306 lines
36 KiB
XML
306 lines
36 KiB
XML
<document id="59C4807E644B93C6A4653C8E49826221" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. & Van Wyk, Abraham E." docDate="2017" docId="03F687CD462DFFC2FF2AC8F3FBF954D3" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis nitida H. M. Steyn 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="113" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="111" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B2F94A77B83BF338418AB975CC6E139C">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
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<mods:name id="7E26C6F6B837A4DD84A9642EA8211D38" type="personal">
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<mods:roleTerm id="583D2F248E8D504E79C5C27A3362C930">Author</mods:roleTerm>
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<mods:namePart id="67BBB0A3FB8513CB7C574F299CD89339">Steyn, Hester M.</mods:namePart>
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<mods:affiliation id="2831BE680915409F181A9BBE4171FBBF">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
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<mods:namePart id="E1486456C0EA223BF14BA23D5A078BE1">Van Wyk, Abraham E.</mods:namePart>
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<mods:affiliation id="8929A8C75C4109A6F67FF7CC49216826">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
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<mods:typeOfResource id="66351D9CBA922DD089EBD5186DBFD962">text</mods:typeOfResource>
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<mods:title id="12C30591F9F379C877C3CB4017D0CE5C">Phytotaxa</mods:title>
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<mods:part id="8CA0875265AD9EBF7AA04CC55A62B164">
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<mods:date id="660DF3C623B2E423BC5F7968C5A4E729">2017</mods:date>
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<mods:number id="B9349A33974EE83B5ADC260BF2960B0F">2017-10-10</mods:number>
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<mods:number id="413DB1B335FF9CC8584E4DC4DF121DFC">324</mods:number>
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<mods:url id="3B104EEBBEEF98B37F77DD659D141E8F">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
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<mods:classification id="E681FE30945B1CBA7164A3093B9F437E">journal article</mods:classification>
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<mods:identifier id="24B723F4D887C3B46378980C61BACA52" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
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<mods:identifier id="A675F239871B13DC2A81FA0267BB4AB2" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03F687CD462DFFC2FF2AC8F3FBF954D3" ID-DOI="http://doi.org/10.5281/zenodo.13697382" ID-Zenodo-Dep="13697382" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3" httpUri="http://treatment.plazi.org/id/03F687CD462DFFC2FF2AC8F3FBF954D3" lastPageId="12" lastPageNumber="113" pageId="10" pageNumber="111">
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<subSubSection id="C3456550462DFFC4FF2AC8F3FCBE529D" box="[136,807,1411,1438]" pageId="10" pageNumber="111" type="nomenclature">
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<paragraph id="8BE036DB462DFFC4FF2AC8F3FCBE529D" blockId="10.[136,1452,1411,1617]" box="[136,807,1411,1438]" pageId="10" pageNumber="111">
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<heading id="D0A881B7462DFFC4FF2AC8F3FCBE529D" box="[136,807,1411,1438]" fontSize="11" level="2" pageId="10" pageNumber="111" reason="4">
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<emphasis id="B92BEAC9462DFFC4FF2AC8F3FF04529D" bold="true" box="[136,157,1412,1438]" pageId="10" pageNumber="111">3.</emphasis>
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<taxonomicName id="4C5F4D58462DFFC4FF01C8F5FE66529D" ID-CoL="8ZLG" authority="H.M.Steyn" authorityName="H. M. Steyn" authorityYear="2017" box="[163,511,1411,1438]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="nitida" status="sp. nov.">
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<emphasis id="B92BEAC9462DFFC4FF01C8F5FEE2529D" bold="true" box="[163,379,1411,1438]" italics="true" pageId="10" pageNumber="111">Acanthopsis nitida</emphasis>
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H.M.Steyn
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857B2462DFFC4FDA8C8F3FDF8529D" box="[522,609,1413,1438]" pageId="10" pageNumber="111" rank="species">
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<emphasis id="B92BEAC9462DFFC4FDA8C8F3FDC0529D" box="[522,601,1413,1438]" italics="true" pageId="10" pageNumber="111">sp. nov</emphasis>
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.
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||
</taxonomicNameLabel>
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(
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<figureCitation id="13642A5E462DFFC4FDD2C8F3FD24529D" box="[624,701,1413,1438]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunberg’s route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="10" pageNumber="111">Figs. 1</figureCitation>
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,
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<figureCitation id="13642A5E462DFFC4FD69C8F3FD75529D" box="[715,748,1413,1438]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="10" pageNumber="111">3C</figureCitation>
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&
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<figureCitation id="13642A5E462DFFC4FCB2C8F3FC86529D" box="[784,799,1413,1438]" captionStart="FIGURE 6" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[151,1436,353,1933]" captionTargetId="figure-89@11.[151,1436,353,1933]" captionTargetPageId="11" captionText="FIGURE 6. Acanthopsis nitida. A & B. Habit. C & D. Leaf indumentum (abaxial). E & F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: C–I = Steyn 1884 (PRE), J = Burgoyne 1514 (PRE), K = Steyn 2129 (PRE). Photographed by: M. Koekemoer (A & B) and H.M. Steyn (C–K)." figureDoi="http://doi.org/10.5281/zenodo.13697372" httpUri="https://zenodo.org/record/13697372/files/figure.png" pageId="10" pageNumber="111">6</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3456550462DFFC4FF2AC8DAFD1D52E6" pageId="10" pageNumber="111" type="materials_examined">
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<paragraph id="8BE036DB462DFFC4FF2AC8DAFD1D52E6" blockId="10.[136,1452,1411,1617]" pageId="10" pageNumber="111">
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<emphasis id="B92BEAC9462DFFC4FF2AC8DAFF5B52C2" bold="true" box="[136,194,1452,1473]" pageId="10" pageNumber="111">Type:</emphasis>
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—
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<collectingCountry id="F348764B462DFFC4FF79C8DAFE1552C2" box="[219,396,1452,1473]" name="South Africa" pageId="10" pageNumber="111">SOUTH AFRICA</collectingCountry>
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.
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<collectingRegion id="499BF839462DFFC4FE35C8DAFDB352C2" box="[407,554,1452,1473]" country="South Africa" name="Northern Cape" pageId="10" pageNumber="111">Northern Cape</collectingRegion>
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: Namaqualand District. Anenous Pass, S of road,
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<quantity id="4CA79B3E462DFFC4FBB7C8DAFBCF52C2" box="[1045,1110,1452,1473]" metricMagnitude="2" metricUnit="m" metricValue="7.09" pageId="10" pageNumber="111" unit="m" value="709.0">709 m</quantity>
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, (2917
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<collectionCode id="ED4EAE1E462DFFC4FB39C8DAFB2752C2" box="[1179,1214,1452,1473]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="10" pageNumber="111" type="Herbarium">BA</collectionCode>
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),
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<date id="FFE1101B462DFFC4FB73C8DAFAF552C2" box="[1233,1388,1452,1473]" pageId="10" pageNumber="111" value="2015-10-09">9 October 2015</date>
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,
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<emphasis id="B92BEAC9462DFFC4FAD5C8DAFF6D52E6" italics="true" pageId="10" pageNumber="111">Steyn 2129,</emphasis>
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(
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<typeStatus id="54E48879462DFFC4FEA2C8A6FECF52E6" box="[256,342,1488,1509]" pageId="10" pageNumber="111" type="holotype">holotype</typeStatus>
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<collectionCode id="ED4EAE1E462DFFC4FEF9C8A6FE1352E6" box="[347,394,1488,1509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
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0866187!,
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<typeStatus id="54E48879462DFFC4FE4DC8A6FDAD52E6" box="[495,564,1488,1509]" pageId="10" pageNumber="111" type="isotype">isotype</typeStatus>
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||
<collectionCode id="ED4EAE1E462DFFC4FD99C8A6FDEF52E6" box="[571,630,1488,1509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="10" pageNumber="111" type="Herbarium">NBG</collectionCode>
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!).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3456550462DFFC5FF2AC882FF65564A" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111" type="diagnosis">
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<paragraph id="8BE036DB462DFFC4FF2AC882FD855152" blockId="10.[136,1452,1411,1617]" pageId="10" pageNumber="111">
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<emphasis id="B92BEAC9462DFFC4FF2AC882FF6A510A" bold="true" box="[136,243,1524,1545]" pageId="10" pageNumber="111">Diagnosis:</emphasis>
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—
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<taxonomicName id="4C5F4D58462DFFC4FEA9C882FE5B510A" authorityName="H. M. Steyn" authorityYear="2017" box="[267,450,1524,1545]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="nitida">
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<emphasis id="B92BEAC9462DFFC4FEA9C882FE5B510A" box="[267,450,1524,1545]" italics="true" pageId="10" pageNumber="111">Acanthopsis nitida</emphasis>
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</taxonomicName>
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differs from all other members of the group with semi-dense spikes and 5-fid bracts in its glossy appearance (
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<emphasis id="B92BEAC9462DFFC4FE95CB6EFECF512E" box="[311,342,1560,1581]" italics="true" pageId="10" pageNumber="111">not</emphasis>
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matt), yellow-green puberulent to sub-glabrous leaves (
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<emphasis id="B92BEAC9462DFFC4FCD0CB6EFC08512E" box="[882,913,1560,1581]" italics="true" pageId="10" pageNumber="111">not</emphasis>
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grey or green strigose to villose leaves) and in having a glabrous (
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<emphasis id="B92BEAC9462DFFC4FEBCCB4AFEA45152" box="[286,317,1596,1617]" italics="true" pageId="10" pageNumber="111">not</emphasis>
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variously hairy) calyx.
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</paragraph>
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<paragraph id="8BE036DB462DFFC5FF2ACB0CFF65564A" blockId="10.[136,1452,1658,2079]" lastBlockId="11.[136,1452,160,329]" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111">
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Perennial, compact subshrub,
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<quantity id="4CA79B3E462DFFC4FE42CB0CFDDA5190" box="[480,579,1658,1683]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="10" pageNumber="111" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 cm</quantity>
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tall with very short internodes.
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<emphasis id="B92BEAC9462DFFC4FC14CB0CFB9D5190" box="[950,1028,1658,1683]" italics="true" pageId="10" pageNumber="111">Leaves</emphasis>
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sessile, base petiole-like with spines
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<quantity id="4CA79B3E462DFFC4FF2ACBE8FF7151B4" box="[136,232,1694,1719]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="10" pageNumber="111" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
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long; lamina oblanceolate, 40–60(–90) ×
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<quantity id="4CA79B3E462DFFC4FD6DCBE8FCA651B4" box="[719,831,1694,1719]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="111" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
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, often with yellow-green appearance, puberulent with strigose to spreading short hairs, also with subsessile glandular hairs; apex acute, base attenuate, margin undulate, dentate-spinose, spines rigid, 3–5(–8) mm long, yellow to brown. Inflorescences semi-dense, cylindrical, subsessile,
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||
<quantity id="4CA79B3E462DFFC4FF2ACA7CFE995020" box="[136,256,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="10" pageNumber="111" unit="mm" value="50.0" valueMax="55.0" valueMin="45.0">45–55 mm</quantity>
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long, (8–)
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||
<quantity id="4CA79B3E462DFFC4FED3CA7CFE735020" box="[369,490,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="10" pageNumber="111" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
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in diameter. Bracts broadly obovate to wedge-shaped, (15–)
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||
<quantity id="4CA79B3E462DFFC4FB22CA7CFB655020" box="[1152,1276,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="111" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 mm</quantity>
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long (including spines), lamina
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||
<quantity id="4CA79B3E462DFFC4FE9ECA58FE3E5044" box="[316,423,1838,1863]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="10" pageNumber="111" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
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||
long, base obtuse to attenuate; lamina and spines glabrous to puberulent with short white, spreading, hairs and isolated subsessile glandular hairs abaxially, silky-hairy adaxially; middle and upper bracts ending in 5(–7) primary spines; central primary spine often compound with one pair of marginal secondary spines, often simple; lateral primary spines very often with 1 long basal secondary spine (shorter than primary spine) on side facing central primary spine and a short marginal secondary spine on opposite side; spines divergent in flower, spreading to recurved in fruit. Bracteoles linear, 8(–12) mm long, silky-hairy adaxially. Calyx brown-tipped with dorsal sepal ovate, acuminate, 13–15(–16) mm long, glabrous abaxially, silky-hairy adaxially, 7–9(–11)-veined; ventral sepal ovate, 12– 13(–15) mm long, glabrous abaxially, silky-hairy adaxially, 5–7(–9)-veined; lateral sepals 8–9(–10) mm long, glabrous abaxially, silky-hairy adaxially, especially towards base. Corolla lilac to purple with lemon-yellow throat, 25–27 ×
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<quantity id="4CA79B3E462CFFC5FF2ACD9EFE9D5602" box="[136,260,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="11" pageNumber="112" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
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, tube 8(–9) mm long, central lobe as wide as long, constricted at the base, emarginate. Filaments
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||
<quantity id="4CA79B3E462CFFC5FAEDCD9EFA355602" box="[1359,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="112" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
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long, glandular; anthers beige,
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<quantity id="4CA79B3E462CFFC5FE40CC7AFDBA5626" box="[482,547,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
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||
long. Style covered with glandular hairs at the base. Capsules 8–9 ×
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<quantity id="4CA79B3E462CFFC5FA85CC7AFAF35626" box="[1319,1386,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
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. Seed 3 ×
|
||
<quantity id="4CA79B3E462CFFC5FF16CC46FF61564A" box="[180,248,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF206653462CFFC5FF2ACAD0FB505F00" ID-DOI="http://doi.org/10.5281/zenodo.13697372" ID-Zenodo-Dep="13697372" httpUri="https://zenodo.org/record/13697372/files/figure.png" pageId="11" pageNumber="112" startId="11.[136,229,1958,1979]" targetBox="[151,1436,353,1933]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BE036DB462CFFC5FF2ACAD0FB505F00" blockId="11.[136,1452,1958,2051]" pageId="11" pageNumber="112">
|
||
<emphasis id="B92BEAC9462CFFC5FF2ACAD0FF6650B8" bold="true" box="[136,255,1958,1979]" pageId="11" pageNumber="112">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C5F4D58462CFFC5FEA5CAD0FE2550B8" authorityName="H. M. Steyn" authorityYear="2017" box="[263,444,1958,1979]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="112" phylum="Tracheophyta" rank="species" species="nitida">
|
||
<emphasis id="B92BEAC9462CFFC5FEA5CAD0FE2550B8" box="[263,444,1958,1979]" italics="true" pageId="11" pageNumber="112">Acanthopsis nitida</emphasis>
|
||
</taxonomicName>
|
||
. A & B. Habit. C & D. Leaf indumentum (abaxial). E & F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: C–I =
|
||
<emphasis id="B92BEAC9462CFFC5FB7ACABCFAD850DC" box="[1240,1345,1994,2015]" italics="true" pageId="11" pageNumber="112">Steyn 1884</emphasis>
|
||
(PRE), J =
|
||
<emphasis id="B92BEAC9462CFFC5FF2ACA98FE835F00" box="[136,282,2030,2051]" italics="true" pageId="11" pageNumber="112">Burgoyne 1514</emphasis>
|
||
(PRE), K =
|
||
<emphasis id="B92BEAC9462CFFC5FE31CA98FE645F00" box="[403,509,2030,2051]" italics="true" pageId="11" pageNumber="112">Steyn 2129</emphasis>
|
||
(PRE). Photographed by: M. Koekemoer (A & B) and H.M. Steyn (C–K).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3456550462BFFC2FF1FCDE9FEEB57DF" pageId="12" pageNumber="113" type="etymology">
|
||
<paragraph id="8BE036DB462BFFC2FF1FCDE9FEEB57DF" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
|
||
<emphasis id="B92BEAC9462BFFC2FF1FCDE9FED157BB" bold="true" box="[189,328,159,184]" pageId="12" pageNumber="113">Etymology:</emphasis>
|
||
—The specific epithet
|
||
<taxonomicName id="4C5F4D58462BFFC2FDEECDE9FD1457BB" authorityName="H. M. Steyn" authorityYear="2017" box="[588,653,159,184]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
|
||
<emphasis id="B92BEAC9462BFFC2FDEECDE9FD1457BB" box="[588,653,159,184]" italics="true" pageId="12" pageNumber="113">nitida</emphasis>
|
||
</taxonomicName>
|
||
, meaning “shiny/glossy”, refers to the glossy leaves, floral bracts and calyx of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3456550462BFFC2FF1FCD91FBF954D3" pageId="12" pageNumber="113" type="distribution">
|
||
<paragraph id="8BE036DB462BFFC2FF1FCD91FE4A56FF" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
|
||
<emphasis id="B92BEAC9462BFFC2FF1FCD91FD1A5603" bold="true" box="[189,643,231,256]" pageId="12" pageNumber="113">Distribution, ecology and phenology:</emphasis>
|
||
—
|
||
<taxonomicName id="4C5F4D58462BFFC2FD3DCD91FCED5603" authorityName="H. M. Steyn" authorityYear="2017" box="[671,884,231,256]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
|
||
<emphasis id="B92BEAC9462BFFC2FD3DCD91FCED5603" box="[671,884,231,256]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the
|
||
<collectingRegion id="499BF839462BFFC2FBFACD91FA9B5603" box="[1112,1282,231,256]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
|
||
and is known from three localities in an area to the west of Steinkopf (
|
||
<figureCitation id="13642A5E462BFFC2FCB5CC7DFCC25627" box="[791,859,267,292]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunberg’s route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="12" pageNumber="113">Fig. 1</figureCitation>
|
||
). It is centred in the core part of the Gariep Centre of Endemism (
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FE93CC59FDD4564B" author="Van Wyk, A. E. & Smith, G. F." box="[305,589,303,328]" pageId="12" pageNumber="113" refId="ref13657" refString="Van Wyk, A. E. & Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Pretoria, 160 pp." type="book" year="2001">Van Wyk & Smith 2001</bibRefCitation>
|
||
), a biogeographical region rich in restricted-range plants and animals. This distribution falls within the Succulent Karoo Biome (
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FD4BCC25FBB7566F" author="Rutherford, M. C. & Westfall, R. H." box="[745,1070,339,364]" pageId="12" pageNumber="113" refId="ref13271" refString="Rutherford, M. C. & Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford & Westfall 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FB99CC25FAB3566F" author="Low, A. B. & Rebelo, A. G." box="[1083,1322,339,364]" pageId="12" pageNumber="113" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs & Tourism, Pretoria, 85 pp." type="book" year="1996">Low & Rebelo 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FA95CC25FEDF5693" author="Mucina, L. & Rutherford, M. C." pageId="12" pageNumber="113" refId="ref13030" refString="Mucina, L. & Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) in the Richtersveld Bioregion, Anenous Plateau Shrubland (
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FBA3CC01FAA75693" author="Mucina, L. & Rutherford, M. C." box="[1025,1342,375,400]" pageId="12" pageNumber="113" refId="ref13030" refString="Mucina, L. & Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) where it receives predominantly winter rains with an average of ±
|
||
<quantity id="4CA79B3E462BFFC2FCBECCEDFCE356B7" box="[796,890,411,436]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="113" unit="mm" value="150.0">150 mm</quantity>
|
||
per year. The specimens were collected on stony mountain slopes in well-drained sandy soils at elevations of
|
||
<quantity id="4CA79B3E462BFFC2FCEACCC9FC4356DB" box="[840,986,447,472]" metricMagnitude="2" metricUnit="m" metricValue="8.95" metricValueMax="11.0" metricValueMin="6.9" pageId="12" pageNumber="113" unit="m" value="895.0" valueMax="1100.0" valueMin="690.0">690–1100 m</quantity>
|
||
. Specimens with flowers were collected between August and October.
|
||
</paragraph>
|
||
<paragraph id="8BE036DB462BFFC2FF1FCF71FEE4558F" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
|
||
<emphasis id="B92BEAC9462BFFC2FF1FCF71FE2B5523" bold="true" box="[189,434,519,544]" pageId="12" pageNumber="113">Conservation status:</emphasis>
|
||
—
|
||
<taxonomicName id="4C5F4D58462BFFC2FE6CCF71FD045523" authorityName="H. M. Steyn" authorityYear="2017" box="[462,669,519,544]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
|
||
<emphasis id="B92BEAC9462BFFC2FE6CCF71FD045523" box="[462,669,519,544]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
|
||
</taxonomicName>
|
||
is a range-restricted species (EOO
|
||
<quantity id="4CA79B3E462BFFC2FB82CF71FBFF5523" box="[1056,1126,519,544]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="113" unit="km" value="42.0">42 km</quantity>
|
||
²) known from three locations within a small area. It is potentially threatened by habitat loss and degradation due to mining and overgrazing and is categorised as Endangered B1ab(iii) according to the IUCN Red List Category and Criteria (
|
||
<bibRefCitation id="EFCE4B2A462BFFC2FB18CF39FADC556B" author="IUCN" box="[1210,1349,591,616]" pageId="12" pageNumber="113" refId="ref12795" refString="IUCN (2012) IUCN Red List categories and criteria version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
|
||
) (L. von Staden, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="8BE036DB462BFFC2FF1FCFE1FE665443" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
|
||
<materialsCitation id="3B373C86462BFFC2FF1FCFE1FBF555D7" collectingDate="2012-08-22" collectionCode="BA, PRE" collectorName="Namaqualand District. Anenous Pass, S" country="South Africa" elevation="691" location="Northern Cape" pageId="12" pageNumber="113" specimenCount="2" stateProvince="Northern Cape">
|
||
<emphasis id="B92BEAC9462BFFC2FF1FCFE1FD7F55B3" bold="true" box="[189,742,663,688]" pageId="12" pageNumber="113">
|
||
<typeStatus id="54E48879462BFFC2FF1FCFE1FDDD55B3" box="[189,580,663,688]" isOtherTypeStatus="true" pageId="12" pageNumber="113">Additional specimens examined</typeStatus>
|
||
(
|
||
<typeStatus id="54E48879462BFFC2FDF8CFE1FD4055B3" box="[602,729,663,688]" pageId="12" pageNumber="113" type="paratype">Paratypes</typeStatus>
|
||
):
|
||
</emphasis>
|
||
—
|
||
<collectingCountry id="F348764B462BFFC2FCA0CFE1FC4C55B3" box="[770,981,663,688]" name="South Africa" pageId="12" pageNumber="113">SOUTH AFRICA</collectingCountry>
|
||
.
|
||
<collectingRegion id="499BF839462BFFC2FC44CFE1FB0E55B3" box="[998,1175,663,688]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
|
||
:
|
||
<collectorName id="26AA530D462BFFC2FB05CFE1FEA655D7" pageId="12" pageNumber="113">Namaqualand District. Anenous Pass, S</collectorName>
|
||
slope of mountain,
|
||
<quantity id="4CA79B3E462BFFC2FDBACFCDFDF955D7" box="[536,608,699,724]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="12" pageNumber="113" unit="m" value="691.0">
|
||
<elevation id="0072D1E8462BFFC2FDBACFCDFDF955D7" box="[536,608,699,724]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="12" pageNumber="113" unit="m" value="691.0">691 m</elevation>
|
||
</quantity>
|
||
, (2917
|
||
<collectionCode id="ED4EAE1E462BFFC2FD0FCFCDFD4F55D7" box="[685,726,699,724]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="12" pageNumber="113" type="Herbarium">BA</collectionCode>
|
||
),
|
||
<date id="FFE1101B462BFFC2FD4ACFCDFC0055D7" box="[744,921,699,724]" pageId="12" pageNumber="113" value="2012-08-22">
|
||
<collectingDate id="EFA5E9F3462BFFC2FD4ACFCDFC0055D7" box="[744,921,699,724]" pageId="12" pageNumber="113" value="2012-08-22">22 August 2012</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B92BEAC9462BFFC2FC01CFCDFB8455D7" box="[931,1053,699,724]" italics="true" pageId="12" pageNumber="113">Steyn 1884</emphasis>
|
||
(
|
||
<collectionCode id="ED4EAE1E462BFFC2FB88CFCDFBFA55D7" box="[1066,1123,699,724]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B373C86462BFFC2FBD4CFCDFAD955D7" box="[1142,1344,699,724]" country="South Africa" location="Steinkopf District" pageId="12" pageNumber="113" specimenCount="1">
|
||
<location id="8E806000462BFFC2FBD4CFCDFAD955D7" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FBD4CFCDFAD955D7" box="[1142,1344,699,724]" country="South Africa" name="Steinkopf District" pageId="12" pageNumber="113">Steinkopf District</location>
|
||
</materialsCitation>
|
||
; Anenous Pass;
|
||
<materialsCitation id="3B373C86462BFFC2FF6ACFA9FD0C55FB" box="[200,661,735,760]" country="South Africa" location="17 km West of Steinkopf" pageId="12" pageNumber="113" specimenCount="1">
|
||
Approximately
|
||
<location id="8E806000462BFFC2FEDACFA9FD0C55FB" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FEDACFA9FD0C55FB" box="[376,661,735,760]" country="South Africa" name="17 km West of Steinkopf" pageId="12" pageNumber="113">
|
||
<quantity id="4CA79B3E462BFFC2FEDACFA9FE5955FB" box="[376,448,735,760]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="113" unit="km" value="17.0">17 km</quantity>
|
||
West of Steinkopf
|
||
</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B373C86462BFFC2FD02CFA9FDF0541F" collectingDate="1992-12-20" collectionCode="BC, PRE" collectorName="Pass" country="South Africa" elevation="1100" pageId="12" pageNumber="113" specimenCount="1">
|
||
hills on southern side of the
|
||
<collectorName id="26AA530D462BFFC2FC42CFA9FB8B55FB" box="[992,1042,735,760]" pageId="12" pageNumber="113">Pass</collectorName>
|
||
on the old road,
|
||
<quantity id="4CA79B3E462BFFC2FB70CFA9FAB155FB" box="[1234,1320,735,760]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="113" unit="m" value="1100.0">
|
||
<elevation id="0072D1E8462BFFC2FB70CFA9FAB155FB" box="[1234,1320,735,760]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="113" unit="m" value="1100.0">1100 m</elevation>
|
||
</quantity>
|
||
, (2917
|
||
<collectionCode id="ED4EAE1E462BFFC2FADACFA9FA0655FB" box="[1400,1439,735,760]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="12" pageNumber="113" type="Herbarium">BC</collectionCode>
|
||
),
|
||
<date id="FFE1101B462BFFC2FF2ACE75FEF8541F" box="[136,353,771,796]" pageId="12" pageNumber="113" value="1992-12-20">
|
||
<collectingDate id="EFA5E9F3462BFFC2FF2ACE75FEF8541F" box="[136,353,771,796]" pageId="12" pageNumber="113" value="1992-12-20">20 December 1992</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B92BEAC9462BFFC2FECFCE75FD81541F" box="[365,536,771,796]" italics="true" pageId="12" pageNumber="113">Burgoyne 1514</emphasis>
|
||
(
|
||
<collectionCode id="ED4EAE1E462BFFC2FD8ACE75FDF8541F" box="[552,609,771,796]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B373C86462BFFC2FDD7CE75FE625443" collectingDate="2015-10-10" collectionCode="BC, PRE" country="South Africa" elevation="838" location="Namaqualand District" pageId="12" pageNumber="113" specimenCount="1">
|
||
<location id="8E806000462BFFC2FDD7CE75FCE9541F" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FDD7CE75FCE9541F" box="[629,880,771,796]" country="South Africa" name="Namaqualand District" pageId="12" pageNumber="113">Namaqualand District</location>
|
||
. On dirt road
|
||
<locationDeviation id="9A522FF9462BFFC2FBAECE75FB36541F" box="[1036,1199,771,796]" location="Eyams" pageId="12" pageNumber="113">NW of Eyams</locationDeviation>
|
||
,
|
||
<quantity id="4CA79B3E462BFFC2FB19CE75FA9C541F" box="[1211,1285,771,796]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="12" pageNumber="113" unit="m" value="838.0">
|
||
<elevation id="0072D1E8462BFFC2FB19CE75FA9C541F" box="[1211,1285,771,796]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="12" pageNumber="113" unit="m" value="838.0">838 m</elevation>
|
||
</quantity>
|
||
, (2917
|
||
<collectionCode id="ED4EAE1E462BFFC2FAF6CE75FAE2541F" box="[1364,1403,771,796]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="12" pageNumber="113" type="Herbarium">BC</collectionCode>
|
||
),
|
||
<date id="FFE1101B462BFFC2FA2DCE75FEBD5443" pageId="12" pageNumber="113" value="2015-10-10">
|
||
<collectingDate id="EFA5E9F3462BFFC2FA2DCE75FEBD5443" pageId="12" pageNumber="113" value="2015-10-10">10 October 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B92BEAC9462BFFC2FE8DCE51FE325443" box="[303,427,807,832]" italics="true" pageId="12" pageNumber="113">Steyn 2138</emphasis>
|
||
(
|
||
<collectionCode id="ED4EAE1E462BFFC2FE18CE51FE6A5443" box="[442,499,807,832]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE036DB462BFFC2FF1FCE3DFBF954D3" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
|
||
<emphasis id="B92BEAC9462BFFC2FF1FCE3DFE905467" bold="true" box="[189,265,843,868]" pageId="12" pageNumber="113">Notes:</emphasis>
|
||
—
|
||
<taxonomicName id="4C5F4D58462BFFC2FE84CE3DFE6F5467" authorityName="H. M. Steyn" authorityYear="2017" box="[294,502,843,868]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
|
||
<emphasis id="B92BEAC9462BFFC2FE84CE3DFE6F5467" box="[294,502,843,868]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from the other subshrubs with semi-dense inflorescences in the genus by its glossy appearance (
|
||
<emphasis id="B92BEAC9462BFFC2FDECCE19FDEB548B" box="[590,626,879,904]" italics="true" pageId="12" pageNumber="113">not</emphasis>
|
||
matt) and yellow-green (
|
||
<emphasis id="B92BEAC9462BFFC2FC39CE19FC26548B" box="[923,959,879,904]" italics="true" pageId="12" pageNumber="113">not</emphasis>
|
||
green or grey-green) leaves. Although the central primary spine could be simple or compound (one pair of short, marginal secondary spines), the lateral primary spines always have a basal secondary spine on the side facing the central primary spine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |