406 lines
55 KiB
XML
406 lines
55 KiB
XML
<document id="93CEB63FFA962554DC35066CF7B22CA3" ID-CLB-Dataset="304558" ID-DOI="10.11646/zootaxa.5519.3.3" ID-GBIF-Dataset="420544ff-4b36-493d-8fa7-53470b12dde7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915935" ID-ZooBank="988A65E0-4B68-4E8F-B52C-112FEAE8F06C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1728586691339" checkinUser="plazi" docAuthor="Pavel, Filip & Purchart, Luboš" docDate="2024" docId="03925419FFF9FFF113E9FABDFAAFACC1" docLanguage="en" docName="zootaxa.5519.3.3.pdf" docOrigin="Zootaxa 5519 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Biflabelotillus bicolorus Pavel & Purchart, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="393" masterDocId="FFAB2C61FFFCFFF8137EFFE4FFCDAA52" masterDocTitle="Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)" masterLastPageNumber="394" masterPageNumber="384" pageNumber="389" updateTime="1729080790271" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E552259E087DBFAAB589DEF1BCB61A45">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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<mods:name id="E7F1E6D0CC788ED9CC74DAE3C659AC44" type="personal">
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<mods:namePart id="F3B1EFBF0B63DF33D1AC6D87E3F92B9D">Pavel, Filip</mods:namePart>
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<mods:namePart id="84E65EBA2690408C2CB926A51EE4CCD5">Purchart, Luboš</mods:namePart>
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<mods:title id="9D4B948CE28958816874BF92C488EA92">Zootaxa</mods:title>
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<mods:part id="F956574E1B4C9E8C57164231DA4C365A">
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<mods:date id="64D88E8D7E2D48001BF963DDDC4F3FCC">2024</mods:date>
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<mods:number id="6C10932D51BFD8E359D4BE368A2BE12C">2024-10-09</mods:number>
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<treatment id="03925419FFF9FFF113E9FABDFAAFACC1" ID-DOI="http://doi.org/10.5281/zenodo.13939793" ID-Zenodo-Dep="13939793" LSID="urn:lsid:plazi:treatment:03925419FFF9FFF113E9FABDFAAFACC1" httpUri="http://treatment.plazi.org/id/03925419FFF9FFF113E9FABDFAAFACC1" lastPageId="9" lastPageNumber="393" pageId="5" pageNumber="389">
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<subSubSection id="C321B684FFF9FFFD13E9FABDFDD2AF26" box="[151,543,1369,1396]" pageId="5" pageNumber="389" type="nomenclature">
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<paragraph id="8B84E50FFFF9FFFD13E9FABDFDD2AF26" blockId="5.[151,543,1369,1432]" box="[151,543,1369,1396]" pageId="5" pageNumber="389">
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<heading id="D0CC5263FFF9FFFD13E9FABDFDD2AF26" bold="true" box="[151,543,1369,1396]" fontSize="11" level="1" pageId="5" pageNumber="389" reason="1">
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<emphasis id="B94F391DFFF9FFFD13E9FABDFDD2AF26" bold="true" box="[151,543,1369,1396]" pageId="5" pageNumber="389">
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<taxonomicName id="4C3B9E8CFFF9FFFD13E9FABDFE77AF26" authorityName="Pavel & Purchart" authorityYear="2024" box="[151,442,1369,1396]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
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<emphasis id="B94F391DFFF9FFFD13E9FABDFE77AF26" bold="true" box="[151,442,1369,1396]" italics="true" pageId="5" pageNumber="389">Biflabelotillus bicolorus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27C8466FFF9FFFD12BFFABEFDD2AF26" box="[449,543,1370,1396]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321B684FFF9FFFD13E9FA9AFE5EAFCA" box="[151,403,1406,1432]" pageId="5" pageNumber="389" type="description">
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<paragraph id="8B84E50FFFF9FFFD13E9FA9AFE5EAFCA" blockId="5.[151,543,1369,1432]" box="[151,403,1406,1432]" pageId="5" pageNumber="389">
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<figureCitation id="1300F98AFFF9FFFD13E9FA9AFE8DAFCA" box="[151,320,1406,1432]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 3, 6, 9, 12</figureCitation>
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, 33–37
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</paragraph>
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</subSubSection>
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<subSubSection id="C321B684FFF9FFFD13E9FA22FB8BACEA" pageId="5" pageNumber="389" type="materials_examined">
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<paragraph id="8B84E50FFFF9FFFD13E9FA22FF20AC56" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="389">
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<materialsCitation id="3B53EF52FFF9FFFD13E9FA22FF24AC56" collectionCode="RN" country="Madagascar" location="Moramanga" pageId="5" pageNumber="389" specimenCode="RN 2" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
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<emphasis id="B94F391DFFF9FFFD13E9FA22FEFAAF8D" bold="true" box="[151,311,1477,1504]" pageId="5" pageNumber="389">
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<typeStatus id="54805BADFFF9FFFD13E9FA22FF1FAFB2" box="[151,210,1478,1504]" pageId="5" pageNumber="389">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F32CA59FFFF9FFFD1241FA22FE05AFB2" box="[319,456,1478,1504]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
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,
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W of
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<collectingRegion id="49FF2BEDFFF9FFFD111AFA22FD32AFB2" box="[612,767,1478,1504]" country="Madagascar" name="Antananarivo" pageId="5" pageNumber="389">Antananarivo</collectingRegion>
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,
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<quantity id="4CC348EAFFF9FFFD1072FA22FC99AFB2" box="[780,852,1478,1504]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="389" unit="km" value="20.0">20 km</quantity>
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||
E-NE of
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<location id="8EE4B3D4FFF9FFFD10BAFA22FB83AFB2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFF113E9FABDFAAFACC1:8EE4B3D4FFF9FFFD10BAFA22FB83AFB2" box="[964,1102,1478,1504]" country="Madagascar" name="Moramanga" pageId="5" pageNumber="389" stateProvince="Antananarivo">Moramanga</location>
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,
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<locationDeviation id="9A36FC2DFFF9FFFD1724FA22FADAAFB2" box="[1114,1303,1478,1505]" location="Route" pageId="5" pageNumber="389">
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<quantity id="4CC348EAFFF9FFFD1724FA22FB58AFB2" box="[1114,1173,1478,1505]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="389" unit="km" value="5.0">5 km</quantity>
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S of Route
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</locationDeviation>
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nationale 2 (
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<specimenCode id="DB9D4D74FFF9FFFD13E1FA0EFF12AC56" box="[159,223,1514,1540]" collectionCode="RN" pageId="5" pageNumber="389">RN 2</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B84E50FFFF9FFFD13B9F9EAFE62ACC6" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="389">
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<emphasis id="B94F391DFFF9FFFD13B9F9EAFEB0AC75" bold="true" box="[199,381,1549,1576]" pageId="5" pageNumber="389">Type material.</emphasis>
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<materialsCitation id="3B53EF52FFF9FFFD12F5F9EAFDF3AC22" collectionCode="DATE" country="Madagascar" location="Route" pageId="5" pageNumber="389" specimenCode="DATE 1-70" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54805BADFFF9FFFD12F5F9EAFE3AAC7A" box="[395,503,1550,1576]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
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. (
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||
<specimenCount id="9D3D2E86FFF9FFFD116AF9EAFD89AC7A" box="[532,580,1550,1576]" count="1" pageId="5" pageNumber="389" type="male">1 ♂</specimenCount>
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||
FPC): [white, printed label]
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<collectingCountry id="F32CA59FFFF9FFFD10D3F9EAFBB9AC7A" box="[941,1140,1550,1576]" name="Madagascar" pageId="5" pageNumber="389">MADAGASCAR</collectingCountry>
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EST // [white, printed + handwritten label]
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<location id="8EE4B3D4FFF9FFFD1214F9D6FE60AC1E" LSID="urn:lsid:plazi:treatment:03925419FFF9FFF113E9FABDFAAFACC1:8EE4B3D4FFF9FFFD1214F9D6FE60AC1E" box="[362,429,1586,1612]" country="Madagascar" name="Route" pageId="5" pageNumber="389">Route</location>
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ANOSIBE / P.K. 40 /
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||
<specimenCode id="DB9D4D74FFF9FFFD11D6F9D6FCE3AC1E" box="[680,814,1586,1612]" collectionCode="DATE" pageId="5" pageNumber="389">DATE 1–70</specimenCode>
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// [white, printed label] DUJARDIN-DELACOUR G. / ENTOMOLOGIE/ TANANARIVE //
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</materialsCitation>
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||
<typeStatus id="54805BADFFF9FFFD1138F9B2FD21AC22" box="[582,748,1622,1648]" pageId="5" pageNumber="389" type="holotype">HOLOTYPUS</typeStatus>
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<taxonomicName id="4C3B9E8CFFF9FFFD118AF9B2FBC9AC22" authorityName="Pavel & Purchart" authorityYear="2024" box="[756,1028,1622,1648]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
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<emphasis id="B94F391DFFF9FFFD118AF9B2FBC9AC22" box="[756,1028,1622,1648]" italics="true" pageId="5" pageNumber="389">Biflabelotillus bicolorus</emphasis>
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</taxonomicName>
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<emphasis id="B94F391DFFF9FFFD1775F9B2FBABAC22" bold="true" box="[1035,1126,1622,1648]" pageId="5" pageNumber="389">
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<taxonomicNameLabel id="A27C8466FFF9FFFD1775F9B2FBABAC22" box="[1035,1126,1622,1648]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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det. F. Pavel & L. Purchart 2024 [red, printed label].
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</paragraph>
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<paragraph id="8B84E50FFFF9FFFD13B9F979FB8BACEA" blockId="5.[151,1437,1477,2008]" box="[199,1094,1693,1720]" pageId="5" pageNumber="389">
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<emphasis id="B94F391DFFF9FFFD13B9F979FE52ACEA" bold="true" box="[199,415,1693,1720]" pageId="5" pageNumber="389">
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Condition of
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<typeStatus id="54805BADFFF9FFFD121AF97AFE54ACEA" box="[356,409,1694,1720]" pageId="5" pageNumber="389">type</typeStatus>
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.
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</emphasis>
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Right front leg missing. Right antennomeres IV-XI missing.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321B684FFF9FFF013B9F925FD21AA87" lastPageId="8" lastPageNumber="392" pageId="5" pageNumber="389" type="diagnosis">
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<paragraph id="8B84E50FFFF9FFF013B9F925FD21AA87" blockId="5.[151,1437,1477,2008]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="392" pageId="5" pageNumber="389">
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<emphasis id="B94F391DFFF9FFFD13B9F925FE1CAC89" bold="true" box="[199,465,1729,1755]" pageId="5" pageNumber="389">Differential diagnosis.</emphasis>
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Both
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||
<taxonomicName id="4C3B9E8CFFF9FFFD1162F927FD64AC8E" authorityName="Pavel & Purchart" authorityYear="2024" box="[540,681,1730,1756]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF9FFFD1162F927FD64AC8E" box="[540,681,1730,1756]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B94F391DFFF9FFFD11CDF926FCDDAC8E" bold="true" box="[691,784,1730,1756]" pageId="5" pageNumber="389">
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||
<taxonomicNameLabel id="A27C8466FFF9FFFD11CDF926FCDDAC8E" box="[691,784,1730,1756]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName id="4C3B9E8CFFF9FFFD1033F927FC2EAC8E" authorityName="Pic" authorityYear="1949" box="[845,995,1730,1756]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
|
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<emphasis id="B94F391DFFF9FFFD1033F927FC2EAC8E" box="[845,995,1730,1756]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
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</taxonomicName>
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can be distinguished reliably from
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<taxonomicName id="4C3B9E8CFFF9FFFD16FAF927FEC4AD52" authorityName="Menier" authorityYear="1979" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
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<emphasis id="B94F391DFFF9FFFD16FAF927FEC4AD52" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
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</taxonomicName>
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mainly by elytral surface, as they have smooth elytral interstriae (tubercles present in elytral interstriae in
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<taxonomicName id="4C3B9E8CFFF9FFFD13C9F8EFFE87AD76" authorityName="Menier" authorityYear="1979" box="[183,330,1803,1828]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
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||
<emphasis id="B94F391DFFF9FFFD13C9F8EFFE87AD76" box="[183,330,1803,1828]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
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), by bituberculate foveae which are lacking in apical fourth in
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<taxonomicName id="4C3B9E8CFFF9FFFD1757F8EFFB73AD76" authorityName="Pic" authorityYear="1949" box="[1065,1214,1802,1828]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
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||
<emphasis id="B94F391DFFF9FFFD1757F8EFFB73AD76" box="[1065,1214,1802,1828]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
|
||
</taxonomicName>
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and apical third of elytra in
|
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<taxonomicName id="4C3B9E8CFFF9FFFD1385F8CBFE48AD1A" authorityName="Pavel & Purchart" authorityYear="2024" box="[251,389,1838,1864]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus">
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||
<emphasis id="B94F391DFFF9FFFD1385F8CBFE48AD1A" box="[251,389,1838,1864]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
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</taxonomicName>
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, respectively (foveae cover whole surface in
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<taxonomicName id="4C3B9E8CFFF9FFFD10FDF8CBFBD8AD1A" authorityName="Menier" authorityYear="1979" box="[899,1045,1839,1864]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF9FFFD10FDF8CBFBD8AD1A" box="[899,1045,1839,1864]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
), and space between foveae which is equal or larger than their diameter (smaller in
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD11C9F8B7FC84AD3E" authorityName="Menier" authorityYear="1979" box="[695,841,1875,1900]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF9FFFD11C9F8B7FC84AD3E" box="[695,841,1875,1900]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
). Last but not least, ultimate antennomere is more or less securiform in
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD121FF893FE24ADC2" authorityName="Pavel & Purchart" authorityYear="2024" box="[353,489,1910,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF9FFFD121FF893FE24ADC2" box="[353,489,1910,1936]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF9FFFD1291F892FD8AADC2" bold="true" box="[495,583,1910,1936]" pageId="5" pageNumber="389">
|
||
<taxonomicNameLabel id="A27C8466FFF9FFFD1291F892FD8AADC2" box="[495,583,1910,1936]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD1105F893FCC1ADC2" authorityName="Pic" authorityYear="1949" box="[635,780,1910,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFF9FFFD1105F893FCC1ADC2" box="[635,780,1910,1936]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1300F98AFFF9FFFD1067F892FCB6ADC2" box="[793,891,1910,1936]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 4, 6</figureCitation>
|
||
) whereas it is elongate-oval in
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD17AFF893FAACADC2" authorityName="Menier" authorityYear="1979" box="[1233,1377,1911,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF9FFFD17AFF893FAACADC2" box="[1233,1377,1911,1936]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1300F98AFFF9FFFD1611F892FF6AADE7" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Fig. 5</figureCitation>
|
||
). The new species most resembles
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD114FF87FFD0EADE6" authorityName="Pic" authorityYear="1949" box="[561,707,1946,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFF9FFFD114FF87FFD0EADE6" box="[561,707,1946,1972]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
but differs in following characteristics:
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD17FBF87FFAC3ADE6" authorityName="Pavel & Purchart" authorityYear="2024" box="[1157,1294,1946,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF9FFFD17FBF87FFAC3ADE6" box="[1157,1294,1946,1972]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF9FFFD166BF87EFAA2ADE6" bold="true" box="[1301,1391,1946,1972]" pageId="5" pageNumber="389">
|
||
<taxonomicNameLabel id="A27C8466FFF9FFFD166BF87EFAA2ADE6" box="[1301,1391,1946,1972]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a smaller species with bicoloured elytra and non-parallel sided pronotum, the latter widest before middle (
|
||
<figureCitation id="1300F98AFFF9FFFD165BF85AFAA0AD8A" box="[1317,1389,1982,2008]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 9</figureCitation>
|
||
, 33, 34) whereas
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF1257F925FE70AC88" authorityName="Pic" authorityYear="1949" box="[297,445,1728,1754]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFFBFFFF1257F925FE70AC88" box="[297,445,1728,1754]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
is a larger species with uniformly coloured elytra and parallel sided pronotum (
|
||
<figureCitation id="1300F98AFFFBFFFF1633F924FA5AAC89" box="[1357,1431,1728,1755]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 7</figureCitation>
|
||
, 14–17). Additionally, these species can be distinguished as follows: in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF10B2F901FB95ACAC" authorityName="Pavel & Purchart" authorityYear="2024" box="[972,1112,1764,1790]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFFBFFFF10B2F901FB95ACAC" box="[972,1112,1764,1790]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFFBFFFF171FF900FB73ACAC" bold="true" box="[1121,1214,1764,1790]" pageId="7" pageNumber="391">
|
||
<taxonomicNameLabel id="A27C8466FFFBFFFF171FF900FB73ACAC" box="[1121,1214,1764,1790]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the eyes are larger and less emarginated anteriorly, the space between eyes (vertex) very narrow with HW / FW = 4.26 (
|
||
<figureCitation id="1300F98AFFFBFFFF1666F8ECFA43AD70" box="[1304,1422,1800,1826]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 9, 12</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF1262F8C9FE7DAD14" authorityName="Pic" authorityYear="1949" box="[284,432,1836,1862]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFFBFFFF1262F8C9FE7DAD14" box="[284,432,1836,1862]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
the eyes are smaller, more emarginated anteriorly and the space between eyes is much wider with HW / FW = 2.99–3.31 (
|
||
<figureCitation id="1300F98AFFFBFFFF115DF8B4FD5AAD38" box="[547,663,1872,1899]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 7, 10</figureCitation>
|
||
). The terminal antennomere of
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF1087F8B5FB4FAD38" authorityName="Pavel & Purchart" authorityYear="2024" box="[1017,1154,1872,1898]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFFBFFFF1087F8B5FB4FAD38" box="[1017,1154,1872,1898]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFFBFFFF17F7F8B4FB2EAD38" bold="true" box="[1161,1251,1872,1898]" pageId="7" pageNumber="391">
|
||
<taxonomicNameLabel id="A27C8466FFFBFFFF17F7F8B4FB2EAD38" box="[1161,1251,1872,1898]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is broadened on one side (2.75x longer than broad) and somewhat resembling the shape of a securiform shape of a palpus (
|
||
<figureCitation id="1300F98AFFFBFFFF1637F890FA43ADDC" box="[1353,1422,1908,1934]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Fig. 6</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF126BF87DFE6BADE0" authorityName="Pic" authorityYear="1949" box="[277,422,1944,1970]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFFBFFFF126BF87DFE6BADE0" box="[277,422,1944,1970]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
it is more elongate (3.25x longer than broad) and not securiform in shape (
|
||
<figureCitation id="1300F98AFFFBFFFF179CF87CFAEAADE0" box="[1250,1319,1944,1970]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Fig. 4</figureCitation>
|
||
). Shape of antennomeres III-V is different, more tooth-like widened in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF104EF859FC0CAD84" authorityName="Pic" authorityYear="1949" box="[816,961,1980,2006]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFFBFFFF104EF859FC0CAD84" box="[816,961,1980,2006]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1300F98AFFFBFFFF10B1F858FBFCAD84" box="[975,1073,1980,2006]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 1, 3</figureCitation>
|
||
). Scutellum almost squared with slightly rounded sides in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF12CDF805FDF0ADA8" authorityName="Pavel & Purchart" authorityYear="2024" box="[435,573,2016,2042]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFFBFFFF12CDF805FDF0ADA8" box="[435,573,2016,2042]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFFBFFFF113BF804FD52ADA8" bold="true" box="[581,671,2016,2042]" pageId="7" pageNumber="391">
|
||
<taxonomicNameLabel id="A27C8466FFFBFFFF113BF804FD52ADA8" box="[581,671,2016,2042]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
whereas it is almost rounded in
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF1770F805FB6CADA8" authorityName="Pic" authorityYear="1949" box="[1038,1185,2016,2042]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFFBFFFF1770F805FB6CADA8" box="[1038,1185,2016,2042]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
with the basal margin distinctly cut-out in middle (
|
||
<figureCitation id="1300F98AFFF4FFF012A4FF73FDFCAAE3" box="[474,561,151,177]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="8" pageNumber="392">Figs 19</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFF4FFF01140FF73FD93AAE3" box="[574,606,151,177]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="8" pageNumber="392">36</figureCitation>
|
||
). Prosternal inter-coxal process narrower in
|
||
<taxonomicName id="4C3B9E8CFFF4FFF0172DFF73FB10AAE3" authorityName="Pavel & Purchart" authorityYear="2024" box="[1107,1245,151,177]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF4FFF0172DFF73FB10AAE3" box="[1107,1245,151,177]" italics="true" pageId="8" pageNumber="392">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF4FFF0179AFF73FAF2AAE3" bold="true" box="[1252,1343,151,177]" pageId="8" pageNumber="392">
|
||
<taxonomicNameLabel id="A27C8466FFF4FFF0179AFF73FAF2AAE3" box="[1252,1343,151,177]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(WPC / WPP = 3.25) than in
|
||
<taxonomicName id="4C3B9E8CFFF4FFF012FDFF5FFDD8AA87" authorityName="Pic" authorityYear="1949" box="[387,533,187,213]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFF4FFF012FDFF5FFDD8AA87" box="[387,533,187,213]" italics="true" pageId="8" pageNumber="392">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
(WPC / WPP = 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF44B587FFFAFFFE13E9F873FD58ADA5" ID-DOI="http://doi.org/10.5281/zenodo.13915941" ID-Zenodo-Dep="13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="6" pageNumber="390" startId="6.[151,250,1943,1967]" targetBox="[225,1371,187,1919]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B84E50FFFFAFFFE13E9F873FD58ADA5" blockId="6.[151,1436,1943,2039]" pageId="6" pageNumber="390">
|
||
<emphasis id="B94F391DFFFAFFFE13E9F873FE85ADFD" bold="true" box="[151,328,1943,1967]" pageId="6" pageNumber="390">FIGURE 21–27.</emphasis>
|
||
<taxonomicName id="4C3B9E8CFFFAFFFE122EF873FD12ADFD" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[336,735,1943,1967]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="390" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFFAFFFE122EF873FD82ADFD" box="[336,591,1943,1967]" italics="true" pageId="6" pageNumber="390">Biflabelotillus peyrierasi</emphasis>
|
||
<bibRefCitation id="EFAA98FEFFFAFFFE1129F873FD12ADFD" author="Menier, J. J." box="[599,735,1943,1967]" pageId="6" pageNumber="390" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF44B587FFFBFFFF13E9F9D6FE20ACC0" ID-DOI="http://doi.org/10.5281/zenodo.13915943" ID-Zenodo-Dep="13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="7" pageNumber="391" startId="7.[151,250,1586,1610]" targetBox="[170,1418,184,1562]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B84E50FFFFBFFFF13E9F9D6FE20ACC0" blockId="7.[151,1436,1586,1682]" pageId="7" pageNumber="391">
|
||
<emphasis id="B94F391DFFFBFFFF13E9F9D6FE84AC18" bold="true" box="[151,329,1586,1610]" pageId="7" pageNumber="391">FIGURE 28–32.</emphasis>
|
||
<taxonomicName id="4C3B9E8CFFFBFFFF122FF9D6FD28AC18" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[337,741,1586,1610]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFFBFFFF122FF9D6FD9DAC18" box="[337,592,1586,1610]" italics="true" pageId="7" pageNumber="391">Biflabelotillus peyrierasi</emphasis>
|
||
<bibRefCitation id="EFAA98FEFFFBFFFF1127F9D6FD28AC18" author="Menier, J. J." box="[601,741,1586,1610]" pageId="7" pageNumber="391" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF44B587FFF4FFF013E9F96BFF12ACBD" ID-DOI="http://doi.org/10.5281/zenodo.13915945" ID-Zenodo-Dep="13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="8" pageNumber="392" startId="8.[151,250,1679,1703]" targetBox="[173,1409,286,1655]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B84E50FFFF4FFF013E9F96BFF12ACBD" blockId="8.[151,1436,1679,1775]" pageId="8" pageNumber="392">
|
||
<emphasis id="B94F391DFFF4FFF013E9F96BFE85ACF5" bold="true" box="[151,328,1679,1703]" pageId="8" pageNumber="392">FIGURE 33–37.</emphasis>
|
||
<taxonomicName id="4C3B9E8CFFF4FFF01231F96BFD8AACF5" authorityName="Pavel & Purchart" authorityYear="2024" box="[335,583,1679,1703]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF4FFF01231F96BFD8AACF5" box="[335,583,1679,1703]" italics="true" pageId="8" pageNumber="392">Biflabelotillus bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF4FFF01131F96BFD6CACF5" bold="true" box="[591,673,1679,1703]" pageId="8" pageNumber="392">
|
||
<taxonomicNameLabel id="A27C8466FFF4FFF01131F96BFD6CACF5" box="[591,673,1679,1703]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C321B684FFF4FFF113B9F8F9FBB3AC19" lastPageId="9" lastPageNumber="393" pageId="8" pageNumber="392" type="description">
|
||
<paragraph id="8B84E50FFFF4FFF013B9F8F9FE41AD09" blockId="8.[151,1437,1821,1991]" pageId="8" pageNumber="392">
|
||
<emphasis id="B94F391DFFF4FFF013B9F8F9FE94AD65" bold="true" box="[199,345,1821,1847]" pageId="8" pageNumber="392">Description.</emphasis>
|
||
Size of
|
||
<typeStatus id="54805BADFFF4FFF012CBF8F9FDDAAD65" box="[437,535,1821,1847]" pageId="8" pageNumber="392" type="holotype">holotype</typeStatus>
|
||
9.0 x
|
||
<quantity id="4CC348EAFFF4FFF01123F8F9FD7BAD6A" box="[605,694,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="392" unit="mm" value="2.5">2.5 mm</quantity>
|
||
. Winged, black, with apical half of elytra and tarsi rust-brown and abdomen pale-brown.
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF4FFF013B9F881FE0DAD95" blockId="8.[151,1437,1821,1991]" pageId="8" pageNumber="392">
|
||
<emphasis id="B94F391DFFF4FFF013B9F881FEC3AD2D" bold="true" box="[199,270,1893,1919]" pageId="8" pageNumber="392">Head.</emphasis>
|
||
Head including eyes broader than pronotum (DV) in ratio 1:0.7 (HW =
|
||
<quantity id="4CC348EAFFF4FFF01736F881FB7CAD2D" box="[1096,1201,1893,1919]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="8" pageNumber="392" unit="mm" value="2.13">2.13 mm</quantity>
|
||
). The space between eyes (frons) very narrow (HW / FW = 4.26) (
|
||
<figureCitation id="1300F98AFFF4FFF011F4F86DFD11ADF1" box="[650,732,1929,1955]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="8" pageNumber="392">Fig. 12</figureCitation>
|
||
). Antennae 1.47x longer than pronotum. Ultimate antennomere axe-like, widened (
|
||
<figureCitation id="1300F98AFFF4FFF01210F849FE7FAD95" box="[366,434,1965,1991]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="8" pageNumber="392">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF44B587FFF5FFF113E9FBBAFE60AEC8" ID-DOI="http://doi.org/10.5281/zenodo.13915947" ID-Zenodo-Dep="13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="9" pageNumber="393" startId="9.[151,250,1118,1142]" targetBox="[489,1099,181,1094]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B84E50FFFF5FFF113E9FBBAFE60AEC8" blockId="9.[151,1437,1118,1178]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113E9FBBAFED3AE24" bold="true" box="[151,286,1118,1142]" pageId="9" pageNumber="393">FIGURE 38.</emphasis>
|
||
Distribution map of the genus
|
||
<taxonomicName id="4C3B9E8CFFF5FFF1112EFBBAFD2DAE24" authorityName="Pic" authorityYear="1949" box="[592,736,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F391DFFF5FFF1112EFBBAFD2DAE24" box="[592,736,1118,1142]" italics="true" pageId="9" pageNumber="393">Biflabelotillus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B94F391DFFF5FFF1118EFBBBFC44AE24" box="[752,905,1118,1142]" italics="true" pageId="9" pageNumber="393">
|
||
<taxonomicName id="4C3B9E8CFFF5FFF1118EFBBBFCBDAE24" authorityName="Pic" authorityYear="1949" box="[752,880,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus">B. foveolatus</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
red triangle,
|
||
<taxonomicName id="4C3B9E8CFFF5FFF11779FBBBFB45AE24" authorityName="Menier" authorityYear="1979" box="[1031,1160,1119,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF5FFF11779FBBBFB45AE24" box="[1031,1160,1119,1142]" italics="true" pageId="9" pageNumber="393">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
—white circle,
|
||
<taxonomicName id="4C3B9E8CFFF5FFF1165CFBBBFA50AE24" authorityName="Pavel & Purchart" authorityYear="2024" box="[1314,1437,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF5FFF1165CFBBBFA50AE24" box="[1314,1437,1118,1142]" italics="true" pageId="9" pageNumber="393">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF5FFF113E9FB66FF25AEC8" bold="true" box="[151,232,1154,1178]" pageId="9" pageNumber="393">
|
||
<taxonomicNameLabel id="A27C8466FFF5FFF113E9FB66FF25AEC8" box="[151,232,1154,1178]" pageId="9" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
—yellow rhombus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B84E50FFFF5FFF113B9FB2DFA4DAF1D" blockId="9.[151,1437,1224,1683]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9FB2DFE8AAEB1" bold="true" box="[199,327,1225,1251]" pageId="9" pageNumber="393">Pronotum.</emphasis>
|
||
Longer than wide in ratio 1:1.5 (PL =
|
||
<quantity id="4CC348EAFFF5FFF1106EFB2CFCB5AEB1" box="[784,888,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="393" unit="mm" value="2.25">2.25 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CC348EAFFF5FFF110ADFB2CFBFDAEB1" box="[979,1072,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="393" unit="mm" value="1.5">1.5 mm</quantity>
|
||
), subcylindrical but not parallel sided, widest before middle, sides of pronotum narrowing in basal third from pronotal base toward the middle part of pronotum, where it is distinctly widening and then slightly narrowing toward anterior edge of pronotum, surface roughly wrinkled, irregularly in dorsal part, more or less longitudinally coarsely wrinkled in lateral part (
|
||
<figureCitation id="1300F98AFFF5FFF11653FAD0FABFAF1D" box="[1325,1394,1332,1359]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF5FFF113B9FABDFE3DAC50" blockId="9.[151,1437,1224,1683]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9FABDFED7AF21" bold="true" box="[199,282,1369,1395]" pageId="9" pageNumber="393">Elytra.</emphasis>
|
||
Subparallel, 2.7x longer than their width (EL =
|
||
<quantity id="4CC348EAFFF5FFF1103FFABCFC65AF21" box="[833,936,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" pageId="9" pageNumber="393" unit="mm" value="6.75">6.75 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CC348EAFFF5FFF1177EFABCFB96AF21" box="[1024,1115,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="393" unit="mm" value="2.5">2.5 mm</quantity>
|
||
), with posterior half slightly widening posteriad. Anterior half of elytra black, posterior half rust-brown (
|
||
<figureCitation id="1300F98AFFF5FFF11773FA98FBA8AFC5" box="[1037,1125,1404,1431]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 34</figureCitation>
|
||
). Black part of elytra with bituberculate foveae arranged in regular rows and ending at interface of black and rust-brown area, whereas the latter is smooth with only several bituberculate foveae close to black area. Space between foveae approx. 1–3x larger than diameter of foveae.
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF5FFF113B9F9E8FC49AC74" blockId="9.[151,1437,1224,1683]" box="[199,900,1548,1575]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9F9E8FE8BAC74" bold="true" box="[199,326,1548,1574]" pageId="9" pageNumber="393">Scutellum.</emphasis>
|
||
Almost square with slightly curved sides (
|
||
<figureCitation id="1300F98AFFF5FFF1105DF9E8FCB5AC74" box="[803,888,1548,1574]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF5FFF113B9F9D5FBB3AC19" blockId="9.[151,1437,1224,1683]" box="[199,1150,1584,1611]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9F9D5FEB7AC18" bold="true" box="[199,378,1584,1611]" pageId="9" pageNumber="393">Lower surface.</emphasis>
|
||
Ventrite I brown to pale-brown, ventrites II-VI pale brown (
|
||
<figureCitation id="1300F98AFFF5FFF11763F9D4FBBCAC19" box="[1053,1137,1584,1611]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF5FFF113B9F9B0FAEDAC3D" box="[199,1312,1620,1647]" pageId="9" pageNumber="393" type="distribution">
|
||
<paragraph id="8B84E50FFFF5FFF113B9F9B0FAEDAC3D" blockId="9.[151,1437,1224,1683]" box="[199,1312,1620,1647]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9F9B0FEACAC3C" bold="true" box="[199,353,1620,1646]" pageId="9" pageNumber="393">Distribution.</emphasis>
|
||
<collectingCountry id="F32CA59FFFF5FFF11219F9B0FE3DAC3C" box="[359,496,1620,1646]" name="Madagascar" pageId="9" pageNumber="393">Madagascar</collectingCountry>
|
||
, Moramanga district. Known only from the
|
||
<typeStatus id="54805BADFFF5FFF110A0F9B1FBC3AC3D" box="[990,1038,1621,1647]" pageId="9" pageNumber="393">type</typeStatus>
|
||
locality so far (
|
||
<figureCitation id="1300F98AFFF5FFF117C1F9B0FADEAC3D" box="[1215,1299,1620,1647]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="9" pageNumber="393">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF5FFF113B9F99DFAAFACC1" box="[199,1378,1656,1683]" pageId="9" pageNumber="393" type="etymology">
|
||
<paragraph id="8B84E50FFFF5FFF113B9F99DFAAFACC1" blockId="9.[151,1437,1224,1683]" box="[199,1378,1656,1683]" pageId="9" pageNumber="393">
|
||
<emphasis id="B94F391DFFF5FFF113B9F99DFE83ACC1" bold="true" box="[199,334,1657,1683]" pageId="9" pageNumber="393">Etymology.</emphasis>
|
||
Latin expression for having two colors or to be of two colors. Adjective –
|
||
<taxonomicName id="4C3B9E8CFFF5FFF117ECF99CFB31ACC0" authorityName="Pavel & Purchart" authorityYear="2024" box="[1170,1276,1656,1682]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus">
|
||
<emphasis id="B94F391DFFF5FFF117ECF99CFB31ACC0" box="[1170,1276,1656,1682]" italics="true" pageId="9" pageNumber="393">bicolorus</emphasis>
|
||
</taxonomicName>
|
||
, -
|
||
<emphasis id="B94F391DFFF5FFF1166AF99DFAEFACC0" box="[1300,1314,1657,1682]" italics="true" pageId="9" pageNumber="393">a</emphasis>
|
||
, -
|
||
<emphasis id="B94F391DFFF5FFF11647F99DFA96ACC0" box="[1337,1371,1657,1682]" italics="true" pageId="9" pageNumber="393">um</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |