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<mods:classification id="B3C01DB5BF5AB1DCB4EF4525F338F78C">journal article</mods:classification>
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|
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<mods:identifier id="CF2B0F77DB77DFDB57DEC42766CC83FF" type="CLB-Dataset">301345</mods:identifier>
|
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<mods:identifier id="B4C6DD0872286F296EF27B827D324812" type="DOI">10.1093/zoolinnean/zlad123</mods:identifier>
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|
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|
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<treatment id="03B30955FFC0FFE4FF7147FBFDD3F87F" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F" httpUri="http://treatment.plazi.org/id/03B30955FFC0FFE4FF7147FBFDD3F87F" lastPageId="27" lastPageNumber="474" pageId="23" pageNumber="470">
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||||
<treatment id="03B30955FFC0FFE4FF7147FBFDD3F87F" ID-DOI="http://doi.org/10.5281/zenodo.13302856" ID-Zenodo-Dep="13302856" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F" httpUri="http://treatment.plazi.org/id/03B30955FFC0FFE4FF7147FBFDD3F87F" lastPageId="27" lastPageNumber="474" pageId="23" pageNumber="470">
|
||||
<subSubSection id="C300EBC8FFC0FFE8FF7147FBFD35F828" box="[179,697,1921,1948]" pageId="23" pageNumber="470" type="nomenclature">
|
||||
<paragraph id="8BA5B843FFC0FFE8FF7147FBFD35F828" blockId="23.[113,697,1921,1984]" box="[179,697,1921,1948]" pageId="23" pageNumber="470">
|
||||
<taxonomicName id="4C1AC3C0FFC0FFE8FF7147FBFD35F828" ID-CoL="3WQ3G" authority="Schmidt, 1919" authorityName=", Schmidt" authorityYear="1919" box="[179,697,1921,1948]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="depressus">
|
||||
|
|
@ -133,9 +136,9 @@
|
|||
<subSubSection id="C300EBC8FFC0FFE8FFB347D2FD3EF874" box="[113,690,1960,1984]" pageId="23" pageNumber="470" type="description">
|
||||
<paragraph id="8BA5B843FFC0FFE8FFB347D2FD3EF874" blockId="23.[113,697,1921,1984]" box="[113,690,1960,1984]" pageId="23" pageNumber="470">
|
||||
(
|
||||
<figureCitation id="1321A4C6FFC0FFE8FFBE47D2FF61F874" box="[124,237,1960,1984]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStart-2="Figure 15" captionStart-3="Figure 16" captionStartId-0="24.[129,194,1924,1948]" captionStartId-1="25.[113,178,1107,1131]" captionStartId-2="26.[129,194,1836,1860]" captionStartId-3="27.[112,177,961,985]" captionTargetBox-0="[132,1470,1394,1896]" captionTargetBox-1="[115,1458,147,1078]" captionTargetBox-2="[198,1405,144,1808]" captionTargetBox-3="[124,1455,157,932]" captionTargetId-0="figure-744@24.[129,1473,1388,1897]" captionTargetId-1="figure-474@25.[113,1460,144,1079]" captionTargetId-2="figure-10@26.[198,1405,144,1808]" captionTargetId-3="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionText-0="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." captionText-1="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." captionText-2="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-3="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="23" pageNumber="470">Figs 13–16</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC0FFE8FFBE47D2FF61F874" box="[124,237,1960,1984]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStart-2="Figure 15" captionStart-3="Figure 16" captionStartId-0="24.[129,194,1924,1948]" captionStartId-1="25.[113,178,1107,1131]" captionStartId-2="26.[129,194,1836,1860]" captionStartId-3="27.[112,177,961,985]" captionTargetBox-0="[132,1470,1394,1896]" captionTargetBox-1="[115,1458,147,1078]" captionTargetBox-2="[198,1405,144,1808]" captionTargetBox-3="[124,1455,157,932]" captionTargetId-0="figure-744@24.[129,1473,1388,1897]" captionTargetId-1="figure-474@25.[113,1460,144,1079]" captionTargetId-2="figure-10@26.[198,1405,144,1808]" captionTargetId-3="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionText-0="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." captionText-1="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." captionText-2="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-3="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." figureDoi-0="http://doi.org/10.5281/zenodo.13302879" figureDoi-1="http://doi.org/10.5281/zenodo.13302883" figureDoi-2="http://doi.org/10.5281/zenodo.13302887" figureDoi-3="http://doi.org/10.5281/zenodo.13302895" httpUri-0="https://zenodo.org/record/13302879/files/figure.png" httpUri-1="https://zenodo.org/record/13302883/files/figure.png" httpUri-2="https://zenodo.org/record/13302887/files/figure.png" httpUri-3="https://zenodo.org/record/13302895/files/figure.png" pageId="23" pageNumber="470">Figs 13–16</figureCitation>
|
||||
,
|
||||
<tableCitation id="C6988DF8FFC0FFE8FF3547D2FECEF874" box="[247,322,1960,1984]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="23" pageNumber="470">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFC0FFE8FF3547D2FECEF874" box="[247,322,1960,1984]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="23" pageNumber="470" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
; Supporting Information, Table S6)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -254,9 +257,9 @@ Malonza
|
|||
, from the rainforest of the
|
||||
<collectingRegion id="49DE76A1FFCFFFE7FF424157FF4AFEF1" box="[128,198,301,325]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="24" pageNumber="471">Congo</collectingRegion>
|
||||
Basin (
|
||||
<figureCitation id="1321A4C6FFCFFFE7FED04157FEC0FEF1" box="[274,332,301,325]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." pageId="24" pageNumber="471">Figs 1</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FED04157FEC0FEF1" box="[274,332,301,325]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." figureDoi="http://doi.org/10.5281/zenodo.13302837" httpUri="https://zenodo.org/record/13302837/files/figure.png" pageId="24" pageNumber="471">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="1321A4C6FFCFFFE7FE944157FEEFFEF1" box="[342,355,301,325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="24" pageNumber="471">6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FE944157FEEFFEF1" box="[342,355,301,325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="24" pageNumber="471">6</figureCitation>
|
||||
) that differs by
|
||||
<emphasis id="B96E6451FFCFFFE7FDC24157FD85FEF0" box="[512,521,301,324]" italics="true" pageId="24" pageNumber="471">c</emphasis>
|
||||
. 9.04% (
|
||||
|
|
@ -308,7 +311,7 @@ north-east from the previous locality) and identified by de Witte in 1923 (RMCA
|
|||
<emphasis id="B96E6451FFCFFFE7FDE34467FD16FB80" box="[545,666,1052,1076]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
, show the same two dorsal patterns as the previous specimens mentioned above (
|
||||
<figureCitation id="1321A4C6FFCFFFE7FF0E4421FE9CFBC7" box="[204,272,1115,1139]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." pageId="24" pageNumber="471">Fig. 13</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FF0E4421FE9CFBC7" box="[204,272,1115,1139]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." figureDoi="http://doi.org/10.5281/zenodo.13302879" httpUri="https://zenodo.org/record/13302879/files/figure.png" pageId="24" pageNumber="471">Fig. 13</figureCitation>
|
||||
). Two additional specimens (RMCA 1038/B and 1038/C) were collected from the Ituri Forest (without a precise locality within this region, but in the same region as the
|
||||
<typeStatus id="54A106E1FFCFFFE7FD1944E0FC85FB06" box="[731,777,1178,1202]" pageId="24" pageNumber="471">type</typeStatus>
|
||||
locality of
|
||||
|
|
@ -334,15 +337,15 @@ and
|
|||
, making it impossible to differentiate them from the
|
||||
<typeStatus id="54A106E1FFCFFFE7FA9640B5FA0EFF53" box="[1364,1410,207,231]" pageId="24" pageNumber="471">type</typeStatus>
|
||||
series (
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC864094FC1DFEB1" box="[836,913,237,262]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="24" pageNumber="471">Figs 15</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC864094FC1DFEB1" box="[836,913,237,262]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." figureDoi="http://doi.org/10.5281/zenodo.13302887" httpUri="https://zenodo.org/record/13302887/files/figure.png" pageId="24" pageNumber="471">Figs 15</figureCitation>
|
||||
,
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC634094FC37FEB2" box="[929,955,238,262]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="24" pageNumber="471">16</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC634094FC37FEB2" box="[929,955,238,262]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." figureDoi="http://doi.org/10.5281/zenodo.13302895" httpUri="https://zenodo.org/record/13302895/files/figure.png" pageId="24" pageNumber="471">16</figureCitation>
|
||||
). The specimen collected from
|
||||
<collectingCountry id="F30DF8D3FFCFFFE7FAD14094FAD0FEB2" box="[1299,1372,238,262]" name="Angola" pageId="24" pageNumber="471">Angola</collectingCountry>
|
||||
(MNCN 50772) has a dark blueish coloration
|
||||
<emphasis id="B96E6451FFCFFFE7FB074174FB71FE91" box="[1221,1277,269,293]" italics="true" pageId="24" pageNumber="471">in life</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCFFFE7FAD24177FAE7FE91" box="[1296,1387,269,293]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="24" pageNumber="471">Fig. 15G</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FAD24177FAE7FE91" box="[1296,1387,269,293]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." figureDoi="http://doi.org/10.5281/zenodo.13302887" httpUri="https://zenodo.org/record/13302887/files/figure.png" pageId="24" pageNumber="471">Fig. 15G</figureCitation>
|
||||
), as described in
|
||||
<taxonomicName id="4C1AC3C0FFCFFFE7FC644157FB97FEF0" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[934,1051,300,324]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
|
||||
<emphasis id="B96E6451FFCFFFE7FC644157FB97FEF0" box="[934,1051,300,324]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
|
||||
|
|
@ -353,7 +356,7 @@ series (
|
|||
<taxonomicNameLabel id="A25DD92AFFCFFFE7FC124116FC67FE37" box="[976,1003,364,387]" pageId="24" pageNumber="471" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC394111FBBFFE37" box="[1019,1075,363,387]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." pageId="24" pageNumber="471">Fig. 1</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC394111FBBFFE37" box="[1019,1075,363,387]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." figureDoi="http://doi.org/10.5281/zenodo.13302837" httpUri="https://zenodo.org/record/13302837/files/figure.png" pageId="24" pageNumber="471">Fig. 1</figureCitation>
|
||||
), represents
|
||||
<taxonomicName id="4C1AC3C0FFCFFFE7FB7B4116FAA1FE37" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[1209,1325,363,387]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
|
||||
<emphasis id="B96E6451FFCFFFE7FB7B4116FAA1FE37" box="[1209,1325,363,387]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
|
||||
|
|
@ -389,7 +392,7 @@ rivers might belong to an undescribed taxon.
|
|||
<typeStatus id="54A106E1FFCFFFE7FCFB437FFC1EFCA9" box="[825,914,773,797]" pageId="24" pageNumber="471" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC65437FFC60FCAA" box="[935,1004,773,798]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." pageId="24" pageNumber="471">Fig. 14</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCFFFE7FC65437FFC60FCAA" box="[935,1004,773,798]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." figureDoi="http://doi.org/10.5281/zenodo.13302883" httpUri="https://zenodo.org/record/13302883/files/figure.png" pageId="24" pageNumber="471">Fig. 14</figureCitation>
|
||||
):
|
||||
<specimenCode id="DBBC1038FFCFFFE7FBC6437CFB11FCA9" box="[1028,1181,773,798]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="471">AMNH 10345</specimenCode>
|
||||
, adult male with original tail, collected at
|
||||
|
|
@ -603,7 +606,7 @@ and
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF65E8CBFFCFFFE7FF4347FEFD3DF80D" pageId="24" pageNumber="471" startId="24.[129,194,1924,1948]" targetBox="[132,1470,1394,1896]" targetPageId="24" targetType="figure">
|
||||
<caption id="DF65E8CBFFCFFFE7FF4347FEFD3DF80D" ID-DOI="http://doi.org/10.5281/zenodo.13302879" ID-Zenodo-Dep="13302879" httpUri="https://zenodo.org/record/13302879/files/figure.png" pageId="24" pageNumber="471" startId="24.[129,194,1924,1948]" targetBox="[132,1470,1394,1896]" targetPageId="24" targetType="figure">
|
||||
<paragraph id="8BA5B843FFCFFFE7FF4347FEFD3DF80D" blockId="24.[129,1446,1924,1980]" pageId="24" pageNumber="471">
|
||||
<emphasis id="B96E6451FFCFFFE7FF4347FEFF69F828" bold="true" box="[129,229,1924,1948]" pageId="24" pageNumber="471">Figure 13.</emphasis>
|
||||
Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of
|
||||
|
|
@ -613,7 +616,7 @@ Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) a
|
|||
, collected in sympatry at Flandria, Équateur Province, DRC.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF65E8CBFFCEFFE6FFB34429FB85FB17" pageId="25" pageNumber="472" startId="25.[113,178,1107,1131]" targetBox="[115,1458,147,1078]" targetPageId="25" targetType="figure">
|
||||
<caption id="DF65E8CBFFCEFFE6FFB34429FB85FB17" ID-DOI="http://doi.org/10.5281/zenodo.13302883" ID-Zenodo-Dep="13302883" httpUri="https://zenodo.org/record/13302883/files/figure.png" pageId="25" pageNumber="472" startId="25.[113,178,1107,1131]" targetBox="[115,1458,147,1078]" targetPageId="25" targetType="figure">
|
||||
<paragraph id="8BA5B843FFCEFFE6FFB34429FB85FB17" blockId="25.[113,1444,1107,1187]" pageId="25" pageNumber="472">
|
||||
<emphasis id="B96E6451FFCEFFE6FFB34429FF5AFBDF" bold="true" box="[113,214,1107,1131]" pageId="25" pageNumber="472">Figure 14.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -642,7 +645,7 @@ with a maximum SVL
|
|||
<emphasis id="B96E6451FFCEFFE6FE2D459DFDC6FA4B" box="[495,586,1511,1535]" italics="true" pageId="25" pageNumber="472">gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
group. Seven to nine supralabials and 6–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate in original tails. Large triangular mental followed by 2–3 symmetric postmental scales (
|
||||
<figureCitation id="1321A4C6FFCEFFE6FE44461FFE42F9C9" box="[390,462,1637,1661]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="25" pageNumber="472">Fig. 16</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCEFFE6FE44461FFE42F9C9" box="[390,462,1637,1661]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." figureDoi="http://doi.org/10.5281/zenodo.13302895" httpUri="https://zenodo.org/record/13302895/files/figure.png" pageId="25" pageNumber="472">Fig. 16</figureCitation>
|
||||
). Males with 7–8 precloacal pores. Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually with small denticulation posteriorly. Digits elongated with five terminal scansors on the fourth toe (Supporting Information, Table S6).
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFCEFFE6FF4F477BFDF3F802" blockId="25.[113,763,1449,1974]" pageId="25" pageNumber="472">
|
||||
|
|
@ -659,7 +662,7 @@ group members and from other members of the
|
|||
<emphasis id="B96E6451FFCEFFE6FE4B473BFE66F8EC" box="[393,490,1857,1880]" italics="true" pageId="25" pageNumber="472">picturatus</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup by the gular pattern and dorsal colour pattern (
|
||||
<figureCitation id="1321A4C6FFCEFFE6FE6C4725FE6AF8C3" box="[430,486,1887,1911]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="25" pageNumber="472">Fig. 5</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCEFFE6FE6C4725FE6AF8C3" box="[430,486,1887,1911]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." figureDoi="http://doi.org/10.5281/zenodo.13302850" httpUri="https://zenodo.org/record/13302850/files/figure.png" pageId="25" pageNumber="472">Fig. 5</figureCitation>
|
||||
). It should be noted that
|
||||
<taxonomicName id="4C1AC3C0FFCEFFE6FD24471AFF47F823" authorityName=", Schmidt" authorityYear="1919" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
|
||||
<emphasis id="B96E6451FFCEFFE6FD24471AFF47F823" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
|
||||
|
|
@ -717,7 +720,7 @@ as follows: eye proportionally smaller in
|
|||
in
|
||||
<emphasis id="B96E6451FFCEFFE6FC8A4610FC08F936" box="[840,900,1642,1666]" italics="true" pageId="25" pageNumber="472">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCEFFE6FC5B4610FC59F936" box="[921,981,1642,1666]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="25" pageNumber="472">Fig. 2</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCEFFE6FC5B4610FC59F936" box="[921,981,1642,1666]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="25" pageNumber="472">Fig. 2</figureCitation>
|
||||
), and habitat (rainforest for
|
||||
<taxonomicName id="4C1AC3C0FFCEFFE6FAC94611FA0FF936" authorityName=", Schmidt" authorityYear="1919" box="[1291,1411,1642,1666]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
|
||||
<emphasis id="B96E6451FFCEFFE6FAC94611FA0FF936" box="[1291,1411,1642,1666]" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
|
||||
|
|
@ -744,7 +747,7 @@ by having fewer precloacal pores (7–8 vs.
|
|||
<paragraph id="8BA5B843FFCEFFE4FCE84725FDDCFABF" blockId="25.[810,1459,1887,1974]" lastBlockId="27.[113,763,1048,1291]" lastPageId="27" lastPageNumber="474" pageId="25" pageNumber="472">
|
||||
<emphasis id="B96E6451FFCEFFE6FCE84725FC51F8C3" box="[810,989,1887,1911]" italics="true" pageId="25" pageNumber="472">Coloration: In life</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCEFFE6FC2C4725FBA2F8C3" box="[1006,1070,1887,1911]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="25" pageNumber="472">Fig.15</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCEFFE6FC2C4725FBA2F8C3" box="[1006,1070,1887,1911]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." figureDoi="http://doi.org/10.5281/zenodo.13302887" httpUri="https://zenodo.org/record/13302887/files/figure.png" pageId="25" pageNumber="472">Fig.15</figureCitation>
|
||||
), dorsal colour is highly variable; predominantly dark grey, with lighter grey blotches surrounded by black dots on the flanks, which can form a diffuse dorsolateral band; dark
|
||||
<emphasis id="B96E6451FFCCFFE4FF2F4463FF71FB84" box="[237,253,1049,1072]" italics="true" pageId="27" pageNumber="474">V</emphasis>
|
||||
-shaped line between the eyes (absent in darker specimens), and black line from snout to anterior insertion of the forelimb; tail with diffuse bars of lighter grey; throat usually with white background and black chevrons; venter vivid yellow, orange, or cream from tail tip to posterior part of the gular region.
|
||||
|
|
@ -752,10 +755,10 @@ by having fewer precloacal pores (7–8 vs.
|
|||
(
|
||||
<typeStatus id="54A106E1FFCCFFE4FE8444CFFE26FB79" box="[326,426,1205,1229]" pageId="27" pageNumber="474" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFCCFFE4FE7644CFFE76FB79" box="[436,506,1205,1229]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." pageId="27" pageNumber="474">Fig. 14</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCCFFE4FE7644CFFE76FB79" box="[436,506,1205,1229]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." figureDoi="http://doi.org/10.5281/zenodo.13302883" httpUri="https://zenodo.org/record/13302883/files/figure.png" pageId="27" pageNumber="474">Fig. 14</figureCitation>
|
||||
), dorsum grey with dark brown dots from head to midbody; venter uniform light cream to yellow with whitish colouration on all digits.
|
||||
</paragraph>
|
||||
<caption id="DF65E8CBFFCDFFE5FF434756FC23F82C" pageId="26" pageNumber="473" startId="26.[129,194,1836,1860]" targetBox="[198,1405,144,1808]" targetPageId="26" targetType="figure">
|
||||
<caption id="DF65E8CBFFCDFFE5FF434756FC23F82C" ID-DOI="http://doi.org/10.5281/zenodo.13302887" ID-Zenodo-Dep="13302887" httpUri="https://zenodo.org/record/13302887/files/figure.png" pageId="26" pageNumber="473" startId="26.[129,194,1836,1860]" targetBox="[198,1405,144,1808]" targetPageId="26" targetType="figure">
|
||||
<paragraph id="8BA5B843FFCDFFE5FF434756FC23F82C" blockId="26.[129,1451,1836,1944]" pageId="26" pageNumber="473">
|
||||
<emphasis id="B96E6451FFCDFFE5FF434756FF69F8F0" bold="true" box="[129,229,1836,1860]" pageId="26" pageNumber="473">Figure 15.</emphasis>
|
||||
Specimens of
|
||||
|
|
@ -765,7 +768,7 @@ Specimens of
|
|||
in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G. (A–C), C.K. (D), P.V.P. (E–G), and J.L.R. (H).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF65E8CBFFCCFFE4FFB243BBFEF8FC41" pageId="27" pageNumber="474" startId="27.[112,177,961,985]" targetBox="[124,1455,157,932]" targetPageId="27" targetType="figure">
|
||||
<caption id="DF65E8CBFFCCFFE4FFB243BBFEF8FC41" ID-DOI="http://doi.org/10.5281/zenodo.13302895" ID-Zenodo-Dep="13302895" httpUri="https://zenodo.org/record/13302895/files/figure.png" pageId="27" pageNumber="474" startId="27.[112,177,961,985]" targetBox="[124,1455,157,932]" targetPageId="27" targetType="figure">
|
||||
<paragraph id="8BA5B843FFCCFFE4FFB243BBFEF8FC41" blockId="27.[112,1456,961,1013]" pageId="27" pageNumber="474">
|
||||
<emphasis id="B96E6451FFCCFFE4FFB243BBFF58FC6D" bold="true" box="[112,212,961,985]" pageId="27" pageNumber="474">Figure 16.</emphasis>
|
||||
Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in
|
||||
|
|
@ -778,7 +781,7 @@ Dorsal and ventral views of head showing gular ornamentation and pholidosis vari
|
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<paragraph id="8BA5B843FFCCFFE4FFB34557FDB6F9B5" blockId="27.[113,762,1325,1537]" pageId="27" pageNumber="474">
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<emphasis id="B96E6451FFCCFFE4FFB34557FF58FAF0" box="[113,212,1325,1348]" italics="true" pageId="27" pageNumber="474">Variation:</emphasis>
|
||||
Meristic and morphometric data are summarized and depicted in the Supporting Information (Table S6) and
|
||||
<figureCitation id="1321A4C6FFCCFFE4FD6D4536FF3FFA37" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStartId-0="26.[129,194,1836,1860]" captionStartId-1="27.[112,177,961,985]" captionTargetBox-0="[198,1405,144,1808]" captionTargetBox-1="[124,1455,157,932]" captionTargetId-0="figure-10@26.[198,1405,144,1808]" captionTargetId-1="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-1="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="27" pageNumber="474">Figures 15–16</figureCitation>
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||||
<figureCitation id="1321A4C6FFCCFFE4FD6D4536FF3FFA37" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStartId-0="26.[129,194,1836,1860]" captionStartId-1="27.[112,177,961,985]" captionTargetBox-0="[198,1405,144,1808]" captionTargetBox-1="[124,1455,157,932]" captionTargetId-0="figure-10@26.[198,1405,144,1808]" captionTargetId-1="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-1="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." figureDoi-0="http://doi.org/10.5281/zenodo.13302887" figureDoi-1="http://doi.org/10.5281/zenodo.13302895" httpUri-0="https://zenodo.org/record/13302887/files/figure.png" httpUri-1="https://zenodo.org/record/13302895/files/figure.png" pageId="27" pageNumber="474">Figures 15–16</figureCitation>
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||||
.
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||||
<taxonomicName id="4C1AC3C0FFCCFFE4FF024511FE2CFA37" authorityName=", Schmidt" authorityYear="1919" box="[192,416,1387,1411]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="depressus">
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||||
<emphasis id="B96E6451FFCCFFE4FF024511FE2CFA37" box="[192,416,1387,1411]" italics="true" pageId="27" pageNumber="474">Lygodactylus depressus</emphasis>
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|
|
@ -788,9 +791,9 @@ is the most variable taxon within the
|
|||
<emphasis id="B96E6451FFCCFFE4FF5545F1FE87FA17" box="[151,267,1419,1443]" italics="true" pageId="27" pageNumber="474">L. gutturalis</emphasis>
|
||||
</taxonomicName>
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||||
group with respect to coloration, particularly the gular pattern (
|
||||
<figureCitation id="1321A4C6FFCCFFE4FEC645D0FEC6FA76" box="[260,330,1450,1474]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="27" pageNumber="474">Fig. 16</figureCitation>
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||||
<figureCitation id="1321A4C6FFCCFFE4FEC645D0FEC6FA76" box="[260,330,1450,1474]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." figureDoi="http://doi.org/10.5281/zenodo.13302895" httpUri="https://zenodo.org/record/13302895/files/figure.png" pageId="27" pageNumber="474">Fig. 16</figureCitation>
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||||
). Some specimens lose the dorsal pattern, resulting in a uniform dorsal coloration. However, other specimens have a mottled dorsal pattern (
|
||||
<figureCitation id="1321A4C6FFCCFFE4FE264592FDA5F9B5" box="[484,553,1512,1537]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." pageId="27" pageNumber="474">Fig. 13</figureCitation>
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||||
<figureCitation id="1321A4C6FFCCFFE4FE264592FDA5F9B5" box="[484,553,1512,1537]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." figureDoi="http://doi.org/10.5281/zenodo.13302879" httpUri="https://zenodo.org/record/13302879/files/figure.png" pageId="27" pageNumber="474">Fig. 13</figureCitation>
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||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -798,9 +801,9 @@ group with respect to coloration, particularly the gular pattern (
|
|||
<paragraph id="8BA5B843FFCCFFE4FFB34658FD2FF942" blockId="27.[113,763,1570,1782]" pageId="27" pageNumber="474">
|
||||
<emphasis id="B96E6451FFCCFFE4FFB34658FEEDF98E" box="[113,353,1570,1594]" italics="true" pageId="27" pageNumber="474">Habitat and distribution</emphasis>
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||||
(
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||||
<figureCitation id="1321A4C6FFCCFFE4FEBA4658FE39F98E" box="[376,437,1570,1594]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="27" pageNumber="474">Figs 6</figureCitation>
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<figureCitation id="1321A4C6FFCCFFE4FEBA4658FE39F98E" box="[376,437,1570,1594]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="27" pageNumber="474">Figs 6</figureCitation>
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,
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||||
<figureCitation id="1321A4C6FFCCFFE4FE024658FE56F98E" box="[448,474,1570,1594]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="27" pageNumber="474">15</figureCitation>
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<figureCitation id="1321A4C6FFCCFFE4FE024658FE56F98E" box="[448,474,1570,1594]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." figureDoi="http://doi.org/10.5281/zenodo.13302887" httpUri="https://zenodo.org/record/13302887/files/figure.png" pageId="27" pageNumber="474">15</figureCitation>
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): This species is widely distributed within the
|
||||
<collectingCountry id="F30DF8D3FFCCFFE4FE81463BFE05F9ED" box="[323,393,1601,1625]" name="Democratic Republic of the Congo" pageId="27" pageNumber="474">Congo</collectingCountry>
|
||||
Basin, from the Ituri Forest in the north-east, through the heart of the
|
||||
|
|
|
|||
|
|
@ -1,125 +1,128 @@
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|||
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<treatment id="03B30955FFC2FFE8FF0C470DFDDEF8D6" LSID="urn:lsid:plazi:treatment:03B30955FFC2FFE8FF0C470DFDDEF8D6" httpUri="http://treatment.plazi.org/id/03B30955FFC2FFE8FF0C470DFDDEF8D6" lastPageId="23" lastPageNumber="470" pageId="21" pageNumber="468">
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<treatment id="03B30955FFC2FFE8FF0C470DFDDEF8D6" ID-DOI="http://doi.org/10.5281/zenodo.13302844" ID-Zenodo-Dep="13302844" LSID="urn:lsid:plazi:treatment:03B30955FFC2FFE8FF0C470DFDDEF8D6" httpUri="http://treatment.plazi.org/id/03B30955FFC2FFE8FF0C470DFDDEF8D6" lastPageId="23" lastPageNumber="470" pageId="21" pageNumber="468">
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<taxonomicName id="4C1AC3C0FFC2FFEAFF0C470DFD11F825" ID-CoL="C7JCR" authority="(Perret, 1963)" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[206,669,1911,1937]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
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|
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@ -129,9 +132,9 @@
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</emphasis>
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(
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<figureCitation id="1321A4C6FFC2FFEAFEA547E7FE20F801" box="[359,428,1949,1973]" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." pageId="21" pageNumber="468">Fig. 12</figureCitation>
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<figureCitation id="1321A4C6FFC2FFEAFEA547E7FE20F801" box="[359,428,1949,1973]" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302877" httpUri="https://zenodo.org/record/13302877/files/figure.png" pageId="21" pageNumber="468">Fig. 12</figureCitation>
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;
|
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<tableCitation id="C6988DF8FFC2FFEAFE7B47E7FD88F801" box="[441,516,1949,1973]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="21" pageNumber="468">Table 3</tableCitation>
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<tableCitation id="C6988DF8FFC2FFEAFE7B47E7FD88F801" box="[441,516,1949,1973]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="21" pageNumber="468" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
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)
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|
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|
|
@ -189,7 +192,7 @@
|
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</emphasis>
|
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<caption id="DF65E8CBFFC1FFE9FF4244C9FB66FAB7" pageId="22" pageNumber="469" startId="22.[128,193,1203,1227]" targetBox="[129,1473,144,1175]" targetPageId="22" targetType="figure">
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<paragraph id="8BA5B843FFC1FFE9FF4244C9FB66FAB7" blockId="22.[128,1474,1203,1284]" pageId="22" pageNumber="469">
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<emphasis id="B96E6451FFC1FFE9FF4244C9FF68FB7F" bold="true" box="[128,228,1203,1227]" pageId="22" pageNumber="469">Figure 11.</emphasis>
|
||||
Specimens of
|
||||
|
|
@ -299,7 +302,7 @@ a.s.l.
|
|||
<paragraph id="8BA5B843FFC1FFE8FCFB4630FE23F987" blockId="22.[825,1475,1610,1978]" lastBlockId="23.[113,763,1250,1587]" lastPageId="23" lastPageNumber="470" pageId="22" pageNumber="469">
|
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<emphasis id="B96E6451FFC1FFE9FCFB4630FC15F9D5" box="[825,921,1610,1633]" italics="true" pageId="22" pageNumber="469">Diagnosis</emphasis>
|
||||
(
|
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<figureCitation id="1321A4C6FFC1FFE9FC6D4630FC76F9D6" box="[943,1018,1610,1634]" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." pageId="22" pageNumber="469">Fig. 12</figureCitation>
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||||
<figureCitation id="1321A4C6FFC1FFE9FC6D4630FC76F9D6" box="[943,1018,1610,1634]" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302877" httpUri="https://zenodo.org/record/13302877/files/figure.png" pageId="22" pageNumber="469">Fig. 12</figureCitation>
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): As noted by
|
||||
<bibRefCitation id="EF8BC5B2FFC1FFE9FB6A4630FAB0F9D6" author="Perret JL" box="[1192,1340,1610,1634]" pageId="22" pageNumber="469" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret (1963)</bibRefCitation>
|
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, the species differs from the
|
||||
|
|
@ -351,7 +354,7 @@ vs. 33.3 ±
|
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<emphasis id="B96E6451FFC0FFE8FF4E451AFE8FFAC3" box="[140,259,1375,1399]" italics="true" pageId="23" pageNumber="470">L. gutturalis</emphasis>
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|
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], with a paedomorphic skull that has an unfused frontal bone, braincase, and compound bone + surangular (
|
||||
<figureCitation id="1321A4C6FFC0FFE8FD164505FFF2FA02" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="23" pageNumber="470">Fig. 5</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC0FFE8FD164505FFF2FA02" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." figureDoi="http://doi.org/10.5281/zenodo.13302850" httpUri="https://zenodo.org/record/13302850/files/figure.png" pageId="23" pageNumber="470">Fig. 5</figureCitation>
|
||||
). However, the well-developed state of the precloacal pores suggest sexual maturity of this specimen (
|
||||
<bibRefCitation id="EF8BC5B2FFC0FFE8FDE245C7FD4AFA61" author="Rhen T & Sakata JT & Crews D" box="[544,710,1469,1493]" pageId="23" pageNumber="470" pagination="289 - 96" refId="ref36740" refString="Rhen T, Sakata JT, Crews D. Effects of gonadal sex and incubation temperature on the ontogeny of gonadal steroid concentrations and secondary sex structures in leopard geckos, Eublepharis macularius. General and Comparative Endocrinology 2005; 142: 289 - 96. https: // doi. org / 10.1016 / j. ygcen. 2005.01.018" type="journal article" year="2005">
|
||||
Rhen
|
||||
|
|
@ -359,7 +362,7 @@ Rhen
|
|||
2005
|
||||
</bibRefCitation>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFC0FFE8FD1645C7FF16FA41" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." pageId="23" pageNumber="470">Fig. 12B</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC0FFE8FD1645C7FF16FA41" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302877" httpUri="https://zenodo.org/record/13302877/files/figure.png" pageId="23" pageNumber="470">Fig. 12B</figureCitation>
|
||||
). Finally,
|
||||
<taxonomicName id="4C1AC3C0FFC0FFE8FEC245A7FEFDFA41" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[256,369,1501,1525]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
|
||||
<emphasis id="B96E6451FFC0FFE8FEC245A7FEFDFA41" box="[256,369,1501,1525]" italics="true" pageId="23" pageNumber="470">L. dysmicus</emphasis>
|
||||
|
|
@ -369,11 +372,11 @@ can be differentiated from
|
|||
<emphasis id="B96E6451FFC0FFE8FD4645A7FD77FA41" box="[644,763,1501,1525]" italics="true" pageId="23" pageNumber="470">L. gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
based on its distribution in lowland Congolian rainforest vs. the sub-Saharan savannah (
|
||||
<figureCitation id="1321A4C6FFC0FFE8FEA44661FE13F987" box="[358,415,1563,1587]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="23" pageNumber="470">Fig. 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC0FFE8FEA44661FE13F987" box="[358,415,1563,1587]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="23" pageNumber="470">Fig. 6</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF65E8CBFFC0FFE8FFB34418FF48FB06" pageId="23" pageNumber="470" startId="23.[113,178,1122,1146]" targetBox="[114,1458,144,1094]" targetPageId="23" targetType="figure">
|
||||
<caption id="DF65E8CBFFC0FFE8FFB34418FF48FB06" ID-DOI="http://doi.org/10.5281/zenodo.13302877" ID-Zenodo-Dep="13302877" httpUri="https://zenodo.org/record/13302877/files/figure.png" pageId="23" pageNumber="470" startId="23.[113,178,1122,1146]" targetBox="[114,1458,144,1094]" targetPageId="23" targetType="figure">
|
||||
<paragraph id="8BA5B843FFC0FFE8FFB34418FF48FB06" blockId="23.[113,1455,1122,1202]" pageId="23" pageNumber="470">
|
||||
<emphasis id="B96E6451FFC0FFE8FFB34418FF5AFBCE" bold="true" box="[113,214,1122,1146]" pageId="23" pageNumber="470">Figure 12.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -387,7 +390,7 @@ Holotype of
|
|||
<paragraph id="8BA5B843FFC0FFE8FFB3462FFDA6F8BD" blockId="23.[113,762,1620,1801]" pageId="23" pageNumber="470">
|
||||
<emphasis id="B96E6451FFC0FFE8FFB3462FFEF9F9D9" box="[113,373,1621,1645]" italics="true" pageId="23" pageNumber="470">Habitat and distribution</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFC0FFE8FE4D462FFE5EF9D8" box="[399,466,1620,1645]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="23" pageNumber="470">Fig. 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC0FFE8FE4D462FFE5EF9D8" box="[399,466,1620,1645]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="23" pageNumber="470">Fig. 6</figureCitation>
|
||||
):
|
||||
<taxonomicName id="4C1AC3C0FFC0FFE8FE36462FFD56F9D9" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[500,730,1621,1645]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
|
||||
<emphasis id="B96E6451FFC0FFE8FE36462FFD56F9D9" box="[500,730,1621,1645]" italics="true" pageId="23" pageNumber="470">Lygodactylus dysmicus</emphasis>
|
||||
|
|
|
|||
|
|
@ -1,125 +1,128 @@
|
|||
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<mods:namePart id="D54C12D1E3CC7F7CCD7B1F696D426495">Kusamba, Chifundera</mods:namePart>
|
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@ -135,11 +138,11 @@
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<subSubSection id="C300EBC8FFCAFFE2FFB3468BFE16F8D5" pageId="29" pageNumber="476" type="description">
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<paragraph id="8BA5B843FFCAFFE2FFB3468BFD76F8BD" blockId="29.[113,762,1777,1801]" box="[113,762,1777,1801]" pageId="29" pageNumber="476">
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(
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<figureCitation id="1321A4C6FFCAFFE2FFBE468BFF67F8BD" box="[124,235,1777,1801]" captionStart-0="Figure 19" captionStart-1="Figure 20" captionStartId-0="31.[115,180,1898,1922]" captionStartId-1="32.[128,193,1898,1922]" captionTargetBox-0="[114,1458,920,1870]" captionTargetBox-1="[129,1473,449,1870]" captionTargetId-0="figure-423@31.[114,1458,920,1870]" captionTargetId-1="figure-170@32.[129,1473,449,1870]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="29" pageNumber="476">Figs 19–20</figureCitation>
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<figureCitation id="1321A4C6FFCAFFE2FFBE468BFF67F8BD" box="[124,235,1777,1801]" captionStart-0="Figure 19" captionStart-1="Figure 20" captionStartId-0="31.[115,180,1898,1922]" captionStartId-1="32.[128,193,1898,1922]" captionTargetBox-0="[114,1458,920,1870]" captionTargetBox-1="[129,1473,449,1870]" captionTargetId-0="figure-423@31.[114,1458,920,1870]" captionTargetId-1="figure-170@32.[129,1473,449,1870]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi-0="http://doi.org/10.5281/zenodo.13302905" figureDoi-1="http://doi.org/10.5281/zenodo.13302907" httpUri-0="https://zenodo.org/record/13302905/files/figure.png" httpUri-1="https://zenodo.org/record/13302907/files/figure.png" pageId="29" pageNumber="476">Figs 19–20</figureCitation>
|
||||
,
|
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<tableCitation id="C6988DF8FFCAFFE2FF37468BFEB3F8BD" box="[245,319,1777,1801]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="29" pageNumber="476">Table 3</tableCitation>
|
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<tableCitation id="C6988DF8FFCAFFE2FF37468BFEB3F8BD" box="[245,319,1777,1801]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="29" pageNumber="476" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
; Supporting Information,
|
||||
<figureCitation id="1321A4C6FFCAFFE2FD8B468BFD01F8BD" box="[585,653,1777,1801]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="29" pageNumber="476">Fig. S7</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCAFFE2FD8B468BFD01F8BD" box="[585,653,1777,1801]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="29" pageNumber="476">Fig. S7</figureCitation>
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; Table S8)
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</paragraph>
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<paragraph id="8BA5B843FFCAFFE2FFB34750FE16F8D5" blockId="29.[113,762,1834,1889]" pageId="29" pageNumber="476">
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|
|
@ -198,7 +201,7 @@ in clade (B1), from which it differs by
|
|||
mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
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<emphasis id="B96E6451FFCAFFE2FC86458DFC0DF9BB" box="[836,897,1527,1551]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
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(
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<figureCitation id="1321A4C6FFCAFFE2FC50458DFC77F9BB" box="[914,1019,1527,1551]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCAFFE2FC50458DFC77F9BB" box="[914,1019,1527,1551]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||||
). The new species differs from
|
||||
<taxonomicName id="4C1AC3C0FFCAFFE2FAFF4582FA3EF9BB" ID-CoL="3WQ3T" box="[1341,1458,1527,1551]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="gutturalis">
|
||||
<emphasis id="B96E6451FFCAFFE2FAFF4582FA3EF9BB" box="[1341,1458,1527,1551]" italics="true" pageId="29" pageNumber="476">L. gutturalis</emphasis>
|
||||
|
|
@ -214,7 +217,7 @@ by
|
|||
(uncorrected p-distance) mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
|
||||
<emphasis id="B96E6451FFCAFFE2FB6A462FFB69F9D9" box="[1192,1253,1621,1645]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCAFFE2FB37462FFAD0F9D9" box="[1269,1372,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCAFFE2FB37462FFAD0F9D9" box="[1269,1372,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -349,7 +352,7 @@ and
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF65E8CBFFC9FFE1FF434412FEC7FB60" pageId="30" pageNumber="477" startId="30.[129,194,1128,1152]" targetBox="[129,1473,144,1100]" targetPageId="30" targetType="figure">
|
||||
<caption id="DF65E8CBFFC9FFE1FF434412FEC7FB60" ID-DOI="http://doi.org/10.5281/zenodo.13302899" ID-Zenodo-Dep="13302899" httpUri="https://zenodo.org/record/13302899/files/figure.png" pageId="30" pageNumber="477" startId="30.[129,194,1128,1152]" targetBox="[129,1473,144,1100]" targetPageId="30" targetType="figure">
|
||||
<paragraph id="8BA5B843FFC9FFE1FF434412FEC7FB60" blockId="30.[129,1430,1127,1236]" pageId="30" pageNumber="477">
|
||||
<emphasis id="B96E6451FFC9FFE1FF434412FF69FB34" bold="true" box="[129,229,1128,1152]" pageId="30" pageNumber="477">Figure 18.</emphasis>
|
||||
Specimens of
|
||||
|
|
@ -392,9 +395,9 @@ that is also a large-sized
|
|||
<emphasis id="B96E6451FFC9FFE1FEBE4644FE79F9E2" box="[380,501,1598,1622]" italics="true" pageId="30" pageNumber="477">L. gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup. Seven to eight supralabials and 5–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate on original tails. Large triangular mental followed by usually two (or occasionally three) symmetric postmental scales (Supporting Information,
|
||||
<figureCitation id="1321A4C6FFC9FFE1FECE46A0FED9F946" box="[268,341,1754,1778]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="30" pageNumber="477">Fig. S7</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC9FFE1FECE46A0FED9F946" box="[268,341,1754,1778]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="30" pageNumber="477">Fig. S7</figureCitation>
|
||||
). Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually denticulated posteriorly. Six terminal scansors on the tail tip. Digits elongated with five terminal scansors on the fourth toe (
|
||||
<tableCitation id="C6988DF8FFC9FFE1FDDC4742FDE4F8E4" box="[542,616,1848,1872]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="30" pageNumber="477">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFC9FFE1FDDC4742FDE4F8E4" box="[542,616,1848,1872]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="30" pageNumber="477" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFC9FFE1FF5E4722FA4FFACE" blockId="30.[129,778,1503,1967]" lastBlockId="30.[825,1475,1283,1965]" pageId="30" pageNumber="477">
|
||||
|
|
@ -524,7 +527,7 @@ Description of the
|
|||
<typeStatus id="54A106E1FFC8FFE0FE8241C5FE1EFE63" box="[320,402,447,471]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFC8FFE0FE7E41C5FD80FE63" box="[444,524,447,471]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="31" pageNumber="478">Fig. 19</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC8FFE0FE7E41C5FD80FE63" box="[444,524,447,471]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="31" pageNumber="478">Fig. 19</figureCitation>
|
||||
): Measurements and meristic characters of the
|
||||
<typeStatus id="54A106E1FFC8FFE0FEB841A4FE5AFE42" box="[378,470,478,502]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
|
||||
are presented in Supporting Information, Table S8. Adult male, with a snout–vent length (
|
||||
|
|
@ -558,11 +561,11 @@ with circular pupil. Ear to eye distance twice the orbital diameter (
|
|||
<paragraph id="8BA5B843FFC8FFDFFCE842A4FDE8FEF1" blockId="31.[810,1459,734,852]" lastBlockId="32.[129,778,144,325]" lastPageId="32" lastPageNumber="479" pageId="31" pageNumber="478">
|
||||
<emphasis id="B96E6451FFC8FFE0FCE842A4FC60FD42" box="[810,1004,734,758]" italics="true" pageId="31" pageNumber="478">Coloration: In life</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFC8FFE0FBC142A4FBC2FD42" box="[1027,1102,734,758]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="31" pageNumber="478">Fig. 20</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC8FFE0FBC142A4FBC2FD42" box="[1027,1102,734,758]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="31" pageNumber="478">Fig. 20</figureCitation>
|
||||
), dorsal coloration from light to dark brown to greyish or almost black (
|
||||
<figureCitation id="1321A4C6FFC8FFE0FB0B4287FAAFFCA1" box="[1225,1315,765,789]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="31" pageNumber="478">Fig. 20D</figureCitation>
|
||||
<figureCitation id="1321A4C6FFC8FFE0FB0B4287FAAFFCA1" box="[1225,1315,765,789]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="31" pageNumber="478">Fig. 20D</figureCitation>
|
||||
), with lighter cream-beige lines with five or six lighter cream dots surrounded by black flanks. Some specimens have lighter cream-beige dots along the vertebral line, giving them a granitic appearance (
|
||||
<figureCitation id="1321A4C6FFF7FFDFFD2140EAFF27FF73" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="32" pageNumber="479">Fig. 20A</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF7FFDFFD2140EAFF27FF73" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479">Fig. 20A</figureCitation>
|
||||
). Black line from nostril to the anterior insertion of the forelimb. Gular region white and two or three black,
|
||||
<emphasis id="B96E6451FFF7FFDFFDAE40B5FDF1FF53" bold="true" box="[620,637,207,231]" pageId="32" pageNumber="479">∩</emphasis>
|
||||
-shaped chevrons. Venter uniformly orange or white. Tip of tail and digits dark brown.
|
||||
|
|
@ -570,10 +573,10 @@ with circular pupil. Ear to eye distance twice the orbital diameter (
|
|||
(
|
||||
<typeStatus id="54A106E1FFF7FFDFFEAE4177FE5CFE91" box="[364,464,269,293]" pageId="32" pageNumber="479" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFF7FFDFFE194177FDAEFE91" box="[475,546,269,293]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="32" pageNumber="479">Fig. 19</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF7FFDFFE194177FDAEFE91" box="[475,546,269,293]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="32" pageNumber="479">Fig. 19</figureCitation>
|
||||
): dorsum dark brown and venter with uniform light cream coloration.
|
||||
</paragraph>
|
||||
<caption id="DF65E8CBFFC8FFE0FFB14710FDDAF80E" pageId="31" pageNumber="478" startId="31.[115,180,1898,1922]" targetBox="[114,1458,920,1870]" targetPageId="31" targetType="figure">
|
||||
<caption id="DF65E8CBFFC8FFE0FFB14710FDDAF80E" ID-DOI="http://doi.org/10.5281/zenodo.13302905" ID-Zenodo-Dep="13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="31" pageNumber="478" startId="31.[115,180,1898,1922]" targetBox="[114,1458,920,1870]" targetPageId="31" targetType="figure">
|
||||
<paragraph id="8BA5B843FFC8FFE0FFB14710FDDAF80E" blockId="31.[113,1443,1898,1979]" pageId="31" pageNumber="478">
|
||||
<emphasis id="B96E6451FFC8FFE0FFB14710FF54F837" bold="true" box="[115,216,1898,1923]" pageId="31" pageNumber="478">Figure 19.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -589,15 +592,15 @@ Holotype of
|
|||
<paragraph id="8BA5B843FFF7FFDFFF43411DFAEAFF53" blockId="32.[129,778,358,413]" lastBlockId="32.[825,1474,144,231]" pageId="32" pageNumber="479">
|
||||
<emphasis id="B96E6451FFF7FFDFFF43411DFF68FECA" box="[129,228,359,382]" italics="true" pageId="32" pageNumber="479">Variation:</emphasis>
|
||||
The meristic and morphometric data are summarized in Supporting Information, Table S8. Coloration of this species seems to be consistent, with gular coloration slightly lighter in females. Number and shape of postmentals and internasals vary between specimens (Supporting Information,
|
||||
<figureCitation id="1321A4C6FFF7FFDFFAD240B5FADAFF53" box="[1296,1366,207,231]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="32" pageNumber="479">Fig. S7</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF7FFDFFAD240B5FADAFF53" box="[1296,1366,207,231]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="32" pageNumber="479">Fig. S7</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF7FFDEFCFB4172FE7BFE91" blockId="32.[825,1474,263,414]" lastBlockId="33.[113,762,144,293]" lastPageId="33" lastPageNumber="480" pageId="32" pageNumber="479">
|
||||
<emphasis id="B96E6451FFF7FFDFFCFB4172FBA1FE94" box="[825,1069,264,288]" italics="true" pageId="32" pageNumber="479">Habitat and distribution</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF7FFDFFB8A4172FB0BFE94" box="[1096,1159,264,288]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="32" pageNumber="479">Figs 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF7FFDFFB8A4172FB0BFE94" box="[1096,1159,264,288]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="32" pageNumber="479">Figs 6</figureCitation>
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,
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||||
<figureCitation id="1321A4C6FFF7FFDFFB564172FB22FE94" box="[1172,1198,264,288]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="32" pageNumber="479">20</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF7FFDFFB564172FB22FE94" box="[1172,1198,264,288]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479">20</figureCitation>
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<emphasis id="B96E6451FFF7FFDFFB6C4172FA4EFEAB" box="[1198,1474,263,287]" italics="true" pageId="32" pageNumber="479">
|
||||
):
|
||||
<taxonomicName id="4C1AC3C0FFF7FFDFFB0B417DFA4EFEAB" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1225,1474,263,287]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="479" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||||
|
|
@ -621,7 +624,7 @@ also belong to this species. If confirmed, this species would have a distributio
|
|||
. However, we recommend caution until molecular data become available.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF65E8CBFFF7FFDFFF424710FDD7F80E" pageId="32" pageNumber="479" startId="32.[128,193,1898,1922]" targetBox="[129,1473,449,1870]" targetPageId="32" targetType="figure">
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<caption id="DF65E8CBFFF7FFDFFF424710FDD7F80E" ID-DOI="http://doi.org/10.5281/zenodo.13302907" ID-Zenodo-Dep="13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479" startId="32.[128,193,1898,1922]" targetBox="[129,1473,449,1870]" targetPageId="32" targetType="figure">
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<paragraph id="8BA5B843FFF7FFDFFF424710FDD7F80E" blockId="32.[128,1444,1898,1979]" pageId="32" pageNumber="479">
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<emphasis id="B96E6451FFF7FFDFFF424710FF69F837" bold="true" box="[128,229,1898,1923]" pageId="32" pageNumber="479">Figure 20.</emphasis>
|
||||
Specimens of
|
||||
|
|
@ -646,7 +649,7 @@ A diurnal and arboreal species that is always found on trees.
|
|||
.
|
||||
<quantity id="4CE215A6FFF6FFDEFEC041DFFEA5FE08" box="[258,297,421,445]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="480" unit="m" value="1.0">1 m</quantity>
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above ground in an open clearing of grass and rocky outcrops that were surrounded by dry forest (
|
||||
<figureCitation id="1321A4C6FFF6FFDEFDB541BEFD45FE68" box="[631,713,452,476]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="33" pageNumber="480">Fig. 20F</figureCitation>
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<figureCitation id="1321A4C6FFF6FFDEFDB541BEFD45FE68" box="[631,713,452,476]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="33" pageNumber="480">Fig. 20F</figureCitation>
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||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,125 +1,128 @@
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<treatment id="03B30955FFCCFFE2FC6E4462FF66F92B" ID-DOI="http://doi.org/10.5281/zenodo.13302862" ID-Zenodo-Dep="13302862" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B" httpUri="http://treatment.plazi.org/id/03B30955FFCCFFE2FC6E4462FF66F92B" lastPageId="29" lastPageNumber="476" pageId="27" pageNumber="474">
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<paragraph id="8BA5B843FFCCFFE4FC6E4462FABDFB86" blockId="27.[810,1459,1047,1110]" box="[940,1329,1047,1074]" pageId="27" pageNumber="474">
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@ -133,11 +136,11 @@
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<subSubSection id="C300EBC8FFCCFFE4FCE84444FBD7FB1B" pageId="27" pageNumber="474" type="description">
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<paragraph id="8BA5B843FFCCFFE4FCE84444FA3FFBE2" blockId="27.[810,1459,1047,1110]" box="[810,1459,1086,1110]" pageId="27" pageNumber="474">
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(
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<figureCitation id="1321A4C6FFCCFFE4FCF74444FC28FBE2" box="[821,932,1086,1110]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStartId-0="29.[113,178,1122,1146]" captionStartId-1="30.[129,194,1128,1152]" captionTargetBox-0="[114,1458,144,1094]" captionTargetBox-1="[129,1473,144,1100]" captionTargetId-0="figure-359@29.[114,1458,144,1094]" captionTargetId-1="figure-444@30.[129,1473,144,1100]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." captionText-1="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="27" pageNumber="474">Figs 17–18</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCCFFE4FCF74444FC28FBE2" box="[821,932,1086,1110]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStartId-0="29.[113,178,1122,1146]" captionStartId-1="30.[129,194,1128,1152]" captionTargetBox-0="[114,1458,144,1094]" captionTargetBox-1="[129,1473,144,1100]" captionTargetId-0="figure-359@29.[114,1458,144,1094]" captionTargetId-1="figure-444@30.[129,1473,144,1100]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." captionText-1="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." figureDoi-0="http://doi.org/10.5281/zenodo.13302897" figureDoi-1="http://doi.org/10.5281/zenodo.13302899" httpUri-0="https://zenodo.org/record/13302897/files/figure.png" httpUri-1="https://zenodo.org/record/13302899/files/figure.png" pageId="27" pageNumber="474">Figs 17–18</figureCitation>
|
||||
,
|
||||
<tableCitation id="C6988DF8FFCCFFE4FC6C4444FC74FBE2" box="[942,1016,1086,1110]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="27" pageNumber="474">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFCCFFE4FC6C4444FC74FBE2" box="[942,1016,1086,1110]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="27" pageNumber="474" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
; Supporting Information,
|
||||
<figureCitation id="1321A4C6FFCCFFE4FAC34444FACAFBE2" box="[1281,1350,1086,1110]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="27" pageNumber="474">Fig. S6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCCFFE4FAC34444FACAFBE2" box="[1281,1350,1086,1110]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="27" pageNumber="474">Fig. S6</figureCitation>
|
||||
; Table S7)
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFCCFFE4FCE84402FBD7FB1B" blockId="27.[810,1459,1144,1199]" pageId="27" pageNumber="474">
|
||||
|
|
@ -187,7 +190,7 @@ and several highlands of
|
|||
uncorrected p-distance (Table 2), and a lack of nuclear haplotype sharing in
|
||||
<emphasis id="B96E6451FFCCFFE4FC564634FC5CF9D2" box="[916,976,1614,1638]" italics="true" pageId="27" pageNumber="474">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCCFFE4FC224634FBC4F9D2" box="[992,1096,1614,1638]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="27" pageNumber="474">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCCFFE4FC224634FBC4F9D2" box="[992,1096,1614,1638]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="27" pageNumber="474">Fig. 2B–C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -382,7 +385,7 @@ A large
|
|||
<emphasis id="B96E6451FFCBFFE3FE1D421BFDDBFDCD" box="[479,599,609,633]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup. It has 7–9 supralabials and 5–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate on original tails. Large triangular mental followed by usually three (occasionally two) symmetrical postmental scales (Supporting Information,
|
||||
<figureCitation id="1321A4C6FFCBFFE3FECD4287FED7FCA1" box="[271,347,765,789]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. S6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FECD4287FED7FCA1" box="[271,347,765,789]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="28" pageNumber="475">Fig. S6</figureCitation>
|
||||
). Nostril never in contact with rostral. Ventral pholidosis with large, flattened, imbricate scales. Five to six terminal scansors on the tail tip. Digits elongated with five terminal scansors on the fourth toe (Supporting Information, Table S7).
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFCBFFE3FF5E43E0FD21FB78" blockId="28.[129,778,546,1228]" pageId="28" pageNumber="475">
|
||||
|
|
@ -509,9 +512,9 @@ based on gular pattern always having three thick
|
|||
<emphasis id="B96E6451FFCBFFE3FA8C40CAFA4EFF73" box="[1358,1474,175,199]" italics="true" pageId="28" pageNumber="475">L. depressus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCBFFE3FC8640B4FCF0FF52" box="[836,892,206,230]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." pageId="28" pageNumber="475">Fig. 4</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FC8640B4FCF0FF52" box="[836,892,206,230]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." figureDoi="http://doi.org/10.5281/zenodo.13302846" httpUri="https://zenodo.org/record/13302846/files/figure.png" pageId="28" pageNumber="475">Fig. 4</figureCitation>
|
||||
;
|
||||
<tableCitation id="C6988DF8FFCBFFE3FC4B40B4FC5FFF52" box="[905,979,206,230]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="28" pageNumber="475">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFCBFFE3FC4B40B4FC5FFF52" box="[905,979,206,230]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="28" pageNumber="475" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
). The new species occurs in mid-elevation moist forest, agricultural fields, and human habitations, vs. lowland dry sub-Saharan savannah (
|
||||
<taxonomicName id="4C1AC3C0FFCBFFE3FBF34174FB24FE91" box="[1073,1192,269,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
|
||||
<emphasis id="B96E6451FFCBFFE3FBF34174FB24FE91" box="[1073,1192,269,293]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
|
||||
|
|
@ -523,7 +526,7 @@ Basin (
|
|||
<emphasis id="B96E6451FFCBFFE3FC364157FBE9FEF0" box="[1012,1125,300,324]" italics="true" pageId="28" pageNumber="475">L. dysmicus</emphasis>
|
||||
</taxonomicName>
|
||||
; see
|
||||
<figureCitation id="1321A4C6FFCBFFE3FB554156FB43FEF0" box="[1175,1231,300,324]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FB554156FB43FEF0" box="[1175,1231,300,324]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="28" pageNumber="475">Fig. 6</figureCitation>
|
||||
). For a distinction with other species described below, see their respective diagnoses.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -553,7 +556,7 @@ Description of the
|
|||
<typeStatus id="54A106E1FFCBFFE3FBC54266FBD7FD80" box="[1031,1115,540,564]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCBFFE3FB414266FB5EFD80" box="[1155,1234,540,564]" captionStart="Figure 17" captionStartId="29.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-359@29.[114,1458,144,1094]" captionTargetPageId="29" captionText="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." pageId="28" pageNumber="475">Fig. 17</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FB414266FB5EFD80" box="[1155,1234,540,564]" captionStart="Figure 17" captionStartId="29.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-359@29.[114,1458,144,1094]" captionTargetPageId="29" captionText="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." figureDoi="http://doi.org/10.5281/zenodo.13302897" httpUri="https://zenodo.org/record/13302897/files/figure.png" pageId="28" pageNumber="475">Fig. 17</figureCitation>
|
||||
): Measurements and meristic characters of the
|
||||
<typeStatus id="54A106E1FFCBFFE3FB834241FB13FDE7" box="[1089,1183,571,595]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
|
||||
are presented in Supporting Information, Table S7. Adult male, with a snout–vent length (
|
||||
|
|
@ -587,7 +590,7 @@ and a slightly larger original tail length (TL =
|
|||
<paragraph id="8BA5B843FFCBFFE3FCFB45B2FA1BF94E" blockId="28.[825,1474,1480,1786]" pageId="28" pageNumber="475">
|
||||
<emphasis id="B96E6451FFCBFFE3FCFB45B2FC74FA54" box="[825,1016,1480,1504]" italics="true" pageId="28" pageNumber="475">Coloration: In life</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCBFFE3FBCE45B2FBDAFA55" box="[1036,1110,1480,1505]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="28" pageNumber="475">Fig. 18</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FBCE45B2FBDAFA55" box="[1036,1110,1480,1505]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." figureDoi="http://doi.org/10.5281/zenodo.13302899" httpUri="https://zenodo.org/record/13302899/files/figure.png" pageId="28" pageNumber="475">Fig. 18</figureCitation>
|
||||
), dorsal coloration brownish grey with light cream-beige lines from nape to tail on each side of the dorsum interspersed by five or six lighter cream dots surrounded by black flanks. Black line from nostril to the anterior insertion of the forelimb. Gular region with white coloration and three black
|
||||
<emphasis id="B96E6451FFCBFFE3FCFB461FFCC6F9C9" bold="true" box="[825,842,1637,1661]" pageId="28" pageNumber="475">∩</emphasis>
|
||||
-shaped chevrons. First and second chevrons in contact. Venter uniform orange from second chevrons to anterior portion of the tail and extending onto the hind limbs. Ventral surface of tail tip and digits whitish.
|
||||
|
|
@ -595,19 +598,19 @@ and a slightly larger original tail length (TL =
|
|||
(
|
||||
<typeStatus id="54A106E1FFCBFFE3FB6746B9FA85F96F" box="[1189,1289,1731,1755]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFCBFFE3FAD446B9FAD3F96F" box="[1302,1375,1731,1755]" captionStart="Figure 17" captionStartId="29.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-359@29.[114,1458,144,1094]" captionTargetPageId="29" captionText="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." pageId="28" pageNumber="475">Fig. 17</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FAD446B9FAD3F96F" box="[1302,1375,1731,1755]" captionStart="Figure 17" captionStartId="29.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-359@29.[114,1458,144,1094]" captionTargetPageId="29" captionText="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." figureDoi="http://doi.org/10.5281/zenodo.13302897" httpUri="https://zenodo.org/record/13302897/files/figure.png" pageId="28" pageNumber="475">Fig. 17</figureCitation>
|
||||
) dorsum dark brown and venter with uniform light cream coloration.
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFCBFFE2FCFB475AFE56F9BC" blockId="28.[825,1474,1823,1973]" lastBlockId="29.[113,763,1300,1544]" lastPageId="29" lastPageNumber="476" pageId="28" pageNumber="475">
|
||||
<emphasis id="B96E6451FFCBFFE3FCFB475AFC10F883" box="[825,924,1824,1847]" italics="true" pageId="28" pageNumber="475">Variation:</emphasis>
|
||||
The meristic characters of the head and body of this species are variable (see Supporting Information, Table S7). Coloration of this species seems to be consistent, with the gular coloration slightly lighter in females. First and second gular chevrons may be in contact or not (Supporting Information,
|
||||
<figureCitation id="1321A4C6FFCBFFE3FAAF47E6FA3DF800" box="[1389,1457,1948,1972]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. S6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCBFFE3FAAF47E6FA3DF800" box="[1389,1457,1948,1972]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="28" pageNumber="475">Fig. S6</figureCitation>
|
||||
).
|
||||
<emphasis id="B96E6451FFCAFFE2FFB3456FFEE2FA99" box="[113,366,1301,1325]" italics="true" pageId="29" pageNumber="476">Habitat and distribution</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFCAFFE2FE50456FFE59FA99" box="[402,469,1301,1325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="29" pageNumber="476">Figs 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCAFFE2FE50456FFE59FA99" box="[402,469,1301,1325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="29" pageNumber="476">Figs 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="1321A4C6FFCAFFE2FE25456FFD8DFA99" box="[487,513,1301,1325]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="29" pageNumber="476">18</figureCitation>
|
||||
<figureCitation id="1321A4C6FFCAFFE2FE25456FFD8DFA99" box="[487,513,1301,1325]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." figureDoi="http://doi.org/10.5281/zenodo.13302899" httpUri="https://zenodo.org/record/13302899/files/figure.png" pageId="29" pageNumber="476">18</figureCitation>
|
||||
):
|
||||
<emphasis id="B96E6451FFCAFFE2FDE2456EFF44FAF8" bold="true" pageId="29" pageNumber="476">
|
||||
<taxonomicName id="4C1AC3C0FFCAFFE2FDE2456EFD76FA98" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[544,762,1300,1324]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||||
|
|
@ -630,7 +633,7 @@ and
|
|||
, was found on a tree near a small river between secondary gallery forest and grassland.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF65E8CBFFCAFFE2FFB34418FE4AFB06" pageId="29" pageNumber="476" startId="29.[113,178,1122,1146]" targetBox="[114,1458,144,1094]" targetPageId="29" targetType="figure">
|
||||
<caption id="DF65E8CBFFCAFFE2FFB34418FE4AFB06" ID-DOI="http://doi.org/10.5281/zenodo.13302897" ID-Zenodo-Dep="13302897" httpUri="https://zenodo.org/record/13302897/files/figure.png" pageId="29" pageNumber="476" startId="29.[113,178,1122,1146]" targetBox="[114,1458,144,1094]" targetPageId="29" targetType="figure">
|
||||
<paragraph id="8BA5B843FFCAFFE2FFB34418FE4AFB06" blockId="29.[113,1442,1122,1202]" pageId="29" pageNumber="476">
|
||||
<emphasis id="B96E6451FFCAFFE2FFB34418FF5AFBCE" bold="true" box="[113,214,1122,1146]" pageId="29" pageNumber="476">Figure 17.</emphasis>
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Holotype of
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|
|
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|||
|
|
@ -1,125 +1,128 @@
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<treatment id="03B30955FFF3FFD9FF3444C6FC56F8A3" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3" httpUri="http://treatment.plazi.org/id/03B30955FFF3FFD9FF3444C6FC56F8A3" lastPageId="38" lastPageNumber="485" pageId="36" pageNumber="483">
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<treatment id="03B30955FFF3FFD9FF3444C6FC56F8A3" ID-DOI="http://doi.org/10.5281/zenodo.13302903" ID-Zenodo-Dep="13302903" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3" httpUri="http://treatment.plazi.org/id/03B30955FFF3FFD9FF3444C6FC56F8A3" lastPageId="38" lastPageNumber="485" pageId="36" pageNumber="483">
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<subSubSection id="C300EBC8FFF3FFDBFF3444C6FD18FB62" box="[246,660,1212,1238]" pageId="36" pageNumber="483" type="nomenclature">
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<paragraph id="8BA5B843FFF3FFDBFF3444C6FD18FB62" blockId="36.[246,660,1212,1238]" box="[246,660,1212,1238]" pageId="36" pageNumber="483">
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<heading id="D0ED0F2FFFF3FFDBFF3444C6FD18FB62" allCaps="true" bold="true" box="[246,660,1212,1238]" centered="true" fontSize="11" level="1" pageId="36" pageNumber="483" reason="1">
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||||
|
|
@ -135,9 +138,9 @@
|
|||
<subSubSection id="C300EBC8FFF3FFDBFF5C4499FD66FA39" pageId="36" pageNumber="483" type="description">
|
||||
<paragraph id="8BA5B843FFF3FFDBFF5C4499FD60FB4F" blockId="36.[158,748,1251,1275]" box="[158,748,1251,1275]" pageId="36" pageNumber="483">
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF3FFDBFF6B4499FE95FB4F" box="[169,281,1251,1275]" captionStart-0="Figure 23" captionStart-1="Figure 24" captionStartId-0="37.[113,178,1204,1228]" captionStartId-1="38.[129,194,1232,1256]" captionTargetBox-0="[114,1458,144,1176]" captionTargetBox-1="[129,1473,144,1205]" captionTargetId-0="figure-366@37.[114,1458,144,1176]" captionTargetId-1="figure-368@38.[129,1473,144,1205]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." captionText-1="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="36" pageNumber="483">Figs 23–24</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFF6B4499FE95FB4F" box="[169,281,1251,1275]" captionStart-0="Figure 23" captionStart-1="Figure 24" captionStartId-0="37.[113,178,1204,1228]" captionStartId-1="38.[129,194,1232,1256]" captionTargetBox-0="[114,1458,144,1176]" captionTargetBox-1="[129,1473,144,1205]" captionTargetId-0="figure-366@37.[114,1458,144,1176]" captionTargetId-1="figure-368@38.[129,1473,144,1205]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." captionText-1="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi-0="http://doi.org/10.5281/zenodo.13302917" figureDoi-1="http://doi.org/10.5281/zenodo.13302919" httpUri-0="https://zenodo.org/record/13302917/files/figure.png" httpUri-1="https://zenodo.org/record/13302919/files/figure.png" pageId="36" pageNumber="483">Figs 23–24</figureCitation>
|
||||
,
|
||||
<tableCitation id="C6988DF8FFF3FFDBFEE64499FEE3FB4F" box="[292,367,1251,1275]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="36" pageNumber="483">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFF3FFDBFEE64499FEE3FB4F" box="[292,367,1251,1275]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="36" pageNumber="483" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
; Supporting Information, Table S10)
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF3FFDBFF434566FE38FAE7" blockId="36.[129,777,1308,1363]" pageId="36" pageNumber="483">
|
||||
|
|
@ -182,11 +185,11 @@ mitochondrial gene (Table 2) from its sister taxon
|
|||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFE58462FFE61F9D9" box="[410,493,1621,1645]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE3F462FFDC7F9D9" box="[509,587,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="36" pageNumber="483">Fig. 2A</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE3F462FFDC7F9D9" box="[509,587,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="36" pageNumber="483">Fig. 2A</figureCitation>
|
||||
) and lacks nuclear haplotype sharing in
|
||||
<emphasis id="B96E6451FFF3FFDBFE94460FFE1FF939" box="[342,403,1653,1677]" italics="true" pageId="36" pageNumber="483">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE61460FFD87F939" box="[419,523,1653,1677]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="36" pageNumber="483">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE61460FFD87F939" box="[419,523,1653,1677]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="36" pageNumber="483">Fig. 2B–C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -570,7 +573,7 @@ subgroup by the following combination of characters: dorsal coloration lacking d
|
|||
<emphasis id="B96E6451FFF3FFDBFAC5465DFAF7F98A" box="[1287,1403,1574,1598]" italics="true" pageId="36" pageNumber="483">L. dysmicus</emphasis>
|
||||
</taxonomicName>
|
||||
), with two parallel dorsolateral lines from head to forelimbs, the lower line extending on to the anterior side of the forelimb, reaching the cubital fossa (
|
||||
<figureCitation id="1321A4C6FFF3FFDBFC3046FEFBC0F928" box="[1010,1100,1668,1692]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="36" pageNumber="483">Fig. 25B</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFC3046FEFBC0F928" box="[1010,1100,1668,1692]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="36" pageNumber="483">Fig. 25B</figureCitation>
|
||||
); males usually with three
|
||||
<emphasis id="B96E6451FFF3FFDBFAA046FEFAFFF928" bold="true" box="[1378,1395,1668,1692]" pageId="36" pageNumber="483">∩</emphasis>
|
||||
<emphasis id="B96E6451FFF3FFDBFAB146FFFAF6F928" box="[1395,1402,1669,1692]" italics="true" pageId="36" pageNumber="483">-</emphasis>
|
||||
|
|
@ -615,7 +618,7 @@ shaped chevrons (vs. one or two in
|
|||
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC56475BFC65F88D" box="[916,1001,1825,1849]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
always has three postmental scales, with the lateral pair separated by a posterior extension of the mental (
|
||||
<figureCitation id="1321A4C6FFF3FFDBFC864725FC2DF8CC" box="[836,929,1887,1912]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="36" pageNumber="483">Fig. 25A</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFC864725FC2DF8CC" box="[836,929,1887,1912]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="36" pageNumber="483">Fig. 25A</figureCitation>
|
||||
). It can be differentiated from
|
||||
<emphasis id="B96E6451FFF3FFDBFB1D471AFA31F8CC" bold="true" box="[1247,1469,1887,1912]" pageId="36" pageNumber="483">
|
||||
<taxonomicName id="4C1AC3C0FFF3FFDBFB1D471AFAEEF8C3" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1247,1378,1887,1911]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||||
|
|
@ -649,7 +652,7 @@ by having a proportionally larger tibia (CL/
|
|||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFE24454EFDB4FAF8" box="[486,568,1332,1356]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, females lack gular patterning (present in all the other members of the group,
|
||||
<figureCitation id="1321A4C6FFF2FFDAFD164529FF2DFA3E" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="37" pageNumber="484">Fig. 25A</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF2FFDAFD164529FF2DFA3E" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302923" httpUri="https://zenodo.org/record/13302923/files/figure.png" pageId="37" pageNumber="484">Fig. 25A</figureCitation>
|
||||
).
|
||||
<emphasis id="B96E6451FFF2FFDAFF704508FE61FA3E" bold="true" box="[178,493,1394,1418]" pageId="37" pageNumber="484">
|
||||
<taxonomicName id="4C1AC3C0FFF2FFDAFF704508FE1AFA3E" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[178,406,1394,1418]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||||
|
|
@ -665,10 +668,10 @@ can be differentiated from
|
|||
<taxonomicNameLabel id="A25DD92AFFF2FFDAFED045E8FEE8FA1E" box="[274,356,1426,1450]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
by having a lower number of precloacal pores (7–9 vs. 10) and by subtle morphometric differences (
|
||||
<tableCitation id="C6988DF8FFF2FFDAFFBE45AAFF4BFA5C" box="[124,199,1488,1512]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="37" pageNumber="484">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFF2FFDAFFBE45AAFF4BFA5C" box="[124,199,1488,1512]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="37" pageNumber="484" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF65E8CBFFF2FFDAFFB344CEFADFFAB0" pageId="37" pageNumber="484" startId="37.[113,178,1204,1228]" targetBox="[114,1458,144,1176]" targetPageId="37" targetType="figure">
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<caption id="DF65E8CBFFF2FFDAFFB344CEFADFFAB0" ID-DOI="http://doi.org/10.5281/zenodo.13302917" ID-Zenodo-Dep="13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="37" pageNumber="484" startId="37.[113,178,1204,1228]" targetBox="[114,1458,144,1176]" targetPageId="37" targetType="figure">
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<paragraph id="8BA5B843FFF2FFDAFFB344CEFADFFAB0" blockId="37.[113,1458,1203,1284]" pageId="37" pageNumber="484">
|
||||
<emphasis id="B96E6451FFF2FFDAFFB344CEFF5AFB78" bold="true" box="[113,214,1204,1228]" pageId="37" pageNumber="484">Figure 23.</emphasis>
|
||||
A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of
|
||||
|
|
@ -698,7 +701,7 @@ Description of the
|
|||
<typeStatus id="54A106E1FFF2FFDAFE8246D6FE1EF970" box="[320,402,1708,1732]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF2FFDAFE7F46D6FD81F970" box="[445,525,1708,1732]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="37" pageNumber="484">Fig. 24</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF2FFDAFE7F46D6FD81F970" box="[445,525,1708,1732]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="37" pageNumber="484">Fig. 24</figureCitation>
|
||||
): Measurements and meristic characters of the
|
||||
<typeStatus id="54A106E1FFF2FFDAFEB846B1FE59F957" box="[378,469,1739,1763]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
|
||||
are presented in Supporting Information, Table S10. Adult male, with a snout–vent length (
|
||||
|
|
@ -725,7 +728,7 @@ with circular pupil. Ear to eye distance slightly larger than the orbit diameter
|
|||
<collectionCode id="ED0B2086FFF1FFD9FD89452EFDD0FADF" box="[587,604,1364,1387]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="485" type="Museum">V</collectionCode>
|
||||
<III <IV (pes).
|
||||
</paragraph>
|
||||
<caption id="DF65E8CBFFF1FFD9FF4344AAFABCFAB0" pageId="38" pageNumber="485" startId="38.[129,194,1232,1256]" targetBox="[129,1473,144,1205]" targetPageId="38" targetType="figure">
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||||
<caption id="DF65E8CBFFF1FFD9FF4344AAFABCFAB0" ID-DOI="http://doi.org/10.5281/zenodo.13302919" ID-Zenodo-Dep="13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="38" pageNumber="485" startId="38.[129,194,1232,1256]" targetBox="[129,1473,144,1205]" targetPageId="38" targetType="figure">
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||||
<paragraph id="8BA5B843FFF1FFD9FF4344AAFABCFAB0" blockId="38.[129,1456,1232,1285]" pageId="38" pageNumber="485">
|
||||
<emphasis id="B96E6451FFF1FFD9FF4344AAFF69FB5D" bold="true" box="[129,229,1232,1257]" pageId="38" pageNumber="485">Figure 24.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -741,7 +744,7 @@ Holotype of
|
|||
<paragraph id="8BA5B843FFF1FFD9FF4345F7FD3FF96A" blockId="38.[129,778,1421,1758]" pageId="38" pageNumber="485">
|
||||
<emphasis id="B96E6451FFF1FFD9FF4345F7FEA2FA11" box="[129,302,1421,1445]" italics="true" pageId="38" pageNumber="485">Coloration: In life</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF1FFD9FEFF45F7FE52FA11" box="[317,478,1421,1445]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="38" pageNumber="485">Fig. 23A–D, F–I</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF1FFD9FEFF45F7FE52FA11" box="[317,478,1421,1445]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="38" pageNumber="485">Fig. 23A–D, F–I</figureCitation>
|
||||
), dorsal colour light brown to grey, with dark brown mottled pattern in the female; male with three rows of interrupted, but conspicuous black stripes on lateral surface from head to neck, converging at the anterior insertion of the forearm, giving the impression of a single large blotch; male and female with a black-brown line from snout to ear aperture; venter uniform orange-yellowish from posterior part of gular to cloaca, and cream to white elsewhere; gular immaculate white in female, and white in male with three thick and black
|
||||
<emphasis id="B96E6451FFF1FFD9FD6846F2FD37F914" bold="true" box="[682,699,1672,1696]" pageId="38" pageNumber="485">∩</emphasis>
|
||||
<emphasis id="B96E6451FFF1FFD9FD7946F3FD4EF914" box="[699,706,1673,1696]" italics="true" pageId="38" pageNumber="485">-</emphasis>
|
||||
|
|
@ -750,7 +753,7 @@ shaped chevrons.
|
|||
(
|
||||
<typeStatus id="54A106E1FFF1FFD9FEB946DDFE52F90B" box="[379,478,1703,1727]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFF1FFD9FE2446DDFDA6F90B" box="[486,554,1703,1727]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="38" pageNumber="485">Fig. 24</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF1FFD9FE2446DDFDA6F90B" box="[486,554,1703,1727]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302919" httpUri="https://zenodo.org/record/13302919/files/figure.png" pageId="38" pageNumber="485">Fig. 24</figureCitation>
|
||||
): similar coloration as in life, slightly darker, with gular pattern more brownish.
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF1FFD9FF43477BFC3CFA3F" blockId="38.[129,778,1792,1972]" lastBlockId="38.[825,1474,1332,1419]" pageId="38" pageNumber="485">
|
||||
|
|
@ -782,9 +785,9 @@ to this conclusion.
|
|||
<paragraph id="8BA5B843FFF1FFD9FCFB45D6FC19F9D5" blockId="38.[825,1474,1452,1633]" pageId="38" pageNumber="485">
|
||||
<emphasis id="B96E6451FFF1FFD9FCFB45D6FBA5FA70" box="[825,1065,1452,1476]" italics="true" pageId="38" pageNumber="485">Habitat and distribution</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF1FFD9FBFD45D6FBF0FA70" box="[1087,1148,1452,1476]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="38" pageNumber="485">Figs 6</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF1FFD9FBFD45D6FBF0FA70" box="[1087,1148,1452,1476]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="38" pageNumber="485">Figs 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="1321A4C6FFF1FFD9FB4A45D7FB2EFA71" box="[1160,1186,1453,1477]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="38" pageNumber="485">23</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF1FFD9FB4A45D7FB2EFA71" box="[1160,1186,1453,1477]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="38" pageNumber="485">23</figureCitation>
|
||||
):
|
||||
<emphasis id="B96E6451FFF1FFD9FB7845D6FCEBFA50" bold="true" pageId="38" pageNumber="485">
|
||||
<taxonomicName id="4C1AC3C0FFF1FFD9FB7845D6FA12FA70" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1210,1438,1452,1476]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
|
||||
|
|
|
|||
|
|
@ -1,125 +1,128 @@
|
|||
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<mods:namePart id="0534EBFB61363506DD7407D845D60877">Pinto, Pedro Vaz</mods:namePart>
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(
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<figureCitation id="1321A4C6FFF5FFDDFCAA41DFFC54FE09" box="[872,984,421,445]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStartId-0="35.[113,178,1896,1920]" captionStartId-1="37.[113,178,1204,1228]" captionTargetBox-0="[114,1458,918,1868]" captionTargetBox-1="[114,1458,144,1176]" captionTargetId-0="figure-448@35.[114,1458,918,1868]" captionTargetId-1="figure-366@37.[114,1458,144,1176]" captionTargetPageId-0="35" captionTargetPageId-1="37" captionText-0="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="34" pageNumber="481">Figs 22–23</figureCitation>
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<figureCitation id="1321A4C6FFF5FFDDFCAA41DFFC54FE09" box="[872,984,421,445]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStartId-0="35.[113,178,1896,1920]" captionStartId-1="37.[113,178,1204,1228]" captionTargetBox-0="[114,1458,918,1868]" captionTargetBox-1="[114,1458,144,1176]" captionTargetId-0="figure-448@35.[114,1458,918,1868]" captionTargetId-1="figure-366@37.[114,1458,144,1176]" captionTargetPageId-0="35" captionTargetPageId-1="37" captionText-0="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi-0="http://doi.org/10.5281/zenodo.13302915" figureDoi-1="http://doi.org/10.5281/zenodo.13302917" httpUri-0="https://zenodo.org/record/13302915/files/figure.png" httpUri-1="https://zenodo.org/record/13302917/files/figure.png" pageId="34" pageNumber="481">Figs 22–23</figureCitation>
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,
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<tableCitation id="C6988DF8FFF5FFDDFC2141DFFBA2FE09" box="[995,1070,421,445]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="34" pageNumber="481">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFF5FFDDFC2141DFFBA2FE09" box="[995,1070,421,445]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="34" pageNumber="481" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
; Supporting Information, Table S9)
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF5FFDDFCFB41A4FBE5FDA1" blockId="34.[825,1474,478,721]" pageId="34" pageNumber="481">
|
||||
|
|
@ -165,7 +168,7 @@ subgroup. It also differs by a minimum of
|
|||
uncorrected p-distance (Table 2) from related taxa, and it lacks nuclear haplotype sharing in
|
||||
<emphasis id="B96E6451FFF5FFDDFA4742E0FA4EFD06" box="[1413,1474,666,690]" italics="true" pageId="34" pageNumber="481">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF5FFDDFC8642C3FC20FD65" box="[836,940,697,721]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="34" pageNumber="481">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFC8642C3FC20FD65" box="[836,940,697,721]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="34" pageNumber="481">Fig. 2B–C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF5FFDDFCFB4289FA88FC3C" blockId="34.[825,1474,754,904]" pageId="34" pageNumber="481">
|
||||
|
|
@ -316,7 +319,7 @@ This species represents a moderately sized
|
|||
<emphasis id="B96E6451FFF5FFDDFA6C4525FC18FA21" italics="true" pageId="34" pageNumber="481">L. gutturalis</emphasis>
|
||||
</taxonomicName>
|
||||
group (described below,
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB514507FB47FA21" box="[1171,1227,1405,1429]" captionStart="Figure 3" captionStartId="14.[129,194,1167,1191]" captionTargetBox="[131,1462,158,1136]" captionTargetId="figure-369@14.[126,1476,144,1139]" captionTargetPageId="14" captionText="Figure 3. Comparative figure including dorsal (or dorsolateral) and ventral views of preserved holotypes (with exception of Lygodactylus gutturalis that is represented by a paratype) of all species within the L.gutturalis subgroup used in the current study." pageId="34" pageNumber="481">Fig. 3</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB514507FB47FA21" box="[1171,1227,1405,1429]" captionStart="Figure 3" captionStartId="14.[129,194,1167,1191]" captionTargetBox="[131,1462,158,1136]" captionTargetId="figure-369@14.[126,1476,144,1139]" captionTargetPageId="14" captionText="Figure 3. Comparative figure including dorsal (or dorsolateral) and ventral views of preserved holotypes (with exception of Lygodactylus gutturalis that is represented by a paratype) of all species within the L.gutturalis subgroup used in the current study." figureDoi="http://doi.org/10.5281/zenodo.13302842" httpUri="https://zenodo.org/record/13302842/files/figure.png" pageId="34" pageNumber="481">Fig. 3</figureCitation>
|
||||
). Females without gular pattern. Eight supralabials and 6–7 infralabials. Males with seven precloacal pores.
|
||||
</paragraph>
|
||||
<paragraph id="8BA5B843FFF5FFDCFC9745A1FDE4FF73" blockId="34.[825,1475,1311,1994]" lastBlockId="35.[113,763,144,669]" lastPageId="35" lastPageNumber="482" pageId="34" pageNumber="481">
|
||||
|
|
@ -345,7 +348,7 @@ and the unique pattern of
|
|||
<emphasis id="B96E6451FFF5FFDDFA6C46E2FC01F97A" italics="true" pageId="34" pageNumber="481">L. baptistai</emphasis>
|
||||
</taxonomicName>
|
||||
described above; see
|
||||
<figureCitation id="1321A4C6FFF5FFDDFBB746CCFB23F97A" box="[1141,1199,1718,1742]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="34" pageNumber="481">Fig. 5</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFBB746CCFB23F97A" box="[1141,1199,1718,1742]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." figureDoi="http://doi.org/10.5281/zenodo.13302850" httpUri="https://zenodo.org/record/13302850/files/figure.png" pageId="34" pageNumber="481">Fig. 5</figureCitation>
|
||||
). The species can be confused with
|
||||
<taxonomicName id="4C1AC3C0FFF5FFDDFC6446ACFB8BF95A" box="[934,1031,1750,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="heeneni">
|
||||
<emphasis id="B96E6451FFF5FFDDFC6446ACFB8BF95A" box="[934,1031,1750,1774]" italics="true" pageId="34" pageNumber="481">L. heeneni</emphasis>
|
||||
|
|
@ -366,15 +369,15 @@ based on the broken gular ornamentation; however, it can be differentiated from
|
|||
<emphasis id="B96E6451FFF5FFDDFBD9472EFBF0F8DF" box="[1051,1148,1875,1899]" italics="true" pageId="34" pageNumber="481">L. heeneni</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB494729FB4FF8DF" box="[1163,1219,1875,1899]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 7</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB494729FB4FF8DF" box="[1163,1219,1875,1899]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="34" pageNumber="481">Fig. 7</figureCitation>
|
||||
), similar to
|
||||
<taxonomicName id="4C1AC3C0FFF5FFDDFAF9472EFA4EF8DF" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[1339,1474,1875,1899]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="paurospilus">
|
||||
<emphasis id="B96E6451FFF5FFDDFAF9472EFA4EF8DF" box="[1339,1474,1875,1899]" italics="true" pageId="34" pageNumber="481">L. paurospilus</emphasis>
|
||||
</taxonomicName>
|
||||
(based on preserved specimens;
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB464708FB37F83E" box="[1156,1211,1906,1930]" captionStart="Figure 8" captionStartId="19.[115,180,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-377@19.[114,1458,144,1094]" captionTargetPageId="19" captionText="Figure 8. Paratype of L. paurospilus [RMCA (=MRAC) R.27409] from Haute Lubitshako, South Kivu Province, DRC. A, Ventral and dorsal views of body. B, Details of head in ventral, dorsal, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of right manus. Photos by J.L.R." pageId="34" pageNumber="481">Fig. 8</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFB464708FB37F83E" box="[1156,1211,1906,1930]" captionStart="Figure 8" captionStartId="19.[115,180,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-377@19.[114,1458,144,1094]" captionTargetPageId="19" captionText="Figure 8. Paratype of L. paurospilus [RMCA (=MRAC) R.27409] from Haute Lubitshako, South Kivu Province, DRC. A, Ventral and dorsal views of body. B, Details of head in ventral, dorsal, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of right manus. Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302860" httpUri="https://zenodo.org/record/13302860/files/figure.png" pageId="34" pageNumber="481">Fig. 8</figureCitation>
|
||||
; Supporting Information,
|
||||
<figureCitation id="1321A4C6FFF5FFDDFCFB47E8FC0CF81E" box="[825,896,1938,1962]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." pageId="34" pageNumber="481">Fig. S4</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFCFB47E8FC0CF81E" box="[825,896,1938,1962]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." figureDoi="http://doi.org/10.5281/zenodo.13302846" httpUri="https://zenodo.org/record/13302846/files/figure.png" pageId="34" pageNumber="481">Fig. S4</figureCitation>
|
||||
); broken pattern that converges anteriorly (vs. a broken pattern that converges posteriorly (
|
||||
<emphasis id="B96E6451FFF5FFDDFB6747C8FB39F87D" box="[1189,1205,1970,1993]" italics="true" pageId="34" pageNumber="481">V</emphasis>
|
||||
-shaped) in
|
||||
|
|
@ -501,18 +504,18 @@ Description of the
|
|||
<typeStatus id="54A106E1FFF4FFDCFE82435FFE1EFC89" box="[320,402,805,829]" pageId="35" pageNumber="482" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF4FFDCFE7E435FFD80FC89" box="[444,524,805,829]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="35" pageNumber="482">Fig. 22</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF4FFDCFE7E435FFD80FC89" box="[444,524,805,829]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302915" httpUri="https://zenodo.org/record/13302915/files/figure.png" pageId="35" pageNumber="482">Fig. 22</figureCitation>
|
||||
): Measurements and meristic characters of the
|
||||
<typeStatus id="54A106E1FFF4FFDCFEB8433EFE5AFCE8" box="[378,470,836,860]" pageId="35" pageNumber="482" type="holotype">holotype</typeStatus>
|
||||
are presented in Supporting Information, Table S9.
|
||||
<tableCitation id="C6988DF8FFF4FFDCFBDE40EAFBE5FF1C" box="[1052,1129,144,168]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="35" pageNumber="482">Table 3</tableCitation>
|
||||
<tableCitation id="C6988DF8FFF4FFDCFBDE40EAFBE5FF1C" box="[1052,1129,144,168]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="35" pageNumber="482" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||||
. Adult male, with a snout–vent length (SVL) of
|
||||
<quantity id="4CE215A6FFF4FFDCFC1B40D5FBBAFF73" box="[985,1078,175,199]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="35" pageNumber="482" unit="mm" value="32.4">32.4 mm</quantity>
|
||||
and a broken tail. Body slender and nape slightly distinct. Head as broad as body, and moderate head length (HW/HL 0.63). Canthus rostralis not prominent. Eye diameter 2.0 mm with circular pupil. Ear to eye distance almost twice the orbit diameter (
|
||||
<quantity id="4CE215A6FFF4FFDCFB424157FB5BFEF1" box="[1152,1239,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="35" pageNumber="482" unit="mm" value="3.4">3.4 mm</quantity>
|
||||
). Snout rounded and moderately pointed. Frontal granular scales larger than occipital scales, with 36 small granular scales between the eyes. Dorsal scales granular from rostral to tail. Rostral undivided, in slight contact with nostril. Three small symmetrical internasals. Eight supralabials and seven infralabials. Prenasal not in contact with the 1st supralabial. Nostril circular, in narrow contact with the rostral and bordered by 1st supralabial, prenasal, one supranasal, and one postnasal. Four rows of scales between supralabials and the orbit. Mental large, triangular, and in contact posteriorly with two rounded and symmetrical postmental scales. Five smaller post-postmental scales. Gular scales granular and smaller than ventral scales. Ventrals imbricate, rounded, and larger than dorsal scales, in 14 scale rows across the venter. Body relatively robust and slightly elongated (TRL/SVL 0.41). Seven precloacal pores. Fore- and hind limbs moderately short and stout; forearm intermediate (FL/SVL 0.15); tibia short (CL/SVL 0.17). Digits elongated, unwebbed, with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw. Relative length of digits: I <II = V <III <IV (manus); I <II <V <III <IV (pes).
|
||||
</paragraph>
|
||||
<caption id="DF65E8CBFFF4FFDCFFB34712FE80F80C" pageId="35" pageNumber="482" startId="35.[113,178,1896,1920]" targetBox="[114,1458,918,1868]" targetPageId="35" targetType="figure">
|
||||
<caption id="DF65E8CBFFF4FFDCFFB34712FE80F80C" ID-DOI="http://doi.org/10.5281/zenodo.13302915" ID-Zenodo-Dep="13302915" httpUri="https://zenodo.org/record/13302915/files/figure.png" pageId="35" pageNumber="482" startId="35.[113,178,1896,1920]" targetBox="[114,1458,918,1868]" targetPageId="35" targetType="figure">
|
||||
<paragraph id="8BA5B843FFF4FFDCFFB34712FE80F80C" blockId="35.[113,1451,1896,1977]" pageId="35" pageNumber="482">
|
||||
<emphasis id="B96E6451FFF4FFDCFFB34712FF5AF834" bold="true" box="[113,214,1896,1920]" pageId="35" pageNumber="482">Figure 22.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -530,7 +533,7 @@ Holotype of
|
|||
(
|
||||
<typeStatus id="54A106E1FFF3FFDBFE894093FE23FEB5" box="[331,431,233,257]" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE794093FDA3FEB5" box="[443,559,233,257]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">Fig. 23A–B</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFE794093FDA3FEB5" box="[443,559,233,257]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="36" pageNumber="483">Fig. 23A–B</figureCitation>
|
||||
), dorsal background colour light grey to brown, with a black reticulate dorsal pattern from snout to anterior insertion of the hind limbs; light cream blotches surrounded by black dots from eye to anterior insertion of the hind limb on each side of dorsum; tail brownish dorsally with light cream ocelli; throat white with two
|
||||
<emphasis id="B96E6451FFF3FFDBFD9841FFFDE7FE29" bold="true" box="[602,619,389,413]" pageId="36" pageNumber="483">∩</emphasis>
|
||||
<emphasis id="B96E6451FFF3FFDBFDA941FCFDFEFE29" box="[619,626,390,413]" italics="true" pageId="36" pageNumber="483">-</emphasis>
|
||||
|
|
@ -539,7 +542,7 @@ shaped broken dark chevrons; venter uniformly white.
|
|||
(
|
||||
<typeStatus id="54A106E1FFF3FFDBFD0D41DFFF3EFE68" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFF3FFDBFF7F41BEFE89FE68" box="[189,261,452,476]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="36" pageNumber="483">Fig. 22</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF3FFDBFF7F41BEFE89FE68" box="[189,261,452,476]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302915" httpUri="https://zenodo.org/record/13302915/files/figure.png" pageId="36" pageNumber="483">Fig. 22</figureCitation>
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||||
), dorsum uniformly dark grey from snout to tail. Venter with silvery appearance, becoming brownish on tail and fore- and hind limbs; black reticulated dorsal pattern visible, with two white paravertebral lines (one on each side) from eyes to midbody; digits dark brown ventrally.
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</paragraph>
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<paragraph id="8BA5B843FFF3FFDBFF434201FD75FC9B" blockId="36.[129,778,635,816]" pageId="36" pageNumber="483">
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|
|
@ -549,7 +552,7 @@ In contrast to the
|
|||
, the female
|
||||
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||||
has a dorsal coloration in life that is entirely black with an orangish tail and dorsolateral dots in paravertebral lines that dissipate posteriorly, without a gular pattern (
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||||
<figureCitation id="1321A4C6FFF3FFDBFE1242A2FDC5FD45" box="[464,585,728,753]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">Fig. 23C–D</figureCitation>
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<figureCitation id="1321A4C6FFF3FFDBFE1242A2FDC5FD45" box="[464,585,728,753]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="36" pageNumber="483">Fig. 23C–D</figureCitation>
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||||
). The lack of gular pattern is corroborated
|
||||
<emphasis id="B96E6451FFF3FFDBFEAC4283FE74FCA4" box="[366,504,761,784]" italics="true" pageId="36" pageNumber="483">in preservative</emphasis>
|
||||
, in the
|
||||
|
|
@ -566,9 +569,9 @@ has a dorsal coloration in life that is entirely black with an orangish tail and
|
|||
<paragraph id="8BA5B843FFF3FFDBFF43432AFF64FB90" blockId="36.[128,778,848,1060]" pageId="36" pageNumber="483">
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<emphasis id="B96E6451FFF3FFDBFF43432AFEE3FCDC" box="[129,367,848,872]" italics="true" pageId="36" pageNumber="483">Habitat and distribution</emphasis>
|
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(
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<figureCitation id="1321A4C6FFF3FFDBFE46432AFE4CFCDC" box="[388,448,848,872]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="36" pageNumber="483">Figs 6</figureCitation>
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<figureCitation id="1321A4C6FFF3FFDBFE46432AFE4CFCDC" box="[388,448,848,872]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="36" pageNumber="483">Figs 6</figureCitation>
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,
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||||
<figureCitation id="1321A4C6FFF3FFDBFE09432BFE69FCDD" box="[459,485,849,873]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">23</figureCitation>
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<figureCitation id="1321A4C6FFF3FFDBFE09432BFE69FCDD" box="[459,485,849,873]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." figureDoi="http://doi.org/10.5281/zenodo.13302917" httpUri="https://zenodo.org/record/13302917/files/figure.png" pageId="36" pageNumber="483">23</figureCitation>
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):
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<emphasis id="B96E6451FFF3FFDBFE3F432AFF55FC3C" bold="true" pageId="36" pageNumber="483">
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<taxonomicName id="4C1AC3C0FFF3FFDBFE3F432AFC86FCDC" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[509,778,848,872]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
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|
|
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|||
|
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@ -1,125 +1,128 @@
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@ -135,9 +138,9 @@
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<paragraph id="8BA5B843FFF6FFDEFE9F4252FD83FDF4" blockId="33.[349,527,552,576]" box="[349,527,552,576]" pageId="33" pageNumber="480">
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(
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<figureCitation id="1321A4C6FFF6FFDEFEA54252FE20FDF4" box="[359,428,552,576]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="33" pageNumber="480">Fig. 21</figureCitation>
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<figureCitation id="1321A4C6FFF6FFDEFEA54252FE20FDF4" box="[359,428,552,576]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302911" httpUri="https://zenodo.org/record/13302911/files/figure.png" pageId="33" pageNumber="480">Fig. 21</figureCitation>
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;
|
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<tableCitation id="C6988DF8FFF6FFDEFE7B4252FD88FDF4" box="[441,516,552,576]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="33" pageNumber="480">Table 3</tableCitation>
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<tableCitation id="C6988DF8FFF6FFDEFE7B4252FD88FDF4" box="[441,516,552,576]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="33" pageNumber="480" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
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)
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</paragraph>
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<paragraph id="8BA5B843FFF6FFDEFFB3421BFE20FD2C" blockId="33.[113,762,609,853]" pageId="33" pageNumber="480">
|
||||
|
|
@ -171,7 +174,7 @@ and
|
|||
(uncorrected p-distance) mitochondrial gene (Table 2), and it lacks nuclear haplotype sharing for
|
||||
<emphasis id="B96E6451FFF6FFDEFF564347FF5DFCE1" box="[148,209,829,853]" italics="true" pageId="33" pageNumber="480">RAG1</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF6FFDEFF234347FEC4FCE1" box="[225,328,829,853]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="33" pageNumber="480">Fig. 2B–C</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF6FFDEFF234347FEC4FCE1" box="[225,328,829,853]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="33" pageNumber="480">Fig. 2B–C</figureCitation>
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).
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||||
</paragraph>
|
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<paragraph id="8BA5B843FFF6FFDEFCE840EAFBAFFE91" blockId="33.[810,1459,144,293]" pageId="33" pageNumber="480">
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|
|
@ -229,9 +232,9 @@ represents a moderately sized
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|||
<emphasis id="B96E6451FFF6FFDEFB9441FCFB41FE29" box="[1110,1229,389,413]" italics="true" pageId="33" pageNumber="480">L. gutturalis</emphasis>
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</taxonomicName>
|
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group. Seven to eight supralabials and six infralabials. Large triangular mental followed by three symmetrical postmental scales (
|
||||
<figureCitation id="1321A4C6FFF6FFDEFACC41BEFADFFE68" box="[1294,1363,452,476]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="33" pageNumber="480">Fig. 21</figureCitation>
|
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<figureCitation id="1321A4C6FFF6FFDEFACC41BEFADFFE68" box="[1294,1363,452,476]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302911" httpUri="https://zenodo.org/record/13302911/files/figure.png" pageId="33" pageNumber="480">Fig. 21</figureCitation>
|
||||
). Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually denticulated posteriorly. Digits elongated with five terminal scansors under the fourth toe (
|
||||
<tableCitation id="C6988DF8FFF6FFDEFB024258FA86FD8E" box="[1216,1290,546,570]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="33" pageNumber="480">Table 3</tableCitation>
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<tableCitation id="C6988DF8FFF6FFDEFB024258FA86FD8E" box="[1216,1290,546,570]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="33" pageNumber="480" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
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).
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</paragraph>
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<paragraph id="8BA5B843FFF6FFDEFC87423BFACCFCA1" blockId="33.[810,1459,326,852]" pageId="33" pageNumber="480">
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|
|
@ -311,7 +314,7 @@ subgroup species described above as follows: more precloacal pores [ten vs. 5–
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<emphasis id="B96E6451FFF5FFDDFF4241D0FF1DFE76" bold="true" box="[128,145,426,450]" pageId="34" pageNumber="481">∩</emphasis>
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-shaped pattern in all the described species above). For comparison with other species not included above, see the diagnoses in the species descriptions below.
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</paragraph>
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|
||||
<caption id="DF65E8CBFFF6FFDEFFB34710FF48F80E" ID-DOI="http://doi.org/10.5281/zenodo.13302911" ID-Zenodo-Dep="13302911" httpUri="https://zenodo.org/record/13302911/files/figure.png" pageId="33" pageNumber="480" startId="33.[113,178,1898,1922]" targetBox="[114,1458,920,1870]" targetPageId="33" targetType="figure">
|
||||
<paragraph id="8BA5B843FFF6FFDEFFB34710FF48F80E" blockId="33.[113,1439,1898,1979]" pageId="33" pageNumber="480">
|
||||
<emphasis id="B96E6451FFF6FFDEFFB34710FF5AF837" bold="true" box="[113,214,1898,1923]" pageId="33" pageNumber="480">Figure 21.</emphasis>
|
||||
Holotype of
|
||||
|
|
@ -339,7 +342,7 @@ Description of the
|
|||
<typeStatus id="54A106E1FFF5FFDDFEEC42C3FE0CFD65" box="[302,384,697,721]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF5FFDDFE5742C3FE55FD65" box="[405,473,697,721]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 21</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFE5742C3FE55FD65" box="[405,473,697,721]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302911" httpUri="https://zenodo.org/record/13302911/files/figure.png" pageId="34" pageNumber="481">Fig. 21</figureCitation>
|
||||
): Measurements and meristic characters of the
|
||||
<typeStatus id="54A106E1FFF5FFDDFEF142A2FE03FD44" box="[307,399,728,752]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
|
||||
are presented in. Adult male, with a snout–vent length (SVL) of
|
||||
|
|
@ -357,7 +360,7 @@ with circular pupil. Ear to eye distance almost double the orbit diameter (
|
|||
(
|
||||
<typeStatus id="54A106E1FFF5FFDDFD86461AFD24F9CC" box="[580,680,1632,1656]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
<figureCitation id="1321A4C6FFF5FFDDFD73461AFD7BF9CC" box="[689,759,1632,1656]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 21</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFD73461AFD7BF9CC" box="[689,759,1632,1656]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302911" httpUri="https://zenodo.org/record/13302911/files/figure.png" pageId="34" pageNumber="481">Fig. 21</figureCitation>
|
||||
): dorsum uniform dark brown-grey with lateral black spots barely visible; black spotted marks on upper and lower lips; fore- and hind limbs slightly lighter dorsally; black line from nostril to anterior insertion of the forelimb; venter uniform light cream all along the body with gular pattern of two lines (one on each side) that do not connect anteriorly, followed by an inverted
|
||||
<emphasis id="B96E6451FFF5FFDDFD694766FD36F887" box="[683,698,1820,1843]" italics="true" pageId="34" pageNumber="481">Y</emphasis>
|
||||
-shaped pattern with middle line between the branches of the
|
||||
|
|
@ -367,7 +370,7 @@ with circular pupil. Ear to eye distance almost double the orbit diameter (
|
|||
<paragraph id="8BA5B843FFF5FFDDFF4247EEFAD5FEB1" blockId="34.[128,777,1939,1995]" lastBlockId="34.[825,1474,144,261]" pageId="34" pageNumber="481">
|
||||
<emphasis id="B96E6451FFF5FFDDFF4247EEFEE3F818" box="[128,367,1940,1964]" italics="true" pageId="34" pageNumber="481">Habitat and distribution</emphasis>
|
||||
(
|
||||
<figureCitation id="1321A4C6FFF5FFDDFE4747EEFE31F81F" box="[389,445,1939,1964]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="34" pageNumber="481">Fig. 5</figureCitation>
|
||||
<figureCitation id="1321A4C6FFF5FFDDFE4747EEFE31F81F" box="[389,445,1939,1964]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." figureDoi="http://doi.org/10.5281/zenodo.13302850" httpUri="https://zenodo.org/record/13302850/files/figure.png" pageId="34" pageNumber="481">Fig. 5</figureCitation>
|
||||
):
|
||||
<materialsCitation id="3B72B21EFFF5FFDDFE1447E9FAD9FEB1" country="Democratic Republic of the Congo" elevation="2" location="The" pageId="34" pageNumber="481" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B96E6451FFF5FFDDFE1447E9FF23F87F" bold="true" pageId="34" pageNumber="481">
|
||||
|
|
|
|||
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Reference in a new issue