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<document id="6B8A3F2EBD23FB5EF1DC5329A160EB46" ID-DOI="10.5281/zenodo.12808371" ID-ISSN="1942-1354" ID-Zenodo-Dep="12808371" ID-ZooBank="0D2107D1-BD4D-426C-ABCF-C4CC3EEA060F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721837453179" checkinUser="felipe" docAuthor="Skelley, Paul E." docDate="2021" docId="03F687B7FFE79569FF56FB75F081F967" docLanguage="en" docName="InsectaMundi.2021.0892.1-7.pdf" docOrigin="Insecta Mundi 2021 (892)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Leptorhyparus brasiliensis Minkina 2020" docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="FFCFFFCFFFE4956DFF96FFAFF47BFF9A" masterDocTitle="A new species of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) in amber from the Dominican Republic, with comments on extant species" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1721840401121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="226CC78A8C472FC1A8CD6FF442F13659">A new species of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) in amber from the Dominican Republic, with comments on extant species</mods:title>
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<mods:affiliation id="DAA98CD8AC405A808A7070A6466FBC22">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100 USA</mods:affiliation>
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<treatment id="03F687B7FFE79569FF56FB75F081F967" LSID="urn:lsid:plazi:treatment:03F687B7FFE79569FF56FB75F081F967" httpUri="http://treatment.plazi.org/id/03F687B7FFE79569FF56FB75F081F967" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<subSubSection id="C345652AFFE7956EFF56FB75F6B6FB6D" box="[192,717,1242,1271]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BE036A1FFE7956EFF56FB75F6B6FB6D" blockId="3.[192,717,1242,1304]" box="[192,717,1242,1271]" pageId="3" pageNumber="2">
<heading id="D0A881CDFFE7956EFF56FB75F6B6FB6D" box="[192,717,1242,1271]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
<taxonomicName id="4C5F4D22FFE7956EFF56FB75F6B6FB6D" ID-CoL="7WXFM" authority="Minkina, 2020" authorityName="Minkina" authorityYear="2020" box="[192,717,1242,1271]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE7956EFF56FB75F6B6FB6D" bold="true" box="[192,717,1242,1271]" pageId="3" pageNumber="2">
<emphasis id="B92BEAB3FFE7956EFF56FB75F676FB6D" bold="true" box="[192,525,1242,1271]" italics="true" pageId="3" pageNumber="2">Leptorhyparus brasiliensis</emphasis>
<bibRefCitation id="EFCE4B50FFE7956EFD85FB75F6B6FB6D" author="Minkina, L." box="[531,717,1242,1271]" pageId="3" pageNumber="2" pagination="34 - 39" refId="ref4351" refString="Minkina, L. 2020. Two new species of Leptorhyparus Howden, 2003 from South America. Folia Heyrovskyana series A 28 (2): 34 - 39." type="journal article" year="2020">Minkina, 2020</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C345652AFFE7956EFF56FB51F6F3FA82" box="[192,648,1278,1304]" pageId="3" pageNumber="2" type="description">
<paragraph id="8BE036A1FFE7956EFF56FB51F6F3FA82" blockId="3.[192,717,1242,1304]" box="[192,648,1278,1304]" pageId="3" pageNumber="2">
<figureCitation id="13642A24FFE7956EFF56FB51F50DFA82" box="[192,374,1278,1304]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="3" pageNumber="2">Figures 1, 4, 68</figureCitation>
(see also
<bibRefCitation id="EFCE4B50FFE7956EFE74FB51F6FAFA82" author="Minkina, L." box="[482,641,1278,1304]" pageId="3" pageNumber="2" pagination="34 - 39" refId="ref4351" refString="Minkina, L. 2020. Two new species of Leptorhyparus Howden, 2003 from South America. Folia Heyrovskyana series A 28 (2): 34 - 39." type="journal article" year="2020">Minkina 2020</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE7956EFF76FA89F6DBFAA4" box="[224,672,1318,1342]" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="8BE036A1FFE7956EFF76FA89F6DBFAA4" blockId="3.[224,672,1318,1342]" box="[224,672,1318,1342]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB4F118FFFE7956EFF76FA89F6DBFAA4" box="[224,672,1318,1342]" pageId="3" pageNumber="2">
<treatmentCitation id="0AFE10B0FFE7956EFF76FA89F6E7FAA4" author="Minkina, L." box="[224,668,1318,1342]" page="34" pageId="3" pageNumber="2" year="2020">
<taxonomicName id="4C5F4D22FFE7956EFF76FA89F6E7FAA4" ID-CoL="7WXFM" authority="Minkina 2020: 34" authorityName="Minkina" authorityPageNumber="34" authorityYear="2020" box="[224,668,1318,1342]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE7956EFF76FA89F599FAA4" box="[224,482,1318,1342]" italics="true" pageId="3" pageNumber="2">Leptorhyparus brasiliensis</emphasis>
<bibRefCitation id="EFCE4B50FFE7956EFE7EFA89F6E7FAA4" author="Minkina, L." box="[488,668,1318,1342]" pageId="3" pageNumber="2" pagination="34 - 39" refId="ref4351" refString="Minkina, L. 2020. Two new species of Leptorhyparus Howden, 2003 from South America. Folia Heyrovskyana series A 28 (2): 34 - 39." type="journal article" year="2020">Minkina 2020: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE7956EFF56FAE2F52DF9EE" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BE036A1FFE7956EFF56FAE2F52DF9EE" blockId="3.[191,1442,1357,1652]" pageId="3" pageNumber="2">
<emphasis id="B92BEAB3FFE7956EFF56FAE2F547FAFD" bold="true" box="[192,316,1357,1383]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="4C5F4D22FFE7956EFED7FAE1F626FAF2" authorityName="Minkina" authorityYear="2020" box="[321,605,1358,1384]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE7956EFED7FAE1F626FAF2" box="[321,605,1358,1384]" italics="true" pageId="3" pageNumber="2">Leptorhyparus brasiliensis</emphasis>
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is readily distinguished from the other members of the genus by the smaller body (length
<quantity id="4CA79B44FFE7956EFEC3FAC0F583FA13" box="[341,504,1391,1417]" metricMagnitude="-3" metricUnit="m" metricValue="2.29" metricValueMax="2.6" metricValueMin="1.98" pageId="3" pageNumber="2" unit="mm" value="2.29" valueMax="2.6" valueMin="1.98">1.982.60 mm</quantity>
); head lacking distinct tufts of setae; sides of clypeus distinctly sinuate before genae; pronotum having a long groove in the interval on the outside margin of the costa that reaches the anterior fovea, distinctly separating the paramedian and discolateral costae (
<figureCitation id="13642A24FFE7956EFC14FA1DF7BFFA57" box="[898,964,1458,1485]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
); pronotum having a small anterior lobe of the discolateral costa; elytra with postdiscal bulbs reduced; elytra with caudal bulbs small and not distinctly visible in ventral view (
<figureCitation id="13642A24FFE7956EFE51FA59F671F98A" box="[455,522,1526,1552]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
); median impression of metaventrite widening posteriorly; epipharynx transverse with deeply emarginate anterior margin (
<figureCitation id="13642A24FFE7956EFD1BF9B8F6B4F9AB" box="[653,719,1559,1585]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="3" pageNumber="2">Fig. 6</figureCitation>
); male genitalia with phallobase tubular (
<figureCitation id="13642A24FFE7956EFB08F9B8F086F9AB" box="[1182,1277,1559,1585]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="3" pageNumber="2">Fig. 78</figureCitation>
); parameres 1/3 length of phallobase, symmetrical broadly rounded in caudal view; and it occurs widely in South America east of the Andes.
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE7956EFF56F92BF7AEF8BE" pageId="3" pageNumber="2" type="description">
<paragraph id="8BE036A1FFE7956EFF56F92BF7AEF8BE" blockId="3.[191,1440,1668,1829]" pageId="3" pageNumber="2">
<emphasis id="B92BEAB3FFE7956EFF56F92BF54DF904" bold="true" box="[192,310,1668,1694]" pageId="3" pageNumber="2">Variation.</emphasis>
<bibRefCitation id="EFCE4B50FFE7956EFEAAF92BF591F904" author="Howden HF" box="[316,490,1668,1694]" pageId="3" pageNumber="2" pagination="391 - 402" refId="ref4096" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden (2003)</bibRefCitation>
noted some variation in
<taxonomicName id="4C5F4D22FFE7956EFD68F92AF738F904" authorityName="Howden" authorityYear="2003" box="[766,835,1668,1695]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE7956EFD68F92AF776F904" box="[766,781,1669,1694]" italics="true" pageId="3" pageNumber="2">L</emphasis>
.
<emphasis id="B92BEAB3FFE7956EFC8CF92BF738F904" box="[794,835,1668,1694]" italics="true" pageId="3" pageNumber="2">gilli</emphasis>
</taxonomicName>
, and similar, subtle variation is seen in the small series available for
<taxonomicName id="4C5F4D22FFE7956EFEDAF908F59BF95A" authorityName="Minkina" authorityYear="2020" box="[332,480,1702,1728]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE7956EFEDAF908F520F95A" box="[332,347,1703,1728]" italics="true" pageId="3" pageNumber="2">L</emphasis>
.
<emphasis id="B92BEAB3FFE7956EFEFEF909F59BF95A" box="[360,480,1702,1728]" italics="true" pageId="3" pageNumber="2">brasiliensis</emphasis>
</taxonomicName>
. Variation was seen in the size and distinctness of dorsal and ventral punctation. Some of this apparent variability is related to the amount of encrustation over the structure. Although not all were dissected, it appears the Brazilian specimens are all female, while the
<collectingCountry id="F3487631FFE7956EFC21F946F024F899" box="[951,1119,1769,1795]" name="French Guiana" pageId="3" pageNumber="2">French Guiana</collectingCountry>
specimens are all male. The Brazilian specimens are slightly larger than those from
<collectingCountry id="F3487631FFE7956EFCBEF8A5F7AAF8BE" box="[808,977,1802,1828]" name="French Guiana" pageId="3" pageNumber="2">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE79569FF56F89BF081F967" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8BE036A1FFE7956EFF56F89BF1DAF8EA" blockId="3.[192,1441,1844,1905]" pageId="3" pageNumber="2">
<emphasis id="B92BEAB3FFE7956EFF56F89BF5D8F8D4" bold="true" box="[192,419,1844,1870]" pageId="3" pageNumber="2">Material examined.</emphasis>
(7)
<materialsCitation id="3B373CFCFFE7956EFE44F89BF686F8EA" collectingDate="1992-04-27" collectionCode="FSCA" collectorName="U. Schmitz" country="Brazil" location="Fazenda Rancho Grande" municipality="Label" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Rondonia">
<collectingMunicipality id="6B84ACDBFFE7956EFE44F89BF674F8D4" box="[466,527,1844,1870]" pageId="3" pageNumber="2">Label</collectingMunicipality>
data:
<emphasis id="B92BEAB3FFE7956EFDC2F89BF742F8D4" bold="true" box="[596,825,1844,1870]" pageId="3" pageNumber="2">
<collectingCountry id="F3487631FFE7956EFDC2F89BF6C1F8D4" box="[596,698,1844,1870]" name="Brazil" pageId="3" pageNumber="2">BRAZIL</collectingCountry>
:
<collectingRegion id="499BF843FFE7956EFD53F89BF742F8D4" box="[709,825,1844,1870]" country="Brazil" name="Rondonia" pageId="3" pageNumber="2">Rondônia</collectingRegion>
</emphasis>
:
<locationDeviation id="9A522F83FFE7956EFCD0F89BF03AF8D4" box="[838,1089,1844,1870]" location="Ariquemes" pageId="3" pageNumber="2">
<quantity id="4CA79B44FFE7956EFCD0F89BF7F6F8D4" box="[838,909,1844,1870]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="2" unit="km" value="62.0">62 km</quantity>
SW Ariquemes
</locationDeviation>
, near
<location id="8E80607AFFE7956EFB13F89BF1E7F8D4" LSID="urn:lsid:plazi:treatment:03F687B7FFE79569FF56FB75F081F967:8E80607AFFE7956EFB13F89BF1E7F8D4" box="[1157,1436,1844,1870]" country="Brazil" municipality="Label" name="Fazenda Rancho Grande" pageId="3" pageNumber="2" stateProvince="Rondonia">Fazenda Rancho Grande</location>
,
<date id="FFE11061FFE7956EFF57F8F9F538F8EA" box="[193,323,1878,1904]" pageId="3" pageNumber="2" value="1992-04-27">
<collectingDate id="EFA5E989FFE7956EFF57F8F9F538F8EA" box="[193,323,1878,1904]" pageId="3" pageNumber="2" value="1992-04-27">27-IV-1992</collectingDate>
</date>
,
<collectorName id="26AA5377FFE7956EFEC6F8F9F5B5F8EA" box="[336,462,1878,1904]" pageId="3" pageNumber="2">U. Schmitz</collectorName>
,
<collectingMethod id="521E4EB6FFE7956EFE4DF8F9F6FCF8EA" box="[475,647,1878,1904]" pageId="3" pageNumber="2">black light trap</collectingMethod>
(1
<collectionCode id="ED4EAE64FFE7956EFD39F8F9F68DF8EA" box="[687,758,1878,1904]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CFCFFE7956EFC9CF8F9F1DAF8EA" box="[778,1441,1878,1905]" collectingDate="1996-12-03" collectingDateMax="1996-12-15" collectingDateMin="1996-12-03" collectionCode="FSCA" collectorName="J. E. Eger" country="Brazil" location="Fazenda Rancho Grande" municipality="Label" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Rondonia">
same locality,
<date id="FFE11061FFE7956EFC3CF8F9F029F8EB" box="[938,1106,1878,1905]" pageId="3" pageNumber="2" value="1996-12-03" valueMax="1996-12-15" valueMin="1996-12-03">
<collectingDate id="EFA5E989FFE7956EFC3CF8F9F029F8EB" box="[938,1106,1878,1905]" pageId="3" pageNumber="2" value="1996-12-03" valueMax="1996-12-15" valueMin="1996-12-03">315-XII-1996</collectingDate>
</date>
,
<collectorName id="26AA5377FFE7956EFBC9F8F9F0B2F8EA" box="[1119,1225,1878,1904]" pageId="3" pageNumber="2">J. E. Eger</collectorName>
,
<collectingMethod id="521E4EB6FFE7956EFB43F8F9F1FAF8EA" box="[1237,1409,1878,1904]" pageId="3" pageNumber="2">black light trap</collectingMethod>
(1
</materialsCitation>
</paragraph>
<caption id="DF206629FFE09569FF56F9A1F014F9F1" ID-DOI="http://doi.org/10.5281/zenodo.12808373" ID-Zenodo-Dep="12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,279,1550,1576]" targetBox="[192,1440,213,1520]" targetPageId="4" targetType="figure">
<paragraph id="8BE036A1FFE09569FF56F9A1F014F9F1" blockId="4.[191,1440,1550,1643]" pageId="4" pageNumber="3">
<emphasis id="B92BEAB3FFE09569FF56F9A1F535F9B2" bold="true" box="[192,334,1550,1576]" pageId="4" pageNumber="3">Figures 18.</emphasis>
<taxonomicName id="4C5F4D22FFE09569FEC5F9A1F589F9B2" authorityName="Howden" authorityYear="2003" box="[339,498,1550,1576]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAB3FFE09569FEC5F9A1F589F9B2" box="[339,498,1550,1576]" italics="true" pageId="4" pageNumber="3">Leptorhyparus</emphasis>
</taxonomicName>
characters.
<emphasis id="B92BEAB3FFE09569FDE2F9A1F6D2F9B2" bold="true" box="[628,681,1550,1576]" pageId="4" pageNumber="3">13)</emphasis>
Head and pronotum.
<emphasis id="B92BEAB3FFE09569FC0CF9A1F7C9F9B2" bold="true" box="[922,946,1550,1576]" pageId="4" pageNumber="3">1)</emphasis>
<taxonomicName id="4C5F4D22FFE09569FC2EF9A0F030F9B2" authorityName="Minkina" authorityYear="2020" box="[952,1099,1550,1576]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE09569FC2EF9A0F7BCF9B2" box="[952,967,1551,1576]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FC45F9A1F030F9B2" box="[979,1099,1550,1576]" italics="true" pageId="4" pageNumber="3">brasiliensis</emphasis>
</taxonomicName>
.
<emphasis id="B92BEAB3FFE09569FBC1F9A1F014F9B2" bold="true" box="[1111,1135,1550,1576]" pageId="4" pageNumber="3">2)</emphasis>
<taxonomicName id="4C5F4D22FFE09569FBE2F9A0F0CCF9B2" authorityName="Howden" authorityYear="2003" box="[1140,1207,1550,1576]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE09569FBE2F9A0F0F8F9B2" box="[1140,1155,1551,1576]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FB18F9A1F0CCF9B2" box="[1166,1207,1550,1576]" italics="true" pageId="4" pageNumber="3">gilli</emphasis>
</taxonomicName>
.
<emphasis id="B92BEAB3FFE09569FB54F9A1F0A0F9B2" bold="true" box="[1218,1243,1550,1576]" pageId="4" pageNumber="3">3)</emphasis>
<taxonomicName id="4C5F4D22FFE09569FB76F9A0F11BF9B2" authorityName="Minkina" authorityYear="2020" box="[1248,1376,1550,1576]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B92BEAB3FFE09569FB76F9A0F094F9B2" box="[1248,1263,1551,1576]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FB6CF9A0F11BF9B2" box="[1274,1376,1551,1576]" italics="true" pageId="4" pageNumber="3">peruanus</emphasis>
</taxonomicName>
.
<emphasis id="B92BEAB3FFE09569FAFAF9A1F1DBF9B2" bold="true" box="[1388,1440,1550,1576]" pageId="4" pageNumber="3">45)</emphasis>
Elytra and pygidium, caudal view.
<emphasis id="B92BEAB3FFE09569FDADF980F628F9D3" bold="true" box="[571,595,1583,1609]" pageId="4" pageNumber="3">4)</emphasis>
<taxonomicName id="4C5F4D22FFE09569FDCFF99FF697F9D3" authorityName="Minkina" authorityYear="2020" box="[601,748,1583,1610]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE09569FDCFF99FF613F9D3" box="[601,616,1584,1609]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FDE2F980F697F9D3" box="[628,748,1583,1609]" italics="true" pageId="4" pageNumber="3">brasiliensis</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAB3FFE09569FD6EF980F76BF9D3" bold="true" box="[760,784,1583,1609]" pageId="4" pageNumber="3">5)</emphasis>
<taxonomicName id="4C5F4D22FFE09569FC80F99FF7EDF9D3" authorityName="Minkina" authorityYear="2020" box="[790,918,1584,1610]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B92BEAB3FFE09569FC80F99FF75EF9D3" box="[790,805,1584,1609]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FCA6F99FF7EDF9D3" box="[816,918,1584,1609]" italics="true" pageId="4" pageNumber="3">peruanus</emphasis>
</taxonomicName>
.
<emphasis id="B92BEAB3FFE09569FC34F980F7B7F9D3" bold="true" box="[930,972,1583,1609]" pageId="4" pageNumber="3">68</emphasis>
) Body parts of
<taxonomicName id="4C5F4D22FFE09569FBE3F99FF172F9D3" authorityName="Minkina" authorityYear="2020" box="[1141,1289,1583,1610]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE09569FBE3F99FF0FFF9D3" box="[1141,1156,1584,1609]" italics="true" pageId="4" pageNumber="3">L</emphasis>
.
<emphasis id="B92BEAB3FFE09569FB07F980F172F9D3" box="[1169,1289,1583,1609]" italics="true" pageId="4" pageNumber="3">brasiliensis</emphasis>
</taxonomicName>
, from French Guiana.
<emphasis id="B92BEAB3FFE09569FE88F9FEF54DF9F1" bold="true" box="[286,310,1617,1643]" pageId="4" pageNumber="3">6)</emphasis>
Epipharynx.
<emphasis id="B92BEAB3FFE09569FE5AF9FEF59FF9F1" bold="true" box="[460,484,1617,1643]" pageId="4" pageNumber="3">7)</emphasis>
Male genitalia, caudal view.
<emphasis id="B92BEAB3FFE09569FCB4F9FEF741F9F1" bold="true" box="[802,826,1617,1643]" pageId="4" pageNumber="3">8)</emphasis>
Male genitalia, lateral view.
</paragraph>
</caption>
<paragraph id="8BE036A1FFE09569FF56F90FF081F967" blockId="4.[190,1440,1696,1790]" pageId="4" pageNumber="3">
<collectionCode id="ED4EAE64FFE09569FF56F90FF56AF920" box="[192,273,1696,1722]" pageId="4" pageNumber="3">CEMT</collectionCode>
);
<materialsCitation id="3B373CFCFFE09569FEB4F90EF08EF967" collectingDate="1997-11-04" collectingDateMax="2016-03-10" collectingDateMin="1997-11-04" collectionCode="FSCA, CSCA, CMNC" collectorName="J. E. Eger &amp; J. B. Heppner" country="Ecuador" elevation="2" latitude="-0.6745" location="Yasuni Research Station" longLatPrecision="55" longitude="-76.397" pageId="4" pageNumber="3" specimenCount="1" stateProvince="Kaw">
same locality,
<date id="FFE11061FFE09569FE2CF90FF622F920" box="[442,601,1696,1722]" pageId="4" pageNumber="3" value="1997-11-04" valueMax="1997-11-16" valueMin="1997-11-04">
<collectingDate id="EFA5E989FFE09569FE2CF90FF622F920" box="[442,601,1696,1722]" pageId="4" pageNumber="3" value="1997-11-04" valueMax="1997-11-16" valueMin="1997-11-04">416-XI-1997</collectingDate>
</date>
,
<collectorName id="26AA5377FFE09569FDF4F90EF6BFF921" box="[610,708,1697,1723]" pageId="4" pageNumber="3">J. E. Eger</collectorName>
,
<collectingMethod id="521E4EB6FFE09569FD5AF90FF709F921" box="[716,882,1696,1723]" pageId="4" pageNumber="3">black light trap</collectingMethod>
(1
<collectionCode id="ED4EAE64FFE09569FC04F90FF7A2F920" box="[914,985,1696,1722]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
).
<emphasis id="B92BEAB3FFE09569FC71F90FF015F920" bold="true" box="[999,1134,1696,1722]" pageId="4" pageNumber="3">
<collectingCountry id="F3487631FFE09569FC71F90FF015F920" box="[999,1134,1696,1722]" name="Ecuador" pageId="4" pageNumber="3">ECUADOR</collectingCountry>
</emphasis>
:
<emphasis id="B92BEAB3FFE09569FBECF90FF09BF920" bold="true" box="[1146,1248,1696,1722]" pageId="4" pageNumber="3">Orellana</emphasis>
:
<location id="8E80607AFFE09569FB7AF90FF574F946" LSID="urn:lsid:plazi:treatment:03F687B7FFE79569FF56FB75F081F967:8E80607AFFE09569FB7AF90FF574F946" country="Ecuador" latitude="-0.6745" longLatPrecision="55" longitude="-76.397" name="Yasuni Research Station" pageId="4" pageNumber="3" stateProvince="Kaw">Yasuni Research Station</location>
,
<geoCoordinate id="EE6B5066FFE09569FE8FF96DF507F947" box="[281,380,1730,1757]" degrees="0.6745" direction="south" orientation="latitude" pageId="4" pageNumber="3" precision="5" value="-0.6745">0.6745°S</geoCoordinate>
,
<geoCoordinate id="EE6B5066FFE09569FE13F96DF589F946" box="[389,498,1730,1756]" degrees="76.397" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="55" value="-76.397">76.397°W</geoCoordinate>
,
<collectingMethod id="521E4EB6FFE09569FE6DF96DF617F946" box="[507,620,1730,1756]" pageId="4" pageNumber="3">Sante trap</collectingMethod>
,
<quantity id="4CA79B44FFE09569FDE3F96DF6E0F946" box="[629,667,1730,1756]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="3" unit="m" value="2.0">
<elevation id="0072D192FFE09569FDE3F96DF6E0F946" box="[629,667,1730,1756]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="3" unit="m" value="2.0">2m</elevation>
</quantity>
, lower jar,
<date id="FFE11061FFE09569FC86F96DF7BCF946" box="[784,967,1730,1756]" pageId="4" pageNumber="3" value="2008-07-12" valueMax="2008-07-16" valueMin="2008-07-12">
<collectingDate id="EFA5E989FFE09569FC86F96DF7BCF946" box="[784,967,1730,1756]" pageId="4" pageNumber="3" value="2008-07-12" valueMax="2008-07-16" valueMin="2008-07-12">1216-VII-2008</collectingDate>
</date>
, LASAM Team (1
<collectionCode id="ED4EAE64FFE09569FB04F96DF0A6F946" box="[1170,1245,1730,1756]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="4" pageNumber="3" type="Museum">CSCA</collectionCode>
).
<emphasis id="B92BEAB3FFE09569FB7BF96EF540F967" bold="true" pageId="4" pageNumber="3">
<collectingCountry id="F3487631FFE09569FB7BF96EF486F967" name="French Guiana" pageId="4" pageNumber="3">FRENCH GUIANA</collectingCountry>
:
<collectingRegion id="499BF843FFE09569FE9EF94CF540F967" box="[264,315,1763,1789]" country="French Guiana" name="Kaw" pageId="4" pageNumber="3">Kaw</collectingRegion>
</emphasis>
: Camp Amazone, Kaw Mtn.,
<date id="FFE11061FFE09569FD13F94CF75EF967" box="[645,805,1763,1789]" pageId="4" pageNumber="3" value="2016-03-06" valueMax="2016-03-10" valueMin="2016-03-06">
<collectingDate id="EFA5E989FFE09569FD13F94CF75EF967" box="[645,805,1763,1789]" pageId="4" pageNumber="3" value="2016-03-06" valueMax="2016-03-10" valueMin="2016-03-06">
610-III-
<quantity id="4CA79B44FFE09569FD7CF94CF75EF967" box="[746,805,1763,1789]" metricMagnitude="6" metricUnit="m" metricValue="2.016307" pageId="4" pageNumber="3" unit="m" value="2016307.0">2016</quantity>
</collectingDate>
</date>
, 307m,
<collectorName id="26AA5377FFE09569FCEDF94BF06BF964" box="[891,1040,1764,1790]" pageId="4" pageNumber="3">J. B. Heppner</collectorName>
(1
<collectionCode id="ED4EAE64FFE09569FBA2F94CF0F2F967" box="[1076,1161,1763,1789]" pageId="4" pageNumber="3">CMNC</collectionCode>
; 3
<collectionCode id="ED4EAE64FFE09569FB31F94CF095F967" box="[1191,1262,1763,1789]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>