regenerated tail with 30 enlarged subcaudal scales, some fragmented. Body relatively robust and slightly elongated (
|
||
<collectionCode id="ED0B2086FFF2FFDAFC274753FB95F8F5" box="[997,1049,1833,1857]" pageId="37" pageNumber="484">TRL</collectionCode>
|
||
/
|
||
<collectionCode id="ED0B2086FFF2FFDAFBE34753FBDFF8F5" box="[1057,1107,1833,1857]" pageId="37" pageNumber="484">SVL</collectionCode>
|
||
0.42). Eight precloacal pores. Fore- and hind limbs moderately short and stout; forearm of medium length (
|
||
<collectionCode id="ED0B2086FFF2FFDAFCB8471DFC16F8CB" box="[890,922,1895,1919]" pageId="37" pageNumber="484">FL</collectionCode>
|
||
/
|
||
<collectionCode id="ED0B2086FFF2FFDAFC60471DFC58F8CB" box="[930,980,1895,1919]" pageId="37" pageNumber="484">SVL</collectionCode>
|
||
0.16); tibia moderately long (CL/
|
||
<collectionCode id="ED0B2086FFF2FFDAFAF9471DFAE1F8CB" box="[1339,1389,1895,1919]" pageId="37" pageNumber="484">SVL</collectionCode>
|
||
0.19). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw.Relative length of digits: I <II =
|
||
<collectionCode id="ED0B2086FFF1FFD9FED4452EFEABFADF" box="[278,295,1364,1387]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="485" type="Museum">V</collectionCode>
|
||
<III <IV (manus); I <II <
|
||
<collectionCode id="ED0B2086FFF1FFD9FD89452EFDD0FADF" box="[587,604,1364,1387]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="485" type="Museum">V</collectionCode>
|
||
<III <IV (pes).
|
||
</paragraph>
|
||
<caption id="DF65E8CBFFF1FFD9FF4344AAFABCFAB0" ID-DOI="http://doi.org/10.5281/zenodo.13302919" ID-Zenodo-Dep="13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="38" pageNumber="485" startId="38.[129,194,1232,1256]" targetBox="[129,1473,144,1205]" targetPageId="38" targetType="figure">
|
||
<paragraph id="8BA5B843FFF1FFD9FF4344AAFABCFAB0" blockId="38.[129,1456,1232,1285]" pageId="38" pageNumber="485">
|
||
<emphasis id="B96E6451FFF1FFD9FF4344AAFF69FB5D" bold="true" box="[129,229,1232,1257]" pageId="38" pageNumber="485">Figure 24.</emphasis>
|
||
Holotype of
|
||
<emphasis id="B96E6451FFF1FFD9FEA344ABFD9BFB5D" bold="true" box="[353,535,1232,1257]" pageId="38" pageNumber="485">
|
||
<taxonomicName id="4C1AC3C0FFF1FFD9FEA344ABFE44FB5C" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[353,456,1232,1256]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF1FFD9FEA344ABFE44FB5C" bold="true" box="[353,456,1232,1256]" italics="true" pageId="38" pageNumber="485">L. gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF1FFD9FE0E44ABFD9BFB5D" box="[460,535,1233,1257]" pageId="38" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(UTEP 22587) from Manono, Tanganyika Province, DRC. A, Dorsal and ventral view of body. B, Details of the cloacal region. C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA5B843FFF1FFD9FF4345F7FD3FF96A" blockId="38.[129,778,1421,1758]" pageId="38" pageNumber="485">
|
||
<emphasis id="B96E6451FFF1FFD9FF4345F7FEA2FA11" box="[129,302,1421,1445]" italics="true" pageId="38" pageNumber="485">Coloration: In life</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF1FFD9FEFF45F7FE52FA11" box="[317,478,1421,1445]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="38" pageNumber="485">Fig. 23A–D, F–I</figureCitation>
|
||
), dorsal colour light brown to grey, with dark brown mottled pattern in the female; male with three rows of interrupted, but conspicuous black stripes on lateral surface from head to neck, converging at the anterior insertion of the forearm, giving the impression of a single large blotch; male and female with a black-brown line from snout to ear aperture; venter uniform orange-yellowish from posterior part of gular to cloaca, and cream to white elsewhere; gular immaculate white in female, and white in male with three thick and black
|
||
<emphasis id="B96E6451FFF1FFD9FD6846F2FD37F914" bold="true" box="[682,699,1672,1696]" pageId="38" pageNumber="485">∩</emphasis>
|
||
<emphasis id="B96E6451FFF1FFD9FD7946F3FD4EF914" box="[699,706,1673,1696]" italics="true" pageId="38" pageNumber="485">-</emphasis>
|
||
shaped chevrons.
|
||
<emphasis id="B96E6451FFF1FFD9FF2446D2FEFCF90B" box="[230,368,1704,1727]" italics="true" pageId="38" pageNumber="485">In preservative</emphasis>
|
||
(
|
||
<typeStatus id="54A106E1FFF1FFD9FEB946DDFE52F90B" box="[379,478,1703,1727]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
|
||
;
|
||
<figureCitation id="1321A4C6FFF1FFD9FE2446DDFDA6F90B" box="[486,554,1703,1727]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="38" pageNumber="485">Fig. 24</figureCitation>
|
||
): similar coloration as in life, slightly darker, with gular pattern more brownish.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF1FFD9FF43477BFC3CFA3F" blockId="38.[129,778,1792,1972]" lastBlockId="38.[825,1474,1332,1419]" pageId="38" pageNumber="485">
|
||
<emphasis id="B96E6451FFF1FFD9FF43477BFF68F8AC" box="[129,228,1793,1816]" italics="true" pageId="38" pageNumber="485">Variation:</emphasis>
|
||
All
|
||
<typeStatus id="54A106E1FFF1FFD9FEDC477AFE0DF8AC" box="[286,385,1792,1816]" pageId="38" pageNumber="485" type="paratype">paratypes</typeStatus>
|
||
largely agree with the
|
||
<typeStatus id="54A106E1FFF1FFD9FDBA477AFD5AF8AC" box="[632,726,1792,1816]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
|
||
, and their measurements are summarized in
|
||
<collectorName id="26EFDD95FFF1FFD9FDCC4765FC89F883" box="[526,773,1823,1847]" pageId="38" pageNumber="485">Supporting Information</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFF1FFD9FF434744FF43F8E3" box="[129,207,1854,1879]" pageId="38" pageNumber="485">Table S</collectorName>
|
||
10.
|
||
<collectorName id="26EFDD95FFF1FFD9FF3A4744FE93F8E2" box="[248,287,1854,1878]" pageId="38" pageNumber="485">The</collectorName>
|
||
specimen
|
||
<specimenCode id="DBBC1038FFF1FFD9FE514745FD96F8E2" box="[403,538,1854,1879]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="38" pageNumber="485" type="Museum">RBINS 6138</specimenCode>
|
||
has a bold dorsal pattern from head to midbody.
|
||
<collectorName id="26EFDD95FFF1FFD9FE5C4724FE22F8C1" box="[414,430,1886,1909]" pageId="38" pageNumber="485">It</collectorName>
|
||
is worth noting that all the female specimens examined from
|
||
<collectorName id="26EFDD95FFF1FFD9FE5A4707FE72F821" box="[408,510,1917,1941]" pageId="38" pageNumber="485">de Witte’s</collectorName>
|
||
(1953) expedition lack a gular pattern, in contradiction to
|
||
<collectorName id="26EFDD95FFF1FFD9FE1D47E6FDC9F800" box="[479,581,1948,1972]" pageId="38" pageNumber="485">de Witte’s</collectorName>
|
||
(1953) statement.
|
||
<collectorName id="26EFDD95FFF1FFD9FCFB454FFC1BFAF8" box="[825,919,1333,1356]" pageId="38" pageNumber="485">However</collectorName>
|
||
, we noted that some young male specimens were catalogued as females, which could have misled
|
||
<collectorName id="26EFDD95FFF1FFD9FAD5452EFAFFFAD8" box="[1303,1395,1364,1388]" pageId="38" pageNumber="485">de Witte</collectorName>
|
||
to this conclusion.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF1FFD9FCFB45D6FC19F9D5" blockId="38.[825,1474,1452,1633]" pageId="38" pageNumber="485">
|
||
<emphasis id="B96E6451FFF1FFD9FCFB45D6FBA5FA70" box="[825,1065,1452,1476]" italics="true" pageId="38" pageNumber="485">Habitat and distribution</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF1FFD9FBFD45D6FBF0FA70" box="[1087,1148,1452,1476]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="38" pageNumber="485">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="1321A4C6FFF1FFD9FB4A45D7FB2EFA71" box="[1160,1186,1453,1477]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="38" pageNumber="485">23</figureCitation>
|
||
):
|
||
<emphasis id="B96E6451FFF1FFD9FB7845D6FCEBFA50" bold="true" pageId="38" pageNumber="485">
|
||
<taxonomicName id="4C1AC3C0FFF1FFD9FB7845D6FA12FA70" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1210,1438,1452,1476]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF1FFD9FB7845D6FA12FA70" bold="true" box="[1210,1438,1452,1476]" italics="true" pageId="38" pageNumber="485">Lygodactylus gamblei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF1FFD9FA6145D7FCEBFA50" pageId="38" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from Manono, Upemba National Park, and other nearby areas in south-eastern
|
||
<collectingCountry id="F30DF8D3FFF1FFD9FBBA4591FB3CF9B7" box="[1144,1200,1515,1539]" name="Democratic Republic of the Congo" pageId="38" pageNumber="485">DRC</collectingCountry>
|
||
. The species is associated with Miombo Woodland forest and gallery forest between 500 and ~
|
||
<quantity id="4CE215A6FFF1FFD9FCB74650FC4BF9F5" box="[885,967,1578,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="485" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l. This species is sympatric with
|
||
<taxonomicName id="4C1AC3C0FFF1FFD9FA834651FA29F9F6" box="[1345,1445,1578,1602]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="heeneni">
|
||
<emphasis id="B96E6451FFF1FFD9FA834651FA29F9F6" box="[1345,1445,1578,1602]" italics="true" pageId="38" pageNumber="485">L. heeneni</emphasis>
|
||
</taxonomicName>
|
||
in this area.
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF1FFD9FCFB46F8FC56F8A3" blockId="38.[825,1474,1666,1816]" pageId="38" pageNumber="485">
|
||
<emphasis id="B96E6451FFF1FFD9FCFB46F8FC55F92E" box="[825,985,1666,1690]" italics="true" pageId="38" pageNumber="485">Natural history:</emphasis>
|
||
A diurnal and arboreal species. The
|
||
<typeStatus id="54A106E1FFF1FFD9FAA446F8FA4EF92E" box="[1382,1474,1666,1690]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
|
||
(UTEP 22587) was collected active during the daytime with an air temperature of 45 °C and one of the
|
||
<typeStatus id="54A106E1FFF1FFD9FACB46B8FAE0F96E" box="[1289,1388,1730,1754]" pageId="38" pageNumber="485" type="paratype">paratypes</typeStatus>
|
||
(UTEP 22584) was found scampering around a tree close to the ground in an open area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |