751 lines
77 KiB
XML
751 lines
77 KiB
XML
<document id="B724A725FF67F25286C3C1142D2D72BA" ID-DOI="10.3897/zookeys.1218.120744" ID-ZooBank="094646BE-BF9F-4890-BD59-FF26FA95A3FE" ID-publisher-id="120744" URI-arpha="6E4DF739-56EF-5B8D-AEB7-2839D486DC50" URI-zoobank="http://zoobank.org/094646BE-BF9F-4890-BD59-FF26FA95A3FE" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946914" checkinUser="pensoft" docAuthor="Murányi, Dávid & Kovács, Tibor" docDate="2024" docId="83CD7F7B778158F0B107C65E2DED1906" docLanguage="en" docName="ZooKeys 1218: 49-79" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/120744/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Leuctra visitor Murányi & Kovács, 2024, sp. nov." docType="treatment" docUuid="6E2C2B5D-5228-4A6A-B501-C2FB65A162EC" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="79" masterDocId="6E4DF73956EF5B8DAEB72839D486DC50" masterDocTitle="Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)" masterLastPageNumber="79" masterPageNumber="49" pageNumber="49" updateTime="1731697136958" updateUser="admin">
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<mods:title id="646DEB54F6DE551B9BCDC0D87C748374">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:name id="059FB83678B36C178BD3994CD9ADD469" type="personal">
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<mods:roleTerm id="0F7A3727B2BE3860AD79D6E11C9E9F60">Author</mods:roleTerm>
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<mods:namePart id="DC0259D5DFB293D0A2BE10CA9CEFA85E">Murányi, Dávid</mods:namePart>
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<mods:affiliation id="C001C5007DD3CFDD4C0FF513112E41DC">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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<mods:name id="723CD24E8CD22B90F4B5769303B7F030" type="personal">
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<mods:namePart id="06C480F14C987248374907FBB21EB743">Kovács, Tibor</mods:namePart>
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<mods:nameIdentifier id="057D42880ABC1AE42B0DA1799AE591B4" type="ORCID">0000-0003-4577-0394</mods:nameIdentifier>
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<mods:affiliation id="7C1D5E239684AD4B091DFDDCCBCA0782">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<mods:typeOfResource id="613DA0C24F10294BB638E3B7722A1F35">text</mods:typeOfResource>
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<mods:title id="D0D8EC6900A5A98CAFBE4287DB07B682">ZooKeys</mods:title>
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<mods:part id="113C8F2CBED10395A79E21214E68472D">
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<mods:date id="CCB853DA6C78A0929ABEE6889CD72278">2024</mods:date>
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<mods:detail id="628280B994A3359B632296934C4B21BC" type="pubDate">
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<mods:number id="06B4274CB22F21D336727A089B037E14">2024-11-15</mods:number>
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<mods:number id="3C6FEA9A8FF05B5B426DAE2D2E214D49">1218</mods:number>
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<mods:start id="829D529FB6AD24050177A5715284E04C">49</mods:start>
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<mods:identifier id="B1E97533C747E3DFC8B819397D41A36D" type="DOI">10.3897/zookeys.1218.120744</mods:identifier>
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<mods:identifier id="2007B0F9A0566A72DF5B13CC602A9D03" type="ZooBank">094646BE-BF9F-4890-BD59-FF26FA95A3FE</mods:identifier>
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<treatment id="83CD7F7B778158F0B107C65E2DED1906" ID-arpha="83CD7F7B-7781-58F0-B107-C65E2DED1906" ID-zoobank="https://zoobank.org/6E2C2B5D-5228-4A6A-B501-C2FB65A162EC" LSID="urn:lsid:zoobank.org:act:6E2C2B5D-5228-4A6A-B501-C2FB65A162EC" LSID-ZBK="urn:lsid:zoobank.org:act:6E2C2B5D-5228-4A6A-B501-C2FB65A162EC" httpUri="http://treatment.plazi.org/id/83CD7F7B778158F0B107C65E2DED1906">
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<subSubSection id="DEDE5E51A593F20F62B3CDC39D72BCE1" type="nomenclature">
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<paragraph id="8FAD5DAB665EAA36D7074E9392CB1346">
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<taxonomicName id="5A1BE46EC1319A9A1492A508785D951D" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
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<emphasis id="C54CF1AB0092CE7B2A439EF6C047865C" italics="true">Leuctra visitor</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="8D6FE0D215FABB7A6DDDB1B1D915887E" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7A9C0F0BF7694FEBC7F6CDCBEE4C4720" type="description">
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<paragraph id="6E68330531BCEA3DBCE177D18B0F1107">
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<figureCitation id="26356180B3068B4FAC3A45F4138F5312" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">Figs 1</figureCitation>
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,
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||
<figureCitation id="800011FA3073B52D079A4BD983B467F3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10</figureCitation>
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||
,
|
||
<figureCitation id="483FBE3EF102F294EA7CCBB5066AD4A4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11</figureCitation>
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,
|
||
<figureCitation id="6999D7581D1540DE76B8FFE9B1B3CD49" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 B, F</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="SECID0EBRBG" type="type material">
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||
<paragraph id="B0D875763183135E44DFB495D85D9C77">
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||
<heading id="3F6E7AFE99FAA70AD73D0E4FE2DF96C9" reason="title">Type material.</heading>
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||
</paragraph>
|
||
<paragraph id="457A98E39FBC1540EE506A1D1FFBB294">
|
||
<materialsCitation id="0054425AB4586AFB267E8E9434BEAF46" collectingDate="2023-05-05" collectionCode="MM" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCode="2023-49, PLETYP-37" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="19A12D0F17FDED284C3412F2D1096AC7" bold="true" italics="true">
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||
<typeStatus id="038D498027E3EF93D53E0BAF1DB94AD6" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="DF875CD840A4AF186767B2AFA48E9AFB" type="male">male</specimenCount>
|
||
:
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||
<collectingCountry id="DDAD585E5F4F9AC5C4A1840B376F1552" name="Serbia">Montenegro</collectingCountry>
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||
: •
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||
<collectingMunicipality id="E35A71C15BFA9B3B2751CEE29F8BF967">Plav municipality</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="C8816EDE5D906D725894089D3BD19536">Visitor Mts</collectingMunicipality>
|
||
,
|
||
<location id="F3D3BF253E0ADA4EA1121C59B3E3152F" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:F3D3BF253E0ADA4EA1121C59B3E3152F" country="Montenegro" county="Katun" latitude="42.6308" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" name="forest springs and their outlet along the Katun road">forest springs and their outlet along the Katun road</location>
|
||
,
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||
<quantity id="D4E57FBF035814E0796ACC5275708FE6" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">
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||
<elevation id="3166346C3364884F6B449000A6356E45" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">1730 m</elevation>
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||
</quantity>
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||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPRBG" specific-use="{"type":"Point","coordinates":[19.835800,42.630800]}">
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<geoCoordinate id="E26BDA1A4B6AD818036AE5ED732428A7" degrees="42.6308" direction="north" orientation="latitude" precision="5" value="42.6308">42.6308°N</geoCoordinate>
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,
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||
<geoCoordinate id="9A3AC4BE59A3AC86905E8232A4B3E566" degrees="19.8358" direction="east" orientation="longitude" precision="5" value="19.8358">19.8358°E</geoCoordinate>
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||
</named-content>
|
||
,
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||
<date id="840BB767EF82AD1BCE18E3F28364F090" value="2023-05-05">
|
||
<collectingDate id="9391A0A646EE035F12EDFE6F64B6BA89" value="2023-05-05">5.v.2023</collectingDate>
|
||
</date>
|
||
(field number: loc. 13), leg.
|
||
<collectorName id="FDF421C846F2010525ED42F4A8B74202">T. Kovács</collectorName>
|
||
,
|
||
<collectorName id="309DEC2BDDADE94DC7A24DC8D099E5C5">D. Murányi</collectorName>
|
||
(
|
||
<named-content id="4D8763B3BEB8C2FF25A0C1068168BC8B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
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||
<collectionCode id="7DF58FDB202024CBD9E4DDA754E01538" country="France" name="University of Montpellier">MM</collectionCode>
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</named-content>
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||
:
|
||
<specimenCode id="0B899EAAAC44573A148F3670C047E1E2">2023-49</specimenCode>
|
||
,
|
||
<specimenCode id="9B747CB31C2FF1EC2044D0E69DDAF9A2">PLETYP-37</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="0F0F0E193286D76A919D9C2EB657C9C3" collectingDate="2023-05-05" collectionCode="BYU" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="DEBC873F79BFB2A0F1FEC516A8BEE17D" bold="true" italics="true">
|
||
<typeStatus id="7AB1F2A7C04330467FFC45EAB6CEF80A" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
: • same locality and date:
|
||
<specimenCount id="D2C553FB9DF10F69D30F659429AE5023" count="1" type="male">1 ♂</specimenCount>
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<specimenCount id="520A2F0E13581503ED9C15BD9F744080" count="1" type="female">1 ♀</specimenCount>
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(
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<named-content id="E6BB938FFF00F4DE79B2ACDFFD4E11B2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/monte-l-bean-life-science-museum" xlink_title="Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, US">
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<collectionCode id="FA9A184B0D23B386411D52B44F700C3B" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" type="University or college">BYU</collectionCode>
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</named-content>
|
||
)
|
||
</materialsCitation>
|
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•
|
||
<materialsCitation id="E0EB637C4CDC5E7294CAAC4E330262B2" collectingDate="2023-05-05" collectionCode="CGV" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
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<specimenCount id="D7FA167DF4F2BFE2D15197057821F24B" count="1" type="male">1 ♂</specimenCount>
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<specimenCount id="82006D631148CF59A500A48966297171" count="1" type="female">1 ♀</specimenCount>
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(
|
||
<abbrev content-type="institution" id="ABBRID0EBSBG" xlink_title="Gilles Vinçon Collection, Grenoble, France">
|
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<collectionCode id="8E26273B9B6D2F4F2F22DE95653A3CA4">CGV</collectionCode>
|
||
</abbrev>
|
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)
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</materialsCitation>
|
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•
|
||
<materialsCitation id="C190BDAD34B4B34F09FFC9294BED93B3" collectingDate="2023-05-05" collectionCode="EKCU" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCode="PLP 5706" specimenCount="19" specimenCount-female="8" specimenCount-male="11">
|
||
<specimenCount id="B406BE9F0E609278985C0CF37C3DBC46" count="11" type="male">11 ♂</specimenCount>
|
||
<specimenCount id="7EEF8C93889149EAA843E4181A29757A" count="8" type="female">8 ♀</specimenCount>
|
||
(3 pairs in copula) (
|
||
<abbrev content-type="institution" id="ABBRID0EGSBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
|
||
<collectionCode id="23EF534BBA60DA22D456AD48C86E7288">EKCU</collectionCode>
|
||
</abbrev>
|
||
:
|
||
<specimenCode id="19E414D72252DFEDC70EC0222D02DE40" collectionCode="EKCU">PLP 5706</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="64D7E2C3C7FE14BF4D847EF7812C4860" collectingDate="2023-05-05" collectionCode="MM" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCode="2023-49, PLETYP-38" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
||
<specimenCount id="0FF57A6CD9A0FC8B55864F8E360EF56B" count="1" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="CCDC074B4938B93774A067A55E81A322" count="2" type="female">2 ♀</specimenCount>
|
||
(
|
||
<named-content id="DADF0120711B2F4C2BDEBC14FEA0E208" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||
<collectionCode id="D7F6A0D886F8FB02C04EDC0DE9417A4D" country="France" name="University of Montpellier">MM</collectionCode>
|
||
</named-content>
|
||
:
|
||
<specimenCode id="19D45309748DC1E88AAD02B976CE52C3">2023-49</specimenCode>
|
||
,
|
||
<specimenCode id="18EC84A86471A10471B9A5E5A143830A">PLETYP-38</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="874DE4B6A20B9E6C4E72B4FDDFE7672D" collectingDate="2023-05-05" collectionCode="SMNS" collectorName="T. Kovacs & D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="5A55B27C3ABFC5366306EB68A1FC6C92" count="1" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="FDC220A0A7EACD5A992123A80031F3A9" count="1" type="female">1 ♀</specimenCount>
|
||
(
|
||
<named-content id="3DACA575285EE2ABC234C7D364E04EEB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-fuer-naturkund-stuttgart" xlink_title="Stuttgart State Museum of Natural History, Stuttgart, Germany">
|
||
<collectionCode id="AA88C77959FBC2A9745C0CDFD57C5557" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" type="">SMNS</collectionCode>
|
||
</named-content>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="642CC0E789FD15E3C23F23C1F7C312E6" collectingDate="2022-06-11" collectionCode="EKCU" collectorName="A. Hunyadi & T. Kovacs & D. Muranyi & P. Olajos" country="Serbia" county="Katun" elevation="1845" latitude="42.6185" location="partly open brook and spring along the Katun road" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" specimenCode="PLP 5584" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
|
||
<collectingMunicipality id="BD9E68DC1811DEC837849A2C14DF0399">Plav municipality</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="78F803975256E3A4045FF055B78646B2">Visitor Mts</collectingMunicipality>
|
||
,
|
||
<location id="C97D8D5F029AF91752767115EFEB5CE0" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:C97D8D5F029AF91752767115EFEB5CE0" country="Serbia" county="Katun" latitude="42.6185" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" name="partly open brook and spring along the Katun road">partly open brook and spring along the Katun road</location>
|
||
,
|
||
<quantity id="85CF9027D67FC9C3142AA08FA1CC42BA" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">
|
||
<elevation id="ED9BD750267B3888F67063990D5D4815" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">1845 m</elevation>
|
||
</quantity>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYSBG" specific-use="{"type":"Point","coordinates":[19.837800,42.618500]}">
|
||
<geoCoordinate id="73CC9376D83EF3DEFB823BEE64D9168E" degrees="42.6185" direction="north" orientation="latitude" precision="5" value="42.6185">42.6185°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AFF32FD41E4BE3A74C8A88F70E2EFA25" degrees="19.8378" direction="east" orientation="longitude" precision="5" value="19.8378">19.8378°E</geoCoordinate>
|
||
</named-content>
|
||
,
|
||
<date id="BD219651E8D49768D2C6920B692E3CA1" value="2022-06-11">
|
||
<collectingDate id="C0EFC641586B2CCA625E8C426983ABEB" value="2022-06-11">11.vi.2022</collectingDate>
|
||
</date>
|
||
(loc. 24), leg.
|
||
<collectorName id="8DFD1A1E3FC6EAA624544881D07E3C0D">A. Hunyadi</collectorName>
|
||
,
|
||
<collectorName id="D313D2E5D349BBF31F32E8F55CD89B9C">T. Kovács</collectorName>
|
||
,
|
||
<collectorName id="040F81C5C9B51EA63C48590F7CFA00FD">D. Murányi</collectorName>
|
||
,
|
||
<collectorName id="EDBE98D8B2F31E00DF97DA8643D62071">P. Olajos</collectorName>
|
||
:
|
||
<specimenCount id="A5E93276699B4394596527E516A30F52" count="5" type="male">5 ♂</specimenCount>
|
||
<specimenCount id="1251AE8D1921418F1A8A4A03A8FD7920" count="3" type="female">3 ♀</specimenCount>
|
||
(
|
||
<abbrev content-type="institution" id="ABBRID0E4SBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
|
||
<collectionCode id="053A908A0833F051DF55097473D5753B">EKCU</collectionCode>
|
||
</abbrev>
|
||
:
|
||
<specimenCode id="BC85F061FC7AD64265690ADC99FC36FD" collectionCode="EKCU">PLP 5584</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="BF9E8A786ECC08B65833B42E2D046167" collectingDate="2022-06-11" collectionCode="MM" collectorName="A. Hunyadi & T. Kovacs & D. Muranyi & P. Olajos" country="Serbia" county="Katun" elevation="1845" latitude="42.6185" location="partly open brook and spring along the Katun road" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" specimenCode="2022-90, PLETYP-39" specimenCount="17" specimenCount-female="9" specimenCount-male="8">
|
||
<specimenCount id="A3116784D2FDAC242A80A209621A6E48" count="8" type="male">8 ♂</specimenCount>
|
||
<specimenCount id="76E67D2CCD8A18DF26DD0FAC01738A4B" count="9" type="female">9 ♀</specimenCount>
|
||
(1 pair in copula) (
|
||
<named-content id="2C62FBD2EF9BAFA6D4E3ACCAA8A825F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||
<collectionCode id="AD1204AF4BFD7DB68866A319A4F6EBFF" country="France" name="University of Montpellier">MM</collectionCode>
|
||
</named-content>
|
||
:
|
||
<specimenCode id="1767C9A9E4E92FF9D4D1068F87E8EEAE">2022-90</specimenCode>
|
||
,
|
||
<specimenCode id="1BABC28478128932A652B25D66B66751">PLETYP-39</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="57BA5ADDE1F4746D21AC2DD4D9D1FEB3" collectingDate="2022-06-11" collectionCode="EKCU" collectorName="A. Hunyadi & T. Kovacs & D. Muranyi & P. Olajos" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="PLP 5572" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<collectingMunicipality id="17A940B1A78D3E0D3557436F5F8567FC">Plav municipality</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="EE14053A8094F9BE8ECC00FB698523BA">Visitor Mts</collectingMunicipality>
|
||
,
|
||
<location id="77C9C54CFC45F14DD2E95B3F75699CE7" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:77C9C54CFC45F14DD2E95B3F75699CE7" country="Serbia" county="Katun" latitude="42.6294" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" name="forest streams and brooks along the Katun road">forest streams and brooks along the Katun road</location>
|
||
,
|
||
<quantity id="99E51062DE18C1FB96F1C9A7717BAA70" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">
|
||
<elevation id="EC1D464FC85771792ADA706B3FA4CE10" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">1515 m</elevation>
|
||
</quantity>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKTBG" specific-use="{"type":"Point","coordinates":[19.844200,42.629400]}">
|
||
<geoCoordinate id="7C2570353B6F1BB37BEF61CB3F39330A" degrees="42.6294" direction="north" orientation="latitude" precision="5" value="42.6294">42.6294°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3ECAC83EEE5866B06F41CFA8467A4210" degrees="19.8442" direction="east" orientation="longitude" precision="5" value="19.8442">19.8442°E</geoCoordinate>
|
||
</named-content>
|
||
,
|
||
<date id="7EFEFED731BE6392A88D2918A0B7ACA2" value="2022-06-11">
|
||
<collectingDate id="BE827A1E1419F69FC514593E2BFFFC2C" value="2022-06-11">11.vi.2022</collectingDate>
|
||
</date>
|
||
(loc. 22), leg.
|
||
<collectorName id="70F2FFF8B4458336B34ECC490E490E9D">A. Hunyadi</collectorName>
|
||
,
|
||
<collectorName id="FF81DBDF2CDBA77D3EBC7F7DA0D3B3F1">T. Kovács</collectorName>
|
||
,
|
||
<collectorName id="4FD697D3FDCEB4614D844F154A4B3E95">D. Murányi</collectorName>
|
||
,
|
||
<collectorName id="B9A9DDFC5F6EB362C81441411A7E19C3">P. Olajos</collectorName>
|
||
:
|
||
<specimenCount id="274CA913DE95444369BCD4D8DDF5767D" count="1" type="female">1 ♀</specimenCount>
|
||
(
|
||
<abbrev content-type="institution" id="ABBRID0EPTBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
|
||
<collectionCode id="8C676EAB099845F15C0D596C8EAC4D4B">EKCU</collectionCode>
|
||
</abbrev>
|
||
:
|
||
<specimenCode id="BC3EA7D7C6573329E5D1CF6B6FFB9C6D" collectionCode="EKCU">PLP 5572</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="67A17445E9C8C2F81E4AEDB6140355F8" collectingDate="2023-05-05" collectionCode="EKCU" collectorName="L. P. Kolcsar & T. Kovacs & D. Muranyi" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="PLP 5701" specimenCount="15" specimenCount-female="7" specimenCount-male="8">
|
||
same locality,
|
||
<date id="8244337572CEF53DBDF0864FE5468230" value="2023-05-05">
|
||
<collectingDate id="FF8BCB823C87B604C878E070222002A0" value="2023-05-05">5.v.2023</collectingDate>
|
||
</date>
|
||
(loc. 12), leg.
|
||
<collectorName id="D439F5561939F513C6C9868BD75E5F6F">L. P. Kolcsár</collectorName>
|
||
,
|
||
<collectorName id="151EA1847C827F98C00B33826A31F8CE">T. Kovács</collectorName>
|
||
,
|
||
<collectorName id="15C28C8047CAB4F50103BE2CC58AE815">D. Murányi</collectorName>
|
||
:
|
||
<specimenCount id="C0D257A5E2E5B658EEBB599223E886E1" count="8" type="male">8 ♂</specimenCount>
|
||
<specimenCount id="C05B866FE52A457B8E2A69970DAA9071" count="7" type="female">7 ♀</specimenCount>
|
||
(2 pairs in copula) (
|
||
<abbrev content-type="institution" id="ABBRID0EUTBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
|
||
<collectionCode id="B5F02FAB642D3BD83E804E8CE443E846">EKCU</collectionCode>
|
||
</abbrev>
|
||
:
|
||
<specimenCode id="3406D87548C609707634E7DCE967C79F" collectionCode="EKCU">PLP 5701</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="D4309814D3288D8805572060E904F19C" collectingDate="2023-05-05" collectionCode="MM" collectorName="L. P. Kolcsar & T. Kovacs & D. Muranyi" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="2023-48, PLETYP-40" specimenCount="14" specimenCount-female="8" specimenCount-male="6">
|
||
<specimenCount id="D8BF9B31422C257BD091F31E0519C639" count="6" type="male">6 ♂</specimenCount>
|
||
<specimenCount id="2307CDCA3BD2629985A3AF07247C687E" count="8" type="female">8 ♀</specimenCount>
|
||
(1 pair in copula) (
|
||
<named-content id="8319A630D72EDCD8CB9A5F3B7FE00B0E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
|
||
<collectionCode id="C19C7AB5CBAFE8164D80A9A0E8FDCEE2" country="France" name="University of Montpellier">MM</collectionCode>
|
||
</named-content>
|
||
:
|
||
<specimenCode id="EE61A8828BC2F4BFAEB3147B93916E61">2023-48</specimenCode>
|
||
,
|
||
<specimenCode id="77E01B9CDA41FED892BD0E1BD06F33EA">PLETYP-40</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E5TBG" type="diagnosis">
|
||
<paragraph id="7745A045E34DBBE337057C5A0160F72B">
|
||
<heading id="74CEC6603DC5E584686D155D8FA35D3B" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="6B0C07A1EB1DA52F48896456DEB3CE3E">Brachypterous in both sexes. Male tergite VII with small membranous portion; tergite VIII with strong posteromedial process that is not erect in side view and bear angular teeth, membranous area longer than 1 / 2 segment length with a crossband, the posterior margin indented in a rounded membranous field between the posteromedial process and segment sides; tergite IX with short but wide posteromedial sclerite; tergite X posterior margin with a wide V-shaped notch; sternite IX bears a vesicle much shorter than 1 / 2 segment length; specillum slightly longer than paraproct, tip blunt behind subterminal constriction. Female subgenital plate large, trapezoid, with an incision between the long lobes slightly widening towards the median bulge but not forming a triangular field; median bulge is wide in ventral view and distinctly raised in lateral view, with distinct, erect setation; spermathecal sclerite ring-shaped, with large and converging posterior teeth.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDUBG" type="description">
|
||
<paragraph id="CD089914392FCCAA88BD45DADAD52DA5">
|
||
<heading id="03EF998358DE571A538BCFB17B44AE80" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="0FBCC8906DA33134CBE067D8E63DA49E">
|
||
Medium sized, robust species, both sexes brachypterous (Fig.
|
||
<figureCitation id="96AA9A0DFFC4242E2B74664CBF5DFF4E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 A – C</figureCitation>
|
||
). Forewing length:
|
||
<typeStatus id="19D5FDBB040318672B0869D780222F46" type="holotype">holotype</typeStatus>
|
||
<quantity id="953570C42BA04073814F6579F1803043" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
|
||
, male
|
||
<typeStatus id="50B8666C29443CD52991BF9B14C420A1" type="paratype">paratypes</typeStatus>
|
||
3.0–
|
||
<quantity id="C6EBE731E2697F45C24CB9D0FF5F36B1" metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, female
|
||
<typeStatus id="52505A0D89EECA02D9C3477876ED6EFF" type="paratype">paratypes</typeStatus>
|
||
<quantity id="2038165EE469136CA90BFFD8BDF863C5" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" unit="mm" value="4.7" valueMax="5.2" valueMin="4.2">4.2–5.2 mm</quantity>
|
||
; body length:
|
||
<typeStatus id="20C543D28703D235A6C7E914491696EB" type="holotype">holotype</typeStatus>
|
||
<quantity id="E8A47D07863D35D2426D09B446917F5A" metricMagnitude="-3" metricUnit="m" metricValue="6.2" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, male
|
||
<typeStatus id="ECAB02EF9E3F79BA9EC354A82BFE94DF" type="paratype">paratypes</typeStatus>
|
||
5.4–7.0 mm, female
|
||
<typeStatus id="D26D2CA822727759B44F94769E7C8AE2" type="paratype">paratypes</typeStatus>
|
||
6.8–9.0 mm. Setation generally short and dense. General colour dark brown to blackish. Head, antennae, and palpi dark brown. Pronotum dark brown, as wide as long and having rounded corners, rugosities distinct. Legs brown to dark brown. Wings brownish but hyaline, venation brown.
|
||
</paragraph>
|
||
<paragraph id="951C26E2DBBA2A7482AB5138BEE99A92">
|
||
Male abdomen (Figs
|
||
<figureCitation id="E7FFEE92625ED227ED73ABD8BC6BF582" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10 A – D</figureCitation>
|
||
,
|
||
<figureCitation id="A4E6F37345C0B7FC4FDAA927F1810D1A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 B</figureCitation>
|
||
): Tergite I membranous in its whole medial portion, tergite II membranous anteromedially behind antecosta. Transverse row of four pigmented spots distinct on terga I – VII. Terga II and III with medially divided antecosta, terga IV – VII with entire antecosta and full sclerotised, tergite VII with small posterior membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends in a small triangular plate, and the membranous area between them is bridged by a weakly sclerotised crossband; width of the membranous area is less than 1 / 2 the segment width, and reaches below 1 / 2 the segment length above the posteromedial process, posterior margin of the membranous portion nearly straight; the surrounding sclerotisation of the membranous area paler than remainder of the segment; posteromedial process paler than the lateral areas of the segment, not erect in side view and slightly overhangs the segment posterior edge, as wide as the membranous area above, bilobed with angular teeth slightly pointed laterad, the space between the teeth is deeply indented and concave; posterior margin between the posteromedial process and segment sides indented in a shallow, rounded membranous field but not lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 2 of its length; posteromedial sclerite short but wide, medial areas usually slightly darker, the medial portion usually not narrowed. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep, V-shaped notch, 1 / 2 as wide as tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk relatively long. Cercus short, covered with long setae. Sterna II – VIII simple, sternite IX bears a vesicle much shorter than 1 / 2 segment length, its width more than 1 / 2 its length; the area surrounding the vesicle is pale, posterior edge with two pale and weakly sclerotised areas. Paraproct with moderately wide base, abruptly narrowed after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, short, blunt triangular process. Specillum slightly longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, blunt tip.
|
||
</paragraph>
|
||
<caption id="4D7C9FEADB58CC7520FBD70BA49246CA" ID-DOI="10.3897/zookeys.1218.120744.figure10" ID-arpha="2C375FA0-3562-5AAA-9381-95F4188FEC09" httpUri="https://binary.pensoft.net/fig/1175461" startId="F10">
|
||
<paragraph id="10691B74F9BDE423E9692267691AC299">
|
||
<label id="E8D12188D5EC14FF9816D311BBFDFD2F">Figure 10.</label>
|
||
</paragraph>
|
||
<paragraph id="6917B7A7BB38EB0B81C35D288A0E7909">
|
||
Terminalia of
|
||
<taxonomicName id="009F4B15AD2AB9FFD5029DF30B58B062" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
|
||
<emphasis id="67BB788B7DF7AF345CCADE766D2B0210" italics="true">Leuctra visitor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90416D15AF6C653C28C2CD790E217F61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="5D4453C581F15C9215F82ED7CBF35780" bold="true">A</emphasis>
|
||
male terminalia, dorsal view
|
||
<emphasis id="710153C13C0A021B472826A0CD25713D" bold="true">B</emphasis>
|
||
same, lateral view
|
||
<emphasis id="23812668778E8617AD813341900BB461" bold="true">C</emphasis>
|
||
same, ventral view
|
||
<emphasis id="911D93C9F164711847A213E524EE1419" bold="true">D</emphasis>
|
||
male paraprocts and specillae, caudal view
|
||
<emphasis id="9F6865CFDDF9BE36A4B954A54ACA1453" bold="true">E</emphasis>
|
||
female terminalia, ventral view
|
||
<emphasis id="A324E1A98F6B0FD893D209C30D6AE545" bold="true">F</emphasis>
|
||
same, lateral view. Setation omitted with the exception of subgenital plate on
|
||
<emphasis id="8DC9B2D7F2FC138701465E4289FFAFE4" bold="true">F</emphasis>
|
||
. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F08BD09240EDE018EF698EDDB7CEA6F6" ID-DOI="10.3897/zookeys.1218.120744.figure11" ID-arpha="680DCBF2-C9BE-5907-B2CB-A9055C8270FB" httpUri="https://binary.pensoft.net/fig/1175462" startId="F11">
|
||
<paragraph id="A20904142E00D77BF0F0E026084B2E6B">
|
||
<label id="3CC4DE726E014D47AF6D1C735168A1AD">Figure 11.</label>
|
||
</paragraph>
|
||
<paragraph id="893773C691D3DDAB95D16E236D5E0B41">
|
||
Habitus and habitat of
|
||
<taxonomicName id="B93B018737BA3D641BFE10FA68C8ABE8" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
|
||
<emphasis id="69670862E94EF06B6D545DDB18955D96" italics="true">Leuctra visitor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3E9344E641F806FD8830F174CC842E48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="1FDEB634A4F4EEC64975586E4FFCD774" bold="true">A</emphasis>
|
||
male habitus
|
||
<emphasis id="EB9C8BF335F621D829481AFC761B72D3" bold="true">B</emphasis>
|
||
female habitus
|
||
<emphasis id="0DB99A2520C27F4448678232A35A854D" bold="true">C</emphasis>
|
||
copula
|
||
<emphasis id="4E749354253E1F0BFED869F3DDDD60F8" bold="true">D</emphasis>
|
||
partly open brook at 1845 m in the Visitor Mts.
|
||
<emphasis id="C2F511E997ABC8B9E0A4ECCB6492622D" bold="true">E</emphasis>
|
||
type locality: forest springs and their outlet at 1730 m in the Visitor Mts.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2E482067E55FA955FE302EF3AF5CE5BE" ID-DOI="10.3897/zookeys.1218.120744.figure12" ID-arpha="58F889BE-E1CD-54C3-985B-73213274A2E5" httpUri="https://binary.pensoft.net/fig/1175463" startId="F12">
|
||
<paragraph id="2FBD7E8ABFE07BA3140525A926347B1E">
|
||
<label id="40DE752A54B73D809D14DF8E040395BF">Figure 12.</label>
|
||
</paragraph>
|
||
<paragraph id="D60C7EFC73BFFC22E23CE8F06D99A819">
|
||
Terminalia of five Balkan members of the
|
||
<taxonomicName id="E8BF02B21D939A34EB26F169E0E7415C" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="C0469E434357CFE38812DB32117328DC" italics="true">Leuctra prima</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
<emphasis id="3AC78F880E41C9B40AB71802658E5358" bold="true">A – D, I, K</emphasis>
|
||
male terminalia, dorsal view
|
||
<emphasis id="780C1888B7CFC9EA4AA0489CFC418B8B" bold="true">E – H, J, L</emphasis>
|
||
female terminalia, ventral view
|
||
<emphasis id="45FFCB7D6CF9B56EB2DD09919B53EF03" bold="true">A, E</emphasis>
|
||
<taxonomicName id="CC61EEC2042FA632F72BD26D41D39B1E" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="puskasi" status="sp. nov.">
|
||
<emphasis id="7B27045A59C9B5CC8BD73AB158A0F578" italics="true">L. puskasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A3BF7804DC4E4F21591AE5DB4C14E1A2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Kozara Mts, Bosnia & Herzegovina
|
||
<emphasis id="D65AF1A5954FA41A0C87AF7897629F9B" bold="true">B, F</emphasis>
|
||
<taxonomicName id="93CA346F20A9A98A1A9A3CA3D351075A" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
|
||
<emphasis id="8D8E138F0910898CF3CF1D327D4726F6" italics="true">L. visitor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8235A8AE1528A7C8D8A2E03E46E91D4E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Visitor Mts, Montenegro
|
||
<emphasis id="E1211E7681E0864D26E45008199BAD85" bold="true">C, G</emphasis>
|
||
<taxonomicName id="8DF01D015B60083BF2A1BEF7C053FF73" authority="Vincon & Muranyi, 2007" authorityName="Vincon & Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="A8D9E553557423677C3FB52633F39863" italics="true">L. dalmoni</emphasis>
|
||
<bibRefCitation id="EA0CE98699B0DB3D89E7AC168CC41EAA" author="Vinçon G & Murányi D" issue="3" journalOrPublisher="Nouvelle Revue d’Entomologie" pagination="237-248" refId="B82" refString="Vincon G, Muranyi D (2007) Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera). Nouvelle Revue d'Entomologie 23 (3): 237 - 248." title="Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera)." volume="23" year="2007">Vinçon & Murányi, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, Visitor Mts, Montenegro
|
||
<emphasis id="3F202F744E834315C44E7B3863B91017" bold="true">D, H</emphasis>
|
||
<taxonomicName id="16750FCB5DE867757F658FC086DF2298" authority=", Golija Mts" authorityName="Golija Mts" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="5525039FCBF5B12527D92942BAA9F35D" italics="true">L. dalmoni</emphasis>
|
||
, Golija Mts
|
||
</taxonomicName>
|
||
, Serbia
|
||
<emphasis id="C440F3D6BAE671ED4E1C6CABF41BC5CD" bold="true">I – J</emphasis>
|
||
<taxonomicName id="52A933EBE40824EF47DF873AC679DB8D" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="62485E51826F15147E96E3DCB63982FB" italics="true">L. prima</emphasis>
|
||
<bibRefCitation id="349E484119062605EE4D209DA8B689DA" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
, Börzsöny Mts, Hungary
|
||
<emphasis id="B4FACE2D44C4051760A501A5173FCBA6" bold="true">K – L</emphasis>
|
||
<taxonomicName id="322364AFD6E2F646549851BCEFE16C21" authority="Aubert, 1954" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
|
||
<emphasis id="1A6D3F3B119017689462E112209E0BA0" italics="true">L. pseudosignifera</emphasis>
|
||
<bibRefCitation id="AAD5147A2DF0ABA04DA0D178602231B7" DOI="10.5169/seals-401211" author="Aubert J" issue="2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="124-136" refId="B2" refString="Aubert J (1954) Contribution a l'etude du genre Leuctra Stephens et description de quelques especes nouvelles de ce genre. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 27 (2): 124 - 136. https: // doi. org / 10.5169 / seals- 401211" title="Contribution à l’étude du genre Leuctra Stephens et description de quelques espèces nouvelles de ce genre." volume="27" year="1954">Aubert, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, Poljana Mts, Slovakia. Scale bars: 1 mm (
|
||
<emphasis id="C96C03473DA712FECF8B3AC09B9864FE" bold="true">D</emphasis>
|
||
for
|
||
<emphasis id="42E561F08BBF4573513BD7021F71CAD2" bold="true">A – D, I, K; E</emphasis>
|
||
for
|
||
<emphasis id="4BFF980DE183E6EE6FC4B70189F99764" bold="true">E – H, J, L</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D290DBB28C6F117BC2994F8F6A333D40">
|
||
Female abdomen (Figs
|
||
<figureCitation id="8AEB333271813C1467DBA4D94DEAD137" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10 E, F</figureCitation>
|
||
,
|
||
<figureCitation id="87C7F6CADD900EBC2D0B6FF141438CA7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 F</figureCitation>
|
||
): Terga I – VIII with transverse row of four pigmented spots clearly visible; terga I – VIII mostly membranous but with lateral sclerites, terga II, III, and VI – VIII, sometimes also tergite V, bear a patch-like medial sclerite, largest on tergite VIII; tergite IX membranous only anterolaterally, tergite X fully sclerotised. Sterna I – VII simple, sterna II – VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large wide lobes, slightly darker brown than remainder of the plate; posterior margin convex, incision between the lobes is a membranous strip, slightly widened towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is wide in ventral view, distinctly raised, nose-shaped in lateral view, with distinct erect setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus, and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging posterior teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZ1BG" type="Affinities">
|
||
<paragraph id="D5A73455103211B7BFE26FBE5F88C6A0">
|
||
<heading id="974B8785FFE8B99FD3FD7FCDD12AF52B" reason="title">Affinities.</heading>
|
||
</paragraph>
|
||
<paragraph id="CAA18A149D315EDD1DA546E30FDB2A69">
|
||
The males are morphologically closest to
|
||
<taxonomicName id="6E9A988BD8D9543D276821D647D62AC8" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="CA5D84D46B6FA1FEBF830C19E081A347" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
(compare with Fig.
|
||
<figureCitation id="25F6356425B5D3BEB4B0429ED08BBCB9" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 C, D</figureCitation>
|
||
) while the female is closest to
|
||
<taxonomicName id="4C201A3A1D06853DD7BC56AF08183B2E" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="35AAF42BE7E35E087969ECB058C40165" italics="true">L. prima</emphasis>
|
||
</taxonomicName>
|
||
(compare with Fig.
|
||
<figureCitation id="A611E44242B7F28CF9E9DCD972BBFC79" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 J</figureCitation>
|
||
). Males can be distinguished from
|
||
<taxonomicName id="CFEE0C55A5BE6534973C4A2E551EA910" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="A8E58FF19F07FD86C00E54699864AB9D" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the following characters: a weak crossband on the membranous area of tergite VIII lacking in
|
||
<taxonomicName id="97461D5890CD70A67FC0F85ACC2B04CB" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="B8AB1BD80C24FF8DC1886E09F0FAC02C" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
, angular teeth of the posteromedial process of tergite VIII stronger than those of
|
||
<taxonomicName id="B2922D707A71D70E521C3C052771AB4E" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="948E347F4ACA5B46A29F01A22DD761FF" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
; the posterior indentation of tergite X is V-shaped, while it is rounded in
|
||
<taxonomicName id="3BC2B0B9749D67087850E9ACE79B1F2C" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="E4AA85190E39CA04603F01BEC54E6A7D" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
; the ventral vesicle of sternite IX is shorter than 1 / 2 the segment length, while clearly longer than 1 / 2 the segment length in
|
||
<taxonomicName id="70337ED9A4B145864F9C0AB324ED8F57" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="A48A1F66B13344F3E2DBF4C01F61614D" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
. The males of the new species can be easily distinguished from
|
||
<taxonomicName id="06715D5F819D2225D21E9EB9668F93BA" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="0DE93DFCAB063A6D685C81126446C4F6" italics="true">L. prima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2DAFBEEF6B99E872B45EEC5C9A7265D2" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
|
||
<emphasis id="5B9599C0C1D3F367348F8FAAC98538B4" italics="true">L. pseudosignifera</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="09806C6A065CEB87C875414F88D247E3" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
|
||
<emphasis id="101499A749CDFBFE9AAFC80391ABECEA" italics="true">L. pseudosignifera</emphasis>
|
||
</taxonomicName>
|
||
is characterised by a pair of rounded sclerotised expansions, located between the posteromedial process and segment sides of tergite VIII (Fig.
|
||
<figureCitation id="886B6365343A5CC9A5A7B93A41B41BB9" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 K</figureCitation>
|
||
); the posteromedial process of tergite VIII is erect in lateral view in
|
||
<taxonomicName id="117CBC981FFC5D5B5FA97B0DE8B42294" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="8B0CF75586C73F8A2E44C7FDA9986F85" italics="true">L. prima</emphasis>
|
||
</taxonomicName>
|
||
; furthermore, this species has a divided posteromedial sclerite on tergite IX (Fig.
|
||
<figureCitation id="4C66AA10B2758C5919C1762529AEE155" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 I</figureCitation>
|
||
). Females can be distinguished from
|
||
<taxonomicName id="A8D5C866B957C719E40AE56F9764F6DB" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="B8550F770306850A5B18EFEB64290FE6" italics="true">L. prima</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the distinct, erect hairs of the subgenital plate as seen in lateral view, and the nose-shape of the bulge of the subgenital plate; the bulge is rounded in
|
||
<taxonomicName id="C140B4466DD13A9739F43DD93E5E46D7" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
|
||
<emphasis id="2733C977CBAB9D5BC4F2896C2A6FF34F" italics="true">L. prima</emphasis>
|
||
</taxonomicName>
|
||
and the subgenital plate lacks distinct setation. The wide bulge of the subgenital plate (in ventral view), and the erect hairs (in lateral view) distinguish the new species from
|
||
<taxonomicName id="5F0F3F680865A495D1D02C50DFB422B2" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="84418B16184B3CC2218A05C0EA7796F2" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4083E7513D79DA3F0D225A0430B7DDA4" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
|
||
<emphasis id="393FA3209847260EA91E0DD11B526B5A" italics="true">L. pseudosignifera</emphasis>
|
||
</taxonomicName>
|
||
; furthermore,
|
||
<taxonomicName id="5FE9D11FA0FEBBA87B04896D0347F824" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
|
||
<emphasis id="E235F4C69FD5CF9BEF51BD8C08FACDDA" italics="true">L. pseudosignifera</emphasis>
|
||
</taxonomicName>
|
||
is characterised by a triangular membranous field at the end of the incision between the lobes of the subgenital plate (Fig.
|
||
<figureCitation id="455C7A2757C9588CA5B755C0BB61B258" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 L</figureCitation>
|
||
). Distinction of both sexes from
|
||
<taxonomicName id="DCF39647F37E8A44A3CFF5FCCF5981CF" authorityName="Murányi & Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="puskasi">
|
||
<emphasis id="1C76928453E7E69C03331BCBBA2CBDA3" italics="true">L. puskasi</emphasis>
|
||
</taxonomicName>
|
||
was described in detail above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EPBAI" type="distribution">
|
||
<paragraph id="065551569A4C4FBEBDC9CAA05B8DAF51">
|
||
<heading id="6FEF3BF9056C80D77336CFFBFA203D33" reason="title">Distribution and ecology.</heading>
|
||
</paragraph>
|
||
<paragraph id="BC08BCD120216946CB2FAEB678C0166C">
|
||
The species was collected at three high-elevation brooks and spring outlets in the Visitor Mts of eastern
|
||
<collectingCountry id="E2B3D20546F2427466CFEB6037CE6C5F" name="Serbia">Montenegro</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="155558EDF86FEE9AF0FEAA1C97477F7B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">1</figureCitation>
|
||
). The Visitor is a mostly volcanic range of small area, with its highest peak reaching more than 2200 meters, surrounded by the notably higher limestone chains of the Prokletije Mts. The Visitor receives high precipitation and is covered with dense forest, resulting in a rich pattern of different waterflows. The streams where the new species was found are running in mixed forest or partly open meadows, the lentic parts usually with dense
|
||
<taxonomicName id="B02BB6A66B454AC15A5F7EBFE45FA92C" class="Magnoliopsida" family="Ranunculaceae" genus="Caltha" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="F0AF79D53B297D96A7750098222D7C9C" italics="true">Caltha</emphasis>
|
||
</taxonomicName>
|
||
growth (Fig.
|
||
<figureCitation id="E5737B3EB5E12E54733218CE67D3096E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 D, E</figureCitation>
|
||
). The brooks have variable flow rates but are permanent streams, the substrate is stony mixed with mud and plenty of dead wood. The specimens were emerging together with
|
||
<taxonomicName id="B88FE4D6F9C52EB4C536E768320CA269" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="AE4DB61C9E6DA8221BE01CF0072566C2" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="350350234022DDC8CD3B62F5C44F8399" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A – D, I, K male terminalia, dorsal view E – H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia & Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon & Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I – J L. prima Kempny, 1899, Börzsöny Mts, Hungary K – L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A – D, I, K; E for E – H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 C, G</figureCitation>
|
||
) and both species were present with many mating pairs in May, but we found no interbreeding couples (Fig.
|
||
<figureCitation id="87FD4B89A46BEACAF328A9A01A245F85" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 C</figureCitation>
|
||
). We visited the area three times: the new species was more numerous than
|
||
<taxonomicName id="BE947D33ABA02722E14C397D6F80BE8F" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="3812CA66113414FBE658EF81F1C6A658" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
in early
|
||
<date id="D9F49ED0DEACA63C330E6F4A2235467D" value="2023-05">May 2023</date>
|
||
, both scarce in early
|
||
<date id="B6896E2A14E89B0F9C91895BB00766B7" value="2022-06">June 2022</date>
|
||
, and both absent by late
|
||
<date id="EBA4BCDDD5D868E782B8E8C0C5C928B1" value="2023-06">June 2023</date>
|
||
. In May, the upper site (
|
||
<quantity id="9E0BF113DA1A6B643F3B2EBE463CCCB6" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">1845 m</quantity>
|
||
) was not available, and even the lowest site (
|
||
<quantity id="3A8FFA6305176D5EB7F4A91B5386B0C2" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">1515 m</quantity>
|
||
) was partly covered by snow, but all were snow-free by late June. We found a rich accompanying stonefly fauna at the lowest site:
|
||
<taxonomicName id="DD48C4F87FD5B9117A495A1A96F43F8B" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
|
||
<emphasis id="7D045A2B6443AB82505B7783242DFE83" italics="true">Brachyptera helenica</emphasis>
|
||
</taxonomicName>
|
||
(adults and matured larvae, early June),
|
||
<taxonomicName id="6FE5F8F313730AF2147B2FCE51325A9D" authority="Berthelemy, 1971" authorityName="Berthelemy" authorityYear="1971" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="graeca">
|
||
<emphasis id="2409A5E9B5E2605D9D45E034161513D9" italics="true">B. graeca</emphasis>
|
||
<bibRefCitation id="CDE093F91C9CD253D98458BDB200429E" author="Berthélemy C" issue="1" journalOrPublisher="Biologia Gallo-Hellenica" pagination="23-56" refId="B6" refString="Berthelemy C (1971) Plecopteres de Grece Centrale et d'Eubee. Biologia Gallo-Hellenica 3 (1): 23 - 56." title="Plécoptères de Grèce Centrale et d’Eubée." volume="3" year="1971">Berthélemy, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
(adults, late June),
|
||
<taxonomicName id="0DA1D48AC6F7B6F290D611DDA8E77241" baseAuthorityName="Klapalek" baseAuthorityYear="1902" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="seticornis">
|
||
<emphasis id="2C5CCD8DC72FD72DF7CB7137366375A2" italics="true">B. seticornis</emphasis>
|
||
</taxonomicName>
|
||
(adults, early June),
|
||
<taxonomicName id="75934D95CFF62D8C93A11C94DF4E3212" authorityName="Pictet" authorityYear="1841" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="C6D87CAC9E4F1CC1EAA792F7BA3E98FB" italics="true">Capnia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDDF631E5A4C587AACA5512C6F7034EF" sensu="lato">s. l.</taxonomicNameLabel>
|
||
<emphasis id="FA549CD6270AEB2532867943510289FC" italics="true">vidua rilensis</emphasis>
|
||
(adults, May and early June),
|
||
<taxonomicName id="DA428B8DA01A0AD228A764EBB9379C11" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="170A038E30186E5616AFFED049154A80" italics="true">Leuctra nigra</emphasis>
|
||
</taxonomicName>
|
||
(adults, May and June),
|
||
<taxonomicName id="02C5DFE114621CFDCB7A35192EEEBC5E" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="C64E7C2DC022179F3DDAF93AE1CF38A5" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
(adults, May and early June),
|
||
<taxonomicName id="C8704489FD2E7C9B1DB23667C1C3C603" authority="Aubert, 1956 b" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
|
||
<emphasis id="3486936DC74B6D231EBF69A889D58221" italics="true">L. metsovonica</emphasis>
|
||
Aubert, 1956 b
|
||
</taxonomicName>
|
||
(adults, early June),
|
||
<taxonomicName id="894A978CC086F884BCA3E32E99A40AC5" authority="Illies, 1954 b" authorityName="Illies" authorityYear="1954" class="Insecta" family="Nemouridae" genus="Protonemura" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="auberti">
|
||
<emphasis id="FA551445A21891E033A87DD17F68DCD8" italics="true">Protonemura</emphasis>
|
||
cf.
|
||
<emphasis id="A5C9E768F01E71DD82271AB8ED8A69F7" italics="true">auberti</emphasis>
|
||
Illies, 1954 b
|
||
</taxonomicName>
|
||
(adults and matured larvae, early June),
|
||
<taxonomicName id="C423729F2F5D7557D2072A9F53F4AB9E" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
|
||
<emphasis id="D7F2E5C4AEBA05A6D7BD95B3B1F161F7" italics="true">Nemurella pictetii</emphasis>
|
||
</taxonomicName>
|
||
(adults, May and June),
|
||
<taxonomicName id="5599EF53973BF50994E8AB4DDD700FFA" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
|
||
<emphasis id="CD4B5763DC462A4B16707AD5AB3F0D1B" italics="true">Nemoura cinerea cinerea</emphasis>
|
||
</taxonomicName>
|
||
(adults, early June),
|
||
<taxonomicName id="03C8CF6157734147B483DC11236A3A53" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
|
||
<emphasis id="6B34C54D13F66C17479213CB553ADC4B" italics="true">Nemoura marginata</emphasis>
|
||
</taxonomicName>
|
||
(adults, May and early June), a yet unnamed
|
||
<taxonomicName id="173E7DBBE25AFA13780A0093908A533B" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F18E0A04742C351753657D0963169CD7" italics="true">Nemoura</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="6151C2CFD1E975027FD4A6BD0FF45819" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
|
||
<emphasis id="8B8DE4448B75D309AF3DB9F579412A91" italics="true">marginata</emphasis>
|
||
</taxonomicName>
|
||
group (adults, May and early June),
|
||
<taxonomicName id="ED7841E01AE8D6A7CD8455005943F4CF" authority="Despax, 1934" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
|
||
<emphasis id="4299179A632190A1BBA1EF5B3466929B" italics="true">N. uncinata</emphasis>
|
||
<bibRefCitation id="0741902B260B38F224D4F5DD2993F288" author="Despax R" issue="2" journalOrPublisher="Bulletin de la Société d’histoire Naturelle de Toulouse" pagination="255-270" refId="B15" refString="Despax R (1934) Plecopteres pyreneens. VII. Etude et description de quelques formes de Nemoures apparentees a Nemoura marginata (Pict.). Bulletin de la Societe d'histoire Naturelle de Toulouse 66 (2): 255 - 270." title="Plécoptères pyrénéens. VII. Etude et description de quelques formes de Nemoures apparentées à Nemoura marginata (Pict.)." volume="66" year="1934">Despax, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
(adults, May and early June),
|
||
<taxonomicName id="1A9765BADA5CF48B39C19C72F84C703E" authority="(McLachlan, 1872)" baseAuthorityName="McLachlan" baseAuthorityYear="1872" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dichroa">
|
||
<emphasis id="0D911BA1724C56F38C1F5DB96FCF04AF" italics="true">Arcynopteryx dichroa</emphasis>
|
||
(
|
||
<bibRefCitation id="BEC3C26269CBE82CB2FBDB560EC262F7" author="McLachlan MR" refId="B42" refString="McLachlan MR (1872) Materiaux pour une Faune Nevropterologique de l'Asie Septentrionale. Seconde partie. Non-Odonates. Annales de la Societe Entomologique Belgique 15: 47 - 77 [+ Pl I - II]. https: // www. biodiversitylibrary. org / item / 44639 # page / 345 / mode / 1 up" url="https://www.biodiversitylibrary.org/item/44639#page/345/mode/1up" year="1872">McLachlan, 1872</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(adults and matured larvae, June), a yet unnamed
|
||
<taxonomicName id="92E2D54F65CB9307712B4D863F42AA98" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BF05F4D88D9A1A0D0C4116DA859D8F16" italics="true">Isoperla</emphasis>
|
||
</taxonomicName>
|
||
related to
|
||
<taxonomicName id="A9434D4203216BB2FEFF257EDDBE3A80" authority="Ikonomov, 1980" authorityName="Ikonomov" authorityYear="1980" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="breviptera">
|
||
<emphasis id="B0B32366453D3E2B1C2C9718225FAC8D" italics="true">I. breviptera</emphasis>
|
||
<bibRefCitation id="8D5B229117C4BBCF9BE71C3250C04E97" author="Ikonomov P" issue="4" journalOrPublisher="Fragmenta Balcanica" pagination="19-31" refId="B23" refString="Ikonomov P (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica 11 (4): 19 - 31." title="Nouvelles espèces de Plécoptères (Insecta, Plecoptera) de Macedoine. II." volume="11" year="1980">Ikonomov, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
(adults and matured larvae, June),
|
||
<taxonomicName id="6560C470B35358458E34D6A44030C15D" authority="Illies, 1954 a" authorityName="Illies" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="tripartita">
|
||
<emphasis id="C2F03F58BB6DF062B5F9D7C9C5DB3D8C" italics="true">I.</emphasis>
|
||
cf.
|
||
<emphasis id="350278A2B960EA2581C223E79B15CC80" italics="true">tripartita</emphasis>
|
||
Illies, 1954 a
|
||
</taxonomicName>
|
||
(larvae, June), and unidentified larvae of
|
||
<taxonomicName id="50E4D8922937B8093D5A29C9630D4C10" authorityName="Zwick" authorityYear="1967" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF01DA61DCE5B14D8C8F177A67756BB2" italics="true">Siphonoperla</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="912B1C73E7A6A6B955DF81E9545F8CD1" authorityName="Newman" authorityYear="1836" class="Insecta" family="Chloroperlidae" genus="Chloroperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2711B112D349CE9C69513B48E69075E" italics="true">Chloroperla</emphasis>
|
||
</taxonomicName>
|
||
(early June). Some of these were probably not developing in the same stream section since the site is placed at a junction of three different brooks. The type locality (sampled only in May) was inhabited by the new species,
|
||
<taxonomicName id="E52237FC5D176B31CCA822B725536469" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="626502107001B9CFB050422DB3DEE77B" italics="true">L. nigra</emphasis>
|
||
</taxonomicName>
|
||
(adults),
|
||
<taxonomicName id="7944E6EB762189FFFF04D8A179825141" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="180FB8333F12430AAFE82274EAF991B3" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
(adults), the unnamed
|
||
<taxonomicName id="73FB3CBD58431AA556AFC805ED495671" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="055FF7C4B7B7C54F9ADFF38175643A48" italics="true">Nemoura</emphasis>
|
||
</taxonomicName>
|
||
(adults and putative larvae), an unidentified
|
||
<taxonomicName id="C25BDB2A01E8012E81B223F44A8FCEBF" authorityName="Kempny" authorityYear="1898" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DECE3D441B488E6C08386D08E659E4FD" italics="true">Protonemura</emphasis>
|
||
</taxonomicName>
|
||
(larvae), and the unnamed
|
||
<taxonomicName id="3FBAAA392CBB4DAB0EC1DEB4A47183AC" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BA64DF64179A8105FF2F91525319508A" italics="true">Isoperla</emphasis>
|
||
</taxonomicName>
|
||
(larvae). At the highest site (sampled only in June), the new species was found together with
|
||
<taxonomicName id="FFDC65054F7BDF717646491F5213339A" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="1572F4A464A5491B4FB71C94C479A42C" italics="true">Leuctra nigra</emphasis>
|
||
</taxonomicName>
|
||
(adults),
|
||
<taxonomicName id="18644824EB4427F453D11083E0D38886" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="EE46DF706F551CB566A3CD6B193C0C67" italics="true">L. dalmoni</emphasis>
|
||
</taxonomicName>
|
||
(adults), the unnamed
|
||
<taxonomicName id="51CF757CCD7FBCA7766B0AEB917309AE" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02B40628AB06D830B9173E5B5B001371" italics="true">Nemoura</emphasis>
|
||
</taxonomicName>
|
||
(adults),
|
||
<taxonomicName id="0C25CA801038F31E4D339C1CC6E3A9A8" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
|
||
<emphasis id="9748A27E3CC5E8BE9F857F885F759C70" italics="true">Nemurella pictetii</emphasis>
|
||
</taxonomicName>
|
||
(adults),
|
||
<taxonomicName id="7A8CF850FC7169B54C9E5D75FE925029" baseAuthorityName="McLachlan" baseAuthorityYear="1872" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dichroa">
|
||
<emphasis id="ED640DC5B1FA83EC8BDE816B29854774" italics="true">Arcynopteryx dichroa</emphasis>
|
||
</taxonomicName>
|
||
(larvae), and the unnamed
|
||
<taxonomicName id="2C9E226BCA0B30B3742630C524244F80" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E2394672E1ACF4BDC38AB4D462EEC0E8" italics="true">Isoperla</emphasis>
|
||
</taxonomicName>
|
||
(adults).
|
||
<taxonomicName id="0FA3C487240C94270A75B1331193E273" authorityName="Pictet" authorityYear="1841" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="BC623DD730D0E48D660DB2950315E30D" italics="true">Capnia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="84063059A86F432DFCA463949E6003E5" sensu="lato">s. l.</taxonomicNameLabel>
|
||
<emphasis id="A1EBB56E7F428E411608102E2B9E7162" italics="true">vidua rilensis</emphasis>
|
||
,
|
||
<taxonomicName id="3D01587F41233D08821768DC44401C7B" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
|
||
<emphasis id="D9F4F564CE8B58BE0DC8123E148C977C" italics="true">Leuctra dalmoni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D4DED9B6BFD84D73681DCA9C0B820C29" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
|
||
<emphasis id="93B2FCA78ED12B0642A338A12F68EAC7" italics="true">L. metsovonica</emphasis>
|
||
</taxonomicName>
|
||
are new records for
|
||
<collectingCountry id="72A27A035B61EF21A60A4E9C60210FED" name="Serbia">Montenegro</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EHOAI" type="etymology">
|
||
<paragraph id="1D2ED37F908BE3B6844D22B741E5173D">
|
||
<heading id="599A6BAE8E221A8ECF8660F756BB3C96" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="9F4BF4A614825BDBE86B169A41CBAEA3">
|
||
The name
|
||
<emphasis id="09E38B3F1BA492B4D8C7D6186AF75304" italics="true">visitor</emphasis>
|
||
is derived from the Visitor Mountains of
|
||
<collectingCountry id="595FF87CF29996ECFE57CEEED6223810" name="Serbia">Montenegro</collectingCountry>
|
||
, where the new species was found and probably restricted to the high elevations of the range. Used as a noun, gender neutral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |