Add updates up until 2024-11-15 18:59:06

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<document id="F2A9A4DC92B6028F972E5DF15DD38461" ID-DOI="10.3897/zookeys.1218.120744" ID-ZooBank="094646BE-BF9F-4890-BD59-FF26FA95A3FE" ID-publisher-id="120744" URI-arpha="6E4DF739-56EF-5B8D-AEB7-2839D486DC50" URI-zoobank="http://zoobank.org/094646BE-BF9F-4890-BD59-FF26FA95A3FE" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731693946914" checkinUser="pensoft" docAuthor="Murányi, Dávid &amp; Kovács, Tibor" docDate="2024" docId="42DFF82501A653BDAAA879B5D3C1D0E1" docLanguage="en" docName="ZooKeys 1218: 49-79" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/120744/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Leuctra golija Murányi &amp; Kovács, 2024, sp. nov." docType="treatment" docUuid="776454AB-1086-4B10-A56B-636E13224914" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="79" masterDocId="6E4DF73956EF5B8DAEB72839D486DC50" masterDocTitle="Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)" masterLastPageNumber="79" masterPageNumber="49" pageNumber="49" updateTime="1731697136958" updateUser="admin">
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<mods:title id="653D2388ADAD63FCD27434C95635E9C2">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:namePart id="F383D841F2FB0F59FE4B45A529FA82CC">Murányi, Dávid</mods:namePart>
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<mods:affiliation id="DAA41CD427B3FF8B0C70639D006F01A8">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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<mods:namePart id="0CD98EC9A5AB6167B42539ECF24D956E">Kovács, Tibor</mods:namePart>
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<mods:affiliation id="E69DAD8E14F17347984A8D0252716AF2">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<paragraph id="7890C5F30FCBF4417CA4BB8A0BF7DFD4">
<taxonomicName id="8935BFE06D20EF1D4125ACD041DA804A" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="golija" status="sp. nov.">
<emphasis id="29C0A77A1ECCF93F418FE1D64AE6E34D" italics="true">Leuctra golija</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8F9315BF2C56AF956242A10300E2A89" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="895226D7C10D44ACB5BEE0FF4BF28FE0" type="description">
<paragraph id="D56F9454DE1BEF3847402CFC3B342058">
<figureCitation id="24F0255DC1F43F29D58DAA0F567719A0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">Figs 1</figureCitation>
,
<figureCitation id="7CAD5C865DABADF0C80663536706EAEB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminalia of Leuctra golija sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure5" httpUri="https://binary.pensoft.net/fig/1175456">5</figureCitation>
,
<figureCitation id="DCCFFB0764D5DB1ACF2F3EF68EFB6B83" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitat of Leuctra golija sp. nov.: Moravica Stream in the Golija Mts (type locality, 23. x. 2023)." figureDoi="10.3897/zookeys.1218.120744.figure6" httpUri="https://binary.pensoft.net/fig/1175457">6</figureCitation>
,
<figureCitation id="591C03E9A8D09939691F07BF2CC61846" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 B, F</figureCitation>
</paragraph>
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<paragraph id="808349F0A297C7670FCF3E4932DEAAF3">
<heading id="98160F23D605912C73923A9474C80430" reason="title">Type material.</heading>
</paragraph>
<paragraph id="08C6DD5E89A9BE43C26E5CE504401A2D">
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<emphasis id="171CDA8DD7AD661CAD25BAC85377DFE5" bold="true" italics="true">
<typeStatus id="AB144CFA191050CF071C65528511CD21" type="holotype">Holotype</typeStatus>
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<specimenCount id="EDD93F33EDC1773D6AFD26EC0AD96D75" type="male">male</specimenCount>
:
<collectingCountry id="319784ADCD88EB61A52BC38D34EB06AA" name="Serbia">Serbia</collectingCountry>
: •
<collectingCounty id="801ABB6044F8BEEDE3CF8C211855DB00">Moravica district</collectingCounty>
,
<collectingMunicipality id="629BCC7976676C1A47A7275D0833F8C9">Ivanjica municipality</collectingMunicipality>
,
<collectingMunicipality id="C31806DB33C9156B9612ED1BFAF99740">Golija Mts</collectingMunicipality>
,
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,
<location id="CA39D33FC5DF093B4CAFF474CDC3C669" LSID="urn:lsid:plazi:treatment:42DFF82501A653BDAAA879B5D3C1D0E1:CA39D33FC5DF093B4CAFF474CDC3C669" country="Serbia" county="Moravica district" latitude="43.3384" longLatPrecision="6" longitude="20.2507" municipality="Ivanjica municipality" name="Moravica Stream at Milosica" stateProvince="Ivanjica">Moravica Stream at Milošica</location>
,
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(field number: loc. 16), leg.
<collectorName id="7498F6D509298613161F0A1118069B28">L. P. Kolcsár</collectorName>
,
<collectorName id="9977A70E0F5C9C71BAB5D326B342E918">T. Kovács</collectorName>
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<collectorName id="7E15264275D8A109104489565089D251">D. Murányi</collectorName>
(
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<specimenCode id="593A1A60F4A3CA26E6A8DD741D64CB74">PLETYP-32</specimenCode>
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<typeStatus id="4E27EAD3DDA6F1E41DE4249ADC1722E0" type="paratype">Paratypes</typeStatus>
</emphasis>
: • same locality and date:
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<specimenCount id="8A39E743874ADEF6F21FDAFC89BA4300" count="1" type="female">1 ♀</specimenCount>
(
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<collectionCode id="E637849BBA314CBA0CCA994A191FEAD0" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" type="University or college">BYU</collectionCode>
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(
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)
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<specimenCount id="154627FC4C7ED9657DA9AAEC92F0E257" count="14" type="male">14 ♂</specimenCount>
<specimenCount id="ECA591881468B9AFEE087960C598E43C" count="6" type="female">6 ♀</specimenCount>
(
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:
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<specimenCount id="6A380DE2E8EE4248775A7E8CD92EC4FA" count="10" type="male">10 ♂</specimenCount>
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(
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<collectionCode id="398801AB5CA3A325ECCA4CED3B70A732" country="France" name="University of Montpellier">MM</collectionCode>
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,
<specimenCode id="2EFAFD4C8CAADFB638F0C72B8B1EB6EF">PLETYP-33</specimenCode>
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<specimenCount id="50817BAA7A5B548E7E68DCD9690B147F" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="F8B2297A8B1CBB1A3EFD93E862C18E27" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-fuer-naturkund-stuttgart" xlink_title="Stuttgart State Museum of Natural History, Stuttgart, Germany">
<collectionCode id="FEB0B7CCEFFDD01F6F26A5A30DF7F07C" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" type="">SMNS</collectionCode>
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)
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same locality,
<date id="C8E387BE351F174BF09253F4A3001382" value="2018-06-26">
<collectingDate id="1E1CC43CEF5AC3EA4F347259CAAE9C21" value="2018-06-26">26.vi.2018</collectingDate>
</date>
(loc. 1), leg.
<collectorName id="F007F91137741FDB624F418ADC5A09B0">P. Juhász</collectorName>
,
<collectorName id="BE7B2919F08E4B0FF57DD39886CCAAE1">T. Kovács</collectorName>
,
<collectorName id="BC3FDE9CB6DEE64DC8534457054318B8">D. Murányi</collectorName>
:
<specimenCount id="001B0DECEBAC855E8465AB6761C8DFCC" count="1" type="female">1 ♀</specimenCount>
(
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:
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<specimenCount id="D4AC9404A9D531D83D4984EC513B31AF" count="1" type="male">1 ♂</specimenCount>
(
<named-content id="63EF1C6A94D5BDBD73E005F978F28354" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="D1A05DC57D8363A6E601F064E41CA507" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="AD43A512F1C1FF3B84BEDA84F3124E8E">2018-43</specimenCode>
,
<specimenCode id="B74E2A72E4AB3F1E41186EC0746BF575">PLETYP-34</specimenCode>
)
</materialsCitation>
<materialsCitation id="A46B32B43FC8C53E6AC33D9C605711B4" collectingDate="2023-05-06" collectionCode="T, EKCU" collectorName="L. P. Kolcsar &amp; Kovacs, D &amp; Muranyi" country="Serbia" elevation="1575" latitude="43.2743" location="Radaljica" longLatPrecision="6" longitude="20.3467" municipality="Golija Mts" specimenCode="PLP 5727" specimenCount="1" specimenCount-male="1" stateProvince="Raska district, Novi Pazar" typeStatus="paratype">
<collectingRegion id="548243B1171D8D6CB4A041B2F886EB9D" isEnumeration="true">Raška district, Novi Pazar</collectingRegion>
municipality,
<collectingMunicipality id="29C5DE64040011FFCF90EB223B3960A5">Golija Mts</collectingMunicipality>
,
<location id="DD9D2E6BF2132999B8464B69CFA219E4" LSID="urn:lsid:plazi:treatment:42DFF82501A653BDAAA879B5D3C1D0E1:DD9D2E6BF2132999B8464B69CFA219E4" country="Serbia" latitude="43.2743" longLatPrecision="6" longitude="20.3467" municipality="Golija Mts" name="Radaljica" stateProvince="Raska district, Novi Pazar">Radaljica</location>
, spring brooks in forest edge,
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<elevation id="680BC6AD3CB88A97D3CFB964137B39CF" metricMagnitude="3" metricUnit="m" metricValue="1.575" unit="m" value="1575.0">1575 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E21AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[20.346700,43.274300]}">
<geoCoordinate id="5A0DFFFF63799B63A02FF7D527425FC3" degrees="43.2743" direction="north" orientation="latitude" precision="5" value="43.2743">43.2743°N</geoCoordinate>
,
<geoCoordinate id="AC5D41AFF08CF459D15E62A8EC18CA0E" degrees="20.3467" direction="east" orientation="longitude" precision="5" value="20.3467">20.3467°E</geoCoordinate>
</named-content>
,
<date id="76C959A45D8D72E62DE1E3F6CE3FAF80" value="2023-05-06">
<collectingDate id="DB13F0D2D3F5C13A29F8518B2C9D5F32" value="2023-05-06">6.v.2023</collectingDate>
</date>
(loc. 17), leg.
<collectorName id="601BC06B49FE690A71AD5185FF1F1049">L. P. Kolcsár</collectorName>
, T.
<collectorName id="EAA0D7DE770253AADCB4E907567391DA">Kovács, D</collectorName>
.
<collectorName id="64C5D04E91223D77C47D23B65C9C6C6F">Murányi</collectorName>
:
<specimenCount id="8F1ABABE2B5A77951F197428329FFA89" count="1" type="male">1 ♂</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EA2AE" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
<collectionCode id="B536B21E42D10E79A401A74F4440B896">EKCU</collectionCode>
</abbrev>
:
<specimenCode id="0C2E63659E56F20955F8EF457B8BB18B" collectionCode="EKCU">PLP 5727</specimenCode>
)
</materialsCitation>
<materialsCitation id="4C017F62036A38E54A0185AE217F8C82" collectingDate="2023-05-06" collectionCode="MM" collectorName="L. P. Kolcsar &amp; Kovacs, D &amp; Muranyi" country="Serbia" elevation="1575" latitude="43.2743" location="Radaljica" longLatPrecision="6" longitude="20.3467" municipality="Golija Mts" specimenCode="2023-52, PLETYP-35" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Raska district, Novi Pazar" typeStatus="paratype">
<specimenCount id="BA45F9300D965AC699A3DE0F16C364AA" count="2" type="male">2 ♂</specimenCount>
<specimenCount id="0B203E10DDFC6AC1413979650DEB591B" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="BA122A6587E8FB0544A036B0759C9F72" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="27A47AE51D07F8FE09984BE0D23BDFAD" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="7FA840A7E2BD87C8AAAF272C967355D1">2023-52</specimenCode>
,
<specimenCode id="62549CC6E41AEFD6CCBE2296C790E897">PLETYP-35</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK2AE" type="diagnosis">
<paragraph id="02347B7FA6135566979AC7AAB6C44DAA">
<heading id="642592E2C9C8578CBFC8757B5D69A83C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D027438E5E9B5A3CE6B5C125B29B6E14">Macropterous in both sexes. Male tergite VII mostly membranous, with bicoloured antecosta; tergite VIII with converging pair of slender, pale brown processes, medial membranous area rounded between the processes, triangular between the processes and the lateral sclerotised portions; tergite IX with small posteromedial sclerite, divided into leaf-like portions; tergite X posterior margin with wide and deep notch; epiproct sclerotised only at its sides, stalk nearly as long as the rounded apex; sternite IX bears a vesicle 1 / 2 as long as segment length; specillum slightly longer than paraproct, tip of paraproct acute, tip of specillum blunt. Female subgenital plate large and bicoloured, lobes large, triangular, and not converging, slightly raised in lateral view and the notch between them is triangular; spermathecal sclerite ring-shaped, with large and converging posterior teeth.</paragraph>
</subSubSection>
<subSubSection id="SECID0EP2AE" type="description">
<paragraph id="50EC6BCD699FD95AA9E2721744E602FB">
<heading id="29A1A976EC6C78C8225CF18B2CF76D71" reason="title">Description.</heading>
</paragraph>
<paragraph id="10D71D3254538CD1CC64420A4B31155D">
Medium sized, slender species, both sexes macropterous. Forewing length:
<typeStatus id="7FB46BCFF11FFD630D7DF4886326C0EA" type="holotype">holotype</typeStatus>
<quantity id="F95EE12A1E0343888711E0EACA070D8C" metricMagnitude="-3" metricUnit="m" metricValue="5.6" unit="mm" value="5.6">5.6 mm</quantity>
, male
<typeStatus id="05227C63546C0EF204DD157422CA776B" type="paratype">paratypes</typeStatus>
5.46.0 mm, female
<typeStatus id="7A47EB959A24DA14678DB1115B0B8DC0" type="paratype">paratypes</typeStatus>
6.67.0 mm; body length:
<typeStatus id="3C4B21E4BA9A02E61094BE64C07CB979" type="holotype">holotype</typeStatus>
<quantity id="9B70373978265956494F3374004FC2FF" metricMagnitude="-3" metricUnit="m" metricValue="5.4" unit="mm" value="5.4">5.4 mm</quantity>
, male
<typeStatus id="C572C7A7EBAF14B7ECAD37CA60107475" type="paratype">paratypes</typeStatus>
<quantity id="945769A2B6C517573DBBA18EDB7D1BB9" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.4" metricValueMin="5.2" unit="mm" value="5.8" valueMax="6.4" valueMin="5.2">5.26.4 mm</quantity>
, female
<typeStatus id="12898487A27FE2E87AD5F0EB7759C4A8" type="paratype">paratypes</typeStatus>
<quantity id="7B69C2A5FF5824DA0A8C38AA126A5925" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.2" metricValueMin="5.5" unit="mm" value="6.35" valueMax="7.2" valueMin="5.5">5.57.2 mm</quantity>
. Setation generally short and dense. General colour brown to dark brown. Head and antennae dark brown, palpi brown. Pronotum brown to dark brown, as wide as long or slightly wider, having rounded corners, rugosities distinct. Legs brown to dark brown. Wings brownish but hyaline, venation brown.
</paragraph>
<paragraph id="1B002C8FA1A854022E5E46E0B104D9D0">
Male abdomen (Figs
<figureCitation id="062653045C283DE3EBB6DDBDC2432999" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminalia of Leuctra golija sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure5" httpUri="https://binary.pensoft.net/fig/1175456">5 A D</figureCitation>
,
<figureCitation id="8512A7204352EBE1156582BAB6155795" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 B</figureCitation>
): Tergite I membranous medially, with two medial spots; terga II and III poorly sclerotised anteriorly. Transverse row of four pigmented spots distinct on terga II VII. Terga II III with widely divided antecosta, terga IV VII with entire antecosta, terga IV VI full sclerotised. Tergite VII mostly membranous, sclerotised only laterally and in the widened antecosta that is distinctly bicoloured: dark brown laterally and pale brown medially. Tergite VIII: antecosta medially divided nearly in the 1 / 2 width of the segment; paired processes originated from the end of antecosta and reaches the posterior end of the segment, pale brown coloured, converging posteriorly, very slender, and only slightly widened apically, raised in lateral view; the medial membranous area well limited, rounded between the processes and with two sclerotised spots, triangular between the processes and the lateral sclerotised portion of the segment. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 2 of its length; posteromedial sclerite small, divided into leaf-like portions. Anterior margin of tergite X bilobed anteriorly; posterior margin with deep and wide notch, as wide as 1 / 2 tergite width. Epiproct large, sclerotised only at its sides, stalk nearly as long as the rounded apex. Cercus simple, covered with long setae. Sterna II VIII simple. Sternite IX bears a long, tongue-shaped vesicle 1 / 2 as long as segment length; posterolateral portions of the sternum pale coloured and weakly sclerotised, connected to pale area around the base of vesicle. Paraproct with moderately wide base, narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards an acute tip. Base of paraproct connected to a subrectangular lateral expansion, having a small inner apical tip. Specillum longer than the paraproct, gently curved in lateral view and ending in a blunt tip.
</paragraph>
<caption id="F64D0EC180FC00DFDAAC289F55E11CA8" ID-DOI="10.3897/zookeys.1218.120744.figure5" ID-arpha="34A974ED-A90F-59CF-A2E5-6F8FB75AE32E" httpUri="https://binary.pensoft.net/fig/1175456" startId="F5">
<paragraph id="8CB4F9BB929DD2EAB8F9770CABFD4B49">
<label id="E1CB35B7584FEA36FFA6DC080B34A060">Figure 5.</label>
</paragraph>
<paragraph id="2F146C2945DCCA2825AC7F4F3357E85A">
Terminalia of
<taxonomicName id="7079FD7238CACA61BB1A1DAE7714861D" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="golija" status="sp. nov.">
<emphasis id="534EF2BC6A24BBD205044E914E8CC875" italics="true">Leuctra golija</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5E8D75652164709605E1F90791FE036" rank="species">sp. nov.</taxonomicNameLabel>
with setation omitted
<emphasis id="98D733D650C793E54A3263E66E154F9A" bold="true">A</emphasis>
male terminalia, dorsal view
<emphasis id="930ED327B3307A0C7C84AA2E748F7EB0" bold="true">B</emphasis>
same, lateral view
<emphasis id="1C10E60C6F00764DA78B97B60E5B9C25" bold="true">C</emphasis>
same, ventral view
<emphasis id="50DA9B75E65A16DF6C8F7AAF40741B8B" bold="true">D</emphasis>
male paraprocts and specillae, caudal view
<emphasis id="4F3A1E550B77404E764DADF4BD76DA57" bold="true">E</emphasis>
female terminalia, ventral view
<emphasis id="D68B99B7795E47D500F1BB461D449AC9" bold="true">F</emphasis>
same, lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8C22B25C830A3EFF92A92244289EB918">
Female abdomen (Figs
<figureCitation id="B8D88C763A3716DA82F4066ADC03D382" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminalia of Leuctra golija sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure5" httpUri="https://binary.pensoft.net/fig/1175456">5 E, F</figureCitation>
,
<figureCitation id="9E531088D5F9983606F1A931436C2F2C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 F</figureCitation>
): Terga I VIII with transverse row of four pigmented spots clearly visible; terga I VIII mostly membranous but with lateral sclerites; small medial sclerite is present on terga VII VIII, sometimes also on tergite II; tergite IX mostly, tergite X fully sclerotised. Sterna I VII simple, sterna II VII with one subrectangular median sclerite and two small anterior sclerites that are fused with median sclerite only on sternite VII. Subgenital plate of sternite VIII large, setation not distinctive; not fused with lateral sclerites but touching with, a small sclerotised portion is visible on the integument anterior to the plate, as well a wider, pale sclerite on the posterior edge of segment. The subgenital plate with two large and well-defined lobes, the plate is distinctly bicoloured: lateral sides and lobes dark brown, while the rest of the plate pale brown. The lobes are triangular, not swollen, slightly raised in lateral view, not converging in ventral view and overhang the segment end, the notch between them has wavy sides but the proximal part is triangular. The central plate is smooth, only very slightly and evenly bulging in lateral view. Sternite IX without anterior indentation but the anterior 1 / 3 is membranous. Paraproct, cercus and epiproct simple. Spermathecal sclerite thin, ring-shaped, with small anterior teeth and converging, longer posterior teeth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EM4AE" type="Affinities">
<paragraph id="351B5E5BCADC7E2E1AD46BE1DA1C4F50">
<heading id="4C4A28B062484E5041A581BF1876B91D" reason="title">Affinities.</heading>
</paragraph>
<paragraph id="991ED85FD35FF250528DBBC0E798E068">
The new species is a member of the narrowly defined
<taxonomicName id="94E7726A65D8D79579AE5E6B4E95FCCD" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="8BA81BF8A8AF4079E42322DED5B7FD7A" italics="true">hippopus</emphasis>
</taxonomicName>
group (
<taxonomicName id="3E512A1C7322006CAD39AF3DE53C9952" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="88651CF185B8B68571C44214C5D99C50" italics="true">hippopus</emphasis>
</taxonomicName>
subgroup, sensu, e. g.,
<bibRefCitation id="78956EE16A7FE78F01184E73497DB71F" author="Ravizza C" journalOrPublisher="Lauterbornia" pagination="1-42" refId="B69" refString="Ravizza C (2002 b) Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species. Lauterbornia 44: 142. https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" title="Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species." url="https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" volume="44">Ravizza 2002 b</bibRefCitation>
), and morphologically closest to the West Balkan endemic
<taxonomicName id="72944B4267FE0E3E7CE6E402EBB01777" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="5299C9F8E8FF9AD35E0AB3F62AFCB397" italics="true">L. hippopoides</emphasis>
</taxonomicName>
and the East Balkan endemic
<taxonomicName id="C77BC10D72D52A2B33EB5774B267BDC5" authority="Rauser, 1965" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudohippopus">
<emphasis id="F2B07345075BF5877D5B89DEE519594C" italics="true">L. pseudohippopus</emphasis>
<bibRefCitation id="30CB29AA5B86DC7CEB96ED7A813E1AB8" author="Raušer J" issue="92" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="125-138" refId="B67" refString="Rauser J (1965) Plecoptera bulgarica - II. Acta Faunistica Entomologica Musei Nationalis Pragae 10 (92): 125 - 138. https: // www. aemnp. eu / data / article- 113 / 94 - 10 _ 0 _ 125. pdf" title="Plecoptera bulgarica - II." url="https://www.aemnp.eu/data/article-113/94-10_0_125.pdf" volume="10" year="1965">Raušer, 1965</bibRefCitation>
</taxonomicName>
. The male can be distinguished from both on the basis of a nearly fully membranous tergite VII, convergent pair of processes of tergite VIII that delimit the rounded median membranous area, the membranous area triangular lateral to the processes, the small posteromedial sclerite of tergite IX divided into leaf-like portions, and the epiproct sclerotised only at its sides, but with long stalk. Both
<taxonomicName id="937A6AD05C76BD22C42279D86358E76F" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="2B300D86523A3C8DE2A7AC0EF181099C" italics="true">L. hippopoides</emphasis>
</taxonomicName>
and
<taxonomicName id="268E05F5D681A77F7486143D99C4FBD4" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudohippopus">
<emphasis id="E14423BF470BB85DD304474329439263" italics="true">L. pseudohippopus</emphasis>
</taxonomicName>
have a bell-shaped membranous area on tergite VII, the pair of processes not converging on tergite VIII and delimiting a square median membranous area, while the membranous area is rounded laterad to the processes, their epiproct more heavily sclerotised and with a short stalk; the posteromedial sclerite of tergite IX is divided into triangular portions in
<taxonomicName id="C632A89E8FA6728AB7E41E88EAFB4BDC" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudohippopus">
<emphasis id="C7986BCEC2D841F46FBEDF1074001A5F" italics="true">L. pseudohippopus</emphasis>
</taxonomicName>
, while the undivided sclerite is large and trapezoidal in
<taxonomicName id="B95F3AB29F6DF6CA5B078E5783D317C0" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="44E261AE37E8CCA209E92B3A965F438B" italics="true">L. hippopoides</emphasis>
</taxonomicName>
. The island endemic
<taxonomicName id="B9F43B99646F70A411405EF8FA942CE1" authority="Vincon, 2015" authorityName="Vincon" authorityYear="2015" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pavesii">
<emphasis id="6A806A83475E0934B1436DE73D679AA0" italics="true">L. pavesii</emphasis>
<bibRefCitation id="1FAE49F09216489F7A165FFD757B1679" author="Vinçon G" issue="1-2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="73-76" refId="B81" refString="Vincon G (2015) A new stonefly from Greece, Leuctra pavesii sp. n. (Plecoptera: Leuctridae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 88 (1 - 2): 73 - 76. https: // www. e-periodica. ch / digbib / view? pid = seg- 001 % 3 A 2015 % 3 A 88 % 3 A % 3 A 6 # 82" title="A new stonefly from Greece, Leuctra pavesii sp. n. (Plecoptera: Leuctridae)." url="https://www.e-periodica.ch/digbib/view?pid=seg-001%3A2015%3A88%3A%3A6#82" volume="88" year="2015">Vinçon, 2015</bibRefCitation>
</taxonomicName>
from Cephalonia,
<collectingCountry id="EA859F9CF1783FFA0A896B07F7DA9BD8" name="Greece">Greece</collectingCountry>
, resembling
<taxonomicName id="2AAB37624133997E488E266EE98BEE96" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="AB99D0A3D1E7225DA5C4B916D167E6A6" italics="true">L. hippopoides</emphasis>
</taxonomicName>
but differs by having a straight specillum in lateral view, contrary to the gentle curve for males of all other Balkan species of the species group. The widespread
<taxonomicName id="A74A14875024313EA5AD16F5624F4F95" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="38DDD4766C0117B5CC17BA30D8808FC0" italics="true">L. hippopus</emphasis>
<bibRefCitation id="4C246B5EA30F497FCBF002BCBCA8F514" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
</taxonomicName>
, co-occurring with the new species, is morphologically less related, and the male can easily be distinguished by its mostly sclerotised tergite VII, robust paired processes of tergite VIII, large and complex posteromedial sclerite of tergite IX, and heavily sclerotised epiproct with a short stalk. The female of the new species distinctly differs from all members of the narrowly defined
<taxonomicName id="9593F8349BE39956BF6B007145C16787" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="6EE16B61964EFAD53619CB3977AAAE9C" italics="true">hippopus</emphasis>
</taxonomicName>
group by having triangular lobes of the subgenital plate, contrary to the bulging, rounded, square, or finger-like lobes of the other species. The subgenital plate most resembles the not closely related
<taxonomicName id="A2F48694C23A4A5F6159149874D751F1" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="C347B93B7D460C4D0F40A4FB2DEA7269" italics="true">L. armata</emphasis>
<bibRefCitation id="F42250CE7B21E2E9A5F447D335ED06DA" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
</taxonomicName>
, which has a small central lobule between the base of the lobes.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDCAG" type="distribution">
<paragraph id="D70EC6BD3B2522E6AA6301572C6B10E9">
<heading id="F7D8F177B96B6829E8914D2C77EA43A0" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="7CA86476469D98BD2F64C3FC75B83E13">
The species was collected at two high elevation localities of the Golija Mts of southwestern
<collectingCountry id="A3EF61DF88CD82AF578C08F0CE515090" name="Serbia">Serbia</collectingCountry>
(Fig.
<figureCitation id="7F662EA024DF6D30AA949D1A11936CA5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">1</figureCitation>
). The Golija is a south-eastern range of the Dinarids, a mostly volcanic area with a large plateau, its highest peak reaching more than 1800 meters. Most of the specimens were found at the Moravica Stream, 12 meters wide, &lt;
<quantity id="FDD00CA8C9A20EE160FB08DE9650BF2E" metricMagnitude="-1" metricUnit="m" metricValue="5.0" unit="m" value="0.5">0.5 meter</quantity>
deep, running in alder forest (Fig.
<figureCitation id="7A7A3CB4C3E517C544EB2997915D32F6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitat of Leuctra golija sp. nov.: Moravica Stream in the Golija Mts (type locality, 23. x. 2023)." figureDoi="10.3897/zookeys.1218.120744.figure6" httpUri="https://binary.pensoft.net/fig/1175457">6</figureCitation>
). The other habitat, where only a few specimens were found, is a spring brook that belongs to the watershed of the Ibar River, being rather steep and only
<quantity id="8DD15306738DBD32555A96457B168668" metricMagnitude="-1" metricUnit="m" metricValue="5.0" unit="m" value="0.5">0.5 meter</quantity>
wide with a few centimetres depth, running at the edge of a beech forest. Both streams have stony substrates mixed with gravel and small sandy patches. We visited the localities three times (
<bibRefCitation id="DF3BCAEC1C8E1DE9D38802F3860A3B9C" author="Muranyi and Kovacs" firstAuthor="Muranyi" refId="B49" refString="Murányi D, Kovács T (2024) Contribution to the Plecoptera fauna of Serbia. Folia historico-naturalia Musei Matraensis 48. [in press]" year="2024">Murányi and Kovács 2024</bibRefCitation>
): the new species was swarming in early May (
<date id="8AC661D9AC873C9CCA573121DA27BEF9" value="2023-05-06">6. v. 2023</date>
, leg. L. P. Kolcsár, T. Kovács, D. Murányi), only a few individuals were found in late June (
<date id="75E7E27CC97B1F75321C948751E310C1" value="2018-06-26">26. vi. 2018</date>
, leg. P. Juhász, T. Kovács, D. Murányi), while no specimens were present in October (
<date id="90EC478DF752ED0286B6115F89FE4AD2" value="2023-10-23">23. x. 2023</date>
, leg. T. Kovács, D. Murányi, D. Pifkó). At the Moravica Stream, the following species were found to be cohabiting:
<taxonomicName id="B520ADDBE701F7B278B53A87A906A59E" authority="Rauser, 1962" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="bulgarica">
<emphasis id="E3EF6A14748446ACE4FF01BBF0986F6E" italics="true">Brachyptera bulgarica</emphasis>
<bibRefCitation id="BFEDA1F0BE490E68CBD99840931991F0" author="Raušer J" issue="70" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="67-82" refId="B66" refString="Rauser J (1962) Plecoptera bulgarica - I. Acta Faunistica Entomologica Musei Nationalis Pragae 8 (70): 67 - 82. https: // www. aemnp. eu / data / article- 91 / 72 - 8 _ 0 _ 67. pdf" title="Plecoptera bulgarica - I." url="https://www.aemnp.eu/data/article-91/72-8_0_67.pdf" volume="8" year="1962">Raušer, 1962</bibRefCitation>
</taxonomicName>
(adults and larvae, May),
<taxonomicName id="C0673E8FCCF5EBD59B75EC472DFF68B7" authority="(Klapalek, 1902)" baseAuthorityName="Klapalek" baseAuthorityYear="1902" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="428854BCBC1C01580F30FDB645F3FB63" italics="true">B. seticornis</emphasis>
(
<bibRefCitation id="88057916D75806C6B1990AD81F4B3FCB" author="Klapálek F" issue="1-2" journalOrPublisher="Természetrajzi Füzetek" pagination="161-180" refId="B37" refString="Klapalek F (1902) Zur kenntniss der Neuropteroiden von Ungarn, Bosnien und Herzegovina. Termeszetrajzi Fuzetek 25 (1 - 2): 161 - 180. https: // epa. oszk. hu / 02300 / 02370 / 00061 / pdf / EPA 02370 _ termszetrajzif 25 magy _ 01 - 02 _ 161 - 180. pdf" title="Zur kenntniss der Neuropteroiden von Ungarn, Bosnien und Herzegovina." url="https://epa.oszk.hu/02300/02370/00061/pdf/EPA02370_termszetrajzif25magy_01-02_161-180.pdf" volume="25" year="1902">Klapálek, 1902</bibRefCitation>
)
</taxonomicName>
(adults, larvae, and exuviae, May),
<taxonomicName id="4A24BF03E0D3AE6B15814F5B51132BC9" authorityName="Pictet" authorityYear="1841" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="D5709A985747F1E0B2A3A98D48377BDB" italics="true">Capnia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8EEF1A651E2EDFC87AB05DC9246D4D4" sensu="lato">s. l.</taxonomicNameLabel>
<emphasis id="61CEE48067D58CE993A773567F6ABE70" italics="true">vidua rilensis</emphasis>
<bibRefCitation id="FDB301AC2D043121688D10E18CE098E7" author="Raušer J" issue="70" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="67-82" refId="B66" refString="Rauser J (1962) Plecoptera bulgarica - I. Acta Faunistica Entomologica Musei Nationalis Pragae 8 (70): 67 - 82. https: // www. aemnp. eu / data / article- 91 / 72 - 8 _ 0 _ 67. pdf" title="Plecoptera bulgarica - I." url="https://www.aemnp.eu/data/article-91/72-8_0_67.pdf" volume="8" year="1962">Raušer, 1962</bibRefCitation>
(adults, May),
<taxonomicName id="E1349982D2C37FC7750C36E9A2F80E85" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="4874259698533B4AE0D6A3AE3C371BBA" italics="true">Leuctra nigra</emphasis>
</taxonomicName>
(adults, May and June),
<taxonomicName id="CA5761CE650EACBDA59A8C86D7E81616" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="cingulata">
<emphasis id="0026BA516D643F2FE7ACA375EC0DBD64" italics="true">L. cingulata</emphasis>
<bibRefCitation id="2BC5C5C17BE6AFC0CC7C9C10A6EC9E71" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
</taxonomicName>
(adults, October),
<taxonomicName id="0FF8DE9DEA28EF5ED28B707D30B57F93" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="8FE2194EC43DD6A1FF0018438E44B8EA" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(adults, May) (Fig.
<figureCitation id="F3DC33865906172C6B177D289F7DBDFA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 D, H</figureCitation>
),
<taxonomicName id="53232A16A7D69A8CEB694B781E55F8A4" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="F1523A794094F21058790F8AF5294823" italics="true">L. prima</emphasis>
<bibRefCitation id="3311FD040E7B12C9B635CDECCB9ACB3D" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
</taxonomicName>
(adults, May),
<taxonomicName id="19D2B50A695E7C237ABC73EF19A4F5DC" authority="Rauser, 1965" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="F5E8D047A91BA8930C91A978C1DCEA9B" italics="true">L.</emphasis>
cf.
<emphasis id="BB7B8A998E70D4AB5FBA806E1551AA8F" italics="true">balcanica</emphasis>
<bibRefCitation id="043882971B27C525BD313B6F111E3709" author="Raušer J" issue="92" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="125-138" refId="B67" refString="Rauser J (1965) Plecoptera bulgarica - II. Acta Faunistica Entomologica Musei Nationalis Pragae 10 (92): 125 - 138. https: // www. aemnp. eu / data / article- 113 / 94 - 10 _ 0 _ 125. pdf" title="Plecoptera bulgarica - II." url="https://www.aemnp.eu/data/article-113/94-10_0_125.pdf" volume="10" year="1965">Raušer, 1965</bibRefCitation>
</taxonomicName>
(adults, June),
<taxonomicName id="25400F769371D773BBB48737E2B81C90" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
<emphasis id="F4AAB003F1AC274B5012CF18CBDE42A6" italics="true">Nemoura cinerea cinerea</emphasis>
</taxonomicName>
(adults, June),
<taxonomicName id="CF1EC3ED9A5FB2D104364719AEA8C2EF" authority="Despax, 1934" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="3BD605ED749FBE33BD4D68D558606109" italics="true">N. uncinata</emphasis>
<bibRefCitation id="9417005F07BE116DCC9ADCCF3F69EDD8" author="Despax R" issue="2" journalOrPublisher="Bulletin de la Société dhistoire Naturelle de Toulouse" pagination="255-270" refId="B15" refString="Despax R (1934) Plecopteres pyreneens. VII. Etude et description de quelques formes de Nemoures apparentees a Nemoura marginata (Pict.). Bulletin de la Societe d'histoire Naturelle de Toulouse 66 (2): 255 - 270." title="Plécoptères pyrénéens. VII. Etude et description de quelques formes de Nemoures apparentées à Nemoura marginata (Pict.)." volume="66" year="1934">Despax, 1934</bibRefCitation>
</taxonomicName>
(adults, May),
<taxonomicName id="8524DB152E8324C29A83219DE986AD90" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="80E22CE782F3E76B6889B06CE59D5BE9" italics="true">N. marginata</emphasis>
</taxonomicName>
(adults, June), an undescribed
<taxonomicName id="CCC0AA41D1E3D1379EA86D7A664783CC">
<emphasis id="F9792497FC5A38FFFC847DB080118E04" italics="true">Nemoura</emphasis>
species
</taxonomicName>
of the
<taxonomicName id="0CA821A2F24D209C227B98C1DFAD37F0" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="343B9210DF702850CCB884611C9320DB" italics="true">marginata</emphasis>
</taxonomicName>
group (adults, May), an undescribed
<taxonomicName id="9E63882C19B5A4ACDADB2EDE0B089EF9" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F22CD548913D5C86B512461D2E00E66C" italics="true">Nemoura</emphasis>
</taxonomicName>
related to
<taxonomicName id="C8E47643950852D205003A18B73336D8" authority="Rauser, 1962" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="bulgarica">
<emphasis id="8DC3B3227B9DD04DCCF45BC95851B751" italics="true">N. bulgarica</emphasis>
<bibRefCitation id="0BEC9813E6E53AE6424901B6E96B8A28" author="Raušer J" issue="70" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="67-82" refId="B66" refString="Rauser J (1962) Plecoptera bulgarica - I. Acta Faunistica Entomologica Musei Nationalis Pragae 8 (70): 67 - 82. https: // www. aemnp. eu / data / article- 91 / 72 - 8 _ 0 _ 67. pdf" title="Plecoptera bulgarica - I." url="https://www.aemnp.eu/data/article-91/72-8_0_67.pdf" volume="8" year="1962">Raušer, 1962</bibRefCitation>
</taxonomicName>
(adults, June),
<taxonomicName id="46D0AE72122E2C4F4B2DCAD5419D987F" authority="(Morton, 1894)" baseAuthorityName="Morton" baseAuthorityYear="1894" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="praecox" subSpecies="praecox">
<emphasis id="CA44A16E12A6D40888197B579E906E10" italics="true">Protonemura praecox praecox</emphasis>
(
<bibRefCitation id="5E9EA40473F0C39A0D0F54CFA241343B" author="Morton KJ" refId="B43" refString="Morton KJ (1894) Palaearctic Nemourae. Transactions of the Entomological Society of London 1894 (23): 557 - 574 [Pl. XIII - XIV]. https: // www. biodiversitylibrary. org / item / 50996 # page / 599 / mode / 1 up" url="https://www.biodiversitylibrary.org/item/50996#page/599/mode/1up" year="1894">Morton, 1894</bibRefCitation>
)
</taxonomicName>
(adults, May),
<taxonomicName id="05175FD3A5A7F093407E6169C2610BFD" authority="(Ris, 1902)" baseAuthorityName="Ris" baseAuthorityYear="1902" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="intricata" subSpecies="intricata">
<emphasis id="CD3FA1259CC1EFBC7CDB7BA23C2CF4C5" italics="true">P. intricata intricata</emphasis>
(Ris, 1902)
</taxonomicName>
(adults, June),
<taxonomicName id="BA20C183D57A1D3C8E197A8E1393B5A6" authority="Rauser, 1956" authorityName="Rauser" authorityYear="1956" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hrabei">
<emphasis id="9FA08317122BA6D9321E8B6412399CE4" italics="true">P. hrabei</emphasis>
<bibRefCitation id="FEDE1133370DFE0064797E286493AD0B" author="Raušer J" refId="B65" refString="Rauser J (1956) K poznani ceskoslovenskych larev rodu Protonemura. Acta Academiae Scientiarum Cechoslovenicae Basis Brunensis 28 (9): 449 - 498." year="1956">Raušer, 1956</bibRefCitation>
</taxonomicName>
(adults, October),
<taxonomicName id="A8AD160DC53C3760D717C214504ED9B0" authority="(Pictet, 1836)" baseAuthorityName="Pictet" baseAuthorityYear="1836" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E75BD7BDA02510E6ECA422A6BCD424BE" italics="true">P. nitida</emphasis>
(
<bibRefCitation id="E2E98F4907C91A64995CB08A42D0C2A1" author="Pictet FJ" issue="1" journalOrPublisher="Memoires de la Société de physique et dhistoire naturelle de Genève" pagination="173-190" refId="B61" refString="Pictet FJ (1836) Description de quelques nouvelles especes d'Insectes du Bassin du Leman. Memoires de la Societe de physique et d'histoire naturelle de Geneve 7 (1): 173 - 190. https: // www. biodiversitylibrary. org / item / 39755 # page / 201 / mode / 1 up" title="Description de quelques nouvelles espéces dInsectes du Bassin du Léman." url="https://www.biodiversitylibrary.org/item/39755#page/201/mode/1up" volume="7" year="1836">Pictet, 1836</bibRefCitation>
)
</taxonomicName>
(adults, October),
<taxonomicName id="7CF5E39B09477733BAA62FC1735A4EDF" authorityName="Guerin-Meneville" authorityYear="1843" class="Insecta" family="Perlidae" genus="Perla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="E6C2F7A209EE856E8647F9554154323C" italics="true">Perla</emphasis>
cf.
<emphasis id="B860D3DB22C2435EF5B2F55F1B904F6C" italics="true">pallida</emphasis>
</taxonomicName>
(larvae, May to October),
<taxonomicName id="00E2BB3EB73F064B06A7FD5D50765F31" authority="Sowa, 1970" authorityName="Sowa" authorityYear="1970" class="Insecta" family="Perlodidae" genus="Isoperla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="russevi">
<emphasis id="9E22F30CA1050E725C8AED0A0DE4EADE" italics="true">Isoperla</emphasis>
cf.
<emphasis id="52C6B53379187D7A6D3634B1E49AFFB1" italics="true">russevi</emphasis>
<bibRefCitation id="3343B44A99340D475BEF8D6421A11761" author="Sowa R" issue="12" journalOrPublisher="Bulletin de lAcadémie Polonaise des Sciences" pagination="767-772" refId="B77" refString="Sowa R (1970) Deux Plecopteres nouveaux de Bulgarie. Bulletin de l'Academie Polonaise des Sciences 18 (12): 767 - 772." title="Deux Plécoptères nouveaux de Bulgarie." volume="18" year="1970">Sowa, 1970</bibRefCitation>
</taxonomicName>
(adults, June),
<taxonomicName id="3F8B8630DE93467EDFF94EAE36ED07C0" authority="Braasch, 1969" authorityName="Braasch" authorityYear="1969" class="Insecta" family="Chloroperlidae" genus="Chloroperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="russevi">
<emphasis id="82A59463CAE00905D42893EAE654577D" italics="true">Chloroperla russevi</emphasis>
<bibRefCitation id="746B9538E836009099B62B642B3FB424" author="Braasch D" refId="B9" refString="Braasch D (1969) Chloroperla russevi n. sp. und Chloroperla kosarovi n. sp. aus Bulgarien. Mitteilungen der Deutschen Entomologischen Gesellschaft 28 (5 / 6): 51 - 54." year="1969">Braasch, 1969</bibRefCitation>
</taxonomicName>
(adults, June) and
<taxonomicName id="104EE4BBE725C2EC1BFFC65A1D3FEB6F" baseAuthorityName="Rostock" baseAuthorityYear="1881" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="0565B5F5B9D43559DC1B7B7B7F7BFA90" italics="true">Siphonoperla neglecta</emphasis>
</taxonomicName>
(adults, June). At the spring brook, a different set of species were collected:
<taxonomicName id="119355AAAAEA7C5E688917EACA8EB385" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="bulgarica">
<emphasis id="43C2F644995193DF216C698C35198610" italics="true">Brachyptera bulgarica</emphasis>
</taxonomicName>
(adults, May),
<taxonomicName id="0DAED3F7A035E2BCC84EBAEFD0E87195" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1C6875A169CCF13B55DBA962C5AA3986" italics="true">Leuctra nigra</emphasis>
</taxonomicName>
(adults, May and June),
<taxonomicName id="5B04A4F9D09986B9AA0D47B11EA9011F" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="fusca" subSpecies="fusca">
<emphasis id="D5CE53AA984D6CAD27FEAC7AFC4EE8B6" italics="true">L. fusca fusca</emphasis>
(
<bibRefCitation id="442D492F0BBCA41E9C89B697F48A71EA" DOI="10.5962/bhl.title.542" author="Linnaeus C" journalOrPublisher="Laurentii Salvii, Holmi" refId="B41" refString="Linnaeus C (1758) Systema naturae per regna tria natur, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmi, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" title="Systema naturae per regna tria natur, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata." year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
(adults, October),
<taxonomicName id="4BD599C9612E290078521D96B0E0559B" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="cingulata">
<emphasis id="579B9AE07D238B2B71F6DD76DA06CA30" italics="true">L. cingulata</emphasis>
</taxonomicName>
(adults, October),
<taxonomicName id="6BA637809B11EA7C29D9BB7713E25FF8" authorityName="Bogoesco &amp; Tabacaru" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="BF1FD7CBDA4840FBD16B28DEC15DCC20" italics="true">L. hirsuta</emphasis>
</taxonomicName>
(adults, October),
<taxonomicName id="0F62FB541AB73E16D7C42BB98498DAE0" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="02E57E01FB21AC04C04D35032684B835" italics="true">L. hippopus</emphasis>
</taxonomicName>
(adults, May),
<taxonomicName id="6EDBFFF64C28AFEFF923B9075BC83114" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="144C3185827A2BC9822B113B060AB710" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(adults, May),
<taxonomicName id="EFD280EC517D2F05E00DFF9F264C2BAF" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B0142D254FA4694941503DD03631D676" italics="true">L. prima</emphasis>
</taxonomicName>
(adults, May),
<taxonomicName id="A2629EF0A5C83D603BE796D721C741B8" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="F58AE68F3B82CC979A0F809869AB6B17" italics="true">L.</emphasis>
cf.
<emphasis id="D8805AC501058D99759CA1AF024E6639" italics="true">balcanica</emphasis>
</taxonomicName>
(adults, June),
<taxonomicName id="6AC08007421F5A9B89CD76C757EBA4BB" authority="(Ris, 1902)" baseAuthorityName="Ris" baseAuthorityYear="1902" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="standfussi">
<emphasis id="665F8E4AA6EBADA935C0B9EA5744A575" italics="true">Amphinemura standfussi</emphasis>
(Ris, 1902)
</taxonomicName>
(adults, June),
<taxonomicName id="71BB40F06CD21E02B6E57FC0D016E385" authorityName="Rauser" authorityYear="1956" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hrabei">
<emphasis id="A42995AA8C470B0ED85DE21A42DD55AF" italics="true">Protonemura hrabei</emphasis>
</taxonomicName>
(adults, October), an undescribed
<taxonomicName id="D376ECCFA6891027DFDA79E38585EEBA">
<emphasis id="37555F10710BAC98F7711876DA40B724" italics="true">Protonemura</emphasis>
species
</taxonomicName>
of the
<taxonomicName id="163CBB53319040A96E4841693C81E5C0" authorityName="Illies" authorityYear="1954" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="auberti">
<emphasis id="5ACCAE1575CD275E96BB5E7530BBCAEB" italics="true">auberti</emphasis>
</taxonomicName>
group (adults in October, larvae in May and June),
<taxonomicName id="F4C5439B97635221689AC683D6FC731C" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
<emphasis id="581B26BC575D80CBEE216DBF195EAE4E" italics="true">Nemoura cinerea cinerea</emphasis>
</taxonomicName>
(adults, June),
<taxonomicName id="19388709AB3534382FB36FDA37A49C3C" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="350B5CF9388A8CCD7DFD93FD70A26EB3" italics="true">N. marginata</emphasis>
</taxonomicName>
(adults, June), the undescribed
<taxonomicName id="0A2075C54665C6CB5997FBCB42EE187A">
<emphasis id="5846275C0BDDDF4A00EC06EB92BA3E9A" italics="true">Nemoura</emphasis>
species
</taxonomicName>
of the
<taxonomicName id="B87A9EED63B408BB6BA3253F999B1D33" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="1FFCDC165AE49077D96874D16706F77D" italics="true">marginata</emphasis>
</taxonomicName>
group (adults, May), the undescribed
<taxonomicName id="DC08444756FB9F3F42E9C048DAB2879C" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="A6A8E0835A6C039977AC5E1D666D4CC3" italics="true">Nemoura</emphasis>
</taxonomicName>
related to
<taxonomicName id="B4FDE7915E622F093ACA323BCF0A551E" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="bulgarica">
<emphasis id="7B0893ED78A647FAF85397B0E5673BA5" italics="true">N. bulgarica</emphasis>
</taxonomicName>
(adults, June),
<taxonomicName id="587C4034D59419B2271202FAD0C99032" authorityName="Guerin-Meneville" authorityYear="1843" class="Insecta" family="Perlidae" genus="Perla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="7679A555DE094402A13AAEF058483A30" italics="true">Perla</emphasis>
cf.
<emphasis id="CC9123BAF57FFE6C35E7B6C07B65CDA3" italics="true">pallida</emphasis>
</taxonomicName>
(larvae, October),
<taxonomicName id="2DF045F271E329A70895D320D13B32F7" authorityName="Sowa" authorityYear="1970" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="russevi">
<emphasis id="D97228C2E10915C8DCC7A2438E258B50" italics="true">Isoperla russevi</emphasis>
</taxonomicName>
(adults, June) and an unidentified
<taxonomicName id="E3E10D27DC0C48D9C6AC69DC68930BE7" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="19BBEC5CA1B6000344A8DA009CA8C0EB" italics="true">Isoperla</emphasis>
</taxonomicName>
(adult female, June).
</paragraph>
<caption id="0A9745E96590CF02CE80AE81FC0B2555" ID-DOI="10.3897/zookeys.1218.120744.figure6" ID-arpha="554A4F53-10A8-513A-98BA-B506657D2C61" httpUri="https://binary.pensoft.net/fig/1175457" startId="F6">
<paragraph id="2DEDEB9EED9D35D90D709052B0F337DA">
<label id="795F7CA77FC4262A1F9EB956AB6E9C5F">Figure 6.</label>
</paragraph>
<paragraph id="FD9937F924B0D13826FFBE8A45E375D2">
Habitat of
<taxonomicName id="B372B18A61E152482E4E8A48EC962C8E" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="golija" status="sp. nov.">
<emphasis id="FF0F6E653C5831427F42342A31DF61B3" italics="true">Leuctra golija</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9C141482EF4E4FBA88F11850F90BA504" rank="species">sp. nov.</taxonomicNameLabel>
: Moravica Stream in the Golija Mts (type locality, 23. x. 2023).
</paragraph>
</caption>
<caption id="8F4F66EE739C7EF64DBD67B57111F97E" ID-DOI="10.3897/zookeys.1218.120744.figure7" ID-arpha="21292C49-853C-56E7-AEDB-85FF30F331EF" httpUri="https://binary.pensoft.net/fig/1175458" startId="F7">
<paragraph id="61F3E053AACDD1E412524A7EED0FFA36">
<label id="3BAF2CF48131CC19E1EECB3F4AB86DA4">Figure 7.</label>
</paragraph>
<paragraph id="690C4CE502ECDFF49FFAABAF22585D94">
Terminalia of four Balkan members of the
<taxonomicName id="88A1E05237C5D511193102AE34636299" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="7F27F74355E2FF330BCDA476A42982E0" italics="true">Leuctra hippopus</emphasis>
</taxonomicName>
group
<emphasis id="E62943A6B7E78ACA402F728145DF2399" bold="true">A D</emphasis>
male terminalia, dorsal view
<emphasis id="82236B43222BAEAD9690011AEF4ABB4F" bold="true">E H</emphasis>
female terminalia, ventral view.
<emphasis id="57E194651334C9FA563D7D709E032931" bold="true">A, E</emphasis>
<taxonomicName id="6BC8C67B0FD3C3F9FBB206229F539F39" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="2086CD6D677EC270175914972C932D36" italics="true">L. enigma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DFA7E9102CF35C6FFE667576B109BA5" rank="species">sp. nov.</taxonomicNameLabel>
, Çermenikë Mts, Albania
<emphasis id="FFDEA1DE895EFCF3F81EC3CAB9FC40DB" bold="true">B, F</emphasis>
<taxonomicName id="20CC44E2BA8BFA583F68E187F6A022B2" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="golija" status="sp. nov.">
<emphasis id="529CB32CD7450094AC8071B02ABB2B66" italics="true">L. golija</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E62DF13173B002EA8507764BF7BC759" rank="species">sp. nov.</taxonomicNameLabel>
, Golija Mts, Serbia
<emphasis id="C8DF253314FEFB453076613092E7A7BD" bold="true">C, G</emphasis>
<taxonomicName id="5798C5BBFCFF55935270C15CFD3D224C" authority="Rauser, 1965" authorityName="Rauser" authorityYear="1965" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudohippopus">
<emphasis id="234DDB1884451FDC26D4161FFC8527B6" italics="true">L. pseudohippopus</emphasis>
<bibRefCitation id="737105FFFB3C85BFE6D8A9DD874159D1" author="Raušer J" issue="92" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="125-138" refId="B67" refString="Rauser J (1965) Plecoptera bulgarica - II. Acta Faunistica Entomologica Musei Nationalis Pragae 10 (92): 125 - 138. https: // www. aemnp. eu / data / article- 113 / 94 - 10 _ 0 _ 125. pdf" title="Plecoptera bulgarica - II." url="https://www.aemnp.eu/data/article-113/94-10_0_125.pdf" volume="10" year="1965">Raušer, 1965</bibRefCitation>
</taxonomicName>
, Stara Planina, Serbia
<emphasis id="88467D8E264986BC819A5164DBC421AE" bold="true">D, H</emphasis>
<taxonomicName id="14760664E3558886AD09A90B0B8B2642" authority="Kacanski &amp; Zwick, 1970" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="DFBDBDF5D2BDB2BFE2E42E8F9EE0B97A" italics="true">L. hippopoides</emphasis>
<bibRefCitation id="EB586B41D3BC30126A1E6DB983A44E2E" author="Kaćanski D &amp; Zwick P" issue="1" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="1-16" refId="B30" refString="Kacanski D, Zwick P (1970) Neue und wenig bekannte Plecopteren aus Jugoslawien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 43 (1): 1 - 16. https: // www. e-periodica. ch / cntmng? pid = seg- 001 % 3 A 1970 % 3 A 43 % 3 A % 3 A 14" title="Neue und wenig bekannte Plecopteren aus Jugoslawien." url="https://www.e-periodica.ch/cntmng?pid=seg-001%3A1970%3A43%3A%3A14" volume="43" year="1970">Kaćanski &amp; Zwick, 1970</bibRefCitation>
</taxonomicName>
, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (
<emphasis id="6AA66A51CA70F5EB3F2B53A7AC8FFEF2" bold="true">D</emphasis>
for
<emphasis id="478D3CCE07E62479DC4520AFB1180544" bold="true">A D</emphasis>
,
<emphasis id="2FA5CC744EDABBFC38CF8B480B369B1A" bold="true">H</emphasis>
for
<emphasis id="E616AE1C67995262E7668B6991F7AEC5" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBWAG" type="etymology">
<paragraph id="2D07D121FCC07F703F9CFF3F59D4F484">
<heading id="3567847C7DFACD8D0F28C58E902E0421" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="2788EC3AEE344B23E4BE37AFF7F31E8E">
The name
<emphasis id="A85E82C2F618472933A257AFF7A2C96B" italics="true">golija</emphasis>
is derived from the Golija Mountains of
<collectingCountry id="804A4A965C3677BDEA60071D3DC88436" name="Serbia">Serbia</collectingCountry>
, where the new species was found and is probably restricted to these mountains and the nearby ranges. Used as a noun, gender neutral.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F42684CED5C427132B257192BAB18C80">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:number id="AEE386F99FF5F6DC30E08838879EDEDE">1218</mods:number>
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<mods:extent id="52AE17462B93F38B559056D99B936179" unit="page">
<mods:start id="34BE0394F136F0CDE8DA0092328764DC">49</mods:start>
<mods:end id="227EA58B02D22394151D52CA6B8D13F9">79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="631350A34C096DFEFD586B4113C9FC56">journal article</mods:classification>
<mods:identifier id="D36BD12119C3D5FC24A00FF0366C9017" type="DOI">10.3897/zookeys.1218.120744</mods:identifier>
<mods:identifier id="A9B24A08CF8E454C4E1AE38C97C0EC07" type="ZooBank">094646BE-BF9F-4890-BD59-FF26FA95A3FE</mods:identifier>
</mods:mods>
<treatment id="6EAE8429DA2E58F48A813DD7F30E156A" ID-arpha="6EAE8429-DA2E-58F4-8A81-3DD7F30E156A" ID-zoobank="https://zoobank.org/F0FF7DC4-7EA8-4BDE-9042-34D956A63609" LSID="urn:lsid:zoobank.org:act:F0FF7DC4-7EA8-4BDE-9042-34D956A63609" LSID-ZBK="urn:lsid:zoobank.org:act:F0FF7DC4-7EA8-4BDE-9042-34D956A63609" httpUri="http://treatment.plazi.org/id/6EAE8429DA2E58F48A813DD7F30E156A">
<subSubSection id="32CF0813C499F1BD6A0B9CB12F1E03CB" type="nomenclature">
<paragraph id="7C5BCD1E5AB54FC8A17EC8F8A99FE788">
<taxonomicName id="3FF8E843145769EDEB98A2286FF0055A" authority="Kovacs &amp; Muranyi" authorityName="Kovacs &amp; Muranyi" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="FA2DC3D910A24176CFA3B7CA1E017EAF" italics="true">Leuctra enigma</emphasis>
Kovács &amp; Murányi
</taxonomicName>
<taxonomicNameLabel id="B8FB01E5CD52087A1CB7AD1E202B9B60" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6A6C55823B55DA35F410D0696AEE6EB6" type="description">
<paragraph id="2E1D148F61D7C62435AC510823309942">
<figureCitation id="00FD09877E6DEB2739D6BD0C7135AF63" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">Figs 1</figureCitation>
,
<figureCitation id="D5B96BB91E404DEECC7F2E16407F8D09" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2</figureCitation>
,
<figureCitation id="640789A2AB8EEFC8D94718E1D815EB55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Putative larva of Leuctra enigma sp. nov. A head and pronotum, dorsal view B meso- and metathorax, dorsal view C hind leg, dorsal view D terminalia, dorsal view. Scale bars: 1 m." figureDoi="10.3897/zookeys.1218.120744.figure3" httpUri="https://binary.pensoft.net/fig/1175454">3</figureCitation>
,
<figureCitation id="E5ACE05C9B71F11AB6AF1F3104752662" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4</figureCitation>
,
<figureCitation id="4BB965E034B5FD1F716FE02E8FDB6097" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 A, E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEOAC" type="materials_examined">
<paragraph id="0126ABC37F466E8BAEB890E0EAD32356">
<heading id="560A4010925A5D09914B3840A1EECADC" reason="title">Type material.</heading>
</paragraph>
<paragraph id="ABD2D5596D695672A99626FB19745397">
<materialsCitation id="3C5355A2E3B122303F1EDAD02825C78D" collectingDate="2012-10-10" collectionCode="MM" collectorName="P. Juhasz &amp; T. Kovacs &amp; D. Muranyi &amp; G. Puskas" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2012-178, PLETYP-30" specimenCount="1" specimenCount-male="1" stateProvince="Diber county" typeStatus="holotype">
<emphasis id="7F13AE7AC28B93A9EA10E67BC34FA02F" bold="true" italics="true">
<typeStatus id="A741F6D46ADC696A5BB2485E08EB3DF4" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="C7C1B6D1BFEDA82260C4D4456C464848" count="1" type="male">male</specimenCount>
:
<collectingCountry id="DB6C208F616F41F94444EC7F7199E1C5" name="Albania">Albania</collectingCountry>
: •
<collectingRegion id="27330240143CAC28C1698E8FA411425B" country="Albania" name="Diber">Dibër county</collectingRegion>
,
<collectingMunicipality id="B672AFCC40CFCEAB27510F71E0309FD3">Bulqizë municipality</collectingMunicipality>
,
<collectingMunicipality id="5F8D3AD7EC20EA823AEFC5FFF871D832">Çermenikë Mts</collectingMunicipality>
,
<location id="D9124DA58F03C65614D2A7EADB75807A" LSID="urn:lsid:plazi:treatment:6EAE8429DA2E58F48A813DD7F30E156A:D9124DA58F03C65614D2A7EADB75807A" country="Albania" county="Bulqize" latitude="41.38665" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" name="brooks in open forest beneath Mt. Kaptine" stateProvince="Diber county">brooks in open forest beneath Mt. Kaptinë</location>
,
<quantity id="1E04BEDFD7C6DE0401409AABFC33C2D1" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="A3A8926D39E6CC4D4FF0C882831DA717" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESOAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[20.288967,41.386650]}">
<geoCoordinate id="827D837D81A5F0B00432A2D4DE17C060" degrees="41" direction="north" minutes="23.199" orientation="latitude" precision="1" value="41.38665">41°23.199'N</geoCoordinate>
,
<geoCoordinate id="35B0D6825BE8418142353A3FC2077CAB" degrees="20" direction="east" minutes="17.338" orientation="longitude" precision="1" value="20.288967">20°17.338'E</geoCoordinate>
</named-content>
,
<date id="F6B118D5C55656BC6456D9E85D064F5F" value="2012-10-10">
<collectingDate id="2D4C0261269DD943B2CA6580199D38F9" value="2012-10-10">10.x.2012</collectingDate>
</date>
(field number: loc. 25), leg.
<collectorName id="5905C8F6D06B9D7F03EFEBABE4E40436">P. Juhász</collectorName>
,
<collectorName id="E0DE69E4214469A65437F0B4AF40D493">T. Kovács</collectorName>
,
<collectorName id="19BD116B691D3BAE221B8E90A1D5C7CE">D. Murányi</collectorName>
,
<collectorName id="82F77C00D9AAD8264B9F500A3071462F">G. Puskás</collectorName>
(
<named-content id="BA406BC178241D40ADA535E75CC81C76" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="EE54FE3A1A7BDDFCB525554EF5668C4A" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="50F536263C045D9558395F656BDC2AC5">2012-178</specimenCode>
,
<specimenCode id="63E5A67E0C00A28CD313330F0B8E7876">PLETYP-30</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3C203646BECE370B2315EE7069790EF3" collectingDate="2012-10-10" collectionCode="BYU" collectorName="P. Juhasz &amp; Kovacs, D &amp; Muranyi, G" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
<emphasis id="9E5D63E817B9D3F85DD1F27E50C14E63" bold="true" italics="true">
<typeStatus id="D25EB46D3C22182F1353E50EFCD5735A" type="paratype">Paratypes</typeStatus>
</emphasis>
: • same locality and date:
<specimenCount id="3C18C539F49C20305DD4F01E3FA27278" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="8251C2078C4F266B1D94E97A11D957F7" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="4F5E622C03253AFD5F8EBEEEB7CAA12B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/monte-l-bean-life-science-museum" xlink_title="Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, US">
<collectionCode id="29ECEDC6A0A44EE0E95A11D810FB73E0" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" type="University or college">BYU</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="0359E18E5FAB06CE4FD5BA099E29E610" collectingDate="2012-10-10" collectionCode="CGV" collectorName="P. Juhasz &amp; Kovacs, D &amp; Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
<specimenCount id="EF39CA5B7BBE1E294B787BCA65003EF2" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="CDA4FA16E24C57AAE778FD1190202F8A" count="1" type="female">1 ♀</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EEPAC" xlink_title="Gilles Vinçon Collection, Grenoble, France">
<collectionCode id="18BB392891DCCA3A2FDE1052F1923BA5">CGV</collectionCode>
</abbrev>
)
</materialsCitation>
<materialsCitation id="84E2BA28AA8D024C193AD38B7F8DE9E3" collectingDate="2012-10-10" collectionCode="HNHM" collectorName="P. Juhasz &amp; Kovacs, D &amp; Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCode="PLP 4184" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Diber" typeStatus="paratype">
<specimenCount id="91630917EEED0ACEE938599BE808050D" count="5" type="male">5 ♂</specimenCount>
<specimenCount id="47AEB02B438FA870B706F6115E28CE35" count="6" type="female">6 ♀</specimenCount>
(
<named-content id="B50D0B257AC67A3ED14D647EAF59AAD4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Collection of Smaller Insect Orders, Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="53BB43BB699D933327052F17CE599E76" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
:
<specimenCode id="DEA222A88988C6FECFF0D41EC4CED7A7" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">PLP 4184</specimenCode>
)
</materialsCitation>
<materialsCitation id="EF2B236853CDE8A511AC46E47022A4DD" collectingDate="2012-10-10" collectionCode="MM" collectorName="P. Juhasz &amp; Kovacs, D &amp; Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCode="2012-178, PLETYP-31" specimenCount="15" specimenCount-female="10" specimenCount-male="5" stateProvince="Diber" typeStatus="paratype">
<specimenCount id="1C87400453B25986E670B85A3443D38E" count="5" type="male">5 ♂</specimenCount>
<specimenCount id="FCCA2BA04FAD2F11A7BD3D25DA2C3165" count="10" type="female">10 ♀</specimenCount>
(
<named-content id="257858127369A044A0054F73A02C50EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="1EF113F034501570CB3AE40D5FC26152" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="D5CE0A55919D5071644E3C2A9459393D">2012-178</specimenCode>
,
<specimenCode id="36B5E556FC960624EC3E849A2B65B20F">PLETYP-31</specimenCode>
)
</materialsCitation>
<materialsCitation id="FDFBEC229D01B683D50070DC39A3480F" collectingDate="2012-10-10" collectionCode="SMNS" collectorName="P. Juhasz &amp; Kovacs, D &amp; Muranyi, G" country="Albania" county="Bulqize" elevation="1600" isTailEnumeration="true" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Cermenike Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Diber" typeStatus="paratype">
<specimenCount id="2061A9EFC1EE8C9BED20734DE2F7FBB7" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="DC6CF87D2E038DCF4A6EDFA2EEAABFD5" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="FFB8AA8B216CC59C715C7C878FCFB2F0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-fuer-naturkund-stuttgart" xlink_title="Stuttgart State Museum of Natural History, Stuttgart, Germany">
<collectionCode id="59EC5B8FE10781CFB2E8874212C9A851" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" type="">SMNS</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5CEC01EA0728C2E6F03118AF8B50D91B">
<heading id="680E8E18EA394FA39A695CCBFA32DCC1" reason="title">Other materials.</heading>
</paragraph>
<paragraph id="775487E14E9A8830C1829CBDE394D659">
<materialsCitation id="2A19483FAF48F90044602AC4BB2B7A4B" collectingDate="2012-06-21" collectionCode="HNHM" collectorName="Z. Feher &amp; T. Kovacs &amp; D. Muranyi" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="PLP 4099" specimenCount="3" specimenCount-larva="3" stateProvince="Diber county">
Same locality,
<date id="AB44B9CA1A5243BBEC803F2B2793EDD1" value="2012-06-21">
<collectingDate id="794A5F440C4004ABE58D5BD8DEBA3AB4" value="2012-06-21">21.vi.2012</collectingDate>
</date>
(loc. 58), leg.
<collectorName id="1A1BA45815EE9C823D3C9CC182F9DDDF">Z. Fehér</collectorName>
,
<collectorName id="A778FD7C8EFA460BB6A3A548855AAF84">T. Kovács</collectorName>
,
<collectorName id="B5CC7B7AB1751C9FCE48202326DF3E2F">D. Murányi</collectorName>
:
<specimenCount id="81C64CD763AB53457E63C5BADC424CEC" count="3" type="larva">3 putative larvae</specimenCount>
(
<named-content id="0D98F68556253705BB32CDD70433E15B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Collection of Smaller Insect Orders, Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="EEC0B5785C423A9B80E8BBCBE39A57CB" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
:
<specimenCode id="25911FB961AA049AAB38E3D26DA44394">PLP 4099</specimenCode>
)
</materialsCitation>
<materialsCitation id="DFD8A18159A178F3F92F04AFC446415B" collectingDate="2012-06-21" collectionCode="MM" collectorName="Z. Feher &amp; T. Kovacs &amp; D. Muranyi" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2012-1" specimenCount="2" specimenCount-larva="2" stateProvince="Diber county">
<specimenCount id="7558581FBA14482ABF7DBAC85A28E6EB" count="2" type="pupa">2 putative larvae</specimenCount>
(
<named-content id="7F5C705FA24469DD5E0DE764705201CA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="F937F2AF3B3F95EE120668EFCE3FF174" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="585AEB4FFA56BED0B3C8F7114DE1F1AC">2012-1</specimenCode>
)
</materialsCitation>
<materialsCitation id="2E96475B8AA9D537569B700F954E8D52" collectingDate="2013-05-27" collectionCode="MM" collectorName="P. Juhasz &amp; T. Kovacs &amp; G. Magos &amp; G. Puskas" country="Albania" county="Bulqize" elevation="1600" latitude="41.38665" location="brooks in open forest beneath Mt. Kaptine" longLatPrecision="1" longitude="20.288967" municipality="Bulqize municipality" specimenCode="2013-19" specimenCount="2" specimenCount-larva="2" stateProvince="Diber county">
same locality,
<date id="1C9F813D5E1224A66711FA3025EA8E47" value="2013-05-27">
<collectingDate id="66112502A9D9E6EB32CD463B75AF76CF" value="2013-05-27">27.v.2013</collectingDate>
</date>
(loc. 19), leg.
<collectorName id="5E778670F8D3375B8DF94A66036BC0F4">P. Juhász</collectorName>
,
<collectorName id="DAD424E9366B51FC5CC04AF89887CA32">T. Kovács</collectorName>
,
<collectorName id="E9A80D47C57C3F0A4BAF0299D2AA72EB">G. Magos</collectorName>
,
<collectorName id="F3F1A53D8F2F52D5A54BA1379DCF397A">G. Puskás</collectorName>
:
<specimenCount id="23FC3240D9C4DFB83C0845D10B14531A" count="2" type="larva">2 putative larvae</specimenCount>
(
<named-content id="4B4C930BF6017CCEB3440A860DC8FDE3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="AF8A5184FE81265141FCDC9BA42822A8" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="14977F826BEE3873F549ED111CF7AB0E">2013-19</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOAAE" type="diagnosis">
<paragraph id="F295F25D32DE87B9121FE44C46A386FE">
<heading id="D185C4FAD07D75A16AA0DC1EFC7A2E7D" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="45A473D00F8A660D232A47FC19D73135">Macropterous in both sexes. Male tergite VII with huge membranous portion; tergite VIII with large, triangular posteromedial process, terminating in bi- or trilobed tip; tergite IX with large, lobed posteromedial sclerite supported by anterior sclerotised spots; tergite X posterior margin with deep and wide notch; sternite IX bears a vesicle shorter than 1 / 5 segment length; paraproct tip pointed, specillum slightly longer than paraproct, tip blunt. Female terga I VIII with medial sclerite; subgenital plate with large lobes terminating in swollen and raised apex, central plate with anterolateral swellings and pale medial portion, posterior portion trapezoid in ventral view and nose-shaped in lateral view, well past by the lobes. Larva stout and with long setation, eyes small, clypeus lacks pointed corners.</paragraph>
</subSubSection>
<subSubSection id="SECID0ETAAE" type="description">
<paragraph id="EDF3965B6579F5BB7CBC04A97A33168C">
<heading id="E5FD55298CB17BD8D8998A8D09689085" reason="title">Description.</heading>
</paragraph>
<paragraph id="A223C02EE9D7C9037CCCB4D2EFABBAF2">
Medium sized, robust species, both sexes macropterous. Forewing length:
<typeStatus id="EEEC63E844C50C043ED702BCAFFF79C4" type="holotype">holotype</typeStatus>
6.0 mm, male
<typeStatus id="7BEAD6A3A8CE8D24D65D3E7ACC7B9ABA" type="paratype">paratypes</typeStatus>
<quantity id="34CBFA8E80568BF60D98332C5D1F45E5" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.86.4 mm</quantity>
, female
<typeStatus id="9E507600E0EED1092595DEAA2D7173E0" type="paratype">paratypes</typeStatus>
<quantity id="772733596F3C0897617FB5E5624D611A" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.8" metricValueMin="6.5" unit="mm" value="7.15" valueMax="7.8" valueMin="6.5">6.57.8 mm</quantity>
; body length:
<typeStatus id="BE4D0B8769CE69A1D0761D98F227196A" type="holotype">holotype</typeStatus>
<quantity id="0DBE9FADC0DD1E73A45848E2C8B8E225" metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
, male
<typeStatus id="B5C94B151BCA70BB3BD75F142443EE4E" type="paratype">paratypes</typeStatus>
6.0
<quantity id="3AC6A75237AA7AD10DE179D7B4FEB317" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
, female
<typeStatus id="4BD690CF0C472CDE709EFA0F6A8C1DC8" type="paratype">paratypes</typeStatus>
8.0
<quantity id="29E59B02CC18DDF6771822A3A3E9B912" metricMagnitude="-3" metricUnit="m" metricValue="8.8" unit="mm" value="8.8">8.8 mm</quantity>
. Setation generally short and dense. General colour dark brown. Head brown with distinct M-line and occipital rugosities, compound eyes relatively small; antennae dark brown, palpi brown. Pronotum brown to dark brown, wider than long and having rounded corners, rugosities distinct. Legs brown to dark brown, tarsi slightly darker. Wings brownish but hyaline, venation brown.
</paragraph>
<paragraph id="2572AE07402953658E21F98D301A8EAE">
Male abdomen (Figs
<figureCitation id="028B93260787AD6B44B103F050534AE6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2 A E</figureCitation>
,
<figureCitation id="ED80133E17CB1D8ACF44962122766431" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 A</figureCitation>
): Tergite I membranous posteriorly and medially, with two medial spots; tergite II poorly sclerotised anteriorly. Transverse row of four pigmented spots distinct on terga II VIII. Terga II III with poorly developed antecosta, terga IV VII with entire antecosta, terga III VI full sclerotised. Tergite VII with huge medial membranous portion, a rounded or arrow-shaped sclerotised area extends between the medial pair of pigmented spots. Tergite VIII: antecosta nearly entire, gradually weakened in the medial 1 / 5, sometimes interrupted by a narrow medial notch; the posteromedial process is triangular, originated in the anterior 1 / 3 of the segment and reach the posterior 1 / 3 by its weakly bilobed or trilobed tip, medial portion with small membranous portion, medial pair of pigmented spots recognisable by the lateral sides of the process; the process is not darker than the lateral areas of the segment, wider than 1 / 3 segment width, slightly erect in side view; lateral edges of the medial membranous area obscure, while the anterior edge is sharply defined. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 of its length; posteromedial sclerite large, weakly bilobed anteriorly while posterior portion with long, diverging lobes, the sclerite is anteriorly supported by two sclerotised spots. Anterior margin of tergite X bilobed anteriorly but sclerotisation of lobes obscure; posterior margin with deep notch, as wide as 1 / 2 tergite width; medial portion of the notch with small protrusion at the origin of epiproct. Epiproct drop-shaped, sclerotised only at its sides, stalk gradually widening. Cercus simple, covered with long setae. Sterna II VIII simple, sternite IX bears a very small, rounded vesicle shorter than 1 / 5 of segment length; posterolateral portions of the sternum pale coloured and weakly sclerotised. Paraproct with moderately wide base, narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a pointed tip. Base of paraproct connected to a subrectangular lateral expansion. Specillum longer than the paraproct, gently curved in lateral view and ending in a blunt tip.
</paragraph>
<caption id="C6F74C891334CA4FB42D9636CADFCC95" ID-DOI="10.3897/zookeys.1218.120744.figure2" ID-arpha="D29E1921-6368-5CBB-9AFB-8E4A219564F2" httpUri="https://binary.pensoft.net/fig/1175453" startId="F2">
<paragraph id="68311BF8843F9FF8571D31596AD4D613">
<label id="3C63737BB61DB7AFBD11DACB06E22E48">Figure 2.</label>
</paragraph>
<paragraph id="74E17A853095798D140081A493D14EF0">
Terminalia of
<taxonomicName id="6D5494B556CCAF59F6476D587153CAA5" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="D87AA987BB612E507289054A73C8881C" italics="true">Leuctra enigma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7E95C8CAB3A981111C29D441C25310F" rank="species">sp. nov.</taxonomicNameLabel>
with setation omitted
<emphasis id="9F4705E1230B07538BB53E95053496F4" bold="true">A</emphasis>
male terminalia, dorsal view
<emphasis id="D8DB4DEEE3C7384F5C4298FF43525F8C" bold="true">B</emphasis>
same, lateral view
<emphasis id="39BD783A513D71BAC14D89F7C51F305F" bold="true">C</emphasis>
same, ventral view
<emphasis id="A0841E6B3AE0329FE3300802813416DC" bold="true">D</emphasis>
variations of male tergite 8, dorsal view
<emphasis id="15F1692F8B358D740DBE6405038DB197" bold="true">E</emphasis>
male paraprocts and specillae, caudal view
<emphasis id="EF2E1BCFB445DC1D8463E49334978803" bold="true">F</emphasis>
female terminalia, ventral view
<emphasis id="F825201A0BBF94194A023EC16F11348C" bold="true">G</emphasis>
same, lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="D13F9AF2745E4692691CC060A36E789F">
Female abdomen (Figs
<figureCitation id="84911C26077950CE2F2263191655CC96" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Terminalia of Leuctra enigma sp. nov. with setation omitted A male terminalia, dorsal view B same, lateral view C same, ventral view D variations of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure2" httpUri="https://binary.pensoft.net/fig/1175453">2 F G</figureCitation>
,
<figureCitation id="695AEA7C28E90041CF43D3ADBF33E3FC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Terminalia of four Balkan members of the Leuctra hippopus group A D male terminalia, dorsal view E H female terminalia, ventral view. A, E L. enigma sp. nov., Çermenikë Mts, Albania B, F L. golija sp. nov., Golija Mts, Serbia C, G L. pseudohippopus Raušer, 1965, Stara Planina, Serbia D, H L. hippopoides Kaćanski &amp; Zwick, 1970, Sutjeska, Bosnia &amp; Herzegovina. Scale bars: 1 mm (D for A D, H for E H)." figureDoi="10.3897/zookeys.1218.120744.figure7" httpUri="https://binary.pensoft.net/fig/1175458">7 E</figureCitation>
): Terga I VIII with transverse row of four pigmented spots clearly visible; terga I VIII mostly membranous but with lateral sclerites and a medial sclerite as well; medial sclerite is a longitudinal stripe, more widened on terga II and III; tergite IX mostly, tergite X fully sclerotised. Sterna I VII simple, sterna II VII with one subrectangular median sclerite and two small anterior sclerites that are not fused. Subgenital plate of sternite VIII large and complex, not fused with other sclerites, lacks distinct setation. It has two large and well-defined lobes, darker brown than the remainder of the plate; the lobes terminate in a swollen, rounded apex that are paler than the base of the lobes, raised in lateral view and converging in ventral view. The central plate has two anterolateral swellings, medial portion pale and lightly sclerotised; the plate terminates in a trapezoidal bulge, raised and nose-shaped in lateral view, clearly separated from the lateral lobes that extend beyond it. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus, and epiproct simple. Spermathecal sclerite thin, ring-shaped, with small anterior teeth and long posterior teeth.
</paragraph>
<paragraph id="7C25993C8C3BF898B493C0EFAD5DCDEE">
Putative larva (Fig.
<figureCitation id="A73B88BF9CB7FA0271C78C4EC17811B3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Putative larva of Leuctra enigma sp. nov. A head and pronotum, dorsal view B meso- and metathorax, dorsal view C hind leg, dorsal view D terminalia, dorsal view. Scale bars: 1 m." figureDoi="10.3897/zookeys.1218.120744.figure3" httpUri="https://binary.pensoft.net/fig/1175454">3</figureCitation>
): Body stout, length of the penultimate or younger larvae 7.0
<quantity id="795125E323BED14F4B18022BD0A81C30" metricMagnitude="-3" metricUnit="m" metricValue="7.5" unit="mm" value="7.5">7.5 mm</quantity>
. General colour pale brown, without distinctive markings, pilosity long and distinct. Head much wider than long, eye small, shorter than the widely rounded occipital region, ocelli distinct, clypeus not modified. Palpi simple, antenna longer than 1 / 2 body length. Pronotum much wider than long. Wing pads under development, posterior edge of meso- and metathorax between wing pads rounded, not like the typical
<emphasis id="F60092C8A418DDDDB3D2CFEE36FB0C36" italics="true">prima-hippopus-inermis</emphasis>
group shape. Legs short, tibiae not longer than femora, stretched hind leg reaches back to the seventh abdominal segment. Abdomen stout, integument matt pale brown, paraproct triangular. Cercus nearly as long as abdomen, with 25 cylindrical segments, medial segments twice longer than wide. Setation of the larva: antenna and palpi with short setae but head and clypeus with dense and erect setation, longest setae as long as the distance between hind ocelli; marginal setae continuous on pronotum, as long as the setae of the head; wing pads with similar long, erect and dense setae; femora and tibiae with hairs as long as the width of each femora, distributed on the whole surface but not forming swimming fringe; tarsi with short and scarce setation; abdominal segments evenly setose, with erect setae approximately as long as the setae of the head; cercomeres bald besides the erect apical whorl, slightly shorter than abdominal setae.
</paragraph>
<caption id="AAB0889B5304D2E8B98789495C3C1901" ID-DOI="10.3897/zookeys.1218.120744.figure3" ID-arpha="00379A01-9C9E-5DE2-9BF7-DD7563E6CB57" httpUri="https://binary.pensoft.net/fig/1175454" startId="F3">
<paragraph id="8BA2825452430B0BCE8C43EF0AEAF91C">
<label id="791AA4B18D15D7B28D5ADC69E0CED2D0">Figure 3.</label>
</paragraph>
<paragraph id="B9BC152434A262E1FDCB8FF01BA11A8D">
Putative larva of
<taxonomicName id="E5EDDFAFDEB4230B415505CE1135B1B7" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="1340B40A4FC9E1DD97FBA4CD0912581D" italics="true">Leuctra enigma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5B33ED51049E5866E131D6DD702697C9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7CD88880B28C60180EE234A4235B4011" bold="true">A</emphasis>
head and pronotum, dorsal view
<emphasis id="5493C952A555BCB9BA7A99C4ADA73615" bold="true">B</emphasis>
meso- and metathorax, dorsal view
<emphasis id="C188AC4EFD090E3F21C8CC9F46DD5D36" bold="true">C</emphasis>
hind leg, dorsal view
<emphasis id="358B790E117E9830EFA839DF18E493D8" bold="true">D</emphasis>
terminalia, dorsal view. Scale bars: 1 m.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMEAE" type="Affinities">
<paragraph id="6F5A7C7C84858BA3DD5A3768809F9784">
<heading id="C3F04B272615CD629A67D5A995B34B35" reason="title">Affinities.</heading>
</paragraph>
<paragraph id="51E6B10CE08D94ADECA0FA6008FDCEDE">
The new species is an isolated member of the
<taxonomicName id="A855CA7D39ADF2DE6BA8748E5A1C2052" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="A6F614163113C96DE5DEEF42F5E5E77B" italics="true">hippopus</emphasis>
</taxonomicName>
group. The male is rather distinctive, the only
<taxonomicName id="AF2D8B3CBC513CD23E904FED48D3B918" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="9A9EEB7E24C5594CDCB3BB76A33D631A" italics="true">Leuctra</emphasis>
</taxonomicName>
with comparable posteromedial process of tergite VIII is the Caucasian-Anatolian
<taxonomicName id="88165E5618477302FADF9144C520AE9A" authority="Zhiltzova, 1960" authorityName="Zhiltzova" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="martynovi">
<emphasis id="9659771FA458234E78E2827C428F5099" italics="true">L. martynovi</emphasis>
<bibRefCitation id="658B9CABA39350D4E515D98437AD3A9B" author="Zhiltzova LA" issue="1" journalOrPublisher="Entomologicheskoe Obozrenie" pagination="156-171" refId="B84" refString="Zhiltzova LA (1960) K poznanyu vesnyanok (Plecoptera) Kavkaza. 4. Novye vidy Leuctridae. Entomologicheskoe Obozrenie 39 (1): 156 - 171." title="K poznanyu vesnyanok (Plecoptera) Kavkaza. 4. Novye vidy Leuctridae." volume="39" year="1960">Zhiltzova, 1960</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="E31E2BA1B27EA1E22D84E22B9A788BEC" authorityName="Zhiltzova" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="martynovi">
<emphasis id="4E2E017F9C3505410E345CA7D611BD72" italics="true">L. martynovi</emphasis>
</taxonomicName>
differs by larger posteromedial sclerite of tergite IX, shallow posterior notch of tergite X, and larger vesicle of sternite IX. The subgenital plate of the female shows similarity with the narrowly defined
<taxonomicName id="59DC3945CB751FDBC3794C50170D14C4" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="14102CD7658965A8ED02AC9479685AFC" italics="true">hippopus</emphasis>
</taxonomicName>
group (
<taxonomicName id="AFA67400EA5E7CDD4F04EF6420AB4E43" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopus">
<emphasis id="35CE7F348E5F816718A2D33C4EFFB3B7" italics="true">hippopus</emphasis>
</taxonomicName>
subgroup, sensu, e. g.,
<bibRefCitation id="8B45A0B49E6358704BD487B65C4F44CD" author="Ravizza C" journalOrPublisher="Lauterbornia" pagination="1-42" refId="B69" refString="Ravizza C (2002 b) Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species. Lauterbornia 44: 142. https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" title="Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species." url="https://www.zobodat.at/pdf/Lauterbornia_2002_45_0001-0042.pdf" volume="44">Ravizza 2002 b</bibRefCitation>
), having large lobes and a pale central plate. However, the combination of bulging lobes of the subgenital plate, central plate with anterolateral swellings and trapezoid, bulging posterior portion, already distinguish it from all other
<taxonomicName id="1857BC0F2F015C821C56F6C0B09FB992" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="58C3E862010CA04C3E8F7318259ED8AA" italics="true">Leuctra</emphasis>
</taxonomicName>
. The putative larva is distinctive by its stout body and dense, long setation. Among the Balkanian
<taxonomicName id="FAA3ED4E4EAC5EAEB97850F692C114E4" authorityName="Stephens" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="912B9F485A2F8D9AB2C9CE7C498454C5" italics="true">Leuctra</emphasis>
</taxonomicName>
, only
<taxonomicName id="8D1011899E36158E0B68EC38AF1A063F" authority="(Olivier, 1811)" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B329A83A9C6D474BC5B6BA7D2986854E" italics="true">L. nigra</emphasis>
(
<bibRefCitation id="32BA3B7AE0BA7C23F8F307E3D9956721" author="Olivier GA" journalOrPublisher="Encyclopédie méthodique, ou par ordre de matiéres" pagination="185-187" refId="B57" refString="Olivier GA (1811) Nemoure. Encyclopedie methodique, ou par ordre de matieres 8: 185 - 187. https: // archive. org / details / encyclopdiemthod 08 oliv / page / 186 / mode / 2 up" title="Némoure." url="https://archive.org/details/encyclopdiemthod08oliv/page/186/mode/2up" volume="8" year="1811">Olivier, 1811</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="549EF697D19D85155A86D7540224A985" authority="Bogoesco &amp; Tabacaru, 1960" authorityName="Bogoesco &amp; Tabacaru" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="033C91CB60A60BF60590D7E88AE19CB1" italics="true">L. hirsuta</emphasis>
<bibRefCitation id="6EC5496CE4D1D44AA0679ED29B2EC070" DOI="10.5169/seals-401378" author="Bogoesco C &amp; Tabacaru I" issue="1-2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="91-96" refId="B8" refString="Bogoesco C, Tabacaru I (1960) Contribution a letude des Leuctra (Plecopteres) des Carpathes roumaines. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 33 (1 - 2): 91 - 96. https: // doi. org / 10.5169 / seals- 401378" title="Contribution à ľétude des Leuctra (Plécoptères) des Carpathes roumaines." volume="33" year="1960">Bogoesco &amp; Tabacaru, 1960</bibRefCitation>
</taxonomicName>
are having similar long setation, however, their body is less stout, and their eyes are larger. The Alpine-Carpathian
<taxonomicName id="0E930CB682ED633D1FCAFCC0497CA67E" authority="Kempny, 1898" authorityName="Kempny" authorityYear="1898" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="braueri">
<emphasis id="298D06D51EC988E3E74BD513EF1AECD3" italics="true">L. braueri</emphasis>
<bibRefCitation id="FD740C0FBDB8A551A0A65ED3244FA7DF" author="Kempny P" refId="B33" refString="Kempny P (1898) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. I. Theil. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 48: 213 - 221 + Pl. III. https: // www. biodiversitylibrary. org / part / 39234" url="https://www.biodiversitylibrary.org/part/39234" year="1898">Kempny, 1898</bibRefCitation>
</taxonomicName>
has similarly stout and hairy larva, but its clypeus is armed with prominent pointed corners.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAIAE" type="distribution">
<paragraph id="F6E67EB96A4E7181E731FF659F519C0D">
<heading id="2D2CF29CA637C602E580E7BDCE9D725A" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="4D2A369FA6A90594C83CF126C6DA117E">
The species was collected at a single high elevation site of the Çermenikë Mts, central
<collectingCountry id="A29BE67EE8FE99DC3766E4BA32A3474F" name="Albania">Albania</collectingCountry>
(Fig.
<figureCitation id="49E3F9D2BD38FB03E9E6739E188550F9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">1</figureCitation>
). It was not found at any further localities, despite of intensive collecting efforts during the last twenty years conducted in central
<collectingCountry id="DA8E16D7F57DA965236B84B2AE33109A" name="Albania">Albania</collectingCountry>
. The habitat is disturbed by partial deforestation, most sections of the two confluent brooks run open or in bush of young beech (Fig.
<figureCitation id="8EB7FC15C785F8F0AD41290B6AD793FA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4 A, B</figureCitation>
). The brooks run moderately fast,
<quantity id="489B17EF23FDB669606D23DDFA8378A0" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.51 meter</quantity>
wide and not deeper than 20 centimetres (both in June and October). The substrate is stony, mixed with small sandy patches, few patches of dead wood and partly submerged plants (Fig.
<figureCitation id="C842B61835E58C0DCE64E63D43B6CAE1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Leuctra enigma sp. nov. A type locality: partly open brook in the Çermenikë Mts. B section at forest edge C substrate of the type locality." figureDoi="10.3897/zookeys.1218.120744.figure4" httpUri="https://binary.pensoft.net/fig/1175455">4 B, C</figureCitation>
). The specimens were swarming together with
<taxonomicName id="9D43477EB2527EAD87484521703235E2" authorityName="Bogoesco &amp; Tabacaru" authorityYear="1960" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="5AD26731567414651A6639209B431E76" italics="true">L. hirsuta</emphasis>
</taxonomicName>
in
<date id="D1035615169C006A6D9192D598859626" value="2012-10">October 2012</date>
. When we visited the locality in late spring and early summer, a more diverse fauna was found:
<date id="ABC002AA39179D628260C0B528D9B5DC" value="2012-06-21">21. vi. 2012</date>
(leg. Z. Fehér, T. Kovács, D. Murányi): adults of
<taxonomicName id="B1CA003067590B4D194952BF43101F04" authority="Aubert, 1956 b" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
<emphasis id="E0E937E89FBA9703678FF8201B8740FF" italics="true">Brachyptera helenica</emphasis>
Aubert, 1956 b
</taxonomicName>
,
<taxonomicName id="5031065F8ABB2E085CD07C55CBA762F7" authority="Kacanski &amp; Zwick, 1970" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="F8236301CAC40959DAF72B617E2B733E" italics="true">Leuctra hippopoides</emphasis>
<bibRefCitation id="4AD3651C6B8D6F2F332903621721C0FB" author="Kaćanski D &amp; Zwick P" issue="1" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="1-16" refId="B30" refString="Kacanski D, Zwick P (1970) Neue und wenig bekannte Plecopteren aus Jugoslawien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 43 (1): 1 - 16. https: // www. e-periodica. ch / cntmng? pid = seg- 001 % 3 A 1970 % 3 A 43 % 3 A % 3 A 14" title="Neue und wenig bekannte Plecopteren aus Jugoslawien." url="https://www.e-periodica.ch/cntmng?pid=seg-001%3A1970%3A43%3A%3A14" volume="43" year="1970">Kaćanski &amp; Zwick, 1970</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C4E17D06B602AB539CE5BBEA954FF134" authority="Aubert, 1956" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
<emphasis id="6355BE2DF1C31C2607A121C4756D56D0" italics="true">L. metsovonica</emphasis>
Aubert, 1956
</taxonomicName>
,
<taxonomicName id="23189C516F836AA75794A383AD3E40DB" authority="(Klapalek, 1900)" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
<emphasis id="08B675B9DB0B58C0222C8A66440CCEFC" italics="true">Nemurella pictetii</emphasis>
(
<bibRefCitation id="2F401CFA3D2FEF7E84913A91A0E9200A" author="Klapálek F" issue="20" journalOrPublisher="Slovesnost a Uměni" pagination="1-34" refId="B36" refString="Klapalek F (1900) Plekopterologicke Studie. Rozpravy Ceske Akademie Cisare Frantiska Josefa pro Vedy. Slovesnost a Umeni 9 (20): 1 - 34. [+ Pl. I - II.]" title="Plekopterologické Studie. Rozpravy České Akademie Cisaře Františka Josefa pro Vědy." volume="9" year="1900">Klapálek, 1900</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="754DC00D11D0FB59970A09A1ED071DA1" authority="(Retzius, 1783)" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
<emphasis id="E52FF16D84E81CBB4782DB668D3AE3E3" italics="true">Nemoura cinerea cinerea</emphasis>
(
<bibRefCitation id="14E3730D496549FD6EF86C783AA50D9B" author="Retzius AJ" journalOrPublisher="Cruse, Lipsiae" refId="B72" refString="Retzius AJ (1783) Caroli de Geer genera et species insectorum. Cruse, Lipsiae, 220 pp. https: // www. biodiversitylibrary. org / item / 80958 # page / 68 / mode / 1 up" title="Caroli de Geer genera et species insectorum." url="https://www.biodiversitylibrary.org/item/80958#page/68/mode/1up" year="1783">Retzius, 1783</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4DE46F805528AFDC594CE2DD0AE2B7C1" authority="Pictet, 1836" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="D81EB68957DBDD309E4E7ECEF0DEC34C" italics="true">N. marginata</emphasis>
<bibRefCitation id="4E6FD3B00914EF00037DB6B3856D7B17" author="Pictet FJ" issue="1" journalOrPublisher="Memoires de la Société de physique et dhistoire naturelle de Genève" pagination="173-190" refId="B61" refString="Pictet FJ (1836) Description de quelques nouvelles especes d'Insectes du Bassin du Leman. Memoires de la Societe de physique et d'histoire naturelle de Geneve 7 (1): 173 - 190. https: // www. biodiversitylibrary. org / item / 39755 # page / 201 / mode / 1 up" title="Description de quelques nouvelles espéces dInsectes du Bassin du Léman." url="https://www.biodiversitylibrary.org/item/39755#page/201/mode/1up" volume="7" year="1836">Pictet, 1836</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="DB0C4198EFECC59A07F64DE60DA24D9B" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" class="Insecta" family="Perlidae" genus="Perla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="A42155A39A1DE79BA41BB4441DF07341" italics="true">Perla</emphasis>
cf.
<emphasis id="37E1C56F90579208955DFDC8787817F9" italics="true">pallida</emphasis>
<bibRefCitation id="E13CD5A981C94F58B6D7153698A1B632" author="Guérin-Méneville FE" refId="B21" refString="Guerin-Meneville FE (1843) Genre Perle. In: Guerin-Meneville FE (Ed.) Iconographie de Regne Animal de G. Cuvier. Insectes. Bailliere, Paris 393 - 395. https: // www. biodiversitylibrary. org / item / 88601 # page / 399 / mode / 1 up" url="https://www.biodiversitylibrary.org/item/88601#page/399/mode/1up" year="1843">Guérin-Méneville, 1843</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B01FF173BA21C95DCFC5097275AC27A9" authority="Rauser, 1962" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="buresi">
<emphasis id="B930A4995E921877BA0F07258ED49310" italics="true">Isoperla buresi</emphasis>
<bibRefCitation id="C62486DB9011D81952D011D5D9EC92A3" author="Raušer J" issue="70" journalOrPublisher="Acta Faunistica Entomologica Musei Nationalis Pragae" pagination="67-82" refId="B66" refString="Rauser J (1962) Plecoptera bulgarica - I. Acta Faunistica Entomologica Musei Nationalis Pragae 8 (70): 67 - 82. https: // www. aemnp. eu / data / article- 91 / 72 - 8 _ 0 _ 67. pdf" title="Plecoptera bulgarica - I." url="https://www.aemnp.eu/data/article-91/72-8_0_67.pdf" volume="8" year="1962">Raušer, 1962</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="79A22F90F72ABF12B78CB174536FC0B8" authority="(Rostock, 1881)" baseAuthorityName="Rostock" baseAuthorityYear="1881" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="E01DB81F32872006B7ADEF7CF2E0B35D" italics="true">Siphonoperla neglecta</emphasis>
(
<bibRefCitation id="D9DAA556ED76D8EC07596E01E0E86683" author="Rostock M" issue="15" journalOrPublisher="Entomologische Nachrichten" pagination="217-228" refId="B73" refString="Rostock M (1881) Verzeichniss der Neuropteren Deutschlands (1), Oesterreichs (2) und der Schweiz (3). Entomologische Nachrichten 7 (15): 217 - 228. https: // www. biodiversitylibrary. org / item / 110225 # page / 221 / mode / 1 up" title="Verzeichniss der Neuropteren Deutschlands (1), Oesterreichs (2) und der Schweiz (3)." url="https://www.biodiversitylibrary.org/item/110225#page/221/mode/1up" volume="7" year="1881">Rostock, 1881</bibRefCitation>
)
</taxonomicName>
;
<date id="19E05683D69CAB6D57A8F62CE5569290" value="2013-05-27">27. v. 2013</date>
(leg. P. Juhász, T. Kovács, G. Magos, G. Puskás): adults of
<taxonomicName id="6A4FE3EA20CAD035AFAC5EE358E7ECD1" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
<emphasis id="F00F0AC0B87B5F163B3FDED5AC3E2D6B" italics="true">B. helenica</emphasis>
</taxonomicName>
,
<taxonomicName id="4E1D8E48C9D672A6F4524C1A80BF308E" authorityName="Kacanski &amp; Zwick" authorityYear="1970" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="hippopoides">
<emphasis id="B263FC181753FA31B9BCC06142AC1739" italics="true">L. hippopoides</emphasis>
</taxonomicName>
,
<taxonomicName id="44B8CE2D5488ED1168BA1A47C5CBED80" authority="Vincon &amp; Muranyi, 2007" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="A04547F6D8DDD48A43AF5C2B0B1CBB52" italics="true">L.</emphasis>
cf.
<emphasis id="B83C8CAD2724BFF191A184F2946A2FA8" italics="true">dalmoni</emphasis>
<bibRefCitation id="018B05331D9BD2D1E9379F48BF9FF837" author="Vinçon G &amp; Murányi D" issue="3" journalOrPublisher="Nouvelle Revue dEntomologie" pagination="237-248" refId="B82" refString="Vincon G, Muranyi D (2007) Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera). Nouvelle Revue d'Entomologie 23 (3): 237 - 248." title="Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera)." volume="23" year="2007">Vinçon &amp; Murányi, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="80913B4B1B2D6AE12EBA48DEA419A09D" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
<emphasis id="B8C2B79820AB8C9CDEBFE4A326862350" italics="true">Nemoura cinerea cinerea</emphasis>
</taxonomicName>
,
<taxonomicName id="0E40FC64D12715B608660D5D0389CE9F" authority="Despax, 1934" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="943AF154AB664A78EF51AF27BD097531" italics="true">N. uncinata</emphasis>
<bibRefCitation id="B6F6E8F3123F42D70F94732666B04A66" author="Despax R" issue="2" journalOrPublisher="Bulletin de la Société dhistoire Naturelle de Toulouse" pagination="255-270" refId="B15" refString="Despax R (1934) Plecopteres pyreneens. VII. Etude et description de quelques formes de Nemoures apparentees a Nemoura marginata (Pict.). Bulletin de la Societe d'histoire Naturelle de Toulouse 66 (2): 255 - 270." title="Plécoptères pyrénéens. VII. Etude et description de quelques formes de Nemoures apparentées à Nemoura marginata (Pict.)." volume="66" year="1934">Despax, 1934</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="DA218B67B88F9990FBBF70A2CA1F4383" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="F855F3AE815534D6DD11C97897BC39CA" italics="true">N. marginata</emphasis>
</taxonomicName>
,
<taxonomicName id="923121EB61F39E9C3FD54465F44B2523" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
<emphasis id="B6E362DB5BE1A38C3A1BEF31392D234A" italics="true">Nemurella pictetii</emphasis>
</taxonomicName>
,
<taxonomicName id="C2C61A34212082A4F37B7A173EFA2856" baseAuthorityName="Rostock" baseAuthorityYear="1881" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="670F5D5021FBF8BD0F35C8050CBE5B00" italics="true">Siphonoperla neglecta</emphasis>
</taxonomicName>
and
<taxonomicName id="70B7BFA1E367F777E1F46370417C4CF3" authority="(Aubert, 1956)" baseAuthorityName="Aubert" baseAuthorityYear="1956" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="7800E1251F991AFCDD7DE8F3E4025899" italics="true">S. graeca</emphasis>
(Aubert, 1956)
</taxonomicName>
, and putative larvae of the new species were found.
<taxonomicName id="5AC9E7ABB5840518FCC2E1143D350015" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="BE2691385E5499FA856BA96C68025012" italics="true">Leuctra</emphasis>
cf.
<emphasis id="6B606F0801D4B6744D7F5AE575A3D0A8" italics="true">dalmoni</emphasis>
</taxonomicName>
,
<taxonomicName id="3C7A3AFFAB2EC1075BBFB772097251E5" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="8E38907E8C8112A0046CF9A851558814" italics="true">Nemoura uncinata</emphasis>
</taxonomicName>
,
<taxonomicName id="178676AFA9BB2B5E9221B58A315F06D0" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="59C181CCC2DF19E1EF614F14833C383E" italics="true">N. marginata</emphasis>
</taxonomicName>
and
<taxonomicName id="F90CFD8D187A303381AAB4098E59973F" authorityName="Rauser" authorityYear="1962" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="buresi">
<emphasis id="13C9E211C60536F4437D41BF1BD8671A" italics="true">Isoperla buresi</emphasis>
</taxonomicName>
are new findings for
<collectingCountry id="7237F6113BEBBB14A4679FC610DEB376" name="Albania">Albania</collectingCountry>
. Records of other aquatic insects collected at the type locality were already published:
<taxonomicName id="FAF377EF32C69C189CCAE4FC8F9421B7" authority="(Kovacs and Muranyi 2013)" baseAuthorityName="Kovacs and Muranyi" baseAuthorityYear="2013" class="Insecta" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="order">
Ephemeroptera (
<bibRefCitation id="F3457D02FC142EB3B60554A3B7B65BDD" author="Kovacs and Muranyi" firstAuthor="Kovacs" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="15-19" refId="B38" refString="Kovács T, Murányi D (2013) New country data of some mayflies (Ephemeroptera) from Europe. Folia historico-naturalia Musei Matraensis 37: 1519. http://www.nhmus.hu/sites/default/files/balkan/literature/Ephemeroptera/Kovacs_Muranyi_Fh-nMM_15_2013.pdf" title="New country data of some mayflies (Ephemeroptera) from Europe." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Ephemeroptera/Kovacs_Muranyi_Fh-nMM_15_2013.pdf" volume="37" year="2013">Kovács and Murányi 2013</bibRefCitation>
)
</taxonomicName>
:
<taxonomicName id="18D8EF744D04B92B588112D77C4DB5DA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="2BF44927C61715D2359780C52740A76B" italics="true">Baetis muticus</emphasis>
</taxonomicName>
,
<taxonomicName id="2DD6BA9C057747F7C4D52829C273555E" authorityName="McLachlan" authorityYear="1884" class="Insecta" family="Leptophlebiidae" genus="Habrophlebia" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="lauta">
<emphasis id="18CC2CB0AFFF8C30F2AB4BC0EE80AE2C" italics="true">Habrophlebia lauta</emphasis>
</taxonomicName>
;
<taxonomicName id="D806CF5B3C24F3904C819F3CA9EE2E99" authority="(Muranyi and Kovacs 2013)" baseAuthorityName="Muranyi and Kovacs" baseAuthorityYear="2013" class="Insecta" kingdom="Animalia" order="Odonata" phylum="Arthropoda" rank="order">
Odonata (
<bibRefCitation id="7A5B3284AEE9F8D3C68F160112240D77" author="Muranyi and Kovacs" firstAuthor="Muranyi" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="29-41" refId="B48" refString="Murányi D, Kovács T (2013) Contribution to the Odonata fauna of Albania and Montenegro. Folia historico-naturalia Musei Matraensis 37: 2941. http://www.nhmus.hu/sites/default/files/balkan/literature/Odonata/Muranyi_Kovacs_2013.pdf" title="Contribution to the Odonata fauna of Albania and Montenegro." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Odonata/Muranyi_Kovacs_2013.pdf" volume="37" year="2013">Murányi and Kovács 2013</bibRefCitation>
)
</taxonomicName>
:
<taxonomicName id="993C21E416868E5B411206A7C1814A9A" authorityName="Selys" authorityYear="1843" class="Insecta" family="Cordulegastridae" genus="Cordulegaster" kingdom="Animalia" order="Odonata" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="15D18A3A955D7E57F46D9BBE16DC1421" italics="true">Cordulegaster bidentata</emphasis>
</taxonomicName>
;
<taxonomicName id="165FA26D4636BB04F95D4D7FA8E7644C" class="Insecta" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
(
<bibRefCitation id="1B93B582B68DFC2BA5560E87825C4EA5" author="Oláh J &amp; Kovács T" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="89-104" refId="B54" refString="Oláh J, Kovács T (2012 b) New species and records of autumnal Trichoptera form Albania. Folia historico-naturalia Musei Matraensis 36: 89104. http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_Fh-nMM_89_2012.pdf" title="New species and records of autumnal Trichoptera form Albania." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_Fh-nMM_89_2012.pdf" volume="36">Oláh and Kovács 2012 b</bibRefCitation>
,
<bibRefCitation id="DAA80A8EEDF8D8741F10159D84106C3F" author="Olah and Kovacs" firstAuthor="Olah" journalOrPublisher="Folia historico-naturalia Musei Matraensis" pagination="109-121" refId="B55" refString="Oláh J, Kovács T (2013) New species and new records of Balkan Trichoptera II. Folia historico-naturalia Musei Matraensis 37: 109121. http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_2013.pdf" title="New species and new records of Balkan Trichoptera II." url="http://www.nhmus.hu/sites/default/files/balkan/literature/Trichoptera/Olah_Kovacs_2013.pdf" volume="37" year="2013">2013</bibRefCitation>
):
<taxonomicName id="F7900F4623E58251B15B6D7EE9CD0CCF" baseAuthorityName="McLachlan" baseAuthorityYear="1879" class="Insecta" family="Glossosomatidae" genus="Agapetus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="iridipennis">
<emphasis id="733610DD353D3F47041EAA2D534CAFDC" italics="true">Agapetus iridipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="41819550EA61828BDE137DC2AC45772B" authorityName="Marinkovic-Gospodnetic" authorityYear="1959" class="Insecta" family="Limnephilidae" genus="Chaetopteryx" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="bosniaca">
<emphasis id="3A772455527FD3B8EC3CA8129C496CA5" italics="true">Chaetopteryx bosniaca</emphasis>
</taxonomicName>
,
<taxonomicName id="A6B8F1632D411E60F6011DD4200705A5" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Psychomyiidae" genus="Psychomyia" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="05BDFA19C9CD3E321BA1226DD7C90F7C" italics="true">Psychomyia pusilla</emphasis>
</taxonomicName>
,
<taxonomicName id="92E3C1251A2E513EF8A9C1FA3F092F83" baseAuthorityName="Curtis" baseAuthorityYear="1834" class="Insecta" family="Polycentropodidae" genus="Cyrnus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="D84842B959EAFF2E83A011B1A1D05C57" italics="true">Cyrnus trimaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="FD8B1C968CC28F7BA0B232F5BB619EAC" baseAuthorityName="Fabricius" baseAuthorityYear="1775" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="hirtum">
<emphasis id="9FD7B7A4A53843888415D2A766EC73E0" italics="true">Lepidostoma hirtum</emphasis>
</taxonomicName>
,
<taxonomicName id="938DD4EC3E58B9F4E63F0E051FF8EC53" authorityName="Radovanovic" authorityYear="1942" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="plicatus">
<emphasis id="188E128DC127B404EB363EAF72C1C3C9" italics="true">Drusus plicatus</emphasis>
</taxonomicName>
,
<taxonomicName id="3C22A32B44D4E751C21A8AF6581F6E42" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" class="Insecta" family="Leptoceridae" genus="Mystacides" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="azureus">
<emphasis id="6384D7EB7066747EDBB799BB2F17CDEC" italics="true">Mystacides azureus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="7197664E3E06018A98E32208A27A4FCA" ID-DOI="10.3897/zookeys.1218.120744.figure4" ID-arpha="A4EE17C3-ABA5-5EA7-B6E3-CBD7B4B8796E" httpUri="https://binary.pensoft.net/fig/1175455" startId="F4">
<paragraph id="7C6A4ADFA8BB6DF4AE28ECF491E4D57F">
<label id="37D3644262E9DB742CAFA0F0AB03F299">Figure 4.</label>
</paragraph>
<paragraph id="569C3E12E4CCBCDD6E2AA4F253202DD7">
Habitat of
<taxonomicName id="4075DD850E8B65937F60249371526DE4" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="enigma" status="sp. nov.">
<emphasis id="FBAFE96B5794780A3EDFAD0A1A60342F" italics="true">Leuctra enigma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48CEC4B234897FDBB9FA2A5CE584F936" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7F286AD9D4E26DC3AC78A5FF6D23F0C1" bold="true">A</emphasis>
type locality: partly open brook in the Çermenikë Mts.
<emphasis id="4DCB5F2DC35BAE63C0BD06E6E0E382B2" bold="true">B</emphasis>
section at forest edge
<emphasis id="DF334195AF19C2B43FDA4B3DA9ED2057" bold="true">C</emphasis>
substrate of the type locality.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIWAE" type="etymology">
<paragraph id="9DD90B641DE90E886F592B8A2612B990">
<heading id="811205C22358A62CCD7A7DCE4F4A3A37" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="D034F2BC25B5BE653F581DF011CA8418">
The name
<emphasis id="C015550C96019F475BEACBA980E8E70A" italics="true">enigma</emphasis>
(Latin; enigma or riddle) refers to the strange fact that such a distinctive new species was found only at a single locality. Used as a noun, gender neutral.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,751 @@
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<mods:title id="646DEB54F6DE551B9BCDC0D87C748374">Four new species of Leuctra Stephens, 1836 from the Balkans (Plecoptera, Leuctridae)</mods:title>
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<mods:namePart id="DC0259D5DFB293D0A2BE10CA9CEFA85E">Murányi, Dávid</mods:namePart>
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<mods:affiliation id="7C1D5E239684AD4B091DFDDCCBCA0782">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<subSubSection id="DEDE5E51A593F20F62B3CDC39D72BCE1" type="nomenclature">
<paragraph id="8FAD5DAB665EAA36D7074E9392CB1346">
<taxonomicName id="5A1BE46EC1319A9A1492A508785D951D" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
<emphasis id="C54CF1AB0092CE7B2A439EF6C047865C" italics="true">Leuctra visitor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D6FE0D215FABB7A6DDDB1B1D915887E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="7A9C0F0BF7694FEBC7F6CDCBEE4C4720" type="description">
<paragraph id="6E68330531BCEA3DBCE177D18B0F1107">
<figureCitation id="26356180B3068B4FAC3A45F4138F5312" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">Figs 1</figureCitation>
,
<figureCitation id="800011FA3073B52D079A4BD983B467F3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10</figureCitation>
,
<figureCitation id="483FBE3EF102F294EA7CCBB5066AD4A4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11</figureCitation>
,
<figureCitation id="6999D7581D1540DE76B8FFE9B1B3CD49" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 B, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBRBG" type="type material">
<paragraph id="B0D875763183135E44DFB495D85D9C77">
<heading id="3F6E7AFE99FAA70AD73D0E4FE2DF96C9" reason="title">Type material.</heading>
</paragraph>
<paragraph id="457A98E39FBC1540EE506A1D1FFBB294">
<materialsCitation id="0054425AB4586AFB267E8E9434BEAF46" collectingDate="2023-05-05" collectionCode="MM" collectorName="T. Kovacs &amp; D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCode="2023-49, PLETYP-37" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="19A12D0F17FDED284C3412F2D1096AC7" bold="true" italics="true">
<typeStatus id="038D498027E3EF93D53E0BAF1DB94AD6" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="DF875CD840A4AF186767B2AFA48E9AFB" type="male">male</specimenCount>
:
<collectingCountry id="DDAD585E5F4F9AC5C4A1840B376F1552" name="Serbia">Montenegro</collectingCountry>
: •
<collectingMunicipality id="E35A71C15BFA9B3B2751CEE29F8BF967">Plav municipality</collectingMunicipality>
,
<collectingMunicipality id="C8816EDE5D906D725894089D3BD19536">Visitor Mts</collectingMunicipality>
,
<location id="F3D3BF253E0ADA4EA1121C59B3E3152F" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:F3D3BF253E0ADA4EA1121C59B3E3152F" country="Montenegro" county="Katun" latitude="42.6308" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" name="forest springs and their outlet along the Katun road">forest springs and their outlet along the Katun road</location>
,
<quantity id="D4E57FBF035814E0796ACC5275708FE6" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">
<elevation id="3166346C3364884F6B449000A6356E45" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">1730 m</elevation>
</quantity>
,
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<geoCoordinate id="E26BDA1A4B6AD818036AE5ED732428A7" degrees="42.6308" direction="north" orientation="latitude" precision="5" value="42.6308">42.6308°N</geoCoordinate>
,
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<collectingDate id="9391A0A646EE035F12EDFE6F64B6BA89" value="2023-05-05">5.v.2023</collectingDate>
</date>
(field number: loc. 13), leg.
<collectorName id="FDF421C846F2010525ED42F4A8B74202">T. Kovács</collectorName>
,
<collectorName id="309DEC2BDDADE94DC7A24DC8D099E5C5">D. Murányi</collectorName>
(
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<collectionCode id="7DF58FDB202024CBD9E4DDA754E01538" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
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,
<specimenCode id="9B747CB31C2FF1EC2044D0E69DDAF9A2">PLETYP-37</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="0F0F0E193286D76A919D9C2EB657C9C3" collectingDate="2023-05-05" collectionCode="BYU" collectorName="T. Kovacs &amp; D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="DEBC873F79BFB2A0F1FEC516A8BEE17D" bold="true" italics="true">
<typeStatus id="7AB1F2A7C04330467FFC45EAB6CEF80A" type="paratype">Paratypes</typeStatus>
</emphasis>
: • same locality and date:
<specimenCount id="D2C553FB9DF10F69D30F659429AE5023" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="520A2F0E13581503ED9C15BD9F744080" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="E6BB938FFF00F4DE79B2ACDFFD4E11B2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/monte-l-bean-life-science-museum" xlink_title="Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, US">
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<specimenCount id="D7FA167DF4F2BFE2D15197057821F24B" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="82006D631148CF59A500A48966297171" count="1" type="female">1 ♀</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EBSBG" xlink_title="Gilles Vinçon Collection, Grenoble, France">
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<specimenCount id="B406BE9F0E609278985C0CF37C3DBC46" count="11" type="male">11 ♂</specimenCount>
<specimenCount id="7EEF8C93889149EAA843E4181A29757A" count="8" type="female">8 ♀</specimenCount>
(3 pairs in copula) (
<abbrev content-type="institution" id="ABBRID0EGSBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
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:
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<specimenCount id="0FF57A6CD9A0FC8B55864F8E360EF56B" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="CCDC074B4938B93774A067A55E81A322" count="2" type="female">2 ♀</specimenCount>
(
<named-content id="DADF0120711B2F4C2BDEBC14FEA0E208" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="D7F6A0D886F8FB02C04EDC0DE9417A4D" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
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,
<specimenCode id="18EC84A86471A10471B9A5E5A143830A">PLETYP-38</specimenCode>
)
</materialsCitation>
<materialsCitation id="874DE4B6A20B9E6C4E72B4FDDFE7672D" collectingDate="2023-05-05" collectionCode="SMNS" collectorName="T. Kovacs &amp; D. Muranyi" country="Montenegro" county="Katun" elevation="1730" latitude="42.6308" location="forest springs and their outlet along the Katun road" longLatPrecision="6" longitude="19.8358" municipality="Plav municipality" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="5A55B27C3ABFC5366306EB68A1FC6C92" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="FDC220A0A7EACD5A992123A80031F3A9" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="3DACA575285EE2ABC234C7D364E04EEB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-fuer-naturkund-stuttgart" xlink_title="Stuttgart State Museum of Natural History, Stuttgart, Germany">
<collectionCode id="AA88C77959FBC2A9745C0CDFD57C5557" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" type="">SMNS</collectionCode>
</named-content>
)
</materialsCitation>
<materialsCitation id="642CC0E789FD15E3C23F23C1F7C312E6" collectingDate="2022-06-11" collectionCode="EKCU" collectorName="A. Hunyadi &amp; T. Kovacs &amp; D. Muranyi &amp; P. Olajos" country="Serbia" county="Katun" elevation="1845" latitude="42.6185" location="partly open brook and spring along the Katun road" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" specimenCode="PLP 5584" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<collectingMunicipality id="BD9E68DC1811DEC837849A2C14DF0399">Plav municipality</collectingMunicipality>
,
<collectingMunicipality id="78F803975256E3A4045FF055B78646B2">Visitor Mts</collectingMunicipality>
,
<location id="C97D8D5F029AF91752767115EFEB5CE0" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:C97D8D5F029AF91752767115EFEB5CE0" country="Serbia" county="Katun" latitude="42.6185" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" name="partly open brook and spring along the Katun road">partly open brook and spring along the Katun road</location>
,
<quantity id="85CF9027D67FC9C3142AA08FA1CC42BA" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">
<elevation id="ED9BD750267B3888F67063990D5D4815" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">1845 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYSBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.837800,42.618500]}">
<geoCoordinate id="73CC9376D83EF3DEFB823BEE64D9168E" degrees="42.6185" direction="north" orientation="latitude" precision="5" value="42.6185">42.6185°N</geoCoordinate>
,
<geoCoordinate id="AFF32FD41E4BE3A74C8A88F70E2EFA25" degrees="19.8378" direction="east" orientation="longitude" precision="5" value="19.8378">19.8378°E</geoCoordinate>
</named-content>
,
<date id="BD219651E8D49768D2C6920B692E3CA1" value="2022-06-11">
<collectingDate id="C0EFC641586B2CCA625E8C426983ABEB" value="2022-06-11">11.vi.2022</collectingDate>
</date>
(loc. 24), leg.
<collectorName id="8DFD1A1E3FC6EAA624544881D07E3C0D">A. Hunyadi</collectorName>
,
<collectorName id="D313D2E5D349BBF31F32E8F55CD89B9C">T. Kovács</collectorName>
,
<collectorName id="040F81C5C9B51EA63C48590F7CFA00FD">D. Murányi</collectorName>
,
<collectorName id="EDBE98D8B2F31E00DF97DA8643D62071">P. Olajos</collectorName>
:
<specimenCount id="A5E93276699B4394596527E516A30F52" count="5" type="male">5 ♂</specimenCount>
<specimenCount id="1251AE8D1921418F1A8A4A03A8FD7920" count="3" type="female">3 ♀</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0E4SBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
<collectionCode id="053A908A0833F051DF55097473D5753B">EKCU</collectionCode>
</abbrev>
:
<specimenCode id="BC85F061FC7AD64265690ADC99FC36FD" collectionCode="EKCU">PLP 5584</specimenCode>
)
</materialsCitation>
<materialsCitation id="BF9E8A786ECC08B65833B42E2D046167" collectingDate="2022-06-11" collectionCode="MM" collectorName="A. Hunyadi &amp; T. Kovacs &amp; D. Muranyi &amp; P. Olajos" country="Serbia" county="Katun" elevation="1845" latitude="42.6185" location="partly open brook and spring along the Katun road" longLatPrecision="6" longitude="19.8378" municipality="Plav municipality" specimenCode="2022-90, PLETYP-39" specimenCount="17" specimenCount-female="9" specimenCount-male="8">
<specimenCount id="A3116784D2FDAC242A80A209621A6E48" count="8" type="male">8 ♂</specimenCount>
<specimenCount id="76E67D2CCD8A18DF26DD0FAC01738A4B" count="9" type="female">9 ♀</specimenCount>
(1 pair in copula) (
<named-content id="2C62FBD2EF9BAFA6D4E3ACCAA8A825F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="AD1204AF4BFD7DB68866A319A4F6EBFF" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="1767C9A9E4E92FF9D4D1068F87E8EEAE">2022-90</specimenCode>
,
<specimenCode id="1BABC28478128932A652B25D66B66751">PLETYP-39</specimenCode>
)
</materialsCitation>
<materialsCitation id="57BA5ADDE1F4746D21AC2DD4D9D1FEB3" collectingDate="2022-06-11" collectionCode="EKCU" collectorName="A. Hunyadi &amp; T. Kovacs &amp; D. Muranyi &amp; P. Olajos" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="PLP 5572" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingMunicipality id="17A940B1A78D3E0D3557436F5F8567FC">Plav municipality</collectingMunicipality>
,
<collectingMunicipality id="EE14053A8094F9BE8ECC00FB698523BA">Visitor Mts</collectingMunicipality>
,
<location id="77C9C54CFC45F14DD2E95B3F75699CE7" LSID="urn:lsid:plazi:treatment:83CD7F7B778158F0B107C65E2DED1906:77C9C54CFC45F14DD2E95B3F75699CE7" country="Serbia" county="Katun" latitude="42.6294" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" name="forest streams and brooks along the Katun road">forest streams and brooks along the Katun road</location>
,
<quantity id="99E51062DE18C1FB96F1C9A7717BAA70" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">
<elevation id="EC1D464FC85771792ADA706B3FA4CE10" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">1515 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKTBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.844200,42.629400]}">
<geoCoordinate id="7C2570353B6F1BB37BEF61CB3F39330A" degrees="42.6294" direction="north" orientation="latitude" precision="5" value="42.6294">42.6294°N</geoCoordinate>
,
<geoCoordinate id="3ECAC83EEE5866B06F41CFA8467A4210" degrees="19.8442" direction="east" orientation="longitude" precision="5" value="19.8442">19.8442°E</geoCoordinate>
</named-content>
,
<date id="7EFEFED731BE6392A88D2918A0B7ACA2" value="2022-06-11">
<collectingDate id="BE827A1E1419F69FC514593E2BFFFC2C" value="2022-06-11">11.vi.2022</collectingDate>
</date>
(loc. 22), leg.
<collectorName id="70F2FFF8B4458336B34ECC490E490E9D">A. Hunyadi</collectorName>
,
<collectorName id="FF81DBDF2CDBA77D3EBC7F7DA0D3B3F1">T. Kovács</collectorName>
,
<collectorName id="4FD697D3FDCEB4614D844F154A4B3E95">D. Murányi</collectorName>
,
<collectorName id="B9A9DDFC5F6EB362C81441411A7E19C3">P. Olajos</collectorName>
:
<specimenCount id="274CA913DE95444369BCD4D8DDF5767D" count="1" type="female">1 ♀</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EPTBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
<collectionCode id="8C676EAB099845F15C0D596C8EAC4D4B">EKCU</collectionCode>
</abbrev>
:
<specimenCode id="BC3EA7D7C6573329E5D1CF6B6FFB9C6D" collectionCode="EKCU">PLP 5572</specimenCode>
)
</materialsCitation>
<materialsCitation id="67A17445E9C8C2F81E4AEDB6140355F8" collectingDate="2023-05-05" collectionCode="EKCU" collectorName="L. P. Kolcsar &amp; T. Kovacs &amp; D. Muranyi" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="PLP 5701" specimenCount="15" specimenCount-female="7" specimenCount-male="8">
same locality,
<date id="8244337572CEF53DBDF0864FE5468230" value="2023-05-05">
<collectingDate id="FF8BCB823C87B604C878E070222002A0" value="2023-05-05">5.v.2023</collectingDate>
</date>
(loc. 12), leg.
<collectorName id="D439F5561939F513C6C9868BD75E5F6F">L. P. Kolcsár</collectorName>
,
<collectorName id="151EA1847C827F98C00B33826A31F8CE">T. Kovács</collectorName>
,
<collectorName id="15C28C8047CAB4F50103BE2CC58AE815">D. Murányi</collectorName>
:
<specimenCount id="C0D257A5E2E5B658EEBB599223E886E1" count="8" type="male">8 ♂</specimenCount>
<specimenCount id="C05B866FE52A457B8E2A69970DAA9071" count="7" type="female">7 ♀</specimenCount>
(2 pairs in copula) (
<abbrev content-type="institution" id="ABBRID0EUTBG" xlink_title="Department of Zoology, Eszterházy Károly Catholic University, Eger, Hungary">
<collectionCode id="B5F02FAB642D3BD83E804E8CE443E846">EKCU</collectionCode>
</abbrev>
:
<specimenCode id="3406D87548C609707634E7DCE967C79F" collectionCode="EKCU">PLP 5701</specimenCode>
)
</materialsCitation>
<materialsCitation id="D4309814D3288D8805572060E904F19C" collectingDate="2023-05-05" collectionCode="MM" collectorName="L. P. Kolcsar &amp; T. Kovacs &amp; D. Muranyi" country="Serbia" county="Katun" elevation="1515" latitude="42.6294" location="forest streams and brooks along the Katun road" longLatPrecision="6" longitude="19.8442" municipality="Plav municipality" specimenCode="2023-48, PLETYP-40" specimenCount="14" specimenCount-female="8" specimenCount-male="6">
<specimenCount id="D8BF9B31422C257BD091F31E0519C639" count="6" type="male">6 ♂</specimenCount>
<specimenCount id="2307CDCA3BD2629985A3AF07247C687E" count="8" type="female">8 ♀</specimenCount>
(1 pair in copula) (
<named-content id="8319A630D72EDCD8CB9A5F3B7FE00B0E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="C19C7AB5CBAFE8164D80A9A0E8FDCEE2" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
:
<specimenCode id="EE61A8828BC2F4BFAEB3147B93916E61">2023-48</specimenCode>
,
<specimenCode id="77E01B9CDA41FED892BD0E1BD06F33EA">PLETYP-40</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5TBG" type="diagnosis">
<paragraph id="7745A045E34DBBE337057C5A0160F72B">
<heading id="74CEC6603DC5E584686D155D8FA35D3B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6B0C07A1EB1DA52F48896456DEB3CE3E">Brachypterous in both sexes. Male tergite VII with small membranous portion; tergite VIII with strong posteromedial process that is not erect in side view and bear angular teeth, membranous area longer than 1 / 2 segment length with a crossband, the posterior margin indented in a rounded membranous field between the posteromedial process and segment sides; tergite IX with short but wide posteromedial sclerite; tergite X posterior margin with a wide V-shaped notch; sternite IX bears a vesicle much shorter than 1 / 2 segment length; specillum slightly longer than paraproct, tip blunt behind subterminal constriction. Female subgenital plate large, trapezoid, with an incision between the long lobes slightly widening towards the median bulge but not forming a triangular field; median bulge is wide in ventral view and distinctly raised in lateral view, with distinct, erect setation; spermathecal sclerite ring-shaped, with large and converging posterior teeth.</paragraph>
</subSubSection>
<subSubSection id="SECID0EDUBG" type="description">
<paragraph id="CD089914392FCCAA88BD45DADAD52DA5">
<heading id="03EF998358DE571A538BCFB17B44AE80" reason="title">Description.</heading>
</paragraph>
<paragraph id="0FBCC8906DA33134CBE067D8E63DA49E">
Medium sized, robust species, both sexes brachypterous (Fig.
<figureCitation id="96AA9A0DFFC4242E2B74664CBF5DFF4E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 A C</figureCitation>
). Forewing length:
<typeStatus id="19D5FDBB040318672B0869D780222F46" type="holotype">holotype</typeStatus>
<quantity id="953570C42BA04073814F6579F1803043" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
, male
<typeStatus id="50B8666C29443CD52991BF9B14C420A1" type="paratype">paratypes</typeStatus>
3.0
<quantity id="C6EBE731E2697F45C24CB9D0FF5F36B1" metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
, female
<typeStatus id="52505A0D89EECA02D9C3477876ED6EFF" type="paratype">paratypes</typeStatus>
<quantity id="2038165EE469136CA90BFFD8BDF863C5" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" unit="mm" value="4.7" valueMax="5.2" valueMin="4.2">4.25.2 mm</quantity>
; body length:
<typeStatus id="20C543D28703D235A6C7E914491696EB" type="holotype">holotype</typeStatus>
<quantity id="E8A47D07863D35D2426D09B446917F5A" metricMagnitude="-3" metricUnit="m" metricValue="6.2" unit="mm" value="6.2">6.2 mm</quantity>
, male
<typeStatus id="ECAB02EF9E3F79BA9EC354A82BFE94DF" type="paratype">paratypes</typeStatus>
5.47.0 mm, female
<typeStatus id="D26D2CA822727759B44F94769E7C8AE2" type="paratype">paratypes</typeStatus>
6.89.0 mm. Setation generally short and dense. General colour dark brown to blackish. Head, antennae, and palpi dark brown. Pronotum dark brown, as wide as long and having rounded corners, rugosities distinct. Legs brown to dark brown. Wings brownish but hyaline, venation brown.
</paragraph>
<paragraph id="951C26E2DBBA2A7482AB5138BEE99A92">
Male abdomen (Figs
<figureCitation id="E7FFEE92625ED227ED73ABD8BC6BF582" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10 A D</figureCitation>
,
<figureCitation id="A4E6F37345C0B7FC4FDAA927F1810D1A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 B</figureCitation>
): Tergite I membranous in its whole medial portion, tergite II membranous anteromedially behind antecosta. Transverse row of four pigmented spots distinct on terga I VII. Terga II and III with medially divided antecosta, terga IV VII with entire antecosta and full sclerotised, tergite VII with small posterior membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends in a small triangular plate, and the membranous area between them is bridged by a weakly sclerotised crossband; width of the membranous area is less than 1 / 2 the segment width, and reaches below 1 / 2 the segment length above the posteromedial process, posterior margin of the membranous portion nearly straight; the surrounding sclerotisation of the membranous area paler than remainder of the segment; posteromedial process paler than the lateral areas of the segment, not erect in side view and slightly overhangs the segment posterior edge, as wide as the membranous area above, bilobed with angular teeth slightly pointed laterad, the space between the teeth is deeply indented and concave; posterior margin between the posteromedial process and segment sides indented in a shallow, rounded membranous field but not lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 2 of its length; posteromedial sclerite short but wide, medial areas usually slightly darker, the medial portion usually not narrowed. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep, V-shaped notch, 1 / 2 as wide as tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk relatively long. Cercus short, covered with long setae. Sterna II VIII simple, sternite IX bears a vesicle much shorter than 1 / 2 segment length, its width more than 1 / 2 its length; the area surrounding the vesicle is pale, posterior edge with two pale and weakly sclerotised areas. Paraproct with moderately wide base, abruptly narrowed after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, short, blunt triangular process. Specillum slightly longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, blunt tip.
</paragraph>
<caption id="4D7C9FEADB58CC7520FBD70BA49246CA" ID-DOI="10.3897/zookeys.1218.120744.figure10" ID-arpha="2C375FA0-3562-5AAA-9381-95F4188FEC09" httpUri="https://binary.pensoft.net/fig/1175461" startId="F10">
<paragraph id="10691B74F9BDE423E9692267691AC299">
<label id="E8D12188D5EC14FF9816D311BBFDFD2F">Figure 10.</label>
</paragraph>
<paragraph id="6917B7A7BB38EB0B81C35D288A0E7909">
Terminalia of
<taxonomicName id="009F4B15AD2AB9FFD5029DF30B58B062" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
<emphasis id="67BB788B7DF7AF345CCADE766D2B0210" italics="true">Leuctra visitor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90416D15AF6C653C28C2CD790E217F61" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5D4453C581F15C9215F82ED7CBF35780" bold="true">A</emphasis>
male terminalia, dorsal view
<emphasis id="710153C13C0A021B472826A0CD25713D" bold="true">B</emphasis>
same, lateral view
<emphasis id="23812668778E8617AD813341900BB461" bold="true">C</emphasis>
same, ventral view
<emphasis id="911D93C9F164711847A213E524EE1419" bold="true">D</emphasis>
male paraprocts and specillae, caudal view
<emphasis id="9F6865CFDDF9BE36A4B954A54ACA1453" bold="true">E</emphasis>
female terminalia, ventral view
<emphasis id="A324E1A98F6B0FD893D209C30D6AE545" bold="true">F</emphasis>
same, lateral view. Setation omitted with the exception of subgenital plate on
<emphasis id="8DC9B2D7F2FC138701465E4289FFAFE4" bold="true">F</emphasis>
. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="F08BD09240EDE018EF698EDDB7CEA6F6" ID-DOI="10.3897/zookeys.1218.120744.figure11" ID-arpha="680DCBF2-C9BE-5907-B2CB-A9055C8270FB" httpUri="https://binary.pensoft.net/fig/1175462" startId="F11">
<paragraph id="A20904142E00D77BF0F0E026084B2E6B">
<label id="3CC4DE726E014D47AF6D1C735168A1AD">Figure 11.</label>
</paragraph>
<paragraph id="893773C691D3DDAB95D16E236D5E0B41">
Habitus and habitat of
<taxonomicName id="B93B018737BA3D641BFE10FA68C8ABE8" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
<emphasis id="69670862E94EF06B6D545DDB18955D96" italics="true">Leuctra visitor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E9344E641F806FD8830F174CC842E48" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1FDEB634A4F4EEC64975586E4FFCD774" bold="true">A</emphasis>
male habitus
<emphasis id="EB9C8BF335F621D829481AFC761B72D3" bold="true">B</emphasis>
female habitus
<emphasis id="0DB99A2520C27F4448678232A35A854D" bold="true">C</emphasis>
copula
<emphasis id="4E749354253E1F0BFED869F3DDDD60F8" bold="true">D</emphasis>
partly open brook at 1845 m in the Visitor Mts.
<emphasis id="C2F511E997ABC8B9E0A4ECCB6492622D" bold="true">E</emphasis>
type locality: forest springs and their outlet at 1730 m in the Visitor Mts.
</paragraph>
</caption>
<caption id="2E482067E55FA955FE302EF3AF5CE5BE" ID-DOI="10.3897/zookeys.1218.120744.figure12" ID-arpha="58F889BE-E1CD-54C3-985B-73213274A2E5" httpUri="https://binary.pensoft.net/fig/1175463" startId="F12">
<paragraph id="2FBD7E8ABFE07BA3140525A926347B1E">
<label id="40DE752A54B73D809D14DF8E040395BF">Figure 12.</label>
</paragraph>
<paragraph id="D60C7EFC73BFFC22E23CE8F06D99A819">
Terminalia of five Balkan members of the
<taxonomicName id="E8BF02B21D939A34EB26F169E0E7415C" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="C0469E434357CFE38812DB32117328DC" italics="true">Leuctra prima</emphasis>
</taxonomicName>
group
<emphasis id="3AC78F880E41C9B40AB71802658E5358" bold="true">A D, I, K</emphasis>
male terminalia, dorsal view
<emphasis id="780C1888B7CFC9EA4AA0489CFC418B8B" bold="true">E H, J, L</emphasis>
female terminalia, ventral view
<emphasis id="45FFCB7D6CF9B56EB2DD09919B53EF03" bold="true">A, E</emphasis>
<taxonomicName id="CC61EEC2042FA632F72BD26D41D39B1E" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="puskasi" status="sp. nov.">
<emphasis id="7B27045A59C9B5CC8BD73AB158A0F578" italics="true">L. puskasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3BF7804DC4E4F21591AE5DB4C14E1A2" rank="species">sp. nov.</taxonomicNameLabel>
, Kozara Mts, Bosnia &amp; Herzegovina
<emphasis id="D65AF1A5954FA41A0C87AF7897629F9B" bold="true">B, F</emphasis>
<taxonomicName id="93CA346F20A9A98A1A9A3CA3D351075A" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="visitor" status="sp. nov.">
<emphasis id="8D8E138F0910898CF3CF1D327D4726F6" italics="true">L. visitor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8235A8AE1528A7C8D8A2E03E46E91D4E" rank="species">sp. nov.</taxonomicNameLabel>
, Visitor Mts, Montenegro
<emphasis id="E1211E7681E0864D26E45008199BAD85" bold="true">C, G</emphasis>
<taxonomicName id="8DF01D015B60083BF2A1BEF7C053FF73" authority="Vincon &amp; Muranyi, 2007" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="A8D9E553557423677C3FB52633F39863" italics="true">L. dalmoni</emphasis>
<bibRefCitation id="EA0CE98699B0DB3D89E7AC168CC41EAA" author="Vinçon G &amp; Murányi D" issue="3" journalOrPublisher="Nouvelle Revue dEntomologie" pagination="237-248" refId="B82" refString="Vincon G, Muranyi D (2007) Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera). Nouvelle Revue d'Entomologie 23 (3): 237 - 248." title="Leuctra dalmoni, a new orophilic species with wide distribution in Europe (Plecoptera)." volume="23" year="2007">Vinçon &amp; Murányi, 2007</bibRefCitation>
</taxonomicName>
, Visitor Mts, Montenegro
<emphasis id="3F202F744E834315C44E7B3863B91017" bold="true">D, H</emphasis>
<taxonomicName id="16750FCB5DE867757F658FC086DF2298" authority=", Golija Mts" authorityName="Golija Mts" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="5525039FCBF5B12527D92942BAA9F35D" italics="true">L. dalmoni</emphasis>
, Golija Mts
</taxonomicName>
, Serbia
<emphasis id="C440F3D6BAE671ED4E1C6CABF41BC5CD" bold="true">I J</emphasis>
<taxonomicName id="52A933EBE40824EF47DF873AC679DB8D" authority="Kempny, 1899" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="62485E51826F15147E96E3DCB63982FB" italics="true">L. prima</emphasis>
<bibRefCitation id="349E484119062605EE4D209DA8B689DA" DOI="10.5962/bhl.part.24102" author="Kempny P" refId="B34" refString="Kempny P (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra - Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 49: 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102" year="1899">Kempny, 1899</bibRefCitation>
</taxonomicName>
, Börzsöny Mts, Hungary
<emphasis id="B4FACE2D44C4051760A501A5173FCBA6" bold="true">K L</emphasis>
<taxonomicName id="322364AFD6E2F646549851BCEFE16C21" authority="Aubert, 1954" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
<emphasis id="1A6D3F3B119017689462E112209E0BA0" italics="true">L. pseudosignifera</emphasis>
<bibRefCitation id="AAD5147A2DF0ABA04DA0D178602231B7" DOI="10.5169/seals-401211" author="Aubert J" issue="2" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pagination="124-136" refId="B2" refString="Aubert J (1954) Contribution a l'etude du genre Leuctra Stephens et description de quelques especes nouvelles de ce genre. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 27 (2): 124 - 136. https: // doi. org / 10.5169 / seals- 401211" title="Contribution à létude du genre Leuctra Stephens et description de quelques espèces nouvelles de ce genre." volume="27" year="1954">Aubert, 1954</bibRefCitation>
</taxonomicName>
, Poljana Mts, Slovakia. Scale bars: 1 mm (
<emphasis id="C96C03473DA712FECF8B3AC09B9864FE" bold="true">D</emphasis>
for
<emphasis id="42E561F08BBF4573513BD7021F71CAD2" bold="true">A D, I, K; E</emphasis>
for
<emphasis id="4BFF980DE183E6EE6FC4B70189F99764" bold="true">E H, J, L</emphasis>
).
</paragraph>
</caption>
<paragraph id="D290DBB28C6F117BC2994F8F6A333D40">
Female abdomen (Figs
<figureCitation id="8AEB333271813C1467DBA4D94DEAD137" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Terminalia of Leuctra visitor sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D male paraprocts and specillae, caudal view E female terminalia, ventral view F same, lateral view. Setation omitted with the exception of subgenital plate on F. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure10" httpUri="https://binary.pensoft.net/fig/1175461">10 E, F</figureCitation>
,
<figureCitation id="87C7F6CADD900EBC2D0B6FF141438CA7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 F</figureCitation>
): Terga I VIII with transverse row of four pigmented spots clearly visible; terga I VIII mostly membranous but with lateral sclerites, terga II, III, and VI VIII, sometimes also tergite V, bear a patch-like medial sclerite, largest on tergite VIII; tergite IX membranous only anterolaterally, tergite X fully sclerotised. Sterna I VII simple, sterna II VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large wide lobes, slightly darker brown than remainder of the plate; posterior margin convex, incision between the lobes is a membranous strip, slightly widened towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is wide in ventral view, distinctly raised, nose-shaped in lateral view, with distinct erect setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus, and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging posterior teeth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZ1BG" type="Affinities">
<paragraph id="D5A73455103211B7BFE26FBE5F88C6A0">
<heading id="974B8785FFE8B99FD3FD7FCDD12AF52B" reason="title">Affinities.</heading>
</paragraph>
<paragraph id="CAA18A149D315EDD1DA546E30FDB2A69">
The males are morphologically closest to
<taxonomicName id="6E9A988BD8D9543D276821D647D62AC8" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="CA5D84D46B6FA1FEBF830C19E081A347" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(compare with Fig.
<figureCitation id="25F6356425B5D3BEB4B0429ED08BBCB9" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 C, D</figureCitation>
) while the female is closest to
<taxonomicName id="4C201A3A1D06853DD7BC56AF08183B2E" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="35AAF42BE7E35E087969ECB058C40165" italics="true">L. prima</emphasis>
</taxonomicName>
(compare with Fig.
<figureCitation id="A611E44242B7F28CF9E9DCD972BBFC79" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 J</figureCitation>
). Males can be distinguished from
<taxonomicName id="CFEE0C55A5BE6534973C4A2E551EA910" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="A8E58FF19F07FD86C00E54699864AB9D" italics="true">L. dalmoni</emphasis>
</taxonomicName>
on the basis of the following characters: a weak crossband on the membranous area of tergite VIII lacking in
<taxonomicName id="97461D5890CD70A67FC0F85ACC2B04CB" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="B8AB1BD80C24FF8DC1886E09F0FAC02C" italics="true">L. dalmoni</emphasis>
</taxonomicName>
, angular teeth of the posteromedial process of tergite VIII stronger than those of
<taxonomicName id="B2922D707A71D70E521C3C052771AB4E" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="948E347F4ACA5B46A29F01A22DD761FF" italics="true">L. dalmoni</emphasis>
</taxonomicName>
; the posterior indentation of tergite X is V-shaped, while it is rounded in
<taxonomicName id="3BC2B0B9749D67087850E9ACE79B1F2C" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="E4AA85190E39CA04603F01BEC54E6A7D" italics="true">L. dalmoni</emphasis>
</taxonomicName>
; the ventral vesicle of sternite IX is shorter than 1 / 2 the segment length, while clearly longer than 1 / 2 the segment length in
<taxonomicName id="70337ED9A4B145864F9C0AB324ED8F57" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="A48A1F66B13344F3E2DBF4C01F61614D" italics="true">L. dalmoni</emphasis>
</taxonomicName>
. The males of the new species can be easily distinguished from
<taxonomicName id="06715D5F819D2225D21E9EB9668F93BA" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="0DE93DFCAB063A6D685C81126446C4F6" italics="true">L. prima</emphasis>
</taxonomicName>
and
<taxonomicName id="2DAFBEEF6B99E872B45EEC5C9A7265D2" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
<emphasis id="5B9599C0C1D3F367348F8FAAC98538B4" italics="true">L. pseudosignifera</emphasis>
</taxonomicName>
:
<taxonomicName id="09806C6A065CEB87C875414F88D247E3" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
<emphasis id="101499A749CDFBFE9AAFC80391ABECEA" italics="true">L. pseudosignifera</emphasis>
</taxonomicName>
is characterised by a pair of rounded sclerotised expansions, located between the posteromedial process and segment sides of tergite VIII (Fig.
<figureCitation id="886B6365343A5CC9A5A7B93A41B41BB9" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 K</figureCitation>
); the posteromedial process of tergite VIII is erect in lateral view in
<taxonomicName id="117CBC981FFC5D5B5FA97B0DE8B42294" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="8B0CF75586C73F8A2E44C7FDA9986F85" italics="true">L. prima</emphasis>
</taxonomicName>
; furthermore, this species has a divided posteromedial sclerite on tergite IX (Fig.
<figureCitation id="4C66AA10B2758C5919C1762529AEE155" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 I</figureCitation>
). Females can be distinguished from
<taxonomicName id="A8D5C866B957C719E40AE56F9764F6DB" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B8550F770306850A5B18EFEB64290FE6" italics="true">L. prima</emphasis>
</taxonomicName>
on the basis of the distinct, erect hairs of the subgenital plate as seen in lateral view, and the nose-shape of the bulge of the subgenital plate; the bulge is rounded in
<taxonomicName id="C140B4466DD13A9739F43DD93E5E46D7" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="2733C977CBAB9D5BC4F2896C2A6FF34F" italics="true">L. prima</emphasis>
</taxonomicName>
and the subgenital plate lacks distinct setation. The wide bulge of the subgenital plate (in ventral view), and the erect hairs (in lateral view) distinguish the new species from
<taxonomicName id="5F0F3F680865A495D1D02C50DFB422B2" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="84418B16184B3CC2218A05C0EA7796F2" italics="true">L. dalmoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4083E7513D79DA3F0D225A0430B7DDA4" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
<emphasis id="393FA3209847260EA91E0DD11B526B5A" italics="true">L. pseudosignifera</emphasis>
</taxonomicName>
; furthermore,
<taxonomicName id="5FE9D11FA0FEBBA87B04896D0347F824" authorityName="Aubert" authorityYear="1954" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pseudosignifera">
<emphasis id="E235F4C69FD5CF9BEF51BD8C08FACDDA" italics="true">L. pseudosignifera</emphasis>
</taxonomicName>
is characterised by a triangular membranous field at the end of the incision between the lobes of the subgenital plate (Fig.
<figureCitation id="455C7A2757C9588CA5B755C0BB61B258" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 L</figureCitation>
). Distinction of both sexes from
<taxonomicName id="DCF39647F37E8A44A3CFF5FCCF5981CF" authorityName="Murányi &amp; Kovács" authorityYear="2024" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="puskasi">
<emphasis id="1C76928453E7E69C03331BCBBA2CBDA3" italics="true">L. puskasi</emphasis>
</taxonomicName>
was described in detail above.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPBAI" type="distribution">
<paragraph id="065551569A4C4FBEBDC9CAA05B8DAF51">
<heading id="6FEF3BF9056C80D77336CFFBFA203D33" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="BC08BCD120216946CB2FAEB678C0166C">
The species was collected at three high-elevation brooks and spring outlets in the Visitor Mts of eastern
<collectingCountry id="E2B3D20546F2427466CFEB6037CE6C5F" name="Serbia">Montenegro</collectingCountry>
(Fig.
<figureCitation id="155558EDF86FEE9AF0FEAA1C97477F7B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the four new Leuctra species." figureDoi="10.3897/zookeys.1218.120744.figure1" httpUri="https://binary.pensoft.net/fig/1175452">1</figureCitation>
). The Visitor is a mostly volcanic range of small area, with its highest peak reaching more than 2200 meters, surrounded by the notably higher limestone chains of the Prokletije Mts. The Visitor receives high precipitation and is covered with dense forest, resulting in a rich pattern of different waterflows. The streams where the new species was found are running in mixed forest or partly open meadows, the lentic parts usually with dense
<taxonomicName id="B02BB6A66B454AC15A5F7EBFE45FA92C" class="Magnoliopsida" family="Ranunculaceae" genus="Caltha" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="genus">
<emphasis id="F0AF79D53B297D96A7750098222D7C9C" italics="true">Caltha</emphasis>
</taxonomicName>
growth (Fig.
<figureCitation id="E5737B3EB5E12E54733218CE67D3096E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 D, E</figureCitation>
). The brooks have variable flow rates but are permanent streams, the substrate is stony mixed with mud and plenty of dead wood. The specimens were emerging together with
<taxonomicName id="B88FE4D6F9C52EB4C536E768320CA269" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="AE4DB61C9E6DA8221BE01CF0072566C2" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="350350234022DDC8CD3B62F5C44F8399" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 C, G</figureCitation>
) and both species were present with many mating pairs in May, but we found no interbreeding couples (Fig.
<figureCitation id="87FD4B89A46BEACAF328A9A01A245F85" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus and habitat of Leuctra visitor sp. nov. A male habitus B female habitus C copula D partly open brook at 1845 m in the Visitor Mts. E type locality: forest springs and their outlet at 1730 m in the Visitor Mts." figureDoi="10.3897/zookeys.1218.120744.figure11" httpUri="https://binary.pensoft.net/fig/1175462">11 C</figureCitation>
). We visited the area three times: the new species was more numerous than
<taxonomicName id="BE947D33ABA02722E14C397D6F80BE8F" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="3812CA66113414FBE658EF81F1C6A658" italics="true">L. dalmoni</emphasis>
</taxonomicName>
in early
<date id="D9F49ED0DEACA63C330E6F4A2235467D" value="2023-05">May 2023</date>
, both scarce in early
<date id="B6896E2A14E89B0F9C91895BB00766B7" value="2022-06">June 2022</date>
, and both absent by late
<date id="EBA4BCDDD5D868E782B8E8C0C5C928B1" value="2023-06">June 2023</date>
. In May, the upper site (
<quantity id="9E0BF113DA1A6B643F3B2EBE463CCCB6" metricMagnitude="3" metricUnit="m" metricValue="1.845" unit="m" value="1845.0">1845 m</quantity>
) was not available, and even the lowest site (
<quantity id="3A8FFA6305176D5EB7F4A91B5386B0C2" metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">1515 m</quantity>
) was partly covered by snow, but all were snow-free by late June. We found a rich accompanying stonefly fauna at the lowest site:
<taxonomicName id="DD48C4F87FD5B9117A495A1A96F43F8B" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="helenica">
<emphasis id="7D045A2B6443AB82505B7783242DFE83" italics="true">Brachyptera helenica</emphasis>
</taxonomicName>
(adults and matured larvae, early June),
<taxonomicName id="6FE5F8F313730AF2147B2FCE51325A9D" authority="Berthelemy, 1971" authorityName="Berthelemy" authorityYear="1971" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="2409A5E9B5E2605D9D45E034161513D9" italics="true">B. graeca</emphasis>
<bibRefCitation id="CDE093F91C9CD253D98458BDB200429E" author="Berthélemy C" issue="1" journalOrPublisher="Biologia Gallo-Hellenica" pagination="23-56" refId="B6" refString="Berthelemy C (1971) Plecopteres de Grece Centrale et d'Eubee. Biologia Gallo-Hellenica 3 (1): 23 - 56." title="Plécoptères de Grèce Centrale et dEubée." volume="3" year="1971">Berthélemy, 1971</bibRefCitation>
</taxonomicName>
(adults, late June),
<taxonomicName id="0DA1D48AC6F7B6F290D611DDA8E77241" baseAuthorityName="Klapalek" baseAuthorityYear="1902" class="Insecta" family="Taeniopterygidae" genus="Brachyptera" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="2C5CCD8DC72FD72DF7CB7137366375A2" italics="true">B. seticornis</emphasis>
</taxonomicName>
(adults, early June),
<taxonomicName id="75934D95CFF62D8C93A11C94DF4E3212" authorityName="Pictet" authorityYear="1841" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="C6D87CAC9E4F1CC1EAA792F7BA3E98FB" italics="true">Capnia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDDF631E5A4C587AACA5512C6F7034EF" sensu="lato">s. l.</taxonomicNameLabel>
<emphasis id="FA549CD6270AEB2532867943510289FC" italics="true">vidua rilensis</emphasis>
(adults, May and early June),
<taxonomicName id="DA428B8DA01A0AD228A764EBB9379C11" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="170A038E30186E5616AFFED049154A80" italics="true">Leuctra nigra</emphasis>
</taxonomicName>
(adults, May and June),
<taxonomicName id="02C5DFE114621CFDCB7A35192EEEBC5E" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="C64E7C2DC022179F3DDAF93AE1CF38A5" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(adults, May and early June),
<taxonomicName id="C8704489FD2E7C9B1DB23667C1C3C603" authority="Aubert, 1956 b" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
<emphasis id="3486936DC74B6D231EBF69A889D58221" italics="true">L. metsovonica</emphasis>
Aubert, 1956 b
</taxonomicName>
(adults, early June),
<taxonomicName id="894A978CC086F884BCA3E32E99A40AC5" authority="Illies, 1954 b" authorityName="Illies" authorityYear="1954" class="Insecta" family="Nemouridae" genus="Protonemura" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="auberti">
<emphasis id="FA551445A21891E033A87DD17F68DCD8" italics="true">Protonemura</emphasis>
cf.
<emphasis id="A5C9E768F01E71DD82271AB8ED8A69F7" italics="true">auberti</emphasis>
Illies, 1954 b
</taxonomicName>
(adults and matured larvae, early June),
<taxonomicName id="C423729F2F5D7557D2072A9F53F4AB9E" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
<emphasis id="D7F2E5C4AEBA05A6D7BD95B3B1F161F7" italics="true">Nemurella pictetii</emphasis>
</taxonomicName>
(adults, May and June),
<taxonomicName id="5599EF53973BF50994E8AB4DDD700FFA" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="subSpecies" species="cinerea" subSpecies="cinerea">
<emphasis id="CD4B5763DC462A4B16707AD5AB3F0D1B" italics="true">Nemoura cinerea cinerea</emphasis>
</taxonomicName>
(adults, early June),
<taxonomicName id="03C8CF6157734147B483DC11236A3A53" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="6B34C54D13F66C17479213CB553ADC4B" italics="true">Nemoura marginata</emphasis>
</taxonomicName>
(adults, May and early June), a yet unnamed
<taxonomicName id="173E7DBBE25AFA13780A0093908A533B" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F18E0A04742C351753657D0963169CD7" italics="true">Nemoura</emphasis>
</taxonomicName>
of the
<taxonomicName id="6151C2CFD1E975027FD4A6BD0FF45819" authorityName="Pictet" authorityYear="1836" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="8B8DE4448B75D309AF3DB9F579412A91" italics="true">marginata</emphasis>
</taxonomicName>
group (adults, May and early June),
<taxonomicName id="ED7841E01AE8D6A7CD8455005943F4CF" authority="Despax, 1934" authorityName="Despax" authorityYear="1934" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="4299179A632190A1BBA1EF5B3466929B" italics="true">N. uncinata</emphasis>
<bibRefCitation id="0741902B260B38F224D4F5DD2993F288" author="Despax R" issue="2" journalOrPublisher="Bulletin de la Société dhistoire Naturelle de Toulouse" pagination="255-270" refId="B15" refString="Despax R (1934) Plecopteres pyreneens. VII. Etude et description de quelques formes de Nemoures apparentees a Nemoura marginata (Pict.). Bulletin de la Societe d'histoire Naturelle de Toulouse 66 (2): 255 - 270." title="Plécoptères pyrénéens. VII. Etude et description de quelques formes de Nemoures apparentées à Nemoura marginata (Pict.)." volume="66" year="1934">Despax, 1934</bibRefCitation>
</taxonomicName>
(adults, May and early June),
<taxonomicName id="1A9765BADA5CF48B39C19C72F84C703E" authority="(McLachlan, 1872)" baseAuthorityName="McLachlan" baseAuthorityYear="1872" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="0D911BA1724C56F38C1F5DB96FCF04AF" italics="true">Arcynopteryx dichroa</emphasis>
(
<bibRefCitation id="BEC3C26269CBE82CB2FBDB560EC262F7" author="McLachlan MR" refId="B42" refString="McLachlan MR (1872) Materiaux pour une Faune Nevropterologique de l'Asie Septentrionale. Seconde partie. Non-Odonates. Annales de la Societe Entomologique Belgique 15: 47 - 77 [+ Pl I - II]. https: // www. biodiversitylibrary. org / item / 44639 # page / 345 / mode / 1 up" url="https://www.biodiversitylibrary.org/item/44639#page/345/mode/1up" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
(adults and matured larvae, June), a yet unnamed
<taxonomicName id="92E2D54F65CB9307712B4D863F42AA98" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="BF05F4D88D9A1A0D0C4116DA859D8F16" italics="true">Isoperla</emphasis>
</taxonomicName>
related to
<taxonomicName id="A9434D4203216BB2FEFF257EDDBE3A80" authority="Ikonomov, 1980" authorityName="Ikonomov" authorityYear="1980" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="breviptera">
<emphasis id="B0B32366453D3E2B1C2C9718225FAC8D" italics="true">I. breviptera</emphasis>
<bibRefCitation id="8D5B229117C4BBCF9BE71C3250C04E97" author="Ikonomov P" issue="4" journalOrPublisher="Fragmenta Balcanica" pagination="19-31" refId="B23" refString="Ikonomov P (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica 11 (4): 19 - 31." title="Nouvelles espèces de Plécoptères (Insecta, Plecoptera) de Macedoine. II." volume="11" year="1980">Ikonomov, 1980</bibRefCitation>
</taxonomicName>
(adults and matured larvae, June),
<taxonomicName id="6560C470B35358458E34D6A44030C15D" authority="Illies, 1954 a" authorityName="Illies" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" isUncertain="true" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="tripartita">
<emphasis id="C2F03F58BB6DF062B5F9D7C9C5DB3D8C" italics="true">I.</emphasis>
cf.
<emphasis id="350278A2B960EA2581C223E79B15CC80" italics="true">tripartita</emphasis>
Illies, 1954 a
</taxonomicName>
(larvae, June), and unidentified larvae of
<taxonomicName id="50E4D8922937B8093D5A29C9630D4C10" authorityName="Zwick" authorityYear="1967" class="Insecta" family="Chloroperlidae" genus="Siphonoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="FF01DA61DCE5B14D8C8F177A67756BB2" italics="true">Siphonoperla</emphasis>
</taxonomicName>
and
<taxonomicName id="912B1C73E7A6A6B955DF81E9545F8CD1" authorityName="Newman" authorityYear="1836" class="Insecta" family="Chloroperlidae" genus="Chloroperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F2711B112D349CE9C69513B48E69075E" italics="true">Chloroperla</emphasis>
</taxonomicName>
(early June). Some of these were probably not developing in the same stream section since the site is placed at a junction of three different brooks. The type locality (sampled only in May) was inhabited by the new species,
<taxonomicName id="E52237FC5D176B31CCA822B725536469" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="626502107001B9CFB050422DB3DEE77B" italics="true">L. nigra</emphasis>
</taxonomicName>
(adults),
<taxonomicName id="7944E6EB762189FFFF04D8A179825141" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="180FB8333F12430AAFE82274EAF991B3" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(adults), the unnamed
<taxonomicName id="73FB3CBD58431AA556AFC805ED495671" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="055FF7C4B7B7C54F9ADFF38175643A48" italics="true">Nemoura</emphasis>
</taxonomicName>
(adults and putative larvae), an unidentified
<taxonomicName id="C25BDB2A01E8012E81B223F44A8FCEBF" authorityName="Kempny" authorityYear="1898" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="DECE3D441B488E6C08386D08E659E4FD" italics="true">Protonemura</emphasis>
</taxonomicName>
(larvae), and the unnamed
<taxonomicName id="3FBAAA392CBB4DAB0EC1DEB4A47183AC" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="BA64DF64179A8105FF2F91525319508A" italics="true">Isoperla</emphasis>
</taxonomicName>
(larvae). At the highest site (sampled only in June), the new species was found together with
<taxonomicName id="FFDC65054F7BDF717646491F5213339A" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="1572F4A464A5491B4FB71C94C479A42C" italics="true">Leuctra nigra</emphasis>
</taxonomicName>
(adults),
<taxonomicName id="18644824EB4427F453D11083E0D38886" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="EE46DF706F551CB566A3CD6B193C0C67" italics="true">L. dalmoni</emphasis>
</taxonomicName>
(adults), the unnamed
<taxonomicName id="51CF757CCD7FBCA7766B0AEB917309AE" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="02B40628AB06D830B9173E5B5B001371" italics="true">Nemoura</emphasis>
</taxonomicName>
(adults),
<taxonomicName id="0C25CA801038F31E4D339C1CC6E3A9A8" baseAuthorityName="Klapalek" baseAuthorityYear="1900" class="Insecta" family="Nemouridae" genus="Nemurella" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="pictetii">
<emphasis id="9748A27E3CC5E8BE9F857F885F759C70" italics="true">Nemurella pictetii</emphasis>
</taxonomicName>
(adults),
<taxonomicName id="7A8CF850FC7169B54C9E5D75FE925029" baseAuthorityName="McLachlan" baseAuthorityYear="1872" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="ED640DC5B1FA83EC8BDE816B29854774" italics="true">Arcynopteryx dichroa</emphasis>
</taxonomicName>
(larvae), and the unnamed
<taxonomicName id="2C9E226BCA0B30B3742630C524244F80" authorityName="Banks" authorityYear="1906" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus">
<emphasis id="E2394672E1ACF4BDC38AB4D462EEC0E8" italics="true">Isoperla</emphasis>
</taxonomicName>
(adults).
<taxonomicName id="0FA3C487240C94270A75B1331193E273" authorityName="Pictet" authorityYear="1841" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="BC623DD730D0E48D660DB2950315E30D" italics="true">Capnia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84063059A86F432DFCA463949E6003E5" sensu="lato">s. l.</taxonomicNameLabel>
<emphasis id="A1EBB56E7F428E411608102E2B9E7162" italics="true">vidua rilensis</emphasis>
,
<taxonomicName id="3D01587F41233D08821768DC44401C7B" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="D9F4F564CE8B58BE0DC8123E148C977C" italics="true">Leuctra dalmoni</emphasis>
</taxonomicName>
, and
<taxonomicName id="D4DED9B6BFD84D73681DCA9C0B820C29" authorityName="Aubert" authorityYear="1956" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="metsovonica">
<emphasis id="93B2FCA78ED12B0642A338A12F68EAC7" italics="true">L. metsovonica</emphasis>
</taxonomicName>
are new records for
<collectingCountry id="72A27A035B61EF21A60A4E9C60210FED" name="Serbia">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHOAI" type="etymology">
<paragraph id="1D2ED37F908BE3B6844D22B741E5173D">
<heading id="599A6BAE8E221A8ECF8660F756BB3C96" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9F4BF4A614825BDBE86B169A41CBAEA3">
The name
<emphasis id="09E38B3F1BA492B4D8C7D6186AF75304" italics="true">visitor</emphasis>
is derived from the Visitor Mountains of
<collectingCountry id="595FF87CF29996ECFE57CEEED6223810" name="Serbia">Montenegro</collectingCountry>
, where the new species was found and probably restricted to the high elevations of the range. Used as a noun, gender neutral.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,46 +1,46 @@
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<mods:namePart id="F8C3357207642BAAB55B9BF0BE148E73">Murányi, Dávid</mods:namePart>
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<mods:affiliation id="DC8E4650C235BD4893866161C92CEFD8">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
<mods:namePart id="A78D410C617F74A9D64869E8F72C82F2">Murányi, Dávid</mods:namePart>
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<mods:affiliation id="92F241CE0B0AC2CBF194754C22A511C9">Department of Zoology, Eszterházy Károly Catholic University, Leányka út 6, Eger H- 3300, Hungary</mods:affiliation>
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<mods:namePart id="9CEF06FD7D30556F8C9477F7FB272930">Kovács, Tibor</mods:namePart>
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<mods:affiliation id="337143A993AAA40582F505F238B2B8B4">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
<mods:namePart id="27C91C7F22F1CA4824E8763FB907649D">Kovács, Tibor</mods:namePart>
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<mods:affiliation id="2D395B05A44AE09C4CFFCF4B5B6BABC3">Mátra Museum of the Hungarian Natural History Museum, Kossuth Lajos u. 40, Gyöngyös H- 3200, Hungary</mods:affiliation>
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<subSubSection id="3001862477E418D038970AA1086F9B3F" type="nomenclature">
@ -68,39 +68,40 @@ Murányi &amp; Kovács
<heading id="BA9EC575EF4E8F152AD14DC6D41BABAE" reason="title">Type material.</heading>
</paragraph>
<paragraph id="518CC0EEF550530E9046C2530F82CE6A">
<materialsCitation id="2DC30931E8455274475C51977B9A5079" collectingDate="2012-03-16" collectionCode="T, HNHM" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 4185" specimenCount="1" typeStatus="holotype">
<materialsCitation id="2DC30931E8455274475C51977B9A5079" collectingDate="2012-03-16" collectionCode="HNHM" collectorName="T. Kovacs &amp; D. Muranyi &amp; G. Puskas" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 4185" specimenCount="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="holotype">
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<typeStatus id="1ADBCAF2A43C45EFC0FFE7F21EA9AC43" type="holotype">Holotype</typeStatus>
</emphasis>
male:
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:
<collectingCountry id="7EFD3E67E65C5EBFBA3C7118524902BF" name="Bosnia and Herzegovina">Bosnia &amp; Herzegovina</collectingCountry>
<location id="E0A69505027825BDE8D44DB4A7B4F8E2" LSID="urn:lsid:plazi:treatment:B7518924CBFD591FA01FC896BA1EA93B:E0A69505027825BDE8D44DB4A7B4F8E2" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" latitude="45.0414" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" name="Republika Srpska">Republika Srpska</location>
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,
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, forest brook along the
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road,
,
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,
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<elevation id="F5EB13A793FC0EC4FAF8491114F5E4F2" metricMagnitude="2" metricUnit="m" metricValue="5.95" unit="m" value="595.0">595 m</elevation>
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,
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<geoCoordinate id="18E24858F79CDDEA5EE7FF2C5230A8D5" degrees="45.0414" direction="north" orientation="latitude" precision="5" value="45.0414">45.0414 ° N</geoCoordinate>
<geoCoordinate id="18E24858F79CDDEA5EE7FF2C5230A8D5" degrees="45.0414" direction="north" orientation="latitude" precision="5" value="45.0414">45.0414°N</geoCoordinate>
,
<geoCoordinate id="4DDC80F53122873B849A22BDD41106B2" degrees="16.9040" direction="east" orientation="longitude" precision="5" value="16.904">16.9040 ° E</geoCoordinate>
<geoCoordinate id="4DDC80F53122873B849A22BDD41106B2" degrees="16.9040" direction="east" orientation="longitude" precision="5" value="16.904">16.9040°E</geoCoordinate>
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,
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</date>
(field number: loc. 6), leg.
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.
<collectorName id="CF3557FDF07EF46BF89555C330C0E954">Kovács, D</collectorName>
.
<collectorName id="4F464BC0E6B9123FEC8555435450A9A8">Murányi, G</collectorName>
. Puskás (
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,
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,
<collectorName id="5F4C1EA53C8CED358FBFAB9240774660">G. Puskás</collectorName>
(
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<collectionCode id="BA1C2A6CDDDAC1042A99C03CF672257C" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
@ -109,7 +110,7 @@ road,
)
</materialsCitation>
.
<materialsCitation id="5030C2568A25D6ED99411158393277D2" collectingDate="2012-03-16" collectionCode="BYU" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<materialsCitation id="5030C2568A25D6ED99411158393277D2" collectingDate="2012-03-16" collectionCode="BYU" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
<emphasis id="48B47BFD1CA335F40C08A7B289E0B813" bold="true" italics="true">
<typeStatus id="94CC5C7768A6158C84D0EEF56F1DB841" type="paratype">Paratypes</typeStatus>
</emphasis>
@ -123,7 +124,7 @@ road,
)
</materialsCitation>
<materialsCitation id="DA18DFC1D5883CF45CDD2ABA8C4DE8E1" collectingDate="2012-03-16" collectionCode="CGV" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="Republika Srpska" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<materialsCitation id="DA18DFC1D5883CF45CDD2ABA8C4DE8E1" collectingDate="2012-03-16" collectionCode="CGV" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
<specimenCount id="6A4C0A57638671E71C373013F2DF843F" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="9F54000A6F14D6C31859E2D15F11BA09" count="1" type="female">1 ♀</specimenCount>
(
@ -133,7 +134,7 @@ road,
)
</materialsCitation>
<materialsCitation id="6556CEE5986DFD6D3E12021A7C19DBAC" collectionCode="HNHM" country="Bosnia and Herzegovina" specimenCode="PLP 3853" specimenCount="14" specimenCount-female="4" specimenCount-male="10" typeStatus="paratype">
<materialsCitation id="6556CEE5986DFD6D3E12021A7C19DBAC" collectingDate="2012-03-16" collectionCode="HNHM" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="PLP 3853" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Republika Srpska" typeStatus="paratype">
<specimenCount id="FFC84BB845E61235B3F0038CA61BFF8E" count="10" type="male">10 ♂</specimenCount>
<specimenCount id="3698B56407C9DD52C3FFDAD5D8D0A30C" count="4" type="female">4 ♀</specimenCount>
(
@ -145,17 +146,21 @@ road,
)
</materialsCitation>
<materialsCitation id="6B8B5A63AD77561F29857FADE5E08B82" collectionCode="MM" country="Bosnia and Herzegovina" specimenCount="17" specimenCount-female="5" specimenCount-male="12" typeStatus="paratype">
<materialsCitation id="6B8B5A63AD77561F29857FADE5E08B82" collectingDate="2012-03-16" collectionCode="MM" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCode="2012-10, PLETYP-36" specimenCount="17" specimenCount-female="5" specimenCount-male="12" stateProvince="Republika Srpska" typeStatus="paratype">
<specimenCount id="CB5B5E993D7B2F8FDA66D67736658085" count="12" type="male">12 ♂</specimenCount>
<specimenCount id="14B57FDBAC4668AD07D3467C10FE6694" count="5" type="female">5 ♀</specimenCount>
(
<named-content id="45AD23B29E824D51AC88FEF3324A183B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum" xlink_title="Mátra Museum of the Hungarian Natural History Museum, Gyöngyös, Hungary">
<collectionCode id="56F001848ED40A23AB887F5892D637E1" country="France" name="University of Montpellier">MM</collectionCode>
</named-content>
: 2012-10, PLETYP- 36)
:
<specimenCode id="B543C76CE86B469B25AEFC1A1B9C6088">2012-10</specimenCode>
,
<specimenCode id="7C673B3EAD26FE246198F37385A23DDE">PLETYP-36</specimenCode>
)
</materialsCitation>
<materialsCitation id="AB95F7C73A3D87FC085811582410602B" collectionCode="SMNS" country="Bosnia and Herzegovina" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<materialsCitation id="AB95F7C73A3D87FC085811582410602B" collectingDate="2012-03-16" collectionCode="SMNS" collectorName="Kovacs, D &amp; Muranyi, G" country="Bosnia and Herzegovina" county="Gornji Podgradci-Kozarac" elevation="595" isTailEnumeration="true" latitude="45.0414" location="forest brook along the Gornji Podgradci-Kozarac road" longLatPrecision="6" longitude="16.904" municipality="Kozara Mts" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Republika Srpska" typeStatus="paratype">
<specimenCount id="F9F8FA8E5AC4048F4354E85EEB471194" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="34596C539C68B38D1C57B5B9B3B611C9" count="1" type="female">1 ♀</specimenCount>
(
@ -207,17 +212,7 @@ Male abdomen (Figs
<figureCitation id="99A73570751D5C5177EDC13C0E5DBFDA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Terminalia of Leuctra puskasi sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D variation of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Setation omitted with the exception of subgenital plate on G. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure8" httpUri="https://binary.pensoft.net/fig/1175459">8 A E</figureCitation>
,
<figureCitation id="D8F873418A09FE79297D9F18F5952505" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 A</figureCitation>
): Tergite I sclerotised only anteriorly and in lateral stripes, with two medial spots. Transverse row of four pigmented spots distinct on terga
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VII. Terga
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and
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with medially divided antecosta, tergite
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with triangular membranous area anteriorly. Terga IV VII with entire antecosta and full sclerotised, tergite VII with weakly sclerotised posterior edge and small membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends into a small triangular plate; the membranous area is ~ 1 / 3 segment width, and usually do not reach the 1 / 2 of the segment length but in some specimens it reach down to 2 / 3 above the posteromedial process, posterior margin of the membranous portion wavy; the surrounding sclerotisation of the membranous area lighter than the rest of the segment; posteromedial process not darker than the lateral areas of the segment, not erect in side view and not or only slightly overhangs the segment posterior edge, nearly as wide as the membranous area above, bilobed with rounded lobes slightly pointed laterad, the space between the lobes is concave to nearly straight; posterior margin between the posteromedial process and segment sides nearly straight, not indenting nor distinctly lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 to 1 / 2 of its length; posteromedial sclerite short but wide, sublateral areas darker, the medial portion usually narrower than the lateral portions. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep notch, not wider than 1 / 3 tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk short. Cercus simple, covered with long setae. Sterna
<collectionCode id="1DB1A43BE5D2C2D7B810E5A53BBE8CC1">II</collectionCode>
VIII simple, sternite IX bears a vesicle shorter than 1 / 2 segment length, its width more than 1 / 2 its length; posteromedial 1 / 2 of the sternum is delimited by a pale and weakly sclerotised area. Paraproct with moderately wide base, abruptly narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, blunt triangular process. Specillum longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, sharp tip.
): Tergite I sclerotised only anteriorly and in lateral stripes, with two medial spots. Transverse row of four pigmented spots distinct on terga II VII. Terga II and III with medially divided antecosta, tergite II with triangular membranous area anteriorly. Terga IV VII with entire antecosta and full sclerotised, tergite VII with weakly sclerotised posterior edge and small membranous portion. Tergite VIII: antecosta interrupted by a wide membranous area, each part ends into a small triangular plate; the membranous area is ~ 1 / 3 segment width, and usually do not reach the 1 / 2 of the segment length but in some specimens it reach down to 2 / 3 above the posteromedial process, posterior margin of the membranous portion wavy; the surrounding sclerotisation of the membranous area lighter than the rest of the segment; posteromedial process not darker than the lateral areas of the segment, not erect in side view and not or only slightly overhangs the segment posterior edge, nearly as wide as the membranous area above, bilobed with rounded lobes slightly pointed laterad, the space between the lobes is concave to nearly straight; posterior margin between the posteromedial process and segment sides nearly straight, not indenting nor distinctly lobed. Tergite IX mostly membranous, antecosta interrupted in the medial 1 / 3 to 1 / 2 of its length; posteromedial sclerite short but wide, sublateral areas darker, the medial portion usually narrower than the lateral portions. Anterior margin of tergite X bilobed anteriorly, posterior margin with deep notch, not wider than 1 / 3 tergite width. Epiproct large, posteriorly rounded, sclerotised only at its sides, stalk short. Cercus simple, covered with long setae. Sterna II VIII simple, sternite IX bears a vesicle shorter than 1 / 2 segment length, its width more than 1 / 2 its length; posteromedial 1 / 2 of the sternum is delimited by a pale and weakly sclerotised area. Paraproct with moderately wide base, abruptly narrowing after basal 1 / 3, apex gently curved in lateral view and tapering towards a sharp tip. Base of paraproct connected to a subrectangular lateral expansion with an apical, blunt triangular process. Specillum longer than the paraproct, gently curved in lateral view and ending in a subterminally constricted, sharp tip.
</paragraph>
<caption id="6404BCBEEF92B2192C1250928B7726F6" ID-DOI="10.3897/zookeys.1218.120744.figure8" ID-arpha="74A8422E-EEA4-5FD7-B500-E04DB1941AC0" httpUri="https://binary.pensoft.net/fig/1175459" startId="F8">
<paragraph id="065AD1DE22E29BB8D344F05846D9F354">
@ -274,13 +269,7 @@ Female abdomen (Figs
<figureCitation id="3BA23C40155AC557F5A6E77793D018C9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Terminalia of Leuctra puskasi sp. nov. A male terminalia, dorsal view B same, lateral view C same, ventral view D variation of male tergite 8, dorsal view E male paraprocts and specillae, caudal view F female terminalia, ventral view G same, lateral view. Setation omitted with the exception of subgenital plate on G. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1218.120744.figure8" httpUri="https://binary.pensoft.net/fig/1175459">8 F, G</figureCitation>
,
<figureCitation id="AC8F464624BD2164516EBEBDC32BAF59" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Terminalia of five Balkan members of the Leuctra prima group A D, I, K male terminalia, dorsal view E H, J, L female terminalia, ventral view A, E L. puskasi sp. nov., Kozara Mts, Bosnia &amp; Herzegovina B, F L. visitor sp. nov., Visitor Mts, Montenegro C, G L. dalmoni Vinçon &amp; Murányi, 2007, Visitor Mts, Montenegro D, H L. dalmoni, Golija Mts, Serbia I J L. prima Kempny, 1899, Börzsöny Mts, Hungary K L L. pseudosignifera Aubert, 1954, Poljana Mts, Slovakia. Scale bars: 1 mm (D for A D, I, K; E for E H, J, L)." figureDoi="10.3897/zookeys.1218.120744.figure12" httpUri="https://binary.pensoft.net/fig/1175463">12 E</figureCitation>
): Terga I VIII with transverse row of four pigmented spots clearly visible; terga I IV mostly membranous but with lateral sclerites, terga
<collectionCode id="3102681D03D3F090D8AC27E79FF36778" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
VIII with medial sclerite as well; medial sclerite is a longitudinal stripe on tergite
<collectionCode id="025B1D2EBB3F8DBE1E074BD3C1CD57A7" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
, gradually widens on the following segments and as wide as 1 / 2 segment width on tergite VIII; terga IX X fully sclerotised. Sterna I VII simple, sterna
<collectionCode id="CB70F36E4200B92027B319F98E11BD53">II</collectionCode>
VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large and wide lobes, slightly darker brown than the rest of the plate; posterior margin convex, incision between the lobes is a membranous strip, widening towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is narrow in ventral view, distinctly raised, nose shaped to rounded in lateral view, lacking distinct setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging the posterior teeth.
): Terga I VIII with transverse row of four pigmented spots clearly visible; terga I IV mostly membranous but with lateral sclerites, terga V VIII with medial sclerite as well; medial sclerite is a longitudinal stripe on tergite V, gradually widens on the following segments and as wide as 1 / 2 segment width on tergite VIII; terga IX X fully sclerotised. Sterna I VII simple, sterna II VII with one subrectangular median sclerite and two small anterior sclerites that are fused with the median sclerite on sternite VII. Subgenital plate of sternite VIII large and trapezoid, not fused with other sclerites. It has two large and wide lobes, slightly darker brown than the rest of the plate; posterior margin convex, incision between the lobes is a membranous strip, widening towards the median bulge near the centre of the plate but not forming a triangular field; median bulge is narrow in ventral view, distinctly raised, nose shaped to rounded in lateral view, lacking distinct setation. Sternite IX with wide but shallow anterior indentation. Paraproct, cercus and epiproct simple. Spermathecal sclerite ring-shaped, with small anterior teeth and large, converging the posterior teeth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMEBG" type="Affinities">
@ -373,9 +362,7 @@ The species was collected at a single forest brook in the Kozara Mts of northwes
<figureCitation id="F781F301F816534F036C58F141AF81FA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus and habitat of Leuctra puskasi sp. nov. A female habitus B, C male habitus D type locality: forest brook in the Kozara Mts. E substrate of the type locality. Photographs A C by Gellért Puskás." figureDoi="10.3897/zookeys.1218.120744.figure9" httpUri="https://binary.pensoft.net/fig/1175460">9 D</figureCitation>
). It has a stony substrate mixed with mud and plenty of dead wood and fallen leaves (Fig.
<figureCitation id="9A38933F6D79A9E4FF724D8B26CEFCC9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus and habitat of Leuctra puskasi sp. nov. A female habitus B, C male habitus D type locality: forest brook in the Kozara Mts. E substrate of the type locality. Photographs A C by Gellért Puskás." figureDoi="10.3897/zookeys.1218.120744.figure9" httpUri="https://binary.pensoft.net/fig/1175460">9 E</figureCitation>
). This habitat
<typeStatus id="D94A788395B2A5E4B0187E7DC807597C">type</typeStatus>
is very different from those of the closely related
). This habitat type is very different from those of the closely related
<taxonomicName id="66CCCB5CE61FB8AF55C5A1225BEE1A49" authorityName="Vincon &amp; Muranyi" authorityYear="2007" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="dalmoni">
<emphasis id="A5C5E9475DFC830B3B5E384030C3BB74" italics="true">L. dalmoni</emphasis>
</taxonomicName>
@ -391,9 +378,7 @@ species complex was found in the Kozara, despite the fact that some other habita
<taxonomicName id="4F82D6887444F8D224621E025341FACF" authorityName="Kempny" authorityYear="1899" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="30D60B3AC1FB733872D61F4EC2402BE5" italics="true">L. prima</emphasis>
</taxonomicName>
. The
<typeStatus id="EDDFE9D59C7B7791D734BE1B071D5213">type</typeStatus>
specimens were the only adult stoneflies found there (in partial snow cover), but last instar larvae of
. The type specimens were the only adult stoneflies found there (in partial snow cover), but last instar larvae of
<taxonomicName id="F6FC02AE5EFBF81B463110794F2782C0" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="456DF089029E01262EEA28D99B1CA975" italics="true">Leuctra nigra</emphasis>
</taxonomicName>
@ -408,13 +393,9 @@ specimens were the only adult stoneflies found there (in partial snow cover), bu
</taxonomicName>
group member were found in the brook. We visited the locality twice more, once in May (
<date id="D3E10F98837D924D384DBA1C4F5338AF" value="2012-05-24">24. v. 2012</date>
, leg.
<collectionCode id="9AF131AA3B8AE65457CB3DC4FC193A60" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. Kovács, G. Puskás) and once in November (
, leg. T. Kovács, G. Puskás) and once in November (
<date id="67132132AA6DD5D2A496C6C4E77EBF0D" value="2012-11-07">7. xi. 2012</date>
, leg.
<collectionCode id="1A8FC093D45AD93C1927C1E98674916C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. Kovács, G. Magos). In May, adults of
, leg. T. Kovács, G. Magos). In May, adults of
<taxonomicName id="D83F0FF46958D0A414B527E9E54C1A77" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="D30EA5B9DA79CFF10B5E741A8E2F94BA" italics="true">Leuctra nigra</emphasis>
</taxonomicName>