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<mods:title id="28DB170B49FADC9DB031E4F694262AF6">The Lace Web Spider Genus Vytfutia Deeleman-Reinhold (Araneae, Phyxelididae) in the Indo-Pacific Region</mods:title>
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<treatment id="03B487984B62FFB0293F927FE663F4FF" ID-DOI="http://doi.org/10.5281/zenodo.12692113" ID-GBIF-Taxon="234981480" ID-Zenodo-Dep="12692113" LSID="urn:lsid:plazi:treatment:03B487984B62FFB0293F927FE663F4FF" httpUri="http://treatment.plazi.org/id/03B487984B62FFB0293F927FE663F4FF" lastPageId="11" lastPageNumber="340" pageId="10" pageNumber="339">
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<paragraph id="8BA2368E4B62FFB1293F927FE032F75F" blockId="10.[123,1174,541,655]" box="[123,595,541,565]" pageId="10" pageNumber="339">
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<taxonomicName id="4C1D4D0D4B62FFB1293F927FE032F75F" ID-CoL="5BSNK" authority="Deeleman-Reinhold, 1989" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[123,595,541,565]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="339" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B969EA9C4B62FFB1293F927FE032F75F" bold="true" box="[123,595,541,565]" pageId="10" pageNumber="339">
<emphasis id="B969EA9C4B62FFB1293F927FE349F75F" bold="true" box="[123,296,541,565]" italics="true" pageId="10" pageNumber="339">Vytfutia pallens</emphasis>
<bibRefCitation id="EF8C4B7F4B62FFB1286B927CE032F75F" author="DEELEMAN-REINHOLD, C. L." box="[303,595,542,565]" pageId="10" pageNumber="339" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30765054B62FFB1293F925CE32FF739" box="[123,334,574,595]" pageId="10" pageNumber="339" type="description">
<paragraph id="8BA2368E4B62FFB1293F925CE32FF739" blockId="10.[123,1174,541,655]" box="[123,334,574,595]" pageId="10" pageNumber="339">
<figureCitation id="13262A0B4B62FFB1293F925CE369F739" box="[123,264,574,595]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figures 4549</figureCitation>
,
<figureCitation id="13262A0B4B62FFB12856925CE32FF739" box="[274,334,574,595]" captionStart-0="FIGURES" captionStart-1="FIGURE" captionStartId-0="25.[155,167,1487,1506]" captionStartId-1="26.[155,167,1071,1090]" captionTargetBox-0="[153,1147,252,1446]" captionTargetBox-1="[120,1176,216,1056]" captionTargetId-0="figure-24@25.[120,1176,216,1475]" captionTargetId-1="figure-19@26.[120,1176,216,1056]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 5661. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." captionText-1="FIGURE 62. Vytfutia spp. Distributions on map of Eurasia and Australasia. Filled circle. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. Triangle. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia. Diamond. Vytfutia halandrefana new species, Madagascar. Star. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia." figureDoi-0="http://doi.org/10.5281/zenodo.11513207" figureDoi-1="http://doi.org/10.5281/zenodo.11513211" httpUri-0="https://zenodo.org/record/11513207/files/figure.png" httpUri-1="https://zenodo.org/record/11513211/files/figure.png" pageId="10" pageNumber="339">6062</figureCitation>
</paragraph>
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<subSubSection id="C30765054B62FFB1293F9200E0AAF7E5" pageId="10" pageNumber="339" type="reference_group">
<paragraph id="8BA2368E4B62FFB1293F9200E0AAF7E5" blockId="10.[123,1174,541,655]" pageId="10" pageNumber="339">
<taxonomicName id="4C1D4D0D4B62FFB1293F9200E386F71F" ID-CoL="5BSNK" authority="Deeleman-Reinhold, 1989" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[123,487,610,629]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="339" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B969EA9C4B62FFB1293F9200E29EF71F" box="[123,255,610,629]" italics="true" pageId="10" pageNumber="339">Vytfutia pallens</emphasis>
<bibRefCitation id="EF8C4B7F4B62FFB128479200E386F71F" author="DEELEMAN-REINHOLD, C. L." box="[259,487,610,629]" pageId="10" pageNumber="339" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989</bibRefCitation>
</taxonomicName>
:
<specimenCount id="9D1BFD074B62FFB128AA9200E39FF71F" box="[494,510,610,629]" count="1" pageId="10" pageNumber="339" type="male"></specimenCount>
M
<specimenCount id="9D1BFD074B62FFB12B559200E07FF71F" box="[529,542,610,629]" count="1" pageId="10" pageNumber="339" type="female"></specimenCount>
F
<typeStatus id="54A6882C4B62FFB12B759200E0E2F71F" box="[561,643,610,629]" pageId="10" pageNumber="339" type="paratype">paratypes</typeStatus>
from
<location id="8EC260554B62FFB12BF29200E173F71F" LSID="urn:lsid:plazi:treatment:03B487984B62FFB0293F927FE663F4FF:8EC260554B62FFB12BF29200E173F71F" box="[694,786,610,629]" country="Malaysia" name="Niah Cave" pageId="10" pageNumber="339">Niah Cave</location>
,
<collectingCountry id="F30A761E4B62FFB12A5E9200E107F71F" box="[794,870,610,629]" name="Malaysia" pageId="10" pageNumber="339">Sarawak</collectingCountry>
,
<collectingCountry id="F30A761E4B62FFB12A299200E1DCF71F" box="[877,957,610,629]" name="Malaysia" pageId="10" pageNumber="339">Malaysia</collectingCountry>
,
<date id="FFA3104E4B62FFB12A819200E65CF71F" box="[965,1085,610,629]" pageId="10" pageNumber="339" value="1984-04-10">
<collectingDate id="EFE7E9A64B62FFB12A819200E65CF71F" box="[965,1085,610,629]" pageId="10" pageNumber="339" value="1984-04-10">10 April 1984</collectingDate>
</date>
, C. Deeleman and C. Hug,
<collectionCode id="ED0CAE4B4B62FFB12807921EE3E4F7E5" box="[323,389,636,655]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="339" type="Museum">RMNH</collectionCode>
, examined.
<bibRefCitation id="EF8C4B7F4B62FFB128A9921EE0A6F7E5" author="GRISWOLD, C. E. &amp; RAMIREZ, M. J. &amp; CODDINGTON, J. A. &amp; AND PLATNICK, N. I." box="[493,711,636,655]" pageId="10" pageNumber="339" pagination="1 - 324" refId="ref12029" refString="GRISWOLD, C. E., RAMIREZ, M. J., CODDINGTON, J. A. AND PLATNICK, N. I. 2005. Atlas of phylogenetic data for entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny. Proceedings of the California Academy of Sciences, ser. 4, 56 (Suppl. II): 1 - 324." type="journal article" year="2005">
Griswold
<emphasis id="B969EA9C4B62FFB12B06921EE009F7E5" box="[578,616,636,655]" italics="true" pageId="10" pageNumber="339">et al</emphasis>
., 2005: 90
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30765054B62FFB029EF92FDE1A0F43E" lastPageId="11" lastPageNumber="340" pageId="10" pageNumber="339" type="diagnosis">
<paragraph id="8BA2368E4B62FFB129EF92FDE3B7F6F2" blockId="10.[123,1175,671,1725]" pageId="10" pageNumber="339">
<emphasis id="B969EA9C4B62FFB129EF92FDE378F7DC" bold="true" box="[171,281,671,694]" pageId="10" pageNumber="339">Diagnosis</emphasis>
: Males with palpal tibia with elongate apical spine (
<figureCitation id="13262A0B4B62FFB12A1E92FDE180F7DC" box="[858,993,671,694]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figs. 46, 47</figureCitation>
), embolus short, straight, arises on prolateral side of bulb (near 8 oclock), median apophysis hooked at apex (
<figureCitation id="13262A0B4B62FFB129C792BDE367F79C" box="[131,262,735,758]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figs. 45, 46</figureCitation>
); females with epigynum with copulatory opening rims oblique, diverging anteriorly, posterior margin straight to slightly convex (
<figureCitation id="13262A0B4B62FFB12BCF9362E082F67D" box="[651,739,768,791]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Fig. 48</figureCitation>
), vulva simple, with paired lobes of spermathecae extending far anteriad of copulatory openings (
<figureCitation id="13262A0B4B62FFB12A469342E130F65D" box="[770,849,800,823]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Fig. 49</figureCitation>
); largest
<taxonomicName id="4C1D4D0D4B62FFB12AF49342E663F65D" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[944,1026,800,823]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="339" phylum="Arthropoda" rank="genus">
<emphasis id="B969EA9C4B62FFB12AF49342E663F65D" box="[944,1026,800,823]" italics="true" pageId="10" pageNumber="339">Vytfutia</emphasis>
</taxonomicName>
(see
<figureCitation id="13262A0B4B62FFB12D7D9342E2F9F63D" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 5661. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="10" pageNumber="339">Figs. 60, 61</figureCitation>
), total length of males 4.405.00, of females 6.207.70; markings pale; legs long, male metatarsus I with base of concavity at more than 0.45 metatarsus length, spur high, height greater than 0.09 metatarsus length (
<figureCitation id="13262A0B4B62FFB1280293E3E3ABF6F2" box="[326,458,897,920]" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 5661. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="10" pageNumber="339">Figs. 60, 61</figureCitation>
).
</paragraph>
<paragraph id="8BA2368E4B62FFB029EF93C3E16AF479" blockId="10.[123,1175,671,1725]" lastBlockId="11.[123,1174,220,405]" lastPageId="11" lastPageNumber="340" pageId="10" pageNumber="339">
<emphasis id="B969EA9C4B62FFB129EF93C3E284F6D2" bold="true" box="[171,229,929,952]" pageId="10" pageNumber="339">Male</emphasis>
(
<typeStatus id="54A6882C4B62FFB129B093C3E335F6D2" box="[244,340,929,952]" pageId="10" pageNumber="339" type="paratype">paratype</typeStatus>
, after
<bibRefCitation id="EF8C4B7F4B62FFB128DC93C3E140F6D2" author="DEELEMAN-REINHOLD, C. L." box="[408,801,929,952]" pageId="10" pageNumber="339" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989: 622-623</bibRefCitation>
): Total length 4.40. Carapace light grey-brown, ocular area slightly darker; chelicerae dark brown; labium, palpal coxae, sternum, and legs light grey-brown; abdomen pale with 3 snowy granulations on dorsal surface. Carapace 2.20 long, 1.70 wide; carapace length 1.29 times width;
<specimenCode id="DBBB9EF54B62FFB12BE99463E177F172" box="[685,790,1025,1048]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="10" pageNumber="339" type="Herbarium">PER 1.14</specimenCode>
times
<collectionCode id="ED0CAE4B4B62FFB12A1B9463E1F6F172" box="[863,919,1025,1048]" pageId="10" pageNumber="339">AER</collectionCode>
,
<specimenCode id="DBBB9EF54B62FFB12AE09463E66DF172" box="[932,1036,1025,1048]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="10" pageNumber="339" type="Herbarium">PER 2.28</specimenCode>
times OAL; OAL 0.57 times OQL; OQA 0.625 times OQP; ratio of eyes
<collectionCode id="ED0CAE4B4B62FFB12A729440E104F153" box="[822,869,1058,1081]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
:AL:
<collectionCode id="ED0CAE4B4B62FFB12AD69440E1DCF153" box="[914,957,1058,1081]" country="USA" name="Pratt Museum" pageId="10" pageNumber="339">PM</collectionCode>
:
<collectionCode id="ED0CAE4B4B62FFB12A849440E182F153" box="[960,995,1058,1081]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="339" type="Museum">PL</collectionCode>
, 1.0:2.0:3.0:3.0; AM-AM, AM-AL equal to
<collectionCode id="ED0CAE4B4B62FFB128EE9420E3B4F133" box="[426,469,1090,1113]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
diameter;
<collectionCode id="ED0CAE4B4B62FFB12B099420E0C7F133" box="[589,678,1090,1113]" pageId="10" pageNumber="339">PM-PM</collectionCode>
,
<collectionCode id="ED0CAE4B4B62FFB12BF09420E163F133" box="[692,770,1090,1113]" pageId="10" pageNumber="339">PM-PL</collectionCode>
equal to
<collectionCode id="ED0CAE4B4B62FFB12A2A9420E1F4F133" box="[878,917,1090,1113]" country="USA" name="Pratt Museum" pageId="10" pageNumber="339">PM</collectionCode>
diameter; AL touching
<collectionCode id="ED0CAE4B4B62FFB1293F9400E2FCF113" box="[123,157,1122,1145]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="339" type="Museum">PL</collectionCode>
. Sternum length 1.26 times width; labium length 0.75 times width; palpal coxae length 2.0 times width. Legs with typical spination; femur I length 1.25 times carapace width; metatarsus I as in
<figureCitation id="13262A0B4B62FFB129DC94C0E37CF1D3" box="[152,285,1186,1209]" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 5661. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="10" pageNumber="339">Figs. 60, 61</figureCitation>
, clasping spine extends beyond median spur, spur long, length nearly ½ segment width. Leg measurements: I: 2.3 + 0.9 + 2.2 + 1.9 + 1.2 = [8.5]; II: 2.1 + 0.7 + 1.8 + 1.8 + 1.0 = [7.4];
<collectionCode id="ED0CAE4B4B62FFB129F89481E2BBF190" box="[188,218,1251,1274]" pageId="10" pageNumber="339">III</collectionCode>
: 2.1 + 0.8 + 1.5 + 1.7 + 0.9 = [7.0];
<collectionCode id="ED0CAE4B4B62FFB12B1F9481E018F190" box="[603,633,1251,1274]" pageId="10" pageNumber="339">IV</collectionCode>
: 2.3 + 0.9 + 1.9 + 1.9 + 1.0 = [8.0]; palp: 1.2 + 0.4 + 0.6 + NA + 0.6 = [2.8]; leg formula 1423. Male palp as in
<figureCitation id="13262A0B4B62FFB12A509561E1F6F070" box="[788,919,1283,1306]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figs. 4547</figureCitation>
; tibia length 1.21 times cymbial length, RTA length 0.23 times tibia (
<figureCitation id="13262A0B4B62FFB12B1C9541E0B8F050" box="[600,729,1315,1338]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figs. 46, 47</figureCitation>
), MA origin 0.36 distance to tegular base, MA length 0.83 times that of tegulum, MA length 7.86 times width (
<figureCitation id="13262A0B4B62FFB12A159526E1B3F031" box="[849,978,1348,1371]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Figs. 45, 46</figureCitation>
). Variation (N=3): Total length 4.40 to 5.00.
<emphasis id="B969EA9C4B62FFB128E29506E398F011" bold="true" box="[422,505,1380,1403]" pageId="10" pageNumber="339">Female</emphasis>
(
<typeStatus id="54A6882C4B62FFB12B489506E00FF011" box="[524,622,1380,1403]" pageId="10" pageNumber="339" type="paratype">Paratype</typeStatus>
, after
<bibRefCitation id="EF8C4B7F4B62FFB12BFD9506E628F011" author="DEELEMAN-REINHOLD, C. L." box="[697,1097,1380,1403]" pageId="10" pageNumber="339" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989: 622-623</bibRefCitation>
):
<collectorName id="26E853584B62FFB12D1B9506E6F4F011" box="[1119,1173,1380,1403]" pageId="10" pageNumber="339">Total</collectorName>
length 6.70.
<collectorName id="26E853584B62FFB129BB95E6E304F0F1" box="[255,357,1412,1435]" pageId="10" pageNumber="339">Markings</collectorName>
as in male.
<collectorName id="26E853584B62FFB128A695E6E024F0F1" box="[482,581,1412,1435]" pageId="10" pageNumber="339">Carapace</collectorName>
3.30 long, 2.30 wide, carapace length 1.43 times width, carapace height 0.47 width;
<specimenCode id="DBBB9EF54B62FFB128EF95C6E070F0D1" box="[427,529,1444,1467]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="10" pageNumber="339" type="Herbarium">PER 1.11</specimenCode>
times
<collectionCode id="ED0CAE4B4B62FFB12B1395C6E0EEF0D1" box="[599,655,1444,1467]" pageId="10" pageNumber="339">AER</collectionCode>
,
<specimenCode id="DBBB9EF54B62FFB12BDE95C6E163F0D1" box="[666,770,1444,1467]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="10" pageNumber="339" type="Herbarium">PER 2.50</specimenCode>
times OAL; OAL equals OQL; OQA 0.59 times OQP; clypeal height 1.25 times
<collectionCode id="ED0CAE4B4B62FFB12BC895A6E0D6F0B1" box="[652,695,1476,1499]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
diameter; ratio of eyes
<collectionCode id="ED0CAE4B4B62FFB12AA095A6E672F0B1" box="[996,1043,1476,1499]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
:AL:
<collectionCode id="ED0CAE4B4B62FFB12D0595A6E60DF0B1" box="[1089,1132,1476,1499]" country="USA" name="Pratt Museum" pageId="10" pageNumber="339">PM</collectionCode>
:
<collectionCode id="ED0CAE4B4B62FFB12D2B95A6E6F3F0B1" box="[1135,1170,1476,1499]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="339" type="Museum">PL</collectionCode>
, 1.0:1.6:1.6:1.2; AM-AM 0.6 times
<collectionCode id="ED0CAE4B4B62FFB12B459587E04DF096" box="[513,556,1509,1532]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
diameter, AM-AL equals
<collectionCode id="ED0CAE4B4B62FFB12A179587E11FF096" box="[851,894,1509,1532]" country="Australia" name="Australian Museum" pageId="10" pageNumber="339" type="Museum">AM</collectionCode>
diameter;
<specimenCode id="DBBB9EF54B62FFB12ABC9587E6F4F096" box="[1016,1173,1509,1532]" collectionCode="PM-PM" pageId="10" pageNumber="339">PM-PM 1.125</specimenCode>
times
<collectionCode id="ED0CAE4B4B62FFB129849667E286F376" box="[192,231,1541,1564]" country="USA" name="Pratt Museum" pageId="10" pageNumber="339">PM</collectionCode>
diameter,
<specimenCode id="DBBB9EF54B62FFB1281B9667E386F376" box="[351,487,1541,1564]" collectionCode="PM-PL" pageId="10" pageNumber="339">PM-PL 1.25</specimenCode>
times
<collectionCode id="ED0CAE4B4B62FFB12B739667E001F376" box="[567,608,1541,1564]" country="USA" name="Pratt Museum" pageId="10" pageNumber="339">PM</collectionCode>
, AL touching
<collectionCode id="ED0CAE4B4B62FFB12A409667E147F376" box="[772,806,1541,1564]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="339" type="Museum">PL</collectionCode>
.
<collectorName id="26E853584B62FFB12A719667E1C6F376" box="[821,935,1541,1564]" pageId="10" pageNumber="339">Chelicerae</collectorName>
1.30 long; cheliceral length 12.0 times clypeal height; sternum length 1.34 times width; labium length 1.09 times width; palpal coxae length 1.75 times width.
<collectorName id="26E853584B62FFB12B659627E034F336" box="[545,597,1605,1628]" pageId="10" pageNumber="339">Legs</collectorName>
with typical spination.
<collectorName id="26E853584B62FFB12A1F9627E1D5F336" box="[859,948,1605,1628]" pageId="10" pageNumber="339">Femur I</collectorName>
1.26 times carapace width.
<collectorName id="26E853584B62FFB129829604E28EF317" box="[198,239,1638,1661]" pageId="10" pageNumber="339">Leg</collectorName>
measurements: I: 2.9 + 1.2 + 2.5 + 2.4 + 1.3 = [10.30]; II: 2.7 + 1.1 + 2.2 + 2.0 + 1.0 = [9.00];
<collectionCode id="ED0CAE4B4B62FFB1299B96E4E29CF3F7" box="[223,253,1670,1693]" pageId="10" pageNumber="339">III</collectionCode>
: 2.4 + 1.0 + 1.7 + 1.7 + 1.0 = [7.80];
<collectionCode id="ED0CAE4B4B62FFB12BC896E4E0CBF3F7" box="[652,682,1670,1693]" pageId="10" pageNumber="339">IV</collectionCode>
: 2.7 + 1.1 + 1.9 + 1.9 + 1.0 = [8.60]; palp: 1.1 + 0.25 + 0.70 + NA + 1.0 = [3.05], leg formula 1243.
<collectorName id="26E853584B62FFB12B9496C4E17EF3D7" box="[720,799,1702,1725]" pageId="10" pageNumber="339">Female</collectorName>
genitalia as in
<figureCitation id="13262A0B4B62FFB12A8096C4E670F3D7" box="[964,1041,1702,1725]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="10" pageNumber="339">Fig. 48</figureCitation>
(epigynum) and
<figureCitation id="13262A0B4B63FFB029ED90BEE297F599" box="[169,246,220,243]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 4549. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 4547. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="11" pageNumber="340">Fig. 49</figureCitation>
(vulva). Epigynum width 2.66 times length; epigynum MLW 2.20 times MLL, MLW 1.50 times LLW. Variation (N=12): Total length 6.20 to 7.70.
</paragraph>
<paragraph id="8BA2368E4B63FFB029EF917EE1A0F43E" blockId="11.[123,1174,220,405]" pageId="11" pageNumber="340">
<emphasis id="B969EA9C4B63FFB029EF917EE301F459" bold="true" box="[171,352,284,307]" pageId="11" pageNumber="340">Natural History</emphasis>
:
<bibRefCitation id="EF8C4B7F4B63FFB0282B917EE0A5F459" author="DEELEMAN-REINHOLD, C. L." box="[367,708,284,307]" pageId="11" pageNumber="340" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold (1989: 619</bibRefCitation>
, 620, 624) describes this species from Niah Cave,
<collectingCountry id="F30A761E4B63FFB029FB915FE37FF43E" box="[191,286,317,340]" name="Malaysia" pageId="11" pageNumber="340">Sarawak</collectingCountry>
,
<collectingCountry id="F30A761E4B63FFB0286D915FE3ECF43E" box="[297,397,317,340]" name="Malaysia" pageId="11" pageNumber="340">Malaysia</collectingCountry>
, a cave famous for the diversity of endemic species.
</paragraph>
</subSubSection>
<subSubSection id="C30765054B63FFB029EF913FE147F41E" box="[171,806,349,372]" pageId="11" pageNumber="340" type="distribution">
<paragraph id="8BA2368E4B63FFB029EF913FE147F41E" blockId="11.[123,1174,220,405]" box="[171,806,349,372]" pageId="11" pageNumber="340">
<emphasis id="B969EA9C4B63FFB029EF913FE357F41E" bold="true" box="[171,310,349,372]" pageId="11" pageNumber="340">Distribution</emphasis>
: Known only from the
<typeStatus id="54A6882C4B63FFB02B70913FE003F41E" box="[564,610,349,372]" pageId="11" pageNumber="340">type</typeStatus>
locality (
<figureCitation id="13262A0B4B63FFB02B8D913FE178F41E" box="[713,793,349,372]" captionStart="FIGURE" captionStartId="26.[155,167,1071,1090]" captionTargetBox="[120,1176,216,1056]" captionTargetId="figure-19@26.[120,1176,216,1056]" captionTargetPageId="26" captionText="FIGURE 62. Vytfutia spp. Distributions on map of Eurasia and Australasia. Filled circle. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. Triangle. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia. Diamond. Vytfutia halandrefana new species, Madagascar. Star. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.11513211" httpUri="https://zenodo.org/record/11513211/files/figure.png" pageId="11" pageNumber="340">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30765054B63FFB029EF911CE663F4FF" box="[171,1026,382,405]" pageId="11" pageNumber="340" type="materials_examined">
<paragraph id="8BA2368E4B63FFB029EF911CE663F4FF" blockId="11.[123,1174,220,405]" box="[171,1026,382,405]" pageId="11" pageNumber="340">
<emphasis id="B969EA9C4B63FFB029EF911CE3E8F4FF" bold="true" box="[171,393,382,405]" pageId="11" pageNumber="340">Material Examined</emphasis>
:
<materialsCitation id="3B753CD34B63FFB028DC911CE19EF4FF" ID-GBIF-Occurrence="4903682308" box="[408,1023,382,405]" collectionCode="RMNH" country="Malaysia" location="Niah Cave" pageId="11" pageNumber="340" specimenCount="1" typeStatus="paratype">
Only the
<typeStatus id="54A6882C4B63FFB028B8911CE003F4FF" box="[508,610,382,405]" pageId="11" pageNumber="340" type="paratype">paratypes</typeStatus>
from
<location id="8EC260554B63FFB02BE1911CE178F4FF" LSID="urn:lsid:plazi:treatment:03B487984B62FFB0293F927FE663F4FF:8EC260554B63FFB02BE1911CE178F4FF" box="[677,793,382,405]" country="Malaysia" name="Niah Cave" pageId="11" pageNumber="340">Niah Cave</location>
,
<collectingCountry id="F30A761E4B63FFB02A61911CE1E5F4FF" box="[805,900,382,405]" name="Malaysia" pageId="11" pageNumber="340">Sarawak</collectingCountry>
, in
<collectionCode id="ED0CAE4B4B63FFB02AEF911CE19EF4FF" box="[939,1023,382,405]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="340" type="Museum">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>