treatments-xml/data/D3/6A/82/D36A8250FFD6FF88FF71FBD8FCDA190F.xml

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<mods:title>Two new species of Aleocharinae (Coleoptera, Staphylinidae) found in fungus gardens of Odontotermes termites (Isoptera, Termitidae, Macrotermitinae) in Khao Yai National Park, Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kanao, Taisuke</mods:namePart>
<mods:affiliation>Kyushu University ,, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maruyama, Munetoshi</mods:namePart>
<mods:affiliation>The Kyushu University Museum, Fukuoka, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sakchoowong, Watana</mods:namePart>
<mods:affiliation>Forest Entomology and Microbiology Group, National Park, Wildlife and Plant Conservation Department, Bangkok ,, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-06-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>49</mods:number>
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<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>86</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.49.468</mods:identifier>
<mods:identifier type="GBIF-Dataset">aabca6d4-7333-4bb0-861c-cb6e63ee3518</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576677</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788133" ID-GBIF-Taxon="163747296" ID-Zenodo-Dep="3788133" LSID="urn:lsid:plazi:treatment:D36A8250FFD6FF88FF71FBD8FCDA190F" httpUri="http://treatment.plazi.org/id/D36A8250FFD6FF88FF71FBD8FCDA190F" lastPageId="8" lastPageNumber="85" pageId="4" pageNumber="81">
<subSubSection pageId="4" pageNumber="81" type="nomenclature">
<paragraph blockId="4.[140,476,1046,1073]" box="[140,476,1046,1073]" pageId="4" pageNumber="81">
<heading box="[140,476,1046,1073]" fontSize="11" level="2" pageId="4" pageNumber="81" reason="3">
<emphasis bold="true" box="[140,476,1046,1073]" pageId="4" pageNumber="81">
Genus 
<taxonomicName authority="Kistner" authorityName="Kistner" authorityYear="1958" box="[223,476,1046,1073]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[223,379,1046,1073]" italics="true" pageId="4" pageNumber="81">Odontoxenus</emphasis>
Kistner
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[140,1108,1117,1179]" pageId="4" pageNumber="81">
<bibRefCitation author="Kistner DH" box="[140,349,1117,1144]" pageId="4" pageNumber="81" pagination="1 - 198" refId="ref3315" refString="Kistner DH (1958) The Evolution of the Pygostenini (Coleoptera Staphylinidae). Annalen Du Musee Royal du Congo Belge Tervuren (Belgique). 68: 1 - 198." type="journal article" year="1958">Kistner 1958: 104</bibRefCitation>
(original description); 
<bibRefCitation author="Jacobson HR &amp; Kistner DH" box="[613,981,1117,1144]" pageId="4" pageNumber="81" pagination="201 - 335" refId="ref3196" refString="Jacobson HR, Kistner DH (1975) A manual for the indication of the Pygostenini, the natural history of the myrmecophilous tribe Pygostenini. Sociobiology 1: 201 - 335." type="journal article" year="1975">Jacobson and Kistner 1975: 293</bibRefCitation>
(key, diagnosis).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="81" type="discussion">
<paragraph blockId="4.[140,1108,1222,1390]" pageId="4" pageNumber="81">
<emphasis bold="true" box="[140,279,1222,1248]" pageId="4" pageNumber="81">
<emphasis box="[140,274,1222,1248]" inLineHeading="true" pageId="4" pageNumber="81" reason="1">Comments</emphasis>
.
</emphasis>
This genus in general appearance and body size is similar to 
<taxonomicName authorityName="Wasmann" authorityYear="1898" box="[975,1108,1223,1249]" class="Insecta" family="Staphylinidae" genus="Doryloxenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis box="[975,1108,1223,1249]" italics="true" pageId="4" pageNumber="81">Doryloxenus</emphasis>
</taxonomicName>
among the Asian genera of 
<taxonomicName authorityName="Fauvel" authorityYear="1899" box="[461,596,1258,1284]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Pygostenini">Pygostenini</taxonomicName>
, but distinguished from it by the shape and position of eyes having part of their surface on the anterior margin of the head, the relatively long mesosternum, and the reduced 4-segmented tarsi (
<bibRefCitation author="Jacobson HR &amp; Kistner DH" pageId="4" pageNumber="81" pagination="201 - 335" refId="ref3196" refString="Jacobson HR, Kistner DH (1975) A manual for the indication of the Pygostenini, the natural history of the myrmecophilous tribe Pygostenini. Sociobiology 1: 201 - 335." type="journal article" year="1975">Jacobson and Kistner 1975</bibRefCitation>
).
</paragraph>
<paragraph blockId="4.[140,983,1468,1566]" pageId="4" pageNumber="81">
<emphasis bold="true" box="[140,785,1468,1495]" pageId="4" pageNumber="81">
<taxonomicName authority="Kanao &amp; Maruyama" authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[140,707,1468,1495]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="thailandicus" status="sp. n.">
<emphasis bold="true" box="[140,451,1468,1495]" italics="true" pageId="4" pageNumber="81">
<emphasis box="[140,296,1468,1495]" inLineHeading="true" pageId="4" pageNumber="81" reason="1">Odontoxenus</emphasis>
thailandicus
</emphasis>
Kanao &amp; Maruyama
</taxonomicName>
, 
<taxonomicNameLabel box="[720,785,1469,1495]" pageId="4" pageNumber="81" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<uri box="[140,983,1504,1531]" pageId="4" pageNumber="81">
urn:lsid:zoobank.org:act: 
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</uri>
</paragraph>
<paragraph blockId="4.[140,983,1468,1566]" box="[140,288,1539,1566]" pageId="4" pageNumber="81">
<figureCitation box="[140,208,1539,1566]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="4" pageNumber="81">Figs 3</figureCitation>
, 
<figureCitation box="[221,238,1540,1566]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="4" pageNumber="81">4</figureCitation>
, 
<figureCitation box="[251,288,1539,1566]" captionStart="Figure 5" captionStartId="7.[140,212,920,942]" captionTargetBox="[152,1096,176,887]" captionTargetId="figure@7.[140,1108,165,900]" captionTargetPageId="7" captionText="Figure 5. A Odontotermes proformosana, queen, workers and soldiers B the fungus garden of O. proformosana C Discoxenus katayamai, on fungus garden D Odontoxenus thailandicus, on fungus garden. Photos © Y. Katayama (5A–5B), T. Komatsu (5C–5D), 2007." figureDoi="http://doi.org/10.5281/zenodo.3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="4" pageNumber="81">5D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81" type="materials_examined">
<paragraph blockId="4.[140,1107,1609,1672]" lastBlockId="5.[140,1108,1328,1672]" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81">
<emphasis bold="true" box="[140,279,1609,1636]" pageId="4" pageNumber="81">Type series.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608811495" collectingDate="2007-09-26" collectingDateMax="2007-09-28" collectingDateMin="2007-09-26" collectorName="Katayama Y. &amp; Palatypes" county="Thai" elevation="700" lastPageId="5" lastPageNumber="82" location="Mo Sing To" municipality="Khao Yai National Park" pageId="4" pageNumber="81" specimenCount="4" specimenCount-female="4" stateProvince="Nakhon Nayok" typeStatus="holotype">
<typeStatus box="[288,399,1610,1636]" pageId="4" pageNumber="81">Holotype</typeStatus>
: ♁, “ 
<collectingCounty box="[460,516,1610,1636]" pageId="4" pageNumber="81">Thai</collectingCounty>
: 
<collectingRegion box="[528,708,1610,1636]" country="Thailand" name="Nakhon Nayok" pageId="4" pageNumber="81">Nakhon Nayok</collectingRegion>
, 
<collectingMunicipality box="[721,996,1610,1636]" pageId="4" pageNumber="81">Khao Yai National Park</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:D36A8250FFD6FF88FF71FBD8FCDA190F:5E1C659DFFD6FF84FC0DF984FF6B1CBC" county="Thai" municipality="Khao Yai National Park" name="Mo Sing To" pageId="4" pageNumber="81" stateProvince="Nakhon Nayok">Mo Sing To</location>
(
<quantity box="[186,262,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="81" unit="m" value="700.0">
<elevation box="[186,262,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="81" unit="m" value="700.0">700 m</elevation>
</quantity>
alt.), 
<date box="[329,465,1645,1672]" pageId="4" pageNumber="81" value="2007-09-28">
<collectingDate box="[329,465,1645,1672]" pageId="4" pageNumber="81" value="2007-09-28">28 IX 2007</collectingDate>
</date>
, Maruyama M. and 
<collectorName box="[699,841,1646,1672]" pageId="4" pageNumber="81">Katayama Y.</collectorName>
leg. // 
<typeStatus box="[921,1028,1645,1671]" pageId="4" pageNumber="81">Holotype</typeStatus>
<emphasis italics="true" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81">
Odon- toxenus 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[231,364,1328,1354]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="thailandicus">thailandicus</taxonomicName>
</emphasis>
des. Kanao &amp; Maruyama, 2010”. 
<collectorName box="[781,889,1328,1354]" pageId="5" pageNumber="82">Palatypes</collectorName>
: 
<specimenCount box="[904,933,1328,1355]" pageId="5" pageNumber="82" type="female">2♀</specimenCount>
, same data as holotype; 2♁, 
<specimenCount box="[305,335,1364,1390]" pageId="5" pageNumber="82" type="female">2♀</specimenCount>
, ditto, but 
<date box="[472,607,1363,1390]" pageId="5" pageNumber="82" value="2007-09-26">
<collectingDate box="[472,607,1363,1390]" pageId="5" pageNumber="82" value="2007-09-26">26 IX 2007</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767165" ID-Zenodo-Dep="3767165" box="[140,898,1227,1249]" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="5" pageNumber="82" startId="5.[140,212,1227,1249]" targetBox="[140,1108,165,1207]" targetPageId="5">
<paragraph blockId="5.[140,898,1227,1249]" box="[140,898,1227,1249]" pageId="5" pageNumber="82">
<emphasis bold="true" box="[140,238,1227,1249]" pageId="5" pageNumber="82">Figure 3.</emphasis>
Habitus of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[349,578,1227,1248]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis box="[349,578,1227,1248]" italics="true" pageId="5" pageNumber="82">Odontoxenus thailandicus</emphasis>
</taxonomicName>
. 
<emphasis bold="true" box="[591,610,1227,1249]" pageId="5" pageNumber="82">A</emphasis>
dorsal view and 
<emphasis bold="true" box="[765,782,1227,1249]" pageId="5" pageNumber="82">B</emphasis>
lateral view.
</paragraph>
</caption>
<paragraph blockId="5.[140,1108,1328,1672]" box="[185,856,1398,1425]" pageId="5" pageNumber="82">
<emphasis bold="true" box="[185,372,1398,1425]" pageId="5" pageNumber="82">
<emphasis box="[185,306,1398,1424]" inLineHeading="true" pageId="5" pageNumber="82" reason="1">Symbiotic</emphasis>
host:
</emphasis>
<taxonomicName authority="Ahmad, 1965" authorityName="Ahmad" authorityYear="1965" box="[379,856,1398,1425]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="proformosanus">
<emphasis box="[379,693,1399,1425]" italics="true" pageId="5" pageNumber="82">Odontotermes proformosanus</emphasis>
<bibRefCitation author="Ahmad M" box="[700,856,1398,1425]" pageId="5" pageNumber="82" pagination="1 - 111" refId="ref3009" refString="Ahmad M (1965) Termites of Thailand. Bulletin of the American Museum of Natural History 131: 1 - 111." type="journal article" year="1965">Ahmad, 1965</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[185,663,1434,1460]" pageId="5" pageNumber="82" type="etymology">
<paragraph blockId="5.[140,1108,1328,1672]" box="[185,663,1434,1460]" pageId="5" pageNumber="82">
<emphasis bold="true" box="[185,320,1434,1460]" pageId="5" pageNumber="82">
<emphasis box="[185,315,1434,1460]" inLineHeading="true" pageId="5" pageNumber="82" reason="1">Etymology</emphasis>
.
</emphasis>
Named after the 
<typeStatus box="[520,568,1434,1460]" pageId="5" pageNumber="82">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="82" type="diagnosis">
<paragraph blockId="5.[140,1108,1328,1672]" pageId="5" pageNumber="82">
<emphasis bold="true" box="[185,311,1469,1495]" pageId="5" pageNumber="82">
<emphasis box="[185,306,1469,1495]" inLineHeading="true" pageId="5" pageNumber="82" reason="1">Diagnosis</emphasis>
.
</emphasis>
Most similar to 
<taxonomicName authority="(Wasmann, 1916)" baseAuthorityName="Wasmann" baseAuthorityYear="1916" box="[500,820,1469,1496]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="butteri">
<emphasis box="[500,608,1469,1496]" italics="true" pageId="5" pageNumber="82">O. butteri</emphasis>
(
<bibRefCitation author="Wasmann E" box="[624,812,1469,1496]" pageId="5" pageNumber="82" pagination="169 - 210" refId="ref3630" refString="Wasmann E (1916) Wissenshaftliche Ergebnisse einer Forschungsreise nach Ostindien, V, Termitophile und Myrmekophile Coleopteren, gesammelt von Herrn Prof. Dr. v. Buttel- Reepen, 19911 - 1912. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 39: 169 - 210." type="journal article" year="1916">Wasmann, 1916</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName authority="Kistner, 2005" authorityName="Kistner" authorityYear="2005" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis box="[878,1045,1469,1496]" italics="true" pageId="5" pageNumber="82">O. malaysianus</emphasis>
<bibRefCitation author="Kistner DH" pageId="5" pageNumber="82" pagination="209 - 213" refId="ref3467" refString="Kistner DH (2005) A new species of Odontoxenus from Malaysia (Coleoptera: Staphylinidae: Aleocharinae: Pygostenini). Sociobiology 45: 209 - 213." type="journal article" year="2005">Kistner, 2005</bibRefCitation>
</taxonomicName>
through the pronotum with a row of macrosetae at the lateral margin, but distinguished from them by the macrochaetotaxy of the tergites II–VIII: 0, 2, 2, 2, 4, 6, 8.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="85" pageId="5" pageNumber="82" type="description">
<paragraph blockId="5.[140,1108,1328,1672]" lastBlockId="6.[140,1108,1152,1672]" lastPageId="6" lastPageNumber="83" pageId="5" pageNumber="82">
<emphasis bold="true" box="[185,335,1574,1600]" pageId="5" pageNumber="82">
<emphasis box="[185,330,1574,1600]" inLineHeading="true" pageId="5" pageNumber="82" reason="1">Description</emphasis>
.
</emphasis>
Body (
<figureCitation box="[414,479,1575,1601]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="5" pageNumber="82">Figs 3</figureCitation>
A–3B) almost uniformly reddish brown; head and elytra slightly darker. Dorsal surfaces of head, pronotum and elytra (
<figureCitation box="[840,926,1610,1636]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="5" pageNumber="82">Fig. 3A</figureCitation>
) smooth, glossy and glabrous. Head to elytra (
<figureCitation box="[472,556,1645,1671]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="5" pageNumber="82">Fig. 3B</figureCitation>
) well convex, laterally curved in shape of half cir- cle. Pronotum (
<figureCitation box="[315,382,1152,1178]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="6" pageNumber="83">Figs 3</figureCitation>
A–3B, 4B) much narrower than elytra, with a row of 5 macrosetae on lateral margin. Elytra (
<figureCitation box="[466,532,1187,1214]" captionStart="Figure 3" captionStartId="5.[140,212,1227,1249]" captionTargetBox="[140,1108,165,1207]" captionTargetId="figure@5.[140,1108,165,1207]" captionTargetPageId="5" captionText="Figure 3. Habitus of Odontoxenus thailandicus. A dorsal view and B lateral view." figureDoi="http://doi.org/10.5281/zenodo.3767165" httpUri="https://zenodo.org/record/3767165/files/figure.png" pageId="6" pageNumber="83">Figs 3</figureCitation>
A–3B, 4A) with inflexed lateral margins, and about 10 macrosetae on lateral margins. Macrochaetotaxy of abdominal tergites II–VIII: 0, 2, 2, 2, 4, 6, 8; male abdominal tergite VIII (
<figureCitation box="[672,761,1258,1284]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4C</figureCitation>
) slightly truncate at apex and slightly wrinkled on dorsal surface, with 3 pairs of macrosetae near apex (1 laterally, 2 dorsally), and with 1 pair of flattened setae at apex; sternite VIII (
<figureCitation box="[924,1015,1328,1354]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4D</figureCitation>
) with 2 pairs of macrosetae on dorsal surface and 3 pairs at apex; female tergite VIII (
<figureCitation box="[1016,1099,1363,1390]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4F</figureCitation>
) with basal projections and macrosetae shorter than in male. Tergite IX (
<figureCitation box="[955,1040,1399,1425]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4E</figureCitation>
) with 7 pairs of macrosetae laterally; tergite X (
<figureCitation box="[615,702,1434,1460]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4E</figureCitation>
) with 3 macrosetae around middle and 2 pairs of macrosetae at apex. Median lobe of aedeagus (
<figureCitation box="[824,912,1469,1495]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4G</figureCitation>
) with apical lobe almost straight, much narrower than basal capsule; copulatory piece membranous; basal capsule swollen, half as long as apical lobe. Spermatheca (
<figureCitation box="[851,941,1539,1566]" captionStart="Figure 4" captionStartId="6.[140,212,1014,1036]" captionTargetBox="[316,936,177,987]" captionTargetId="figure@6.[302,945,165,994]" captionTargetPageId="6" captionText="Figure 4. Odontoxenus thailandicus. A elytron B pronotum C male abdominal tergite VIII D male abdominal sternite VIII E male abdominal tergite IV &amp; V F female abdominal tergite VIII G median lobe of aedeagus, and H spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83">Fig. 4H</figureCitation>
) curved twice, S-shaped; apical part swollen, its inner wall hollowed at apex, densely wrinkled from apex to around apical 1/3; basal part 2.5 times as long as apical part, roundly curved around basal 1/3 and gently curved around apex.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767167" ID-Zenodo-Dep="3767167" httpUri="https://zenodo.org/record/3767167/files/figure.png" pageId="6" pageNumber="83" startId="6.[140,212,1014,1036]" targetBox="[316,936,177,987]" targetPageId="6">
<paragraph blockId="6.[140,1107,1014,1100]" pageId="6" pageNumber="83">
<emphasis bold="true" box="[140,239,1014,1036]" pageId="6" pageNumber="83">Figure 4.</emphasis>
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[245,475,1014,1035]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis box="[245,475,1014,1035]" italics="true" pageId="6" pageNumber="83">Odontoxenus thailandicus</emphasis>
</taxonomicName>
. 
<emphasis bold="true" box="[488,507,1014,1036]" pageId="6" pageNumber="83">A</emphasis>
elytron 
<emphasis bold="true" box="[585,602,1014,1036]" pageId="6" pageNumber="83">B</emphasis>
pronotum 
<emphasis bold="true" box="[709,728,1014,1036]" pageId="6" pageNumber="83">C</emphasis>
male abdominal tergite VIII 
<emphasis bold="true" box="[1002,1021,1014,1036]" pageId="6" pageNumber="83">D</emphasis>
male abdominal sternite VIII 
<emphasis bold="true" box="[344,359,1046,1068]" pageId="6" pageNumber="83">E</emphasis>
male abdominal tergite IV &amp; V 
<emphasis bold="true" box="[664,678,1046,1068]" pageId="6" pageNumber="83">F</emphasis>
female abdominal tergite VIII 
<emphasis bold="true" box="[967,987,1046,1068]" pageId="6" pageNumber="83">G</emphasis>
median lobe of aedeagus, and 
<emphasis bold="true" box="[298,318,1078,1100]" pageId="6" pageNumber="83">H</emphasis>
spermatheca.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767169" ID-Zenodo-Dep="3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="7" pageNumber="84" startId="7.[140,212,920,942]" targetBox="[152,1096,176,887]" targetPageId="7">
<paragraph blockId="7.[140,1108,920,1006]" pageId="7" pageNumber="84">
<emphasis bold="true" box="[140,264,920,942]" pageId="7" pageNumber="84">Figure 5. A</emphasis>
<taxonomicName box="[270,519,920,941]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="proformosana">
<emphasis box="[270,519,920,941]" italics="true" pageId="7" pageNumber="84">Odontotermes proformosana</emphasis>
</taxonomicName>
, queen, workers and soldiers 
<emphasis bold="true" box="[795,812,920,942]" pageId="7" pageNumber="84">B</emphasis>
the fungus garden of 
<taxonomicName class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="proformosana">
<emphasis italics="true" pageId="7" pageNumber="84">O. proformosana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[212,231,952,974]" pageId="7" pageNumber="84">C</emphasis>
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[236,431,952,974]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis box="[236,431,952,974]" italics="true" pageId="7" pageNumber="84">Discoxenus katayamai</emphasis>
</taxonomicName>
, on fungus garden 
<emphasis bold="true" box="[607,626,952,974]" pageId="7" pageNumber="84">D</emphasis>
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[632,861,952,973]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis box="[632,861,952,973]" italics="true" pageId="7" pageNumber="84">
<taxonomicName box="[632,747,952,973]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="genus">Odontoxenus</taxonomicName>
thailandicus
</emphasis>
</taxonomicName>
, on fungus garden. Photos © Y. Katayama (
<emphasis bold="true" box="[294,367,984,1006]" pageId="7" pageNumber="84">5A–5B</emphasis>
), T. Komatsu (
<emphasis bold="true" box="[508,584,984,1006]" pageId="7" pageNumber="84">5C–5D</emphasis>
), 2007.
</paragraph>
</caption>
<paragraph blockId="7.[140,1108,1046,1144]" pageId="7" pageNumber="84">
<emphasis bold="true" box="[185,365,1047,1073]" pageId="7" pageNumber="84">
<emphasis box="[185,360,1047,1073]" inLineHeading="true" pageId="7" pageNumber="84" reason="1">Measurements</emphasis>
.
</emphasis>
Body length:ca.1.7–2.0;pronotal length:0.44–0.51 (0.47±0.031); pronotal width: 0.64–0.73 (0.68±0.037); antennal length: 0.29–0.30 (0.30±0.012). N=5.
</paragraph>
<paragraph blockId="7.[140,356,1237,1264]" box="[140,356,1237,1264]" pageId="7" pageNumber="84">
<emphasis bold="true" box="[140,356,1237,1264]" pageId="7" pageNumber="84">Biological notes</emphasis>
</paragraph>
<paragraph blockId="7.[140,1108,1293,1672]" lastBlockId="8.[140,1107,166,228]" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
These new species were found in the fungus gardens of 
<taxonomicName authorityName="Ahmad" authorityYear="1965" box="[786,1107,1293,1319]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="proformosanus">
<emphasis box="[786,1107,1293,1319]" italics="true" pageId="7" pageNumber="84">Odontotermes proformosanus</emphasis>
</taxonomicName>
(
<figureCitation box="[147,219,1328,1355]" captionStart="Figure 5" captionStartId="7.[140,212,920,942]" captionTargetBox="[152,1096,176,887]" captionTargetId="figure@7.[140,1108,165,900]" captionTargetPageId="7" captionText="Figure 5. A Odontotermes proformosana, queen, workers and soldiers B the fungus garden of O. proformosana C Discoxenus katayamai, on fungus garden D Odontoxenus thailandicus, on fungus garden. Photos © Y. Katayama (5A–5B), T. Komatsu (5C–5D), 2007." figureDoi="http://doi.org/10.5281/zenodo.3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="7" pageNumber="84">Figs 5</figureCitation>
A–5B). The scuttle flies 
<emphasis box="[507,770,1328,1354]" italics="true" pageId="7" pageNumber="84">Clitelloxenus perdosetae</emphasis>
<bibRefCitation author="Disney RHL &amp; Kistner DH" box="[779,938,1328,1354]" pageId="7" pageNumber="84" pagination="3 - 118" refId="ref3073" refString="Disney RHL, Kistner DH (1997) Revision of the oriental Termitoxeniinae (Diptera: Phoridae). Sociobiology 29: 3 - 118." type="journal article" year="1997">Disney, 1997</bibRefCitation>
, 
<taxonomicName authority="Disney, 1997" authorityName="Disney" authorityYear="1997" class="Insecta" family="Staphylinidae" genus="Compactopedia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandae">
<emphasis box="[953,1107,1328,1355]" italics="true" pageId="7" pageNumber="84">C. thailandae</emphasis>
<bibRefCitation author="Disney RHL &amp; Kistner DH" box="[140,301,1363,1390]" pageId="7" pageNumber="84" pagination="3 - 118" refId="ref3073" refString="Disney RHL, Kistner DH (1997) Revision of the oriental Termitoxeniinae (Diptera: Phoridae). Sociobiology 29: 3 - 118." type="journal article" year="1997">Disney, 1997</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authorityName="Schmitz" authorityYear="1931" box="[317,433,1364,1389]" class="Insecta" family="Phoridae" genus="Franssenia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis box="[317,433,1364,1389]" italics="true" pageId="7" pageNumber="84">Franssenia</emphasis>
</taxonomicName>
sp., 
<taxonomicName box="[497,583,1363,1389]" class="Insecta" family="Phoridae" genus="Crasilla" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis box="[497,583,1363,1389]" italics="true" pageId="7" pageNumber="84">Crasilla</emphasis>
</taxonomicName>
sp. and 
<taxonomicName authorityName="Schmitz" authorityYear="1931" box="[693,853,1364,1389]" class="Insecta" family="Phoridae" genus="Dicranopteron" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis box="[693,853,1364,1389]" italics="true" pageId="7" pageNumber="84">Dicranopteron</emphasis>
</taxonomicName>
sp. were also caught at the same time. The individual number of each species was very low, compared to 
<taxonomicName authorityName="Kistner" authorityYear="2005" box="[176,352,1434,1460]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis box="[176,352,1434,1460]" italics="true" pageId="7" pageNumber="84">D. malaysianus</emphasis>
</taxonomicName>
collected in a high number from a few 
<emphasis box="[851,1017,1434,1460]" italics="true" pageId="7" pageNumber="84">Odonototermes</emphasis>
<bibRefCitation author="Holmgren N" pageId="7" pageNumber="84" pagination="1 - 166" refId="ref3167" refString="Holmgren N (1912) Termiten Studien. der Termiten. Die Familie Metatermitidae. Svenska Vetensk. Akad. Handl (4): 1 - 166." type="journal article" year="1912">Holmgren, 1912</bibRefCitation>
nests in 
<collectingCountry box="[368,469,1469,1495]" name="Malaysia" pageId="7" pageNumber="84">Malaysia</collectingCountry>
(
<bibRefCitation author="Kistner DH" box="[485,643,1469,1495]" pageId="7" pageNumber="84" pagination="165 - 186" refId="ref3428" refString="Kistner DH (1982) A revision of the termitophilous genus Discoxenus with a study of the relationships of the genus and notes on its behavior (Coleoptera, Staphylinidae). Sociobiology 7: 165 - 186." type="journal article" year="1982">Kistner 1982</bibRefCitation>
). Only a few specimens of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[954,1107,1469,1495]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis box="[954,1107,1469,1495]" italics="true" pageId="7" pageNumber="84">D. katayamai</emphasis>
</taxonomicName>
(
<figureCitation box="[148,240,1504,1531]" captionStart="Figure 5" captionStartId="7.[140,212,920,942]" captionTargetBox="[152,1096,176,887]" captionTargetId="figure@7.[140,1108,165,900]" captionTargetPageId="7" captionText="Figure 5. A Odontotermes proformosana, queen, workers and soldiers B the fungus garden of O. proformosana C Discoxenus katayamai, on fungus garden D Odontoxenus thailandicus, on fungus garden. Photos © Y. Katayama (5A–5B), T. Komatsu (5C–5D), 2007." figureDoi="http://doi.org/10.5281/zenodo.3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="7" pageNumber="84">Fig. 5C</figureCitation>
) were found in one nest, which generally contained one to ten, fist-sized fungus gardens, and none or one specimen of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[669,841,1539,1566]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis box="[669,841,1539,1566]" italics="true" pageId="7" pageNumber="84">O. thailandicus</emphasis>
</taxonomicName>
was found in one nest, though density of the host termites was very high at Mo Sing To, the 
<typeStatus box="[965,1015,1575,1601]" pageId="7" pageNumber="84">type</typeStatus>
locality of the present new species. The behavior of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[651,805,1610,1636]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis box="[651,805,1610,1636]" italics="true" pageId="7" pageNumber="84">D. katayamai</emphasis>
</taxonomicName>
was almost the same as it was reported for 
<taxonomicName box="[338,509,1645,1671]" class="Insecta" family="Phoridae" genus="Dicranopteron" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="malaysiensis">
<emphasis box="[338,509,1645,1671]" italics="true" pageId="7" pageNumber="84">D. malaysiensis</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Kistner DH" box="[554,730,1645,1671]" pageId="7" pageNumber="84" pagination="165 - 186" refId="ref3428" refString="Kistner DH (1982) A revision of the termitophilous genus Discoxenus with a study of the relationships of the genus and notes on its behavior (Coleoptera, Staphylinidae). Sociobiology 7: 165 - 186." type="journal article" year="1982">Kistner (1982)</bibRefCitation>
. The behavior of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[934,1108,1645,1672]" class="Insecta" family="Staphylinidae" genus="Odontoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis box="[934,955,1646,1672]" italics="true" pageId="7" pageNumber="84">O</emphasis>
. 
<emphasis box="[972,1108,1645,1671]" italics="true" pageId="7" pageNumber="84">thailandicus</emphasis>
</taxonomicName>
(
<figureCitation box="[148,242,166,193]" captionStart="Figure 5" captionStartId="7.[140,212,920,942]" captionTargetBox="[152,1096,176,887]" captionTargetId="figure@7.[140,1108,165,900]" captionTargetPageId="7" captionText="Figure 5. A Odontotermes proformosana, queen, workers and soldiers B the fungus garden of O. proformosana C Discoxenus katayamai, on fungus garden D Odontoxenus thailandicus, on fungus garden. Photos © Y. Katayama (5A–5B), T. Komatsu (5C–5D), 2007." figureDoi="http://doi.org/10.5281/zenodo.3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="8" pageNumber="85">Fig. 5D</figureCitation>
) was similar to that of 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" box="[522,677,167,193]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis box="[522,677,167,193]" italics="true" pageId="8" pageNumber="85">D. katayamai</emphasis>
</taxonomicName>
, but it moved much slower than 
<taxonomicName authorityName="Kanao &amp; Maruyama &amp; Sakchoowong" authorityYear="2010" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis italics="true" pageId="8" pageNumber="85">D. katayamai</emphasis>
</taxonomicName>
and it often stopped.
</paragraph>
<paragraph blockId="8.[140,381,322,349]" box="[140,381,322,349]" pageId="8" pageNumber="85">
<emphasis bold="true" box="[140,381,322,349]" pageId="8" pageNumber="85">Acknowledgment</emphasis>
</paragraph>
<paragraph blockId="8.[140,1108,378,827]" pageId="8" pageNumber="85">
We would like to express our thanks to Professor Osamu Tadauchi, Associate Professor Satoshi Kamitani, Associate Professor Layne Westover for reviewing the manuscript. We are also indebted to Mr. Narong Mahannop, the superintendent of Khao Yai National Park, for supporting and provisioning our fieldwork. Permission for sampling in this project was granted by the National Park, Wildlife, and Plant Conservation Department, 
<collectingCountry box="[298,398,554,580]" name="Thailand" pageId="8" pageNumber="85">Thailand</collectingCountry>
(No. 0907/14255). T. Kanao and M. Maruyama thank Dr. Yoko Takemastu for identification of the termite species. M. Maruyama thanks Alfred F. Newton and Margaret K. Thayer for allowing use of the Microptics system at the Field Museum of Natural History, and Y. Katayama and T. Komatsu for their assistance in the first author’s field work and for taking excellent pictures (
<figureCitation box="[899,965,694,721]" captionStart="Figure 5" captionStartId="7.[140,212,920,942]" captionTargetBox="[152,1096,176,887]" captionTargetId="figure@7.[140,1108,165,900]" captionTargetPageId="7" captionText="Figure 5. A Odontotermes proformosana, queen, workers and soldiers B the fungus garden of O. proformosana C Discoxenus katayamai, on fungus garden D Odontoxenus thailandicus, on fungus garden. Photos © Y. Katayama (5A–5B), T. Komatsu (5C–5D), 2007." figureDoi="http://doi.org/10.5281/zenodo.3767169" httpUri="https://zenodo.org/record/3767169/files/figure.png" pageId="8" pageNumber="85">Figs 5</figureCitation>
A–5D). This study was partially supported by a JSPS Postdoctoral Fellow for Research Abroad to M. Maruyama. This is a Contribution from the Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka (Ser. 6, No. 86).
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