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<document ID-DOI="http://doi.org/10.5281/zenodo.5190422" ID-ISSN="1639-4798" ID-Zenodo-Dep="5190422" approvalRequired="126" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="69" approvalRequired_for_treatments="56" checkinTime="1628790964299" checkinUser="carolina" docAuthor="Janssen, Thomas &amp; Rakotondrainibe, France" docDate="2008" docId="03D3163AFF0BFF233EFE4D6F15A96AEE" docLanguage="en" docName="Adansonia.30.2.221-376.pdf" docOrigin="Adansonia (3) 30 (2)" docStyle="DocumentStyle:7E81836D30F26F6CECD5F7ABFAAC2CA1.1:Adansonia.2005-2008.journal_article" docStyleId="7E81836D30F26F6CECD5F7ABFAAC2CA1" docStyleName="Adansonia.2005-2008.journal_article" docStyleVersion="1" docTitle="Cyathea goudotii Kunze" docType="treatment" docVersion="2" lastPageId="144" lastPageNumber="365" masterDocId="FFEA6E42FF84FFB33E014B25157E6861" masterDocTitle="A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles" masterLastPageNumber="376" masterPageNumber="221" pageId="143" pageNumber="364" updateTime="1628797601358" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janssen, Thomas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakotondrainibe, France</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Adansonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="series">
<mods:title>3</mods:title>
</mods:detail>
<mods:detail type="volume">
<mods:number>30</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>221</mods:start>
<mods:end>376</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.5190422</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
<mods:identifier type="Zenodo-Dep">5190422</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03D3163AFF0BFF233EFE4D6F15A96AEE" httpUri="http://treatment.plazi.org/id/03D3163AFF0BFF233EFE4D6F15A96AEE" lastPageId="144" lastPageNumber="365" pageId="143" pageNumber="364">
<subSubSection pageId="143" pageNumber="364" type="nomenclature">
<paragraph blockId="143.[255,505,1610,1637]" box="[255,505,1610,1637]" pageId="143" pageNumber="364">
<heading box="[255,505,1610,1637]" centered="true" fontSize="11" level="2" pageId="143" pageNumber="364" reason="6">
<taxonomicName authority="Kunze" box="[255,505,1610,1637]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[255,429,1610,1636]" italics="true" pageId="143" pageNumber="364">Cyathea goudotii</emphasis>
Kunze
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="143.[136,646,1675,1699]" box="[136,646,1675,1699]" pageId="143" pageNumber="364">
<emphasis box="[136,320,1675,1699]" italics="true" pageId="143" pageNumber="364">Botanische Zeitung</emphasis>
(
<collectingRegion box="[335,399,1675,1699]" country="Germany" name="Berlin" pageId="143" pageNumber="364">Berlin</collectingRegion>
) 2: 283, in obs. (1844);
</paragraph>
</subSubSection>
<subSubSection pageId="143" pageNumber="364" type="description">
<paragraph blockId="143.[678,1210,215,265]" pageId="143" pageNumber="364">
<materialsCitation country="Madagascar" location="Goudot" pageId="143" pageNumber="364" specimenCount="1">
Moore,
<treatmentCitation author="Moore" box="[758,1025,215,239]" page="269" pageId="143" pageNumber="364" score="0.00114607548378871" volumeTitle="Index Filicum" year="1884">
<taxonomicName authority="Filicum" authorityName="Filicum" box="[758,893,215,239]" class="Mollusca (awaiting allocation)" family="Mollusca (awaiting allocation)" genus="Index" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mollusca (awaiting allocation)" pageId="143" pageNumber="364" phylum="Mollusca" rank="genus">
<emphasis box="[758,893,215,239]" italics="true" pageId="143" pageNumber="364">Index Filicum</emphasis>
</taxonomicName>
: 269 (1884)
</treatmentCitation>
, nomen. —
<typeStatus box="[1154,1206,215,239]" pageId="143" pageNumber="364">Type</typeStatus>
:
<collectingCountry box="[678,794,241,265]" name="Madagascar" pageId="143" pageNumber="364">Madagascar</collectingCountry>
,
<emphasis box="[804,913,241,265]" italics="true" pageId="143" pageNumber="364">
<location LSID="urn:lsid:plazi:treatment:03D3163AFF0BFF233EFE4D6F15A96AEE:8EA5F1F7FF0BFF3C3D254BD416146968" box="[804,874,241,265]" country="Madagascar" name="Goudot" pageId="143" pageNumber="364">Goudot</location>
s.n.
</emphasis>
(holo-, LZ† in hb. Kunze)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="144" lastPageNumber="365" pageId="143" pageNumber="364" type="discussion">
<paragraph blockId="143.[674,1210,311,1700]" box="[677,777,311,337]" pageId="143" pageNumber="364">REMARKS</paragraph>
<paragraph blockId="143.[674,1210,311,1700]" pageId="143" pageNumber="364">
<taxonomicName authority="Kunze" authorityName="Kunze" box="[677,927,342,370]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[677,851,342,368]" italics="true" pageId="143" pageNumber="364">Cyathea goudotii</emphasis>
Kunze
</taxonomicName>
is based on a fragmentary differential description comparing a specimen in hb. Kunze (LZ) to
<emphasis box="[890,1142,406,432]" italics="true" pageId="143" pageNumber="364">Sieber fl. mixt. exs. 304</emphasis>
(
<taxonomicName authority="(Hook.) Bonap." authorityName="Bonap." baseAuthorityName="Hook." class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="variety" species="borbonica" variety="latifolia">
<emphasis italics="true" pageId="143" pageNumber="364">Cyathea borbonica</emphasis>
Desv. var.
<emphasis box="[938,1018,437,463]" italics="true" pageId="143" pageNumber="364">latifolia</emphasis>
(Hook.) Bonap.
</taxonomicName>
). Although this description only includes characters of the veins, it must nevertheless be regarded as a valid publication of the name.
</paragraph>
<paragraph blockId="143.[674,1210,311,1700]" pageId="143" pageNumber="364">
Kunzes herbarium at LZ, including the
<typeStatus box="[1116,1209,564,591]" pageId="143" pageNumber="364">holotype</typeStatus>
of
<taxonomicName box="[703,816,596,622]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[703,816,596,622]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
</taxonomicName>
, has been destroyed. We located three potential isotype specimens that might serve for lectotypfication. The first,
<emphasis box="[971,1094,659,685]" italics="true" pageId="143" pageNumber="364">Goudot s.n.</emphasis>
(G-DEL!, and filed there as
<typeStatus box="[861,956,691,718]" pageId="143" pageNumber="364">holotype</typeStatus>
of
<taxonomicName box="[986,1100,691,717]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[986,1100,691,717]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
</taxonomicName>
), includes the annotation “
<taxonomicName authority="Janssen &amp; Rakotondrainibe, 2008" authorityName="Janssen &amp; Rakotondrainibe" authorityYear="2008" box="[847,937,723,750]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="genus" status="n. sp.">Cyathea</taxonomicName>
<taxonomicNameLabel box="[942,999,723,750]" pageId="143" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
Goudotii Kze et affinitate C.
<taxonomicName authority="Hook." authorityName="Hook." box="[792,941,754,781]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="375" phylum="Tracheophyta" rank="species" species="burkei">Burkei Hook.</taxonomicName>
sed ad specimina perfectiora desorbenda”, most likely in Kunzes hand. It contains a leaf apex of
<taxonomicName authority="Kunze." box="[921,1155,818,845]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="dregei">
<emphasis box="[921,1072,818,844]" italics="true" pageId="143" pageNumber="364">Cyathea dregei</emphasis>
Kunze.
</taxonomicName>
</paragraph>
<paragraph blockId="143.[674,1210,311,1700]" pageId="143" pageNumber="364">
The second,
<emphasis box="[825,937,849,875]" italics="true" pageId="143" pageNumber="364">Goudot s.n.</emphasis>
(K!,nr.
<quantity box="[1017,1069,849,876]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="143" pageNumber="364" unit="in" value="27.0">27 in</quantity>
hb.T.Moore) has been marked “don de M. Delessert” and is thus most likely from the same collection as the specimen in G-DEL, although it has been determined as
<taxonomicName class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="variety" species="boivinii" variety="humblotii">
<emphasis italics="true" pageId="143" pageNumber="364">C.boivinii</emphasis>
var.
<emphasis box="[776,878,976,1002]" italics="true" pageId="143" pageNumber="364">humblotii</emphasis>
</taxonomicName>
and cited as
<typeStatus box="[1015,1061,976,1003]" pageId="143" pageNumber="364">type</typeStatus>
of
<taxonomicName box="[1095,1209,976,1003]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[1095,1209,976,1003]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
</taxonomicName>
by
<bibRefCitation author="CHRISTENSEN C." box="[710,963,1008,1035]" pageId="143" pageNumber="364" pagination="1 - 253" refId="ref116813" refString="CHRISTENSEN C. 1932. - Pteridophyta of Madagascar. Dansk Botanisk Arkiv 7: 1 - 253." type="journal article" year="1932">Christensen (1932: 34)</bibRefCitation>
. Pinnule size, however, points to
<taxonomicName box="[776,865,1039,1066]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="dregei">
<emphasis box="[776,865,1039,1066]" italics="true" pageId="143" pageNumber="364">C. dregei</emphasis>
</taxonomicName>
, but determination must remain doubtful for such a fragmentary specimen.
</paragraph>
<paragraph blockId="143.[674,1210,311,1700]" pageId="143" pageNumber="364">
The third,
<emphasis box="[819,941,1103,1129]" italics="true" pageId="143" pageNumber="364">Goudot s.n.</emphasis>
(P!, in hb. Luerssen no. 10416), is without doubt a fragment of
<taxonomicName authority="Mett. ex Kuhn. Luerssen" authorityName="Kuhn. Luerssen" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="decrescens">
<emphasis box="[1084,1209,1134,1161]" italics="true" pageId="143" pageNumber="364">C. decrescens</emphasis>
Mett. ex Kuhn. Luerssen
</taxonomicName>
indicates on the envelope containing one pinna with a rachis fragment, that this is the “real”
<taxonomicName box="[903,1019,1229,1256]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[903,1019,1229,1256]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
</taxonomicName>
(“Ist die echte
<taxonomicName class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="365" phylum="Tracheophyta" rank="species" species="goudotii">C. Goudotii</taxonomicName>
Kze.”). On the specimen it is stated that it has been taken from hb. Kunze and that it has therein been determined as
<taxonomicName box="[981,1113,1324,1351]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="decrescens">
<emphasis box="[981,1113,1324,1351]" italics="true" pageId="143" pageNumber="364">C. decrescens</emphasis>
</taxonomicName>
by Mettenius (“
<taxonomicName authority="Goudotii Kze" authorityName="Goudotii Kze" box="[765,1011,1356,1383]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="genus">Cyathea Goudotii Kze</taxonomicName>
fragm. origin. hb. Kze. dort von mett. im hb. Kze. als decrescens bezeichnet.
<collectingCountry box="[767,897,1420,1447]" name="Madagascar" pageId="143" pageNumber="364">Madagascar</collectingCountry>
: Goudot leg.”). Although this is second hand information and no autograph of Kunze is associated to it, this specimen is the most promising candidate for lectotypification.
</paragraph>
<paragraph blockId="143.[674,1210,311,1700]" pageId="143" pageNumber="364">
In summary, bipinnate and tripinnate candidate specimens are available for the lectotypification of
<taxonomicName authority="Kunze." authorityName="Kunze." box="[756,955,1610,1637]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[756,871,1610,1636]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
Kunze.
</taxonomicName>
In his diagnosis, Kunze compares
<taxonomicName box="[785,902,1641,1668]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="143" pageNumber="364" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[785,902,1641,1668]" italics="true" pageId="143" pageNumber="364">C. goudotii</emphasis>
</taxonomicName>
to
<emphasis box="[939,1184,1641,1668]" italics="true" pageId="143" pageNumber="364">Sieber fl. mixt. exs. 304</emphasis>
, a clearly bipinnate specimen.
<bibRefCitation author="KUHN M." box="[969,1109,1673,1700]" pageId="143" pageNumber="364" refId="ref117757" refString="KUHN M. 1868. - Filices Africanae. W. Engelmann, Lipsiae, 233 p." type="book" year="1868">Kuhn (1868)</bibRefCitation>
notes the
</paragraph>
<paragraph blockId="144.[111,648,216,655]" pageId="144" pageNumber="365">
similarity of
<taxonomicName box="[255,374,216,242]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="144" pageNumber="365" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[255,374,216,242]" italics="true" pageId="144" pageNumber="365">C. goudotii</emphasis>
</taxonomicName>
to
<taxonomicName authority="Desv." authorityName="Desv. (Christensen" authorityYear="1906" box="[414,623,216,243]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="144" pageNumber="365" phylum="Tracheophyta" rank="species" species="borbonica">
<emphasis box="[414,551,216,242]" italics="true" pageId="144" pageNumber="365">C. borbonica</emphasis>
Desv.
</taxonomicName>
, a clearly bipinnate taxon. It is hence likely that the bipinnate specimen in hb. Luerssen is part of the original material of
<taxonomicName box="[335,454,311,337]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="144" pageNumber="365" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[335,454,311,337]" italics="true" pageId="144" pageNumber="365">C. goudotii</emphasis>
</taxonomicName>
and that Kunzes autograph became associated with the tripinnate specimen in G-DEL by error.
</paragraph>
<paragraph blockId="144.[111,648,216,655]" pageId="144" pageNumber="365">
As this cannot be proven with certainty and as information on the specimen in hb. Luerssen is from second hand, we claim this case to be unsolvable with the evidence currently at hand and leave it to the judgment of future monographers whether the name
<taxonomicName authority="Kunze" authorityName="Kunze" box="[181,376,564,591]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="144" pageNumber="365" phylum="Tracheophyta" rank="species" species="goudotii">
<emphasis box="[181,299,564,591]" italics="true" pageId="144" pageNumber="365">C. goudotii</emphasis>
Kunze
</taxonomicName>
can be lectotypified and replace the widely used name
<taxonomicName authority="Mett. ex Kuhn." authorityName="Kuhn." class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="144" pageNumber="365" phylum="Tracheophyta" rank="species" species="decrescens">
<emphasis box="[444,577,596,622]" italics="true" pageId="144" pageNumber="365">C. decrescens</emphasis>
Mett. ex Kuhn.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>