1297 lines
104 KiB
XML
1297 lines
104 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.98.89413" ID-Pensoft-Pub="1860-0743-2-327" ID-Pensoft-UUID="1C115BD7E10C5139A21E322F336B5A5C" ID-ZooBank="78BD2D176B3446D6816385C87B9652BF" ModsDocID="1860-0743-98-2-327" checkinTime="1662649616165" checkinUser="pensoft" docAuthor="Depra, Gabriel de Carvalho, Aguilera, Gaston, Faustino-Fuster, Dario R., Katz, Axel M. & Azevedo-Santos, Valter M." docDate="2022" docId="4923CD45E6FD55FC804F0BEEDA430F73" docLanguage="en" docName="ZoosystEvol 98(2): 327-343" docOrigin="Zoosystematics and Evolution 98 (2)" docPubDate="2022-09-08" docSource="http://dx.doi.org/10.3897/zse.98.89413" docTitle="Heptapterus carmelitanorum sp. nov., Azevedo-Santos, Deprá, Aguilera, Faustino-Fuster & Katz" docType="treatment" docVersion="2" id="1C115BD7E10C5139A21E322F336B5A5C" lastPageNumber="327" masterDocId="1C115BD7E10C5139A21E322F336B5A5C" masterDocTitle="Redefinition of Heptapterus (Heptapteridae) and description of Heptapterus carmelitanorum, a new species from the upper Parana River basin in Brazil" masterLastPageNumber="343" masterPageNumber="327" pageNumber="327" updateTime="1662649917466" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Redefinition of Heptapterus (Heptapteridae) and description of Heptapterus carmelitanorum, a new species from the upper Parana River basin in Brazil</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Depra, Gabriel de Carvalho</mods:namePart>
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<mods:affiliation>Universidade Estadual de Maringa. Av. Colombo, 5790, 87020 - 900, Maringa, Parana, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aguilera, Gaston</mods:namePart>
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<mods:affiliation>Unidad Ejecutora Lillo (CONICET) - Fundacion Miguel Lillo. Miguel Lillo 251, San Miguel de Tucuman (CP 4000), Tucuman, Argentina</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Faustino-Fuster, Dario R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-3495</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Porto Alegre, Brazil & Departamento de Ictiologia, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru & Escuela de Ingenieria y Gestion Ambiental, Facultad de Ingenieria y Gestion, Universidad Nacional Autonoma de Huanta, Huanta, Peru</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Katz, Axel M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2933-7163</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Azevedo-Santos, Valter M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8986-6406</mods:nameIdentifier>
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<mods:affiliation>Faculdade Eduvale de Avare, Avare, Sao Paulo, Brazil & Nucleo de Ecologia Aquatica e Pesca da Amazonia, Grupo de Ecologia Aquatica, Universidade Federal do Para, Belem, Para, Brazil & Programa de Pos-Graduacao em Biodiversidade, Ecologia e Conservacao, Universidade Federal do Tocantins, Porto Nacional, Tocantins, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">valter.ecologia@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-08</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>98</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>327</mods:start>
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<mods:end>343</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.98.89413</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.89413</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-327</mods:identifier>
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<mods:identifier type="ZooBank">78BD2D176B3446D6816385C87B9652BF</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1C115BD7E10C5139A21E322F336B5A5C</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="327" type="results">
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<treatment LSID="urn:lsid:zoobank.org:act:995EA984-A1D8-4A4E-8224-7D5D9637AFBF" httpUri="http://treatment.plazi.org/id/4923CD45E6FD55FC804F0BEEDA430F73" lastPageNumber="327" pageId="0" pageNumber="327">
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<subSubSection pageId="0" pageNumber="327" type="nomenclature">
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<paragraph pageId="0" pageNumber="327">
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<taxonomicName LSID="https://zoobank.org/995EA984-A1D8-4A4E-8224-7D5D9637AFBF" authority="sp. nov., Azevedo-Santos, Depra, Aguilera, Faustino-Fuster & Katz" authorityName="sp. nov., Azevedo-Santos, Deprá, Aguilera, Faustino-Fuster & Katz" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
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Heptapterus carmelitanorum sp. nov., Azevedo-Santos,
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<normalizedToken originalValue="Deprá">Depra</normalizedToken>
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, Aguilera, Faustino-Fuster & Katz
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="327" type="description">
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<paragraph pageId="0" pageNumber="327">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heptapterus carmelitanorum, new species, holotype, MNRJ 53174, 144.3 mm SL; a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/zse.98.89413.figure3" httpUri="https://binary.pensoft.net/fig/739960" pageId="0" pageNumber="327">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heptapterus carmelitanorum, new species, holotype, coloration in life." figureDoi="10.3897/zse.98.89413.figure4" httpUri="https://binary.pensoft.net/fig/739961" pageId="0" pageNumber="327">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="327" type="reference_group">
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<paragraph pageId="0" pageNumber="327">
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<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Heptapterus</taxonomicName>
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'
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<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Heptapterus</taxonomicName>
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' sp.: -
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<bibRefCitation DOI="https://doi.org/10.1590/1676-0611-bn-2018-0617" author="Azevedo-Santos, VM" journalOrPublisher="Brazilian Journal of Biology" pageId="0" pageNumber="327" refId="B8" refString="Azevedo-Santos, VM, Britski, HA, Oliveira, C, Benine, RC, 2019. Ichthyofauna of streams of the Rio Sapucai basin, upper Rio Parana system, Minas Gerais, Brazil. Biota Neotropica 19(1): e20180617. https://doi.org/10.1590/1676-0611-bn-2018-0617" title="Ichthyofauna of streams of the Rio Sapucai basin, upper Rio Parana system, Minas Gerais, Brazil. Biota Neotropica 19 (1): e 20180617." url="https://doi.org/10.1590/1676-0611-bn-2018-0617" year="2019">Azevedo-Santos et al. (2019)</bibRefCitation>
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(listed in a survey).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="327" type="holotype">
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<paragraph pageId="0" pageNumber="327">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="327">
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MNRJ 53174, 144.3 mm SL; Brazil: Minas Gerais State: limit of Carmo do Rio Claro and
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<normalizedToken originalValue="Ilicínea">Ilicinea</normalizedToken>
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municipalities: Unknown named stream tributary of
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<normalizedToken originalValue="Itací">Itaci</normalizedToken>
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stream, tributary of
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<normalizedToken originalValue="Sapucaí">Sapucai</normalizedToken>
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River (stretch flooded by Furnas reservoir), Grande River Drainage,
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<normalizedToken originalValue="Paraná">Parana</normalizedToken>
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River basin, ~
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<geoCoordinate degrees="20" direction="south" minutes="54" orientation="latitude" precision="15" seconds="57" value="-20.915833">20°54'57"S</geoCoordinate>
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,
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<geoCoordinate degrees="45" direction="west" minutes="56" orientation="longitude" precision="15" seconds="21" value="-45.939167">45°56'21"W</geoCoordinate>
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, altitude about 830 m asl; A. M. Katz and V. M. Azevedo-Santos, 31 October 2021.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="327" type="paratypes">
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<paragraph pageId="0" pageNumber="327">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="327">LBP 26570, 1, 95.7 mm SL; same locality as holotype; V. M. Azevedo-Santos and P. N. Coelho, 22 July 2017; LBP 26575, 1, 89.1 mm SL, same locality as holotype; V. M. Azevedo-Santos and P. N. Coelho, 25 May 2018; LBP 23577, 1, 104.4 mm SL, same locality as holotype; V. M. Azevedo-Santos and P. N. Coelho, 10 April 2017.</paragraph>
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<caption doi="10.3897/zse.98.89413.figure3" httpUri="https://binary.pensoft.net/fig/739960" pageId="0" pageNumber="327" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="327">
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<emphasis bold="true" pageId="0" pageNumber="327">Figure 3.</emphasis>
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<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
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<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
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</taxonomicName>
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, new species, holotype, MNRJ 53174, 144.3 mm SL;
|
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<emphasis bold="true" pageId="0" pageNumber="327">a.</emphasis>
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Dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
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Lateral view;
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<emphasis bold="true" pageId="0" pageNumber="327">c.</emphasis>
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Ventral view.
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</paragraph>
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</caption>
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<caption doi="10.3897/zse.98.89413.figure4" httpUri="https://binary.pensoft.net/fig/739961" pageId="0" pageNumber="327" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="327">
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<emphasis bold="true" pageId="0" pageNumber="327">Figure 4.</emphasis>
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<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
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<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
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</taxonomicName>
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, new species, holotype, coloration in life.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="327" type="diagnosis">
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<paragraph pageId="0" pageNumber="327">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="327">
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<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
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<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
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</taxonomicName>
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differs from all congeners by possessing the anal-fin insertion less than one eye diameter posterior to a vertical through the adipose-fin insertion (vs. more than one eye diameter posterior). From all congeners, except
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
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</taxonomicName>
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, by an isognathous mouth (vs. slightly to moderately retrognathous). It differs from all other congeners except
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
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<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
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</taxonomicName>
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, by the keel formed by ventral procurrent caudal-fin rays shallow, far from reaching anal-fin base (vs. keel formed by ventral procurrent caudal-fin rays deep, continuing almost to the anal-fin base, even though its anterior portion is devoid of fin rays) (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Schematic representation of the different degrees of proximity between the anal and caudal fins in Heptapterus. a. Keel formed by rigid connective tissue with imbedded ventral procurrent caudal-fin rays not much developed, its anterior end distant from anal-fin base (Heptapterus borodini, H. carmelitanorum and H. hollandi); b. Keel well developed, its anterior end reaching or almost reaching anal-fin base (remaining Heptapterus species)." figureDoi="10.3897/zse.98.89413.figure5" httpUri="https://binary.pensoft.net/fig/739962" pageId="0" pageNumber="327">5</figureCitation>
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). It differs from both
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
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<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
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</taxonomicName>
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by having an almost elliptical caudal fin (vs. lanceolate in
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
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</taxonomicName>
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, obliquely truncate to falcate in
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
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<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
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</taxonomicName>
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; Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Different caudal-fin shapes in Heptapterus. a. H. borodini, NUP 14882, 74.3 mm; b. H. hollandi, young; c. H. hollandi, adult, NUP 5978, 199.1 mm SL." figureDoi="10.3897/zse.98.89413.figure6" httpUri="https://binary.pensoft.net/fig/739963" pageId="0" pageNumber="327">6</figureCitation>
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), the length of its dorsal lobe 18.3-19.3% SL (vs. 24.4-43.3% SL in
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
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</taxonomicName>
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). Additionally,
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
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<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
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</taxonomicName>
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differs from all other congeners, except
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carnatus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carnatus">
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<emphasis italics="true" pageId="0" pageNumber="327">H. carnatus</emphasis>
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</taxonomicName>
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,
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mbya" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mbya">
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<emphasis italics="true" pageId="0" pageNumber="327">H. mbya</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. qenqo" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="qenqo">
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<emphasis italics="true" pageId="0" pageNumber="327">H. qenqo</emphasis>
|
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</taxonomicName>
|
|
, and some specimens of
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
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<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
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</taxonomicName>
|
|
, by having inconspicuous dorsal bars (vs. conspicuous). From
|
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<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
|
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<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
|
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</taxonomicName>
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,
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carnatus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carnatus">
|
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<emphasis italics="true" pageId="0" pageNumber="327">H. carnatus</emphasis>
|
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</taxonomicName>
|
|
,
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. exilis" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="exilis">
|
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<emphasis italics="true" pageId="0" pageNumber="327">H. exilis</emphasis>
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</taxonomicName>
|
|
,
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|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
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<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
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</taxonomicName>
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,
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mustelinus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mustelinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. mustelinus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. ornaticeps" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="ornaticeps">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. ornaticeps</emphasis>
|
|
</taxonomicName>
|
|
, by having 14-15 anal-fin rays (vs. 10-12 in
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
|
</taxonomicName>
|
|
; 18-21 in
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carnatus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carnatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. carnatus</emphasis>
|
|
</taxonomicName>
|
|
; 16-19 in
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. exilis" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="exilis">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. exilis</emphasis>
|
|
</taxonomicName>
|
|
; 18-23 in
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mustelinus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mustelinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. mustelinus</emphasis>
|
|
</taxonomicName>
|
|
; and 19 in
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. ornaticeps" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="ornaticeps">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. ornaticeps</emphasis>
|
|
</taxonomicName>
|
|
). Differs from
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. exilis" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="exilis">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. exilis</emphasis>
|
|
</taxonomicName>
|
|
by the complete lateral line (in adults), continuous to base of hypural plate (vs. incomplete, not reaching dorsal-fin insertion).
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
further differs from
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
|
</taxonomicName>
|
|
by having i,6 dorsal-fin rays (vs. i,7).
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure5" httpUri="https://binary.pensoft.net/fig/739962" pageId="0" pageNumber="327" start="Figure 5" startId="F5">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 5.</emphasis>
|
|
Schematic representation of the different degrees of proximity between the anal and caudal fins in
|
|
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<emphasis bold="true" pageId="0" pageNumber="327">a.</emphasis>
|
|
Keel formed by rigid connective tissue with imbedded ventral procurrent caudal-fin rays not much developed, its anterior end distant from anal-fin base (
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus borodini" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="borodini">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus borodini</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
|
</taxonomicName>
|
|
);
|
|
<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
|
|
Keel well developed, its anterior end reaching or almost reaching anal-fin base (remaining
|
|
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
|
|
</taxonomicName>
|
|
species).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zse.98.89413.figure6" httpUri="https://binary.pensoft.net/fig/739963" pageId="0" pageNumber="327" start="Figure 6" startId="F6">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 6.</emphasis>
|
|
Different caudal-fin shapes in
|
|
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<emphasis bold="true" pageId="0" pageNumber="327">a.</emphasis>
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. borodini" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="borodini">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. borodini</emphasis>
|
|
</taxonomicName>
|
|
, NUP 14882, 74.3 mm;
|
|
<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
|
</taxonomicName>
|
|
, young;
|
|
<emphasis bold="true" pageId="0" pageNumber="327">c.</emphasis>
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
|
|
</taxonomicName>
|
|
, adult, NUP 5978, 199.1 mm SL.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="327" type="description">
|
|
<paragraph pageId="0" pageNumber="327">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">General morphology</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heptapterus carmelitanorum, new species, holotype, MNRJ 53174, 144.3 mm SL; a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/zse.98.89413.figure3" httpUri="https://binary.pensoft.net/fig/739960" pageId="0" pageNumber="327">3</figureCitation>
|
|
-
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heptapterus carmelitanorum, new species, holotype, coloration in life." figureDoi="10.3897/zse.98.89413.figure4" httpUri="https://binary.pensoft.net/fig/739961" pageId="0" pageNumber="327">4</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7</figureCitation>
|
|
; Suppl. material 1: Figs S1-S3).
|
|
</emphasis>
|
|
Available specimens (holotype and three paratypes) ranging from 89.1-144.3 mm SL; morphometric data in Table
|
|
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data of the type specimens of Heptapterus carmelitanorum sp. nov." httpUri="http://table.plazi.org/id/D8C33DF75F8C32276B484F505F72302A" pageId="0" pageNumber="327" tableUuid="D8C33DF75F8C32276B484F505F72302A">1</tableCitation>
|
|
. General shape of body presented in photographs of preserved and live specimens. Dorsal profile convex from premaxillary symphysis to end of dorsal-fin base; slightly convex from that point to adipose-fin insertion; slightly convex along adipose-fin base. Caudal-fin base rounded. Ventral profile convex from dentary symphysis to isthmus; straight or slightly convex from that point to anal opening; straight along anal-fin base; concave from its end to caudal-fin base. In dorsal view, mouth rim gently arched, convex; lateral profile of head convex due to well-developed
|
|
<emphasis italics="true" pageId="0" pageNumber="327">adductor mandibulae</emphasis>
|
|
muscle; lateral profile of body straight to slightly convex along abdomen, tapering gently to about half adipose-fin base, then tapering more abruptly to caudal-fin base.
|
|
</paragraph>
|
|
<caption ID-Table-UUID="D8C33DF75F8C32276B484F505F72302A" httpUri="http://table.plazi.org/id/D8C33DF75F8C32276B484F505F72302A" pageId="0" pageNumber="327" start="Table 1" startId="T1">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Table 1.</emphasis>
|
|
Morphometric data of the type specimens of
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<table pageId="0" pageNumber="327">
|
|
<tr pageId="0" pageNumber="327">
|
|
<th colspan="1" isEmpty="true" pageId="0" pageNumber="327" rowspan="1">-</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">Paratype (LBP 23577)</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">Paratype (LBP 26575)</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">Paratype (LBP 26570)</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">Holotype (MNRJ 53174)</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">x</th>
|
|
<th colspan="1" pageId="0" pageNumber="327" rowspan="1">SD</th>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Total length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">121.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">106.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">114.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">144.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">121.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.296</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Standard length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">104.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">89.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">95.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">120.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">102.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">13.670</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="327" rowspan="1">-</td>
|
|
<td colspan="4" pageId="0" pageNumber="327" rowspan="1">As percentages of SL</td>
|
|
<td colspan="2" isEmpty="true" pageId="0" pageNumber="327" rowspan="1">-</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Body depth at dorsal-fin origin</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.014</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Body depth at adipose-fin origin</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Caudal-fin depth</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.017</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Body width at dorsal-fin origin</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Cleithral width</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.008</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Head length to base of supra-occipital process</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">17.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Lateral head length (to posteriormost point of opercle)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">21.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.004</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Maxillary-barbel length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">82.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">78.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">92.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">72.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">81.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.084</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Outer mental-barbel length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">44.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">44.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">52.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">45.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">46.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.041</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Inner mental-barbel length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">29.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">29.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">27.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">31.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">29.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.014</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Predorsal length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">37.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">38.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">38.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.008</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and terminus of dorsal-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">49.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">51.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">50.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">51.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">50.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and dorsal-fin distal end, adpressed</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">58.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">60.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">59.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">60.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">59.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.010</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Dorsal fin to adipose fin</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Dorsal-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.003</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of first dorsal-fin ray (unbranched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of stiffened part of first dorsal-fin ray</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">4.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">4.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">4.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">4.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of second dorsal-fin ray (first branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.015</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of third dorsal-fin ray (second branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">13.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">13.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">13.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of last dorsal-fin ray</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Prepectoral length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.002</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and terminus of pectoral-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">23.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">23.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.004</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and pectoral-fin distal end, adpressed</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">31.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">32.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">33.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">32.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">32.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of first pectoral-fin ray (unbranched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">7.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of stiffened part of first pectoral-fin ray</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">4.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of second pectoral-fin ray (first branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of third pectoral-fin ray (second branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Pectoral to pelvic-fin distance</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">21.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Prepelvic length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">38.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">38.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and terminus of pelvic-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">42.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">42.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">41.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">41.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.010</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and pelvic-fin distal end, adpressed</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">50.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">54.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">54.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">54.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">53.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.018</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between pelvic fins</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">5.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">5.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.003</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of first pelvic-fin ray (unbranched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">7.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.010</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of second pelvic-fin ray (first branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.011</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Length of third pelvic-fin ray (second branched)</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">13.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.008</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Pelvic to anal-fin distance</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">27.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.008</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Anal-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">17.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Preanal length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">64.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">69.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">68.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">68.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">68.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.021</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and terminus of anal-fin base</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">84.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">84.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">84.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">85.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">84.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">First branched anal-fin ray length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">7.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">6.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.004</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and anal-fin distal end, adpressed</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">90.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">90.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">91.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">91.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">91.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Adipose-fin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">27.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">29.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">28.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Preadipose length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">65.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">67.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">66.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">66.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">66.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and adipose-fin base end</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">93.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">94.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">96.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">95.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">95.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.014</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Adipose-fin depth</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">2.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">2.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">2.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">1.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">2.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.004</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Caudal-peduncle length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.001</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Caudal-peduncle depth at adipose-fin terminus</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.002</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Snout-anus distance</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">44.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">46.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">46.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">45.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">45.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.010</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Snout-urogenital papilla distance</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">47.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">49.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">48.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">49.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">49.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.010</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Anus-urogenital papilla distance</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">2.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">3.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.002</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Dorsal lobe of caudal fin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">18.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.004</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Ventral lobe of caudal fin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="327" rowspan="1">-</td>
|
|
<td colspan="4" pageId="0" pageNumber="327" rowspan="1">As percentages of HL (lateral)</td>
|
|
<td colspan="2" isEmpty="true" pageId="0" pageNumber="327" rowspan="1">-</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Head depth</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">46.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">41.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">43.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">43.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">43.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.020</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Head width</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">76.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">72.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">74.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">78.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">75.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.026</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Eye diameter</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">14.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.005</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Fleshy interorbital</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">15.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">16.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">17.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.017</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Bony interorbital</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">10.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Mouth gape</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">41.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">40.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">39.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">41.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">40.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.009</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Snout length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">33.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">33.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">34.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">33.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">33.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.007</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between snout tip and posterior nare</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">23.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.006</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Distance between posterior nostril and eye</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">8.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">7.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">11.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">9.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.015</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Anterior internarial width</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">23.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">25.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">21.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.026</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Posterior internarial width</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">21.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">19.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.009</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="327">
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">Intranarial length</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">20.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">21.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">24.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">22.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="327" rowspan="1">0.020</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
Head much depressed, flat dorsally and ventrally, rounded laterally. Mouth isognathous. Mouth rictus fleshy, folding ventrally, with large sub-labial groove beneath it (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7a</figureCitation>
|
|
). Lips double, i.e., divided by deep labial slit into outer and inner lip (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7b</figureCitation>
|
|
). Outer dorsal lip thickly and abundantly plicate; outer lower lip thickly, but scarcely plicate; inner dorsal and ventral lips finely and abundantly plicate (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7b</figureCitation>
|
|
). Tubular anterior nostril not reaching mouth rim. Deep skin fold surrounding entire posterior nostril, but with deep posterior notch (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7c</figureCitation>
|
|
). Maxillary barbel groove extending from base of barbel almost to the eye; in dorsal view, rim of groove almost parallel with body axis. Dorsal surface of snout with shallow depression posteriorly to posterior nostril, and elongate depression marking anterior cranial fontanel (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7b</figureCitation>
|
|
). Bulging eyes covered in thick skin with no free rim, almost completely dorsal. Base of inner mental barbel slightly anterior to that of outer mental barbel, distinctly posterior to base of maxillary barbel. Maxillary barbel reaching anterior margin of first pectoral-fin ray. Shallow cleithral skin fold immediately posterior to branchial aperture, posterior terminus medial to base of first pectoral-fin ray (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Superficial structures in Heptapterus carmelitanorum; a. Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead); b. Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips; c. Posterior nostril, evidencing shape of posterior notch (red arrowhead)." figureDoi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327">7a</figureCitation>
|
|
). Abdominal region depressed, distinctly broader than deep; in cross section, something between elliptic and rectangular. Cross section at dorsal-fin base approximately as broad as deep, between round and square. Body compressed from adipose-fin insertion to caudal fin, cross-section distinctly deeper than broad. Vertebrae 43. Ribs 9 (Suppl. material 1: Fig. S4).
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure7" httpUri="https://binary.pensoft.net/fig/739964" pageId="0" pageNumber="327" start="Figure 7" startId="F7">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 7.</emphasis>
|
|
Superficial structures in
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis bold="true" pageId="0" pageNumber="327">a.</emphasis>
|
|
Sub-labial groove (blue arrowhead) and cleithral skinfold (black arrowhead);
|
|
<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
|
|
Labial slit (blue arrowheads) and plicae on the outer (lemon arrowheads) and inner (pink arrowheads) lips;
|
|
<emphasis bold="true" pageId="0" pageNumber="327">c.</emphasis>
|
|
Posterior nostril, evidencing shape of posterior notch (red arrowhead).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">Dorsal fin distal margin convex; i,6*(4) rays (first ray rigid only basally); each branched ray with, at least, tertiary branches; thin membrane between rays. Pelvic-fin insertion at same vertical as base of second (first branched) dorsal-fin ray (2 specimens) or between bases of first and second rays (2*). Adipose fin continuous (i.e., connected) with the anteriormost ray of dorsal portion of caudal fin, originating slightly anteriorly to vertical through anal-fin insertion (distance less than one eye diameter); margin slightly convex. Caudal fin approximately elliptical, rays of dorsal half little longer than ventral ones; xiii,8,8,xi*(1) xv,7,8,xv(1), xvii,6,7,xiv(1), xvii,6,7,xvi(1) rays (Suppl. material 1: Fig. S5); thin membrane between rays. Pectoral fin approximately elliptical, with anterior rays longer than posterior ones; i,7,i(2), i,8*(2) rays on left side (first ray rigid only basally); on right side, i,7,i*(4); each branched ray with, at least, tertiary branches; thin membrane between rays. Pelvic fin approximately elliptical, with anterior rays longer than posterior ones; i,5 (4) rays on both sides; each branched ray with, at least, tertiary branches; thin membrane between rays.</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
Premaxillary toothplate about twice as wide as long, length of lateral margin slightly higher than symphyseal margin; small posterolateral projection present; about six rows of conical teeth (tooth plate virtually identical to the one in
|
|
<bibRefCitation author="Mees, GF" journalOrPublisher="Zoologische Mededeelingen (Leiden)" pageId="0" pageNumber="327" pagination="215 - 229" refId="B37" refString="Mees, GF, 1967. Freshwater fishes of Suriname: The genus Heptapterus (Pimelodidae). Zoologische Mededeelingen (Leiden) 42: 215 - 229" title="Freshwater fishes of Suriname: The genus Heptapterus (Pimelodidae)." volume="42" year="1967">Mees 1967</bibRefCitation>
|
|
, fig. 1c). External gill rakers on first arch 1+6*(3), 1+7(1). Branchiostegal rays 8(2) (Suppl. material 1: Fig. S5).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Laterosensory system</emphasis>
|
|
.
|
|
</emphasis>
|
|
Cephalic laterosensory pores as
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1780.1.1" author="Bockmann, FA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="327" pagination="1 - 54" refId="B17" refString="Bockmann, FA, Miquelarena, AM, 2008. Anatomy and phylogenetic relationships of a new catfish species from northeastern Argentina with comments on the phylogenetic relationships of the genus Rhamdella Eigenmann & Eigenmann 1888 (Siluriformes, Heptapteridae. Zootaxa 1780 (1): 1 - 54, DOI: https://doi.org/10.11646/zootaxa.1780.1.1" title="Anatomy and phylogenetic relationships of a new catfish species from northeastern Argentina with comments on the phylogenetic relationships of the genus Rhamdella Eigenmann & Eigenmann 1888 (Siluriformes, Heptapteridae." url="https://doi.org/10.11646/zootaxa.1780.1.1" volume="1780" year="2008">Bockmann and Miquelarena (2008)</bibRefCitation>
|
|
described for
|
|
<taxonomicName authorityName="Bockmann & Miquelarena" authorityYear="2008" class="Actinopterygii" family="Heptapteridae" genus="Rhamdella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhamdella cainguae" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="cainguae">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Rhamdella cainguae</emphasis>
|
|
</taxonomicName>
|
|
Bockmann & Miquelarena, 2008, except in following details (Fig.
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cephalic laterosensory system of Heptapterus carmelitanorum, new species (based on LBP 23577); abbreviations as in Bockmann and Miquelarena (2008)." figureDoi="10.3897/zse.98.89413.figure8" httpUri="https://binary.pensoft.net/fig/739965" pageId="0" pageNumber="327">8</figureCitation>
|
|
): s2+i2 pore much closer to anterior nostril (vs. at about middle of the distance between anterior and posterior nostrils); s4 pore distinctly more medial than s3 pore (vs. slightly more medial); s8 with two pores (s8a and s8p; vs. s8 with one pore); po3 with two pores (po3a and po3p; vs. po3 with one pore); pm1 pore only slightly posterior to transversal line across pm2 pore (vs. much posterior to it); pm1 directed medially, facing antimere (vs. directed ventrally); pm2 and pm3 pores facing anteroventrally (vs. posteroventrally and ventrally, respectively); pm4 and pm5 pores anteromedial to rictus (vs. posteromedial and posterior to it, respectively); pm10 pore slightly closer to po1+pm11 pore than to pm9 pore (vs. much closer to pm9 pore). Eye also more distant from i5, i6, s6, s7, and s8 pores than in
|
|
<taxonomicName lsidName="R. cainguae" pageId="0" pageNumber="327" rank="species" species="cainguae">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">R. cainguae</emphasis>
|
|
</taxonomicName>
|
|
, seemingly due to anterior displacement of eye in
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
. Lateral line continuous to hypural plate, with 43(1), 46(1), 63(1) pores, or ending on hypural plate, but with large gap between anterior and posterior portions, with 23(1) total pores (smallest specimen, LBP 26575).
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure8" httpUri="https://binary.pensoft.net/fig/739965" pageId="0" pageNumber="327" start="Figure 8" startId="F8">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 8.</emphasis>
|
|
Cephalic laterosensory system of
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
, new species (based on LBP 23577); abbreviations as in
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1780.1.1" author="Bockmann, FA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="327" pagination="1 - 54" refId="B17" refString="Bockmann, FA, Miquelarena, AM, 2008. Anatomy and phylogenetic relationships of a new catfish species from northeastern Argentina with comments on the phylogenetic relationships of the genus Rhamdella Eigenmann & Eigenmann 1888 (Siluriformes, Heptapteridae. Zootaxa 1780 (1): 1 - 54, DOI: https://doi.org/10.11646/zootaxa.1780.1.1" title="Anatomy and phylogenetic relationships of a new catfish species from northeastern Argentina with comments on the phylogenetic relationships of the genus Rhamdella Eigenmann & Eigenmann 1888 (Siluriformes, Heptapteridae." url="https://doi.org/10.11646/zootaxa.1780.1.1" volume="1780" year="2008">Bockmann and Miquelarena (2008)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
<taxonomicName genus="Olfactory" lsidName="Olfactory organ" pageId="0" pageNumber="327" rank="species" species="organ">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Olfactory organ</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
One specimen (LBP 23577) dissected with two longitudinal series of flat, triangular lamellae on right olfactory canal, each series with 32 lamellae (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heptapterus carmelitanorum, LBP 23577, dissected to show the olfactory Olfactory organ (anterior side to the right). Some of the lamellae in the outer (red arrowhead) and inner (green arrowhead) series are outlined in blue to better evidence their shape." figureDoi="10.3897/zse.98.89413.figure9" httpUri="https://binary.pensoft.net/fig/739966" pageId="0" pageNumber="327">9</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure9" httpUri="https://binary.pensoft.net/fig/739966" pageId="0" pageNumber="327" start="Figure 9" startId="F9">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 9.</emphasis>
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
, LBP 23577, dissected to show the olfactory
|
|
<taxonomicName genus="Olfactory" lsidName="Olfactory organ" pageId="0" pageNumber="327" rank="species" species="organ">Olfactory organ</taxonomicName>
|
|
(anterior side to the right). Some of the lamellae in the outer (red arrowhead) and inner (green arrowhead) series are outlined in blue to better evidence their shape.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Epidermal papillae</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
In LBP 23577, external surface of body covered with densely packed, flexible, perpendicularly protruding epidermal
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
(except lips; distal half of barbels, tubular portion of anterior nostril and skin flap of posterior nostril; center of eye; distal margin of branchiostegal membrane; and nearly entire fins). Distance between adjacent
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
~0.15 mm, equal to their maximum length. Papillae slender, rod-like on most of body (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epidermal papillae in Heptapterus carmelitanorum, LBP 23577, paratype. a, b. Slender, rod-like Epidermal papillae are distributed on most of body, such as on the dorsum, between the head and dorsal fin (a) and on the head (b. arrow shows s 6 + s 6 pore); c. Short, club-like Epidermal papillae are distributed on ventral surface of head." figureDoi="10.3897/zse.98.89413.figure10" httpUri="https://binary.pensoft.net/fig/739967" pageId="0" pageNumber="327">10a, b</figureCitation>
|
|
); short, club-like, apparently with widened distal extremity on ventral surface of head (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Epidermal papillae in Heptapterus carmelitanorum, LBP 23577, paratype. a, b. Slender, rod-like Epidermal papillae are distributed on most of body, such as on the dorsum, between the head and dorsal fin (a) and on the head (b. arrow shows s 6 + s 6 pore); c. Short, club-like Epidermal papillae are distributed on ventral surface of head." figureDoi="10.3897/zse.98.89413.figure10" httpUri="https://binary.pensoft.net/fig/739967" pageId="0" pageNumber="327">10c</figureCitation>
|
|
; widened portion possibly attached mucus). Very small
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
on anterior face of first pectoral- and pelvic-fin ray; on base of caudal-fin rays; on margin of eye; on base of tubular portion of anterior nostril; on base of skin flap of posterior nostril; on ventral half of adipose fin. Scarce, but well-developed
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
on urogenital papilla and anus. All epidermal
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
visible only after removal of body mucus.
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure10" httpUri="https://binary.pensoft.net/fig/739967" pageId="0" pageNumber="327" start="Figure 10" startId="F10">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 10.</emphasis>
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
in
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
, LBP 23577, paratype.
|
|
<emphasis bold="true" pageId="0" pageNumber="327">a, b.</emphasis>
|
|
Slender, rod-like
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
are distributed on most of body, such as on the dorsum, between the head and dorsal fin (
|
|
<emphasis bold="true" pageId="0" pageNumber="327">a</emphasis>
|
|
) and on the head (
|
|
<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
|
|
arrow shows s6+s6 pore);
|
|
<emphasis bold="true" pageId="0" pageNumber="327">c.</emphasis>
|
|
Short, club-like
|
|
<taxonomicName genus="Epidermal" lsidName="Epidermal papillae" pageId="0" pageNumber="327" rank="species" species="papillae">Epidermal papillae</taxonomicName>
|
|
are distributed on ventral surface of head.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Color in alcohol</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heptapterus carmelitanorum, new species, holotype, MNRJ 53174, 144.3 mm SL; a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/zse.98.89413.figure3" httpUri="https://binary.pensoft.net/fig/739960" pageId="0" pageNumber="327">3</figureCitation>
|
|
, Suppl. material 1: Figs S1, S2).
|
|
</emphasis>
|
|
Background color greyish-brown, grading to white towards belly and to white beige towards region between anus and anal fin, and ventral side of head; transition between brown and light beige more abrupt on head than in remainder of body. Caudal spot very faint, small, at base of dorsalmost branched caudal-fin ray; DB8 and 7 absent; DB6 through 4 inconspicuous, dark-brown (respectively, at adipose-fin insertion; midway between dorsal and adipose fins; and terminus of dorsal-fin base); DB3 present as roundish dark-brown spot immediately anterior to dorsal fin; DB2 very faint, little posterior to supraoccipital, at vertical through posterior end of pectoral-fin base; DB1 dark brown, extending to opercle; interorbital bar indistinct. Pre-orbital stripe very diffuse, dark-brown. Diffuse, dark-brown humeral spot; faint midlateral stripe present in LBP 26570 specimen; laterodorsal stripe absent.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="327">Color in life</emphasis>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="327">
|
|
Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heptapterus carmelitanorum, new species, holotype, coloration in life." figureDoi="10.3897/zse.98.89413.figure4" httpUri="https://binary.pensoft.net/fig/739961" pageId="0" pageNumber="327">4</figureCitation>
|
|
, Suppl. material 1: Fig. S3).
|
|
</emphasis>
|
|
General pattern of body dark brown, yellowish in the holotype (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heptapterus carmelitanorum, new species, holotype, coloration in life." figureDoi="10.3897/zse.98.89413.figure4" httpUri="https://binary.pensoft.net/fig/739961" pageId="0" pageNumber="327">4</figureCitation>
|
|
). Ventral region from isthmus to anal-fin insertion paler than remainder of body and somewhat pinkish, as well as cheek, branchiostegal membrane, cleithrum and lateral line. All fin rays dark brown. Adipose fin brownish yellow or dark yellowish brown. Interradial membranes of pectoral, anal and caudal fins yellow. Dorsal-fin interradial membrane hyaline, with scattered melanophores on basal third. Barbels dark brown dorsally and beige ventrally.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="327" type="ontogeny">
|
|
<paragraph pageId="0" pageNumber="327">Ontogeny.</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
Strong positive allometry in cleithral width (R2 = 0.997), head length (0.742), fleshy interorbital distance (0.809), mouth width (0.633), and dorsal caudal-fin lobe length (0.593; compare Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heptapterus carmelitanorum, new species, holotype, MNRJ 53174, 144.3 mm SL; a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/zse.98.89413.figure3" httpUri="https://binary.pensoft.net/fig/739960" pageId="0" pageNumber="327">3</figureCitation>
|
|
, Suppl. material 1: Figs S1, S2); moderate positive allometry in ventral caudal-fin lobe length (0.362); moderate negative allometry in bony interorbital distance (0.392), maxillary-barbel length (0.313), first dorsal-fin ray length (0.259), and maximum adipose-fin height (0.317); strong negative allometry in dorsal-adipose distance (0.656), first pectoral-fin ray length (0.993), and first pelvic-fin ray length (0.918). Positive allomery present in the number of branched rays in the dorsal caudal-fin lobe (R2 = 0.5712) and in the number of lateral-line pores (0.899).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="327" type="etymology">
|
|
<paragraph pageId="0" pageNumber="327">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
The specific name is a noun in apposition derived from
|
|
<taxonomicName genus="Carmelitanos" lsidName="Carmelitanos" pageId="0" pageNumber="327" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Carmelitanos</emphasis>
|
|
</taxonomicName>
|
|
(in Portuguese), the local appellation of people born or living in Carmo do Rio Claro (Minas Gerais, Brazil), the city where the species was discovered. The name is in honor of
|
|
<taxonomicName genus="Carmelitanos" lsidName="Carmelitanos" pageId="0" pageNumber="327" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Carmelitanos</emphasis>
|
|
</taxonomicName>
|
|
, especially Ana Maria Vilela Soares,
|
|
<normalizedToken originalValue="José">Jose</normalizedToken>
|
|
<normalizedToken originalValue="Cândido">Candido</normalizedToken>
|
|
de Mello Carvalho, Moara Lemos, and Carlos Roberto Bueno
|
|
<normalizedToken originalValue="Júnior">Junior</normalizedToken>
|
|
, for their contributions to biological science.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="327" type="distribution">
|
|
<paragraph pageId="0" pageNumber="327">Geographical distribution and ecological notes.</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
is recorded only from a single unnamed stream. The watercourse is a tributary of
|
|
<normalizedToken originalValue="Itací">Itaci</normalizedToken>
|
|
stream -
|
|
<normalizedToken originalValue="ribeirão">ribeirao</normalizedToken>
|
|
<normalizedToken originalValue="Itací">Itaci</normalizedToken>
|
|
, in Portuguese - which is an affluent of Furnas reservoir (in the
|
|
<normalizedToken originalValue="Sapucaí">Sapucai</normalizedToken>
|
|
River arm), Grande River basin, in the upper
|
|
<normalizedToken originalValue="Paraná">Parana</normalizedToken>
|
|
River system, in Minas Gerais State, Brazil (Figs
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Heptapterus carmelitanorum." figureDoi="10.3897/zse.98.89413.figure11" httpUri="https://binary.pensoft.net/fig/739968" pageId="0" pageNumber="327">11</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Partial view (i. e., stretch) of the stream where the type specimens of Heptapterus carmelitanorum were sampled." figureDoi="10.3897/zse.98.89413.figure12" httpUri="https://binary.pensoft.net/fig/739969" pageId="0" pageNumber="327">12</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.98.89413.figure11" httpUri="https://binary.pensoft.net/fig/739968" pageId="0" pageNumber="327" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 11.</emphasis>
|
|
Distribution of
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zse.98.89413.figure12" httpUri="https://binary.pensoft.net/fig/739969" pageId="0" pageNumber="327" start="Figure 12" startId="F12">
|
|
<paragraph pageId="0" pageNumber="327">
|
|
<emphasis bold="true" pageId="0" pageNumber="327">Figure 12.</emphasis>
|
|
Partial view (i.e., stretch) of the stream where the type specimens of
|
|
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus carmelitanorum" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
were sampled.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
The stream in which specimens of
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
were collected has its source on a mountain known as
|
|
<normalizedToken originalValue="“Chapadão”">"Chapadao"</normalizedToken>
|
|
(in Portuguese), approximately 1,300 meters a.s.l. Its cannel crosses successive falls (forming waterfalls), including one over 50 meters high. The type locality lies downstream from the waterfalls. According to the classification proposed by
|
|
<bibRefCitation author="Strahler, AN" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="327" refId="B50" refString="Strahler, AN, 1954. Quantitative geomorphology of erosional landscapes, G.-R. 19th Intern. Geol. C//IIf., Algiers, 1952, sec. 13, pt. 3, 341-354." title="Quantitative geomorphology of erosional landscapes, G. - R. 19 th Intern. Geol. C // IIf., Algiers, 1952, sec. 13, pt. 3, 341 - 354." year="1954">Strahler (1954)</bibRefCitation>
|
|
, the stream may be classified as third order. The water was extremely clear (small characids readily observed) and well oxygenated. The stream depth was shallow (not exceeding 1 meter), and its bed was completely formed by rocks. Light penetration was low during samplings. In the reach, submerged tree roots and accumulated leaves and fruits (especially
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
|
|
) formed some microhabitats for some species, notably
|
|
<taxonomicName baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1949" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" lsidName="Trichomycterus candidus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="candidus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Trichomycterus candidus</emphasis>
|
|
</taxonomicName>
|
|
(Miranda Ribeiro, 1949) and
|
|
<taxonomicName authorityName="Schubart & Gomes" authorityYear="1959" class="Actinopterygii" family="Heptapteridae" genus="Cetopsorhamdia" kingdom="Animalia" lsidName="Cetopsorhamdia iheringi" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="iheringi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Cetopsorhamdia iheringi</emphasis>
|
|
</taxonomicName>
|
|
Schubart & Gomes, 1959. The specimens of
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
were captured in environments that combined rocks (generally juxtaposed) and a more turbulent flow (see Fig.
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Partial view (i. e., stretch) of the stream where the type specimens of Heptapterus carmelitanorum were sampled." figureDoi="10.3897/zse.98.89413.figure12" httpUri="https://binary.pensoft.net/fig/739969" pageId="0" pageNumber="327">12</figureCitation>
|
|
). Observation during sampling suggests that the species is demersal.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="327">
|
|
Species collected with
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
|
|
</taxonomicName>
|
|
include
|
|
<taxonomicName lsidName="C. iheringi" pageId="0" pageNumber="327" rank="species" species="iheringi">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">C. iheringi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName baseAuthorityName="Bloch" baseAuthorityYear="1794" class="Actinopterygii" family="Erythrinidae" genus="Hoplias" kingdom="Animalia" lsidName="Hoplias malabaricus" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="malabaricus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Hoplias malabaricus</emphasis>
|
|
</taxonomicName>
|
|
(Bloch, 1794),
|
|
<taxonomicName baseAuthorityName="Eigenmann & Kennedy" baseAuthorityYear="1903" class="Actinopterygii" family="Characidae" genus="Knodus" kingdom="Animalia" lsidName="Knodus moenkhausii" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="moenkhausii">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Knodus moenkhausii</emphasis>
|
|
</taxonomicName>
|
|
(Eigenmann & Kennedy, 1903),
|
|
<taxonomicName authorityName="Chuctaya, Buhrnheim & Malabarba" authorityYear="2018" class="Actinopterygii" family="Characidae" genus="Odontostilbe" kingdom="Animalia" lsidName="Odontostilbe weitzmani" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="weitzmani">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Odontostilbe weitzmani</emphasis>
|
|
</taxonomicName>
|
|
Chuctaya,
|
|
<normalizedToken originalValue="Bührnheim">Buehrnheim</normalizedToken>
|
|
, & Malabarba, 2018,
|
|
<taxonomicName authorityName="Gunther" authorityYear="1864" class="Actinopterygii" family="Characidae" genus="Oligosarcus" kingdom="Animalia" lsidName="Oligosarcus argenteus" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="argenteus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Oligosarcus argenteus</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
|
, 1864,
|
|
<taxonomicName authorityName="Gosline" authorityYear="1947" class="Actinopterygii" family="Loricariidae" genus="Pareiorhina" kingdom="Animalia" lsidName="Pareiorhina" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Pareiorhina</emphasis>
|
|
</taxonomicName>
|
|
sp.,
|
|
<taxonomicName authorityName="Eigenmann" authorityYear="1911" class="Actinopteri" family="Characidae" genus="Psalidodon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psalidodon" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Psalidodon</emphasis>
|
|
</taxonomicName>
|
|
sp.,
|
|
<taxonomicName lsidName="T. candidus" pageId="0" pageNumber="327" rank="species" species="candidus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">T. candidus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="T. septemradiatus" pageId="0" pageNumber="327" rank="species" species="septemradiatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">T. septemradiatus</emphasis>
|
|
</taxonomicName>
|
|
Katz, Barbosa & Costa, 2013 (
|
|
<bibRefCitation DOI="https://doi.org/10.1590/1676-0611-bn-2018-0617" author="Azevedo-Santos, VM" journalOrPublisher="Brazilian Journal of Biology" pageId="0" pageNumber="327" refId="B8" refString="Azevedo-Santos, VM, Britski, HA, Oliveira, C, Benine, RC, 2019. Ichthyofauna of streams of the Rio Sapucai basin, upper Rio Parana system, Minas Gerais, Brazil. Biota Neotropica 19(1): e20180617. https://doi.org/10.1590/1676-0611-bn-2018-0617" title="Ichthyofauna of streams of the Rio Sapucai basin, upper Rio Parana system, Minas Gerais, Brazil. Biota Neotropica 19 (1): e 20180617." url="https://doi.org/10.1590/1676-0611-bn-2018-0617" year="2019">Azevedo-Santos et al. 2019</bibRefCitation>
|
|
). New collections in the same reach resulted in the capture of additional species, such as
|
|
<taxonomicName authorityName="C.H.Eigenmann" authorityYear="1916" class="Actinopterygii" family="Parodontidae" genus="Apareiodon" kingdom="Animalia" lsidName="Apareiodon" order="Characiformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Apareiodon</emphasis>
|
|
</taxonomicName>
|
|
sp. (CICCAA06610) and
|
|
<taxonomicName authorityName="Haseman" authorityYear="1911" class="Actinopterygii" family="Heptapteridae" genus="Rhamdiopsis" kingdom="Animalia" lsidName="Rhamdiopsis" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Rhamdiopsis</emphasis>
|
|
</taxonomicName>
|
|
sp. (CICCAA06611). In addition to fishes, aquatic spiders (e.g.,
|
|
<taxonomicName class="Insecta" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha" order="Trichoptera" pageId="0" pageNumber="327" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="327">Tetragnatha</emphasis>
|
|
</taxonomicName>
|
|
sp.) and insects, including specimens of the order
|
|
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="" order="Trichoptera" pageId="0" pageNumber="327" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
|
|
in cases formed by small gravels, were captured in the stretch.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |