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<mods:title id="BFA891EBBB6F67B0FB7C95C26103883C">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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Key to species of
<taxonomicName id="4C3B9E8CFFF5FFF11210F900FD5CACAC" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[366,657,1764,1791]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFF5FFF11210F900FDD4ACAD" bold="true" box="[366,537,1764,1791]" italics="true" pageId="9" pageNumber="393">Biflabelotillus</emphasis>
<bibRefCitation id="EFAA98FEFFF5FFF1115FF900FD5CACAC" author="Pic, M." box="[545,657,1764,1790]" pageId="9" pageNumber="393" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
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<paragraph id="8B84E50FFFF5FFF113E9F8CEFA51AD0E" blockId="9.[151,1437,1833,2031]" pageId="9" pageNumber="393">
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1. Elytra uniformly coloured (
<figureCitation id="1300F98AFFF5FFF11290F8CDFDADAD6D" box="[494,608,1833,1855]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="9" pageNumber="393">Figs 14, 17</figureCitation>
,
<figureCitation id="1300F98AFFF5FFF11115F8CEFD67AD6D" box="[619,682,1834,1855]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="9" pageNumber="393">21, 24</figureCitation>
) or at most with pale-brown humeral spot (
<figureCitation id="1300F98AFFF5FFF11729F8CDFB6CAD6D" box="[1111,1185,1833,1855]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="9" pageNumber="393">Fig. 28</figureCitation>
). Pronotum parallel-sided (
<figureCitation id="1300F98AFFF5FFF11397F8A3FEF3AD0E" box="[233,318,1863,1885]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Figs 7, 8</figureCitation>
). Space between eyes relatively broad, HW / FW &lt;4................................................. 2
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- Elytra bicoloured with anterior half black and posterior half pale brown. (
<figureCitation id="1300F98AFFF5FFF110E7F880FC2DAD2B" box="[921,992,1892,1914]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 34</figureCitation>
). Sides of pronotum not parallel, widest before middle (
<figureCitation id="1300F98AFFF5FFF1124CF865FEA0ADC5" box="[306,365,1921,1943]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Fig. 9</figureCitation>
). Space between eyes very narrow, HW / FW&gt; 4. Bituberculate foveae lacking in apical third of elytra.........................................................................................
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<taxonomicName id="4C3B9E8CFFF5FFF117AFF87BFA84ADE6" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1233,1353,1950,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF5FFF117AFF87BFA84ADE6" bold="true" box="[1233,1353,1950,1972]" italics="true" pageId="9" pageNumber="393">B. bicolorus</emphasis>
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<taxonomicNameLabel id="A27C8466FFF5FFF1162EF87BFA51ADE6" box="[1360,1436,1951,1972]" pageId="9" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
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2. Elytra pale brown, pronotum and head dark brown. Larger species with body length greater than
<quantity id="4CC348EAFFF5FFF117D2F858FB38AD80" box="[1196,1269,1980,2002]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="393" unit="mm" value="10.0">10 mm</quantity>
. Inner interstriae completely smooth, without any tubercles. Bituberculate foveae lacking in apical fourth of elytra. Space between foveae larger than their diameter..................................................................
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B.
<taxonomicName id="4C3B9E8CFFF6FFF217ABFF73FA51AAFE" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[1237,1436,151,173]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="394" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF6FFF217ABFF73FAF7AAFF" bold="true" box="[1237,1338,151,173]" italics="true" pageId="10" pageNumber="394">foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFF6FFF2163EFF73FA51AAFE" author="Pic, M." box="[1344,1436,151,173]" pageId="10" pageNumber="394" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
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- Whole body black. Smaller species with body length less than
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. All interstriae slightly but distinctly ungulated and with tubercles. Bituberculate foveae present on almost whole elytral surface. Space between foveae smaller than their diameter.................................................................................
<taxonomicName id="4C3B9E8CFFF6FFF217EAFF14FA51AB56" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[1172,1436,239,261]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="394" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF6FFF217EAFF14FA51AB56" bold="true" box="[1172,1436,239,261]" pageId="10" pageNumber="394">
<emphasis id="B94F391DFFF6FFF217EAFF14FADCAB56" bold="true" box="[1172,1297,239,261]" italics="true" pageId="10" pageNumber="394">B. peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFF6FFF21666FF0BFA51AB56" author="Menier, J. J." box="[1304,1436,239,261]" pageId="10" pageNumber="394" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
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