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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng &amp; Schilthuizen, Menno" docDate="2015" docId="929540D335CBCB6C3D4B6FF259639821" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Microcystina microrhynchus Vermeulen, Liew &amp; Schilthuizen, sp. n." docType="treatment" docUuid="5192A533-FC42-49F4-8FE5-A354D53EB8AF" docUuidSource="ZooBank" docVersion="10" lastPageNumber="31" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="28" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liew, Thor-Seng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>531</mods:number>
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<mods:start>1</mods:start>
<mods:end>139</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="144057568" LSID="urn:lsid:zoobank.org:act:5192A533-FC42-49F4-8FE5-A354D53EB8AF" httpUri="http://treatment.plazi.org/id/929540D335CBCB6C3D4B6FF259639821" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
<subSubSection pageId="27" pageNumber="28" type="multiple">
<paragraph pageId="27" pageNumber="28">Taxon classification Animalia Stylommatophora Ariophantidae</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="nomenclature">
<paragraph pageId="27" pageNumber="28">
<taxonomicName LSID="http://zoobank.org/5192A533-FC42-49F4-8FE5-A354D53EB8AF" authority="Vermeulen, Liew &amp; Schilthuizen" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina microrhynchus" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="microrhynchus">Microcystina microrhynchus Vermeulen, Liew &amp; Schilthuizen</taxonomicName>
<taxonomicNameLabel pageId="27" pageNumber="28">sp. n.</taxonomicNameLabel>
Figure 24
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="reference_group">
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina microrhynchus" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="microrhynchus">Microcystina microrhynchus</taxonomicName>
nomen nudum,
<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="27" pageNumber="28" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
: 2761-2762;
<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of the Geological Society, London" pageId="27" pageNumber="28" pagination="539 - 545" title="Species diversity patterns in insular land snail communities of Borneo." url="10.1144/jgs2012-014" volume="170" year="2013">Schilthuizen et al. 2013</bibRefCitation>
: online supplementary data.
</paragraph>
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
sp.
<normalizedToken originalValue="“BO-01”" pageId="27" pageNumber="28">&quot;BO-01&quot;</normalizedToken>
,
<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of Molluscan Studies" pageId="27" pageNumber="28" pagination="255 - 258" title="Microsnails at microscales in Borneo: distributions of Prosobranchia versus Pulmonata." url="10.1093/mollus/68.3.255" volume="68" year="2002">Schilthuizen et al. 2002</bibRefCitation>
a: 258;
<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Bulletin of the Raffles Museum" pageId="27" pageNumber="28" pagination="35 - 42" title="Abundance and diversity of land-snails (Mollusca: Gastropoda) on limestone hills in Borneo." volume="51" year="2002 b">Schilthuizen et al. 2002b</bibRefCitation>
: 41-42;
<bibRefCitation author="Schilthuizen, M" editor="Mohamed, M" journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="27" pageNumber="28" pagination="93 - 98" title="The land snails of the Tabin limestone hill." volumeTitle="Tabin Limestone Scientific Expedition 2000" year="2003">Schilthuizen and Vermeulen 2003</bibRefCitation>
: 96;
<bibRefCitation author="Schilthuizen, M" editor="Mohamed, M" journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="27" pageNumber="28" title="A checklist of the land snails of the Crocker Range." volumeTitle="Crocker Range Scientific Expedition 2002" year="2004">Schilthuizen 2004</bibRefCitation>
: 94;
<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of Tropical Biology and Conservation" pageId="27" pageNumber="28" pagination="1 - 7" title="The land and mangrove snail fauna of the islands of Banggi and Balambangan (Mollusca: Gastropoda)." volume="8" year="2011">Schilthuizen et al. 2011</bibRefCitation>
: 5.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="holotype">
<paragraph pageId="27" pageNumber="28">
<materialsCitation collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectionCode="RMNH" country="Malaysia" httpUri="http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003933" location="Interior Province" municipality="Sabah" specimenCode="RMNH.5003933" specimenCount="1" typeStatus="holotype">
<typeStatus>Holotype</typeStatus>
.
<collectingCountry>Malaysia</collectingCountry>
,
<collectingMunicipality>Sabah</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:929540D335CBCB6C3D4B6FF259639821:F43062E16D2321BE455E1E086FF332D9" country="Malaysia" municipality="Sabah" name="Interior Province">Interior Province</location>
,
<location LSID="urn:lsid:plazi:treatment:929540D335CBCB6C3D4B6FF259639821:552897FC8D39A52E6C42CCC1830F3795" country="Malaysia" municipality="Sabah" name="Gua Pungiton">Gua Pungiton</location>
(
<specimenCode>RMNH.5003933</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28" type="materials_examined">
<paragraph pageId="27" pageNumber="28">Examined material from Sabah.</paragraph>
<paragraph lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
Interior Province. Upper Padas valley, Matang River South of Long Pasia (leg. J.J. Vermeulen, V 9815); Matang River South of Long Pasia, ridge on West bank (leg. J.J. Vermeulen, V 9824); Long Pasia (leg. T.S. Liew &amp; Meckson, BOR/MOL 4355). Tenom (leg. M. Schilthuizen, BOR/MOL 3348). Crocker Range N.P., Gua Laing c. 12 km North of Keningau (leg. J.J. Vermeulen, V 1109, V 1114); West of the km 10 marker on the road Tambunan-Ranau, Mahua Waterfall (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9749; leg. M. Schilthuizen, BOR/MOL 3137; leg. J. Schilthuizen, BOR/MOL 1081). Gunung Trusmadi slopes: Forestry chalet (leg. J.J. Vermeulen, V 13211); Gua Loloposon (leg. J.J. Vermeulen, V 13244). Pinangah valley, Batu Urun (= Bukit Sinobang) (leg. J.J. Vermeulen, V 7997, V 1143, V 1168, BOR/MOL 1067; leg. R. Haegens, V 5633; leg. M. Schilthuizen, BOR/MOL 1085). Pun Batu c. 30 km West of Sepulut (leg. J.J. Vermeulen, V 1284. Sepulut valley, Batu Punggul (leg. J.J. Vermeulen, V 1971; leg. M. Schilthuizen, BOR/MOL 1099); Batu Temurung (leg. J.J. Vermeulen, V 8041, BOR/MOL 1071; leg. M. Schilthuizen, BOR/MOL 1091); Bukit Tinagas, East end of Batu Punggul limestone (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 7629; leg. M. Schilthuizen, BOR/MOL 1096); Gua Pungiton (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 7548, BOR/MOL 1069); Gua Sanaron (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 7678; leg. M. Schilthuizen, BOR/MOL 1082, BOR/MOL 1098). Kudat Province. Balambangan Island, Kok Simpul (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9530); South end, Batu Sireh (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9544; leg. T.H. Liew, Sazilin &amp; Ramlan, BOR/MOL 3689). Banggi Island, South end (leg. J.J. Vermeulen, V 1429; leg. M.A. Rahman, BOR/MOL 1094); Karakit Hill (leg. J.J. Vermeulen, V 1466; leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9470, V 9498); Pitas, Gua Mundau (leg. T.S. Liew &amp; M. Schilthuizen, BOR/MOL 4370); Kg. Magnin (leg. T.S. Liew &amp; M. Schilthuizen, BOR/MOL 4365). Sandakan Province. Meliau Range between 355-470 m (leg. T.S. Liew, M. Schilthuizen &amp; A. van Til, BOR/MOL 3196, BOR/MOL 3197, BOR/MOL 3221). Kinabatangan valley, Batu Batangan (leg. M. Schilthuizen, BOR/MOL 1328); Batu Keruak 2 near Sukau (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9790; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 1858, BOR/MOL 1892; leg. M. Sal
<pageBreakToken pageId="28" pageNumber="29" start="start">verda</pageBreakToken>
&amp; H. van Oosten, BOR/MOL 1322, BOR/MOL 1319); Hill Resang River (leg. M. Schilthuizen, BOR/MOL 1324); Batu Mawas (leg. T.S. Liew &amp; M. Schilthuizen, BOR/MOL 1969, BOR/MOL 2004; leg. M. Schilthuizen, BOR/MOL 1325); Batu
<pageBreakToken pageId="29" pageNumber="30" start="start">Materis</pageBreakToken>
(; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 2094, BOR/MOL 2126); Batu Pangi (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9641; leg. J.P. King, BOR/MOL 1326); Batu Tai (not Bod Tai) near Gomantong (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9582; leg. M. Schilthuizen, BOR/MOL 1321; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 1926); Batu Tomanggong Kecil (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9713); Batu Tulug (Batu Putih) along road Lahad Datu-Sandakan, North of bridge over Kinabatangan River (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1500); Gomantong Hill 30 km South of Sandakan (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1631; leg. M. Schilthuizen, BOR/MOL 1080; leg. A. van Til, BOR/MOL 3122, BOR/MOL 3276, BOR/MOL 3292, BOR/MOL 3302, BOR/MOL 3309, BOR/MOL 3321, BOR/MOL 3344; leg. T.S. Liew &amp; J.P. King, BOR/MOL 3649); Tandu Batu (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9628); Batu Tomanggong Besar (leg. M. Schilthuizen, BOR/MOL 1074, BOR/MOL 1317; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 2263, BOR/MOL 2294); Batu Tomanggong Kecil (leg. M. Salverda &amp; H. van Oosten, BOR/MOL 1327; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 2036, BOR/MOL 2064); Unnamed hill near Sukau Police Station (leg. M. Schilthuizen, BOR/MOL 1320; leg. T.S. Liew, BOR/MOL 2166; leg. T.S. Liew &amp; B. Elahan, BOR/MOL 2197). Segama valley, North end of limestone ridge on East bank Tabin River (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 7780, BOR/MOL 1087); Tabin Wildlife Reserve (leg. T. Kimsin, BOR/MOL 735; leg. M. Schilthuizen, BOR/MOL 1086). Tawau Province. Batu Baturong c. 50 km W.S.W. Lahad Datu (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1853); North slope (leg. J.J. Vermeulen, V 7588). Danum Valley (leg. M. Schilthuizen, BOR/MOL 1068, BOR/MOL 1073, BOR/MOL 1077; leg. UMS students, BOR/MOL 1075; leg. H.A. Rutjes, BOR/MOL 1079, BOR/MOL 1083;; leg. T. Kimsin &amp; H.N. Chai, BOR/MOL 1084). Gua Madai c. 40 km S.S.W. of Lahad Datu (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1724); N.E. End (leg. J.J. Vermeulen, V 7700, BOR/MOL 1093). Segama valley, hill N.W. of crossing road Sandakan-Lahad Datu with the Segama River (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1679);
<normalizedToken originalValue="Kirks" pageId="29" pageNumber="30">'Kirk's</normalizedToken>
<normalizedToken originalValue="Cave" pageId="29" pageNumber="30">Cave'</normalizedToken>
8 km North of Lahad Datu (leg. J.J. Vermeulen, V 1214); limestone hill on North bank Segama River, near bridge of road Sandakan to Lahad Datu (leg. J.J. Vermeulen, V 7501); Sabahmas Cave (leg. J.J. Vermeulen, V 7458, BOR/MOL 1089). Tawau Hills Park (leg. J.P. King, BOR/MOL 1316). Semporna area, Segarong Hills, Bukit Pababola, 25 km E.S.E. of Kunak (leg. J.J. Vermeulen &amp; H. Duistermaat, V 1767); Bod Gaya Island (leg. T.S. Liew, Abdul &amp; Ladja, BOR/MOL 4761, BOR/MOL 4775, BOR/MOL 4789, BOR/MOL 4897, BOR/MOL 4908). Tawau Hills N.P., path up to Bukit Bombalai (leg. J.J. Vermeulen, V 13156). West Coast Province. Crocker Range N.P., km 54 marker on the road Kota Kinabalu-Tambunan, Gunung Mas (leg. J.J. Vermeulen &amp; M. Schilthuizen, V 9766; leg. M. Schilthuizen, BOR/MOL 1090). Kinabalu N.P., Headquarters area near entrance (leg. J.J. Vermeulen, V 9775; leg. M. Schilthuizen, BOR/MOL 1092); Sayap (leg. M. Schilthuizen, J. Schilthuizen &amp; F. Schilthuizen, BOR/MOL 1323); Summit trail between 1616-3089 m (leg. T.S. Liew &amp; J. Lapidin, BOR/MOL 4040; leg. T.S. Liew, BOR/MOL 4063, BOR/MOL 4031, BOR/MOL 4035, BOR/MOL 4066, BOR/MOL
<pageBreakToken pageId="30" pageNumber="31" start="start">4051</pageBreakToken>
, BOR/MOL 4055); Kiau-Spurs route between 3024 and 3088 m (leg. T.S. Liew, J. Lapidin &amp; Safrie, BOR/MOL 4034, BOR/MOL 4069, BOR/MOL 4029);
<normalizedToken originalValue="Kotals" pageId="30" pageNumber="31">Kotal's</normalizedToken>
route between 1952 and 3416 m (leg. T.S. Liew &amp; Jasilin, BOR/MOL 4032, BOR/MOL 4033, BOR/MOL 4059, BOR/MOL 4062, BOR/MOL 4038, BOR/MOL 4039, BOR/MOL 4043); Mesilau trail between 1948 and 2356 m (leg. T.S. Liew &amp; J. Lapidin, BOR/MOL 4054, BOR/MOL 4050, BOR/MOL 4056, BOR/MOL 4058, BOR/MOL 4067; leg. T.S. Liew, BOR/MOL 4028, BOR/MOL 4048, BOR/MOL 4053); Sayap - Nunuhon trail between 960 and 2944 m (leg. T.S. Liew, Dominik, J. Lapidin &amp; Jasilin, BOR/MOL 4037, BOR/MOL 4041, BOR/MOL 4042, BOR/MOL 4046, BOR/MOL 4064); Serinsim (leg. M. Schilthuizen, BOR/MOL 3070); Serinsim-Numbuyukon trail between 700 and 1680 m (leg. T.S. Liew, BOR/MOL 4044, BOR/MOL 4049, BOR/MOL 4057, BOR/MOL 4065, BOR/MOL 4068); Monggis-Tambuyukon trail between 1016 and 2360 m (leg. T.S. Liew, BOR/MOL 4030, BOR/MOL 4045), BOR/MOL 4052). Kota Kinabalu, Kiansom (leg. UMS students, BOR/MOL 1095). Mantanani Group, Pulau Mantanani Besar (leg. T.H. Liew, BOR/MOL 3714); Pulau Mantanani Kecil (leg. T.H. Liew, BOR/MOL 3733).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="31" type="description">
<paragraph pageId="30" pageNumber="31">Description.</paragraph>
<paragraph pageId="30" pageNumber="31">Shell very small, thin, translucent, white, lenticular; spire almost flat or slightly elevated. Surface glossy. Whorls slightly convex. Protoconch: smooth, sometimes with a few inconspicuous, scattered radial riblets only. Teleoconch: Spiral sculpture absent. Radial sculpture teleoconch: growth lines inconspicuous, next to these widely but irregularly spaced, shallow grooves, often alternating with areas of a much finer, more dense but irregularly spaced striation; sometimes the latter striation is predominant. Umbilicus partly or entirely covered by a minute, triangular spur protruding from the columellar corner of the peristome; this spur lacking or inconspicuous in juveniles; umbilical region moderately concave. Dimensions: Height up to 1.7 mm; width up to 3 mm; diameters of the first three whorls 0.5-0.8 mm, 0.9-1.5 mm, 1.5-2.5 mm respectively; number of whorls up to 4 1/8; height aperture up to 1.4 mm; width aperture up to 1.7 mm.</paragraph>
<caption pageId="30" pageNumber="31">
<paragraph pageId="30" pageNumber="31">
Figure 24-26. 24
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina microrhynchus" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="microrhynchus">Microcystina microrhynchus</taxonomicName>
sp. n. 24A Frontal view, shell 1.6 mm high 24B Apical view 24C Umbilical view (Malaysia, Sabah, Sepulut valley, Gua Pungiton; 24A RMNH.5003933, Holotype; 24B, 24CV 7548) 25
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina callifera" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="callifera">Microcystina callifera</taxonomicName>
sp. n. 25A Frontal view, shell 1.4 mm high 25B Apical view 25C Umbilical view (Malaysia, Sabah, Mantanani Group, Pulau Lungisan, RMNH.5003934, holotype) 26
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina striatula" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="striatula">Microcystina striatula</taxonomicName>
sp. n. 26A Frontal view, shell 0.8 mm high 26B Apical view 26C Umbilical view (Malaysia, Sabah, Segama valley, North end of limestone ridge on East bank Tabin River, RMNH.5003935, holotype).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="30" pageNumber="31" type="habitat in sabah and distribution">
<paragraph pageId="30" pageNumber="31">Habitat in Sabah and distribution.</paragraph>
<paragraph pageId="30" pageNumber="31">Rainforest, periodically dry, shrubby woodland, coastal woodland, secondary forest and other degraded vegetation; on limestone, sandstone and granitic soil, up to 3500 m alt. Sabah: common and widespread. Also in Brunei; Sarawak; Kalimantan. Endemic to Borneo.</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="31" type="cross diagnosis">
<paragraph pageId="30" pageNumber="31">Cross diagnosis.</paragraph>
<paragraph pageId="30" pageNumber="31">
Characterized among the Borneo
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
species with white shells by a complete lack of spiral striation. Usually it is also larger and has more rapidly expanding whorls.
</paragraph>
<paragraph pageId="30" pageNumber="31">
Elsewhere,
<taxonomicName genus="Lamprocystus" lsidName="Lamprocystus vitreiformis" pageId="30" pageNumber="31" rank="species" species="vitreiformis">Lamprocystus vitreiformis</taxonomicName>
Von Moellendorff, 1897, from Sumatra, Java, and Bali (see
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Backhuys Publishers &amp; Fauna Malesiana Foundation, Leiden" pageId="30" pageNumber="31" title="Fauna Malesiana Guide to the land snails of Bali" year="1998">Vermeulen and Whitten 1998</bibRefCitation>
: 118) is most similar;
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina microrhynchus" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="microrhynchus">Microcystina microrhynchus</taxonomicName>
is characterized by the minute spur protruding from the columellar corner of the peristome and covering the umbilicus.
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina chionodiscus" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="chionodiscus">Microcystina chionodiscus</taxonomicName>
Vermeulen, 1996, from Bali (see
<bibRefCitation author="Vermeulen, JJ" journalOrPublisher="Backhuys Publishers &amp; Fauna Malesiana Foundation, Leiden" pageId="30" pageNumber="31" title="Fauna Malesiana Guide to the land snails of Bali" year="1998">Vermeulen and Whitten 1998</bibRefCitation>
: 115), has the first whorl consistently smaller (diameter 0.4-0.5 mm) than
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina microrhynchus" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="microrhynchus">Microcystina microrhynchus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="31" type="etymology">
<paragraph pageId="30" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="30" pageNumber="31">The name refers to the spur covering the umbilicus [mikros (Gr.) = small; rhynchos (Gr.) = beak].</paragraph>
</subSubSection>
</treatment>
</document>