309 lines
35 KiB
XML
309 lines
35 KiB
XML
<document id="DC5B1CFA2874F8BED241EB45A5426516" ID-DOI="10.5281/zenodo.191756" ID-GBIF-Dataset="cde479d4-dc42-405a-b3ba-c6e89bfd9c2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="191756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461132017757" checkinUser="plazi" docAuthor="Hall, Sally & Thatje, Sven" docDate="2009" docId="B76B8794FFD3FFE4E7AEFB72FB211FDA" docLanguage="en" docName="zt02302p047.pdf" docOrigin="Zootaxa 2302" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lithodes galapagensis Hall & Thatje, 2009, n. sp." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="4B52FFECFFD8FFEAE739FFA0FFFD183C" masterDocTitle="Four new species of the family Lithodidae (Decapoda: Anomura) from the collections of the National Museum of Natural History, Smithsonian Institution" masterLastPageNumber="47" masterPageNumber="31" pageNumber="42" updateTime="1698597960008" updateUser="plazi">
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<mods:title id="A81F63E9014CBA8EB9E98B0901FA010C">Four new species of the family Lithodidae (Decapoda: Anomura) from the collections of the National Museum of Natural History, Smithsonian Institution</mods:title>
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<mods:name id="144E033A479387FB8D8FD60CE47E4669" type="personal">
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<mods:namePart id="C2CD0F41F615753D111D1D5EF4A24322">Hall, Sally</mods:namePart>
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<mods:namePart id="96CB8D27610A96BC955D279D7D57D6B5">Thatje, Sven</mods:namePart>
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<mods:typeOfResource id="648495C75C53FFAA5345459E884FC576">text</mods:typeOfResource>
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<mods:title id="59607043CC1932DB3085789E1F4C8B0E">Zootaxa</mods:title>
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<mods:part id="AAB3AB2EB9DC19929891219B3378785C">
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<mods:date id="4F9E6B7AEE39202485BF68001CE00DB1">2009</mods:date>
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<mods:number id="C4122CA33ACA488262DFBED3D1EC7D4F">2302</mods:number>
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<mods:identifier id="BCA3CAE79745DDD39783DA3BCC541FD5" type="DOI">10.5281/zenodo.191756</mods:identifier>
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<mods:identifier id="174E857082DF4380BAD7C8BAF7FA09DC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1FD0AF26FE357526BCA8AA3D6A8A05B6" type="Zenodo-Dep">191756</mods:identifier>
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<treatment id="B76B8794FFD3FFE4E7AEFB72FB211FDA" ID-DOI="http://doi.org/10.5281/zenodo.5662611" ID-GBIF-Taxon="119627154" ID-Zenodo-Dep="5662611" LSID="urn:lsid:plazi:treatment:B76B8794FFD3FFE4E7AEFB72FB211FDA" httpUri="http://treatment.plazi.org/id/B76B8794FFD3FFE4E7AEFB72FB211FDA" lastPageId="14" lastPageNumber="45" pageId="11" pageNumber="42">
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<subSubSection id="77D86509FFD3FFE1E7AEFB72FEED1D33" pageId="11" pageNumber="42" type="nomenclature">
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<paragraph id="3F7D3682FFD3FFE1E7AEFB72FE111CD0" blockId="11.[151,492,1234,1295]" box="[151,492,1234,1260]" pageId="11" pageNumber="42">
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<heading id="643581EEFFD3FFE1E7AEFB72FE111CD0" bold="true" box="[151,492,1234,1260]" fontSize="11" level="1" pageId="11" pageNumber="42" reason="1">
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<emphasis id="0DB6EA90FFD3FFE1E7AEFB72FE111CD0" bold="true" box="[151,492,1234,1260]" pageId="11" pageNumber="42">
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<taxonomicName id="F8C24D01FFD3FFE1E7AEFB72FE5F1CD0" ID-CoL="dd432aa4-9bb6-4801-aa2f-9bbc3143a771" box="[151,418,1234,1260]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="galapagensis" status="sp. nov.">
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<emphasis id="0DB6EA90FFD3FFE1E7AEFB72FE5F1CD0" bold="true" box="[151,418,1234,1260]" italics="true" pageId="11" pageNumber="42">Lithodes galapagensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="168557EBFFD3FFE1E693FB72FE111CD0" box="[426,492,1234,1260]" pageId="11" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3F7D3682FFD3FFE1E7AEFB55FEED1D33" blockId="11.[151,492,1234,1295]" box="[151,272,1269,1295]" pageId="11" pageNumber="42">
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(
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<figureCitation id="A7F92A07FFD3FFE1E7A6FB55FF161D33" box="[159,235,1269,1295]" captionStart="FIGURE 7" captionStartId="13.[151,255,1895,1919]" captionTargetBox="[200,1386,197,1870]" captionTargetId="figure@13.[200,1387,197,1871]" captionTargetPageId="13" captionText="FIGURE 7. Lithodes galapagensis n. sp. a – d, male holotype, 114 mm CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island, 00 ° 17 ’ 30 ” S, 091 ° 39 ’ 36 ” W, 648 m. (a) carapace, lateral. (b) dorsal carapace spine, lateral. (c) left walking leg 3, posterior. (d) whole organism, dorsal (note damage to carapace on the left branchial region). Scale bar = 10 mm for a, c, d; 1 mm for b." httpUri="https://zenodo.org/record/191763/files/figure.png" pageId="11" pageNumber="42">Figs 7</figureCitation>
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, 8)
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</paragraph>
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</subSubSection>
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<subSubSection id="77D86509FFD3FFE1E7AEFAE2FA731D96" pageId="11" pageNumber="42" type="materials_examined">
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<paragraph id="3F7D3682FFD3FFE1E7AEFAE2FA731D96" blockId="11.[151,1437,1346,2030]" pageId="11" pageNumber="42">
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<emphasis id="0DB6EA90FFD3FFE1E7AEFAE2FE641D60" bold="true" box="[151,409,1346,1372]" pageId="11" pageNumber="42">Material examined.</emphasis>
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Galapagos Archipelago: Johnson Sea-Link II Cruise station 3101, Cabo Douglas, Fernandina Island,
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<geoCoordinate id="5AF65045FFD3FFE1E6B0FAC9FDE31DBF" box="[393,542,1385,1411]" direction="south" orientation="latitude" pageId="11" pageNumber="42" precision="15" value="-0.29166666">00°17’30”S</geoCoordinate>
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,
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<geoCoordinate id="5AF65045FFD3FFE1E515FAC9FD261DBF" box="[556,731,1385,1411]" direction="west" orientation="longitude" pageId="11" pageNumber="42" precision="15" value="-91.66">091°39’36”W</geoCoordinate>
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,
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<date id="4B7C1042FFD3FFE1E5D0FAC9FC851DBF" box="[745,888,1385,1411]" pageId="11" pageNumber="42" value="1998-07-17">
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17.07.
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<quantity id="F83A9B67FFD3FFE1E40EFAC9FC851DBF" box="[823,888,1385,1411]" metricMagnitude="3" metricUnit="m" metricValue="1.998648" pageId="11" pageNumber="42" unit="m" value="1998.648">1998</quantity>
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</date>
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, 648m: male
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<typeStatus id="E0798820FFD3FFE1E31AFAC9FB691DBF" box="[1059,1172,1385,1411]" pageId="11" pageNumber="42" type="holotype">holotype</typeStatus>
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, CL
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<quantity id="F83A9B67FFD3FFE1E3E8FAC9FAC61DBF" box="[1233,1339,1385,1411]" metricMagnitude="-1" metricUnit="m" metricValue="1.14" pageId="11" pageNumber="42" unit="mm" value="114.0">114 mm</quantity>
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; female
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<typeStatus id="E0798820FFD3FFE1E7AEFA30FF021D96" box="[151,255,1424,1450]" pageId="11" pageNumber="42" type="paratype">paratype</typeStatus>
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, CL
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<quantity id="F83A9B67FFD3FFE1E60EFA30FE701D96" box="[311,397,1424,1450]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="42" unit="mm" value="84.0">84 mm</quantity>
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, Seymour Island,
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<geoCoordinate id="5AF65045FFD3FFE1E566FA30FD111D96" box="[607,748,1424,1450]" direction="south" orientation="latitude" pageId="11" pageNumber="42" precision="15" value="-0.36166665">00°21’42”S</geoCoordinate>
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,
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<geoCoordinate id="5AF65045FFD3FFE1E5C0FA30FC601D96" box="[761,925,1424,1450]" direction="west" orientation="longitude" pageId="11" pageNumber="42" precision="15" value="-90.25">090°15’00”W</geoCoordinate>
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,
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<date id="4B7C1042FFD3FFE1E490FA30FBCD1D96" box="[937,1072,1424,1450]" pageId="11" pageNumber="42" value="1998-07-25">
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25.07.
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<quantity id="F83A9B67FFD3FFE1E4CDFA30FBCD1D96" box="[1012,1072,1424,1450]" metricMagnitude="3" metricUnit="m" metricValue="1.99874" pageId="11" pageNumber="42" unit="m" value="1998.74">1998</quantity>
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</date>
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, 740 m (all
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<collectionCode id="59D3AE47FFD3FFE1E382FA30FAED1D96" LSID="urn:lsid:biocol.org:col:34871" box="[1211,1296,1424,1450]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="42">USNM</collectionCode>
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1122586).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="77D86509FFD3FFE1E7FFFA16FB941DEC" box="[198,1129,1462,1488]" pageId="11" pageNumber="42" type="etymology">
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<paragraph id="3F7D3682FFD3FFE1E7FFFA16FB941DEC" blockId="11.[151,1437,1346,2030]" box="[198,1129,1462,1488]" pageId="11" pageNumber="42">
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<emphasis id="0DB6EA90FFD3FFE1E7FFFA16FEB61DEC" bold="true" box="[198,331,1462,1488]" pageId="11" pageNumber="42">Etymology</emphasis>
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||
. This species is named after its
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<typeStatus id="E0798820FFD3FFE1E5FBFA16FD081DEC" box="[706,757,1462,1488]" pageId="11" pageNumber="42">type</typeStatus>
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locality, the Galapagos Islands.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="77D86509FFD3FFE6E7FFFA7DFDAD1CFA" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42" type="description">
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||
<paragraph id="3F7D3682FFD3FFE1E7FFFA7DFE261F9C" blockId="11.[151,1437,1346,2030]" pageId="11" pageNumber="42">
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||
<emphasis id="0DB6EA90FFD3FFE1E7FFFA7DFDDB1DCB" bold="true" box="[198,550,1501,1527]" pageId="11" pageNumber="42">
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Description of the
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<typeStatus id="E0798820FFD3FFE1E682FA7DFDDB1DCB" box="[443,550,1501,1527]" pageId="11" pageNumber="42" type="holotype">holotype</typeStatus>
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</emphasis>
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||
. Carapace roughly pyriform in outline (
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<figureCitation id="A7F92A07FFD3FFE1E32EFA7DFB9E1DCB" box="[1047,1123,1501,1527]" captionStart="FIGURE 7" captionStartId="13.[151,255,1895,1919]" captionTargetBox="[200,1386,197,1870]" captionTargetId="figure@13.[200,1387,197,1871]" captionTargetPageId="13" captionText="FIGURE 7. Lithodes galapagensis n. sp. a – d, male holotype, 114 mm CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island, 00 ° 17 ’ 30 ” S, 091 ° 39 ’ 36 ” W, 648 m. (a) carapace, lateral. (b) dorsal carapace spine, lateral. (c) left walking leg 3, posterior. (d) whole organism, dorsal (note damage to carapace on the left branchial region). Scale bar = 10 mm for a, c, d; 1 mm for b." httpUri="https://zenodo.org/record/191763/files/figure.png" pageId="11" pageNumber="42">Fig. 7</figureCitation>
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d); as wide as long when measured at maximal width of carapace. Dorsal regions well defined; covered uniformly with small spinules more or less acute at apex, without setae (
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<figureCitation id="A7F92A07FFD3FFE1E5A3F98AFD191E78" box="[666,740,1578,1604]" captionStart="FIGURE 7" captionStartId="13.[151,255,1895,1919]" captionTargetBox="[200,1386,197,1870]" captionTargetId="figure@13.[200,1387,197,1871]" captionTargetPageId="13" captionText="FIGURE 7. Lithodes galapagensis n. sp. a – d, male holotype, 114 mm CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island, 00 ° 17 ’ 30 ” S, 091 ° 39 ’ 36 ” W, 648 m. (a) carapace, lateral. (b) dorsal carapace spine, lateral. (c) left walking leg 3, posterior. (d) whole organism, dorsal (note damage to carapace on the left branchial region). Scale bar = 10 mm for a, c, d; 1 mm for b." httpUri="https://zenodo.org/record/191763/files/figure.png" pageId="11" pageNumber="42">Fig. 7</figureCitation>
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b). Gastric region convex and slightly more inflated than branchial and cardiac regions. One pair of slender spines
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<quantity id="F83A9B67FFD3FFE1E401F9F1FC811E57" box="[824,892,1617,1643]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="42" unit="mm" value="7.0">7 mm</quantity>
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in length, emanating from the mid part of this region — level with hepatic spines on lateral margin. Spinules sparse on apex of gastric region and very few around base of prominent spines. Cardiac region depressed and separated from gastric region by smooth, wide, and saddle-shaped groove. Cardiac region depressed anteriorly, and more inflated posteriorly around single pair of long, slender spines in this region. A pair of acute spinules directly anterior to this pair. Triangular cardiac region separated from branchial regions by grooves which converge posteriorly, and then diverge close to the margin to describe posterior of branchial regions. Branchial regions each with single long, slender spine at apex; a few large, acute spinules posteriorly. A pair of conical spines in intestinal region almost on posterior margin.
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</paragraph>
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<paragraph id="3F7D3682FFD3FFE6E7FFF80DFD5918E4" blockId="11.[151,1437,1346,2030]" lastBlockId="12.[151,1437,152,1338]" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42">
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Exterior orbital spine just surpassing length of eyestalks; anterolateral spine about equal in length or slightly smaller. Hepatic spines slightly inflated at base, with long slender spine reaching
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<quantity id="F83A9B67FFD3FFE1E38CF874FAF11FD2" box="[1205,1292,2004,2030]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="42" unit="mm" value="20.0">20 mm</quantity>
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. Two spines on anterior portion of each branchial lateral margin, and several much smaller, conical spines interspersed between them and on posterolateral margins.
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</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFF45FB8D194F" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">Rostrum with long, straight median projection rising dorsally from surface of carapace and terminating in pair of spines. Mid way along length of median projection emanate a pair of dorsal lateral spines of about the same length as terminal bifurcation. Base of rostrum narrow, without granulation. Ventrally, with long, smooth spine curving gently upward, terminating approximately at level of corneae.</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFE20FEF819FC" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">Eyestalks prominent and without granulation, but with crenulation of dorsal edge of the corneal margin (Fig. 8b).</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFE6DFE0E1A32" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">Second segment of antennal peduncle with long slender spine on exterior aspect. Antennal acicle reduced to very small conical process.</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFDBAFBAC1A94" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">Cheliped merus and carpus with several strong spines on terminal border and poorly defined rows of short spines on dorsal, interior and exterior surfaces. Palm with several poorly defined rows of short spines on dorsal border and two rows on exterior surface leading to articulation with movable finger. Fingers 0.4 × total length of right hand, and 0.5 of left hand. Fingers bearing few tufts of short setae.</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFD15FBB81B8B" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">
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Walking legs long and slender. Merus of pereiopod 3 about 0.8 × length of carapace, and 0.2 × as high as long. Covered densely with spinules on dorsal/posterior surface, and smooth on ventral surface (
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<figureCitation id="A7F92A07FFD4FFE6E3C7FD7CFAB91ACA" box="[1278,1348,732,758]" captionStart="FIGURE 7" captionStartId="13.[151,255,1895,1919]" captionTargetBox="[200,1386,197,1870]" captionTargetId="figure@13.[200,1387,197,1871]" captionTargetPageId="13" captionText="FIGURE 7. Lithodes galapagensis n. sp. a – d, male holotype, 114 mm CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island, 00 ° 17 ’ 30 ” S, 091 ° 39 ’ 36 ” W, 648 m. (a) carapace, lateral. (b) dorsal carapace spine, lateral. (c) left walking leg 3, posterior. (d) whole organism, dorsal (note damage to carapace on the left branchial region). Scale bar = 10 mm for a, c, d; 1 mm for b." httpUri="https://zenodo.org/record/191763/files/figure.png" pageId="12" pageNumber="43">Fig. 7</figureCitation>
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c). Two or three rows of larger conical spines along length of merus and long spine on terminal border. Carpus of walking legs with spinulation on dorsal surfaces and smooth ventral surfaces, as well as two very long spines at proximal and distal ends of dorsal border. Propodus sparsely covered with irregular rows of spinules. Dactyli of walking legs over half length of merus and equal to length of carpus; very sharp, slender spines on dorsal border to tip; few or no spines on ventral border; no tufts of setae present.
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</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FCFC64FD051C6E" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">
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Abdomen of a form typical of
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<taxonomicName id="F8C24D01FFD4FFE6E502FC64FD5E1BE1" box="[571,675,964,989]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="genus">
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||
<emphasis id="0DB6EA90FFD4FFE6E502FC64FD5E1BE1" box="[571,675,964,989]" italics="true" pageId="12" pageNumber="43">Lithodes</emphasis>
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</taxonomicName>
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, with nodules in the medial portion of segments 3–5; separate marginal and lateral plates well-calcified (Fig. 8f). Second abdominal segment with medial and lateral plates fused; marginal plates almost joined or with suture visible. Surface of abdominal plates with several warty tubercles on edges but no spines decorating surface.
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</paragraph>
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<paragraph id="3F7D3682FFD4FFE6E7FFFBFEFDAD1CFA" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">
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<emphasis id="0DB6EA90FFD4FFE6E7FFFBFEFEB31C44" bold="true" box="[198,334,1118,1144]" pageId="12" pageNumber="43">Variations.</emphasis>
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The female
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<typeStatus id="E0798820FFD4FFE6E6C9FBFEFDA81C44" box="[496,597,1118,1144]" pageId="12" pageNumber="43" type="paratype">paratype</typeStatus>
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of this species is slightly smaller than the male. It differs from the
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<typeStatus id="E0798820FFD4FFE6E7AEFB25FF031CA3" box="[151,254,1157,1183]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
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||
in having less acute spinules on the dorsal surface, and the less prominent spines on the proximal and distal angles of the walking-leg carpi.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="77D86509FFD4FFE4E7FFFB72FB211FDA" lastPageId="14" lastPageNumber="45" pageId="12" pageNumber="43" type="discussion">
|
||
<paragraph id="3F7D3682FFD4FFE6E7FFFB72FC971D06" blockId="12.[151,1437,152,1338]" pageId="12" pageNumber="43">
|
||
<emphasis id="0DB6EA90FFD4FFE6E7FFFB72FEC21CD0" bold="true" box="[198,319,1234,1260]" pageId="12" pageNumber="43">Remarks.</emphasis>
|
||
This species is distinguished from all other members of this genus, except
|
||
<taxonomicName id="F8C24D01FFD4FFE6E38DFB73FF291D2F" authority="Haig" authorityName="Haig" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E38DFB73FA611CD0" box="[1204,1436,1235,1260]" italics="true" pageId="12" pageNumber="43">Lithodes wiracocha</emphasis>
|
||
Haig
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="47D57612FFD4FFE6E61AFB59FEA01D2F" box="[291,349,1273,1299]" name="Peru" pageId="12" pageNumber="43">Peru</collectingCountry>
|
||
, in that it has carapace and dorsal surface of the walking legs densely covered in spinules. This species differs from
|
||
<taxonomicName id="F8C24D01FFD4FFE6E6FBFA80FDA01D05" box="[450,605,1312,1337]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E6FBFA80FDA01D05" box="[450,605,1312,1337]" italics="true" pageId="12" pageNumber="43">L. wiracocha</emphasis>
|
||
</taxonomicName>
|
||
in the following ways:
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DCFACDFB1B1DBB" blockId="12.[151,1436,1389,2034]" box="[229,1254,1389,1415]" pageId="12" pageNumber="43">
|
||
The spines on the dorsal surface are less dense in
|
||
<taxonomicName id="F8C24D01FFD4FFE6E415FACDFC1B1DBA" box="[812,998,1389,1414]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E415FACDFC1B1DBA" box="[812,998,1389,1414]" italics="true" pageId="12" pageNumber="43">L. galapagensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="F8C24D01FFD4FFE6E37CFACDFB1D1DBA" box="[1093,1248,1389,1414]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E37CFACDFB1D1DBA" box="[1093,1248,1389,1414]" italics="true" pageId="12" pageNumber="43">L. wiracocha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DCFA34FA661D92" blockId="12.[151,1436,1389,2034]" box="[229,1435,1428,1454]" pageId="12" pageNumber="43">
|
||
The walking legs of
|
||
<taxonomicName id="F8C24D01FFD4FFE6E6EFFA34FD721D91" box="[470,655,1428,1453]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E6EFFA34FD721D91" box="[470,655,1428,1453]" italics="true" pageId="12" pageNumber="43">L. galapagensis</emphasis>
|
||
</taxonomicName>
|
||
have spinules only on the dorsal and not on the ventral surfaces of
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEFA1AFA661DE8" blockId="12.[151,1436,1389,2034]" box="[231,1435,1466,1492]" pageId="12" pageNumber="43">
|
||
the walking-leg segments, whereas
|
||
<taxonomicName id="F8C24D01FFD4FFE6E5B6FA1BFCD61DE8" box="[655,811,1467,1492]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E5B6FA1BFCD61DE8" box="[655,811,1467,1492]" italics="true" pageId="12" pageNumber="43">L. wiracocha</emphasis>
|
||
</taxonomicName>
|
||
has densely packed spinules covering the surface of
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEFA41FE7D1DC7" blockId="12.[151,1436,1389,2034]" box="[231,384,1505,1531]" pageId="12" pageNumber="43">all segments.</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DFF9A8FA661E1E" blockId="12.[151,1436,1389,2034]" box="[230,1435,1544,1570]" pageId="12" pageNumber="43">
|
||
In the
|
||
<typeStatus id="E0798820FFD4FFE6E60BF9A8FE661E1E" box="[306,411,1544,1570]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
, and somewhat on the
|
||
<typeStatus id="E0798820FFD4FFE6E590F9A8FCF31E1E" box="[681,782,1544,1570]" pageId="12" pageNumber="43" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="F8C24D01FFD4FFE6E40FF9A8FC121E1D" box="[822,1007,1544,1569]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E40FF9A8FC121E1D" box="[822,1007,1544,1569]" italics="true" pageId="12" pageNumber="43">L. galapagensis</emphasis>
|
||
</taxonomicName>
|
||
, certain spines on the dorsal surface
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF98EFA9F1E74" blockId="12.[151,1436,1389,2034]" box="[231,1378,1582,1608]" pageId="12" pageNumber="43">
|
||
and on the walking legs are very long and slender, unlike the stout conical spines in
|
||
<taxonomicName id="F8C24D01FFD4FFE6E3F8F98FFAA61E74" box="[1217,1371,1583,1608]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E3F8F98FFAA61E74" box="[1217,1371,1583,1608]" italics="true" pageId="12" pageNumber="43">L. wiracocha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7D0F9F5FA611E52" blockId="12.[151,1436,1389,2034]" box="[233,1436,1621,1647]" pageId="12" pageNumber="43">
|
||
The rostrum of
|
||
<taxonomicName id="F8C24D01FFD4FFE6E68AF9F5FD831E52" box="[435,638,1621,1646]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E68AF9F5FD831E52" box="[435,638,1621,1646]" italics="true" pageId="12" pageNumber="43">L. galapagensis</emphasis>
|
||
</taxonomicName>
|
||
is very long and slender — similar in this respect to
|
||
<taxonomicName id="F8C24D01FFD4FFE6E216F9F5FA611E52" box="[1327,1436,1621,1646]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0DB6EA90FFD4FFE6E216F9F5FA611E52" box="[1327,1436,1621,1646]" italics="true" pageId="12" pageNumber="43">Lithodes</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF9DCFA611EAA" blockId="12.[151,1436,1389,2034]" box="[231,1436,1660,1686]" pageId="12" pageNumber="43">
|
||
<emphasis id="0DB6EA90FFD4FFE6E7DEF9DCFE861EA9" box="[231,379,1660,1685]" italics="true" pageId="12" pageNumber="43">megacantha</emphasis>
|
||
Macpherson, 1991 from the central Pacific; however, the rostrum of
|
||
<taxonomicName id="F8C24D01FFD4FFE6E3F6F9DCFA821EA9" box="[1231,1407,1660,1685]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiraconcha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E3F6F9DCFA821EA9" box="[1231,1407,1660,1685]" italics="true" pageId="12" pageNumber="43">L. wiraconcha</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF902FDDF1E80" blockId="12.[151,1436,1389,2034]" box="[231,546,1698,1724]" pageId="12" pageNumber="43">rather stout in comparison.</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DFF969FA611EDF" blockId="12.[151,1436,1389,2034]" box="[230,1436,1737,1763]" pageId="12" pageNumber="43">
|
||
There are no acute spinules on the dorsal base of the rostrum in
|
||
<emphasis id="0DB6EA90FFD4FFE6E4E0F969FB671EDE" box="[985,1178,1737,1762]" italics="true" pageId="12" pageNumber="43">
|
||
<taxonomicName id="F8C24D01FFD4FFE6E4E0F969FB6B1EDE" box="[985,1174,1737,1762]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">L. galapagensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
whereas spines cover
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF950FC551F36" blockId="12.[151,1436,1389,2034]" box="[231,936,1776,1802]" pageId="12" pageNumber="43">
|
||
the proximal part of all of the lateral spines in
|
||
<taxonomicName id="F8C24D01FFD4FFE6E43FF950FC5C1F35" box="[774,929,1776,1801]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="wiracocha">
|
||
<emphasis id="0DB6EA90FFD4FFE6E43FF950FC5C1F35" box="[774,929,1776,1801]" italics="true" pageId="12" pageNumber="43">L. wiracocha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF8B6FA611F0C" blockId="12.[151,1436,1389,2034]" box="[231,1436,1814,1840]" pageId="12" pageNumber="43">
|
||
No spines on the surface of the abdominal plates in
|
||
<taxonomicName id="F8C24D01FFD4FFE6E46FF8B7FBEF1F0C" box="[854,1042,1815,1840]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E46FF8B7FBEF1F0C" box="[854,1042,1815,1840]" italics="true" pageId="12" pageNumber="43">L. galapagensis</emphasis>
|
||
</taxonomicName>
|
||
. Instead, plates are covered with
|
||
</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7DEF89DFE711F6B" blockId="12.[151,1436,1389,2034]" box="[231,396,1853,1879]" pageId="12" pageNumber="43">low tubercles.</paragraph>
|
||
<paragraph id="3F7D3682FFD4FFE6E7FCF8C4FC281FCE" blockId="12.[151,1436,1389,2034]" pageId="12" pageNumber="43">
|
||
Other species close to this location include
|
||
<taxonomicName id="F8C24D01FFD4FFE6E5DDF8C4FBC11F42" authority="Faxon" authorityName="Faxon" box="[740,1084,1892,1918]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E5DDF8C4FC1A1F41" box="[740,999,1892,1917]" italics="true" pageId="12" pageNumber="43">Lithodes panamensis</emphasis>
|
||
Faxon
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="47D57612FFD4FFE6E3B0F8C4FB171F42" box="[1161,1258,1892,1918]" name="Panama" pageId="12" pageNumber="43">Panama</collectingCountry>
|
||
and
|
||
<collectingCountry id="47D57612FFD4FFE6E21EF8C4FA9E1F42" box="[1319,1379,1892,1918]" name="Peru" pageId="12" pageNumber="43">Peru</collectingCountry>
|
||
, and
|
||
<taxonomicName id="F8C24D01FFD4FFE6E7AEF82BFD8D1F98" authority="Macpherson (1988b)" authorityName="Macpherson" authorityYear="1988" box="[151,624,1930,1956]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="santolla">
|
||
<emphasis id="0DB6EA90FFD4FFE6E7AEF82BFE951F98" box="[151,360,1931,1956]" italics="true" pageId="12" pageNumber="43">Lithodes santolla</emphasis>
|
||
<bibRefCitation id="5B534B73FFD4FFE6E648F82AFDF81F98" author="Macpherson" box="[369,517,1930,1956]" pageId="12" pageNumber="46" refString="Macpherson, E. (1988 b) Revision of the Family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson</bibRefCitation>
|
||
(1988b)
|
||
</taxonomicName>
|
||
, from Patagonia, both of these are readily distinguishable from the present species because of the peculiar spination of
|
||
<taxonomicName id="F8C24D01FFD4FFE6E5CCF811FC011FF6" box="[757,1020,1969,1994]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="0DB6EA90FFD4FFE6E5CCF811FC011FF6" box="[757,1020,1969,1994]" italics="true" pageId="12" pageNumber="43">Lithodes galapagensis</emphasis>
|
||
</taxonomicName>
|
||
in which a dense coverage of acute spinules are combined with the few very long spines and long rostrum.
|
||
</paragraph>
|
||
<caption id="6BBD660AFFD5FFE7E7AEF8C7FD4E1FDA" httpUri="https://zenodo.org/record/191763/files/figure.png" pageId="13" pageNumber="44" targetBox="[200,1386,197,1870]" targetPageId="13">
|
||
<paragraph id="3F7D3682FFD5FFE7E7AEF8C7FD4E1FDA" blockId="13.[151,1436,1895,2022]" pageId="13" pageNumber="44">
|
||
<emphasis id="0DB6EA90FFD5FFE7E7AEF8C7FEE71F43" bold="true" box="[151,282,1895,1919]" pageId="13" pageNumber="44">FIGURE 7.</emphasis>
|
||
<taxonomicName id="F8C24D01FFD5FFE7E61BF8C7FDE91F42" box="[290,532,1895,1918]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="galapagensis" status="sp. nov.">
|
||
<emphasis id="0DB6EA90FFD5FFE7E61BF8C7FDE91F42" box="[290,532,1895,1918]" italics="true" pageId="13" pageNumber="44">Lithodes galapagensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="168557EBFFD5FFE7E522F8C7FDA51F43" box="[539,600,1895,1919]" pageId="13" pageNumber="44" rank="species">
|
||
<emphasis id="0DB6EA90FFD5FFE7E522F8C7FDA51F43" bold="true" box="[539,600,1895,1919]" pageId="13" pageNumber="44">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
a–d, male holotype, 114 mm CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island, 00°17’30”S, 091°39’36”W, 648 m. (a) carapace, lateral. (b) dorsal carapace spine, lateral. (c) left walking leg 3, posterior. (d) whole organism, dorsal (note damage to carapace on the left branchial region). Scale bar = 10 mm for a, c, d; 1 mm for b.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F7D3682FFD6FFE4E7AEF8C4FB211FDA" blockId="14.[151,1437,1892,2022]" pageId="14" pageNumber="45">
|
||
<emphasis id="0DB6EA90FFD6FFE4E7AEF8C4FEE61F40" bold="true" box="[151,283,1892,1916]" pageId="14" pageNumber="45">FIGURE 8.</emphasis>
|
||
<taxonomicName id="F8C24D01FFD6FFE4E61BF8C5FDE91F40" box="[290,532,1893,1916]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="galapagensis" status="sp. nov.">
|
||
<emphasis id="0DB6EA90FFD6FFE4E61BF8C5FDE91F40" box="[290,532,1893,1916]" italics="true" pageId="14" pageNumber="45">Lithodes galapagensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="168557EBFFD6FFE4E522F8C4FDA51F40" box="[539,600,1892,1916]" pageId="14" pageNumber="45" rank="species">
|
||
<emphasis id="0DB6EA90FFD6FFE4E522F8C4FDA51F40" bold="true" box="[539,600,1892,1916]" pageId="14" pageNumber="45">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
a–h, male
|
||
<typeStatus id="E0798820FFD6FFE4E5E9F8C4FCCC1F47" box="[720,817,1892,1915]" pageId="14" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
,
|
||
<quantity id="F83A9B67FFD6FFE4E405F8C4FC681F47" box="[828,917,1892,1915]" metricMagnitude="-1" metricUnit="m" metricValue="1.14" pageId="14" pageNumber="45" unit="mm" value="114.0">114 mm</quantity>
|
||
CL (USNM 1122586), Galapagos Archipelago, Cabo Douglas, Fernandina Island,
|
||
<geoCoordinate id="5AF65045FFD6FFE4E532F827FD711FA2" box="[523,652,1927,1950]" direction="south" orientation="latitude" pageId="14" pageNumber="45" precision="15" value="-0.29166666">00°17’30”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5AF65045FFD6FFE4E5AEF827FCD01FA2" box="[663,813,1927,1950]" direction="west" orientation="longitude" pageId="14" pageNumber="45" precision="15" value="-91.66">091°39’36”W</geoCoordinate>
|
||
,
|
||
<specimenCount id="29C4FD0BFFD6FFE4E401F827FC7E1FA2" box="[824,899,1927,1950]" pageId="14" pageNumber="45" type="generic">
|
||
<quantity id="F83A9B67FFD6FFE4E401F827FC821FA2" box="[824,895,1927,1950]" metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="14" pageNumber="45" unit="m" value="648.0">648 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
(a) left chela, lateral. (b) ocular peduncle, frontal. (c) right chela, lateral. (d) anterior carapace, lateral. (e) second abdominal segment, posterior. (f) abdominal plates. (g) dactylus 3rd walking leg, posterior. (h) rostral spines, dorsal. Scale bar =
|
||
<quantity id="F83A9B67FFD6FFE4E4A0F86FFC191FDA" box="[921,996,1999,2022]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="45" unit="mm" value="10.0">10 mm</quantity>
|
||
for a, c–h;
|
||
<quantity id="F83A9B67FFD6FFE4E367F86FFB661FDA" box="[1118,1179,1999,2022]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="45" unit="mm" value="1.0">1 mm</quantity>
|
||
for b.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |