A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
Author
Janssen, Thomas
Author
Rakotondrainibe, France
text
Adansonia
2008
3
30
2
221
376
journal article
http://doi.org/10.5281/zenodo.5190422
1639-4798
5190422
Cyathea goudotii
Kunze
Botanische Zeitung
(
Berlin
) 2: 283, in obs. (1844);
Moore,
Index Filicum
: 269 (1884)
, nomen. —
Type
:
Madagascar
,
Goudot
s.n.
(holo-, LZ† in hb. Kunze)
.
REMARKS
Cyathea goudotii
Kunze
is based on a fragmentary differential description comparing a specimen in hb. Kunze (LZ) to
Sieber fl. mixt. exs. 304
(
Cyathea borbonica
Desv. var.
latifolia
(Hook.) Bonap.
). Although this description only includes characters of the veins, it must nevertheless be regarded as a valid publication of the name.
Kunze’s herbarium at LZ, including the
holotype
of
C. goudotii
, has been destroyed. We located three potential isotype specimens that might serve for lectotypfication. The first,
Goudot s.n.
(G-DEL!, and filed there as
holotype
of
C. goudotii
), includes the annotation “
Cyathea
n. sp.
Goudotii Kze et affinitate C.
Burkei Hook.
sed ad specimina perfectiora desorbenda”, most likely in Kunze’s hand. It contains a leaf apex of
Cyathea dregei
Kunze.
The second,
Goudot s.n.
(K!,nr.
27 in
hb.T.Moore) has been marked “don de M. Delessert” and is thus most likely from the same collection as the specimen in G-DEL, although it has been determined as
C.boivinii
var.
humblotii
and cited as
type
of
C. goudotii
by
Christensen (1932: 34)
. Pinnule size, however, points to
C. dregei
, but determination must remain doubtful for such a fragmentary specimen.
The third,
Goudot s.n.
(P!, in hb. Luerssen no. 10416), is without doubt a fragment of
C. decrescens
Mett. ex Kuhn. Luerssen
indicates on the envelope containing one pinna with a rachis fragment, that this is the “real”
C. goudotii
(“Ist die echte
C. Goudotii
Kze.”). On the specimen it is stated that it has been taken from hb. Kunze and that it has therein been determined as
C. decrescens
by Mettenius (“
Cyathea Goudotii Kze
fragm. origin. hb. Kze. dort von mett. im hb. Kze. als decrescens bezeichnet.
Madagascar
: Goudot leg.”). Although this is second hand information and no autograph of Kunze is associated to it, this specimen is the most promising candidate for lectotypification.
In summary, bipinnate and tripinnate candidate specimens are available for the lectotypification of
C. goudotii
Kunze.
In his diagnosis, Kunze compares
C. goudotii
to
Sieber fl. mixt. exs. 304
, a clearly bipinnate specimen.
Kuhn (1868)
notes the
similarity of
C. goudotii
to
C. borbonica
Desv.
, a clearly bipinnate taxon. It is hence likely that the bipinnate specimen in hb. Luerssen is part of the original material of
C. goudotii
and that Kunze’s autograph became associated with the tripinnate specimen in G-DEL by error.
As this cannot be proven with certainty and as information on the specimen in hb. Luerssen is from second hand, we claim this case to be unsolvable with the evidence currently at hand and leave it to the judgment of future monographers whether the name
C. goudotii
Kunze
can be lectotypified and replace the widely used name
C. decrescens
Mett. ex Kuhn.