106 lines
14 KiB
XML
106 lines
14 KiB
XML
<document id="D9028DC2F0732D979C0B0CE056D5E789" ID-CLB-Dataset="25983" ID-DOI="10.11646/zootaxa.4657.3.1" ID-GBIF-Dataset="3414318d-7570-49ac-9013-be4e1f1e6347" ID-ISSN="1175-5326" ID-Zenodo-Dep="3371886" ID-ZooBank="86DD93B2-E8F4-4174-B105-9436357CB4B6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566284110186" checkinUser="plazi" docAuthor="Lambert, Gretchen" docDate="2019" docId="6A2E3761A93DFFD51390F9DBDF16FEF6" docLanguage="en" docName="zootaxa.4657.3.1.pdf" docOrigin="Zootaxa 4657 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euherdmania claviformis" docType="treatment" docVersion="11" lastPageNumber="416" masterDocId="96174F19A932FFC51307FFF3DC63FFD6" masterDocTitle="The Ascidiacea collected during the 2017 British Columbia Hakai MarineGEO BioBlitz" masterLastPageNumber="436" masterPageNumber="401" pageNumber="416" updateTime="1698751689998" updateUser="ExternalLinkService">
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<mods:titleInfo id="374315E9EA4080B45224DD92FDE5FEAF">
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<mods:title id="B940CCB47453E003B1E83C35D32097B0">The Ascidiacea collected during the 2017 British Columbia Hakai MarineGEO BioBlitz</mods:title>
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</mods:titleInfo>
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<mods:name id="11568FFED6EA7B32D290E2EB49DF0CE1" type="personal">
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<mods:namePart id="12C77FEDF43AE8F7E7D61B3C1910E826">Lambert, Gretchen</mods:namePart>
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<mods:title id="0154A72830ED4FA36BFCFAE4E590792E">Zootaxa</mods:title>
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<mods:part id="560F19F1CC3E30DA645D0AA33746343C">
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<mods:date id="239A5D8D3BCBDBC0310BF7D02B731CBD">2019</mods:date>
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<mods:number id="A969CA55CD998569E26E05B54BC91527">2019-08-20</mods:number>
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<mods:number id="6ACDDBDFD6423DAB27C1E67EF6D19AE7">4657</mods:number>
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<mods:identifier id="7182FB938F4BCDC9EE62A21405D33D93" type="DOI">10.11646/zootaxa.4657.3.1</mods:identifier>
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<treatment id="6A2E3761A93DFFD51390F9DBDF16FEF6" ID-DOI="http://doi.org/10.5281/zenodo.5941197" ID-GBIF-Taxon="159167968" ID-Zenodo-Dep="5941197" LSID="urn:lsid:plazi:treatment:6A2E3761A93DFFD51390F9DBDF16FEF6" httpUri="http://treatment.plazi.org/id/6A2E3761A93DFFD51390F9DBDF16FEF6" lastPageId="16" lastPageNumber="416" pageId="15" pageNumber="416">
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<subSubSection id="AA9DD5FCA93DFFCA1390F9DBDE19F994" box="[151,634,1576,1603]" pageId="15" pageNumber="416" type="nomenclature">
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<paragraph id="E2388677A93DFFCA1390F9DBDE19F994" blockId="15.[151,634,1576,1638]" box="[151,634,1576,1603]" pageId="15" pageNumber="416">
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<heading id="B970311BA93DFFCA1390F9DBDE19F994" bold="true" box="[151,634,1576,1603]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
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<taxonomicName id="2587FDF4A93DFFCA1390F9DBDE19F994" authority="(Ritter, 1903)" baseAuthorityName="Ritter" baseAuthorityYear="1903" box="[151,634,1576,1603]" class="Ascidiacea" family="Euherdmanniidae" genus="Euherdmania" kingdom="Animalia" order="Enterogona" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="claviformis">
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<emphasis id="D0F35A65A93DFFCA1390F9DBDE19F994" bold="true" box="[151,634,1576,1603]" pageId="15" pageNumber="416">
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<emphasis id="D0F35A65A93DFFCA1390F9DBDDAEF995" bold="true" box="[151,461,1576,1603]" italics="true" pageId="15" pageNumber="416">Euherdmania claviformis</emphasis>
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(
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<bibRefCitation id="8616FB86A93DFFCA12D8F9DBDE11F994" author="Ritter, W. E." box="[479,626,1576,1602]" pageId="15" pageNumber="416" pagination="237 - 261" refId="ref17676" refString="Ritter, W. E. (1903) The structure and affinities of Herdmania claviformis, the type of a new genus and family of ascidians from the coast of California. In: Mark Anniversary Volume 1903, 237 - 261." type="journal article" year="1903">Ritter, 1903</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AA9DD5FCA93DFFD51390F9BFDF16FEF6" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="description">
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<paragraph id="E2388677A93DFFCA1390F9BFDD47F9B0" blockId="15.[151,634,1576,1638]" box="[151,292,1612,1638]" pageId="15" pageNumber="416">
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<figureCitation id="7ABC9AF2A93DFFCA1390F9BFDC97F9B0" box="[151,244,1612,1638]" captionStart="FIGURE 9" captionStartId="16.[151,250,1825,1850]" captionTargetBox="[151,1436,365,1769]" captionTargetId="figure@16.[151,1436,365,1769]" captionTargetPageId="16" captionText="FIGURE 9. Aplousobranchia. A–D: Clavelinidae. A: Clavelina huntsmani. B–D: Pycnoclavella stanleyi. B: expanded orange thoraxes extended beyond sandy tubes. C: thoraxes partially contracted. Photo includes four zooids of orange Metandrocarpa taylori. D: zooids fully contracted; only a bit of orange is visible. E, F: Euherdmaniidae, Euherdmania claviformis. E: zooids fully retracted into long sand-encrusted tubes. F: In this colony the anterior portions of the tubes not sand-encrusted, though the colorless zooid thoraxes are partially contracted. Scale bars: A, 7 mm; B, 2 mm; C, 4 mm; D, 1 cm; E, 1.2 cm; F, 1 cm. A, B, D, F photos by G. Paulay." httpUri="https://zenodo.org/record/3371904/files/figure.png" pageId="15" pageNumber="416">Figure 9</figureCitation>
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E–F
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</paragraph>
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<paragraph id="E2388677A93DFFCA1390F963DFCCF97E" blockId="15.[151,943,1680,1735]" box="[151,943,1680,1705]" pageId="15" pageNumber="416">
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<materialsCitation id="52EF8C2AA93DFFCA1390F963DFCCF97E" ID-GBIF-Occurrence="2336841753" box="[151,943,1680,1705]" collectionCode="IHAK" elevation="19" pageId="15" pageNumber="416" specimenCode="IHAK 60" specimenCount="1">
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<specimenCode id="B2212E0CA93DFFCA1390F963DC95F97F" box="[151,246,1680,1705]" collectionCode="IHAK" pageId="15" pageNumber="416">IHAK 60</specimenCode>
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BHAK 3243, 1738 UF 2549. Rattenbury Pinnacle, Scuba,
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<quantity id="257F2B92A93DFFCA1049F963DFC8F97E" box="[846,939,1680,1704]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="15" pageNumber="416" unit="m" value="18.5" valueMax="20.0" valueMin="17.0">
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<elevation id="69AA6144A93DFFCA1049F963DFC8F97E" box="[846,939,1680,1704]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="15" pageNumber="416" unit="m" value="18.5" valueMax="20.0" valueMin="17.0">17–20 m</elevation>
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</quantity>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="E2388677A93DFFCA1390F95CDF71F911" blockId="15.[151,943,1680,1735]" box="[151,786,1710,1735]" pageId="15" pageNumber="416">
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RHAK 6 BHAK 0995. Low rocky intertidal. Tubes
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<quantity id="257F2B92A93DFFCA11A1F95DDEBBF910" box="[678,728,1710,1735]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="416" unit="cm" value="6.0">6 cm</quantity>
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long.
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</paragraph>
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<paragraph id="E2388677A93DFFD51390F906DF16FEF6" blockId="15.[151,1437,1781,2024]" lastBlockId="16.[151,1437,153,288]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
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Clumps of long thin colorless zooids usually almost completely encrusted and embedded with grey sand, each zooid separate except at the common base formed from tightly interwoven branching basal stolons. Sometimes the anterior individual tubes are clear of sand (
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<figureCitation id="7ABC9AF2A93DFFCA1172F8CEDEA9F881" box="[629,714,1853,1879]" captionStart="FIGURE 9" captionStartId="16.[151,250,1825,1850]" captionTargetBox="[151,1436,365,1769]" captionTargetId="figure@16.[151,1436,365,1769]" captionTargetPageId="16" captionText="FIGURE 9. Aplousobranchia. A–D: Clavelinidae. A: Clavelina huntsmani. B–D: Pycnoclavella stanleyi. B: expanded orange thoraxes extended beyond sandy tubes. C: thoraxes partially contracted. Photo includes four zooids of orange Metandrocarpa taylori. D: zooids fully contracted; only a bit of orange is visible. E, F: Euherdmaniidae, Euherdmania claviformis. E: zooids fully retracted into long sand-encrusted tubes. F: In this colony the anterior portions of the tubes not sand-encrusted, though the colorless zooid thoraxes are partially contracted. Scale bars: A, 7 mm; B, 2 mm; C, 4 mm; D, 1 cm; E, 1.2 cm; F, 1 cm. A, B, D, F photos by G. Paulay." httpUri="https://zenodo.org/record/3371904/files/figure.png" pageId="15" pageNumber="416">Fig. 9F</figureCitation>
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). The thorax is comprised of 12 rows of very short stigmata. The esophageal region is very extended, with the ridged stomach (about six or seven plications) near the posterior end, followed by the testes and the heart.
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<bibRefCitation id="8616FB86A93DFFCA1130F876DEAAF849" author="Ritter, W. E." box="[567,713,1925,1951]" pageId="15" pageNumber="416" pagination="237 - 261" refId="ref17676" refString="Ritter, W. E. (1903) The structure and affinities of Herdmania claviformis, the type of a new genus and family of ascidians from the coast of California. In: Mark Anniversary Volume 1903, 237 - 261." type="journal article" year="1903">Ritter (1903)</bibRefCitation>
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described this species in great detail, so impressed by its difference from all other known colonial ascidians that he established a new family and genus for it. While Ritter himself had to change the family and genus names due to their being preoccupied (
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<bibRefCitation id="8616FB86A93DFFCA10F6F83ED81BF831" author="Ritter, W. E." box="[1009,1144,1997,2023]" pageId="15" pageNumber="416" pagination="650 - 651" refId="ref17720" refString="Ritter, W. E. (1904) Euherdmania vs Herdmania preoccupied. Zoologischer Anzeiger, 27, 650 - 651." type="journal article" year="1904">Ritter 1904</bibRefCitation>
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), this single genus family is still recognized as unusual, and contains only a few species. A detailed morphological description is also given by
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<bibRefCitation id="8616FB86A922FFD513BDFF4EDDE6FF01" author="Van Name, W. G." box="[186,389,188,215]" pageId="16" pageNumber="417" pagination="1 - 476" refId="ref18197" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name (1945)</bibRefCitation>
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;
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<bibRefCitation id="8616FB86A922FFD51297FF4EDE4DFF01" author="Trason, W. B." box="[400,558,188,215]" pageId="16" pageNumber="417" pagination="509 - 545" refId="ref18082" refString="Trason, W. B. (1957) Larval structure and development of the oozooid in the ascidian Euherdmania claviformis. Journal of Morphology, 100, 509 - 545. https: // doi. org / 10.1002 / jmor. 105100030 6" type="journal article" year="1957">Trason (1957)</bibRefCitation>
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described the larva and early post–settlement development. The present study extends the distribution from
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<collectingRegion id="20434895A922FFD512E0FF12DE3AFF2D" box="[487,601,225,251]" country="United States of America" name="California" pageId="16" pageNumber="417">California</collectingRegion>
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(
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<bibRefCitation id="8616FB86A922FFD5116EFF12DFF1FF2D" author="Abbott, D. P. & Newberry, A. T." box="[617,914,225,251]" pageId="16" pageNumber="417" pagination="177 - 226" refId="ref15566" refString="Abbott, D. P. & Newberry, A. T. (1980) Urochordata: the tunicates. In: Morris, R. H., Abbott, D. P. & Haderlie, E. C. (Eds.), Intertidal Invertebrates of California. Stanford, California, Stanford University Press, pp. 177 - 226 + P 57 - P 67." type="journal article" year="1980">Abbott & Newberry 1980</bibRefCitation>
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;
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<bibRefCitation id="8616FB86A922FFD5109AFF12D80CFF2D" author="Abbott, D. P. & Lambert, C. C. & Lambert, G. & Newberry, A. T." box="[925,1135,225,251]" pageId="16" pageNumber="417" pagination="918 - 933" refId="ref15639" refString="Abbott, D. P., Lambert, C. C., Lambert, G. & Newberry, A. T. (2007) Chordata: Ascidiacea. In: Carlton, J. T. (Ed.), The Light & Smith Manual: Intertidal Invertebrates from Central California to Oregon. Berkeley, California, University of California Press, chapter 35, pp. 918 - 933." type="journal article" year="2007">
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Abbott
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<emphasis id="D0F35A65A922FFD510F4FF11D845FF2D" box="[1011,1062,225,251]" italics="true" pageId="16" pageNumber="417">et al</emphasis>
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. 2007
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</bibRefCitation>
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) northward to
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<collectingRegion id="20434895A922FFD51610FF12D9FFFF2D" box="[1303,1436,225,251]" country="United States of America" name="Washington" pageId="16" pageNumber="417">Washington</collectingRegion>
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(unpublished observations) and
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<collectingRegion id="20434895A922FFD512F9FEF6DEDCFEC9" box="[510,703,261,287]" country="Canada" name="British Columbia" pageId="16" pageNumber="417">British Columbia</collectingRegion>
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(present study).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |