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<mods:title id="582F2F8D4EB918CF62A9D9490E06978C">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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<mods:namePart id="8B92E0F0E3AB247C38F09385F932119C">Pavel, Filip</mods:namePart>
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<subSubSection id="C321B684FFFEFFFA13E9FE8AFDF3ABDA" box="[151,574,366,393]" pageId="2" pageNumber="386" type="nomenclature">
<paragraph id="8B84E50FFFFEFFFA13E9FE8AFDF3ABDA" blockId="2.[151,574,366,429]" box="[151,574,366,393]" pageId="2" pageNumber="386">
<heading id="D0CC5263FFFEFFFA13E9FE8AFDF3ABDA" bold="true" box="[151,574,366,393]" fontSize="11" level="1" pageId="2" pageNumber="386" reason="1">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE8AFDF3ABDA" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[151,574,366,393]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE8AFDF3ABDA" bold="true" box="[151,574,366,393]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13E9FE8AFE0BABDB" bold="true" box="[151,454,366,393]" italics="true" pageId="2" pageNumber="386">Biflabelotillus foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFFEFFFA12B3FE8AFDF3ABDA" author="Pic, M." box="[461,574,366,392]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B84E50FFFFEFFFA13E9FE77FE5FABFF" blockId="2.[151,574,366,429]" box="[151,402,403,429]" pageId="2" pageNumber="386">
<figureCitation id="1300F98AFFFEFFFA13E9FE77FE8DABFF" box="[151,320,403,429]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 1, 4, 7, 10</figureCitation>
, 1320
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13E9FE33FEF9A87E" pageId="2" pageNumber="386" type="reference_group">
<paragraph id="8B84E50FFFFEFFFA13E9FE33FCF8ABBD" blockId="2.[151,1436,471,556]" box="[151,821,471,495]" pageId="2" pageNumber="386">
<treatmentCitationGroup id="AB2BC221FFFEFFFA13E9FE33FCF8ABBD" box="[151,821,471,495]" pageId="2" pageNumber="386">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE33FD9CABBD" authority="Pic, 1949: 3" authorityName="Pic" authorityPageNumber="3" authorityYear="1949" box="[151,593,471,495]" class="Insecta" family="Cleridae" genus="Biflabellotilus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE33FEEAABBD" box="[151,295,471,495]" italics="true" pageId="2" pageNumber="386">Biflabellotilus</emphasis>
<treatmentCitation id="0A9AC31EFFFEFFFA1253FE33FD9CABBD" author="Pic, M." box="[301,593,471,495]" page="3" pageId="2" pageNumber="386" year="1949">
[sic!]
<emphasis id="B94F391DFFFEFFFA1217FE33FE1CABBD" box="[361,465,471,495]" italics="true" pageId="2" pageNumber="386">foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFFEFFFA12A9FE33FD9CABBD" author="Pic, M." box="[471,593,471,495]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13E9FE12FEF9A87E" blockId="2.[151,1436,471,556]" pageId="2" pageNumber="386">
<treatmentCitationGroup id="AB2BC221FFFEFFFA13E9FE12FEF9A87E" pageId="2" pageNumber="386">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE12FE18A85C" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[151,469,502,526]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE12FEE3A85C" box="[151,302,502,526]" italics="true" pageId="2" pageNumber="386">Biflabellotillus</emphasis>
[sic!]
<emphasis id="B94F391DFFFEFFFA1213FE12FE18A85C" box="[365,469,502,526]" italics="true" pageId="2" pageNumber="386">foveolatus</emphasis>
</taxonomicName>
<treatmentCitation id="0A9AC31EFFFEFFFA1295FE12FD1CA85C" author="Corporaal, J. B." box="[491,721,502,526]" page="17" pageId="2" pageNumber="386" year="1950">
<bibRefCitation id="EFAA98FEFFFEFFFA1295FE12FD7DA85C" author="Corporaal, J. B." box="[491,688,502,526]" pageId="2" pageNumber="386" refId="ref5062" refString="Corporaal, J. B. (1950) Cleridae. Coleopterorum Catalogus. Supplementa Pars 23. Uitgeverij. Dr. W. Junk, ' s-Gravenhage, 373 pp." type="book" year="1950">CORPORAAL (1950)</bibRefCitation>
: 17
</treatmentCitation>
(catalogue);
<treatmentCitation id="0A9AC31EFFFEFFFA102DFE12FBEEA85C" author="Winkler, J. R." box="[851,1059,502,526]" page="32" pageId="2" pageNumber="386" year="1961">
<bibRefCitation id="EFAA98FEFFFEFFFA102DFE12FC32A85C" author="Winkler, J. R." box="[851,1023,502,526]" pageId="2" pageNumber="386" refId="ref5415" refString="Winkler, J. R. (1961) Die Buntkafer (Cleridae). A. Ziemsen Verlag, Wittenberg, Lutherstadt, 108 pp." type="book" year="1961">WINKLER (1961)</bibRefCitation>
: 32
</treatmentCitation>
;
<treatmentCitation id="0A9AC31EFFFEFFFA1752FE12FB25A85C" author="Menier, J. J." box="[1068,1256,502,526]" page="75" pageId="2" pageNumber="386" year="1979">
<bibRefCitation id="EFAA98FEFFFEFFFA1752FE12FB05A85C" author="Menier, J. J." box="[1068,1224,502,526]" pageId="2" pageNumber="386" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">MENIER (1979)</bibRefCitation>
: 75
</treatmentCitation>
(habitat character drawings).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<materialsCitation id="3B53EF52FFFEFFFA13E9FDBFFB4AA827" box="[151,1159,602,629]" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" stateProvince="Analanjirofo Region">
<emphasis id="B94F391DFFFEFFFA13E9FDBFFEFAA827" bold="true" box="[151,311,602,629]" pageId="2" pageNumber="386">Type locality.</emphasis>
<collectingCountry id="F32CA59FFFFEFFFA1240FDBFFE0AA827" box="[318,455,603,629]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingRegion id="49FF2BEDFFFEFFFA12AEFDBFFD76A827" box="[464,699,603,629]" pageId="2" pageNumber="386">Analanjirofo Region</collectingRegion>
,
<collectingCounty id="62E59D83FFFEFFFA11BBFDBFFC7AA827" box="[709,951,603,629]" pageId="2" pageNumber="386">Maroantsetra District</collectingCounty>
,
<collectingMunicipality id="6BE07F75FFFEFFFA10BFFDBFFB4AA827" box="[961,1159,603,629]" pageId="2" pageNumber="386">Antongonivitsika</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9FD9BFC7EA97B" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FD9BFE2EA8CA" bold="true" box="[199,483,638,665]" pageId="2" pageNumber="386">Type material studied.</emphasis>
<materialsCitation id="3B53EF52FFFEFFFA128CFD9BFC27A8B3" collectingDate="1947-11" collectionCode="MNHN" collectorName="Michel" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Analanjirofo Region" typeStatus="lectotype">
<typeStatus id="54805BADFFFEFFFA128CFD9BFDBDA8CB" box="[498,624,639,665]" pageId="2" pageNumber="386" type="lectotype">LECTOTYPE</typeStatus>
(designated here),
<specimenCount id="9D3D2E86FFFEFFFA1021FD9BFCB9A8CB" box="[863,884,639,665]" count="1" pageId="2" pageNumber="386" type="male"></specimenCount>
(
<collectionCode id="ED2A7DCAFFFEFFFA10F5FD9BFC25A8CB" box="[907,1000,639,665]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="386">MNHN</collectionCode>
): [white, handwritten]
<collectingCountry id="F32CA59FFFFEFFFA1780FD9BFA48A8CB" box="[1278,1413,639,665]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
/
<collectingMunicipality id="6BE07F75FFFEFFFA13E9FD47FE94A8EF" box="[151,345,675,701]" pageId="2" pageNumber="386">Antongonivitsika</collectingMunicipality>
/
<date id="FF85C3CFFFFEFFFA1211FD47FE74A8EF" box="[367,441,675,701]" pageId="2" pageNumber="386" value="1947-11">
<collectingDate id="EFC13A27FFFEFFFA1211FD47FE74A8EF" box="[367,441,675,701]" pageId="2" pageNumber="386" value="1947-11">XI. 47</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA12BAFD47FDD4A8EF" box="[452,537,675,701]" pageId="2" pageNumber="386">Michel</collectorName>
! [white, handwritten] Type // [white, handwritten]
<taxonomicName id="4C3B9E8CFFFEFFFA1731FD47FB22A8EF" authorityName="Pic" authorityYear="1949" box="[1103,1263,675,701]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="genus">Biflabelotillus</taxonomicName>
ng / foveolatus nsp // [white, handwritten] genre pres Paradoxocerus Kr. // [red, printed] ♂
</materialsCitation>
<materialsCitation id="3B53EF52FFFEFFFA108CFD23FB8AA957" collectorName="J. J. MENIER" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" stateProvince="Analanjirofo Region" typeStatus="holotype">
<typeStatus id="54805BADFFFEFFFA108CFD23FB4BA8B3" box="[1010,1158,711,737]" pageId="2" pageNumber="386" type="holotype">HOLOTYPE</typeStatus>
// [blue] // [red, printed] SPECIMEN FIGURE / FAUNE DE MADAG. /
<collectorName id="26CE80D9FFFEFFFA11A5FD0FFCB8A957" box="[731,885,747,773]" pageId="2" pageNumber="386">J.J. MENIER</collectorName>
/ 1933 1991 //
</materialsCitation>
<typeStatus id="54805BADFFFEFFFA172CFD0FFB3FA957" box="[1106,1266,747,773]" pageId="2" pageNumber="386" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C3B9E8CFFFEFFFA1780FD0FFECCA97B" authorityName="Pavel &amp; Purchart" authorityYear="2024" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA1780FD0FFECCA97B" italics="true" pageId="2" pageNumber="386">Biflabelotillus bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1276FCEBFEAFA97B" bold="true" box="[264,354,783,809]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1276FCEBFEAFA97B" box="[264,354,783,809]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
det. F. Pavel &amp; L. Purchart 2024 [red, printed label].
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9FCD6FC85A98F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FCD6FDF8A91E" bold="true" box="[199,565,818,844]" pageId="2" pageNumber="386">Additional material examined.</emphasis>
(
<specimenCount id="9D3D2E86FFFEFFFA1136FCD7FDBCA91F" box="[584,625,819,845]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
IJC):
<materialsCitation id="3B53EF52FFFEFFFA11C6FCD7FB6CA923" collectingDate="2001-11-28" collectorName="Z. Mracek" country="Madagascar" county="Central" location="Ankazobe District" municipality="Analamanga Region" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<collectingCounty id="62E59D83FFFEFFFA11C6FCD7FCC7A91F" box="[696,778,819,845]" pageId="2" pageNumber="386">Central</collectingCounty>
<collectingCountry id="F32CA59FFFFEFFFA106CFCD7FC56A91F" box="[786,923,819,845]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BE07F75FFFEFFFA10DBFCD7FB46A91F" box="[933,1163,819,845]" pageId="2" pageNumber="386">Analamanga Region</collectingMunicipality>
[formerly
<collectingRegion id="49FF2BEDFFFEFFFA167AFCD7FF32A923" country="Madagascar" name="Antananarivo" pageId="2" pageNumber="386">Antananarivo province</collectingRegion>
],
<location id="8EE4B3D4FFFEFFFA126AFCB3FE24A923" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA126AFCB3FE24A923" box="[276,489,855,881]" country="Madagascar" county="Central" municipality="Analamanga Region" name="Ankazobe District" pageId="2" pageNumber="386" stateProvince="Antananarivo">Ankazobe District</location>
,
<location id="8EE4B3D4FFFEFFFA1287FCB3FDB1A923" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA1287FCB3FDB1A923" box="[505,636,855,881]" country="Madagascar" county="Central" municipality="Analamanga Region" name="Manankazo" pageId="2" pageNumber="386" stateProvince="Antananarivo">Manankazo</location>
env.,
<date id="FF85C3CFFFFEFFFA11B7FCB3FC85A923" box="[713,840,855,881]" pageId="2" pageNumber="386" value="2001-11-28">
<collectingDate id="EFC13A27FFFEFFFA11B7FCB3FC85A923" box="[713,840,855,881]" pageId="2" pageNumber="386" value="2001-11-28">28.11.2001</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA1027FCB3FC1EA923" box="[857,979,855,881]" pageId="2" pageNumber="386">Z. Mráček</collectorName>
leg.; (
<specimenCount id="9D3D2E86FFFEFFFA1759FCB3FB98A923" box="[1063,1109,855,881]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
FPC)
</materialsCitation>
:
<materialsCitation id="3B53EF52FFFEFFFA17CEFCB3FB00A9C7" collectorName="G. Dujardin-Delacour" country="Madagascar" county="Alaotra Mangora Region" location="Lakato" municipality="Moramanga District" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F32CA59FFFFEFFFA17CEFCB3FAF4A923" box="[1200,1337,855,881]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingCounty id="62E59D83FFFEFFFA1636FCB3FE94A9C7" pageId="2" pageNumber="386">Alaotra Mangora Region</collectingCounty>
,
<collectingMunicipality id="6BE07F75FFFEFFFA1218FC9FFD80A9C7" box="[358,589,891,917]" pageId="2" pageNumber="386">Moramanga District</collectingMunicipality>
,
<location id="8EE4B3D4FFFEFFFA1124FC9FFD66A9C7" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA1124FC9FFD66A9C7" box="[602,683,891,917]" country="Madagascar" county="Alaotra Mangora Region" municipality="Moramanga District" name="Lakato" pageId="2" pageNumber="386">Lakato</location>
, 1.1969,
<collectorName id="26CE80D9FFFEFFFA106BFC9FFBC1A9C7" box="[789,1036,891,917]" pageId="2" pageNumber="386">G. Dujardin-Delacour</collectorName>
leg.; (
<specimenCount id="9D3D2E86FFFEFFFA1724FC9FFB49A9C7" box="[1114,1156,891,917]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
FPC)
</materialsCitation>
:
<materialsCitation id="3B53EF52FFFEFFFA17A7FC9FFC85A98F" collectingDate="2011-11-29" collectorName="M. Tryzna" country="Madagascar" county="Central" elevation="1530" latitude="-18.197498" location="Ambohitantely Special Reserve" longLatPrecision="21" longitude="47.284164" municipality="Analamanga Region" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1">
same as previous but 2. 1969; (
<specimenCount id="9D3D2E86FFFEFFFA1244FC7BFEA8A9EB" box="[314,357,927,953]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
IJC):
<collectingCounty id="62E59D83FFFEFFFA12CCFC7BFDC9A9EB" box="[434,516,927,953]" pageId="2" pageNumber="386">Central</collectingCounty>
<collectingCountry id="F32CA59FFFFEFFFA1170FC7BFD5AA9EB" box="[526,663,927,953]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BE07F75FFFEFFFA11DAFC7BFC5DA9EB" box="[676,912,927,953]" pageId="2" pageNumber="386">Analamanga Region</collectingMunicipality>
,
<location id="8EE4B3D4FFFEFFFA10E2FC7BFAC5A9EB" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA10E2FC7BFAC5A9EB" box="[924,1288,927,953]" country="Madagascar" county="Central" latitude="-18.197498" longLatPrecision="21" longitude="47.284164" municipality="Analamanga Region" name="Ambohitantely Special Reserve" pageId="2" pageNumber="386">Ambohitantely Special Reserve</location>
,
<geoCoordinate id="EE0F83C8FFFEFFFA1669FC7AFA51A9EB" box="[1303,1436,926,953]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="386" precision="15" seconds="51" value="-18.197498">S18°1151</geoCoordinate>
<geoCoordinate id="EE0F83C8FFFEFFFA13E9FC27FEE9A98F" box="[151,292,963,989]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="2" pageNumber="386" precision="15" seconds="03" value="47.284164">E47°1703</geoCoordinate>
,
<quantity id="4CC348EAFFFEFFFA1251FC26FE4DA98F" box="[303,384,962,989]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="2" pageNumber="386" unit="m" value="1530.0">
<elevation id="0016023CFFFEFFFA1251FC26FE4DA98F" box="[303,384,962,989]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="2" pageNumber="386" unit="m" value="1530.0">1530m</elevation>
</quantity>
, at light, 24.
<date id="FF85C3CFFFFEFFFA1168FC27FD55A98F" box="[534,664,963,989]" pageId="2" pageNumber="386" value="2011-11-29">
<collectingDate id="EFC13A27FFFEFFFA1168FC27FD55A98F" box="[534,664,963,989]" pageId="2" pageNumber="386" value="2011-11-29">29.11.2011</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA11DDFC27FCDAA98F" box="[675,791,963,989]" pageId="2" pageNumber="386">M. Trýzna</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FC03FD77AEC3" pageId="2" pageNumber="386" type="discussion">
<paragraph id="8B84E50FFFFEFFFA13B9FC03FD77AEC3" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FC03FEF6AE53" bold="true" box="[199,315,999,1025]" pageId="2" pageNumber="386">Remarks.</emphasis>
As
<bibRefCitation id="EFAA98FEFFFEFFFA1210FC03FE2AAE53" author="Pic, M." box="[366,487,999,1025]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
made no reference to specimens in his description it cannot be assumed that the nominal species-group name,
<taxonomicName id="4C3B9E8CFFFEFFFA1291FBEFFCC7AE77" authorityName="Pic" authorityYear="1949" box="[495,778,1035,1061]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA1291FBEFFCC7AE77" box="[495,778,1035,1061]" italics="true" pageId="2" pageNumber="386">Biflabelotillus foveolatus</emphasis>
</taxonomicName>
, was based on a single specimen. Despite being labelled holotype (a recent, not original, label) most likely by Menier the type located in the Muséum National dHistoire Naturelle, Paris (MNHN) must be considered a
<typeStatus id="54805BADFFFEFFFA11C3FBB7FCD9AE3F" box="[701,788,1107,1133]" pageId="2" pageNumber="386" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="EFAA98FEFFFEFFFA105AFBB7FC60AE3F" author="ICZN" box="[804,941,1107,1133]" pageId="2" pageNumber="386" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 73.2) and is therefore available for
<typeStatus id="54805BADFFFEFFFA13E9FB93FF33AEC3" box="[151,254,1143,1169]" pageId="2" pageNumber="386" type="lectotype">lectotype</typeStatus>
designation (
<bibRefCitation id="EFAA98FEFFFEFFFA12EAFB93FDD6AEC3" author="ICZN" box="[404,539,1143,1169]" pageId="2" pageNumber="386" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 74.1).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FB7EFEE8AF87" pageId="2" pageNumber="386" type="diagnosis">
<paragraph id="8B84E50FFFFEFFFA13B9FB7EFEE8AF87" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FB7EFE1CAEE6" bold="true" box="[199,465,1178,1204]" pageId="2" pageNumber="386">Differential diagnosis.</emphasis>
<taxonomicName id="4C3B9E8CFFFEFFFA12A2FB7FFDBFAEE7" authorityName="Pic" authorityYear="1949" box="[476,626,1179,1205]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA12A2FB7FFDBFAEE7" box="[476,626,1179,1205]" italics="true" pageId="2" pageNumber="386">B. foveolatus</emphasis>
</taxonomicName>
can easily be distinguished from both
<taxonomicName id="4C3B9E8CFFFEFFFA1743FB7FFB1FAEE6" authorityName="Menier" authorityYear="1979" box="[1085,1234,1179,1204]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1743FB7FFB1FAEE6" box="[1085,1234,1179,1204]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B9E8CFFFEFFFA166EFB7FFA51AEE7" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1296,1436,1179,1205]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA166EFB7FFA51AEE7" box="[1296,1436,1179,1205]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA13E9FB5BFF3CAE8B" bold="true" box="[151,241,1215,1241]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA13E9FB5BFF3CAE8B" box="[151,241,1215,1241]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by brown elytra (black in
<taxonomicName id="4C3B9E8CFFFEFFFA1160FB5BFD7DAE8A" authorityName="Menier" authorityYear="1979" box="[542,688,1215,1240]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1160FB5BFD7DAE8A" box="[542,688,1215,1240]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
and bicoloured in
<taxonomicName id="4C3B9E8CFFFEFFFA10FAFB5BFBC3AE8B" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[900,1038,1215,1241]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA10FAFB5BFBC3AE8B" box="[900,1038,1215,1241]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA176BFB5BFBA2AE8B" bold="true" box="[1045,1135,1215,1241]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA176BFB5BFBA2AE8B" box="[1045,1135,1215,1241]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA1701FB5BFB18AE8B" box="[1151,1237,1215,1241]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 14</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA179FFB5BFACDAE8B" box="[1249,1280,1215,1241]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="2" pageNumber="386">22</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA1672FB5BFAE1AE8B" box="[1292,1324,1215,1241]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">34</figureCitation>
). From
<taxonomicName id="4C3B9E8CFFFEFFFA16FAFB5BFEC4AEAE" authorityName="Menier" authorityYear="1979" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA16FAFB5BFEC4AEAE" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
it further differs by body length which is equal or larger than
<quantity id="4CC348EAFFFEFFFA10B7FB07FBD4AEAF" box="[969,1049,1251,1277]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="386" unit="mm" value="10.0">10 mm</quantity>
(less than
<quantity id="4CC348EAFFFEFFFA17EAFB07FB18AEAF" box="[1172,1237,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="386" unit="mm" value="9.0">9 mm</quantity>
in
<taxonomicName id="4C3B9E8CFFFEFFFA1784FB07FA41AEAE" authorityName="Menier" authorityYear="1979" box="[1274,1420,1251,1276]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1784FB07FA41AEAE" box="[1274,1420,1251,1276]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
), subparallel, slightly widened elytra (parallel in
<taxonomicName id="4C3B9E8CFFFEFFFA11CBFAE3FC85AF72" authorityName="Menier" authorityYear="1979" box="[693,840,1287,1312]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA11CBFAE3FC85AF72" box="[693,840,1287,1312]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
), smooth elytral interstriae (the latter with tubercles in
<taxonomicName id="4C3B9E8CFFFEFFFA13C8FACFFE84AF16" authorityName="Menier" authorityYear="1979" box="[182,329,1323,1348]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA13C8FACFFE84AF16" box="[182,329,1323,1348]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
) and last but not least the shape of ultimate antennomere more or less securiform (elongate-oval in
<taxonomicName id="4C3B9E8CFFFEFFFA13C8FAABFE84AF3A" authorityName="Menier" authorityYear="1979" box="[182,329,1359,1384]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA13C8FAABFE84AF3A" box="[182,329,1359,1384]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
;
<figureCitation id="1300F98AFFFEFFFA1227FAABFE0DAF3B" box="[345,448,1358,1385]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 4, 5</figureCitation>
). From the new species
<taxonomicName id="4C3B9E8CFFFEFFFA11A6FAABFCAEAF3B" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[728,867,1359,1385]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA11A6FAABFCAEAF3B" box="[728,867,1359,1385]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1012FAABFC0AAF3B" bold="true" box="[876,967,1359,1385]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1012FAABFC0AAF3B" box="[876,967,1359,1385]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it can also be distinguished by distinctly securiform ultimate antennomere (more or less securiform in
<taxonomicName id="4C3B9E8CFFFEFFFA1037FA97FC16AFDF" authorityName="Pic" authorityYear="1949" box="[841,987,1395,1421]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA1037FA97FC16AFDF" box="[841,987,1395,1421]" italics="true" pageId="2" pageNumber="386">B. foveolatus</emphasis>
</taxonomicName>
; Figs 4,6), parallel-sided pronotum (not parallel in
<taxonomicName id="4C3B9E8CFFFEFFFA1273FA73FE5BAFE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[269,406,1431,1457]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA1273FA73FE5BAFE3" box="[269,406,1431,1457]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA12E2FA73FE38AFE3" bold="true" box="[412,501,1431,1457]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA12E2FA73FE38AFE3" box="[412,501,1431,1457]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA117DFA73FDA9AFE3" box="[515,612,1431,1457]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 7, 9</figureCitation>
) and dark brown abdomen (distinctly pale-brown in
<taxonomicName id="4C3B9E8CFFFEFFFA17D3FA73FAFBAFE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1197,1334,1431,1457]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA17D3FA73FAFBAFE3" box="[1197,1334,1431,1457]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1642FA73FA59AFE3" bold="true" box="[1340,1428,1431,1457]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1642FA73FA59AFE3" box="[1340,1428,1431,1457]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA13E9FA5FFF21AF87" box="[151,236,1467,1493]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 18</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA1386FA5EFEDAAF87" box="[248,279,1466,1493]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FA3AFAADADD7" pageId="2" pageNumber="386" type="description">
<paragraph id="8B84E50FFFFEFFFA13B9FA3AFF23AC4F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FA3AFEB8AFAA" bold="true" box="[199,373,1502,1528]" pageId="2" pageNumber="386">Redescription.</emphasis>
Size of
<typeStatus id="54805BADFFFEFFFA12A8FA3BFDF5AFAB" box="[470,568,1503,1529]" pageId="2" pageNumber="386" type="holotype">holotype</typeStatus>
10.0 x
<quantity id="4CC348EAFFFEFFFA11EFFA3BFD26AFAB" box="[657,747,1503,1529]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="386" unit="mm" value="2.4">2.4 mm</quantity>
. Winged, dark brown, with elytra and abdomen brown (
<figureCitation id="1300F98AFFFEFFFA1614FA3BFF2CAC4F" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 1418</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F9C3FAF2AC37" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F9C3FEC3AC13" bold="true" box="[199,270,1575,1601]" pageId="2" pageNumber="386">Head.</emphasis>
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
<quantity id="4CC348EAFFFEFFFA1756F9C3FB4CAC13" box="[1064,1153,1575,1601]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="386" unit="mm" value="2.1">2.1 mm</quantity>
). The space between eyes (frons) narrow (HW / FW = 3.5). Ultimate antennomere almost securifom, 3.4x longer then broad (
<figureCitation id="1300F98AFFFEFFFA1793F9AFFAFFAC37" box="[1261,1330,1611,1637]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F98BFC81ACFF" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F98BFE8AACDB" bold="true" box="[199,327,1647,1673]" pageId="2" pageNumber="386">Pronotum.</emphasis>
Longer than wide in ratio 1:1.7 (PL =
<quantity id="4CC348EAFFFEFFFA1070F98AFCA5ACDB" box="[782,872,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="386" unit="mm" value="2.5">2.5 mm</quantity>
, PW =
<quantity id="4CC348EAFFFEFFFA10BCF98AFBD2ACDB" box="[962,1055,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="386" unit="mm" value="1.5">1.5 mm</quantity>
), cylindrical, apical two thirds of pronotum parallel sided, basal third strongly trangled (
<figureCitation id="1300F98AFFFEFFFA1187F977FCF3ACFF" box="[761,830,1683,1709]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F953FC25AD4B" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F953FED7AC83" bold="true" box="[199,282,1719,1745]" pageId="2" pageNumber="386">Elytra.</emphasis>
Subparallel, 2.6x longer than their width (EL =
<quantity id="4CC348EAFFFEFFFA1036F953FC6FAC83" box="[840,930,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="386" unit="mm" value="6.3">6.3 mm</quantity>
, EW =
<quantity id="4CC348EAFFFEFFFA1082F953FB95AC83" box="[1020,1112,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="386" unit="mm" value="2.4">2.4 mm</quantity>
), with posterior half slightly widening posteriad. Three fourths of elytral surface with bitubecrulate foveae, the latter absent in apical fourth (
<figureCitation id="1300F98AFFFEFFFA1611F93FFF2DAD4B" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 14, 17</figureCitation>
). Space between foveae approx. 13x larger than diameter of foveae.
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F8C6FB6DAD6F" blockId="2.[151,1437,602,1997]" box="[199,1184,1826,1853]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F8C6FE8BAD6E" bold="true" box="[199,326,1826,1852]" pageId="2" pageNumber="386">Scutellum.</emphasis>
Almost rounded with the basal margin distinctly cut-out in middle (
<figureCitation id="1300F98AFFFEFFFA1741F8C7FB5EAD6F" box="[1087,1171,1827,1853]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F8A2FAADADD7" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F8A2FE6EAD32" bold="true" box="[199,419,1862,1888]" pageId="2" pageNumber="386">Variability of size.</emphasis>
Examined males (n=2): body length
<quantity id="4CC348EAFFFEFFFA1031F8A2FBCCAD33" box="[847,1025,1862,1889]" metricMagnitude="-2" metricUnit="m" metricValue="1.1824999999999999" metricValueMax="1.19" metricValueMin="1.175" pageId="2" pageNumber="386" unit="mm" value="11.825" valueMax="11.9" valueMin="11.75">11.7511.9 mm</quantity>
, HW =
<quantity id="4CC348EAFFFEFFFA1720F8A3FAC8AD33" box="[1118,1285,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.63" metricValueMin="2.42" pageId="2" pageNumber="386" unit="mm" value="2.525" valueMax="2.63" valueMin="2.42">2.422.63 mm</quantity>
, HW / FW = 2.993.31, PL =
<quantity id="4CC348EAFFFEFFFA122DF88EFE27ADD7" box="[339,490,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" metricValueMax="2.58" metricValueMin="2.5" pageId="2" pageNumber="386" unit="mm" value="2.54" valueMax="2.58" valueMin="2.5">2.52.58 mm</quantity>
, PW =
<quantity id="4CC348EAFFFEFFFA1142F88FFD7DADD7" box="[572,688,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="2" pageNumber="386" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
, EL =
<quantity id="4CC348EAFFFEFFFA1185F88EFC5FADD7" box="[763,914,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="8.475" metricValueMax="8.7" metricValueMin="8.25" pageId="2" pageNumber="386" unit="mm" value="8.475" valueMax="8.7" valueMin="8.25">8.258.7 mm</quantity>
, EW =
<quantity id="4CC348EAFFFEFFFA1098F88EFB41ADD7" box="[998,1164,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.55" metricValueMin="3.25" pageId="2" pageNumber="386" unit="mm" value="3.4" valueMax="3.55" valueMin="3.25">3.253.55 mm</quantity>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9F86AFE8CAD9F" pageId="2" pageNumber="386" type="distribution">
<paragraph id="8B84E50FFFFEFFFA13B9F86AFE8CAD9F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F86AFEACADFA" bold="true" box="[199,353,1934,1960]" pageId="2" pageNumber="386">Distribution.</emphasis>
Central and Eastern
<collectingCountry id="F32CA59FFFFEFFFA112AF86BFD16ADFB" box="[596,731,1935,1961]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
Analamanga, Alaotra Mangora and Sava Regions (
<figureCitation id="1300F98AFFFEFFFA163EF86BFA59ADFB" box="[1344,1428,1935,1961]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="2" pageNumber="386">Fig. 38</figureCitation>
) (
<bibRefCitation id="EFAA98FEFFFEFFFA13E1F857FEF9AD9F" author="Menier, J. J." box="[159,308,1971,1997]" pageId="2" pageNumber="386" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>