518 lines
66 KiB
XML
518 lines
66 KiB
XML
<document id="48A8837942769D9ED65B0948C0CA5BEF" ID-DOI="10.11646/zootaxa.4044.3.4" ID-GBIF-Dataset="e55018d0-72ce-4085-94e2-5a61edd08aaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="232597" ID-ZooBank="DD54E580-B6E7-441E-943C-BDDA2EB193C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461339996114" checkinUser="plazi" docAuthor="Gilgado, José D., Enghoff, Henrik, Tinaut, Alberto, Mauriès, Jean-Paul & Ortuño, Vicente M." docDate="2015" docId="D35887F5FFD7FFB6608E4495FD64FC6E" docLanguage="en" docName="zt04044p410.pdf" docOrigin="Zootaxa 4044 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratosphys cryodeserti Gilgado, Mauries and Enghoff, n. sp." docType="treatment" docVersion="9" lastPageNumber="403" masterDocId="2F61FF8DFFD0FFBA60194064FFBFFFEE" masterDocTitle="Sierra Nevada (Granada, Spain): a high-altitude biogeographical crossroads for millipedes (Diplopoda), with first data on its MSS fauna and description of a new species of the genus Ceratosphys Ribaut, 1920 (Chordeumatida: Opisthocheiridae)" masterLastPageNumber="410" masterPageNumber="391" pageNumber="398" updateTime="1698619943649" updateUser="plazi">
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<mods:title id="44CDD1B1E21E9FB20402CE154F17D0D3">Sierra Nevada (Granada, Spain): a high-altitude biogeographical crossroads for millipedes (Diplopoda), with first data on its MSS fauna and description of a new species of the genus Ceratosphys Ribaut, 1920 (Chordeumatida: Opisthocheiridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E13D17E7CE7E1A437ACC6C3A33EB2AE0" type="personal">
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<mods:roleTerm id="6825A6990BF490D61D227027F2274F3E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6A5C2969C50DDF5CFBE3882967F6EE1">Gilgado, José D.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="3FEB5D6C0FD429BF0E83AEDF47654C84">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3465CB0552E1B19AB5DE7FCC1CAA9C82">Enghoff, Henrik</mods:namePart>
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</mods:name>
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<mods:name id="3BC7AF3F8D35F9FFFB0DD221097007C6" type="personal">
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<mods:roleTerm id="9AF3C977E269F08683E259852A8B7C78">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="34D2DF34C6735256AFFDABDDBD18D771">Tinaut, Alberto</mods:namePart>
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</mods:name>
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<mods:name id="B809E8FD8E5C291AD3671998A97BA598" type="personal">
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<mods:namePart id="A09F286F265D0156B026C4C937FBD17E">Mauriès, Jean-Paul</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="9653EBB8CFA3F5ADB9624192A6C77220">Ortuño, Vicente M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7B664ECB89F7BCDC7775ACE2B66E3311">text</mods:typeOfResource>
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<mods:title id="EB35696510F469C80542FFE545D8090C">Zootaxa</mods:title>
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<mods:date id="9DC1C83B41771E3E8591192E4DBA1B15">2015</mods:date>
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<mods:detail id="640ED5010207214D491DF02ABB8897F1" type="volume">
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<mods:number id="293435C973A149E3B66744E2130331C0">4044</mods:number>
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<mods:classification id="63B37E093CA4F52780CE7B7B310B16A3">journal article</mods:classification>
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<mods:identifier id="22E9010DD5E74826B6C252C2B1C6A7B7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C870FBB497D422F0944BFB55334B06A8" type="ZooBank">DD54E580-B6E7-441E-943C-BDDA2EB193C3</mods:identifier>
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<treatment id="D35887F5FFD7FFB6608E4495FD64FC6E" ID-DOI="http://doi.org/10.5281/zenodo.5618239" ID-GBIF-Taxon="119645072" ID-Zenodo-Dep="5618239" LSID="urn:lsid:plazi:treatment:D35887F5FFD7FFB6608E4495FD64FC6E" httpUri="http://treatment.plazi.org/id/D35887F5FFD7FFB6608E4495FD64FC6E" lastPageId="12" lastPageNumber="403" pageId="7" pageNumber="398">
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<subSubSection id="13EB6568FFD7FFBD608E4495FEBFFAC3" pageId="7" pageNumber="398" type="nomenclature">
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<paragraph id="5B4E36E3FFD7FFBD608E4495FC3BFAE5" blockId="7.[151,900,1265,1325]" box="[151,900,1265,1291]" pageId="7" pageNumber="398">
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<heading id="0006818FFFD7FFBD608E4495FC3BFAE5" bold="true" box="[151,900,1265,1291]" fontSize="11" level="1" pageId="7" pageNumber="398" reason="1">
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<emphasis id="6985EAF1FFD7FFBD608E4495FC3BFAE5" bold="true" box="[151,900,1265,1291]" pageId="7" pageNumber="398">
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||
<taxonomicName id="9CF14D60FFD7FFBD608E4495FC84FAE5" ID-CoL="8479L" authority="Gilgado, Mauries and Enghoff" authorityName="Gilgado, Mauries and Enghoff" box="[151,827,1265,1291]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="398" phylum="Arthropoda" rank="species" species="cryodeserti" status="sp. nov.">
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||
<emphasis id="6985EAF1FFD7FFBD608E4495FE09FAE5" bold="true" box="[151,438,1265,1291]" italics="true" pageId="7" pageNumber="398">Ceratosphys cryodeserti</emphasis>
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||
Gilgado, Mauriès and Enghoff
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||
</taxonomicName>
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||
<taxonomicNameLabel id="72B6578AFFD7FFBD635B4495FC3BFAE5" box="[834,900,1265,1291]" pageId="7" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="5B4E36E3FFD7FFBD608E4570FEBFFAC3" blockId="7.[151,900,1265,1325]" box="[151,256,1300,1325]" pageId="7" pageNumber="398">
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<figureCitation id="C3CA2A66FFD7FFBD608E4570FEBFFAC3" box="[151,256,1300,1325]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="8.[151,250,1504,1526]" captionStartId-1="9.[151,250,1906,1928]" captionStartId-2="10.[151,250,1302,1324]" captionStartId-3="11.[151,250,1627,1649]" captionTargetBox-0="[159,1427,309,1472]" captionTargetBox-1="[157,1428,453,1872]" captionTargetBox-2="[162,1427,190,1265]" captionTargetBox-3="[159,1430,452,1593]" captionTargetId-0="figure@8.[151,1436,301,1483]" captionTargetId-1="figure@9.[151,1436,445,1884]" captionTargetId-2="figure@10.[151,1436,179,1281]" captionTargetId-3="figure@11.[151,1436,445,1606]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." captionText-1="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." captionText-2="FIGURE 5 Anatomical details of Ceratosphys Cryodeserti n. sp. A) Paragonopods (leg 9 of male) in anterior view. Scale bar 0.2 mm. B) Paragonopods (leg 9 of male) in ventral view. Scale bar 0.2 mm. C) Pygidium of the male in ventral view. Scale bar: 0.2 mm. D) Right vulva in posterior view. Scale bar: 0.1 mm. E) Right and left vulvae in ventral view. Scale bar: 0.1 mm." captionText-3="FIGURE 6 Gonopods of Ceratosphys cryodeserti n. sp. A) Ventral view. B) Lateral anterior view. C) Anterior view. D) Posterior view. Scale bars: all 0.2 mm." httpUri-0="https://zenodo.org/record/232600/files/figure.png" httpUri-1="https://zenodo.org/record/232601/files/figure.png" httpUri-2="https://zenodo.org/record/232602/files/figure.png" httpUri-3="https://zenodo.org/record/232603/files/figure.png" pageId="7" pageNumber="398">Figs. 3–6</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="13EB6568FFD7FFBD608E4538FAE8F97A" pageId="7" pageNumber="398" type="materials_examined">
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<paragraph id="5B4E36E3FFD7FFBD608E4538FC68FA76" blockId="7.[151,1436,1372,2009]" pageId="7" pageNumber="398">
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<emphasis id="6985EAF1FFD7FFBD608E4538FED8FA9B" bold="true" box="[151,359,1372,1397]" pageId="7" pageNumber="398">Material studied.</emphasis>
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||
<emphasis id="6985EAF1FFD7FFBD616B4538FD85FA9B" box="[370,570,1372,1397]" italics="true" pageId="7" pageNumber="398">
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||
<typeStatus id="844A8841FFD7FFBD616B4538FE63FA9B" box="[370,476,1372,1397]" pageId="7" pageNumber="398" type="holotype">Holotype</typeStatus>
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||
:
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||
<specimenCount id="4DF7FD6AFFD7FFBD61F54538FD85FA9B" box="[492,570,1372,1397]" pageId="7" pageNumber="398" type="male">1 male</specimenCount>
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</emphasis>
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(
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||
<collectionCode id="3DE0AE26FFD7FFBD62554538FD1FFA9A" box="[588,672,1372,1396]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="398">ZMUC</collectionCode>
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||
00040339), Circo Glaciar del Guarnón (Corral del Veleta), Güejar Sierra,
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||
<collectingCountry id="23E67673FFD7FFBD60F1451BFEF5FA76" box="[232,330,1407,1432]" name="Grenada" pageId="7" pageNumber="398">Granada</collectingCountry>
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||
,
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||
<collectingCountry id="23E67673FFD7FFBD614C451BFE27FA76" box="[341,408,1407,1432]" name="Spain" pageId="7" pageNumber="398">Spain</collectingCountry>
|
||
.
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||
<geoCoordinate id="3EC55024FFD7FFBD61BB45E4FD84FA79" box="[418,571,1408,1432]" direction="north" orientation="latitude" pageId="7" pageNumber="398" precision="15" value="37.06361">37° 03' 49" N</geoCoordinate>
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||
<geoCoordinate id="3EC55024FFD7FFBD625B45E4FD6BFA79" box="[578,724,1408,1432]" direction="west" orientation="longitude" pageId="7" pageNumber="398" precision="15" value="-3.3675">3° 22' 03" W</geoCoordinate>
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||
, 3000 masl.
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||
<date id="2F4F1023FFD7FFBD637945E4FC68FA76" box="[864,983,1408,1432]" pageId="7" pageNumber="398" value="2013-09-09">9-IX-2013</date>
|
||
</paragraph>
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||
<paragraph id="5B4E36E3FFD7FFBD60DE45C1FF69F99E" blockId="7.[151,1436,1372,2009]" pageId="7" pageNumber="398">
|
||
<typeStatus id="844A8841FFD7FFBD60DE45C1FE86FA53" box="[199,313,1445,1469]" pageId="7" pageNumber="398" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<emphasis id="6985EAF1FFD7FFBD615D45C0FD77FA52" box="[324,712,1444,1469]" italics="true" pageId="7" pageNumber="398">
|
||
<specimenCount id="4DF7FD6AFFD7FFBD615D45C0FE15FA53" box="[324,426,1444,1469]" pageId="7" pageNumber="398" type="male">16 males</specimenCount>
|
||
<specimenCount id="4DF7FD6AFFD7FFBD61AD45C0FD85FA53" box="[436,570,1444,1469]" pageId="7" pageNumber="398" type="female">30 Females</specimenCount>
|
||
,
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||
<specimenCount id="4DF7FD6AFFD7FFBD625E45C0FD17FA53" box="[583,680,1444,1469]" pageId="7" pageNumber="398" type="generic">1 subad.</specimenCount>
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M
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||
</emphasis>
|
||
(
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<collectionCode id="3DE0AE26FFD7FFBD62C045C0FC92FA52" box="[729,813,1444,1468]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="398">ZMUC</collectionCode>
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||
00040340-344), Circo Glaciar del Guarnón (Corral del Veleta), Güejar Sierra,
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<collectingCountry id="23E67673FFD7FFBD618745A3FDBFFA0E" box="[414,512,1479,1504]" name="Grenada" pageId="7" pageNumber="398">Granada</collectingCountry>
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||
,
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||
<collectingCountry id="23E67673FFD7FFBD621445A3FDEFFA0E" box="[525,592,1479,1504]" name="Spain" pageId="7" pageNumber="398">Spain</collectingCountry>
|
||
.
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||
<geoCoordinate id="3EC55024FFD7FFBD624545ACFD45FA31" box="[604,762,1480,1504]" direction="north" orientation="latitude" pageId="7" pageNumber="398" precision="15" value="37.06361">37° 03' 49" N</geoCoordinate>
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||
<geoCoordinate id="3EC55024FFD7FFBD631A45ACFC24FA31" box="[771,923,1480,1504]" direction="west" orientation="longitude" pageId="7" pageNumber="398" precision="15" value="-3.3675">3° 22' 03" W</geoCoordinate>
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||
, 3000 masl.
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||
<date id="2F4F1023FFD7FFBD643445ACFB1BFA0E" box="[1069,1188,1480,1504]" pageId="7" pageNumber="398" value="2013-09-09">9-IX-2013</date>
|
||
.—
|
||
<specimenCount id="4DF7FD6AFFD7FFBD64DF45ACFA9EFA0E" box="[1222,1313,1479,1504]" pageId="7" pageNumber="398" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="4DF7FD6AFFD7FFBD653745ACFA23FA0E" box="[1326,1436,1479,1504]" pageId="7" pageNumber="398" type="female">2 females</specimenCount>
|
||
UAH (DZAF-UA/VMO), same data but
|
||
<date id="2F4F1023FFD7FFBD62794588FD47F9EA" box="[608,760,1516,1540]" pageId="7" pageNumber="398" value="2014-08-14">14-VIII-2014</date>
|
||
.—
|
||
<specimenCount id="4DF7FD6AFFD7FFBD63034588FCCCF9EB" box="[794,883,1516,1541]" pageId="7" pageNumber="398" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="4DF7FD6AFFD7FFBD63674588FC55F9EB" box="[894,1002,1516,1541]" pageId="7" pageNumber="398" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="3DE0AE26FFD7FFBD63E04588FBF0F9EA" LSID="urn:lsid:biocol.org:col:35275" box="[1017,1103,1516,1540]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="398">MNCN</collectionCode>
|
||
20.07/
|
||
<date id="2F4F1023FFD7FFBD64B94588FAA4F9EB" box="[1184,1307,1516,1541]" pageId="7" pageNumber="398" value="2005" valueMax="2008">2005-2008</date>
|
||
), same data but
|
||
<date id="2F4F1023FFD7FFBD60DD4674FEE3F9C6" box="[196,348,1552,1576]" pageId="7" pageNumber="398" value="2014-08-14">14-VIII-2014</date>
|
||
.—
|
||
<specimenCount id="4DF7FD6AFFD7FFBD61674674FE67F9C6" box="[382,472,1551,1576]" pageId="7" pageNumber="398" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="4DF7FD6AFFD7FFBD61FF4674FDEBF9C6" box="[486,596,1551,1576]" pageId="7" pageNumber="398" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="3DE0AE26FFD7FFBD627C4674FD00F9C6" LSID="urn:lsid:biocol.org:col:34988" box="[613,703,1552,1576]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="398">MNHN</collectionCode>
|
||
, Collection Myriapodes-Onychophores, DA 280) Same data but
|
||
<date id="2F4F1023FFD7FFBD608E4650FE90F9A2" box="[151,303,1588,1612]" pageId="7" pageNumber="398" value="2014-08-14">14-VIII-2014</date>
|
||
.—
|
||
<specimenCount id="4DF7FD6AFFD7FFBD61484650FE16F9A3" box="[337,425,1588,1613]" pageId="7" pageNumber="398" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="4DF7FD6AFFD7FFBD61AC4650FD9FF9A3" box="[437,544,1588,1613]" pageId="7" pageNumber="398" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="3DE0AE26FFD7FFBD62364650FD36F9A2" LSID="urn:lsid:biocol.org:col:34988" box="[559,649,1588,1612]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="398">MNHN</collectionCode>
|
||
, Collection Myriapodes-Onychophores, DA 280) Same data but
|
||
<date id="2F4F1023FFD7FFBD65474650FF6CF99E" pageId="7" pageNumber="398" value="2013-09-09">9-IX- 2013</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD7FFBD60DE4618FAE8F97A" blockId="7.[151,1436,1372,2009]" box="[199,1367,1660,1685]" pageId="7" pageNumber="398">
|
||
Referred non-type material: 124F, 43 M,
|
||
<specimenCount id="4DF7FD6AFFD7FFBD628D4618FD5BF97A" box="[660,740,1660,1685]" pageId="7" pageNumber="398" type="generic">15 Juv.</specimenCount>
|
||
UAH (DZAF-UA/VMO). Same data but
|
||
<date id="2F4F1023FFD7FFBD64A14618FAECF97A" box="[1208,1363,1660,1684]" pageId="7" pageNumber="398" value="2014-08-14">14-VIII-2014</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13EB6568FFD7FFBD60DE46FBFBEBF956" box="[199,1108,1695,1720]" pageId="7" pageNumber="398" type="etymology">
|
||
<paragraph id="5B4E36E3FFD7FFBD60DE46FBFBEBF956" blockId="7.[151,1436,1372,2009]" box="[199,1108,1695,1720]" pageId="7" pageNumber="398">
|
||
<emphasis id="6985EAF1FFD7FFBD60DE46FBFEF7F956" box="[199,328,1695,1720]" italics="true" pageId="7" pageNumber="398">Etymology:</emphasis>
|
||
the species name means “of the cold desert” and refers to the habitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13EB6568FFD7FFB660DE46A1FD64FC6E" lastPageId="12" lastPageNumber="403" pageId="7" pageNumber="398" type="description">
|
||
<paragraph id="5B4E36E3FFD7FFBD60DE46A1FA99F8CB" blockId="7.[151,1436,1372,2009]" pageId="7" pageNumber="398">
|
||
<emphasis id="6985EAF1FFD7FFBD60DE46A1FEEBF932" box="[199,340,1733,1756]" italics="true" pageId="7" pageNumber="398">Description:</emphasis>
|
||
colour brownish/yellowish. Prozonites paler than metazonites, with the exception of the equally pale paraterga. Adult males and females with 30 ‘segments’ (including collum and telson) and 50 pairs of legs including gonopods in males. Males reaching
|
||
<quantity id="9C099B06FFD7FFBD62834768FD55F8CA" box="[666,746,1804,1828]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="398" unit="mm" value="13.0">13 mm</quantity>
|
||
in length; females slightly bigger, reaching
|
||
<quantity id="9C099B06FFD7FFBD64CE4768FA9DF8CB" box="[1239,1314,1804,1829]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="398" unit="mm" value="15.0">15mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD7FFBD60DE474BFB57F8A6" blockId="7.[151,1436,1372,2009]" box="[199,1256,1839,1864]" pageId="7" pageNumber="398">
|
||
<emphasis id="6985EAF1FFD7FFBD60DE474BFEB2F8A6" bold="true" box="[199,269,1839,1864]" pageId="7" pageNumber="398">Male:</emphasis>
|
||
vertical maximum diameter
|
||
<quantity id="9C099B06FFD7FFBD62484754FD07F8A6" box="[593,696,1840,1864]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="398" unit="mm" value="0.9">0.90 mm</quantity>
|
||
. Horizontal maximum diameter (width)
|
||
<quantity id="9C099B06FFD7FFBD64654754FB5CF8A6" box="[1148,1251,1840,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="398" unit="mm" value="1.24">1.24 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD7FFB260DE4730FD8FFF16" blockId="7.[151,1436,1372,2009]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="399" pageId="7" pageNumber="398">
|
||
Head: no special characters. Width, including strongly convex mandibular stipites reaching
|
||
<quantity id="9C099B06FFD7FFBD64F84730FAF8F882" box="[1249,1351,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="398" unit="mm" value="1.08">1.08 mm</quantity>
|
||
(
|
||
<figureCitation id="C3CA2A66FFD7FFBD65414730FA23F882" box="[1368,1436,1876,1901]" captionStart="FIGURE 3" captionStartId="8.[151,250,1504,1526]" captionTargetBox="[159,1427,309,1472]" captionTargetId="figure@8.[151,1436,301,1483]" captionTargetPageId="8" captionText="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="7" pageNumber="398">Fig. 3</figureCitation>
|
||
A). 22–26 ocelli per eye, arranged 8 vertical rows in a triangular field (
|
||
<figureCitation id="C3CA2A66FFD7FFBD63D04713FBB2F87E" box="[969,1037,1911,1936]" captionStart="FIGURE 3" captionStartId="8.[151,250,1504,1526]" captionTargetBox="[159,1427,309,1472]" captionTargetId="figure@8.[151,1436,301,1483]" captionTargetPageId="8" captionText="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="7" pageNumber="398">Fig. 3</figureCitation>
|
||
B). Numerous relatively long setae scattered over the head (but not as a dense pilosity). Labrum with 3 teeth, 7 + 8 supralabral setae (
|
||
<figureCitation id="C3CA2A66FFD7FFBD64F447F8FA8FF85A" box="[1261,1328,1948,1973]" captionStart="FIGURE 3" captionStartId="8.[151,250,1504,1526]" captionTargetBox="[159,1427,309,1472]" captionTargetId="figure@8.[151,1436,301,1483]" captionTargetPageId="8" captionText="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="7" pageNumber="398">Fig. 3</figureCitation>
|
||
A). Frons slightly concave in males. Area behind antennal insertion with cobblestone paving-like microsculpture like that of
|
||
<taxonomicName id="9CF14D60FFD8FFB2608E40FCFC67FF5E" authority="Reboleira & Enghoff, 2014 (Reboleira & Enghoff 2014)" authorityName="Reboleira & Enghoff, 2014 (Reboleira & Enghoff" authorityYear="2014" box="[151,984,151,176]" class="Diplopoda" family="Opisthocheiridae" genus="Sireuma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="nobile">
|
||
<emphasis id="6985EAF1FFD8FFB2608E40FCFEFCFF5E" box="[151,323,151,176]" italics="true" pageId="8" pageNumber="399">Sireuma nobile</emphasis>
|
||
<bibRefCitation id="3F604B12FFD8FFB2615540F3FD3AFF5E" author="Reboleira" box="[332,645,151,176]" pageId="8" pageNumber="399" refString="Reboleira, A. S. P. S. & Enghoff, H. (2014) Sireuma, a new genus of subterranean millipedes from the Iberian Peninsula (Diplopoda, Chordeumatida, Opisthocheiridae). Zootaxa, 3785 (1), 79 - 86. http: // dx. doi. org / 10.11646 / zootaxa. 3785.1.6" type="journal article" year="2014">Reboleira & Enghoff, 2014</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="3F604B12FFD8FFB2628F40F3FC6FFF5E" author="Reboleira" box="[662,976,151,176]" pageId="8" pageNumber="399" refString="Reboleira, A. S. P. S. & Enghoff, H. (2014) Sireuma, a new genus of subterranean millipedes from the Iberian Peninsula (Diplopoda, Chordeumatida, Opisthocheiridae). Zootaxa, 3785 (1), 79 - 86. http: // dx. doi. org / 10.11646 / zootaxa. 3785.1.6" type="journal article" year="2014">Reboleira & Enghoff 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Antennae with 7 visible antennomeres measuring 0.10, 0.30, 0.54, 0.36, 0.51, 0.28 and
|
||
<quantity id="9C099B06FFD8FFB262A640D8FC99FF3A" box="[703,806,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="399" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, respectively; ‘antennal club’ (antennomeres 4–8) ca. 5 times longer than broad (
|
||
<figureCitation id="C3CA2A66FFD8FFB261B640BBFE51FF16" box="[431,494,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1500,1522]" captionTargetBox="[349,1237,396,1472]" captionTargetId="figure@2.[346,1241,393,1479]" captionTargetPageId="2" captionText="FIGURE 1. Pictures taken at the Veleta Glaciar Cirque: A, B) Aspect of the scree where the MSS was sampled. C) Subterranean Sampling Device (SSD) where the trap was placed, inside a hole dug in the scree where the interstices among the rocks are visible D) The place where the SSD is left, once buried and covered with stones to mark its position. E) Ommatoiulus ilicis photographed alive." httpUri="https://zenodo.org/record/232598/files/figure.png" pageId="8" pageNumber="399">Fig. 1</figureCitation>
|
||
A,C).
|
||
</paragraph>
|
||
<caption id="0F8E666BFFD8FFB2608E4584FDF3F9DA" httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="8" pageNumber="399" targetBox="[159,1427,309,1472]" targetPageId="8">
|
||
<paragraph id="5B4E36E3FFD8FFB2608E4584FDF3F9DA" blockId="8.[151,1436,1504,1588]" pageId="8" pageNumber="399">
|
||
<emphasis id="6985EAF1FFD8FFB2608E4584FEB0FA1B" bold="true" box="[151,271,1504,1526]" pageId="8" pageNumber="399">FIGURE 3</emphasis>
|
||
Anatomy details of the male of
|
||
<taxonomicName id="9CF14D60FFD8FFB2627D4584FCE2FA18" box="[612,861,1504,1526]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="cryodeserti" status="sp. nov.">
|
||
<emphasis id="6985EAF1FFD8FFB2627D4584FCE2FA18" box="[612,861,1504,1526]" italics="true" pageId="8" pageNumber="399">Ceratosphys cryodeserti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6985EAF1FFD8FFB2637F4585FC1EFA18" bold="true" box="[870,929,1505,1526]" pageId="8" pageNumber="399">
|
||
<taxonomicNameLabel id="72B6578AFFD8FFB2637F4585FC1EFA18" box="[870,929,1505,1526]" pageId="8" pageNumber="399" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘segments’ in dorsal view. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B4E36E3FFD8FFB260DE463FFE42F8ED" blockId="8.[151,1437,1627,2012]" pageId="8" pageNumber="399">
|
||
Collum (
|
||
<figureCitation id="C3CA2A66FFD8FFB26129463FFEC6F99A" box="[304,377,1627,1652]" captionStart="FIGURE 3" captionStartId="8.[151,250,1504,1526]" captionTargetBox="[159,1427,309,1472]" captionTargetId="figure@8.[151,1436,301,1483]" captionTargetPageId="8" captionText="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="8" pageNumber="399">Fig. 3</figureCitation>
|
||
B): anterior margin almost semi-circular, although when observed with optic microscopy apparently trapezoidal, reaching a maximum posterior width of
|
||
<quantity id="9C099B06FFD8FFB26367461BFC56F979" box="[894,1001,1663,1688]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="399" unit="mm" value="0.76">0.76 mm</quantity>
|
||
. 3 + 3 macrosetae: lateral seta in the posterior corner; intermediate seta close to the lateral one nearly half-way between anterior and posterior margins of collum; mesal seta approximately mid-way between anterior and posterior margins, a little closer to median suture than to intermediate seta.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD8FFB260DE476BFBC3F856" blockId="8.[151,1437,1627,2012]" pageId="8" pageNumber="399">
|
||
Trunk pleurotergites progressively wider, except for 6 and 7 (
|
||
<quantity id="9C099B06FFD8FFB2639F476BFC4EF8C6" box="[902,1009,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="8" pageNumber="399" unit="mm" value="1.26">1.26 mm</quantity>
|
||
), that are abruptly wider than 5 (
|
||
<quantity id="9C099B06FFD8FFB265734774FF7BF8A2" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="399" unit="mm" value="1.08">1.08 mm</quantity>
|
||
) and 8 (
|
||
<quantity id="9C099B06FFD8FFB2612B4757FE20F8A2" box="[306,415,1843,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="399" unit="mm" value="1.15">1.15 mm</quantity>
|
||
) (
|
||
<figureCitation id="C3CA2A66FFD8FFB261A54757FDBCF8A2" box="[444,515,1843,1868]" captionStart="FIGURE 3" captionStartId="8.[151,250,1504,1526]" captionTargetBox="[159,1427,309,1472]" captionTargetId="figure@8.[151,1436,301,1483]" captionTargetPageId="8" captionText="FIGURE 3 Anatomy details of the male of Ceratosphys cryodeserti n. sp. A) Head in anterior view. Scale bar 0.5 mm. B) Head, collum and second ‘ segment’ in lateral view. Scale bar 0.2 mm. C) Tip of the antenna. Scale bar 0.2 mm. D) First seven ‘ segments’ in dorsal view. Scale bar 1 mm." httpUri="https://zenodo.org/record/232600/files/figure.png" pageId="8" pageNumber="399">Fig. 3</figureCitation>
|
||
D) due to the presence of gonopods. All ‘segments’ with 3+3 medium-sized macrosetae, approximately as long as the distance to the next one. Mesal seta slightly closer to median suture than to intermediate seta, and slightly anterior to it. Intermediate seta located anteriorly and dorsally to the paranotum; lateral seta located ventrally and posteriorly to it. This pattern is repeated along the body.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD8FFB260DE47A0FA23F832" blockId="8.[151,1437,1627,2012]" box="[199,1436,1987,2012]" pageId="8" pageNumber="399">
|
||
Legs 1 and 2 with a single row of densely set setae on ventral face of tarsus, resembling a brush (
|
||
<figureCitation id="C3CA2A66FFD8FFB2652A47A7FAC7F832" box="[1331,1400,1987,2012]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="8" pageNumber="399">Fig. 4</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFD9FFB3608E40F3FA25FE66" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="400">
|
||
Ventral face of tarsus from legs 3 (
|
||
<figureCitation id="C3CA2A66FFD9FFB3622240F3FD3CFF5E" box="[571,643,151,176]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
B) to 40 with a region with multiple tongue-shaped, almost fanlike modified setae, wider at their extreme than in their basis, absent from the last eight pairs of legs. Length of leg 7 and 10: 1.35 and
|
||
<quantity id="9C099B06FFD9FFB361414084FE0FFF19" box="[344,432,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="400" unit="mm" value="1.3">1.3 mm</quantity>
|
||
, respectively. Coxa of leg 7 (
|
||
<figureCitation id="C3CA2A66FFD9FFB362EF40BBFC86FF16" box="[758,825,223,248]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
C,D,E) with a small distal expansion. Prefemur of legs 10 and 11 (
|
||
<figureCitation id="C3CA2A66FFD9FFB361044160FEDCFEF2" box="[285,355,260,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
C,D,F,G) with both a dorsal distal blunt protuberance and another ventral one, bigger in size, lending the prefemur the aspect of a battle axe. All coxae along the body, as well as the median sternal knob, with regions covered by scaly-knobby microsculpture (
|
||
<figureCitation id="C3CA2A66FFD9FFB362DC4128FCB6FE8A" box="[709,777,332,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
C,D). Claws similar along the body (that of leg 11
|
||
<quantity id="9C099B06FFD9FFB365204128FA22FE8A" box="[1337,1437,332,356]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="400" unit="mm" value="0.08">0.08 mm</quantity>
|
||
in length) with two setae dorsally and one ventrally, similar in length, or almost inappreciably larger than the claw.
|
||
</paragraph>
|
||
<caption id="0F8E666BFFD9FFB3608E4716FD19F80A" httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="9" pageNumber="400" targetBox="[157,1428,453,1872]" targetPageId="9">
|
||
<paragraph id="5B4E36E3FFD9FFB3608E4716FD19F80A" blockId="9.[151,1436,1906,2020]" pageId="9" pageNumber="400">
|
||
<emphasis id="6985EAF1FFD9FFB3608E4716FEB2F869" bold="true" box="[151,269,1906,1928]" pageId="9" pageNumber="400">FIGURE 4</emphasis>
|
||
Legs of the male of
|
||
<taxonomicName id="9CF14D60FFD9FFB361F94716FD67F866" box="[480,728,1906,1928]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="cryodeserti" status="sp. nov.">
|
||
<emphasis id="6985EAF1FFD9FFB361F94716FD67F866" box="[480,728,1906,1928]" italics="true" pageId="9" pageNumber="400">Ceratosphys cryodeserti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6985EAF1FFD9FFB362C74716FCA7F869" bold="true" box="[734,792,1906,1927]" pageId="9" pageNumber="400">
|
||
<taxonomicNameLabel id="72B6578AFFD9FFB362C74716FCA7F869" box="[734,792,1906,1927]" pageId="9" pageNumber="400" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0F8E666BFFDAFFB0608E4572FAD9FA84" httpUri="https://zenodo.org/record/232602/files/figure.png" pageId="10" pageNumber="401" targetBox="[162,1427,190,1265]" targetPageId="10">
|
||
<paragraph id="5B4E36E3FFDAFFB0608E4572FAD9FA84" blockId="10.[151,1436,1302,1386]" pageId="10" pageNumber="401">
|
||
<emphasis id="6985EAF1FFDAFFB0608E4572FEB2FAC2" bold="true" box="[151,269,1302,1324]" pageId="10" pageNumber="401">FIGURE 5</emphasis>
|
||
Anatomical details of
|
||
<taxonomicName id="9CF14D60FFDAFFB061EF4572FD4CFAC2" authority="Cryodeserti" authorityName="Cryodeserti" box="[502,755,1302,1324]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6985EAF1FFDAFFB061EF4572FD4CFAC2" box="[502,755,1302,1324]" italics="true" pageId="10" pageNumber="401">Ceratosphys Cryodeserti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6985EAF1FFDAFFB062E34573FC8CFAC2" bold="true" box="[762,819,1303,1324]" pageId="10" pageNumber="401">n. sp.</emphasis>
|
||
A) Paragonopods (leg 9 of male) in anterior view. Scale bar 0.2 mm. B) Paragonopods (leg 9 of male) in ventral view. Scale bar 0.2 mm. C) Pygidium of the male in ventral view. Scale bar: 0.2 mm. D) Right vulva in posterior view. Scale bar: 0.1 mm. E) Right and left vulvae in ventral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B4E36E3FFDAFFB060DE45F5FBD7F9D4" blockId="10.[151,1437,1425,2026]" pageId="10" pageNumber="401">
|
||
Telson (
|
||
<figureCitation id="C3CA2A66FFDAFFB0613D45F5FED4FA44" box="[292,363,1425,1450]" captionStart="FIGURE 5" captionStartId="10.[151,250,1302,1324]" captionTargetBox="[162,1427,190,1265]" captionTargetId="figure@10.[151,1436,179,1281]" captionTargetPageId="10" captionText="FIGURE 5 Anatomical details of Ceratosphys Cryodeserti n. sp. A) Paragonopods (leg 9 of male) in anterior view. Scale bar 0.2 mm. B) Paragonopods (leg 9 of male) in ventral view. Scale bar 0.2 mm. C) Pygidium of the male in ventral view. Scale bar: 0.2 mm. D) Right vulva in posterior view. Scale bar: 0.1 mm. E) Right and left vulvae in ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/232602/files/figure.png" pageId="10" pageNumber="401">Fig. 5</figureCitation>
|
||
C): Epiproct dorsally with 3+3 setae dorsally; posterior margin truncate, with two divergent spinnerets. Paraprocts with three setae each: uppermost seta almost at dorsal edge, second seta slightly above the half of the height of paraproct, and third seta approximately half-way from second to ventral edge of paraproct. Each paraproct with a semicircular crease, starting close to the hypoproct and almost reaching the third seta. Hypoproct semielliptical, with two setae and a narrow flange along its external margin.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDAFFB060DE4621FBC3F9B3" blockId="10.[151,1437,1425,2026]" box="[199,1148,1605,1630]" pageId="10" pageNumber="401">
|
||
Anterior gonopods (
|
||
<figureCitation id="C3CA2A66FFDAFFB061B04621FE53F9B0" box="[425,492,1605,1630]" captionStart="FIGURE 6" captionStartId="11.[151,250,1627,1649]" captionTargetBox="[159,1430,452,1593]" captionTargetId="figure@11.[151,1436,445,1606]" captionTargetPageId="11" captionText="FIGURE 6 Gonopods of Ceratosphys cryodeserti n. sp. A) Ventral view. B) Lateral anterior view. C) Anterior view. D) Posterior view. Scale bars: all 0.2 mm." httpUri="https://zenodo.org/record/232603/files/figure.png" pageId="10" pageNumber="401">Fig. 6</figureCitation>
|
||
) consisting of the three main parts typical of
|
||
<taxonomicName id="9CF14D60FFDAFFB063F14621FBCBF9B0" box="[1000,1140,1605,1630]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6985EAF1FFDAFFB063F14621FBCBF9B0" box="[1000,1140,1605,1630]" italics="true" pageId="10" pageNumber="401">Ceratosphys</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDAFFB060DE460EFB02F8B4" blockId="10.[151,1437,1425,2026]" pageId="10" pageNumber="401">
|
||
a) Paired telopodites (
|
||
<emphasis id="6985EAF1FFDAFFB061DD460DFE68F96C" bold="true" box="[452,471,1641,1666]" pageId="10" pageNumber="401">T</emphasis>
|
||
): Each telopodite is curved and sinuous, anteriorly convex and posteriorly concave, with the distal tips slightly curved. There is a basal bifurcation into two branches: one external (
|
||
<emphasis id="6985EAF1FFDAFFB0651746E9FA97F948" bold="true" box="[1294,1320,1677,1702]" pageId="10" pageNumber="401">Te</emphasis>
|
||
) and one internal (
|
||
<emphasis id="6985EAF1FFDAFFB060E446D5FEA8F924" bold="true" box="[253,279,1713,1738]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
). The distal part of the external branch (
|
||
<emphasis id="6985EAF1FFDAFFB062F846D5FD44F924" bold="true" box="[737,763,1713,1738]" pageId="10" pageNumber="401">Te</emphasis>
|
||
) is hidden below the expanded distal portion of (
|
||
<emphasis id="6985EAF1FFDAFFB0653346D5FAFBF924" bold="true" box="[1322,1348,1713,1738]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
). Close to the tip (
|
||
<emphasis id="6985EAF1FFDAFFB0611346B1FE9BF900" bold="true" box="[266,292,1749,1774]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
) there is a very prominent pointed lateral process (
|
||
<emphasis id="6985EAF1FFDAFFB0634546B1FCCFF900" bold="true" box="[860,880,1749,1774]" pageId="10" pageNumber="401">ts</emphasis>
|
||
). On the lateral margin of (
|
||
<emphasis id="6985EAF1FFDAFFB064BB46B1FB03F900" bold="true" box="[1186,1212,1749,1774]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
), basal to (
|
||
<emphasis id="6985EAF1FFDAFFB0652046B1FAF2F900" bold="true" box="[1337,1357,1749,1774]" pageId="10" pageNumber="401">ts</emphasis>
|
||
), there is a small, barely visible pointed denticle; further basal there is a posteriorly concave, ladle-like process (
|
||
<emphasis id="6985EAF1FFDAFFB0656C469DFA2DF8FC" bold="true" box="[1397,1426,1785,1810]" pageId="10" pageNumber="401">tw</emphasis>
|
||
) projecting towards the gonopod basis. It is between the base of (
|
||
<emphasis id="6985EAF1FFDAFFB063734779FC37F8D8" bold="true" box="[874,904,1821,1846]" pageId="10" pageNumber="401">tw</emphasis>
|
||
) and the main part of (
|
||
<emphasis id="6985EAF1FFDAFFB064954779FB19F8D8" bold="true" box="[1164,1190,1821,1846]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
) that (
|
||
<emphasis id="6985EAF1FFDAFFB064E84779FAB4F8D8" bold="true" box="[1265,1291,1821,1846]" pageId="10" pageNumber="401">Te</emphasis>
|
||
) reaches the anterior side of (
|
||
<emphasis id="6985EAF1FFDAFFB061484725FED4F8B4" bold="true" box="[337,363,1857,1882]" pageId="10" pageNumber="401">Ti</emphasis>
|
||
). The basal posterior part of the telopodite has a protuse stout process (
|
||
<emphasis id="6985EAF1FFDAFFB064924725FB13F8B4" bold="true" box="[1163,1196,1857,1882]" pageId="10" pageNumber="401">Tx</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDAFFB060DE4701FCCBF804" blockId="10.[151,1437,1425,2026]" pageId="10" pageNumber="401">
|
||
The tracheal apodeme has two processes, one wider posterior process (
|
||
<emphasis id="6985EAF1FFDAFFB063FF4701FBBBF890" bold="true" box="[998,1028,1893,1918]" pageId="10" pageNumber="401">U)</emphasis>
|
||
reaching along the posterior face of the basal part of the telopodite; and one anterior process whose internal preapical apophysis (
|
||
<emphasis id="6985EAF1FFDAFFB064B147EDFB09F84C" bold="true" box="[1192,1206,1929,1954]" pageId="10" pageNumber="401">v</emphasis>
|
||
) is articulated to the edge of the synangiocoxite (
|
||
<emphasis id="6985EAF1FFDAFFB061CA47C9FE58F828" bold="true" box="[467,487,1965,1990]" pageId="10" pageNumber="401">A</emphasis>
|
||
), while the apex (
|
||
<emphasis id="6985EAF1FFDAFFB062AB47C9FD79F828" bold="true" box="[690,710,1965,1990]" pageId="10" pageNumber="401">w</emphasis>
|
||
) is free. Also, there is a short basal central process (
|
||
<emphasis id="6985EAF1FFDAFFB0651747C9FAA5F828" bold="true" box="[1294,1306,1965,1990]" pageId="10" pageNumber="401">z</emphasis>
|
||
) projecting distad between the two proximad lobes (
|
||
<emphasis id="6985EAF1FFDAFFB0624447B5FDD4F804" bold="true" box="[605,619,2001,2026]" pageId="10" pageNumber="401">a</emphasis>
|
||
) of the synangiocoxite.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDBFFB160DE40F3FCD0FF16" blockId="11.[151,1436,151,392]" pageId="11" pageNumber="402">
|
||
b) Synangiocoxite (
|
||
<emphasis id="6985EAF1FFDBFFB161BD40F3FE07FF5E" bold="true" box="[420,440,151,176]" pageId="11" pageNumber="402">A</emphasis>
|
||
): Neither lateral horns (
|
||
<emphasis id="6985EAF1FFDBFFB162DE40F3FD6AFF5E" bold="true" box="[711,725,151,176]" pageId="11" pageNumber="402">g</emphasis>
|
||
), nor the “cloison endosqueletique” (
|
||
<emphasis id="6985EAF1FFDBFFB1646040F3FB37FF5E" bold="true" box="[1145,1160,151,176]" pageId="11" pageNumber="402">h</emphasis>
|
||
) as found in
|
||
<taxonomicName id="9CF14D60FFDBFFB1650540FCFEFBFF3B" authority="Mauries 1969" authorityName="Mauries" authorityYear="1969" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="402" phylum="Arthropoda" rank="species" species="soutadei">
|
||
<emphasis id="6985EAF1FFDBFFB1650540FCFA23FF5E" box="[1308,1436,151,176]" italics="true" pageId="11" pageNumber="402">C. soutadei</emphasis>
|
||
(
|
||
<bibRefCitation id="3F604B12FFDBFFB1608740D8FE83FF3B" author="Mauries" box="[158,316,188,213]" pageId="11" pageNumber="402" refString="Mauries, J. P. (1969) Myriapodes de Sierra Nevada (Espagne): Une nouvelle espece du genre Ceratosphys Ribaut, 1920 (Diplopoda). Publicaciones del Instituto de Biologia Aplicada, 47,131 - 138." type="book chapter" year="1969">Mauriès 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
are present. There is a rounded basal lobe (
|
||
<emphasis id="6985EAF1FFDBFFB1633240D8FC86FF3B" bold="true" box="[811,825,188,213]" pageId="11" pageNumber="402">a</emphasis>
|
||
) projecting proximad at each side, with a medial ridge (
|
||
<emphasis id="6985EAF1FFDBFFB160B940BBFF13FF16" bold="true" box="[160,172,223,248]" pageId="11" pageNumber="402">c</emphasis>
|
||
) between them just above process (
|
||
<emphasis id="6985EAF1FFDBFFB1622740BBFDF5FF16" bold="true" box="[574,586,223,248]" pageId="11" pageNumber="402">z</emphasis>
|
||
) of the tracheal apodeme.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDBFFB160DE4160FC05FE66" blockId="11.[151,1436,151,392]" pageId="11" pageNumber="402">
|
||
c) Syncolpocoxite (
|
||
<emphasis id="6985EAF1FFDBFFB161BD4160FE05FEF3" bold="true" box="[420,442,260,285]" pageId="11" pageNumber="402">K</emphasis>
|
||
) with a distal extension (
|
||
<emphasis id="6985EAF1FFDBFFB162C54160FD45FEF3" bold="true" box="[732,762,260,285]" pageId="11" pageNumber="402">ky</emphasis>
|
||
) resembling a “Y” and forming a continuation of a median keel and projecting from the concavity between the denticles (
|
||
<emphasis id="6985EAF1FFDBFFB1637B4143FCC0FEAE" bold="true" box="[866,895,295,320]" pageId="11" pageNumber="402">ka</emphasis>
|
||
).
|
||
<emphasis id="6985EAF1FFDBFFB163814143FC11FEAE" bold="true" box="[920,942,295,320]" pageId="11" pageNumber="402">K</emphasis>
|
||
furthermore with a lateral, curved, pointed process (
|
||
<emphasis id="6985EAF1FFDBFFB160E74128FEA5FE8B" bold="true" box="[254,282,332,357]" pageId="11" pageNumber="402">kc</emphasis>
|
||
) on each side, ca. midway between the basis of
|
||
<emphasis id="6985EAF1FFDBFFB1634D4128FCD5FE8B" bold="true" box="[852,874,332,357]" pageId="11" pageNumber="402">K</emphasis>
|
||
to the
|
||
<emphasis id="6985EAF1FFDBFFB163D94128FC61FE8B" bold="true" box="[960,990,332,357]" pageId="11" pageNumber="402">ky</emphasis>
|
||
extension, at the widest part of
|
||
<emphasis id="6985EAF1FFDBFFB165414128FAD1FE8B" bold="true" box="[1368,1390,332,357]" pageId="11" pageNumber="402">K</emphasis>
|
||
;
|
||
<emphasis id="6985EAF1FFDBFFB165994128FA23FE8B" bold="true" box="[1408,1436,332,357]" pageId="11" pageNumber="402">kc</emphasis>
|
||
curving distad, their tips pointing mesad. Basis of K with two lobes (
|
||
<emphasis id="6985EAF1FFDBFFB16383410BFC16FE66" bold="true" box="[922,937,367,392]" pageId="11" pageNumber="402">k</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="0F8E666BFFDBFFB1608E463FFDA1F97E" httpUri="https://zenodo.org/record/232603/files/figure.png" pageId="11" pageNumber="402" targetBox="[159,1430,452,1593]" targetPageId="11">
|
||
<paragraph id="5B4E36E3FFDBFFB1608E463FFDA1F97E" blockId="11.[151,1435,1627,1680]" pageId="11" pageNumber="402">
|
||
<emphasis id="6985EAF1FFDBFFB1608E463FFEAFF99E" bold="true" box="[151,272,1627,1649]" pageId="11" pageNumber="402">FIGURE 6</emphasis>
|
||
Gonopods of
|
||
<taxonomicName id="9CF14D60FFDBFFB161B5463FFD19F99F" box="[428,678,1627,1649]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="402" phylum="Arthropoda" rank="species" species="cryodeserti" status="sp. nov.">
|
||
<emphasis id="6985EAF1FFDBFFB161B5463FFD19F99F" box="[428,678,1627,1649]" italics="true" pageId="11" pageNumber="402">Ceratosphys cryodeserti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6985EAF1FFDBFFB162A94638FD52F99F" bold="true" box="[688,749,1628,1649]" pageId="11" pageNumber="402">
|
||
<taxonomicNameLabel id="72B6578AFFDBFFB162A94638FD52F99F" box="[688,749,1628,1649]" pageId="11" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) Ventral view. B) Lateral anterior view. C) Anterior view. D) Posterior view. Scale bars: all 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B4E36E3FFDBFFB160DE46D8FA22F8AE" blockId="11.[151,1437,1724,2038]" pageId="11" pageNumber="402">
|
||
Posterior gonopods (paragonopods,
|
||
<figureCitation id="C3CA2A66FFDBFFB1627D46D8FD17F93B" box="[612,680,1724,1749]" captionStart="FIGURE 5" captionStartId="10.[151,250,1302,1324]" captionTargetBox="[162,1427,190,1265]" captionTargetId="figure@10.[151,1436,179,1281]" captionTargetPageId="10" captionText="FIGURE 5 Anatomical details of Ceratosphys Cryodeserti n. sp. A) Paragonopods (leg 9 of male) in anterior view. Scale bar 0.2 mm. B) Paragonopods (leg 9 of male) in ventral view. Scale bar 0.2 mm. C) Pygidium of the male in ventral view. Scale bar: 0.2 mm. D) Right vulva in posterior view. Scale bar: 0.1 mm. E) Right and left vulvae in ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/232602/files/figure.png" pageId="11" pageNumber="402">Fig. 5</figureCitation>
|
||
A,B): Telopodite with three articles, but only distal article clearly delimited: basal article with a conical small sclerotized cuticular appendage with a prickly microsculpture; second article basally wide, then anteriorly concavely narrowing towards tip, with some scattered setae anteriorly, and with a distal rounded protuberance with multiple anterior setae; third article pyriform, darker, with a distal terminal seta.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDBFFB660DE4729FE4EFEAE" blockId="11.[151,1437,1724,2038]" lastBlockId="12.[151,1436,151,896]" lastPageId="12" lastPageNumber="403" pageId="11" pageNumber="402">
|
||
<emphasis id="6985EAF1FFDBFFB160DE4729FE86F888" bold="true" box="[199,313,1869,1894]" pageId="11" pageNumber="402">Female: s</emphasis>
|
||
imilar to male but with some differences, in addition to bigger size. Width across mandibular stipites
|
||
<quantity id="9C099B06FFDBFFB1608E4714FEBEF866" box="[151,257,1904,1929]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="11" pageNumber="402" unit="mm" value="11.6">11.6 mm</quantity>
|
||
. Trunk pleurotergites 6–7 not wider than neighbouring ones. Legs 1 and 2 as in male with a row of densely set setae on the ventral face of metatarsus, but the fanlike setae found on males from leg 3 to 40 are absent. Vulvae (
|
||
<figureCitation id="C3CA2A66FFDBFFB160E347DDFEFEF83C" box="[250,321,1977,2002]" captionStart="FIGURE 5" captionStartId="10.[151,250,1302,1324]" captionTargetBox="[162,1427,190,1265]" captionTargetId="figure@10.[151,1436,179,1281]" captionTargetPageId="10" captionText="FIGURE 5 Anatomical details of Ceratosphys Cryodeserti n. sp. A) Paragonopods (leg 9 of male) in anterior view. Scale bar 0.2 mm. B) Paragonopods (leg 9 of male) in ventral view. Scale bar 0.2 mm. C) Pygidium of the male in ventral view. Scale bar: 0.2 mm. D) Right vulva in posterior view. Scale bar: 0.1 mm. E) Right and left vulvae in ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/232602/files/figure.png" pageId="11" pageNumber="402">Fig. 5</figureCitation>
|
||
C,D): Operculum (
|
||
<emphasis id="6985EAF1FFDBFFB1620747DDFD84F83C" bold="true" box="[542,571,1977,2002]" pageId="11" pageNumber="402">op</emphasis>
|
||
) with an anterior fingerlike expansion (supra-opercular prolongation
|
||
<emphasis id="6985EAF1FFDBFFB1657347DDFA2DF83C" bold="true" box="[1386,1426,1977,2002]" pageId="11" pageNumber="402">sop</emphasis>
|
||
) projecting distad, almost as long as the vulva. Both mesal (
|
||
<emphasis id="6985EAF1FFDBFFB1632147B9FCF2F818" bold="true" box="[824,845,2013,2038]" pageId="11" pageNumber="402">vi</emphasis>
|
||
) and lateral (
|
||
<emphasis id="6985EAF1FFDBFFB163F147B9FBBCF818" bold="true" box="[1000,1027,2013,2038]" pageId="11" pageNumber="402">ve</emphasis>
|
||
) valves flanked by a group or more than 10 long setae, their mesal margins along the apodematic groove liplike. At the distal tip of the vulva, between
|
||
<emphasis id="6985EAF1FFDCFFB6608E40D8FF00FF3B" bold="true" box="[151,191,188,213]" pageId="12" pageNumber="403">sop</emphasis>
|
||
and the apodematic groove, the valves come together in a structure (
|
||
<emphasis id="6985EAF1FFDCFFB663C340D8FC59FF3B" bold="true" box="[986,998,188,213]" pageId="12" pageNumber="403">z</emphasis>
|
||
) with an almost circular emargination with only a narrow opening where
|
||
<emphasis id="6985EAF1FFDCFFB6623F4085FD8EFF16" box="[550,561,225,248]" italics="true" pageId="12" pageNumber="403">z</emphasis>
|
||
embraces the distal end of the apodematic groove. Each vulva further with a posterior foliose postvulvar organ (
|
||
<emphasis id="6985EAF1FFDCFFB6622D4160FDEEFEF3" bold="true" box="[564,593,260,285]" pageId="12" pageNumber="403">pv</emphasis>
|
||
) covering the apodematic groove; length of
|
||
<emphasis id="6985EAF1FFDCFFB6647C4160FB3DFEF3" bold="true" box="[1125,1154,260,285]" pageId="12" pageNumber="403">pv</emphasis>
|
||
varying slightly among specimens, its margin serrated.
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDCFFB660DE4128FAB4FDF6" blockId="12.[151,1436,151,896]" pageId="12" pageNumber="403">
|
||
<emphasis id="6985EAF1FFDCFFB660DE4128FEAEFE8B" bold="true" box="[199,273,332,357]" pageId="12" pageNumber="403">Notes.</emphasis>
|
||
Study of the new species of
|
||
<taxonomicName id="9CF14D60FFDCFFB6626F4128FCBCFE8B" box="[630,771,332,357]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="403" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6985EAF1FFDCFFB6626F4128FCBCFE8B" box="[630,771,332,357]" italics="true" pageId="12" pageNumber="403">Ceratosphys</emphasis>
|
||
</taxonomicName>
|
||
revealed several gonopod differences
|
||
<emphasis id="6985EAF1FFDCFFB664D34129FA96FE8A" box="[1226,1321,333,356]" italics="true" pageId="12" pageNumber="403">vis-a-vis</emphasis>
|
||
the nearsyntopic
|
||
<taxonomicName id="9CF14D60FFDCFFB661194114FE3DFE66" box="[256,386,367,392]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="403" phylum="Arthropoda" rank="species" species="soutadei">
|
||
<emphasis id="6985EAF1FFDCFFB661194114FE3DFE66" box="[256,386,367,392]" italics="true" pageId="12" pageNumber="403">C. soutadei</emphasis>
|
||
</taxonomicName>
|
||
(characters from
|
||
<taxonomicName id="9CF14D60FFDCFFB662484114FD6BFE66" box="[593,724,367,392]" class="Diplopoda" family="Opisthocheiridae" genus="Ceratosphys" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="403" phylum="Arthropoda" rank="species" species="soutadei">
|
||
<emphasis id="6985EAF1FFDCFFB662484114FD6BFE66" box="[593,724,367,392]" italics="true" pageId="12" pageNumber="403">C. soutadei</emphasis>
|
||
</taxonomicName>
|
||
in parentheses,
|
||
<typeStatus id="844A8841FFDCFFB66388410BFC4CFE66" box="[913,1011,367,392]" pageId="12" pageNumber="403" type="holotype">holotype</typeStatus>
|
||
re-examined by JPM): Telopodite is deeply divided into two branches
|
||
<emphasis id="6985EAF1FFDCFFB6622D41F0FDF0FE43" bold="true" box="[564,591,404,429]" pageId="12" pageNumber="403">Te</emphasis>
|
||
and
|
||
<emphasis id="6985EAF1FFDCFFB6628C41F0FD10FE43" bold="true" box="[661,687,404,429]" pageId="12" pageNumber="403">Ti</emphasis>
|
||
(undivided); lateral process (
|
||
<emphasis id="6985EAF1FFDCFFB6640D41F0FB97FE43" bold="true" box="[1044,1064,404,429]" pageId="12" pageNumber="403">ts</emphasis>
|
||
) of
|
||
<emphasis id="6985EAF1FFDCFFB6647C41F0FBC0FE43" bold="true" box="[1125,1151,404,429]" pageId="12" pageNumber="403">Ti</emphasis>
|
||
prominent (tiny); basal telopodital process (
|
||
<emphasis id="6985EAF1FFDCFFB6616441D3FE21FE3E" bold="true" box="[381,414,439,464]" pageId="12" pageNumber="403">Tx</emphasis>
|
||
) more prominent (smaller,
|
||
<emphasis id="6985EAF1FFDCFFB662CE41DDFD5CFE3E" box="[727,739,441,464]" italics="true" pageId="12" pageNumber="403">J</emphasis>
|
||
in
|
||
<bibRefCitation id="3F604B12FFDCFFB6631241D3FC15FE3E" author="Mauries" box="[779,938,439,464]" pageId="12" pageNumber="403" refString="Mauries, J. P. (1969) Myriapodes de Sierra Nevada (Espagne): Une nouvelle espece du genre Ceratosphys Ribaut, 1920 (Diplopoda). Publicaciones del Instituto de Biologia Aplicada, 47,131 - 138." type="book chapter" year="1969">Mauriès 1969</bibRefCitation>
|
||
,
|
||
<figureCitation id="C3CA2A66FFDCFFB663AE41D3FC43FE3E" box="[951,1020,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1982]" captionTargetBox="[225,1367,1165,1929]" captionTargetId="figure@3.[212,1376,1154,1939]" captionTargetPageId="3" captionText="FIGURE 2 Temperature and humidity in the MSS from September 2013 to August 2014. The pale line represents the temperature and the dark line the relative humidity." httpUri="https://zenodo.org/record/232599/files/figure.png" pageId="12" pageNumber="403">Fig. 2</figureCitation>
|
||
); synangiocoxite with neither lateral horns (quite prominent, g in Mauriès’s
|
||
<figureCitation id="C3CA2A66FFDCFFB6624841B8FD2DFE1A" box="[593,658,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1500,1522]" captionTargetBox="[349,1237,396,1472]" captionTargetId="figure@2.[346,1241,393,1479]" captionTargetPageId="2" captionText="FIGURE 1. Pictures taken at the Veleta Glaciar Cirque: A, B) Aspect of the scree where the MSS was sampled. C) Subterranean Sampling Device (SSD) where the trap was placed, inside a hole dug in the scree where the interstices among the rocks are visible D) The place where the SSD is left, once buried and covered with stones to mark its position. E) Ommatoiulus ilicis photographed alive." httpUri="https://zenodo.org/record/232598/files/figure.png" pageId="12" pageNumber="403">Fig. 1</figureCitation>
|
||
), nor the cloison endosqueletique (present, h in Mauriès’ fig. 1), nor the denticles (present,
|
||
<emphasis id="6985EAF1FFDCFFB6618A419BFE1EFDF6" box="[403,417,511,536]" italics="true" pageId="12" pageNumber="403">d</emphasis>
|
||
in Mauriès’ fig. 1); syncolpocoxite (
|
||
<emphasis id="6985EAF1FFDCFFB66324419BFCECFDF6" bold="true" box="[829,851,511,536]" pageId="12" pageNumber="403">K</emphasis>
|
||
) with a distal appendage (
|
||
<emphasis id="6985EAF1FFDCFFB66465419BFB26FDF6" bold="true" box="[1148,1177,511,536]" pageId="12" pageNumber="403">ky</emphasis>
|
||
) (absent).
|
||
</paragraph>
|
||
<paragraph id="5B4E36E3FFDCFFB660DE4240FD64FC6E" blockId="12.[151,1436,151,896]" pageId="12" pageNumber="403">
|
||
Certain other characters of the new species are also worth comment. A ventral row of knifelike modified setae on the ventral side of tarsus 1 and 2 (
|
||
<figureCitation id="C3CA2A66FFDCFFB662374223FDCFFD8E" box="[558,624,583,608]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="12" pageNumber="403">Fig. 4</figureCitation>
|
||
A) is known also from certain other chordeumatidans (
|
||
<bibRefCitation id="3F604B12FFDCFFB664D64223FAEEFD8E" author="Shear" box="[1231,1361,583,608]" pageId="12" pageNumber="403" refString="Shear, W. (2011) Cave millipeds of the United States. XI. Opiona graeningi, n. sp., a troglomorphic caseyid milliped from Siskiyou County, California, with comments on the genus Opiona Chamberlin 1951 (Diplopoda, Chordeumatida, Caseyidae). Zootaxa, 3114, 50 - 56." type="journal article" year="2011">Shear 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F604B12FFDCFFB665454223FEFDFD6A" author="Shear" pageId="12" pageNumber="403" refString="Shear, W. & Krejca, J. K. (2007) Revalidation of the milliped genus Amplaria Chamberlin 1941 (Diplopoda, Chordeumatida, Striariidae), and description of two new species from caves in Sequoia and Kings Canyon National Parks, California. Zootaxa, 1532, 23 - 39." type="journal article" year="2007">Shear & Krejca 2007</bibRefCitation>
|
||
) including females (
|
||
<bibRefCitation id="3F604B12FFDCFFB662334208FCA4FD6A" author="Marti" box="[554,795,620,645]" pageId="12" pageNumber="403" refString="Marti, P. & Wuest, J. (1998) Repartition geographique et morphologie fine de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). Revue Suisse de Zoologie, 105 (1), 15 - 23. http: // dx. doi. org / 10.5962 / bhl. part. 80029" type="journal article" year="1998">Marti & Wüest 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F604B12FFDCFFB663314208FBE4FD6A" author="Reboleira" box="[808,1115,620,645]" pageId="12" pageNumber="403" refString="Reboleira, A. S. P. S. & Enghoff, H. (2014) Sireuma, a new genus of subterranean millipedes from the Iberian Peninsula (Diplopoda, Chordeumatida, Opisthocheiridae). Zootaxa, 3785 (1), 79 - 86. http: // dx. doi. org / 10.11646 / zootaxa. 3785.1.6" type="journal article" year="2014">Reboleira & Enghoff 2014</bibRefCitation>
|
||
). Their function is not clear, although they may possibly be used to clean the antennae and/or mouthparts. The fanlike setae on the ventral side of male tarsus 3 onward (
|
||
<figureCitation id="C3CA2A66FFDCFFB661DD42D0FDB4FD22" box="[452,523,692,717]" captionStart="FIGURE 4" captionStartId="9.[151,250,1906,1928]" captionTargetBox="[157,1428,453,1872]" captionTargetId="figure@9.[151,1436,445,1884]" captionTargetPageId="9" captionText="FIGURE 4 Legs of the male of Ceratosphys cryodeserti n. sp. A) Tarsus of legs 1 and 2 with their ventral row of setae. Scale bar 0.1 mm. B) Tarsus of leg 3 with the ventral fanners. Scale bar 0.1 mm. C) Body in ventral view, legs from 7 to 13 visible. Scale bar 0.2 mm. D) Body in ventral view, legs from 10 to 13 visible. Scale bar 0.2 mm. E) Leg 7. Scale bar 0.3 mm. F) Leg 10. Scale bar 0.4 mm. G) Leg 11. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/232601/files/figure.png" pageId="12" pageNumber="403">Fig. 4</figureCitation>
|
||
B) are known from some other chordeumatidan species (
|
||
<bibRefCitation id="3F604B12FFDCFFB664B942D0FA28FD22" author="Marti" box="[1184,1431,692,717]" pageId="12" pageNumber="403" refString="Marti, P. & Wuest, J. (1998) Repartition geographique et morphologie fine de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). Revue Suisse de Zoologie, 105 (1), 15 - 23. http: // dx. doi. org / 10.5962 / bhl. part. 80029" type="journal article" year="1998">Marti & Wüest 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F604B12FFDCFFB6608E42B3FE3EFD1E" author="Makarov" box="[151,385,727,752]" pageId="12" pageNumber="403" refString="Makarov, S. E., Curcic, B. P., Tomic, V. T., Rada, T., Rada, B., Curcic, S. B, Mitic, B. M. & Lucic, L. R. (2011) Revision of the family Heterolatzeliidae (Diplopoda, Chordeumatida). Zootaxa, 2994, 33 - 44." type="journal article" year="2011">
|
||
Makarov
|
||
<emphasis id="6985EAF1FFDCFFB6611D42BDFE89FD1E" box="[260,310,727,752]" italics="true" pageId="12" pageNumber="403">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="3F604B12FFDCFFB6618F42B3FD30FD1E" box="[406,655,727,752]" pageId="12" pageNumber="403" refString="Makarov, S. E., Curcic, B. P., Tomic, V. T., Rada, T., Rada, B., Curcic, S. B, Mitic, B. M. & Lucic, L. R. (2011) Revision of the family Heterolatzeliidae (Diplopoda, Chordeumatida). Zootaxa, 2994, 33 - 44." type="journal article">
|
||
Makarov
|
||
<emphasis id="6985EAF1FFDCFFB6621D42BDFD89FD1E" box="[516,566,727,752]" italics="true" pageId="12" pageNumber="403">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
call them tarsal papillae, stating that they are common in the family
|
||
<taxonomicName id="9CF14D60FFDCFFB6608E4298FEE6FCFB" box="[151,345,764,789]" class="Diplopoda" family="Heterolatzeliidae" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="403" phylum="Arthropoda" rank="family">Heterolatzeliidae</taxonomicName>
|
||
, but do not assign them a function. Their shape and the fact that they are present only in in males suggest that they are instrumental in grasping the female during mating, as stated by
|
||
<bibRefCitation id="3F604B12FFDCFFB6644B437BFAF0FCD6" author="Marti" box="[1106,1359,799,824]" pageId="12" pageNumber="403" refString="Marti, P. & Wuest, J. (1998) Repartition geographique et morphologie fine de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). Revue Suisse de Zoologie, 105 (1), 15 - 23. http: // dx. doi. org / 10.5962 / bhl. part. 80029" type="journal article" year="1998">Marti & Wüest (1998)</bibRefCitation>
|
||
due to their similarity with analogous structures in males of numerous insects, as for example in the protarsi of males of many
|
||
<taxonomicName id="9CF14D60FFDCFFB660C54303FEE8FC6E" box="[220,343,871,896]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="403" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
<taxonomicName id="9CF14D60FFDCFFB661474303FE6FFC6E" box="[350,464,871,896]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="403" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="3F604B12FFDCFFB6622F4303FD70FC6E" author="Jeannel" box="[566,719,871,896]" pageId="12" pageNumber="403" refString="Jeannel, R. (1941) Coleopteres Carabiques. Deuxieme partie. Faune de France 39. Lechevalier, Paris, 571 pp." type="book" year="1941">Jeannel 1941</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |