<documentid="32A1A24CE81CB323027A5D1D8360F08B"ID-DOI="10.3897/zookeys.1213.133178"ID-ZooBank="B8E6EE1A-023D-4B64-8D3A-C1843F0D8376"ID-publisher-id="133178"URI-arpha="B0B2A119-6CED-5B4F-810D-44C4C6D0F4F3"URI-zoobank="http://zoobank.org/B8E6EE1A-023D-4B64-8D3A-C1843F0D8376"XM.bibliography_approvedBy="admin"XM.materialsCitations_approvedBy="admin"XM.taxonomicNames_approvedBy="admin"XM.treatmentCitations_approvedBy="admin"XM.treatments_approvedBy="admin"article-type="research-article"checkinTime="1727330124077"checkinUser="pensoft"docAuthor="Huber, Bernhard A. & Meng, Guanliang"docDate="2024"docId="933148E97DF357DC8194F5ED2DFB17B1"docLanguage="en"docName="ZooKeys 1213: 95-182"docOrigin="ZooKeys 1213"docSource="https://zookeys.pensoft.net/article/133178/download/xml/"docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic"docStyleName="PensoftTaxPub.0000.journal_article.generic"docTitle="Micropholcus tanomah Huber & Meng 2024, sp. nov."docType="treatment"docUuid="6D679588-8300-427F-89A7-6CA93102C27C"docUuidSource="ZooBank"docVersion="3"dtd-version="3.0"lastPageNumber="182"masterDocId="B0B2A1196CED5B4F810D44C4C6D0F4F3"masterDocTitle="Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae)"masterLastPageNumber="182"masterPageNumber="95"pageNumber="95"updateTime="1727371993187"updateUser="admin">
<mods:titleid="B6A5DF1B8EAF9ECEE7255ACF3CD285A8">Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae)</mods:title>
<taxonomicNameid="1BAD335CDC03FEA1597C7668DA145AA8"authority="Huber"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="tanomah"status="sp. nov.">
<figureCitationid="5DFA37586B5678B3A218BFC77FFA3E92"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH."figureDoi="10.3897/zookeys.1213.133178.figure3"httpUri="https://binary.pensoft.net/fig/1141413">Figs 3 F</figureCitation>
,
<figureCitationid="5C2D43A7216DB988031F7212488EB22A"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)."figureDoi="10.3897/zookeys.1213.133178.figure5"httpUri="https://binary.pensoft.net/fig/1141415">5 A, G</figureCitation>
,
<figureCitationid="F384E412C854120141AF57ED70934AAD"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)."figureDoi="10.3897/zookeys.1213.133178.figure6"httpUri="https://binary.pensoft.net/fig/1141416">6 C</figureCitation>
,
<figureCitationid="E1635E4AFEF66ECE09E53DF99CEC734D"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)."figureDoi="10.3897/zookeys.1213.133178.figure7"httpUri="https://binary.pensoft.net/fig/1141417">7 C – G</figureCitation>
,
<figureCitationid="43E03B6A442833349F2824FD77C98D45"captionStart="Figure 9"captionStartId="F9"captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)."figureDoi="10.3897/zookeys.1213.133178.figure9"httpUri="https://binary.pensoft.net/fig/1141419">9 B, F, G</figureCitation>
,
<figureCitationid="CD939199CC1DA4D4DCFFC5BFA87E6978"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)."figureDoi="10.3897/zookeys.1213.133178.figure10"httpUri="https://binary.pensoft.net/fig/1141420">10 A, B</figureCitation>
,
<figureCitationid="DC95B135D2ABC5E32D14A6D066ACDF4D"captionStart="Figure 11"captionStartId="F11"captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)."figureDoi="10.3897/zookeys.1213.133178.figure11"httpUri="https://binary.pensoft.net/fig/1141421">11 A, F</figureCitation>
,
<figureCitationid="91FD824FDFD38D3D0A712F1A32957D48"captionStart="Figure 12"captionStartId="F12"captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)."figureDoi="10.3897/zookeys.1213.133178.figure12"httpUri="https://binary.pensoft.net/fig/1141422">12 A – D</figureCitation>
,
<figureCitationid="0B05CD424EC796BDBF58CF562AC6672F"captionStart="Figure 39"captionStartId="F39"captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure39"httpUri="https://binary.pensoft.net/fig/1141449">39</figureCitation>
,
<figureCitationid="FF54311868178267A6C21811831FB4D1"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure40"httpUri="https://binary.pensoft.net/fig/1141450">40</figureCitation>
,
<figureCitationid="484161B6F78AF3D8567AE9836588CD95"captionStart="Figure 41"captionStartId="F41"captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure41"httpUri="https://binary.pensoft.net/fig/1141451">41</figureCitation>
,
<figureCitationid="53022C90F450B19D902AABAC9B748F09"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1213.133178.figure42"httpUri="https://binary.pensoft.net/fig/1141452">42</figureCitation>
<materialsCitationid="D3F73AC7B0BC7CCD0E1E34A5BFE68A25"collectingDate="2024-03-19"collectionCode="KSMA"collectorName="B. A. Huber"country="Saudi Arabia"elevation="2250"latitude="19.022"location="Tanomah"longLatPrecision="7"longitude="42.1247"specimenCount="1"specimenCount-male="1"stateProvince="Asir"typeStatus="holotype">
<collectingDateid="6C348759B5C94A6D277387EA0E0D0107"value="2024-03-19">19 Mar. 2024</collectingDate>
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;
<collectorNameid="2E8EB5D07E9310C3F6926924062D1DDE">B. A. Huber</collectorName>
leg.;
<named-contentid="B623BECB3C2A25DBB2EBB97F77DE96E2"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/king-saud-university"xlink_title="Museum of Arthropods, College of Food and Agriculture Sciences, King Saud University">
<materialsCitationid="66145F9953A72EC905CBE42753AAB718"collectingDate="2024-03-19"collectionCode="ZFMK"collectorName="B. A. Huber"country="Saudi Arabia"elevation="2250"latitude="19.022"location="Asir"longLatPrecision="7"longitude="42.1247"specimenCode="Ar 24663"specimenCount="48"specimenCount-female="25"specimenCount-male="22"stateProvince="Asir"typeStatus="paratype">
used for SEM); same collection data as for holotype;
<named-contentid="436AD9031C5C1593C12CA69F4E1110A6"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig"xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">
<collectionCodeid="A3C18CC8F615371A4A74240D206E3B19"country="Germany"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613"lsid="urn:lsid:biocol.org:col:34613"name="Zoologisches Forschungsmuseum Alexander Koenig"type="Museum">ZFMK</collectionCode>
<materialsCitationid="0C18595FA5DC06D31F4FA958EB4AD356"collectingDate="2024-03-19"collectionCode="ZFMK"collectorName="B. A. Huber"country="Saudi Arabia"elevation="2250"latitude="19.022"location="Asir"longLatPrecision="7"longitude="42.1247"specimenCode="SA 100"specimenCount="3"specimenCount-female="3"stateProvince="Asir">
, 4 juvs; in pure ethanol; same collection data as for holotype;
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<collectionCodeid="3718BC9C6BCAD5F6997CF2AF70783C25"country="Germany"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613"lsid="urn:lsid:biocol.org:col:34613"name="Zoologisches Forschungsmuseum Alexander Koenig"type="Museum">ZFMK</collectionCode>
</named-content>
<specimenCodeid="92F9F534DF4D6E2A38F0205E87F1014B"collectionCode="SA"country="France"name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie">SA 100</specimenCode>
Distinguished from similar species in the northern Saudi Arabian group (
<taxonomicNameid="5A2FA45812C755AA67C0F53760C8DD18"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="bashayer"status="sp. nov.">
<taxonomicNameid="9F253F81D547F8FF3EACC19C232DB58B"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="maysaan"status="sp. nov.">
<figureCitationid="7DDB885AABE1EE24B6376DCA3D52A00C"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure40"httpUri="https://binary.pensoft.net/fig/1141450">40 D</figureCitation>
; wider in other species), and by epigynal ‘ knob’ in posterior rather than central position on epigynal plate (Fig.
<figureCitationid="C45F06120E33B365A7A6FD97979D4F73"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1213.133178.figure42"httpUri="https://binary.pensoft.net/fig/1141452">42 A, B</figureCitation>
); from
<taxonomicNameid="74E3E734248412FE992BBAE71803DDDE"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="maysaan"status="sp. nov.">
also by less widened hinged process of procursus (Fig.
<figureCitationid="D089EDBF8CAFA9812D1D97FB5B177530"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure40"httpUri="https://binary.pensoft.net/fig/1141450">40 C</figureCitation>
); from species of the southern Saudi Arabian group and
<taxonomicNameid="FED57653EE7AD372B5954E7EF6738B8F"authorityName="Deeleman-Reinhold & van Harten"authorityYear="2001"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="jacominae">
<figureCitationid="6FA3E802CA21C221614C8FCDC6E8736E"captionStart="Figure 39"captionStartId="F39"captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure39"httpUri="https://binary.pensoft.net/fig/1141449">39 C</figureCitation>
), internal female genitalia with membranous central element rather than distinct arc (Fig.
<figureCitationid="C6987B1BBD2475BCFDCC773969B49BD7"captionStart="Figure 41"captionStartId="F41"captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure41"httpUri="https://binary.pensoft.net/fig/1141451">41 C</figureCitation>
<taxonomicNameid="52C96D0FC386FC74712CEA05646CEA74"authority="Huber"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="tanomah"status="sp. nov.">
<named-contentid="F7F32BFB933C206DF8F2909568728F93"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig"xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
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<named-contentid="CF7225C895430C07A19AC8BDE8829946"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig"xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
<taxonomicNameid="21F36EBC351D4172ED3E59F6A4F67AC2"authority="Huber"authorityName="Huber & Meng"authorityYear="2024"class="Arachnida"family="Pholcidae"genus="Micropholcus"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="species"species="tanomah"status="sp. nov.">
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<named-contentid="6F55301B3450E54CB9C7BACBBD9977AB"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig"xlink_title="Zoologisches Forschungsmuseum Alexander Koenig">ZFMK</named-content>
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; female from Saudi Arabia, ‘ Asir, NW of Tanomah (
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(in ethanol). Carapace pale ochre-yellow with distinct brown mark, ocular area not darkened, clypeus with very indistinct darker pattern; sternum monochromous whitish; legs ochre-yellow to light brown, patella dark brown, tibia-metatarsus joints with small brown ring, femur 1 ventrally proximally brown (less distinct also femur 2); abdomen pale ochre-grey, with indistinct darker internal marks.
<figureCitationid="F9C420BE21853BBAAB61E86E70760365"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH."figureDoi="10.3897/zookeys.1213.133178.figure3"httpUri="https://binary.pensoft.net/fig/1141413">3 F</figureCitation>
.
</specimenCount>
Ocular area slightly raised (distinct in frontal view). Carapace without thoracic groove. Clypeus unmodified. Sternum wider than long (0.74 / 0.58), unmodified. Abdomen oval, approximately twice as long as wide. Gonopore with four epiandrous spigots (Fig.
<figureCitationid="662ECD30C9D2145B6C905A4AC2F0CF4B"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)."figureDoi="10.3897/zookeys.1213.133178.figure10"httpUri="https://binary.pensoft.net/fig/1141420">10 A</figureCitation>
). Spinnerets as in Fig.
<figureCitationid="AC7BE10617A7A854A69E32C4EF917217"captionStart="Figure 9"captionStartId="F9"captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)."figureDoi="10.3897/zookeys.1213.133178.figure9"httpUri="https://binary.pensoft.net/fig/1141419">9 F, G</figureCitation>
<figureCitationid="509FF22D5AFB708CA9A8CC5245DFD24B"captionStart="Figure 41"captionStartId="F41"captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure41"httpUri="https://binary.pensoft.net/fig/1141451">41 A, B</figureCitation>
; with pair of distal apophyses near laminae, each with two cone-shaped hairs (Fig.
<figureCitationid="7EBB358A998EF1D036EAB8329C26B754"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)."figureDoi="10.3897/zookeys.1213.133178.figure6"httpUri="https://binary.pensoft.net/fig/1141416">6 C</figureCitation>
); with pair of distinct proximal frontal apophyses; with pair of very low and indistinct lateral humps.
<figureCitationid="9A26A52CBB6F056C9496F7C1310A1477"captionStart="Figure 39"captionStartId="F39"captionText="Figure 39. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure39"httpUri="https://binary.pensoft.net/fig/1141449">39</figureCitation>
; coxa unmodified; trochanter with long ventral apophysis with distinct proximal retrolateral hump and modified hair on distal tip (Fig.
<figureCitationid="5E514963827432378B9E1750EBD613C5"captionStart="Figure 9"captionStartId="F9"captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)."figureDoi="10.3897/zookeys.1213.133178.figure9"httpUri="https://binary.pensoft.net/fig/1141419">9 B</figureCitation>
); femur distally widened, with subdistal ventral hump; femur-patella joints shifted toward prolateral side; tibia-tarsus joints shifted toward retrolateral side; tarsus with large tarsal organ. Procursus (Fig.
<figureCitationid="BDC4CB9F977C3D08D802352C4F717132"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24663) A – C left procursus in prolateral, dorsal, and retrolateral views D, E left genital bulb in prolateral and retrolateral views. Abbreviation: hp, dorsal hinged process. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure40"httpUri="https://binary.pensoft.net/fig/1141450">40 A – C</figureCitation>
) proximally with sclerotised prolateral hump; at half length with prolateral sclerotised ridge transforming distally into transparent membrane, and brush of dorsal hairs; distally with small retrolateral ridge, large bifid membranous ventral-prolateral flap (Fig.
<figureCitationid="81131BFF724A858C018876A62498217F"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)."figureDoi="10.3897/zookeys.1213.133178.figure7"httpUri="https://binary.pensoft.net/fig/1141417">7 C, D</figureCitation>
), and dorsal hinged process. Genital bulb (Fig 7 F, G, 40 D, E) with strong proximal sclerite; with two sclerotised processes of unclear homology: prolateral process long and slender, with small, pointed branch on retrolateral side; retrolateral process simple, originating from basis of embolus and directed parallel to prolateral process; and mostly semi-transparent short embolus with membranous extensions.
Without spines, without curved hairs, without sexually dimorphic short vertical hairs; retrolateral trichobothrium of tibia 1 at 6 %; prolateral trichobothrium absent on tibia 1; tarsus 1 with> 20 pseudosegments, distally distinct.
<figureCitationid="5813F96FC5290E853C104AB667E46DA0"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)."figureDoi="10.3897/zookeys.1213.133178.figure10"httpUri="https://binary.pensoft.net/fig/1141420">10 B</figureCitation>
,
<figureCitationid="970850BB84392EB97E416D4E94CFD894"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1213.133178.figure42"httpUri="https://binary.pensoft.net/fig/1141452">42 A, B</figureCitation>
) protruding, anterior plate oval, mostly dark brown except medially posteriorly, with small knob-shaped process (Fig.
<figureCitationid="12C80518A22140ECAD80D261361DB374"captionStart="Figure 11"captionStartId="F11"captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)."figureDoi="10.3897/zookeys.1213.133178.figure11"httpUri="https://binary.pensoft.net/fig/1141421">11 A</figureCitation>
) near posterior margin; posterior epigynal plate very short and indistinct, light brown. Internal genitalia (Figs
<figureCitationid="9D07E355239312A971C87F72795C9956"captionStart="Figure 41"captionStartId="F41"captionText="Figure 41. Micropholcus tanomah Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Tanomah A, B male chelicerae, frontal and lateral views (ZFMK Ar 24663) C cleared female genitalia, dorsal view (ZFMK Ar 24664); bold arrow points at membranous central element. Abbreviations: k, epigynal ‘ knob’; pp, pore plate. Scale bars: 0.3 mm."figureDoi="10.3897/zookeys.1213.133178.figure41"httpUri="https://binary.pensoft.net/fig/1141451">41 C</figureCitation>
,
<figureCitationid="2F743812EB666CF625CE5D1E28682BF6"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Micropholcus tanomah Huber, sp. nov.; female from Saudi Arabia, ‘ Asir, NW of Tanomah (ZFMK Ar 24664) A, B abdomen, ventral and lateral views C, D cleared genitalia, ventral and dorsal views. Scale bars: 0.5 mm."figureDoi="10.3897/zookeys.1213.133178.figure42"httpUri="https://binary.pensoft.net/fig/1141452">42 C, D</figureCitation>
) with pair of elongated pore plates in transversal position; with pair of lateral sclerites and complex system of membranous structures.
<figureCitationid="B3C00F21E242789DC4316FFAC4925A2D"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Known distribution of Micropholcus fauroti (Simon) (A) and of Old World Micropholcus in Mindanao (B), on the Arabian Peninsula (C) and in Morocco (D)."figureDoi="10.3897/zookeys.1213.133178.figure13"httpUri="https://binary.pensoft.net/fig/1141423">13 C</figureCitation>
<taxonomicNameid="CBA5F7EBA716EFBFAB72B1CDEDE4298A"authorityName="C. L. Koch"authorityYear="1850"class="Arachnida"family="Pholcidae"kingdom="Animalia"order="Araneae"phylum="Arthropoda"rank="family">Pholcidae</taxonomicName>
). Both species sometimes occurred in very high densities. In one case, a ceiling of a cave was estimated to measure ~
<figureCitationid="90A12EE31F2685ECE703CC3F6406E796"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Section of cave ceiling (16 × 11 cm), showing nine adult specimens of M. tanomah Huber, sp. nov.; from NW of Tanomah, ‘ Asir, Saudi Arabia."figureDoi="10.3897/zookeys.1213.133178.figure15"httpUri="https://binary.pensoft.net/fig/1141425">15</figureCitation>
<quantityid="D2BB239B0A830028DEB1C624AB807D61"metricMagnitude="-4"metricUnit="m"metricValue="7.0"metricValueMax="7.1"metricValueMin="6.8999999999999995"unit="mm"value="0.7"valueMax="0.71"valueMin="0.69">0.69 to 0.71 mm</quantity>
