467 lines
52 KiB
XML
467 lines
52 KiB
XML
<document id="32A1A24CE81CB323027A5D1D8360F08B" ID-DOI="10.3897/zookeys.1213.133178" ID-ZooBank="B8E6EE1A-023D-4B64-8D3A-C1843F0D8376" ID-publisher-id="133178" URI-arpha="B0B2A119-6CED-5B4F-810D-44C4C6D0F4F3" URI-zoobank="http://zoobank.org/B8E6EE1A-023D-4B64-8D3A-C1843F0D8376" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1727330124077" checkinUser="pensoft" docAuthor="Huber, Bernhard A. & Meng, Guanliang" docDate="2024" docId="933148E97DF357DC8194F5ED2DFB17B1" docLanguage="en" docName="ZooKeys 1213: 95-182" docOrigin="ZooKeys 1213" docSource="https://zookeys.pensoft.net/article/133178/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Micropholcus tanomah Huber & Meng 2024, sp. nov." docType="treatment" docUuid="6D679588-8300-427F-89A7-6CA93102C27C" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="182" masterDocId="B0B2A1196CED5B4F810D44C4C6D0F4F3" masterDocTitle="Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae)" masterLastPageNumber="182" masterPageNumber="95" pageNumber="95" updateTime="1727371993187" updateUser="admin">
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<mods:title id="B6A5DF1B8EAF9ECEE7255ACF3CD285A8">Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae)</mods:title>
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<mods:name id="C9D3D40A7502928C8024F3C200CF210A" type="personal">
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<mods:namePart id="096EE3E78318C4EA83EE0AFE510D87A5">Huber, Bernhard A.</mods:namePart>
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<mods:nameIdentifier id="1B12781F60440372AC56BF5A8BCE41AB" type="ORCID">0000-0002-7566-5424</mods:nameIdentifier>
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<mods:affiliation id="C450DCA5493B0DD1F6C72ECFD0272E7F">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:roleTerm id="ACD02C3F09F838EC178124D74A1791EB">Author</mods:roleTerm>
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<mods:namePart id="330A9C154915032BFD25C6C3F2B9BE66">Meng, Guanliang</mods:namePart>
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<mods:nameIdentifier id="279C149DA450BC40D5216D12EF21D2B3" type="ORCID">0000-0002-6488-1527</mods:nameIdentifier>
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<mods:affiliation id="E8FA4789027352A7F38E34FB986AB746">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:typeOfResource id="696D1546A031D2C527EAA001D07A855C">text</mods:typeOfResource>
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<mods:title id="73B4C9DDA643CFA5F6D5D391CF094BB0">ZooKeys</mods:title>
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<mods:part id="6CC8926023E1FDB5EE5A3F4153E66D7E">
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<mods:date id="222585EB9B855E62210ECD9C0A45635E">2024</mods:date>
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<mods:number id="02E296C6ED8EBC430D41575F2F65F91D">2024-09-26</mods:number>
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<mods:number id="5C810787D24FBC01AB4BC35FC67F6F8F">1213</mods:number>
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<treatment id="933148E97DF357DC8194F5ED2DFB17B1" ID-DOI="http://doi.org/10.5281/zenodo.13841462" ID-Zenodo-Dep="13841462" ID-arpha="933148E9-7DF3-57DC-8194-F5ED2DFB17B1" ID-zoobank="https://zoobank.org/6D679588-8300-427F-89A7-6CA93102C27C" LSID="urn:lsid:zoobank.org:act:6D679588-8300-427F-89A7-6CA93102C27C" LSID-ZBK="urn:lsid:zoobank.org:act:6D679588-8300-427F-89A7-6CA93102C27C" httpUri="http://treatment.plazi.org/id/933148E97DF357DC8194F5ED2DFB17B1">
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<subSubSection id="D38F26C57BD4181981E734EB63FC091A" type="nomenclature">
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<paragraph id="FB15A32F00A3FDC6A0326DFD270049C5">
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<taxonomicName id="1BAD335CDC03FEA1597C7668DA145AA8" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
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<emphasis id="CC7896FD70711E7BCB65FAF383DEE740" italics="true">Micropholcus tanomah</emphasis>
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Huber
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</taxonomicName>
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<taxonomicNameLabel id="411097C5FCC4F20953EF3ABBCC3AD7B2" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="350F47EC72B09D8E36B872DEE7CDDE5F" type="description">
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<paragraph id="BB9F33B8FAE9CD7D4851DAA124D2B0C6">
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<figureCitation id="5DFA37586B5678B3A218BFC77FFA3E92" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">Figs 3 F</figureCitation>
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,
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<figureCitation id="5C2D43A7216DB988031F7212488EB22A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)." figureDoi="10.3897/zookeys.1213.133178.figure5" httpUri="https://binary.pensoft.net/fig/1141415">5 A, G</figureCitation>
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,
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<figureCitation id="F384E412C854120141AF57ED70934AAD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)." figureDoi="10.3897/zookeys.1213.133178.figure6" httpUri="https://binary.pensoft.net/fig/1141416">6 C</figureCitation>
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,
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<figureCitation id="E1635E4AFEF66ECE09E53DF99CEC734D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)." figureDoi="10.3897/zookeys.1213.133178.figure7" httpUri="https://binary.pensoft.net/fig/1141417">7 C – G</figureCitation>
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,
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<figureCitation id="43E03B6A442833349F2824FD77C98D45" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9 B, F, G</figureCitation>
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,
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<figureCitation id="CD939199CC1DA4D4DCFFC5BFA87E6978" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)." figureDoi="10.3897/zookeys.1213.133178.figure10" httpUri="https://binary.pensoft.net/fig/1141420">10 A, B</figureCitation>
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,
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<figureCitation id="DC95B135D2ABC5E32D14A6D066ACDF4D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 A, F</figureCitation>
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,
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<figureCitation id="91FD824FDFD38D3D0A712F1A32957D48" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 A – D</figureCitation>
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,
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<figureCitation id="0B05CD424EC796BDBF58CF562AC6672F" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure39" httpUri="https://binary.pensoft.net/fig/1141449">39</figureCitation>
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,
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<figureCitation id="FF54311868178267A6C21811831FB4D1" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure40" httpUri="https://binary.pensoft.net/fig/1141450">40</figureCitation>
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,
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<figureCitation id="484161B6F78AF3D8567AE9836588CD95" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure41" httpUri="https://binary.pensoft.net/fig/1141451">41</figureCitation>
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,
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<figureCitation id="53022C90F450B19D902AABAC9B748F09" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1213.133178.figure42" httpUri="https://binary.pensoft.net/fig/1141452">42</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0ECFAK" type="type material">
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<paragraph id="26389C982B7D3394E74E10CF7BD36BF2">
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<heading id="629241FE995458A96D61B73A366F0F7B" reason="title">Type material.</heading>
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</paragraph>
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<paragraph id="ADBAFCEC170D4D2CF4D1F95EA2BE77C5">
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<materialsCitation id="D3F73AC7B0BC7CCD0E1E34A5BFE68A25" collectingDate="2024-03-19" collectionCode="KSMA" collectorName="B. A. Huber" country="Saudi Arabia" elevation="2250" latitude="19.022" location="Tanomah" longLatPrecision="7" longitude="42.1247" specimenCount="1" specimenCount-male="1" stateProvince="Asir" typeStatus="holotype">
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<emphasis id="D351323B91EB1A0A9BEE5592688DFC5F" bold="true">
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||
<emphasis id="A340569C25741CF1F7FC6FB121E8389A" bold="true" italics="true">
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||
<typeStatus id="51FDDAB0A0BA1061E5FE7DAE6EF297BA" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
.
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</emphasis>
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<collectingCountry id="B6EF32BEEE83C2A9F26E22325014180D" name="Saudi Arabia">Saudi Arabia</collectingCountry>
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||
– ‘
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||
<emphasis id="4211A5906490FB207E86A8DBAD834DC5" bold="true">
|
||
<collectingRegion id="8A37BDF2246BAA0334B9CD122CF50C31" country="Saudi Arabia" name="'Asir">Asir</collectingRegion>
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||
</emphasis>
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||
•
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||
<specimenCount id="1E7F833D17D61BB7DB5E448DC2B493FD" count="1" type="male">♂</specimenCount>
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||
; NW of
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||
<location id="841E25EF81511A70240F0F6EB0E1FA44" LSID="urn:lsid:plazi:treatment:933148E97DF357DC8194F5ED2DFB17B1:841E25EF81511A70240F0F6EB0E1FA44" country="Saudi Arabia" latitude="19.022" longLatPrecision="7" longitude="42.1247" name="Tanomah" stateProvince="Asir">Tanomah</location>
|
||
;
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||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESFAK" specific-use="{"type":"Point","coordinates":[42.124700,19.022000]}">
|
||
<geoCoordinate id="CF113132858B097C73410C81FE431C45" degrees="19.0220" direction="north" orientation="latitude" precision="5" value="19.022">19.0220 ° N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="B9BFE4A9F910EAED2A4405F4827B6B99" degrees="42.1247" direction="east" orientation="longitude" precision="5" value="42.1247">42.1247 ° E</geoCoordinate>
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||
</named-content>
|
||
;
|
||
<elevation id="0AA1621DACFB6A85E2CEE6E7115598C3" metricMagnitude="3" metricUnit="m" metricValue="2.25" unit="m" value="2250.0">
|
||
<quantity id="E94F111CA67007C7DDD379D23EA0110A" metricMagnitude="3" metricUnit="m" metricValue="2.25" unit="m" value="2250.0">2250 m</quantity>
|
||
a. s. l.
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||
</elevation>
|
||
;
|
||
<date id="5298137B58423F91FA61542BE1A03154" value="2024-03-19">
|
||
<collectingDate id="6C348759B5C94A6D277387EA0E0D0107" value="2024-03-19">19 Mar. 2024</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2E8EB5D07E9310C3F6926924062D1DDE">B. A. Huber</collectorName>
|
||
leg.;
|
||
<named-content id="B623BECB3C2A25DBB2EBB97F77DE96E2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/king-saud-university" xlink_title="Museum of Arthropods, College of Food and Agriculture Sciences, King Saud University">
|
||
<collectionCode id="B6EBD55C05AE04B846EA475C279981D7">KSMA</collectionCode>
|
||
</named-content>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="66145F9953A72EC905CBE42753AAB718" collectingDate="2024-03-19" collectionCode="ZFMK" collectorName="B. A. Huber" country="Saudi Arabia" elevation="2250" latitude="19.022" location="Asir" longLatPrecision="7" longitude="42.1247" specimenCode="Ar 24663" specimenCount="48" specimenCount-female="25" specimenCount-male="22" stateProvince="Asir" typeStatus="paratype">
|
||
<emphasis id="B9DD9CECB0247C0D3CF783EF8F03026D" bold="true">
|
||
<emphasis id="9A839AE19D4907873C2E63FD9F87CF35" bold="true" italics="true">
|
||
<typeStatus id="CB5A82B70CAF0A4DB527EE361604EC82" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="B3432A8F858251F1CC0194E5697126C7" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
– ‘
|
||
<emphasis id="668937FDD6DFD828500CFAADEEC1AB47" bold="true">
|
||
<collectingRegion id="BA96EA2C189C216ECC8B32C9DA7BE312" country="Saudi Arabia" name="'Asir">Asir</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="2BE633BA42BBFADB3AC6882AD16F5B0E" count="21" type="male">21 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="F826D3FC54CE5A23F4676F381B768AAC" count="24" type="female">24 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="BFB95E392A57E58B9BB9BDB8B20BDAF4" count="1" type="generic">1 juv.</specimenCount>
|
||
(
|
||
<specimenCount id="9288463DB97EFCAFEF996D4BCC4690EE" count="1" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="30A4417F5FA5DB8A48A2998EAC1B0D20" count="1" type="female">1 ♀</specimenCount>
|
||
used for SEM); same collection data as for holotype;
|
||
<named-content id="436AD9031C5C1593C12CA69F4E1110A6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">
|
||
<collectionCode id="A3C18CC8F615371A4A74240D206E3B19" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
</named-content>
|
||
<specimenCode id="A731E30B228030F0610D46B890C21BEE">Ar 24663</specimenCode>
|
||
to 24664
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C976C0288E3AE10D9E5B984CE22CDDEB">
|
||
<heading id="3913A8EED99C6DE6A6FD8858C067835D" reason="title">Other material.</heading>
|
||
</paragraph>
|
||
<paragraph id="4944BC30891DCC7D495874228C05B312">
|
||
<materialsCitation id="0C18595FA5DC06D31F4FA958EB4AD356" collectingDate="2024-03-19" collectionCode="ZFMK" collectorName="B. A. Huber" country="Saudi Arabia" elevation="2250" latitude="19.022" location="Asir" longLatPrecision="7" longitude="42.1247" specimenCode="SA 100" specimenCount="3" specimenCount-female="3" stateProvince="Asir">
|
||
<collectingCountry id="EF1AA1D3DEBC8FCF4CF4F13A3F211752" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
– ‘
|
||
<emphasis id="6145D7116F7A411D4219AD712BF826F0" bold="true">
|
||
<collectingRegion id="94B6F5B0DF535418F793403DBFB6340E" country="Saudi Arabia" name="'Asir">Asir</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="E6D416627D062AF15E4641F92C7E5AD6" count="3" type="female">3 ♀♀</specimenCount>
|
||
, 4 juvs; in pure ethanol; same collection data as for holotype;
|
||
<named-content id="FEEF3552168DBC42CBD5FD223F7F3A78" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">
|
||
<collectionCode id="3718BC9C6BCAD5F6997CF2AF70783C25" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
</named-content>
|
||
<specimenCode id="92F9F534DF4D6E2A38F0205E87F1014B" collectionCode="SA" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie">SA 100</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ERGAK" type="diagnosis">
|
||
<paragraph id="49E77BF9B30726940CE2298E5A49FD0F">
|
||
<heading id="F1C53B4C77BBD15EAFBC71F780AE46FD" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="CF19A670B0340BC6F40FDE1B0DB65128">
|
||
Distinguished from similar species in the northern Saudi Arabian group (
|
||
<taxonomicName id="5A2FA45812C755AA67C0F53760C8DD18" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bashayer" status="sp. nov.">
|
||
<emphasis id="42526D3BAC005FF3FA5D3C9FEDAB5056" italics="true">M. bashayer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="18C3C8AB3810B864F0CE40C832710641" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="9F253F81D547F8FF3EACC19C232DB58B" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="maysaan" status="sp. nov.">
|
||
<emphasis id="E1672C0C9E5EB1EB2379E62D6341D76E" italics="true">M. maysaan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A8CE85A1DB0F75DBAC2F3E51CCCA1490" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) by very slender main bulbal process (Fig.
|
||
<figureCitation id="7DDB885AABE1EE24B6376DCA3D52A00C" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure40" httpUri="https://binary.pensoft.net/fig/1141450">40 D</figureCitation>
|
||
; wider in other species), and by epigynal ‘ knob’ in posterior rather than central position on epigynal plate (Fig.
|
||
<figureCitation id="C45F06120E33B365A7A6FD97979D4F73" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1213.133178.figure42" httpUri="https://binary.pensoft.net/fig/1141452">42 A, B</figureCitation>
|
||
); from
|
||
<taxonomicName id="74E3E734248412FE992BBAE71803DDDE" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="maysaan" status="sp. nov.">
|
||
<emphasis id="CD2025BA2F0FE3FF1759AAE571EFC107" italics="true">M. maysaan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C69E69226CAC82D3F6B798C09B2C0244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also by less widened hinged process of procursus (Fig.
|
||
<figureCitation id="D089EDBF8CAFA9812D1D97FB5B177530" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure40" httpUri="https://binary.pensoft.net/fig/1141450">40 C</figureCitation>
|
||
); from species of the southern Saudi Arabian group and
|
||
<taxonomicName id="FED57653EE7AD372B5954E7EF6738B8F" authorityName="Deeleman-Reinhold & van Harten" authorityYear="2001" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="jacominae">
|
||
<emphasis id="2B6E3D3FBCD0BF2D2C77F2354BF6C2AE" italics="true">M. jacominae</emphasis>
|
||
</taxonomicName>
|
||
by shorter male palpal trochanter apophysis (Fig.
|
||
<figureCitation id="6FA3E802CA21C221614C8FCDC6E8736E" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure39" httpUri="https://binary.pensoft.net/fig/1141449">39 C</figureCitation>
|
||
), internal female genitalia with membranous central element rather than distinct arc (Fig.
|
||
<figureCitation id="C6987B1BBD2475BCFDCC773969B49BD7" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure41" httpUri="https://binary.pensoft.net/fig/1141451">41 C</figureCitation>
|
||
), and without crescent-shaped structures.
|
||
</paragraph>
|
||
<caption id="FE6566699307B9588B92BF1A24F94457" ID-DOI="10.3897/zookeys.1213.133178.figure39" ID-arpha="0F70A41D-1583-5B66-B405-C395F245F7BD" httpUri="https://binary.pensoft.net/fig/1141449" startId="F39">
|
||
<paragraph id="6BC853DA0157F88D1D941E332E034DA2">
|
||
<label id="6AF6114C83C8BBC12338B07464A86857">Figure 39.</label>
|
||
</paragraph>
|
||
<paragraph id="399B935CA09CCC6A82BCAB52E5D36559">
|
||
<taxonomicName id="52C96D0FC386FC74712CEA05646CEA74" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="80157E8ED70C97BD047860276CA2DDBC" italics="true">Micropholcus tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8B6BA1C7D0677D052A4BDB09AFC90FAA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male from Saudi Arabia, ‘ Asir, NW of Tanomah (
|
||
<named-content id="F7F32BFB933C206DF8F2909568728F93" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
|
||
<specimenCode id="E24FBFC311BFAE06C278B152A89914D1">Ar 24663</specimenCode>
|
||
). Left palp in prolateral (
|
||
<emphasis id="0330DFDFCE40B7020243C55528F2452A" bold="true">A</emphasis>
|
||
), dorsal (
|
||
<emphasis id="B145563A74F0BCA98680C842367925F0" bold="true">B</emphasis>
|
||
), and retrolateral (
|
||
<emphasis id="2173275AFC89CCC57F0C9DDE9AF9A551" bold="true">C</emphasis>
|
||
) views. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="48266A2393C0C82F202ED6B6BCB37499" ID-DOI="10.3897/zookeys.1213.133178.figure40" ID-arpha="40118EDB-10A1-5E1E-AC9F-9A9817CE27A2" httpUri="https://binary.pensoft.net/fig/1141450" startId="F40">
|
||
<paragraph id="977A73DB4A1E567F19F50157AC72900C">
|
||
<label id="1F408DCAC77CDE1A699708ABA4F9A9B8">Figure 40.</label>
|
||
</paragraph>
|
||
<paragraph id="276938D7ECF88E609C833DDBB1AE5D7E">
|
||
<taxonomicName id="EDF96076FCD9D6DD6A447CDF0A091B24" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="C4B601E330E6AA77B2943994EB2A62BA" italics="true">Micropholcus tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4D87C1783EFCA55DC43442C5368663DF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male from Saudi Arabia, ‘ Asir, NW of Tanomah (
|
||
<named-content id="CF7225C895430C07A19AC8BDE8829946" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
|
||
<specimenCode id="C6B68F4D3D39B13BEB1B9E629610EB76">Ar 24663</specimenCode>
|
||
)
|
||
<emphasis id="89D235309CD0D0AF5F2F9B63C6BC31FB" bold="true">A – C</emphasis>
|
||
left procursus in prolateral, dorsal, and retrolateral views
|
||
<emphasis id="7E69327E5AE3A07F5E69B0CB095495AD" bold="true">D, E</emphasis>
|
||
left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="247FB7A16269AD56C49E88C7E82B8EFF" ID-DOI="10.3897/zookeys.1213.133178.figure41" ID-arpha="2C6F8DAB-66ED-5194-B9D0-35C820104C38" httpUri="https://binary.pensoft.net/fig/1141451" startId="F41">
|
||
<paragraph id="EFE54FE1EFD93B278D12076C63ADD4A8">
|
||
<label id="FAF9B996B590DF2183ADEDA841463B6D">Figure 41.</label>
|
||
</paragraph>
|
||
<paragraph id="53174133335E1B988786EF79E51DDB20">
|
||
<taxonomicName id="21F36EBC351D4172ED3E59F6A4F67AC2" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="F08088E89C4A01B6B0078B6D35363B77" italics="true">Micropholcus tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2775D864365A2C0C701DEA98C7B9818B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male from Saudi Arabia, ‘ Asir, NW of Tanomah
|
||
<emphasis id="2A7E4D32FBC2BF7165AC9775D143E6C5" bold="true">A, B</emphasis>
|
||
male chelicerae, frontal and lateral views (
|
||
<named-content id="AA427C77EBE016ACFADC1F854799B2FF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
|
||
<specimenCode id="5CF8C5991F4E999637B883643F24FCE3">Ar 24663</specimenCode>
|
||
)
|
||
<emphasis id="5E9D538591A7F1EC392E09738B4517BC" bold="true">C</emphasis>
|
||
cleared female genitalia, dorsal view (
|
||
<named-content id="6F55301B3450E54CB9C7BACBBD9977AB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
|
||
<specimenCode id="E310AF7A23375F9D1034EE7D2863E78C">Ar 24664</specimenCode>
|
||
); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="579E607C4FC2BD3D2977C731CDC14A06" ID-DOI="10.3897/zookeys.1213.133178.figure42" ID-arpha="75EA0498-443D-5F0B-B489-47E402CAB6F1" httpUri="https://binary.pensoft.net/fig/1141452" startId="F42">
|
||
<paragraph id="2CFDEF311E3BE2E95861D578047FA76F">
|
||
<label id="9C2850B004D86A9D80C907B5ACAC47B4">Figure 42.</label>
|
||
</paragraph>
|
||
<paragraph id="0CCC731E8D0A33A2021A9CD6FACA2547">
|
||
<taxonomicName id="72CE93DAB02F262BABAFDF9CA5AA70F5" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="4AB7EBC83FB6C412B1CD2B5DD210A46B" italics="true">Micropholcus tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6E0FF13B80FC721746AB4ED875528A1B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; female from Saudi Arabia, ‘ Asir, NW of Tanomah (
|
||
<named-content id="53F880611D04CFDD720D825645BD581C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
|
||
<specimenCode id="6CAAA09F37B58F06CB336DDA82556AC9">Ar 24664</specimenCode>
|
||
)
|
||
<emphasis id="C7FB7164DCDE8DAF1353A7414C75D7E7" bold="true">A, B</emphasis>
|
||
abdomen, ventral and lateral views
|
||
<emphasis id="FB388A29828F280620FB6357A13A2315" bold="true">C, D</emphasis>
|
||
cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E4MAK" type="description">
|
||
<paragraph id="DFBBCF2AAE4CCE48294380EFBF25F165">
|
||
<heading id="7CA5F68C2B0D538F0A6D9164BF00543E" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="CD7C5931BF5DC3C4A1C765FA94583436">
|
||
<emphasis id="1D7E5E014F3A46CF120AA48B5E7FD20F" bold="true">Male</emphasis>
|
||
(
|
||
<typeStatus id="C0651881BDB6628CC7E7DDFFE30EB87C" type="holotype">holotype</typeStatus>
|
||
).
|
||
<emphasis id="DCC1731F23A62F153446C4EE9135706C" bold="true">
|
||
<emphasis id="2084795579E4E2910F826D1A5B2721FC" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Total body length 3.4, carapace width 1.2. Distance
|
||
<abbrev id="ABBRID0EJNAK" xlink_title="posterior median eye(s)">PME</abbrev>
|
||
-
|
||
<abbrev id="ABBRID0ENNAK" xlink_title="posterior median eye(s)">PME</abbrev>
|
||
210 µm; diameter
|
||
<abbrev id="ABBRID0ERNAK" xlink_title="posterior median eye(s)">PME</abbrev>
|
||
80 µm; distance
|
||
<abbrev id="ABBRID0EVNAK" xlink_title="posterior median eye(s)">PME</abbrev>
|
||
-
|
||
<abbrev id="ABBRID0EZNAK" xlink_title="anterior lateral eye(s)">ALE</abbrev>
|
||
20 µm; distance
|
||
<abbrev id="ABBRID0E4NAK" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
-
|
||
<abbrev id="ABBRID0EBOAK" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
20 µm; diameter
|
||
<abbrev id="ABBRID0EFOAK" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
45 µm. Leg 1: 28.7 (7.0 + 0.5 + 7.1 + 12.7 + 1.4), tibia 2: 4.5, tibia 3: 2.9, tibia 4: 4.0; tibia 1
|
||
<abbrev id="ABBRID0EJOAK" xlink_title="length/diameter">L / d</abbrev>
|
||
: 71; diameters of leg femora (at half length) 0.11–0.12; of leg tibiae 0.10.
|
||
</paragraph>
|
||
<paragraph id="7EECD656A2CD77C8DA14AF2C09D5CB7F">
|
||
<emphasis id="852F15E30B4550A1AF7FFB03ED851A37" bold="true" italics="true">Colour</emphasis>
|
||
(in ethanol). Carapace pale ochre-yellow with distinct brown mark, ocular area not darkened, clypeus with very indistinct darker pattern; sternum monochromous whitish; legs ochre-yellow to light brown, patella dark brown, tibia-metatarsus joints with small brown ring, femur 1 ventrally proximally brown (less distinct also femur 2); abdomen pale ochre-grey, with indistinct darker internal marks.
|
||
</paragraph>
|
||
<paragraph id="696616EB15BFFF59DCF94A92CD206F71">
|
||
<emphasis id="46FB2AD4EE816ECAD3578904F83DAD0C" bold="true">
|
||
<emphasis id="8AA767E14F091FE5C9AB24D76F949C57" bold="true" italics="true">Body</emphasis>
|
||
.
|
||
</emphasis>
|
||
Habitus as in Fig.
|
||
<specimenCount id="A47B127A3460047309F9895C5BD00F99" count="3" type="generic">
|
||
<figureCitation id="F9C420BE21853BBAAB61E86E70760365" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">3 F</figureCitation>
|
||
.
|
||
</specimenCount>
|
||
Ocular area slightly raised (distinct in frontal view). Carapace without thoracic groove. Clypeus unmodified. Sternum wider than long (0.74 / 0.58), unmodified. Abdomen oval, approximately twice as long as wide. Gonopore with four epiandrous spigots (Fig.
|
||
<figureCitation id="662ECD30C9D2145B6C905A4AC2F0CF4B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)." figureDoi="10.3897/zookeys.1213.133178.figure10" httpUri="https://binary.pensoft.net/fig/1141420">10 A</figureCitation>
|
||
). Spinnerets as in Fig.
|
||
<figureCitation id="AC7BE10617A7A854A69E32C4EF917217" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9 F, G</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E4D0D00B1F0017D6F6C3668831C26768">
|
||
<emphasis id="0A32C8ECDDB1406D561E177E73A77C06" bold="true">
|
||
<emphasis id="88225C3C1B9942BAF1C86B07371EF9FE" bold="true" italics="true">Chelicerae</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in Fig.
|
||
<figureCitation id="509FF22D5AFB708CA9A8CC5245DFD24B" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure41" httpUri="https://binary.pensoft.net/fig/1141451">41 A, B</figureCitation>
|
||
; with pair of distal apophyses near laminae, each with two cone-shaped hairs (Fig.
|
||
<figureCitation id="7EBB358A998EF1D036EAB8329C26B754" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)." figureDoi="10.3897/zookeys.1213.133178.figure6" httpUri="https://binary.pensoft.net/fig/1141416">6 C</figureCitation>
|
||
); with pair of distinct proximal frontal apophyses; with pair of very low and indistinct lateral humps.
|
||
</paragraph>
|
||
<paragraph id="1879DB5852D34B71A75E3FCF0563C54D">
|
||
<emphasis id="4947968DB89140811430AAEE678EC8AA" bold="true">
|
||
<emphasis id="325B10F3EFC6F9E34BD9BE9D2AC9C033" bold="true" italics="true">Palps</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in Fig.
|
||
<figureCitation id="9A26A52CBB6F056C9496F7C1310A1477" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure39" httpUri="https://binary.pensoft.net/fig/1141449">39</figureCitation>
|
||
; coxa unmodified; trochanter with long ventral apophysis with distinct proximal retrolateral hump and modified hair on distal tip (Fig.
|
||
<figureCitation id="5E514963827432378B9E1750EBD613C5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9 B</figureCitation>
|
||
); femur distally widened, with subdistal ventral hump; femur-patella joints shifted toward prolateral side; tibia-tarsus joints shifted toward retrolateral side; tarsus with large tarsal organ. Procursus (Fig.
|
||
<figureCitation id="BDC4CB9F977C3D08D802352C4F717132" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure40" httpUri="https://binary.pensoft.net/fig/1141450">40 A – C</figureCitation>
|
||
) proximally with sclerotised prolateral hump; at half length with prolateral sclerotised ridge transforming distally into transparent membrane, and brush of dorsal hairs; distally with small retrolateral ridge, large bifid membranous ventral-prolateral flap (Fig.
|
||
<figureCitation id="81131BFF724A858C018876A62498217F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)." figureDoi="10.3897/zookeys.1213.133178.figure7" httpUri="https://binary.pensoft.net/fig/1141417">7 C, D</figureCitation>
|
||
), and dorsal hinged process. Genital bulb (Fig 7 F, G, 40 D, E) with strong proximal sclerite; with two sclerotised processes of unclear homology: prolateral process long and slender, with small, pointed branch on retrolateral side; retrolateral process simple, originating from basis of embolus and directed parallel to prolateral process; and mostly semi-transparent short embolus with membranous extensions.
|
||
</paragraph>
|
||
<paragraph id="6F4F1898F4E511D5D918967956AFBE56">
|
||
<emphasis id="A4B49DF7EEAAA58040C9849A11BF352E" bold="true">
|
||
<emphasis id="007F342AACC9610CC67D9086AD0FF7D4" bold="true" italics="true">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Without spines, without curved hairs, without sexually dimorphic short vertical hairs; retrolateral trichobothrium of tibia 1 at 6 %; prolateral trichobothrium absent on tibia 1; tarsus 1 with> 20 pseudosegments, distally distinct.
|
||
</paragraph>
|
||
<paragraph id="AC36B6C8FE6674BE9C684D196BF580E5">
|
||
<emphasis id="D322FCE6B8BE80F01BEBCAE9A8A9491A" bold="true" italics="true">Variation</emphasis>
|
||
(male). Tibia
|
||
<quantity id="7316EC4F9261145ADFE3A6CCBD4CDB8B" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
|
||
21 other males: 5.1–7.3 (mean 6.5). Carapace pattern very consistent. Abdomen usually with large white marks dorsally and laterally.
|
||
</paragraph>
|
||
<paragraph id="BCB95537F26C571633DBBAFB324824DB">
|
||
<emphasis id="EEF7D3DB9BC8F709138CB90A3C1D7D28" bold="true">Female.</emphasis>
|
||
In general very similar to male but anterior leg femora proximally not darkened; ocular area with large median and small lateral brown marks. Tibia
|
||
<quantity id="FDCD6E90EFFAE36107B1E6ABAFEE0411" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
|
||
<specimenCount id="7253639CFF1EC1D106BC99DD12622CDB" count="24" type="female">24 females</specimenCount>
|
||
: 4.4–5.9 (mean 5.2). Epigynum (Figs
|
||
<figureCitation id="5813F96FC5290E853C104AB667E46DA0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)." figureDoi="10.3897/zookeys.1213.133178.figure10" httpUri="https://binary.pensoft.net/fig/1141420">10 B</figureCitation>
|
||
,
|
||
<figureCitation id="970850BB84392EB97E416D4E94CFD894" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1213.133178.figure42" httpUri="https://binary.pensoft.net/fig/1141452">42 A, B</figureCitation>
|
||
) protruding, anterior plate oval, mostly dark brown except medially posteriorly, with small knob-shaped process (Fig.
|
||
<figureCitation id="12C80518A22140ECAD80D261361DB374" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 A</figureCitation>
|
||
) near posterior margin; posterior epigynal plate very short and indistinct, light brown. Internal genitalia (Figs
|
||
<figureCitation id="9D07E355239312A971C87F72795C9956" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure41" httpUri="https://binary.pensoft.net/fig/1141451">41 C</figureCitation>
|
||
,
|
||
<figureCitation id="2F743812EB666CF625CE5D1E28682BF6" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1213.133178.figure42" httpUri="https://binary.pensoft.net/fig/1141452">42 C, D</figureCitation>
|
||
) with pair of elongated pore plates in transversal position; with pair of lateral sclerites and complex system of membranous structures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ETRAK" type="etymology">
|
||
<paragraph id="E077352A5FA32D30AA1C1FE44D4C9374">
|
||
<heading id="AF36AEAB69E9670264DF1A7451928677" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="9F85D5B01EA5FADBAC3144058BBA8820">The species name is derived from the type locality; noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EYRAK" type="distribution">
|
||
<paragraph id="E03A33A31918FE488F86E65995E56F90">
|
||
<heading id="58F11827108E850B09906101939D696B" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="CD4B0F554E4A4BC552108E4264A81163">
|
||
Known from type locality only, in
|
||
<collectingCountry id="76D53E47D87C9E48C44D51162A91225E" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
, ‘
|
||
<collectingRegion id="10ECA8A0F4644C3834303AF905A019D9" country="Saudi Arabia" name="'Asir">Asir Province</collectingRegion>
|
||
(Fig.
|
||
<figureCitation id="B3C00F21E242789DC4316FFAC4925A2D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Known distribution of Micropholcus fauroti (Simon) (A) and of Old World Micropholcus in Mindanao (B), on the Arabian Peninsula (C) and in Morocco (D)." figureDoi="10.3897/zookeys.1213.133178.figure13" httpUri="https://binary.pensoft.net/fig/1141423">13 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ECSAK" type="natural history">
|
||
<paragraph id="8976AAEBF85ACC3F622EA66D39D4568E">
|
||
<heading id="625E12BE66ED70147B24258558D64AA2" reason="title">Natural history.</heading>
|
||
</paragraph>
|
||
<paragraph id="B2903A7A178C6A2787206E6E40925672">
|
||
The spiders were found in caverns among and under boulders, often together with a representative of
|
||
<taxonomicName id="8ED0A481A831AC392BDC0070BC951032" authority="Simon, 1890" authorityName="Simon" authorityYear="1890" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3FCC5B2994F33E77A00017ECA2E2890C" italics="true">Smeringopus</emphasis>
|
||
Simon, 1890
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="DA85B95B5B148DED44B7C42CA7254424" authorityName="Clerck" authorityYear="1757" class="Arachnida" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
|
||
:
|
||
<taxonomicName id="CBA5F7EBA716EFBFAB72B1CDEDE4298A" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
|
||
). Both species sometimes occurred in very high densities. In one case, a ceiling of a cave was estimated to measure ~
|
||
<quantity id="1C1A845778BABFBE5DAAF8F41F64C618" metricMagnitude="0" metricUnit="m" metricValue="3.0" unit="m" value="3.0">3 m</quantity>
|
||
<superScript id="C3CEB1E818E8B0C1C29F15CA4294315C">2</superScript>
|
||
and to contain ~ 250 large (adult and penultimate instar) specimens (i. e., with average distances between specimens of ~
|
||
<quantity id="2E079985336AF5A4221F2E80FDCE5DAE" metricMagnitude="-1" metricUnit="m" metricValue="1.0" unit="cm" value="10.0">10 cm</quantity>
|
||
) (Fig.
|
||
<figureCitation id="90A12EE31F2685ECE703CC3F6406E796" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Section of cave ceiling (16 × 11 cm), showing nine adult specimens of M. tanomah Huber, sp. nov.; from NW of Tanomah, ‘ Asir, Saudi Arabia." figureDoi="10.3897/zookeys.1213.133178.figure15" httpUri="https://binary.pensoft.net/fig/1141425">15</figureCitation>
|
||
).
|
||
<specimenCount id="44BF0766BC2956C93CA2610830770602" count="1" type="egg">One egg</specimenCount>
|
||
sac had a diameter of
|
||
<quantity id="C885D0E5F02E0EF5470587B6B3303C25" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, and contained ~
|
||
<specimenCount id="AC1E7632796BF98BA14AA66B4CB102B9" count="35" type="egg">35 eggs</specimenCount>
|
||
. Egg diameters ranged from
|
||
<quantity id="D2BB239B0A830028DEB1C624AB807D61" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.1" metricValueMin="6.8999999999999995" unit="mm" value="0.7" valueMax="0.71" valueMin="0.69">0.69 to 0.71 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |