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<document id="AFD3AA5389D187E14BDAFA82223FEE1B" ID-CLB-Dataset="9325" ID-DOI="10.11646/zootaxa.4894.2.1" ID-GBIF-Dataset="b084cbe7-0fe6-4dd4-a948-ae519e54292b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4315613" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607639617872" checkinUser="plazi" docAuthor="Waldren, George C., Williams, Kevin A., Cambra, Roberto A. &amp; Pitts, James P." docDate="2020" docId="3C273F3B3059FF9E2BE5FAA156B7FF79" docLanguage="en" docName="zootaxa.4894.2.1.pdf" docOrigin="Zootaxa 4894 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Invreiella chihuahuensis Waldren &amp; Williams &amp; Cambra &amp; Pitts 2020, sp. nov." docType="treatment" docUuid="2270A359-770B-4020-80DA-47F50714A978" docUuidSource="ZooBank" docVersion="11" lastPageNumber="190" masterDocId="C01E4743307FFFB62B72FFB75718FFAD" masterDocTitle="Systematic revision of the North American velvet ant genus Invreiella Suárez (Hymenoptera: Mutillidae) with description of eleven new species" masterLastPageNumber="205" masterPageNumber="151" pageNumber="189" updateTime="1698847345052" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C527CFFE3E92553B5B12EFC822E45B2D">Systematic revision of the North American velvet ant genus Invreiella Suárez (Hymenoptera: Mutillidae) with description of eleven new species</mods:title>
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<mods:namePart id="D055360F02DD4403CE23BFAB050E1DC7">Waldren, George C.</mods:namePart>
<mods:affiliation id="2BD7B6FB551E78294C1C904B770EE341">Utah State University, Department of Biology, Logan, Utah, USA. &amp; gcwaldren @ gmail. com; https: // orcid. org / 0000 - 0002 - 0007 - 1090 &amp; james. pitts @ usu. edu; https: // orcid. org / 0000 - 0002 - 1062 - 3258</mods:affiliation>
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<mods:namePart id="7F0930D612C253696F68DA0217EC6B39">Williams, Kevin A.</mods:namePart>
<mods:affiliation id="C7590843F890A7193DA704394CD83FA3">Plant Pest Diagnostics Center, California Department of Food and Agriculture, Sacramento, California, USA. &amp; kevin. williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
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<mods:namePart id="34AF17D4893C13804DB13440EA78B1EA">Cambra, Roberto A.</mods:namePart>
<mods:affiliation id="0116241C5EA2A10B0119B801AF008EBB">Museo de Invertebrados G. B. Fairchild, Estafeta Universitaria Apartado 00017, Universidad de Panamá, Panamá 0824, República de Panamá. &amp; cambramiup 60 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3799 - 4710</mods:affiliation>
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<mods:namePart id="379B11BE87C773DD7F919BD51481FE6D">Pitts, James P.</mods:namePart>
<mods:affiliation id="F48B8BBCBC1572A4415B3788BFA1FC8C">Utah State University, Department of Biology, Logan, Utah, USA. &amp; gcwaldren @ gmail. com; https: // orcid. org / 0000 - 0002 - 0007 - 1090 &amp; james. pitts @ usu. edu; https: // orcid. org / 0000 - 0002 - 1062 - 3258</mods:affiliation>
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<mods:title id="D0B7AA93021CD6668C0955F6393BC4E3">Zootaxa</mods:title>
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<mods:date id="A0200A88D691284953CDC85081477C29">2020</mods:date>
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<mods:number id="09F2F7FF1DEFF3620B299C871F74A37F">2020-12-09</mods:number>
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<mods:number id="C10BC135E535E7573AF68F92961701CD">4894</mods:number>
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<treatment id="3C273F3B3059FF9E2BE5FAA156B7FF79" ID-DOI="http://doi.org/10.5281/zenodo.4334935" ID-GBIF-Taxon="176085000" ID-Zenodo-Dep="4334935" LSID="urn:lsid:zoobank.org:act:2270A359-770B-4020-80DA-47F50714A978" httpUri="http://treatment.plazi.org/id/3C273F3B3059FF9E2BE5FAA156B7FF79" lastPageId="40" lastPageNumber="190" pageId="38" pageNumber="189">
<subSubSection id="FC94DDA63059FF902BE5FAA155C9FA9C" box="[151,721,1302,1329]" pageId="38" pageNumber="189" type="nomenclature">
<paragraph id="B4318E2D3059FF902BE5FAA155C9FA9C" blockId="38.[151,721,1302,1366]" box="[151,721,1302,1329]" pageId="38" pageNumber="189">
<heading id="EF7939413059FF902BE5FAA155C9FA9C" bold="true" box="[151,721,1302,1329]" fontSize="11" level="1" pageId="38" pageNumber="189" reason="1">
<emphasis id="86FA523F3059FF902BE5FAA155C9FA9C" bold="true" box="[151,721,1302,1329]" pageId="38" pageNumber="189">
10.
<taxonomicName id="738EF5AE3059FF902BB1FAA1557FFA9C" authority="Waldren" authorityName="Waldren &amp; Williams &amp; Cambra &amp; Pitts" authorityYear="2020" box="[195,615,1302,1329]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="chihuahuensis" status="sp. nov.">
<emphasis id="86FA523F3059FF902BB1FAA156EAFA9C" bold="true" box="[195,498,1302,1329]" italics="true" pageId="38" pageNumber="189">Invreiella chihuahuensis</emphasis>
Waldren
</taxonomicName>
,
<taxonomicNameLabel id="9DC9EF443059FF902906FAA055C9FA9C" box="[628,721,1303,1329]" pageId="38" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="FC94DDA63059FF902BE5FA8C55D7FAFB" box="[151,719,1338,1366]" pageId="38" pageNumber="189" type="description">
<paragraph id="B4318E2D3059FF902BE5FA8C55D7FAFB" blockId="38.[151,721,1302,1366]" box="[151,719,1338,1366]" pageId="38" pageNumber="189">
(
<figureCitation id="2CB592A83059FF902BEDFA8C57EEFAF8" box="[159,246,1339,1365]" captionStart="FIGURES 9–23" captionStartId="10.[151,264,1952,1976]" captionTargetBox="[188,1401,202,1928]" captionTargetId="figure-31@10.[181,1408,197,1936]" captionTargetPageId="10" captionText="FIGURES 9–23. Female habitus, dorsal view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4315617" httpUri="https://zenodo.org/record/4315617/files/figure.png" pageId="38" pageNumber="189">Figs 18</figureCitation>
,
<figureCitation id="2CB592A83059FF902A70FA8C566CFAF8" box="[258,372,1339,1365]" captionStart="FIGURES 24–31" captionStartId="11.[151,264,1808,1832]" captionTargetBox="[151,1435,257,1773]" captionTargetId="figure-74@11.[151,1435,257,1781]" captionTargetPageId="11" captionText="FIGURES 24–31. Female clypeus: 24–I. chihuahuensis (0000031), 25–I. breviclypeata (0000003), 26–I. cardinalis (0000018), 27–I. australis (0000001). Female head, left lateral view: 28–I. chihuahuensis (0000031), 29–I. satrapa (0000049). Female pronotum, left lateral view: 30–I. chihuahuensis (0000031), 31–I. satrapa (0000049)." figureDoi="http://doi.org/10.5281/zenodo.4315619" httpUri="https://zenodo.org/record/4315619/files/figure.png" pageId="38" pageNumber="189">24, 28, 30</figureCitation>
,
<figureCitation id="2CB592A83059FF902AF2FA8C5687FAF8" box="[384,415,1339,1365]" captionStart="FIGURES 64–79" captionStartId="15.[151,264,1772,1796]" captionTargetBox="[155,1435,161,1746]" captionTargetId="figure-114@15.[151,1435,151,1756]" captionTargetPageId="15" captionText="FIGURES 64–79. Invreiella australis (0000001): 64–female habitus, lateral view. Female head, left lateral view: 65–I. australis (0000001), 66–I. bimaculata (0000002), 67–I. mesomexicana (0000035), 68–I. acuminata (0000056), 69–I. cardinalis (0000018), 70–I. cuernavaca (0000032), 71–I. cephalargia (0000024), 72–I. manleyi (0000037), 73–I. breviclypeata (0000003), 74–I. chihuahuensis (0000031), 75–I. satrapa (0000049), 76–I. erythrocephala (0000034), 77–I. suarezi (0000051), 78–I. tequila (0000053), 79–Pseudomethoca frigida." figureDoi="http://doi.org/10.5281/zenodo.4315629" httpUri="https://zenodo.org/record/4315629/files/figure.png" pageId="38" pageNumber="189">74</figureCitation>
,
<figureCitation id="2CB592A83059FF902AD8FA8C56D1FAF8" box="[426,457,1339,1365]" captionStart="FIGURES 80–94" captionStartId="16.[151,264,1772,1796]" captionTargetBox="[151,1435,169,1751]" captionTargetId="figure-104@16.[151,1435,169,1757]" captionTargetPageId="16" captionText="FIGURES 80–94. Invreiella australis (0000001): 80–female head, anterior view. Femaleclypeus:81–I.australis (0000001), 82– I. bimaculata (0000002), 83–I. mesomexicana (0000035), 84–I. acuminata (0000056), 85–I. cardinalis (0000018), 86–I. cuernavaca (0000032), 87–I. cephalargia (0000024), 88–I. manleyi (0000037), 89–I. breviclypeata (0000003), 90–I. chihuahuensis (0000031), 91–I. satrapa (0000049), 92–I. erythrocephala (0000034), 93–I. suarezi (0000051), 94–I. tequila (0000053)." figureDoi="http://doi.org/10.5281/zenodo.4315631" httpUri="https://zenodo.org/record/4315631/files/figure.png" pageId="38" pageNumber="189">90</figureCitation>
,
<figureCitation id="2CB592A83059FF902AA6FA8D5518FAF8" box="[468,512,1338,1365]" captionStart="FIGURES 95–110" captionStartId="17.[151,264,1808,1832]" captionTargetBox="[151,1435,151,1752]" captionTargetId="figure-114@17.[151,1435,151,1760]" captionTargetPageId="17" captionText="FIGURES 95–110. Invreiella australis (0000001): 95–female habitus, dorsal view. Female mesosoma, left dorsal view: 96–I. australis (0000001), 97–I. bimaculata (0000002), 98–I. mesomexicana (0000035), 99–I. acuminata (0000056), 100–I. cardinalis (0000018), 101–I. cuernavaca (0000032), 102–I. cephalargia (0000024), 103–I. manleyi (0000037), 104–I. breviclypeata (000003), 105–I. chihuahuensis (0000031), 106–I. satrapa (0000049), 107–I. erythrocephala (0000034), 108–I. suarezi (0000051), 109–I. tequila (0000053), 110–Pseudomethoca frigida." figureDoi="http://doi.org/10.5281/zenodo.4315633" httpUri="https://zenodo.org/record/4315633/files/figure.png" pageId="38" pageNumber="189">105</figureCitation>
,
<figureCitation id="2CB592A83059FF90297EFA8C5520FAF8" box="[524,568,1339,1365]" captionStart="FIGURES 111–125" captionStartId="18.[151,264,1808,1832]" captionTargetBox="[151,1435,151,1772]" captionTargetId="figure-105@18.[151,1435,151,1778]" captionTargetPageId="18" captionText="FIGURES 111–125. Invreiella australis (0000001): 111–female habitus, lateral view. Female mesosoma, left lateral view: 112–I. australis (0000001), 113–I. bimaculata (0000002), 114–I. mesomexicana (0000035), 115–I. acuminata (0000056), 116–I. cardinalis (0000018), 117–I. cuernavaca (0000032), 118–I. cephalargia (0000024), 119–I. manleyi (0000037), 120–I. breviclypeata (0000003), 121–I. chihuahuensis (0000031), 122–I. satrapa (0000049), 123–I. erythrocephala (0000034), 124–I. suarezi (0000051), 125–I. tequila (0000053)." figureDoi="http://doi.org/10.5281/zenodo.4315635" httpUri="https://zenodo.org/record/4315635/files/figure.png" pageId="38" pageNumber="189">121</figureCitation>
,
<figureCitation id="2CB592A83059FF902936FA8D5569FAF8" box="[580,625,1338,1365]" captionStart="FIGURES 134–136" captionStartId="37.[151,264,1772,1796]" captionTargetBox="[149,1433,170,1720]" captionTargetId="figure-43@37.[149,1433,159,1722]" captionTargetPageId="37" captionText="FIGURES 134–136. Female habitus, dorsal view: Invreiella satrapa species-group. Scale bars = 2 mm. MAP 6. Geographic distribution of the Invreiella satrapa species-group." figureDoi="http://doi.org/10.5281/zenodo.4315643" httpUri="https://zenodo.org/record/4315643/files/figure.png" pageId="38" pageNumber="189">135</figureCitation>
; Map 6)
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63059FF902BE5FA3554F4F9FC" pageId="38" pageNumber="189" type="diagnosis">
<paragraph id="B4318E2D3059FF902BE5FA3554F4F9FC" blockId="38.[151,1437,1410,2013]" pageId="38" pageNumber="189">
<emphasis id="86FA523F3059FF902BE5FA355664FA31" bold="true" box="[151,380,1410,1436]" pageId="38" pageNumber="189">Diagnosis (female).</emphasis>
This species is distinguished from other members of the
<taxonomicName id="738EF5AE3059FF902F78FA345361FA31" baseAuthorityName="Gerstaecker" baseAuthorityYear="1874" box="[1034,1145,1411,1436]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="satrapa">
<emphasis id="86FA523F3059FF902F78FA345361FA31" box="[1034,1145,1411,1436]" italics="true" pageId="38" pageNumber="189">I. satrapa</emphasis>
</taxonomicName>
species-group by the following combination of characters: clypeus elongate, extending well-beyond base of mandibular socket (figs 24, 90), with medioapical tubercles basally conjoined and dorsoventrally triangular in shape, genal process spinose and curved (figs 28, 74), pronotal carina not strongly lobate posteriorly and not projecting away from mesosoma, not obscuring dorsal portion of pronotal-mesopleural suture (figs 30, 105, 121), and extreme apical margin of T2 with dark red-brown integument fringed medially with black setae (figs 18, 135).
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63059FF912BB5F9ED5687FA6C" lastPageId="39" lastPageNumber="190" pageId="38" pageNumber="189" type="description">
<paragraph id="B4318E2D3059FF902BB5F9ED541BF9D8" blockId="38.[151,1437,1410,2013]" box="[199,771,1626,1653]" pageId="38" pageNumber="189">
<emphasis id="86FA523F3059FF902BB5F9ED56DAF9D9" bold="true" box="[199,450,1626,1652]" pageId="38" pageNumber="189">Description (female).</emphasis>
Body length 8.0–
<quantity id="737623C83059FF9029F9F9ED55E7F9D8" box="[651,767,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.2449999999999999" pageId="38" pageNumber="189" unit="mm" value="12.45">12.45 mm</quantity>
.
</paragraph>
<paragraph id="B4318E2D3059FF902BB5F9C85358F870" blockId="38.[151,1437,1410,2013]" pageId="38" pageNumber="189">
<emphasis id="86FA523F3059FF902BB5F9C8561CF934" box="[199,260,1663,1689]" italics="true" pageId="38" pageNumber="189">Head</emphasis>
: Head 1.08–1.18 × as wide as mesosoma. Vertex and frons contiguously punctate. Antennal scrobe carina weakly arcuate, slightly more prominent internally, with inner tip of carina well-separated from antennal rim. Frons weakly transversely recessed and concave below antennal scrobe carina, antennal rim consequently slightly recessed and slightly visible when head viewed laterally. Antennal rim apically glabrous, rounded and not tuberculate. F1 1.88–2.3 × as long as F2. Clypeus plate-like and medially flattened, coarsely rugose-granulate, with central cluster of long raised setae, apical margin medially elongate and coming to distinct apex, with two close medioapical tubercles that are anterad-projecting, dorsoventrally triangular in shape, and basally conjoined. Mandible acuminate. Distance from posterior margin of eye to posterolateral corner of head 1.48–2.13 × maximum diameter of eye. Genal process spinose and curved, posterior genal carina nearly straight. Gena with separate yet close moderate punctures laterally, interpunctal space smooth. Postgena transversely rugose-striate.
</paragraph>
<paragraph id="B4318E2D3058FF912BB5FF2056B2FD28" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FF205627FF1D" box="[199,319,151,176]" italics="true" pageId="39" pageNumber="190">Mesosoma</emphasis>
: Mesosoma 1.18–1.26 × as wide as long. Dorsum and posterior face of mesosoma densely, contiguously punctate, surfaces coarse. Anterodorsolateral margin of pronotum outcurved. Pronotal carina prominent, glabrous, visible both dorsally and laterally, not strongly lobate posteriorly and not projecting away from mesosoma, not overlapping and obscuring dorsal portion of pronotal-mesopleural suture. Tubercle anterior of propodeal spiracle present. Width of propodeum greater than distance between propodeal spiracles in dorsal view. Lateral face of pronotum weakly rugose-striate, with scattered moderate punctures, lateral face posteriorly carinate along pronotal-mesopleural suture. Mesopleuron coarsely microgranulate to granulate anteriorly, with few punctures, posteriorly striate-rugose. Mesopleuron with vertical column of punctures weakly tuberculate, not forming anterior or posterior carinae. Mesopleuron posteriorly carinate along ventral two-thirds of mesopleural-metapleural suture, dorsal portion of suture obscure, faintly present. Metapleuron with ventral one-third striate-rugose with scattered punctures, dorsal two-thirds glabrous. Metapleural-propodeal suture with striae perpendicularly overlapping suture along ventral one-third. Lateral face of propodeum posteriorly with ventral one-third to one-half striate-rugose with few large punctures, anteriorly with dorsal one-half to two-thirds glabrous. Coxae coarsely sculptured, with small to large, deep punctures.
</paragraph>
<paragraph id="B4318E2D3058FF912BB5FD385628FC94" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FD385624FD05" box="[199,316,655,680]" italics="true" pageId="39" pageNumber="190">Metasoma</emphasis>
: T1–5 densely, contiguously punctate. Visible portion of pygidial plate not obscured by setal hood rugose-granulate. S2 densely punctate, punctures near contiguous in some areas, basolateral concave area less densely punctate, nearly glabrous between punctures. S3–6 densely punctate, coarsely microgranulate between punctures. Hypopygium with slightly arcuate transverse row of setae near apical margin, lateral setae longer than median setae.
</paragraph>
<paragraph id="B4318E2D3058FF912BB5FCF454DBFC08" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FCF456D9FCF0" box="[199,449,835,861]" italics="true" pageId="39" pageNumber="190">Integument coloration</emphasis>
: Orange, except the following ranging from dark red-brown to black: antenna, antennal rim, medioapical tubercles of clypeus, apical one-fourth to one-third of mandible, pleura, propodeum, legs, T1, extreme apical margins of T2 and S2, T3–6, and apical margins of S3–6.
</paragraph>
<paragraph id="B4318E2D3058FF912BB5FC185518FA30" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FC185653FC64" box="[199,331,943,969]" italics="true" pageId="39" pageNumber="190">Pubescence</emphasis>
: Frons ventrally with orange setae. Frons dorsally and vertex (including posterolateral corner of head) covered with mostly decumbent orange-red setae, with few scattered fuscous setae. Clypeus with long fuscous and whitish setae medially. Mandible with dorsal and ventral longitudinal row of light orange setae. Remainder of head with whitish setae. Dorsum of mesosoma mostly covered with decumbent orange-red setae, dorsal edges (except pronotum) with few raised fuscous setae, particularly at top of posterior face of propodeum. Pronotum without black setae, rather with orange setae. Remainder of mesosoma including legs with whitish setae (except tarsi with dense orange bristles). Anterior face of T1 with pale and fuscous setae. T1 apically fringed with wide band of black setae medially, laterally fringed with whitish setae. Dorsum of T2 covered with orange-red setae. Felt line of T2 and surrounding lateral area with whitish setae. T2 apically fringed with moderately wide band of black setae medially, not coming to median point, laterally fringed with whitish setae. T3–5 mostly covered and apically fringed with whitish setae, with few scattered dark raised setae and apical fringe with median patch of black setae, these black setal patches mostly consistent in width between T3–5, with T5 black setal patch sometimes covering median section of sclerite. T6 with pale yellow-orange setae surrounding pygidial plate, partially obscuring basal half. Remainder of metasoma with whitish setae.
</paragraph>
<paragraph id="B4318E2D3058FF912BB5FA115687FA6C" blockId="39.[151,1437,151,2013]" box="[199,415,1446,1473]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FA11563AFA6D" bold="true" box="[199,290,1446,1472]" pageId="39" pageNumber="190">MALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63058FF912BB5FA7C563DF9A4" pageId="39" pageNumber="190" type="etymology">
<paragraph id="B4318E2D3058FF912BB5FA7C563DF9A4" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5FA7C564AFA48" bold="true" box="[199,338,1483,1509]" pageId="39" pageNumber="190">Etymology:</emphasis>
The specific epithet is derived from the ecoregion this species is known to inhabit—the Chihuahuan Desert.
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63058FF912BB5F9A55672F9DB" pageId="39" pageNumber="190" type="distribution">
<paragraph id="B4318E2D3058FF912BB5F9A555EAF980" blockId="39.[151,1437,151,2013]" box="[199,754,1554,1581]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F9A5567BF981" bold="true" box="[199,355,1554,1580]" pageId="39" pageNumber="190">Distribution:</emphasis>
<collectingCountry id="CC99CEBD3058FF912A18F9A456D8F980" box="[362,448,1555,1581]" name="Mexico" pageId="39" pageNumber="190">Mexico</collectingCountry>
(
<collectingRegion id="764A40CF3058FF912ABCF9A45551F980" box="[462,585,1555,1581]" country="Mexico" name="Chihuahua" pageId="39" pageNumber="190">Chihuahua</collectingRegion>
and
<collectingRegion id="764A40CF3058FF91290DF9A455FEF980" box="[639,742,1555,1581]" country="Mexico" name="Durango" pageId="39" pageNumber="190">Durango</collectingRegion>
).
</paragraph>
<paragraph id="B4318E2D3058FF912BB5F9815672F9DB" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F981566FF9FD" bold="true" box="[199,375,1590,1616]" pageId="39" pageNumber="190">Biogeography:</emphasis>
Nearctic region (Chihuahuan Desert province); Mexican transition zone (Sierra Madre Occidental province).
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63058FF912BB5F9C956BBF934" box="[199,419,1662,1689]" pageId="39" pageNumber="190" type="biology_ecology">
<paragraph id="B4318E2D3058FF912BB5F9C956BBF934" blockId="39.[151,1437,151,2013]" box="[199,419,1662,1689]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F9C9563EF935" bold="true" box="[199,294,1662,1688]" pageId="39" pageNumber="190">Host(s):</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63058FF912BB5F9145691F910" box="[199,393,1699,1725]" pageId="39" pageNumber="190" type="discussion">
<paragraph id="B4318E2D3058FF912BB5F9145691F910" blockId="39.[151,1437,151,2013]" box="[199,393,1699,1725]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F9145625F910" bold="true" box="[199,317,1699,1725]" pageId="39" pageNumber="190">Remarks:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="FC94DDA63058FF9E2BB5F97156B7FF79" lastPageId="40" lastPageNumber="191" pageId="39" pageNumber="190" type="materials_examined">
<paragraph id="B4318E2D3058FF912BB5F97155A6F94E" blockId="39.[151,1437,151,2013]" box="[199,702,1734,1763]" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F97155A6F94E" bold="true" box="[199,702,1734,1763]" pageId="39" pageNumber="190">
Material examined (
<taxonomicName id="738EF5AE3058FF912AC8F9705565F94C" authorityName="Waldren &amp; Williams &amp; Cambra &amp; Pitts" authorityYear="2020" box="[442,637,1734,1761]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="chihuahuensis">
<emphasis id="86FA523F3058FF912AC8F9705565F94C" bold="true" box="[442,637,1734,1761]" italics="true" pageId="39" pageNumber="190">I. chihuahuensis</emphasis>
</taxonomicName>
,
<specimenCount id="A28845A43058FF9129F8F97F55B7F94E" box="[650,687,1736,1763]" pageId="39" pageNumber="190" type="female">4♀</specimenCount>
).
</emphasis>
</paragraph>
<paragraph id="B4318E2D3058FF912BB5F95C56AAF838" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="190">
<materialsCitation id="04E684703058FF912BB5F95C56AAF838" ID-GBIF-Occurrence="2995019357" collectingDate="1985-09-25" collectionCode="DGMC" collectorName="R. Huartos" country="Mexico" latitude="31.69" location="Ciudad Juarez" longLatPrecision="72" longitude="-106.424" pageId="39" pageNumber="190" specimenCode="HYM0000031" specimenCount="3" specimenCount-female="3" stateProvince="Chihuahua" typeStatus="holotype">
<emphasis id="86FA523F3058FF912BB5F95C5625F8A8" bold="true" box="[199,317,1771,1797]" pageId="39" pageNumber="190">
<typeStatus id="6B35308F3058FF912BB5F95C5620F8A8" box="[199,312,1771,1797]" pageId="39" pageNumber="190" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="A28845A43058FF912A3BF95C5646F8A8" box="[329,350,1771,1797]" pageId="39" pageNumber="190" type="female">♀</specimenCount>
(0000031–
<collectionCode id="D29F16E83058FF912A91F95C5521F8A8" box="[483,569,1771,1797]" pageId="39" pageNumber="190">DGMC</collectionCode>
) [tarsomere #5 of left mesoleg missing], [label 1 (white):]
<emphasis id="86FA523F3058FF912F8DF95C57C3F884" italics="true" pageId="39" pageNumber="190">CD. JUAREZ CHIH</emphasis>
/
<emphasis id="86FA523F3058FF912B81F8B8567CF884" box="[243,356,1807,1833]" italics="true" pageId="39" pageNumber="190">25/Sep/85</emphasis>
/
<emphasis id="86FA523F3058FF912A09F8B85521F885" box="[379,569,1807,1833]" italics="true" pageId="39" pageNumber="190">Roberto Huartos</emphasis>
[// label 2 (red):]
<typeStatus id="6B35308F3058FF912874F8B85482F884" box="[774,922,1807,1833]" pageId="39" pageNumber="190" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="A28845A43058FF9128D3F8B854AEF884" box="[929,950,1807,1833]" pageId="39" pageNumber="190" type="female">♀</specimenCount>
/
<taxonomicName id="738EF5AE3058FF9128BCF8B853F8F884" authorityName="Waldren &amp; Williams &amp; Cambra &amp; Pitts" authorityYear="2020" box="[974,1248,1807,1833]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="chihuahuensis">Invreiella chihuahuensis</taxonomicName>
/ Waldren, 2018 / GCW_
<specimenCode id="E42826563058FF912B85F8845682F8E0" box="[247,410,1843,1869]" pageId="39" pageNumber="190">HYM0000031</specimenCode>
[// label 3 (yellow):]
<emphasis id="86FA523F3058FF9129FCF8845423F8E0" box="[654,827,1843,1869]" italics="true" pageId="39" pageNumber="190">Compared with</emphasis>
/
<emphasis id="86FA523F3058FF912821F88454F9F8E0" box="[851,993,1843,1869]" italics="true" pageId="39" pageNumber="190">
<typeStatus id="6B35308F3058FF912821F88454F9F8E0" box="[851,993,1843,1869]" pageId="39" pageNumber="190" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
/
<taxonomicName id="738EF5AE3058FF91288AF88453A5F8E0" authorityName="Smith" authorityYear="1879" box="[1016,1213,1843,1869]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="macrocephala">
<emphasis id="86FA523F3058FF91288AF88453A5F8E0" box="[1016,1213,1843,1869]" italics="true" pageId="39" pageNumber="190">M. macrocephala</emphasis>
</taxonomicName>
[// label 4 (white):]
<taxonomicName id="738EF5AE3058FF912BE5F8E0561BF8DC" authorityName=", Suarez" authorityYear="1966" box="[151,259,1879,1905]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="86FA523F3058FF912BE5F8E0561BF8DC" box="[151,259,1879,1905]" italics="true" pageId="39" pageNumber="190">Invreiella</emphasis>
</taxonomicName>
/
<taxonomicName id="738EF5AE3058FF912A6CF8E0566CF8DD" baseAuthorityName="Gerstaecker" baseAuthorityYear="1874" box="[286,372,1879,1904]" class="Insecta" genus="Invreiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="satrapa">
<emphasis id="86FA523F3058FF912A6CF8E0566CF8DD" box="[286,372,1879,1904]" italics="true" pageId="39" pageNumber="190">satrapa</emphasis>
</taxonomicName>
. [(
<emphasis id="86FA523F3058FF912AE4F8E155B7F8DD" bold="true" box="[406,687,1878,1904]" pageId="39" pageNumber="190">
<collectingCountry id="CC99CEBD3058FF912AE4F8E15509F8DD" box="[406,529,1878,1904]" name="Mexico" pageId="39" pageNumber="190">MEXICO</collectingCountry>
:
<collectingRegion id="764A40CF3058FF91296DF8E155B1F8DD" box="[543,681,1878,1904]" country="Mexico" name="Chihuahua" pageId="39" pageNumber="190">Chihuahua</collectingRegion>
:
</emphasis>
<location id="B151D8F63058FF9129CAF8E05444F8DC" LSID="urn:lsid:plazi:treatment:3C273F3B3059FF9E2BE5FAA156B7FF79:B151D8F63058FF9129CAF8E05444F8DC" box="[696,860,1879,1905]" country="Mexico" latitude="31.69" longLatPrecision="72" longitude="-106.424" name="Ciudad Juarez" pageId="39" pageNumber="190" stateProvince="Chihuahua">Ciudad Juárez</location>
, [
<geoCoordinate id="D1BAE8EA3058FF912803F8E054F8F8DC" box="[881,992,1879,1905]" degrees="31.690" direction="north" orientation="latitude" pageId="39" pageNumber="190" precision="55" value="31.69">31.690°N</geoCoordinate>
<geoCoordinate id="D1BAE8EA3058FF912898F8E15368F8DC" box="[1002,1136,1878,1905]" degrees="106.424" direction="west" orientation="longitude" pageId="39" pageNumber="190" precision="55" value="-106.424">106.424°W</geoCoordinate>
],
<date id="C030A8ED3058FF912FF6F8E1520AF8DC" box="[1156,1298,1878,1905]" pageId="39" pageNumber="190" value="1985-09-25">
<collectingDate id="D07451053058FF912FF6F8E1520AF8DC" box="[1156,1298,1878,1905]" pageId="39" pageNumber="190" value="1985-09-25">25.Sep.1985</collectingDate>
</date>
,
<collectorName id="197BEBFB3058FF912E52F8E05285F8DC" box="[1312,1437,1879,1905]" pageId="39" pageNumber="190">R. Huartos</collectorName>
(
<specimenCount id="A28845A43058FF912BEDF8CD57ADF838" box="[159,181,1914,1941]" pageId="39" pageNumber="190" type="female">1♀</specimenCount>
–0000031–
<collectionCode id="D29F16E83058FF912A4FF8CD5681F838" box="[317,409,1914,1941]" pageId="39" pageNumber="190">DGMC</collectionCode>
))]
</materialsCitation>
</paragraph>
<paragraph id="B4318E2D3058FF912BB5F8285295F814" blockId="39.[151,1437,151,2013]" box="[199,1421,1950,1977]" pageId="39" pageNumber="190">
<materialsCitation id="04E684703058FF912BB5F8285291F814" ID-GBIF-Occurrence="2995019382" box="[199,1417,1950,1977]" collectionCode="NHMUK" collectorName="H. Hoge" country="Mexico" latitude="25.544" location="Ciudad Lerdo" longLatPrecision="74" longitude="-103.526" pageId="39" pageNumber="190" specimenCount="1" specimenCount-female="1" stateProvince="Durango" typeStatus="paratype">
<emphasis id="86FA523F3058FF912BB5F8285526F814" bold="true" box="[199,574,1950,1977]" pageId="39" pageNumber="190">
<typeStatus id="6B35308F3058FF912BB5F8285620F814" box="[199,312,1951,1977]" pageId="39" pageNumber="190" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="CC99CEBD3058FF912A36F82956A7F815" box="[324,447,1950,1976]" name="Mexico" pageId="39" pageNumber="190">MEXICO</collectingCountry>
:
<collectingRegion id="764A40CF3058FF912AB9F8285521F814" box="[459,569,1951,1977]" country="Mexico" name="Durango" pageId="39" pageNumber="190">Durango</collectingRegion>
:
</emphasis>
<location id="B151D8F63058FF912937F82855FBF814" LSID="urn:lsid:plazi:treatment:3C273F3B3059FF9E2BE5FAA156B7FF79:B151D8F63058FF912937F82855FBF814" box="[581,739,1951,1977]" country="Mexico" latitude="25.544" longLatPrecision="74" longitude="-103.526" name="Ciudad Lerdo" pageId="39" pageNumber="190" stateProvince="Durango">Ciudad Lerdo</location>
, [
<geoCoordinate id="D1BAE8EA3058FF912986F829547BF814" box="[756,867,1950,1977]" degrees="25.544" direction="north" orientation="latitude" pageId="39" pageNumber="190" precision="55" value="25.544">25.544°N</geoCoordinate>
<geoCoordinate id="D1BAE8EA3058FF912819F82954E9F814" box="[875,1009,1950,1977]" degrees="103.526" direction="west" orientation="longitude" pageId="39" pageNumber="190" precision="55" value="-103.526">103.526°W</geoCoordinate>
],
<collectorName id="197BEBFB3058FF912F70F828537AF814" box="[1026,1122,1951,1977]" pageId="39" pageNumber="190">H. Höge</collectorName>
(
<specimenCount id="A28845A43058FF912F02F829539EF814" box="[1136,1158,1950,1977]" pageId="39" pageNumber="190" type="female">1♀</specimenCount>
–0000057–
<collectionCode id="D29F16E83058FF912E7DF829529AF814" box="[1295,1410,1950,1977]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="190" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B4318E2D3058FF9E2BB5F87556B7FF79" blockId="39.[151,1437,151,2013]" lastBlockId="40.[151,1437,150,212]" lastPageId="40" lastPageNumber="191" pageId="39" pageNumber="190">
<emphasis id="86FA523F3058FF912BB5F87555E0F871" bold="true" box="[199,760,1986,2012]" pageId="39" pageNumber="190">Additional specimens (non-types, examined):</emphasis>
<materialsCitation id="04E684703058FF9E2875F87556B3FF79" ID-GBIF-Occurrence="2995019322" collectingDate="1947-07-05" collectionCode="MIUP" collectorName="Km. &amp; V. den Berghe" country="Mexico" elevation="2164" lastPageId="40" lastPageNumber="191" latitude="28.024" location="Creel Rd." longLatPrecision="74" longitude="-107.597" municipality="San Jose Babicora" pageId="39" pageNumber="190" specimenCount="2" specimenCount-female="2" stateProvince="Chihuahua">
<emphasis id="86FA523F3058FF912875F875533CF871" bold="true" box="[775,1060,1986,2012]" pageId="39" pageNumber="190">
<collectingCountry id="CC99CEBD3058FF912875F875549AF871" box="[775,898,1986,2012]" name="Mexico" pageId="39" pageNumber="190">MEXICO</collectingCountry>
:
<collectingRegion id="764A40CF3058FF9128E7F8755307F871" box="[917,1055,1986,2012]" country="Mexico" name="Chihuahua" pageId="39" pageNumber="190">Chihuahua</collectingRegion>
:
</emphasis>
<collectingMunicipality id="545514573058FF912F41F874520BF870" box="[1075,1299,1987,2013]" pageId="39" pageNumber="190">San José Babícora</collectingMunicipality>
, [
<geoCoordinate id="D1BAE8EA3058FF912E5FF8755284F870" box="[1325,1436,1986,2013]" degrees="29.252" direction="north" orientation="latitude" pageId="39" pageNumber="190" precision="55" value="29.252">29.252°N</geoCoordinate>
<geoCoordinate id="D1BAE8EA3057FF9E2BE5FF215606FF1C" box="[151,286,150,177]" degrees="107.750" direction="west" orientation="longitude" pageId="40" pageNumber="191" precision="55" value="-107.75">107.750°W</geoCoordinate>
],
<quantity id="737623C83057FF9E2A42FF215692FF1D" box="[304,394,150,176]" metricMagnitude="3" metricUnit="m" metricValue="2.16408" pageId="40" pageNumber="191" unit="ft" value="7100.0">
<elevation id="3FA3691E3057FF9E2A42FF215692FF1D" box="[304,394,150,176]" metricMagnitude="3" metricUnit="m" metricValue="2.16408" pageId="40" pageNumber="191" unit="ft" value="7100.0">7100 ft.</elevation>
</quantity>
,
<date id="C030A8ED3057FF9E2AE5FF215502FF1C" box="[407,538,150,177]" pageId="40" pageNumber="191" value="1947-07-05">
<collectingDate id="D07451053057FF9E2AE5FF215502FF1C" box="[407,538,150,177]" pageId="40" pageNumber="191" value="1947-07-05">05.Jul.1947</collectingDate>
</date>
(
<specimenCount id="A28845A43057FF9E2958FF21555AFF1D" box="[554,578,150,176]" pageId="40" pageNumber="191" type="female">1♀</specimenCount>
–
<collectionCode id="D29F16E83057FF9E2921FF2155BEFF1D" box="[595,678,150,176]" pageId="40" pageNumber="191">MIUP</collectionCode>
);
<collectorName id="197BEBFB3057FF9E29C8FF2055F4FF1C" box="[698,748,151,177]" pageId="40" pageNumber="191">Km.</collectorName>
7
<location id="B151D8F63057FF9E287BFF21546DFF1D" LSID="urn:lsid:plazi:treatment:3C273F3B3059FF9E2BE5FAA156B7FF79:B151D8F63057FF9E287BFF21546DFF1D" box="[777,885,150,176]" country="Mexico" latitude="28.024" longLatPrecision="74" longitude="-107.597" municipality="San Jose Babicora" name="Creel Rd." pageId="40" pageNumber="191" stateProvince="Chihuahua">Creel Rd.</location>
[=Carretera a Creel], [28.024°N 107.597°W],
<collectorName id="197BEBFB3057FF9E2EF7FF20560FFF79" pageId="40" pageNumber="191">V. den Berghe</collectorName>
(
<specimenCount id="A28845A43057FF9E2A55FF0D5627FF79" box="[295,319,186,212]" pageId="40" pageNumber="191" type="female">1♀</specimenCount>
–
<collectionCode id="D29F16E83057FF9E2A22FF0D56BBFF79" box="[336,419,186,212]" pageId="40" pageNumber="191">MIUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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