treatments-xml/data/D3/64/73/D364737A2426173AFF60B4A217F31ACF.xml

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<mods:title id="0E48CD70962C6E7E5F47446A35ED2C40">New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies</mods:title>
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<mods:namePart id="F7CF76191D3E919A7CC30A3683691C5A">Kataev, Boris M.</mods:namePart>
<mods:affiliation id="0AC4500DA5D21A36DAC7A99B9590C074">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="EDF74933C1BFF1A2EC8E7B012F383AB5">Hongbin, Liang</mods:namePart>
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<mods:affiliation id="D5E3E6AF52797B5DF155E06440FB6645">Key laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. lianghb @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 9668 - 1167</mods:affiliation>
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<mods:namePart id="9138F94F123EC4C7C3A87F4DBEA2A7BE">Wrase, David W.</mods:namePart>
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<taxonomicName id="9CCDB9EF2426173EFF60B4A212D61BC2" ID-CoL="58LMQ" authorityName="Jedlicka" authorityYear="1949" box="[151,493,474,501]" class="Insecta" family="Carabidae" genus="Trichotichnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="species" species="miwai">
<emphasis id="69B91E7E2426173EFF60B4A2127A1BC3" bold="true" box="[151,321,474,501]" italics="true" pageId="42" pageNumber="343">Trichotichnus</emphasis>
(s. str.) 
<emphasis id="69B91E7E2426173EFE53B4A212D61BC2" bold="true" box="[420,493,474,500]" italics="true" pageId="42" pageNumber="343">miwai</emphasis>
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<emphasis id="69B91E7E2426173EFE02B4A211AA1BC3" bold="true" box="[501,657,474,501]" italics="true" pageId="42" pageNumber="343">continentalis</emphasis>
ssp. n.
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(
<figureCitation id="C3F6DEE92426173EFF68B4861230182E" box="[159,267,510,536]" captionStart="FIGURES 107–108" captionStartId="43.[151,264,1956,1980]" captionTargetBox="[157,1420,650,1924]" captionTargetId="figure-222@43.[153,1433,633,1933]" captionTargetPageId="43" captionText="FIGURES 107–108. Trichotichnus ssp., habitus. 107, T. miwai continentalis ssp. n. (Wuliangshan); 108, T. kavanaughi sp. n. (Kongdang). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="42" pageNumber="343">Figs. 107</figureCitation>
, 
<figureCitation id="C3F6DEE92426173EFEE0B4861242182E" box="[279,377,510,536]" captionStart-0="FIGURES 109–120" captionStart-1="FIGURE 121" captionStartId-0="44.[151,264,1763,1787]" captionStartId-1="46.[151,250,1625,1649]" captionTargetBox-0="[165,1420,194,1727]" captionTargetBox-1="[151,1434,275,1599]" captionTargetId-0="figure-32@44.[151,1436,181,1739]" captionTargetId-1="figure-80@46.[151,1435,273,1602]" captionTargetPageId-0="44" captionTargetPageId-1="46" captionText-0="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118)." captionText-1="FIGURE 121. Trichotichnus miwai continentalis ssp. n., distribution." figureDoi-0="http://doi.org/10.5281/zenodo.6781767" figureDoi-1="http://doi.org/10.5281/zenodo.6781769" httpUri-0="https://zenodo.org/record/6781767/files/figure.png" httpUri-1="https://zenodo.org/record/6781769/files/figure.png" pageId="42" pageNumber="343">114–121</figureCitation>
)
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<emphasis id="69B91E7E2426173EFF60B73E12711856" bold="true" box="[151,330,582,608]" pageId="42" pageNumber="343">Type material.</emphasis>
<materialsCitation id="EBA5C8312426173EFEA0B73E1199189E" ID-GBIF-Occurrence="3821894554" collectingDate="2000-06-28" collectionCode="IOZ" collectorName="D. H. Kavanaugh" country="China" county="Gongshan County" elevation="1500" location="Male" pageId="42" pageNumber="343" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="84767CCE2426173EFEA0B73E12F81856" box="[343,451,582,608]" pageId="42" pageNumber="343" type="holotype">Holotype</typeStatus>
. 
<location id="5E1294B72426173EFE25B73E11351856" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFE25B73E11351856" box="[466,526,582,608]" country="China" county="Gongshan County" name="Male" pageId="42" pageNumber="343" stateProvince="Yunnan">Male</location>
, “ 
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, 
<collectingRegion id="99090C8E2426173EFD60B73F10641856" box="[663,863,582,609]" country="China" name="Yunnan" pageId="42" pageNumber="343">Yunnan Province</collectingRegion>
, Gaoligong Shan, Nujiang Prefecture, 
<collectingCounty id="B213BAE02426173EFADCB73E13D018B2" pageId="42" pageNumber="343">Gongshan County</collectingCounty>
, Gongshan, 
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<elevation id="D0E0255F2426173EFE8EB71212EE18B2" box="[377,469,618,645]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="343" unit="m" value="1500.0">1500 m</elevation>
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,”, “N27.74064° / E98.66557°, 
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<collectingDate id="3F371D442426173EFCCFB71210EF18B2" box="[824,980,618,645]" pageId="42" pageNumber="343" value="2000-06-28">28 June 2000</collectingDate>
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, Stop#00-14, 
<collectorName id="F638A7BA2426173EFB83B713160818B2" box="[1140,1331,618,645]" pageId="42" pageNumber="343">D.H. Kavanaugh</collectorName>
&amp; Liang H.-B. collectors”, “CASENT 1001224” (
<collectionCode id="3DDC5AA92426173EFD96B7F611AE189E" box="[609,661,654,680]" pageId="42" pageNumber="343">IOZ</collectionCode>
).
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<materialsCitation id="EBA5C8312426173EFF30B7CB10B618C6" ID-GBIF-Occurrence="3821894481" collectingDate="2018-06-28" collectorName="I. Belousov &amp; I. Kabak" country="China" elevation="2145" latitude="27.042778" location="Ningnan" longLatPrecision="20" longitude="102.59778" pageId="42" pageNumber="343" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="84767CCE2426173EFF30B7CB120318FB" box="[199,312,691,717]" pageId="42" pageNumber="343" type="paratype">Paratypes</typeStatus>
. 
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<collectingCountry id="23DA82FC2426173EFEB3B7CA12B418FA" box="[324,399,690,716]" name="China" pageId="42" pageNumber="343">China</collectingCountry>
.
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: 
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, W 
<location id="5E1294B72426173EFD5DB7CA103618FA" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFD5DB7CA103618FA" box="[682,781,690,716]" country="China" latitude="27.042778" longLatPrecision="20" longitude="102.59778" name="Ningnan" pageId="42" pageNumber="343" stateProvince="Sichuan">Ningnan</location>
, 
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<quantity id="9C356F892426173EFCECB7CA104218FA" box="[795,889,690,717]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="42" pageNumber="343" unit="km" value="3.15">3.15 km</quantity>
WNW Maojia Heba
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, 
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, 
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, 
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<collectingDate id="3F371D442426173EFF0CB7AE124718C6" box="[251,380,726,752]" pageId="42" pageNumber="343" value="2018-06-28">28.VI.2018</collectingDate>
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, 
<collectorName id="F638A7BA2426173EFE7FB7AF113118C6" box="[392,522,726,753]" pageId="42" pageNumber="343">I. Belousov</collectorName>
&amp; 
<collectorName id="F638A7BA2426173EFDD9B7AF11B418C6" box="[558,655,726,753]" pageId="42" pageNumber="343">I. Kabak</collectorName>
leg. [6_02nt] (cB&amp;K)
</materialsCitation>
. 
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<collectingRegion id="99090C8E2426173EFC60B7AE172B18C6" box="[919,1040,726,752]" country="China" name="Yunnan" pageId="42" pageNumber="343">YUNNAN</collectingRegion>
: 
<specimenCount id="4DCB09E52426173EFBD7B7AE175518C6" box="[1056,1134,726,752]" pageId="42" pageNumber="343" type="male">1 male</specimenCount>
, same data as holotype but 
<geoCoordinate id="3EF9A4AB2426173EFF60B78212241923" box="[151,287,762,789]" degrees="27.74518" direction="north" orientation="latitude" pageId="42" pageNumber="343" precision="1" value="27.74518">27.74518°N</geoCoordinate>
/ 
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, 
<collectorName id="F638A7BA2426173EFE26B78211331922" box="[465,520,762,788]" pageId="42" pageNumber="343">Stop</collectorName>
#00-13, “CASENT 1001191” (
<collectionCode id="3DDC5AA92426173EFC90B78210981922" box="[871,931,762,788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="42" pageNumber="343" type="Museum">CAS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="EBA5C8312426173EFC4EB7821160196A" ID-GBIF-Occurrence="3821894423" collectingDate="2002-10-07" collectionCode="IOZ" collectorName="A, D. H. Kavanaugh, P. E. &amp; Marek" country="China" county="Gongshan Co." elevation="1350" latitude="27.74843" location="Pula He" longLatPrecision="1" longitude="98.66498" municipality="Cikai Township" pageId="42" pageNumber="343" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52426173EFC4EB782171B1922" box="[953,1056,762,788]" pageId="42" pageNumber="343" type="female">1 female</specimenCount>
, 
<collectingCounty id="B213BAE02426173EFBD9B78217E81922" box="[1070,1235,762,788]" pageId="42" pageNumber="343">Gongshan Co.</collectingCounty>
, 
<collectingMunicipality id="BB1658162426173EFB16B78216A31922" box="[1249,1432,762,788]" pageId="42" pageNumber="343">Cikai Township</collectingMunicipality>
, 
<locationDeviation id="4AC0DB4E2426173EFF60B666128D190E" box="[151,438,798,825]" location="Gongshan" pageId="42" pageNumber="343">
<quantity id="9C356F892426173EFF60B66613CC190E" box="[151,247,798,825]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="42" pageNumber="343" unit="km" value="0.35">0.35 km</quantity>
N of Gongshan
</locationDeviation>
, along 
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, 
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, 
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, stop#DHK-2002-0.24 
<collectorName id="F638A7BA2426173EFA73B66612E5196A" pageId="42" pageNumber="343">A, D.H. Kavanaugh, P.E. Marek</collectorName>
leg. (
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)
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; 
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, 
<collectingCounty id="B213BAE02426173EFD37B63A1058196A" box="[704,867,834,860]" pageId="42" pageNumber="343">Gongshan Co.</collectingCounty>
, 
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, along street, 
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<quantity id="9C356F892426173EFF60B61E13D419B7" box="[151,239,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.515" pageId="42" pageNumber="343" unit="m" value="1515.0">
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, 
<date id="2F73E4AC2426173EFF0CB61E125D19B7" box="[251,358,870,897]" pageId="42" pageNumber="343" value="2002-10-05">
<collectingDate id="3F371D442426173EFF0CB61E125D19B7" box="[251,358,870,897]" pageId="42" pageNumber="343" value="2002-10-05">5.X.2002</collectingDate>
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, 
<collectorName id="F638A7BA2426173EFE85B61F12D419B6" box="[370,495,870,897]" pageId="42" pageNumber="343">H.B. Liang</collectorName>
leg. (
<collectionCode id="3DDC5AA92426173EFDC7B61E115719B6" box="[560,620,870,896]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="42" pageNumber="343" type="Museum">CAS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="EBA5C8312426173EFD77B61E17E01992" ID-GBIF-Occurrence="3821894525" collectingDate="2002-10-09" collectionCode="IOZ" collectorName="Stop &amp; D. Z. Dong" country="China" county="Gongshan Co." elevation="1570" latitude="27.97514" location="Shimengguan" longLatPrecision="1" longitude="98.65502" municipality="Bingzhongluo township" pageId="42" pageNumber="343" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52426173EFD77B61E11F419B6" box="[640,719,870,896]" pageId="42" pageNumber="343" type="male">1 male</specimenCount>
, 
<collectingCounty id="B213BAE02426173EFD2CB61E104519B6" box="[731,894,870,896]" pageId="42" pageNumber="343">Gongshan Co.</collectingCounty>
, 
<collectingMunicipality id="BB1658162426173EFC7DB61E17A119B6" box="[906,1178,870,896]" pageId="42" pageNumber="343">Bingzhongluo township</collectingMunicipality>
, 
<location id="5E1294B72426173EFB52B61E160419B6" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFB52B61E160419B6" box="[1189,1343,870,896]" country="China" county="Gongshan Co." latitude="27.97514" longLatPrecision="1" longitude="98.65502" municipality="Bingzhongluo township" name="Shimengguan" pageId="42" pageNumber="343" stateProvince="Yunnan">Shimengguan</location>
(“ 
<location id="5E1294B72426173EFAAEB61E13EA1992" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFAAEB61E13EA1992" country="China" county="Gongshan Co." latitude="27.97514" longLatPrecision="1" longitude="98.65502" municipality="Bingzhongluo township" name="Stone Gate" pageId="42" pageNumber="343" stateProvince="Yunnan">Stone Gate</location>
”), 
<quantity id="9C356F892426173EFF18B6F2127D1993" box="[239,326,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="42" pageNumber="343" unit="m" value="1570.0">
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, 
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<geoCoordinate id="3EF9A4AB2426173EFE17B6F211531993" box="[480,616,906,933]" degrees="98.65502" direction="east" orientation="longitude" pageId="42" pageNumber="343" precision="1" value="98.65502">98.65502°E</geoCoordinate>
, 
<date id="2F73E4AC2426173EFD84B6F211E61992" box="[627,733,906,932]" pageId="42" pageNumber="343" value="2002-10-09">
<collectingDate id="3F371D442426173EFD84B6F211E61992" box="[627,733,906,932]" pageId="42" pageNumber="343" value="2002-10-09">9.X.2002</collectingDate>
</date>
, 
<collectorName id="F638A7BA2426173EFD1EB6F210261993" box="[745,797,906,933]" pageId="42" pageNumber="343">Stop</collectorName>
#DHK-2002-045, 
<collectorName id="F638A7BA2426173EFC1EB6F3175B1993" box="[1001,1120,907,933]" pageId="42" pageNumber="343">D.Z. Dong</collectorName>
leg. (
<collectionCode id="3DDC5AA92426173EFB68B6F217E81992" box="[1183,1235,906,932]" pageId="42" pageNumber="343">IOZ</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="EBA5C8312426173EFB11B6F216AC19FE" ID-GBIF-Occurrence="3821894476" collectingDate="2004-10-22" collectionCode="CAS" collectorName="D. Kavanaugh" country="China" county="Gongshan" elevation="1520" latitude="27.96817" location="Shuangla He" longLatPrecision="1" longitude="98.66187" municipality="Bingzhongluo" pageId="42" pageNumber="343" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
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, 
<collectingCounty id="B213BAE02426173EFAA2B6F213F619FE" pageId="42" pageNumber="343">Gongshan</collectingCounty>
, 
<collectingMunicipality id="BB1658162426173EFF2FB6D6124119FE" box="[216,378,942,968]" pageId="42" pageNumber="343">Bingzhongluo</collectingMunicipality>
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, beach, 
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<geoCoordinate id="3EF9A4AB2426173EFD0AB6D610BD19FE" box="[765,902,942,968]" degrees="98.66187" direction="east" orientation="longitude" pageId="42" pageNumber="343" precision="1" value="98.66187">98.66187°E</geoCoordinate>
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<quantity id="9C356F892426173EFC67B6D610DD19FF" box="[912,998,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="42" pageNumber="343" unit="m" value="1520.0">
<elevation id="D0E0255F2426173EFC67B6D610DD19FF" box="[912,998,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="42" pageNumber="343" unit="m" value="1520.0">1520 m</elevation>
</quantity>
, 
<date id="2F73E4AC2426173EFC06B6D6175119FE" box="[1009,1130,942,968]" pageId="42" pageNumber="343" value="2004-10-22">
<collectingDate id="3F371D442426173EFC06B6D6175119FE" box="[1009,1130,942,968]" pageId="42" pageNumber="343" value="2004-10-22">22.X.2004</collectingDate>
</date>
, 
<collectorName id="F638A7BA2426173EFB82B6D7162D19FE" box="[1141,1302,942,969]" pageId="42" pageNumber="343">D. Kavanaugh</collectorName>
leg. (
<collectionCode id="3DDC5AA92426173EFAA3B6D616AB19FE" box="[1364,1424,942,968]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="42" pageNumber="343" type="Museum">CAS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="EBA5C8312426173EFF60B6AA16041E26" ID-GBIF-Occurrence="3821894386" collectingDate="2002-09-16" collectingDateMax="2002-09-17" collectingDateMin="2002-09-16" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang" country="China" county="Kunming City" elevation="1945" latitude="25.14035" location="Heilongtan" longLatPrecision="1" longitude="102.74107" municipality="Guandu District" pageId="42" pageNumber="343" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52426173EFF60B6AA13C519DA" box="[151,254,978,1004]" pageId="42" pageNumber="343" type="female">1 female</specimenCount>
, 
<collectingCounty id="B213BAE02426173EFEFBB6AA128B19DA" box="[268,432,978,1004]" pageId="42" pageNumber="343">Kunming City</collectingCounty>
, 
<collectingMunicipality id="BB1658162426173EFE49B6AA114319DA" box="[446,632,978,1004]" pageId="42" pageNumber="343">Guandu District</collectingMunicipality>
, 
<location id="5E1294B72426173EFD72B6AA103F19DA" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFD72B6AA103F19DA" box="[645,772,978,1004]" country="China" county="Kunming City" latitude="25.14035" longLatPrecision="1" longitude="102.74107" municipality="Guandu District" name="Heilongtan" pageId="42" pageNumber="343" stateProvince="Yunnan">Heilongtan</location>
, 
<location id="5E1294B72426173EFCE5B6AA176519DB" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFCE5B6AA176519DB" box="[786,1118,978,1005]" country="China" county="Kunming City" latitude="25.14035" longLatPrecision="1" longitude="102.74107" municipality="Guandu District" name="Kunming Institute of Botany" pageId="42" pageNumber="343" stateProvince="Yunnan">Kunming Institute of Botany</location>
, 
<location id="5E1294B72426173EFB9BB6AA17EC19DA" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFB9BB6AA17EC19DA" box="[1132,1239,978,1004]" country="China" county="Kunming City" latitude="25.14035" longLatPrecision="1" longitude="102.74107" municipality="Guandu District" name="Botanical" pageId="42" pageNumber="343" stateProvince="Yunnan">Botanical</location>
garden, 
<quantity id="9C356F892426173EFAC9B6AA16AC19DB" box="[1342,1431,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="42" pageNumber="343" unit="m" value="1945.0">
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, 
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, 
<date id="2F73E4AC2426173EFE30B68E11481E26" box="[455,627,1014,1040]" pageId="42" pageNumber="343" value="2002-09-16" valueMax="2002-09-17" valueMin="2002-09-16">
<collectingDate id="3F371D442426173EFE30B68E11481E26" box="[455,627,1014,1040]" pageId="42" pageNumber="343" value="2002-09-16" valueMax="2002-09-17" valueMin="2002-09-16">16–17.IX.2002</collectingDate>
</date>
, 
<collectorName id="F638A7BA2426173EFD89B68E11891E26" box="[638,690,1014,1040]" pageId="42" pageNumber="343">Stop</collectorName>
#DHK-2002-018, 
<collectorName id="F638A7BA2426173EFC89B68F17061E26" box="[894,1085,1014,1041]" pageId="42" pageNumber="343">D.H. Kavanaugh</collectorName>
, 
<collectorName id="F638A7BA2426173EFBBFB68F17FF1E26" box="[1096,1220,1014,1041]" pageId="42" pageNumber="343">H.B. Liang</collectorName>
leg. (
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)
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; 
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<specimenCount id="4DCB09E52426173EFF60B16213C61E02" box="[151,253,1050,1076]" pageId="42" pageNumber="343" type="female">1 female</specimenCount>
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<location id="5E1294B72426173EFEFEB16211241E03" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFEFEB16211241E03" box="[265,543,1050,1077]" country="China" latitude="24.750555" longLatPrecision="21" longitude="100.50667" name="Wuliangshan Mt. Range" pageId="42" pageNumber="343" stateProvince="Yunnan">Wuliangshan Mt. Range</location>
, 
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<quantity id="9C356F892426173EFCA0B162108B1E03" box="[855,944,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="42" pageNumber="343" unit="m" value="2270.0">
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, 
<date id="2F73E4AC2426173EFC49B16217041E02" box="[958,1087,1050,1076]" pageId="42" pageNumber="343" value="2011-06-12">
<collectingDate id="3F371D442426173EFC49B16217041E02" box="[958,1087,1050,1076]" pageId="42" pageNumber="343" value="2011-06-12">12.VI.2011</collectingDate>
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, 
<collectorName id="F638A7BA2426173EFBBBB16317EB1E02" box="[1100,1232,1050,1077]" pageId="42" pageNumber="343">I. Belousov</collectorName>
, 
<collectorName id="F638A7BA2426173EFB2BB163167A1E02" box="[1244,1345,1050,1077]" pageId="42" pageNumber="343">I. Kabak</collectorName>
, 
<collectorName id="F638A7BA2426173EFABAB16213EA1E6E" pageId="42" pageNumber="343">A. Korolev</collectorName>
leg. (cB&amp;K, 
<collectionCode id="3DDC5AA92426173EFE9FB14612A01E6E" box="[360,411,1086,1112]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="42" pageNumber="343" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5B72C26C2426173EFF30B11A10F01E96" blockId="42.[151,1437,582,2013]" pageId="42" pageNumber="343">
<materialsCitation id="EBA5C8312426173EFF30B11A10FC1E96" ID-GBIF-Occurrence="3821894582" collectingDate="2018-10-03" collectingDateMax="2018-10-11" collectingDateMin="2018-10-03" collectionCode="SIEE, ZIN" collectorName="N Vietnam &amp; D. Fedorenko" country="Vietnam" elevation="1675" location="Phia Oac Mt." pageId="42" pageNumber="343" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Cao Bang">
<emphasis id="69B91E7E2426173EFF30B11A120F1E4A" bold="true" box="[199,308,1122,1148]" pageId="42" pageNumber="343">
<collectingCountry id="23DA82FC2426173EFF30B11A12141E4A" box="[199,303,1122,1148]" name="Vietnam" pageId="42" pageNumber="343">Vietnam</collectingCountry>
.
</emphasis>
<collectingRegion id="99090C8E2426173EFECCB11A12E91E4A" box="[315,466,1122,1148]" country="Vietnam" name="Cao Bang" pageId="42" pageNumber="343">CAO BANG</collectingRegion>
: 
<specimenCount id="4DCB09E52426173EFE2BB11A110E1E4A" box="[476,565,1122,1149]" pageId="42" pageNumber="343" type="male">5 males</specimenCount>
, 
<specimenCount id="4DCB09E52426173EFDB6B11A118B1E4A" box="[577,688,1122,1148]" pageId="42" pageNumber="343" type="female">3 females</specimenCount>
, 
<collectorName id="F638A7BA2426173EFD4BB11B10021E4A" box="[700,825,1122,1149]" pageId="42" pageNumber="343">N Vietnam</collectorName>
, 
<locationDeviation id="4AC0DB4E2426173EFCB3B11A177F1E4B" box="[836,1092,1122,1149]" location="Cao Bang" pageId="42" pageNumber="343">
<quantity id="9C356F892426173EFCB3B11A10B01E4A" box="[836,907,1122,1148]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="42" pageNumber="343" unit="km" value="40.0">40 km</quantity>
W of Cao Bang
</locationDeviation>
, 
<location id="5E1294B72426173EFBB8B11A17DC1E4B" LSID="urn:lsid:plazi:treatment:D364737A2426173AFF60B4A217F31ACF:5E1294B72426173EFBB8B11A17DC1E4B" box="[1103,1255,1122,1149]" country="Vietnam" name="Phia Oac Mt." pageId="42" pageNumber="343" stateProvince="Cao Bang">Phia Oac Mt.</location>
, h ~ 
<quantity id="9C356F892426173EFAE9B11A13941E97" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="42" pageNumber="343" unit="m" value="1675.0" valueMax="1700.0" valueMin="1650.0">
<elevation id="D0E0255F2426173EFAE9B11A13941E97" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="42" pageNumber="343" unit="m" value="1675.0" valueMax="1700.0" valueMin="1650.0">1650–1700 m</elevation>
</quantity>
, deciduous forest, 
<date id="2F73E4AC2426173EFE76B1FE112E1E96" box="[385,533,1158,1184]" pageId="42" pageNumber="343" value="2018-10-03" valueMax="2018-10-11" valueMin="2018-10-03">
<collectingDate id="3F371D442426173EFE76B1FE112E1E96" box="[385,533,1158,1184]" pageId="42" pageNumber="343" value="2018-10-03" valueMax="2018-10-11" valueMin="2018-10-03">3–11.X.2018</collectingDate>
</date>
, 
<collectorName id="F638A7BA2426173EFDD7B1FF11801E96" box="[544,699,1158,1185]" pageId="42" pageNumber="343">D. Fedorenko</collectorName>
leg. (
<collectionCode id="3DDC5AA92426173EFD0DB1FE10021E96" box="[762,825,1158,1184]" pageId="42" pageNumber="343">SIEE</collectionCode>
, 
<collectionCode id="3DDC5AA92426173EFCB3B1FF104E1E97" box="[836,885,1159,1185]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="42" pageNumber="343" type="Museum">ZIN</collectionCode>
, cWR)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="13D791E724261739FF30B1D212B41F63" lastPageId="45" lastPageNumber="346" pageId="42" pageNumber="343" type="description">
<paragraph id="5B72C26C2426173EFF30B1D2163C1EF2" blockId="42.[151,1437,582,2013]" box="[199,1287,1194,1221]" pageId="42" pageNumber="343">
<emphasis id="69B91E7E2426173EFF30B1D2126A1EF2" bold="true" box="[199,337,1194,1220]" pageId="42" pageNumber="343">Description</emphasis>
(
<specimenCount id="4DCB09E52426173EFE96B1D212831EF2" box="[353,440,1194,1220]" pageId="42" pageNumber="343" type="male">6 males</specimenCount>
and 
<specimenCount id="4DCB09E52426173EFE18B1D211601EF2" box="[495,603,1194,1220]" pageId="42" pageNumber="343" type="female">2 females</specimenCount>
measured). Body length 8.0– 
<quantity id="9C356F892426173EFC51B1D210C41EF3" box="[934,1023,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="42" pageNumber="343" unit="mm" value="9.4">9.4 mm</quantity>
. Habitus as in 
<figureCitation id="C3F6DEE92426173EFB54B1D216381EF2" box="[1187,1283,1194,1220]" captionStart="FIGURES 107–108" captionStartId="43.[151,264,1956,1980]" captionTargetBox="[157,1420,650,1924]" captionTargetId="figure-222@43.[153,1433,633,1933]" captionTargetPageId="43" captionText="FIGURES 107–108. Trichotichnus ssp., habitus. 107, T. miwai continentalis ssp. n. (Wuliangshan); 108, T. kavanaughi sp. n. (Kongdang). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="42" pageNumber="343">Fig. 107</figureCitation>
.
</paragraph>
<paragraph id="5B72C26C2426173EFF30B1B617AB1F06" blockId="42.[151,1437,582,2013]" pageId="42" pageNumber="343">Body dark brown to black, shiny, iridescent on elytra; labrum, base of mandibles, in many specimens also clypeus (wholly or along anterior margin) and narrow lateral margins of pronotum reddish brown. Palpi, antennae, tibiae and tarsi brownish yellow or light brown; femora infuscate, dark brown to blackish brown.</paragraph>
<paragraph id="5B72C26C2426173EFF30B042111E1DA2" blockId="42.[151,1437,582,2013]" pageId="42" pageNumber="343">
Head moderately sized (HWmax/PWmax 0.72–0.78, HWmin/PWmax 0.56–0.58). Eyes large and convex (HWmax/HWmin 1.28–1.33), in lateral view wide oval. Genae very narrow, narrower than antennomere 2 basally; occasionally eyes touching buccal incisure. Tempora comparatively short, oblique. Fronto-clypeal suture slightly impressed, almost straight. Fronto-ocular furrows reaching supraorbital furrows, deepened at clypeus and superficial at eyes. Supraorbital furrows narrow, touching upper margin of eye. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eye and removed from supraorbital furrows by distance approximately equal to width of antennomere 2 apically; in many specimens one or two very small unsetigerous punctures present medially of each supraorbital seta. Labrum very shallowly concave anteriorly. Clypeus slightly convex, very shallowly emarginate along anterior margin, with a pair of setigerous pores each located at anterior angle. 
<collectingRegion id="99090C8E2426173EFACAB32316A71C43" box="[1341,1436,1627,1653]" country="Myanmar" name="Chin" pageId="42" pageNumber="343">Mentum</collectingRegion>
separated from submentum, with wide and short median tooth; epilobes markedly widened apically; submentum with one pair of long setae. Ligular sclerite widened apically, truncate at apex, with one pair of ventroapical setae. Paraglossae very narrow, about as long as ligular sclerite and separated from it by wide notches. Labial penultimate palpomere about as long as ultimate palpomere. Left mandible sharply bent approximately in the apical quarter, blunted or obliquely truncate at tip. Dorsal microsculpture very obliterated, consisting of indistinct, more or less isodiametric meshes. Antennae slender and long, surpassing pronotal basal edge in male by four, in female by three and a half apical antennomeres, with antennomeres 4–8 about three times as long as wide. Basal antennomere slightly longer than antennomere 3.
</paragraph>
<paragraph id="5B72C26C2426173FFF30B2E712E21857" blockId="42.[151,1437,582,2013]" lastBlockId="43.[151,1437,151,609]" lastPageId="43" lastPageNumber="344" pageId="42" pageNumber="343">Pronotum transverse (PWmax/PL 1.40–1.46), widest slightly before the middle (PWmax/PWmin 1.21–1.32), more strongly narrowed apically than basally, with one lateral seta slightly removed from lateral edge to internal margin of lateral depression and inserted slightly before widest point of pronotum. Sides rounded along apical two thirds; in basal third straightly or rounded converging, occasionally very shallowly sinuate; lateral bead narrow. Apical margin shallowly emarginate, very narrowly bordered along entire length or border interrupted at the middle. Apical angles narrowly rounded, slightly protruding ahead. Basal margin more or less straight, distinctly bordered along entire length, slightly longer than apical margin and markedly shorter than base of elytra between humeral angles. Basal angles obtusangular, with a very small obtuse denticle at tip. Pronotal disc medially moderately convex. Lateral depression beginning from apical angles, narrow in apical third, rather strongly widened in the basal two third and fused with the wide and deep laterobasal depressions; basal foveae located at internal margins of laterobasal depressions, elongate, occasionally indistinct; basal area medially convex but slightly depressed as compared with disc. Median line distinct, slightly impressed. Anterior transverse depression wide and deepened. Surface densely punctate in a wide area along margins, with coarsest punctures in laterobasal depressions; central portion of disc impunctate or with very fine and sparse punctures. Microsculpture highly obliterated, consisting of indistinct transverse meshes.</paragraph>
<caption id="0FB292E42427173FFF60B2DC12D01DD6" ID-DOI="http://doi.org/10.5281/zenodo.6781763" ID-Zenodo-Dep="6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="43" pageNumber="344" startId="43.[151,264,1956,1980]" targetBox="[157,1420,650,1924]" targetPageId="43">
<paragraph id="5B72C26C2427173FFF60B2DC12D01DD6" blockId="43.[151,1436,1956,2017]" pageId="43" pageNumber="344">
<emphasis id="69B91E7E2427173FFF60B2DC12551D8B" bold="true" box="[151,366,1956,1981]" pageId="43" pageNumber="344">FIGURES 107–108.</emphasis>
<taxonomicName id="9CCDB9EF2427173FFE83B2DD12C41D8B" authorityName="Morawitz" authorityYear="1863" box="[372,511,1957,1981]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="69B91E7E2427173FFE83B2DD12C41D8B" box="[372,511,1957,1981]" italics="true" pageId="43" pageNumber="344">Trichotichnus</emphasis>
</taxonomicName>
ssp., habitus. 107, 
<taxonomicName id="9CCDB9EF2427173FFD33B2DE109E1D8B" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[708,933,1957,1981]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="subSpecies" species="miwai" subSpecies="continentalis">
<emphasis id="69B91E7E2427173FFD33B2DE109E1D8B" box="[708,933,1957,1981]" italics="true" pageId="43" pageNumber="344">T. miwai continentalis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2427173FFC5BB2DD10D41D8B" bold="true" box="[940,1007,1957,1981]" pageId="43" pageNumber="344">ssp. n.</emphasis>
(Wuliangshan); 108, 
<taxonomicName id="9CCDB9EF2427173FFB3BB2DE16671D8B" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[1228,1372,1957,1981]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="species" species="kavanaughi" status="sp. nov.">
<emphasis id="69B91E7E2427173FFB3BB2DE16671D8B" box="[1228,1372,1957,1981]" italics="true" pageId="43" pageNumber="344">T. kavanaughi</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2427173FFA94B2DD16A71D8B" bold="true" box="[1379,1436,1957,1981]" pageId="43" pageNumber="344">
<taxonomicNameLabel id="728AA3052427173FFA94B2DD16A71D8B" box="[1379,1436,1957,1981]" pageId="43" pageNumber="344" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Kongdang). Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="0FB292E424201738FF60B39B164E1DBD" ID-DOI="http://doi.org/10.5281/zenodo.6781767" ID-Zenodo-Dep="6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="44" pageNumber="345" startId="44.[151,264,1763,1787]" targetBox="[165,1420,194,1727]" targetPageId="44">
<paragraph id="5B72C26C24201738FF60B39B164E1DBD" blockId="44.[151,1437,1763,1931]" pageId="44" pageNumber="345">
<emphasis id="69B91E7E24201738FF60B39B12561CCD" bold="true" box="[151,365,1763,1787]" pageId="44" pageNumber="345">FIGURES 109–120.</emphasis>
<taxonomicName id="9CCDB9EF24201738FE84B39B10791CCD" authority="Jedlicka, 1949: 109 - 113" authorityName="Jedlicka" authorityPageNumber="109 - 113" authorityYear="1949" box="[371,834,1763,1787]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="345" phylum="Arthropoda" rank="species" species="miwai">
<emphasis id="69B91E7E24201738FE84B39B11791CCD" box="[371,578,1763,1787]" italics="true" pageId="44" pageNumber="345">Trichotichnus miwai</emphasis>
<bibRefCitation id="3F5CBF9D24201738FDBFB39B10791CCD" author="Jedlicka, A." box="[584,834,1763,1787]" pageId="44" pageNumber="345" pagination="108 - 111" refId="ref38451" refString="Jedlicka, A. (1949) Rod Trichotichnus Moraw. Z Ostrova Formosy. Les especes du genre Trichotichnus Moraw. de l'ile de Formose. Casopis Ceskoslovenske Spolecnosti Entomologicke, 46, 108 - 111." type="journal article" year="1949">Jedlička, 1949: 109–113</bibRefCitation>
</taxonomicName>
, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, 
<taxonomicName id="9CCDB9EF24201738FE94B270117E1D29" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[355,581,1799,1823]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="345" phylum="Arthropoda" rank="subSpecies" species="miwai" subSpecies="continentalis">
<emphasis id="69B91E7E24201738FE94B270117E1D29" box="[355,581,1799,1823]" italics="true" pageId="44" pageNumber="345">T. miwai continentalis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24201738FDBBB27F11AB1D29" bold="true" box="[588,656,1799,1823]" pageId="44" pageNumber="345">ssp. n.</emphasis>
(114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118).
</paragraph>
</caption>
<paragraph id="5B72C26C24211739FF30B5EF10FE1BE7" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">Elytra convex, elongate oval (EL/EW 1.54–1.63, EL/PL 3.08–3.16, EW/PWmax 1.33–1.41), widest in the apical third; sides almost rectilinearly diverging at the middle. Humeri rounded, humeral denticle absent or indistinct, at most poorly recognizable in caudal view. Subapical sinuation shallow. Sutural angle acutangular, blunted or very narrowly rounded at tip. Basal edge slightly arched or almost straight; internal humeral angle rounded. Striae impunctate, reaching anteriorly basal edge, impressed along entire length (slightly more deeply apically). Intervals almost flat or faintly convex on disc, markedly narrowed and moderately convex apically. Parascutellar (abbreviate) striole long, with a small setigerous pore basally. Interval 3 with one small setigerous pore adjacent to stria 2 behind the middle. Marginal umbilicate series without distinct gap at middle. Microsculpture strongly obliterated, consisting of indistinct transverse lines recognizable mostly on lateral intervals.</paragraph>
<paragraph id="5B72C26C24211739FF30B4A312F31BC3" blockId="45.[151,1437,150,2013]" box="[199,456,475,501]" pageId="45" pageNumber="346">Wings well developed.</paragraph>
<paragraph id="5B72C26C24211739FF30B48711BD180B" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">Prosternum with very fine short setae. Proepisternum finely punctate. Metepisternum about 1.40–1.60 times as long as wide, strongly narrowed posteriorly.</paragraph>
<paragraph id="5B72C26C24211739FF30B73F116219FF" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
Legs relatively long. Metacoxae without additional setae. Metafemur ventrally with two setae at posterior margin and without setae at anterior margin (with 1–3 short setae just on anterior margin preapically). Protibia on dorsal side without longitudinal sulcus, on outer margin preapically with one stouter and two very thin spines in male and with three comparatively stout, similar sized spines in female. Tarsi dorsally glabrous; pro- and mesotarsomeres 1–4, particularly in male, with dense long ventro-lateral setae (as in 
<figureCitation id="C3F6DEE924211739FC68B7AF10C818C7" box="[927,1011,727,753]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 &amp; 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="45" pageNumber="346">Fig. 20</figureCitation>
); tarsomere 5 with four or three pairs of ventro-lateral setae. Metatarsus slender, about as long as (occasionally slightly shorter or longer than) HWmax, with tarsomeres somewhat weakly widened distally; metatarsomere 1 slightly shorter than metatarsomeres 2 and 3 combined; metatarsomeres 1–4 rather densely setose ventrally. In male, pro- and mesotarsomeres 2–4 moderately widened and with adhesive scales ventrally, mesotarsomere 1 weakly widened and only with a pair of scales ventrally; pro- and mesotarsomeres 2 and 3 longer than wide. In both sexes, mesotarsomere 1 elongate, slightly shorter than mesotarsomeres 2 and 3 combined.
</paragraph>
<paragraph id="5B72C26C24211739FF30B6AB127A1E03" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">Abdominal sternites without additional long setae; apex of last visible abdominal sternite (VII) in male widely rounded or subtruncate, with one pair of marginal setae, in female more narrowly rounded and with two pairs of marginal setae.</paragraph>
<paragraph id="5B72C26C24211739FF30B147107A1F3B" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
Median lobe of aedeagus (
<figureCitation id="C3F6DEE924211739FE1AB14711B61E6F" box="[493,653,1087,1113]" captionStart="FIGURES 109–120" captionStartId="44.[151,264,1763,1787]" captionTargetBox="[165,1420,194,1727]" captionTargetId="figure-32@44.[151,1436,181,1739]" captionTargetPageId="44" captionText="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118)." figureDoi="http://doi.org/10.5281/zenodo.6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="45" pageNumber="346">Figs. 114–118</figureCitation>
) comparatively slender, in lateral view arcuate on dorsal side and with almost straight ventral margin in its medial portion; in dorsal view with mostly equal width basally and preapically; ventral side edged laterally in apical part and depressed between margins; terminal lamella (114, 117) about as wide as long, evenly narrowed posteriorly, slightly concave on dorsal side and narrowly rounded at apex; in lateral view thin, angularly bent ventrally just at apex. Apical orifice in dorsal position, wide apically. Internal sac with a medium-sized elongate spine on the left side behind the middle.
</paragraph>
<paragraph id="5B72C26C24211739FF30B06F12B41F63" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
Female genitalia (
<figureCitation id="C3F6DEE924211739FE63B06F116D1F07" box="[404,598,1303,1329]" captionStart="FIGURES 109–120" captionStartId="44.[151,264,1763,1787]" captionTargetBox="[165,1420,194,1727]" captionTargetId="figure-32@44.[151,1436,181,1739]" captionTargetPageId="44" captionText="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118)." figureDoi="http://doi.org/10.5281/zenodo.6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="45" pageNumber="346">Figs. 119 &amp; 120</figureCitation>
): gonocoxite narrow in ventral view, about 0.6 as long as gonosubcoxite, with rather wide base.
</paragraph>
</subSubSection>
<subSubSection id="13D791E724211739FF30B027166B1C1B" pageId="45" pageNumber="346" type="etymology">
<paragraph id="5B72C26C24211739FF30B02713CB1FAB" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
<emphasis id="69B91E7E24211739FF30B02712751F4F" bold="true" box="[199,334,1375,1401]" pageId="45" pageNumber="346">Etymology.</emphasis>
The subspecific name is a Latin adjective, referring to the continental distribution of the new subspecies.
</paragraph>
<paragraph id="5B72C26C24211739FF30B0DE166B1C1B" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
<emphasis id="69B91E7E24211739FF30B0DE12591FF6" bold="true" box="[199,354,1446,1472]" pageId="45" pageNumber="346">Comparison.</emphasis>
<taxonomicName id="9CCDB9EF24211739FE9CB0DF11D71FF7" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[363,748,1447,1473]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="subSpecies" species="miwai" status="subsp. nov." subSpecies="continentalis">
<emphasis id="69B91E7E24211739FE9CB0DF11D71FF7" box="[363,748,1447,1473]" italics="true" pageId="45" pageNumber="346">Trichotichnus miwai continentalis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24211739FD02B0DF107A1FF7" bold="true" box="[757,833,1447,1473]" pageId="45" pageNumber="346">
<taxonomicNameLabel id="728AA30524211739FD02B0DF107A1FF7" box="[757,833,1447,1473]" pageId="45" pageNumber="346" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
differs from the nominotypical subspecies in having elytra without distinct humeral denticle, femora constantly infuscate, and median lobe of aedeagus with terminal lamella less strongly narrowed apically and with larger spine in internal sac (
<figureCitation id="C3F6DEE924211739FC0DB097160C1C3F" box="[1018,1335,1519,1545]" captionStart="FIGURES 109–120" captionStartId="44.[151,264,1763,1787]" captionTargetBox="[165,1420,194,1727]" captionTargetId="figure-32@44.[151,1436,181,1739]" captionTargetPageId="44" captionText="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118)." figureDoi="http://doi.org/10.5281/zenodo.6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="45" pageNumber="346">Figs. 109–113 and 114–118</figureCitation>
). Among the Chinese species occurring in 
<collectingRegion id="99090C8E24211739FDF0B36B115B1C1B" box="[519,608,1555,1581]" country="China" name="Yunnan" pageId="45" pageNumber="346">Yunnan</collectingRegion>
, this taxon is most similar to 
<taxonomicName id="9CCDB9EF24211739FC5FB36B177C1C1B" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[936,1095,1555,1581]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="kavanaughi" status="sp. nov.">
<emphasis id="69B91E7E24211739FC5FB36B177C1C1B" box="[936,1095,1555,1581]" italics="true" pageId="45" pageNumber="346">T. kavanaughi</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24211739FBBAB36B17B61C1B" bold="true" box="[1101,1165,1555,1581]" pageId="45" pageNumber="346">
<taxonomicNameLabel id="728AA30524211739FBBAB36B17B61C1B" box="[1101,1165,1555,1581]" pageId="45" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described below.
</paragraph>
</subSubSection>
<subSubSection id="13D791E724211739FF30B34E161F1C43" pageId="45" pageNumber="346" type="distribution">
<paragraph id="5B72C26C24211739FF30B34E161F1C43" blockId="45.[151,1437,150,2013]" pageId="45" pageNumber="346">
<emphasis id="69B91E7E24211739FF30B34E12621C66" bold="true" box="[199,345,1590,1616]" pageId="45" pageNumber="346">Distribution</emphasis>
(
<figureCitation id="C3F6DEE924211739FE9EB34F12F71C67" box="[361,460,1591,1617]" captionStart="FIGURE 121" captionStartId="46.[151,250,1625,1649]" captionTargetBox="[151,1434,275,1599]" captionTargetId="figure-80@46.[151,1435,273,1602]" captionTargetPageId="46" captionText="FIGURE 121. Trichotichnus miwai continentalis ssp. n., distribution." figureDoi="http://doi.org/10.5281/zenodo.6781769" httpUri="https://zenodo.org/record/6781769/files/figure.png" pageId="45" pageNumber="346">Fig. 121</figureCitation>
). The new subspecies is known from 
<collectingCountry id="23DA82FC24211739FC81B34F10821C67" box="[886,953,1591,1617]" name="China" pageId="45" pageNumber="346">China</collectingCountry>
(
<collectingRegion id="99090C8E24211739FC3FB34F171B1C67" box="[968,1056,1591,1617]" country="China" name="Yunnan" pageId="45" pageNumber="346">Yunnan</collectingRegion>
and southern 
<collectingRegion id="99090C8E24211739FB48B34F16201C67" box="[1215,1307,1591,1617]" country="China" name="Sichuan" pageId="45" pageNumber="346">Sichuan</collectingRegion>
) and North 
<collectingCountry id="23DA82FC24211739FF60B32313CD1C43" box="[151,246,1627,1653]" name="Vietnam" pageId="45" pageNumber="346">Vietnam</collectingCountry>
where it lives at altitudes of about 
<quantity id="9C356F8924211739FD88B32210261C43" box="[639,797,1626,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.81" metricValueMax="2.27" metricValueMin="1.35" pageId="45" pageNumber="346" unit="m" value="1810.0" valueMax="2270.0" valueMin="1350.0">1350–2270 m</quantity>
. The nominative subspecies occurs in 
<collectingCountry id="23DA82FC24211739FB3DB323161B1C43" box="[1226,1312,1627,1653]" name="Taiwan" pageId="45" pageNumber="346">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="13D791E72421173AFF30B30717F31ACF" lastPageId="46" lastPageNumber="347" pageId="45" pageNumber="346" type="discussion">
<paragraph id="5B72C26C2421173AFF30B30717F31ACF" blockId="45.[151,1437,150,2013]" lastBlockId="46.[151,1437,151,249]" lastPageId="46" lastPageNumber="347" pageId="45" pageNumber="346">
<emphasis id="69B91E7E24211739FF30B30712001CAF" bold="true" box="[199,315,1663,1689]" pageId="45" pageNumber="346">Remarks.</emphasis>
<taxonomicName id="9CCDB9EF24211739FEB2B30711E01CAF" authority="Jedlicka, 1949" authorityName="Jedlicka" authorityYear="1949" box="[325,731,1663,1689]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="miwai">
<emphasis id="69B91E7E24211739FEB2B30711161CAE" box="[325,557,1663,1689]" italics="true" pageId="45" pageNumber="346">Trichotichnus miwai</emphasis>
<bibRefCitation id="3F5CBF9D24211739FDC0B30711E01CAF" author="Jedlicka, A." box="[567,731,1663,1689]" pageId="45" pageNumber="346" pagination="108 - 111" refId="ref38451" refString="Jedlicka, A. (1949) Rod Trichotichnus Moraw. Z Ostrova Formosy. Les especes du genre Trichotichnus Moraw. de l'ile de Formose. Casopis Ceskoslovenske Spolecnosti Entomologicke, 46, 108 - 111." type="journal article" year="1949">Jedlička, 1949</bibRefCitation>
</taxonomicName>
was described from 
<specimenCount id="4DCB09E524211739FC38B30717431CAF" box="[975,1144,1663,1689]" pageId="45" pageNumber="346" type="generic">two specimens</specimenCount>
from 
<collectingCountry id="23DA82FC24211739FB36B30716281CAF" box="[1217,1299,1663,1689]" name="Taiwan" pageId="45" pageNumber="346">Taiwan</collectingCountry>
(“Formose: Arisan”). The brief original description (
<bibRefCitation id="3F5CBF9D24211739FD9CB3DB10301C8B" author="Jedlicka, A." box="[619,779,1699,1725]" pageId="45" pageNumber="346" pagination="108 - 111" refId="ref38451" refString="Jedlicka, A. (1949) Rod Trichotichnus Moraw. Z Ostrova Formosy. Les especes du genre Trichotichnus Moraw. de l'ile de Formose. Casopis Ceskoslovenske Spolecnosti Entomologicke, 46, 108 - 111." type="journal article" year="1949">Jedlička 1949</bibRefCitation>
) was later supplemented by 
<bibRefCitation id="3F5CBF9D24211739FBADB3DB17D01C8B" author="Habu, A." box="[1114,1259,1699,1725]" pageId="45" pageNumber="346" pagination="9 - 19" refId="ref37998" refString="Habu, A. (1980) Some species of Trichotichnus from Formosa (Coleoptera, Carabidae). The Entomological Review of Japan, 34, 9 - 19." type="journal article" year="1980">Habu (1980)</bibRefCitation>
who examined 
<specimenCount id="4DCB09E524211739FF60B3BF12211CD7" box="[151,282,1735,1761]" pageId="45" pageNumber="346" type="male">three males</specimenCount>
from Liu Gui-Xiang (Taiwan) and described the aedeagus and some external distinctive features of this species as compared to 
<taxonomicName id="9CCDB9EF24211739FE39B39311511D32" authorityName="Habu" authorityYear="1954" box="[462,618,1771,1796]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="vespertinus">
<emphasis id="69B91E7E24211739FE39B39311511D32" box="[462,618,1771,1796]" italics="true" pageId="45" pageNumber="346">T. vespertinus</emphasis>
</taxonomicName>
from 
<collectingCountry id="23DA82FC24211739FD5AB39311CB1D33" box="[685,752,1771,1797]" name="Japan" pageId="45" pageNumber="346">Japan</collectingCountry>
. We examined 
<specimenCount id="4DCB09E524211739FC6DB393177B1D33" box="[922,1088,1771,1797]" pageId="45" pageNumber="346" type="generic">two specimens</specimenCount>
from 
<collectingCountry id="23DA82FC24211739FB74B39317E21D33" box="[1155,1241,1771,1797]" name="Taiwan" pageId="45" pageNumber="346">Taiwan</collectingCountry>
: 
<specimenCount id="4DCB09E524211739FB14B393160A1D33" box="[1251,1329,1771,1797]" pageId="45" pageNumber="346" type="male">1 male</specimenCount>
, 
<collectingRegion id="99090C8E24211739FACAB39316A21D33" box="[1341,1433,1771,1797]" country="Taiwan" name="Hualien" pageId="45" pageNumber="346">Hualien</collectingRegion>
, Huojanting, 
<date id="2F73E4AC24211739FED4B277129E1D1F" box="[291,421,1807,1833]" pageId="45" pageNumber="346" value="2007-09-12">12.IX.2007</date>
, Y.-F. Hsu leg. (cMRR), and 
<specimenCount id="4DCB09E524211739FD1EB277100C1D1F" box="[745,823,1807,1833]" pageId="45" pageNumber="346" type="male">1 male</specimenCount>
, 
<collectingRegion id="99090C8E24211739FCB5B27710AC1D1F" box="[834,919,1807,1833]" country="Taiwan" name="Nantou" pageId="45" pageNumber="346">Nantou</collectingRegion>
, Songgang, light, sweeping, 
<date id="2F73E4AC24211739FB2EB27716601D1F" box="[1241,1371,1807,1833]" pageId="45" pageNumber="346" value="2009-06-20">20.VI.2009</date>
, Y.-T. Wang leg. (cMRR), both of which agree well with the original and 
<bibRefCitation id="3F5CBF9D24211739FC66B24B17091D7B" author="Habu, A." box="[913,1074,1843,1869]" pageId="45" pageNumber="346" pagination="9 - 19" refId="ref37998" refString="Habu, A. (1980) Some species of Trichotichnus from Formosa (Coleoptera, Carabidae). The Entomological Review of Japan, 34, 9 - 19." type="journal article" year="1980">Habu’s (1980)</bibRefCitation>
descriptions. Their proportions, respectively: HWmax/PWmax 0.75 and 0.74; HWmin/PWmax 0.58 and 0.57; HWmax/HWmin 1.29 and 1.30; PWmax/PL 1.47 and 1.43; PWmax/PWmin 1.31 and 1.22; EL/EW 1.57 and 1.60; EL/PL 3.15 and 3.06; EW/PWmax 1.36–1.34; body length 9.0 and 
<quantity id="9C356F8924211739FE01B2E711741D8F" box="[502,591,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="45" pageNumber="346" unit="mm" value="9.1">9.1 mm</quantity>
. In both specimens, the dorsum is dark brown, almost black; the appendages are brownish yellow, with the femora not infuscate in the male from 
<collectingRegion id="99090C8E24211739FC6CB2BB10CF1DEB" box="[923,1012,1987,2013]" country="Taiwan" name="Hualien" pageId="45" pageNumber="346">Hualien</collectingRegion>
and infuscate (blackish brown) in the male from 
<collectingRegion id="99090C8E2422173AFEE4B5EF125E1A87" box="[275,357,151,177]" country="Taiwan" name="Nantou" pageId="46" pageNumber="347">Nantou</collectingRegion>
(legs “testacé-rougeâtre” according to the original description). The aedeagi of these specimens are illustrated in 
<figureCitation id="C3F6DEE92422173AFEAFB5C312C71AE3" box="[344,508,187,213]" captionStart="FIGURES 109–120" captionStartId="44.[151,264,1763,1787]" captionTargetBox="[165,1420,194,1727]" captionTargetId="figure-32@44.[151,1436,181,1739]" captionTargetPageId="44" captionText="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 &amp; 110, Nantou; 111 &amp; 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 &amp; 118, Phia Oac, Vietnam; 119 &amp; 120, Kunming). 109, 111, 114 &amp; 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 &amp; 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 &amp; 117), lateral (110, 112, 115, 118 &amp; 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 &amp; 120), B = 1.0 mm (110, 112, 113, 115, 116 &amp; 118)." figureDoi="http://doi.org/10.5281/zenodo.6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="46" pageNumber="347">Figs. 109–113</figureCitation>
. Since the available descriptions of 
<taxonomicName id="9CCDB9EF2422173AFC63B5C310CE1AE2" authorityName="Jedlicka" authorityYear="1949" box="[916,1013,187,212]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="347" phylum="Arthropoda" rank="species" species="miwai">
<emphasis id="69B91E7E2422173AFC63B5C310CE1AE2" box="[916,1013,187,212]" italics="true" pageId="46" pageNumber="347">T. miwai</emphasis>
</taxonomicName>
are rather brief, we describe the new subspecies in more detail. Most of the included characteristics are common for both subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>