The Lace Web Spider Genus Vytfutia Deeleman-Reinhold (Araneae, Phyxelididae) in the Indo-Pacific Region Author Griswold, Charles E. text Proceedings of the California Academy of Sciences 2022 2022-07-29 67 14 329 355 journal article 299622 10.5281/zenodo.11513175 df93398d-58f3-42f9-a866-efb0cd2debcd 0068-547X 11513175 Vytfutia pallens Deeleman-Reinhold, 1989 Figures 45–49 , 60–62 Vytfutia pallens Deeleman-Reinhold, 1989 : ♂ M ♀ F paratypes from Niah Cave , Sarawak , Malaysia , 10 April 1984 , C. Deeleman and C. Hug, RMNH , examined. Griswold et al ., 2005: 90 . Diagnosis : Males with palpal tibia with elongate apical spine ( Figs. 46, 47 ), embolus short, straight, arises on prolateral side of bulb (near 8 o’clock), median apophysis hooked at apex ( Figs. 45, 46 ); females with epigynum with copulatory opening rims oblique, diverging anteriorly, posterior margin straight to slightly convex ( Fig. 48 ), vulva simple, with paired lobes of spermathecae extending far anteriad of copulatory openings ( Fig. 49 ); largest Vytfutia (see Figs. 60, 61 ), total length of males 4.40–5.00, of females 6.20–7.70; markings pale; legs long, male metatarsus I with base of concavity at more than 0.45 metatarsus length, spur high, height greater than 0.09 metatarsus length ( Figs. 60, 61 ). Male ( paratype , after Deeleman-Reinhold, 1989: 622-623 ): Total length 4.40. Carapace light grey-brown, ocular area slightly darker; chelicerae dark brown; labium, palpal coxae, sternum, and legs light grey-brown; abdomen pale with 3 snowy granulations on dorsal surface. Carapace 2.20 long, 1.70 wide; carapace length 1.29 times width; PER 1.14 times AER , PER 2.28 times OAL; OAL 0.57 times OQL; OQA 0.625 times OQP; ratio of eyes AM :AL: PM : PL , 1.0:2.0:3.0:3.0; AM-AM, AM-AL equal to AM diameter; PM-PM , PM-PL equal to PM diameter; AL touching PL . Sternum length 1.26 times width; labium length 0.75 times width; palpal coxae length 2.0 times width. Legs with typical spination; femur I length 1.25 times carapace width; metatarsus I as in Figs. 60, 61 , clasping spine extends beyond median spur, spur long, length nearly ½ segment width. Leg measurements: I: 2.3 + 0.9 + 2.2 + 1.9 + 1.2 = [8.5]; II: 2.1 + 0.7 + 1.8 + 1.8 + 1.0 = [7.4]; III : 2.1 + 0.8 + 1.5 + 1.7 + 0.9 = [7.0]; IV : 2.3 + 0.9 + 1.9 + 1.9 + 1.0 = [8.0]; palp: 1.2 + 0.4 + 0.6 + NA + 0.6 = [2.8]; leg formula 1423. Male palp as in Figs. 45–47 ; tibia length 1.21 times cymbial length, RTA length 0.23 times tibia ( Figs. 46, 47 ), MA origin 0.36 distance to tegular base, MA length 0.83 times that of tegulum, MA length 7.86 times width ( Figs. 45, 46 ). Variation (N=3): Total length 4.40 to 5.00. Female ( Paratype , after Deeleman-Reinhold, 1989: 622-623 ): Total length 6.70. Markings as in male. Carapace 3.30 long, 2.30 wide, carapace length 1.43 times width, carapace height 0.47 width; PER 1.11 times AER , PER 2.50 times OAL; OAL equals OQL; OQA 0.59 times OQP; clypeal height 1.25 times AM diameter; ratio of eyes AM :AL: PM : PL , 1.0:1.6:1.6:1.2; AM-AM 0.6 times AM diameter, AM-AL equals AM diameter; PM-PM 1.125 times PM diameter, PM-PL 1.25 times PM , AL touching PL . Chelicerae 1.30 long; cheliceral length 12.0 times clypeal height; sternum length 1.34 times width; labium length 1.09 times width; palpal coxae length 1.75 times width. Legs with typical spination. Femur I 1.26 times carapace width. Leg measurements: I: 2.9 + 1.2 + 2.5 + 2.4 + 1.3 = [10.30]; II: 2.7 + 1.1 + 2.2 + 2.0 + 1.0 = [9.00]; III : 2.4 + 1.0 + 1.7 + 1.7 + 1.0 = [7.80]; IV : 2.7 + 1.1 + 1.9 + 1.9 + 1.0 = [8.60]; palp: 1.1 + 0.25 + 0.70 + NA + 1.0 = [3.05], leg formula 1243. Female genitalia as in Fig. 48 (epigynum) and Fig. 49 (vulva). Epigynum width 2.66 times length; epigynum MLW 2.20 times MLL, MLW 1.50 times LLW. Variation (N=12): Total length 6.20 to 7.70. Natural History : Deeleman-Reinhold (1989: 619 , 620, 624) describes this species from Niah Cave, Sarawak , Malaysia , a cave famous for the diversity of endemic species. Distribution : Known only from the type locality ( Fig. 62 ). Material Examined : Only the paratypes from Niah Cave , Sarawak , in RMNH .