treatments-xml/data/02/68/12/02681254FFDE28447A8BBA26CB15FB4B.xml

<document id="0284326F6A2B2A6A2BB65575498CED69" ID-CLB-Dataset="8521" ID-DOI="10.11646/zootaxa.4915.3.7" ID-GBIF-Dataset="9213ee25-a9f3-49f6-bf72-8cb456c14b03" ID-ISSN="1175-5326" ID-Zenodo-Dep="4456586" ID-ZooBank="4B0B7EDD-7BED-4C23-948A-7C9467C52C65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611305335637" checkinUser="plazi" docAuthor="Hitchmough, Rodney A., Nielsen, Stuart V., Lysaght, Judith A. &amp; Bauer, Aaron M." docDate="2021" docId="02681254FFDE28447A8BBA26CB15FB4B" docLanguage="en" docName="zootaxa.4915.3.7.pdf" docOrigin="Zootaxa 4915 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naultinus flavirictus Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer 2021, sp. nov." docType="treatment" docUuid="52E975BD-A361-4B5D-B247-0958768B48F4" docUuidSource="ZooBank" docVersion="8" lastPageNumber="398" masterDocId="FE516A2CFFDC284D7A1CBB01CA4EFFA2" masterDocTitle="A new species of Naultinus from the Te Paki area, northern New Zealand" masterLastPageNumber="400" masterPageNumber="389" pageNumber="391" updateTime="1698925167387" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="38240CB9DD5026EDB39C6E4A6BB5960A">A new species of Naultinus from the Te Paki area, northern New Zealand</mods:title>
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<mods:namePart id="3ABBB251834D0265F38DE7A5CBC16879">Hitchmough, Rodney A.</mods:namePart>
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<mods:namePart id="81B56B6984C0690115E9117942EFF362">Nielsen, Stuart V.</mods:namePart>
<mods:affiliation id="1781C363FE4801A9F1EA03FA684D74C6">Florida Museum of Natural History, University of Florida, Gainesville, FL 32601, USA &amp; Department of Biological Sciences, Marquette University, Milwaukee, WI 53233, USA</mods:affiliation>
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<mods:namePart id="B41D0D6CF31CB02B18FAC6166A4ACA9C">Lysaght, Judith A.</mods:namePart>
<mods:affiliation id="FD0A310B4636DDDE8E167DE44356C840">Tea Tree Gully, Adelaide. judith. lysaght @ gmail. com</mods:affiliation>
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<mods:namePart id="532C495468FD9F4869DDB39550D1F4A0">Bauer, Aaron M.</mods:namePart>
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<mods:affiliation id="BC22CC523D525574574CFC0B15F9FF27">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, PA 19085, USA. aaron. bauer @ villanova. edu; https: // orcid. org / 0000 - 0001 - 6839 - 8025</mods:affiliation>
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<paragraph id="8A7EA342FFDE284F7A8BBA26CBB8FEE3" blockId="2.[151,502,295,357]" box="[151,502,295,322]" pageId="2" pageNumber="391">
<heading id="D136142EFFDE284F7A8BBA26CBB8FEE3" bold="true" box="[151,502,295,322]" fontSize="11" level="1" pageId="2" pageNumber="391" reason="1">
<emphasis id="B8B57F50FFDE284F7A8BBA26CBB8FEE3" bold="true" box="[151,502,295,322]" pageId="2" pageNumber="391">
<taxonomicName id="4DC1D8C1FFDE284F7A8BBA26CBDFFEE0" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[151,401,295,322]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDE284F7A8BBA26CBDFFEE0" bold="true" box="[151,401,295,322]" italics="true" pageId="2" pageNumber="391">Naultinus flavirictus</emphasis>
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<taxonomicNameLabel id="A386C22BFFDE284F7B85BA26CBB8FEE3" box="[409,502,295,321]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8A7EA342FFDE284F7A8BBA4ACAA5FEC7" blockId="2.[151,502,295,357]" box="[151,235,331,357]" pageId="2" pageNumber="391">
(
<figureCitation id="12FABFC7FFDE284F7ABCBA4ACAAEFEC7" box="[160,224,331,357]" captionStart="FIGURE 1" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[158,1429,181,1905]" captionTargetId="figure-22@6.[158,1429,181,1920]" captionTargetPageId="6" captionText="FIGURE 1. Adult female holotype of Naultinus flavirictus sp. nov. RE003322 (original field number G.789): A.) dorsal, B.) ventral, and C.) lateral views. Photographs by J-C. Stahl. Scale bar = 1cm." figureDoi="http://doi.org/10.5281/zenodo.4456588" httpUri="https://zenodo.org/record/4456588/files/figure.png" pageId="2" pageNumber="391">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8A7EA342FFDE284F7A8BBA91C8E6FE0A" blockId="2.[151,680,400,454]" box="[151,680,400,424]" pageId="2" pageNumber="391">
<taxonomicName id="4DC1D8C1FFDE284F7A8BBA91CB7BFE05" box="[151,309,400,424]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="grayi">
<emphasis id="B8B57F50FFDE284F7A8BBA91CB7BFE05" box="[151,309,400,424]" italics="true" pageId="2" pageNumber="391">Naultinus grayi</emphasis>
</taxonomicName>
: (part) 
<bibRefCitation id="EE50DEB3FFDE284F7B9EBA91C8EBFE0A" author="Robb, J. &amp; Hitchmough, R. A." box="[386,677,400,424]" pageId="2" pageNumber="391" pagination="189 - 200" refId="ref8947" refString="Robb, J. &amp; Hitchmough, R. A. (1980) Review of the genus Naultinus Gray (Reptilia: Gekkonidae). Records of the Auckland Institute and Museum, 16, 189 - 200." type="journal article" year="1980">Robb &amp; Hitchmough (1980)</bibRefCitation>
.
</paragraph>
<paragraph id="8A7EA342FFDE284F7A8BBAAFC8ECFE64" blockId="2.[151,680,400,454]" box="[151,674,430,454]" pageId="2" pageNumber="391">
<taxonomicName id="4DC1D8C1FFDE284F7A8BBAAFCAB4FE64" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[151,250,430,454]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F7A8BBAAFCAB4FE64" box="[151,250,430,454]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
“North Cape”: 
<bibRefCitation id="EE50DEB3FFDE284F7B85BAAFC8D0FE64" author="Hitchmough, R. A. &amp; Bull, L. &amp; Cromarty, P. &amp; Comps" box="[409,670,430,454]" pageId="2" pageNumber="391" refId="ref8330" refString="Hitchmough, R. A., Bull, L. &amp; Cromarty, P. (Comps.) (2007) New Zealand threat classification system lists 2005. Department of Conservation, Wellington, 194 pp." type="book" year="2007">
Hitchmough 
<emphasis id="B8B57F50FFDE284F7802BAAEC81CFE64" box="[542,594,430,454]" italics="true" pageId="2" pageNumber="391">et al.</emphasis>
(2007)
</bibRefCitation>
.
</paragraph>
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<paragraph id="8A7EA342FFDE284F7A8BBAF4C9DCFD91" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
<materialsCitation id="3AA9A91FFFDE284F7A8BBAF4C9DCFD91" ID-GBIF-Occurrence="3028581302" accessionNumber="RE003322" collectingDate="1972-05" collectionCode="MONZ" collectorName="Te Paki &amp; A. H. Whitaker" country="New Zealand" location="Northland" pageId="2" pageNumber="391" specimenCount="2" stateProvince="Northland" typeStatus="holotype">
<emphasis id="B8B57F50FFDE284F7A8BBAF4CB45FDAD" bold="true" box="[151,267,501,527]" pageId="2" pageNumber="391">
<typeStatus id="557A1DE0FFDE284F7A8BBAF4CB48FDAD" box="[151,262,501,527]" pageId="2" pageNumber="391" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult female 
<typeStatus id="557A1DE0FFDE284F7BB7BAF5C843FDAC" box="[427,525,500,526]" pageId="2" pageNumber="391" type="holotype">holotype</typeStatus>
<collectionCode id="ECD03B87FFDE284F7808BAF4C828FDAD" box="[532,614,501,527]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="2" pageNumber="391" type="Museum">MONZ</collectionCode>
RE003322, original field number G.789 (
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), collected from 
<collectorName id="2734C694FFDE284F7F5DBAF4CFD6FDAC" box="[1345,1432,500,527]" pageId="2" pageNumber="391">Te Paki</collectorName>
, 
<collectingRegion id="48056DA0FFDE284F7A8BB919CB45FD90" box="[151,267,536,562]" country="New Zealand" name="Northland" pageId="2" pageNumber="391">Northland</collectingRegion>
, 
<collectingCountry id="F2D6E3D2FFDE284F7B0BB918CBFEFD90" box="[279,432,536,563]" name="New Zealand" pageId="2" pageNumber="391">New Zealand</collectingCountry>
, collected by 
<collectorName id="2734C694FFDE284F7854B918C8BAFD90" box="[584,756,536,563]" pageId="2" pageNumber="391">A. H. Whitaker</collectorName>
in 
<date id="FE7F8582FFDE284F7904B918C9C3FD91" box="[792,909,537,563]" pageId="2" pageNumber="391" value="1972-05">
<collectingDate id="EE3B7C6AFFDE284F7904B918C9C3FD91" box="[792,909,537,563]" pageId="2" pageNumber="391" value="1972-05">May 1972</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8A7EA342FFDE284F7ADBB93CC84DFD61" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
<materialsCitation id="3AA9A91FFFDE284F7ADBB93CC9BFFDD8" ID-GBIF-Occurrence="3028581301" collectingDate="1986-08" collectionCode="MONZ" country="New Zealand" location="Te Kao District" municipality="Conrad Hepers" pageId="2" pageNumber="391" specimenCode="RAH195" specimenCount="1" stateProvince="Northland" typeStatus="paratype">
<emphasis id="B8B57F50FFDE284F7ADBB93CCB08FDF5" bold="true" box="[199,326,573,599]" pageId="2" pageNumber="391">
<typeStatus id="557A1DE0FFDE284F7ADBB93CCB0FFDF5" box="[199,321,573,599]" pageId="2" pageNumber="391" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Two adults, one male, 
<collectionCode id="ECD03B87FFDE284F7852B93CC8EEFDF5" box="[590,672,573,599]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="2" pageNumber="391" type="Museum">MONZ</collectionCode>
RE.005611 (original field number 
<specimenCode id="DA670B39FFDE284F7E33B93DCED6FDF5" box="[1071,1176,572,599]" collectionCode="RAH" pageId="2" pageNumber="391">RAH195</specimenCode>
), collected from the 
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, 
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, 
<collectingCountry id="F2D6E3D2FFDE284F7BABB960C81DFDD8" box="[439,595,608,635]" name="New Zealand" pageId="2" pageNumber="391">New Zealand</collectingCountry>
, by 
<collectingMunicipality id="6A1A3938FFDE284F789BB961C97DFDD9" box="[647,819,608,635]" pageId="2" pageNumber="391">Conrad Hepers</collectingMunicipality>
in 
<date id="FE7F8582FFDE284F7947B960C9BFFDD8" box="[859,1009,608,635]" pageId="2" pageNumber="391" value="1986-08">
<collectingDate id="EE3B7C6AFFDE284F7947B960C9BFFDD8" box="[859,1009,608,635]" pageId="2" pageNumber="391" value="1986-08">August 1986</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3AA9A91FFFDE284F79E3B961CBB0FD61" ID-GBIF-Occurrence="3028581303" collectingDate="1986-08" collectionCode="MONZ" collectorName="Te Kao &amp; Conrad Hepers" country="New Zealand" location="Northland" pageId="2" pageNumber="391" specimenCode="RAH254" specimenCount="1" stateProvince="Northland" typeStatus="paratype">
and one female, 
<collectionCode id="ECD03B87FFDE284F7EDDB960CF5DFDD9" box="[1217,1299,609,635]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="2" pageNumber="391" type="Museum">MONZ</collectionCode>
RE.005609 (original field number 
<specimenCode id="DA670B39FFDE284F7B8AB985CBB0FD3D" box="[406,510,644,671]" collectionCode="RAH" pageId="2" pageNumber="391">RAH254</specimenCode>
; captive offspring of a female collected from 
<collectorName id="2734C694FFDE284F79E1B984CE1FFD3D" box="[1021,1105,645,671]" pageId="2" pageNumber="391">Te Kao</collectorName>
, 
<collectingRegion id="48056DA0FFDE284F7E40B985CE9EFD3C" box="[1116,1232,644,670]" country="New Zealand" name="Northland" pageId="2" pageNumber="391">Northland</collectingRegion>
, 
<collectingCountry id="F2D6E3D2FFDE284F7EC0B984CF3BFD3C" box="[1244,1397,644,671]" name="New Zealand" pageId="2" pageNumber="391">New Zealand</collectingCountry>
, by 
<collectorName id="2734C694FFDE284F7A8BB9A9CB0EFD61" box="[151,320,680,707]" pageId="2" pageNumber="391">Conrad Hepers</collectorName>
in 
<date id="FE7F8582FFDE284F7B7EB9A8CBB8FD61" box="[354,502,680,707]" pageId="2" pageNumber="391" value="1986-08">
<collectingDate id="EE3B7C6AFFDE284F7B7EB9A8CBB8FD61" box="[354,502,680,707]" pageId="2" pageNumber="391" value="1986-08">August 1986</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2DBF0C9FFDE284F7ADBB9CCCEB4FC8C" pageId="2" pageNumber="391" type="etymology">
<paragraph id="8A7EA342FFDE284F7ADBB9CCCEB4FC8C" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
<emphasis id="B8B57F50FFDE284F7ADBB9CCCB00FD45" bold="true" box="[199,334,717,743]" pageId="2" pageNumber="391">Etymology.</emphasis>
<taxonomicName id="4DC1D8C1FFDE284F7B44B9CDC80CFD44" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[344,578,716,742]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDE284F7B44B9CDC80CFD44" box="[344,578,716,742]" italics="true" pageId="2" pageNumber="391">Naultinus flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDE284F7850B9CCC8E7FD45" bold="true" box="[588,681,717,743]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A386C22BFFDE284F7850B9CCC8E7FD45" box="[588,681,717,743]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(formed from the Lain 
<emphasis id="B8B57F50FFDE284F79D8B9CDCE49FD44" box="[964,1031,716,742]" italics="true" pageId="2" pageNumber="391">flavus</emphasis>
= yellow + 
<emphasis id="B8B57F50FFDE284F7E81B9CCCE93FD44" box="[1181,1245,717,742]" italics="true" pageId="2" pageNumber="391">rictus</emphasis>
= gaping of the mouth) is named for the continuation of the yellow-orange colouring of the jaws and tongue onto the external corners of the mouth, which is diagnostic for the species. The epithet is formed as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C2DBF0C9FFDE284F7ADBB839CEC8F958" pageId="2" pageNumber="391" type="diagnosis">
<paragraph id="8A7EA342FFDE284F7ADBB839CF5AFC38" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
<emphasis id="B8B57F50FFDE284F7ADBB839CB0CFCF0" bold="true" box="[199,322,824,850]" pageId="2" pageNumber="391">Diagnosis.</emphasis>
<taxonomicName id="4DC1D8C1FFDE284F7B57B839C87DFCF0" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[331,563,824,850]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDE284F7B57B839C87DFCF0" box="[331,563,824,850]" italics="true" pageId="2" pageNumber="391">Naultinus flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDE284F7821B838C8D7FCF1" bold="true" box="[573,665,825,851]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A386C22BFFDE284F7821B838C8D7FCF1" box="[573,665,825,851]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a small 
<taxonomicName id="4DC1D8C1FFDE284F7907B839C9C6FCF0" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[795,904,824,850]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F7907B839C9C6FCF0" box="[795,904,824,850]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
(maximum SVL ~ 
<quantity id="4D390EA7FFDE284F7E7DB838CEF6FCF1" box="[1121,1208,825,851]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="391" unit="mm" value="70.0">70 mm</quantity>
), with a short, deep snout, fine granular body scales, flattened scales on the dorsum of the snout, rostral scale divided by a median crease, pale lavender mouth lining, and yellow-orange jaws, tongue and external corners of the mouth.
</paragraph>
<paragraph id="8A7EA342FFDE284F7ADBB8A4C8B1FB19" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
It may be distinguished from all other North Island 
<taxonomicName id="4DC1D8C1FFDE284F790DB8A5C930FC1C" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[785,894,932,958]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F790DB8A5C930FC1C" box="[785,894,932,958]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
species by the pale lavender mouth lining, with yellow-orange jaws and tongue, and by the colour of the jaws extending to the external corners of the mouth, with similar pigment present around the nostrils. A central crease dividing rostral in two is typical in this species, but rare in 
<taxonomicName id="4DC1D8C1FFDE284F7AA8BF10CB56FB88" authorityName="Bell" authorityYear="1843" box="[180,280,1041,1066]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFDE284F7AA8BF10CB56FB88" box="[180,280,1041,1066]" italics="true" pageId="2" pageNumber="391">N. grayii</emphasis>
</taxonomicName>
and 
<emphasis id="B8B57F50FFDE284F7B52BF10CB85FB88" box="[334,459,1040,1066]" italics="true" pageId="2" pageNumber="391">
<taxonomicName id="4DC1D8C1FFDE284F7B52BF10CB89FB88" authorityName="Gray" authorityYear="1842" box="[334,455,1040,1066]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="elegans">N. elegans</taxonomicName>
,
</emphasis>
and partial or absent in other 
<taxonomicName id="4DC1D8C1FFDE284F7907BF11C9C6FB88" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[795,904,1040,1066]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F7907BF11C9C6FB88" box="[795,904,1040,1066]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
. It is distinguished from 
<taxonomicName id="4DC1D8C1FFDE284F7EBEBF10CF4BFB88" authorityName="Bell" authorityYear="1843" box="[1186,1285,1041,1066]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFDE284F7EBEBF10CF4BFB88" box="[1186,1285,1041,1066]" italics="true" pageId="2" pageNumber="391">N. grayii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4DC1D8C1FFDE284F7F20BF10CA83FBEC" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B8B57F50FFDE284F7F20BF10CA83FBEC" italics="true" pageId="2" pageNumber="391">N. punctatus</emphasis>
</taxonomicName>
by much smaller adult size (to ~ 
<quantity id="4D390EA7FFDE284F7825BF34C8C6FBED" box="[569,648,1077,1103]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="391" unit="mm" value="70.0">70 mm</quantity>
vs ~ 
<quantity id="4D390EA7FFDE284F78DFBF35C95CFBED" box="[707,786,1076,1103]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="391" unit="mm" value="95.0">95 mm</quantity>
SVL), and by secondary bluish colouration of adult males including the entire ventral surface rather than being confined to a lower lateral band. It is further distinguished from 
<taxonomicName id="4DC1D8C1FFDE284F7A8BBF7CCAB4FB34" authorityName="Bell" authorityYear="1843" box="[151,250,1149,1174]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFDE284F7A8BBF7CCAB4FB34" box="[151,250,1149,1174]" italics="true" pageId="2" pageNumber="391">N. grayii</emphasis>
</taxonomicName>
by its proportionally shorter, deeper snout (
<figureCitation id="12FABFC7FFDE284F78C2BF7DC96DFB35" box="[734,803,1148,1175]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[246,1313,181,1920]" captionTargetId="figure-17@7.[246,1341,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Comparison of head scalation between relevant Naultinus spp.: A.) N. manukanus (MONZ RE.2013); B.) N. stellatus (RE.005617; original field number RAH466); C.) N. flavirictus sp. nov. (holotype); D.) N. elegans (RE.002658; original field number G.102); and E.) N. grayii (RE.002771; original field number G.231). Photos by SVN." figureDoi="http://doi.org/10.5281/zenodo.4456590" httpUri="https://zenodo.org/record/4456590/files/figure.png" pageId="2" pageNumber="391">Fig. 2</figureCitation>
). Like 
<taxonomicName id="4DC1D8C1FFDE284F796CBF7CC99CFB34" authorityName="Bell" authorityYear="1843" box="[880,978,1149,1174]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFDE284F796CBF7CC99CFB34" box="[880,978,1149,1174]" italics="true" pageId="2" pageNumber="391">N. grayii</emphasis>
</taxonomicName>
, but unlike all other 
<taxonomicName id="4DC1D8C1FFDE284F7EABBF7DCF6AFB34" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[1207,1316,1148,1174]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F7EABBF7DCF6AFB34" box="[1207,1316,1148,1174]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
, the scales on the dorsal surface of the snout are flattened (
<figureCitation id="12FABFC7FFDE284F78B0BFA1C8BCFB19" box="[684,754,1184,1211]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[246,1313,181,1920]" captionTargetId="figure-17@7.[246,1341,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Comparison of head scalation between relevant Naultinus spp.: A.) N. manukanus (MONZ RE.2013); B.) N. stellatus (RE.005617; original field number RAH466); C.) N. flavirictus sp. nov. (holotype); D.) N. elegans (RE.002658; original field number G.102); and E.) N. grayii (RE.002771; original field number G.231). Photos by SVN." figureDoi="http://doi.org/10.5281/zenodo.4456590" httpUri="https://zenodo.org/record/4456590/files/figure.png" pageId="2" pageNumber="391">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8A7EA342FFDE284F7ADBBFC5CEC8F958" blockId="2.[151,1437,500,2039]" pageId="2" pageNumber="391">
The new species may be distinguished from most South Island 
<taxonomicName id="4DC1D8C1FFDE284F799CBFC5C9A3FB7C" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[896,1005,1220,1246]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDE284F799CBFC5C9A3FB7C" box="[896,1005,1220,1246]" italics="true" pageId="2" pageNumber="391">Naultinus</emphasis>
</taxonomicName>
by its fine, granular dorsal body scales (coarser and more conical in South Island species). Among its South Island congeners it is readily distinguished from 
<taxonomicName id="4DC1D8C1FFDE284F7AC9BE0CCBD2FA85" authority="(Fischer)" baseAuthorityName="Fischer" baseAuthorityYear="1881" box="[213,412,1292,1319]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="rudis">
<emphasis id="B8B57F50FFDE284F7AC9BE0CCB7EFA84" box="[213,304,1292,1318]" italics="true" pageId="2" pageNumber="391">N. rudis</emphasis>
(Fischer)
</taxonomicName>
, 
<taxonomicName id="4DC1D8C1FFDE284F7BB6BE0CC818FA84" box="[426,598,1292,1318]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="tuberculatus">
<emphasis id="B8B57F50FFDE284F7BB6BE0CC818FA84" box="[426,598,1292,1318]" italics="true" pageId="2" pageNumber="391">N. tuberculatus</emphasis>
</taxonomicName>
and 
<collectingRegion id="48056DA0FFDE284F7890BE0DC89EFA84" box="[652,720,1292,1318]" country="New Zealand" name="Otago" pageId="2" pageNumber="391">Otago</collectingRegion>
/ 
<collectingRegion id="48056DA0FFDE284F78C4BE0DC904FA84" box="[728,842,1292,1318]" country="New Zealand" name="Southland" pageId="2" pageNumber="391">Southland</collectingRegion>
populations of 
<taxonomicName id="4DC1D8C1FFDE284F79E6BE0CCECCFA84" box="[1018,1154,1293,1318]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="gemmeus">
<emphasis id="B8B57F50FFDE284F79E6BE0CCECCFA84" box="[1018,1154,1293,1318]" italics="true" pageId="2" pageNumber="391">N. gemmeus</emphasis>
</taxonomicName>
by the pale lavender and orange mouth lining. It differs from 
<taxonomicName id="4DC1D8C1FFDE284F7833BE30C89EFAE8" box="[559,720,1328,1354]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="manukanus">
<emphasis id="B8B57F50FFDE284F7833BE30C89EFAE8" box="[559,720,1328,1354]" italics="true" pageId="2" pageNumber="391">N. manukanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4DC1D8C1FFDE284F791ABE30C92FFAE8" baseAuthorityName="Fischer" baseAuthorityYear="1881" box="[774,865,1328,1354]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="rudis">
<emphasis id="B8B57F50FFDE284F791ABE30C92FFAE8" box="[774,865,1328,1354]" italics="true" pageId="2" pageNumber="391">N. rudis</emphasis>
</taxonomicName>
in the absence of any greatly enlarged dorsal body and head scales (although there is a north–south cline in the number and prominence of these enlarged scales within 
<taxonomicName id="4DC1D8C1FFDE284F7A8BBE78CB77FA30" box="[151,313,1400,1426]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="manukanus">
<emphasis id="B8B57F50FFDE284F7A8BBE78CB77FA30" box="[151,313,1400,1426]" italics="true" pageId="2" pageNumber="391">N. manukanus</emphasis>
</taxonomicName>
, and they may be few in number and easily missed in the northernmost population on Stephens Island; this population is morphologically similar to north-western-most populations of 
<taxonomicName id="4DC1D8C1FFDE284F7E52BE9CCE84FA14" authorityName="Hutton" authorityYear="1872" box="[1102,1226,1436,1462]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B8B57F50FFDE284F7E52BE9CCE84FA14" box="[1102,1226,1436,1462]" italics="true" pageId="2" pageNumber="391">N. stellatus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EE50DEB3FFDE284F7EFEBE9DCA9DFA79" author="Bull, P. C. &amp; Whitaker, A. H." pageId="2" pageNumber="391" pagination="231 - 276" refId="ref7712" refString="Bull, P. C. &amp; Whitaker, A. H. (1975) The amphibians, reptiles, birds and mammals. In: Kuschel, G. (Ed.), Biogeography and Ecology in New Zealand. Junk, The Hague, pp. 231 - 276. https: // doi. org / 10.1007 / 978 - 94 - 010 - 1941 - 5 _ 5" type="book chapter" year="1975">Bull &amp; Whitaker 1975</bibRefCitation>
). It may be distinguished from 
<collectingRegion id="48056DA0FFDE284F782EBEC1C8FEFA78" box="[562,688,1472,1498]" country="New Zealand" name="Canterbury" pageId="2" pageNumber="391">Canterbury</collectingRegion>
populations of 
<taxonomicName id="4DC1D8C1FFDE284F797CBEC0C9A7FA78" box="[864,1001,1473,1498]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="gemmeus">
<emphasis id="B8B57F50FFDE284F797CBEC0C9A7FA78" box="[864,1001,1473,1498]" italics="true" pageId="2" pageNumber="391">N. gemmeus</emphasis>
</taxonomicName>
by its smaller adult size, and by males having bluish belly colouration rather than having the green colouration progressively replaced by brown or grey during growth to maturity. It is most similar morphologically to north-western-most populations of 
<emphasis id="B8B57F50FFDE284F7EF1BD08CF3FF980" box="[1261,1393,1544,1570]" italics="true" pageId="2" pageNumber="391">
<taxonomicName id="4DC1D8C1FFDE284F7EF1BD08CF23F980" authorityName="Hutton" authorityYear="1872" box="[1261,1389,1544,1570]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="stellatus">N. stellatus</taxonomicName>
,
</emphasis>
but distinguished by more subimbricate, slightly flattened granular (vs strongly domed, bead-like, tightly juxtaposed) dorsal body scales, less domed canthal scales (
<figureCitation id="12FABFC7FFDE284F78BBBD51C8A2F9C9" box="[679,748,1616,1643]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[246,1313,181,1920]" captionTargetId="figure-17@7.[246,1341,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Comparison of head scalation between relevant Naultinus spp.: A.) N. manukanus (MONZ RE.2013); B.) N. stellatus (RE.005617; original field number RAH466); C.) N. flavirictus sp. nov. (holotype); D.) N. elegans (RE.002658; original field number G.102); and E.) N. grayii (RE.002771; original field number G.231). Photos by SVN." figureDoi="http://doi.org/10.5281/zenodo.4456590" httpUri="https://zenodo.org/record/4456590/files/figure.png" pageId="2" pageNumber="391">Fig. 2</figureCitation>
), presence of an enlarged patch of supratemporal scales, and rows of enlarged, equal-sized scales each side of the canthal row in 
<taxonomicName id="4DC1D8C1FFDE284F7991BD74CE6EF92C" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[909,1056,1652,1678]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDE284F7991BD74CE6EF92C" box="[909,1056,1652,1678]" italics="true" pageId="2" pageNumber="391">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDE284F7E3ABD74CECEF92D" bold="true" box="[1062,1152,1653,1679]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A386C22BFFDE284F7E3ABD74CECEF92D" box="[1062,1152,1653,1679]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(vs irregular-sized scales bordering enlarged canthal row in 
<taxonomicName id="4DC1D8C1FFDE284F783DBD98C8EEF910" authorityName="Hutton" authorityYear="1872" box="[545,672,1688,1714]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B8B57F50FFDE284F783DBD98C8EEF910" box="[545,672,1688,1714]" italics="true" pageId="2" pageNumber="391">N. stellatus</emphasis>
</taxonomicName>
). 
<taxonomicName id="4DC1D8C1FFDE284F78A5BD99C9EFF910" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[697,929,1688,1714]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDE284F78A5BD99C9EFF910" box="[697,929,1688,1714]" italics="true" pageId="2" pageNumber="391">Naultinus flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDE284F79B5BD98CE4AF911" bold="true" box="[937,1028,1689,1715]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A386C22BFFDE284F79B5BD98CE4AF911" box="[937,1028,1689,1715]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from other populations of 
<taxonomicName id="4DC1D8C1FFDE284F7AEDBDBCCB20F974" authorityName="Hutton" authorityYear="1872" box="[241,366,1724,1750]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="391" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B8B57F50FFDE284F7AEDBDBCCB20F974" box="[241,366,1724,1750]" italics="true" pageId="2" pageNumber="391">N. stellatus</emphasis>
</taxonomicName>
by its uniform green background colour, with or without simple, discrete, irregular-shaped spotted or striped markings rather than large, circular blotches with concentric rings of colour.
</paragraph>
</subSubSection>
<subSubSection id="C2DBF0C9FFDE28447ADBBC05C9C4FB97" lastPageId="9" lastPageNumber="398" pageId="2" pageNumber="391" type="description">
<paragraph id="8A7EA342FFDE284E7ADBBC05CF3BFDC3" blockId="2.[151,1437,500,2039]" lastBlockId="3.[151,1437,150,2013]" lastPageId="3" lastPageNumber="392" pageId="2" pageNumber="391">
<emphasis id="B8B57F50FFDE284F7ADBBC05C85AF8BD" bold="true" box="[199,532,1796,1823]" pageId="2" pageNumber="391">
Description of the 
<typeStatus id="557A1DE0FFDE284F7BBABC04C841F8BD" box="[422,527,1797,1823]" pageId="2" pageNumber="391" type="holotype">holotype</typeStatus>
.
</emphasis>
Body moderately slender, not depressed; 
<quantity id="4D390EA7FFDE284F79F3BC05CE1AF8BD" box="[1007,1108,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="6.709999999999999" pageId="2" pageNumber="391" unit="mm" value="67.1">67.1 mm</quantity>
SVL. HeadL 
<quantity id="4D390EA7FFDE284F7EEFBC05CF16F8BD" box="[1267,1368,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="2" pageNumber="391" unit="mm" value="16.6">16.6 mm</quantity>
(25% SVL), HeadW 
<quantity id="4D390EA7FFDE284F7B58BC28CBE4F8E1" box="[324,426,1833,1859]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="2" pageNumber="391" unit="mm" value="12.1">12.1 mm</quantity>
(73% HeadL) and HeadH 
<quantity id="4D390EA7FFDE284F78FCBC28C977F8E1" box="[736,825,1833,1859]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="391" unit="mm" value="8.8">8.8 mm</quantity>
(53% HL). Head subtriangular with a blunt, rounded snout, clearly demarcated from neck; interorbital/frontal region neither concave nor convex, nasal region without indentation; canthus not inflated. SnEye 
<quantity id="4D390EA7FFDE284F787FBC71C8F4F829" box="[611,698,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="391" unit="mm" value="7.5">7.5 mm</quantity>
(45% HeadL), OrbD 
<quantity id="4D390EA7FFDE284F79AEBC70CE47F829" box="[946,1033,1905,1931]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="391" unit="mm" value="3.4">3.4 mm</quantity>
(45% SnEye). Scales on dorsum of anterior snout approximately 3-4 times the diameter of those on occipital region, loreal scales similar, abruptly decreasing in size at level of anterior edge of orbit; patches of enlarged supratemporal scales (smaller than dorsal snout and loreal scales but distinctly larger than occipital granules). OrbD 20% HeadL; pupil oval, margins smooth. Ear opening very small (
<quantity id="4D390EA7FFDF284E7B67BB96CB9AFF13" box="[379,468,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="392" unit="mm" value="1.1">1.1 mm</quantity>
), oval, obliquely oriented; eye to ear distance greater than diameter of eyes (EyeEar 
<quantity id="4D390EA7FFDF284E7F65BB97CA8DFF77" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="392" unit="mm" value="5.6">5.6 mm</quantity>
(165% OrbD). Rostral almost rectangular, much broader (3.0 mm) than high (
<quantity id="4D390EA7FFDF284E7E36BBBACECCFF77" box="[1066,1154,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="392" unit="mm" value="1.3">1.3 mm</quantity>
) completely divided by a strong rostral crease, contacted dorsally by one large round internasal and two large supranasals, each similar in size to the internasal, contacted laterally by first supralabial, excluded from nostril. Nostrils rounded, laterally oriented, and surrounded by two postnasals on right hand side, three on left, first supralabial and first supranasal. Mental subtriangular, broader (
<quantity id="4D390EA7FFDF284E7B80BA4ACBBBFEC7" box="[412,501,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="392" unit="mm" value="2.8">2.8 mm</quantity>
) than tall (
<quantity id="4D390EA7FFDF284E7872BA4AC887FEC7" box="[622,713,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="392" unit="mm" value="1.6">1.6 mm</quantity>
). First infralabials separated by three postmental, the central one largest and oval, longer than wide, the others wider than long. Anterior infralabials and postmental bordered posteriorly by 3–4 series of enlarged hexagonal chin shields, ¼ to ½ the size of central postmental chin shield, changing abruptly to small, rounded scales beneath throat. Scale rows ventral to posterior infralabials on lateral surface of jaw slightly enlarged and elongated relative to ventral throat scales. Eleven (right) and 10 (left) enlarged infralabial scales, posterior most about 4 times size of rictal scales, 10 (left)–12 (right) enlarged supralabial scales, 7 (left) or 10 (right) supralabials to midpoint of orbit; 36 scale rows between supraciliaries (24 between supraocular skin folds), 11 scale rows across frontal bones at midpoint of orbit. Supraciliaries rounded to somewhat conical posteriorly.
</paragraph>
<paragraph id="8A7EA342FFDF284E7ADBB96ACE0FFB97" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="392">
Dorsal scales small, mostly uniform in size with scattered much smaller scales interspersed, homogenous in shape, bead-like, somewhat flattened, granular. Ventral scales roughly twice diameter of dorsals, smooth, flattened, subimbricate, slightly enlarged in precloacal region. Approximately 99 scale rows around mid-body. No ventrolateral skin folds. Scales on dorsal surface of upper forelimb roughly equal in size to dorsum; on forearm slightly smaller. Scales on preaxial surface of thigh similar to those on dorsum, gradually decreasing in size posteriorly to smaller granules on post-axial surface. Scales on palms and soles smooth, flattened and subimbricate. Fore- and hindlimbs short and moderately robust (Forearm 
<quantity id="4D390EA7FFDF284E78A6B842C95CFCFF" box="[698,786,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="392" unit="mm" value="8.3">8.3 mm</quantity>
; ForeaL/SVL 0.12; Crus 
<quantity id="4D390EA7FFDF284E7E35B842CEC1FCFF" box="[1065,1167,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="3" pageNumber="392" unit="mm" value="11.1">11.1 mm</quantity>
; CrusL/SVL 0.17). Digits relatively elongate, all bearing claws. Claws short and moderately robust basally, recurved to a fine point. Basal portion of digits relatively narrow, approximately twice width of distal phalanges. Distal portion of toe strongly compressed, lamellae continuing in uninterrupted series to claw. Apical pads absent. Relative length of digits of manus: I&lt;II&lt;III&lt;V&lt;IV, and of pes: I&lt;II&lt;V&lt;III&lt;IV; no webbing between digits. Lamellae straight or very weakly bowed, unpaired. Lamellar counts (including all scales&gt; twice size of scales at base of digits) from right (and left) sides 7-10-13-14-11 (6-9-14-14-11) manus and 7-13-14-16-15 (7-13-15-17-15) pes.
</paragraph>
<paragraph id="8A7EA342FFDF284E7ADBBF3EC948FB4B" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="392">
Only scattered pitted scales present in precloacal area (c. 5 rows). Small, weakly developed cloacal spurs with 3 enlarged scales on each side just posterior to vent. Tail original but broken (9.5 +79.7 = 
<quantity id="4D390EA7FFDF284E7E8CBF62CEB7FBDF" box="[1168,1273,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="8.92" pageId="3" pageNumber="392" unit="mm" value="89.2">89.2 mm</quantity>
); 133% snoutvent length, muscular, tapering, round in cross-section. Dorsal caudal scales medium, flat, juxtaposed to weakly subimbricate, roughly circular, arranged irregularly. Surface of tail not segmented. Ventral scales slightly larger than dorsal, smooth, somewhat irregular in shape, imbricate.
</paragraph>
<paragraph id="8A7EA342FFDF284E7ADBBFF3CB65FA47" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="392">
<emphasis id="B8B57F50FFDF284E7ADBBFF3CB91FAAE" bold="true" box="[199,479,1266,1292]" pageId="3" pageNumber="392">Colour in preservative.</emphasis>
Dorsum uniform dull dark green with dorso-lateral rows of irregularly shaped pale cream spots, comprising paired longitudinal streaks behind the eyes, seven spots on each side between the neck and the pelvic region; spots continue along the tail where they become elongated into streaks. Uniform whitish ventro-lateral stripes between insertions of fore and hind limbs separating dark green lateral from pale green ventral surface. Small white blotch of c. 7 scales on dorsum of left thigh. Lower lip with strong white marking extending almost to earhole (
<figureCitation id="12FABFC7FFDF284E7B78BEA6CBFEFA63" box="[356,432,1447,1473]" captionStart="FIGURE 1" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[158,1429,181,1905]" captionTargetId="figure-22@6.[158,1429,181,1920]" captionTargetPageId="6" captionText="FIGURE 1. Adult female holotype of Naultinus flavirictus sp. nov. RE003322 (original field number G.789): A.) dorsal, B.) ventral, and C.) lateral views. Photographs by J-C. Stahl. Scale bar = 1cm." figureDoi="http://doi.org/10.5281/zenodo.4456588" httpUri="https://zenodo.org/record/4456588/files/figure.png" pageId="3" pageNumber="392">Figs. 1</figureCitation>
, 
<figureCitation id="12FABFC7FFDF284E7BA1BEA6CB80FA63" box="[445,462,1447,1473]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[246,1313,181,1920]" captionTargetId="figure-17@7.[246,1341,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. Comparison of head scalation between relevant Naultinus spp.: A.) N. manukanus (MONZ RE.2013); B.) N. stellatus (RE.005617; original field number RAH466); C.) N. flavirictus sp. nov. (holotype); D.) N. elegans (RE.002658; original field number G.102); and E.) N. grayii (RE.002771; original field number G.231). Photos by SVN." figureDoi="http://doi.org/10.5281/zenodo.4456590" httpUri="https://zenodo.org/record/4456590/files/figure.png" pageId="3" pageNumber="392">2</figureCitation>
). Faint yellow colouration around nostrils. Undersides of feet and toes similar in colouring to venter.
</paragraph>
<paragraph id="8A7EA342FFDF284E7ADBBEEFCEB4F943" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="392">
<emphasis id="B8B57F50FFDF284E7ADBBEEFCB23F9AA" bold="true" box="[199,365,1518,1544]" pageId="3" pageNumber="392">Colour in life.</emphasis>
(from 
<typeStatus id="557A1DE0FFDF284E7BA0BEEEC864F9AB" box="[444,554,1519,1545]" pageId="3" pageNumber="392" type="paratype">paratypes</typeStatus>
, supplemented by non-vouchered photographs of other wild individuals – 
<figureCitation id="12FABFC7FFDF284E7F6CBEEECAE9F98F" captionStart="FIGURE 3" captionStartId="8.[151,250,1358,1382]" captionTargetBox="[157,1431,181,1319]" captionTargetId="figure-22@8.[157,1431,181,1334]" captionTargetPageId="8" captionText="FIGURE 3. In-life photographs of two N. flavirictus sp. nov. individuals from Te Paki (both male; not vouchered). Photos by SVN (A, B) and D. van Winkel (C)." figureDoi="http://doi.org/10.5281/zenodo.4456592" httpUri="https://zenodo.org/record/4456592/files/figure.png" pageId="3" pageNumber="392">Fig. 3</figureCitation>
). As in preservative, except that the basic colour is very bright green rather than dull dark green. Pattern of spots or stripes is variable among individuals, or can be absent. There is sexual dichromatism, with adult and subadult males having the ventral surface and lower lateral surface very pale blue; these areas are light green in females. In a few individuals the background bright green colouration is replaced by bright yellow (P. Miller pers. comm.; 
<bibRefCitation id="EE50DEB3FFDF284E7F68BD7ECB20F91F" author="van Winkel, D. &amp; Baling, M. &amp; Hitchmough, R." pageId="3" pageNumber="392" refId="ref9042" refString="van Winkel, D., Baling, M. &amp; Hitchmough, R. (2018) Reptiles and Amphibians of New Zealand, a Field Guide. Auckland University Press, Auckland, 366 pp." type="book" year="2018">
van Winkel 
<emphasis id="B8B57F50FFDF284E7AECBDA2CB64F91F" box="[240,298,1699,1725]" italics="true" pageId="3" pageNumber="392">et al.</emphasis>
2018
</bibRefCitation>
). Pale lavender mouth lining, with yellow-orange jaws and tongue, with the colour of the jaws extending to the external corners of the mouth, and with similar pigment present around the nostrils.
</paragraph>
<paragraph id="8A7EA342FFDF284E7ADBBDEACED4F837" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="392">
<emphasis id="B8B57F50FFDF284E7ADBBDEACBE3F8A6" bold="true" box="[199,429,1770,1797]" pageId="3" pageNumber="392">
<typeStatus id="557A1DE0FFDF284E7ADBBDEACB7AF8A7" box="[199,308,1771,1797]" pageId="3" pageNumber="392" type="paratype">Paratype</typeStatus>
variation.
</emphasis>
Both MONZ RE.005611 and MONZ RE.005609 are very similar in appearance to the 
<typeStatus id="557A1DE0FFDF284E7F7DBDEACA85F88B" pageId="3" pageNumber="392" type="holotype">holotype</typeStatus>
. They differ in the extent of pale dorsal markings, and presence of mid-lateral markings in MONZ RE.005611), number and arrangement of internasals and postmentals, and have minor differences in numbers of infralabials and supralabials, scale rows between supraciliaries, scale rows across frontal bones at midpoint of orbit, lamellae counts and numbers of precloacal pores (which are more strongly developed in the male 
<typeStatus id="557A1DE0FFDF284E7E35BC7ACEC1F837" box="[1065,1167,1915,1941]" pageId="3" pageNumber="392" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8A7EA342FFDF28497ADBBC9EC841FD6F" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="393" pageId="3" pageNumber="392">
MONZ RE.005611 has a small, diamond-shaped internasal, posterior to which are a pair of considerably larger, hexagonal internasals, which exclude the small central one from contact with the supranasals. Four postnasals on left, three on right. Two hexagonal postmentals. Chin shields ¼ to same size as postmentals. 11 (left), 10 (right) infralabials. 13 (left), 11 (right) supralabials. 9th (left), 8th (right) in mid-orbital position. 39 scale rows between supraciliaries (29 between supraocular skin folds), 10 scale rows across frontal bones at midpoint of orbit. Dorsal and particularly lateral scales weakly subimbricate. Lamellar counts (including all scales&gt; twice size of scales at base of digits) from right (and left) sides 5-12-13-14-10 (6-11-13-14-12) manus and 7-13-15-14-15 (7-13-14-14-13) pes. Precloacal pores 
<quantity id="4D390EA7FFD828497B90BA4ACB9DFEC7" box="[396,467,331,357]" metricMagnitude="0" metricUnit="m" metricValue="2.8448" pageId="4" pageNumber="393" unit="in" value="112.0">112 in</quantity>
total, in 5 rows, 3 extending onto thigh, 2 reaching mid-point of thigh. Scale row immediately posterior to precloacal spores includes 5 pitted scales – not included in count. Four left and three right, enlarged, raised and sharply pointed cloacal spurs. Colour in preservative deep lavender dorsally, becoming paler on flanks and limbs, body with paired, mostly symmetrical series of cream-coloured markings, fragmented into several elongate lines on left side, series of more variable lines and blotches on right side, extending onto tail as two broken lines; head with a pair of crescentic supratemporal markings, a large marking extending between left and right antorbital positions, narrowing in the interorbital region and expanding again into a blunted arrowhead shape on the parietals. All except last marking outlined by a thin, more or less continuous black line. Narrow cream line extending from infralabials almost to ear opening. Ventrolateral marking as in 
<typeStatus id="557A1DE0FFD828497E26B96ACEEEFD27" box="[1082,1184,619,645]" pageId="4" pageNumber="393" type="holotype">holotype</typeStatus>
. Small scattered white patches on posterior flanks (one on left, two on right). Limbs without markings. Venter immaculate. Dorsum and venter of digits pale straw colour.
</paragraph>
<paragraph id="8A7EA342FFD828497ADBB9D6CEADFBB3" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="393">MONZ RE.005609 has a single hexagonal internasal and three postnasals on both left and right sides. Two irregular postmentals, right three times size of left. Chin shields 1/3 to same size as postmentals. 11 (left), 12 (right) infralabials. 13 (left), 14 (right) supralabials. 10th (left), 11th (right) in mid-orbital position. 37 scale rows between supraciliaries (27 between supraocular skin folds), 11 scale rows across frontal bones at midpoint of orbit. Dorsal and particularly lateral scales weakly subimbricate. Lamellar counts (including all scales&gt; twice size of scales at base of digits) from right (and left) sides 5 (other digits damaged) (5-9-10-11-10) manus and 5-12- damaged-damaged-14 (6-damaged-damaged-14-damaged) pes. Six rows of pitted scales (c. 75 total) only one extending onto base of thigh. Four left and three right, enlarged, raised and sharply pointed cloacal spurs. Colour in preservative pale lavender, only visible dorsal markings are faded, slightly bowed, whitish supratemporal markings.</paragraph>
<paragraph id="8A7EA342FFD828497ADBBF1BCB3AFB48" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="393">
<emphasis id="B8B57F50FFD828497ADBBF1BCBA1FB96" bold="true" box="[199,495,1050,1076]" pageId="4" pageNumber="393">Geographic distribution.</emphasis>
The northern half of the Aupōuri Peninsula, the peninsula at the northern extremity of the North Island of 
<collectingCountry id="F2D6E3D2FFD828497B87BF3EC879FBFB" box="[411,567,1087,1113]" name="New Zealand" pageId="4" pageNumber="393">New Zealand</collectingCountry>
. Most records are from the Te Paki area in the far north (a general distribution map is provided by 
<bibRefCitation id="EE50DEB3FFD828497BB7BF62C8CAFBDF" author="Nielsen, S. V. &amp; Bauer, A. M. &amp; Jackman, T. R. &amp; Hitchmough, R. A. &amp; Daugherty, C. H." box="[427,644,1123,1149]" pageId="4" pageNumber="393" pagination="1 - 22" refId="ref8781" refString="Nielsen, S. V., Bauer, A. M., Jackman, T. R., Hitchmough, R. A. &amp; Daugherty, C. H. (2011) New Zealand geckos (Diplodactylidae): cryptic diversity in a post-Gondwanan lineage with trans-Tasman affinities. Molecular Phylogenetics and Evolution, 59, 1 - 22. https: // doi. org / 10.1016 / j. ympev. 2010.12.007" type="journal article" year="2011">
Nielsen 
<emphasis id="B8B57F50FFD828497814BF62C874FBDF" box="[520,570,1123,1149]" italics="true" pageId="4" pageNumber="393">et al</emphasis>
. 2011
</bibRefCitation>
). The gecko is found on the northern half of the peninsula, from about Houhora northwards. Exact locality details are not provided to discourage illegal collection, which is a well-documented threat to New Zealand’s native herpetofauna (
<bibRefCitation id="EE50DEB3FFD82849791EBFAAC993FB67" author="Nelson, N. J. &amp; Hitchmough, R. A. &amp; Monks, J. M." box="[770,989,1195,1221]" pageId="4" pageNumber="393" pagination="382 - 404" refId="ref8704" refString="Nelson, N. J., Hitchmough, R. A., Monks, J. M. (2014) New Zealand reptiles and their conservation. In: Stow, A., Holwell, G. &amp; Maclean, N. (Eds.), Austral Ark. Cambridge University Press, Cambridge, pp. 382 - 404. https: // doi. org / 10.1017 / CBO 9781139519960.020" type="book chapter" year="2014">
<collectingRegion id="48056DA0FFD82849791EBFAAC91DFB67" box="[770,851,1195,1221]" country="New Zealand" name="Nelson" pageId="4" pageNumber="393">Nelson</collectingRegion>
<emphasis id="B8B57F50FFD828497940BFAAC9D9FB67" box="[860,919,1195,1221]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
2014
</bibRefCitation>
; Hitchmough 
<emphasis id="B8B57F50FFD828497E63BFAACEF4FB67" box="[1151,1210,1195,1221]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
2016) and globally (Bryan 
<emphasis id="B8B57F50FFD828497AF0BFCECB6AFB4B" box="[236,292,1231,1257]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
2006).
</paragraph>
<paragraph id="8A7EA342FFD828497ADBBFF3CE2EF98F" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="393">
<emphasis id="B8B57F50FFD828497ADBBFF3CB3DFAAE" bold="true" box="[199,371,1266,1292]" pageId="4" pageNumber="393">Biogeography.</emphasis>
The paleo-islands at the northern end of the peninsula are formed of Cretaceous ophiolitic (basalt, gabbro and serpentinite) and Lower Miocene marine sedimentary rocks (
<bibRefCitation id="EE50DEB3FFD828497E33BE16CF0BFA93" author="Brook, F. J. &amp; Thrasher, G. P." box="[1071,1349,1303,1329]" pageId="4" pageNumber="393" pagination="1 - 42" refId="ref7675" refString="Brook, F. J. &amp; Thrasher, G. P. (1991) Cretaceous and Cenozoic geology of northernmost New Zealand. New Zealand Geological Survey Record, 41, 1 - 42." type="journal article" year="1991">Brook &amp; Thrasher 1991</bibRefCitation>
). These paleo-islands were probably emergent above sea-level from Late Miocene or Early Pliocene time. During Early to Mid Pleistocene time extensive deposits of marine and aeolian sand accumulated north of Kaitaia; paleo-islands probably existed in the Reinga-North Cape and Houhora areas during interglacial periods (i.e., high sea levels), but a proto-Aupouri Peninsula would have existed during glacial low stands. Dune buildup during Late Pleistocene time formed the present land connection along the Peninsula. Thus, the far northern islands were probably isolated throughout the Pliocene, and were intermittently connected with the rest of 
<collectingRegion id="48056DA0FFD8284979F5BEEECE15F9AB" box="[1001,1115,1519,1545]" country="New Zealand" name="Northland" pageId="4" pageNumber="393">Northland</collectingRegion>
during the Pleistocene, with the connection becoming permanent in the Late Pleistocene (
<bibRefCitation id="EE50DEB3FFD82849795EBD12CE1DF98F" author="Brook, F. J. &amp; Thrasher, G. P." box="[834,1107,1555,1581]" pageId="4" pageNumber="393" pagination="1 - 42" refId="ref7675" refString="Brook, F. J. &amp; Thrasher, G. P. (1991) Cretaceous and Cenozoic geology of northernmost New Zealand. New Zealand Geological Survey Record, 41, 1 - 42." type="journal article" year="1991">Brook &amp; Thrasher 1991</bibRefCitation>
).
</paragraph>
<paragraph id="8A7EA342FFD828497ADBBD36CE18F88B" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="393">
Although we did not perform specific dating analyses per se, if we assume a mutation rate for ND2 of 0.65% per million years (as proposed by Macey 
<emphasis id="B8B57F50FFD828497826BD5AC83CF9D7" box="[570,626,1627,1653]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
1998), the ~4% mitochondrial distance separating 
<taxonomicName id="4DC1D8C1FFD828497EB7BD5ACF73F9D7" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[1195,1341,1627,1653]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFD828497EB7BD5ACF73F9D7" box="[1195,1341,1627,1653]" italics="true" pageId="4" pageNumber="393">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFD828497F5FBD5ACFD2F9D7" bold="true" box="[1347,1436,1627,1653]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A386C22BFFD828497F5FBD5ACFD2F9D7" box="[1347,1436,1627,1653]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from its closest kin (Table S01) correlates to ~2.5 million years. It is therefore reasonable to infer that the ancestors of 
<taxonomicName id="4DC1D8C1FFD828497AA8BDA2CBD7F91F" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[180,409,1699,1725]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFD828497AA8BDA2CBD7F91F" box="[180,409,1699,1725]" italics="true" pageId="4" pageNumber="393">Naultinus flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFD828497B83BDA2CBB9F91F" bold="true" box="[415,503,1699,1725]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A386C22BFFD828497B83BDA2CBB9F91F" box="[415,503,1699,1725]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
reached the Te Paki area while it was an island or series of islands and evolved there in isolation until the formation of the Aupōuri tombolo enabled it to come back into contact with 
<taxonomicName id="4DC1D8C1FFD828497EFBBDC6CF05F942" authorityName="Bell" authorityYear="1843" box="[1255,1355,1735,1760]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFD828497EFBBDC6CF05F942" box="[1255,1355,1735,1760]" italics="true" pageId="4" pageNumber="393">N. grayii</emphasis>
</taxonomicName>
during the Pleistocene. Contact zones with 
<taxonomicName id="4DC1D8C1FFD828497833BDEAC8DDF8A6" authorityName="Bell" authorityYear="1843" box="[559,659,1771,1796]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFD828497833BDEAC8DDF8A6" box="[559,659,1771,1796]" italics="true" pageId="4" pageNumber="393">N. grayii</emphasis>
</taxonomicName>
may have repeatedly formed and then been removed by inter-glacial sea-level rise, and the current contact is likely to have formed in the Late Pleistocene.
</paragraph>
<paragraph id="8A7EA342FFD828497ADBBC33C9EFF837" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="393">
<emphasis id="B8B57F50FFD828497ADBBC33CB3DF8EE" bold="true" box="[199,371,1842,1868]" pageId="4" pageNumber="393">Reproduction.</emphasis>
Animals collected from Te Kao in 
<date id="FE7F8582FFD8284978E5BC32C9C6F8EF" box="[761,904,1843,1869]" pageId="4" pageNumber="393" value="1986-08">August 1986</date>
then held in captivity in 
<collectingRegion id="48056DA0FFD828497E80BC32CF47F8EF" box="[1180,1289,1843,1869]" country="New Zealand" name="Auckland" pageId="4" pageNumber="393">Auckland</collectingRegion>
gave birth to twins in late 
<date id="FE7F8582FFD828497B39BC56CBF5F8D3" box="[293,443,1879,1905]" pageId="4" pageNumber="393" value="1987-01">January 1987</date>
(C. Hepers, pers. comm.). This is almost two months earlier than births of 
<taxonomicName id="4DC1D8C1FFD828497EE6BC56CF13F8D2" authorityName="Bell" authorityYear="1843" box="[1274,1373,1879,1904]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFD828497EE6BC56CF13F8D2" box="[1274,1373,1879,1904]" italics="true" pageId="4" pageNumber="393">N. grayii</emphasis>
</taxonomicName>
, both in the field and in captivity in 
<collectingRegion id="48056DA0FFD828497BF4BC7AC81BF837" box="[488,597,1915,1941]" country="New Zealand" name="Auckland" pageId="4" pageNumber="393">Auckland</collectingRegion>
(
<bibRefCitation id="EE50DEB3FFD82849787FBC7AC9DBF837" author="Robb, J. &amp; Hitchmough, R. A." box="[611,917,1915,1941]" pageId="4" pageNumber="393" pagination="189 - 200" refId="ref8947" refString="Robb, J. &amp; Hitchmough, R. A. (1980) Review of the genus Naultinus Gray (Reptilia: Gekkonidae). Records of the Auckland Institute and Museum, 16, 189 - 200." type="journal article" year="1980">Robb &amp; Hitchmough 1980</bibRefCitation>
).
</paragraph>
<caption id="DEBEF3CAFFD928487A8BB8CCCD41FC7F" ID-Table-UUID="DEBEF3CAFFD928487A8BB8CCCD41FC7F" box="[151,1807,973,989]" httpUri="http://table.plazi.org/id/DEBEF3CAFFD928487A8BB8CCCD41FC7F" pageId="5" pageNumber="394" targetBox="[151,1998,1014,1244]" targetIsTable="true" targetPageId="5">
<paragraph id="8A7EA342FFD928487A8BB8CCCD41FC7F" blockId="5.[151,1807,973,989]" box="[151,1807,973,989]" pageId="5" pageNumber="394">
Ta b le 
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. Unco rre cte d m e a n p a irwise se q ue nce d ive rg e nce (p) va lue s fo r ND 2 a nd 1 6 S, g e ne ra te d in M E G A v.6.0 (Ta m ura e t a l. 2 0 1 3), using se q ue nce d a ta fro m Nie lse n e t a l. (2 0 1 1).
</paragraph>
</caption>
<paragraph id="8A7EA342FFD928487AA2B8F9CD80FB7E" pageId="5" pageNumber="394">
<table id="F8C151E2FFD9D7B27A8BB8F7CD80FB7E" box="[151,1998,1014,1244]" gridcols="11" gridrows="11" pageId="5" pageNumber="394">
<tr id="34F1A100FFD9D7B27A8BB8F7CD80FBAA" box="[151,1998,1014,1032]" gridrow="0" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BB8F7CB86FBAA" box="[151,456,1014,1032]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="5" pageNumber="394">ND 2 \1 6 S N. g e m m e u s</th>
<th id="7720C87CFFD9D7B27BF2B8F7C83DFBAA" box="[494,627,1014,1032]" gridcol="2" gridrow="0" pageId="5" pageNumber="394">
<taxonomicName id="4DC1D8C1FFD928487BF2B8F7C81CFBA4" box="[494,594,1014,1030]" pageId="5" pageNumber="394" rank="subSpecies" species="fla" subSpecies="virictu">N. fla virictu</taxonomicName>
s
</th>
<th id="7720C87CFFD9D7B27885B8F7C951FBAA" box="[665,799,1014,1032]" gridcol="3" gridrow="0" pageId="5" pageNumber="394">N. p u n cta tu s</th>
<th id="7720C87CFFD9D7B27958B8F7C9E3FBAA" box="[836,941,1014,1032]" gridcol="4" gridrow="0" pageId="5" pageNumber="394">N. e le g a n s</th>
<th id="7720C87CFFD9D7B279ECB8F7CE3BFBAA" box="[1008,1141,1014,1032]" gridcol="5" gridrow="0" pageId="5" pageNumber="394">N. g ra yii</th>
<th id="7720C87CFFD9D7B27E87B8F7CF6AFBAA" box="[1179,1316,1014,1032]" gridcol="6" gridrow="0" pageId="5" pageNumber="394">N. tu b e rcu la tu s</th>
<th id="7720C87CFFD9D7B27F5AB8F7CF82FBAA" box="[1350,1484,1014,1032]" gridcol="7" gridrow="0" pageId="5" pageNumber="394">
<taxonomicName id="4DC1D8C1FFD928487F5AB8F7CFDEFBA4" box="[1350,1424,1014,1030]" pageId="5" pageNumber="394" rank="subSpecies" species="ste" subSpecies="lla">N. ste lla</taxonomicName>
tu s
</th>
<th id="7720C87CFFD9D7B27FEEB8F7CC15FBAA" box="[1522,1627,1014,1032]" gridcol="8" gridrow="0" pageId="5" pageNumber="394">N. ru d is</th>
<th id="7720C87CFFD9D7B27C81B8F7CD80FBAA" box="[1693,1998,1014,1032]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="5" pageNumber="394">N. p o e cilo ch lo ru s N. m a n u ka n u s</th>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF0DCD80FBBF" box="[151,1998,1036,1053]" gridrow="1" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF0DCB6EFBBF" box="[151,288,1036,1053]" gridcol="0" gridrow="1" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF0DCB43FBBE" box="[151,269,1036,1052]" italics="true" pageId="5" pageNumber="394">
N. g e m m e u 
<geoCoordinate id="EFF5C585FFD928487B18BF0DCB43FBBE" box="[260,269,1036,1052]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF0DCB86FBBF" box="[350,456,1036,1053]" gridcol="1" gridrow="1" pageId="5" pageNumber="394">0.0 0 2 \0.0 0 0</td>
<td id="7720C87CFFD9D7B27BF2BF0DC83DFBBF" box="[494,627,1036,1053]" gridcol="2" gridrow="1" pageId="5" pageNumber="394">0.0 1 3</td>
<td id="7720C87CFFD9D7B27885BF0DC951FBBF" box="[665,799,1036,1053]" gridcol="3" gridrow="1" pageId="5" pageNumber="394">0.0 1 9</td>
<td id="7720C87CFFD9D7B27958BF0DC9E3FBBF" box="[836,941,1036,1053]" gridcol="4" gridrow="1" pageId="5" pageNumber="394">0.0 2 0</td>
<td id="7720C87CFFD9D7B279ECBF0DCE3BFBBF" box="[1008,1141,1036,1053]" gridcol="5" gridrow="1" pageId="5" pageNumber="394">0.0 2 1</td>
<td id="7720C87CFFD9D7B27E87BF0DCF6AFBBF" box="[1179,1316,1036,1053]" gridcol="6" gridrow="1" pageId="5" pageNumber="394">0.0 1 8</td>
<td id="7720C87CFFD9D7B27F5ABF0DCF82FBBF" box="[1350,1484,1036,1053]" gridcol="7" gridrow="1" pageId="5" pageNumber="394">0.0 1 6</td>
<td id="7720C87CFFD9D7B27FEEBF0DCC15FBBF" box="[1522,1627,1036,1053]" gridcol="8" gridrow="1" pageId="5" pageNumber="394">0.0 2 2</td>
<td id="7720C87CFFD9D7B27C81BF0DCD48FBBF" box="[1693,1798,1036,1053]" gridcol="9" gridrow="1" pageId="5" pageNumber="394">0.0 1 8</td>
<td id="7720C87CFFD9D7B27D78BF0DCD80FBBF" box="[1892,1998,1036,1053]" gridcol="10" gridrow="1" pageId="5" pageNumber="394">0.0 1 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF20CD80FB91" box="[151,1998,1057,1075]" gridrow="2" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF20CB6EFB91" box="[151,288,1057,1075]" gridcol="0" gridrow="2" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF20CB49FB93" box="[151,263,1057,1073]" italics="true" pageId="5" pageNumber="394">
<taxonomicName id="4DC1D8C1FFD928487A8BBF20CAB5FB93" box="[151,251,1057,1073]" pageId="5" pageNumber="394" rank="subSpecies" species="fla" subSpecies="virictu">N. fla virictu</taxonomicName>
<geoCoordinate id="EFF5C585FFD928487AE2BF20CB49FB93" box="[254,263,1057,1073]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF20CB86FB91" box="[350,456,1057,1075]" gridcol="1" gridrow="2" pageId="5" pageNumber="394">0.0 6 9</td>
<td id="7720C87CFFD9D7B27BF2BF20C83DFB91" box="[494,627,1057,1075]" gridcol="2" gridrow="2" pageId="5" pageNumber="394">0.0 0 6 \0.0 0 0</td>
<td id="7720C87CFFD9D7B27885BF20C951FB91" box="[665,799,1057,1075]" gridcol="3" gridrow="2" pageId="5" pageNumber="394">0.0 1 1</td>
<td id="7720C87CFFD9D7B27958BF20C9E3FB91" box="[836,941,1057,1075]" gridcol="4" gridrow="2" pageId="5" pageNumber="394">0.0 0 7</td>
<td id="7720C87CFFD9D7B279ECBF20CE3BFB91" box="[1008,1141,1057,1075]" gridcol="5" gridrow="2" pageId="5" pageNumber="394">0.0 1 0</td>
<td id="7720C87CFFD9D7B27E87BF20CF6AFB91" box="[1179,1316,1057,1075]" gridcol="6" gridrow="2" pageId="5" pageNumber="394">0.0 1 3</td>
<td id="7720C87CFFD9D7B27F5ABF20CF82FB91" box="[1350,1484,1057,1075]" gridcol="7" gridrow="2" pageId="5" pageNumber="394">0.0 0 7</td>
<td id="7720C87CFFD9D7B27FEEBF20CC15FB91" box="[1522,1627,1057,1075]" gridcol="8" gridrow="2" pageId="5" pageNumber="394">0.0 1 5</td>
<td id="7720C87CFFD9D7B27C81BF20CD48FB91" box="[1693,1798,1057,1075]" gridcol="9" gridrow="2" pageId="5" pageNumber="394">0.0 1 1</td>
<td id="7720C87CFFD9D7B27D78BF20CD80FB91" box="[1892,1998,1057,1075]" gridcol="10" gridrow="2" pageId="5" pageNumber="394">0.0 0 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF36CD80FBEA" box="[151,1998,1079,1096]" gridrow="3" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF36CB6EFBEA" box="[151,288,1079,1096]" gridcol="0" gridrow="3" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF36CB47FBE5" box="[151,265,1079,1095]" italics="true" pageId="5" pageNumber="394">
N. p u n cta tu 
<geoCoordinate id="EFF5C585FFD928487B1CBF36CB47FBE5" box="[256,265,1079,1095]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF36CB86FBEA" box="[350,456,1079,1096]" gridcol="1" gridrow="3" pageId="5" pageNumber="394">0.0 7 4</td>
<td id="7720C87CFFD9D7B27BF2BF36C83DFBEA" box="[494,627,1079,1096]" gridcol="2" gridrow="3" pageId="5" pageNumber="394">0.0 4 2</td>
<td id="7720C87CFFD9D7B27885BF36C951FBEA" box="[665,799,1079,1096]" gridcol="3" gridrow="3" pageId="5" pageNumber="394">0.0 1 4 \0.0 0 4</td>
<td id="7720C87CFFD9D7B27958BF36C9E3FBEA" box="[836,941,1079,1096]" gridcol="4" gridrow="3" pageId="5" pageNumber="394">0.0 1 1</td>
<td id="7720C87CFFD9D7B279ECBF36CE3BFBEA" box="[1008,1141,1079,1096]" gridcol="5" gridrow="3" pageId="5" pageNumber="394">0.0 1 4</td>
<td id="7720C87CFFD9D7B27E87BF36CF6AFBEA" box="[1179,1316,1079,1096]" gridcol="6" gridrow="3" pageId="5" pageNumber="394">0.0 1 5</td>
<td id="7720C87CFFD9D7B27F5ABF36CF82FBEA" box="[1350,1484,1079,1096]" gridcol="7" gridrow="3" pageId="5" pageNumber="394">0.0 1 5</td>
<td id="7720C87CFFD9D7B27FEEBF36CC15FBEA" box="[1522,1627,1079,1096]" gridcol="8" gridrow="3" pageId="5" pageNumber="394">0.0 2 5</td>
<td id="7720C87CFFD9D7B27C81BF36CD48FBEA" box="[1693,1798,1079,1096]" gridcol="9" gridrow="3" pageId="5" pageNumber="394">0.0 2 1</td>
<td id="7720C87CFFD9D7B27D78BF36CD80FBEA" box="[1892,1998,1079,1096]" gridcol="10" gridrow="3" pageId="5" pageNumber="394">0.0 1 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF4DCD80FBFC" box="[151,1998,1100,1118]" gridrow="4" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF4DCB6EFBFC" box="[151,288,1100,1118]" gridcol="0" gridrow="4" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF4DCAB4FBFE" box="[151,250,1100,1116]" italics="true" pageId="5" pageNumber="394">
N. e le g a n 
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</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF4DCB86FBFC" box="[350,456,1100,1118]" gridcol="1" gridrow="4" pageId="5" pageNumber="394">0.0 7 4</td>
<td id="7720C87CFFD9D7B27BF2BF4DC83DFBFC" box="[494,627,1100,1118]" gridcol="2" gridrow="4" pageId="5" pageNumber="394">0.0 4 0</td>
<td id="7720C87CFFD9D7B27885BF4DC951FBFC" box="[665,799,1100,1118]" gridcol="3" gridrow="4" pageId="5" pageNumber="394">0.0 3 0</td>
<td id="7720C87CFFD9D7B27958BF4DC9E3FBFC" box="[836,941,1100,1118]" gridcol="4" gridrow="4" pageId="5" pageNumber="394">— \—</td>
<td id="7720C87CFFD9D7B279ECBF4DCE3BFBFC" box="[1008,1141,1100,1118]" gridcol="5" gridrow="4" pageId="5" pageNumber="394">0.0 1 0</td>
<td id="7720C87CFFD9D7B27E87BF4DCF6AFBFC" box="[1179,1316,1100,1118]" gridcol="6" gridrow="4" pageId="5" pageNumber="394">0.0 1 3</td>
<td id="7720C87CFFD9D7B27F5ABF4DCF82FBFC" box="[1350,1484,1100,1118]" gridcol="7" gridrow="4" pageId="5" pageNumber="394">0.0 1 3</td>
<td id="7720C87CFFD9D7B27FEEBF4DCC15FBFC" box="[1522,1627,1100,1118]" gridcol="8" gridrow="4" pageId="5" pageNumber="394">0.0 2 2</td>
<td id="7720C87CFFD9D7B27C81BF4DCD48FBFC" box="[1693,1798,1100,1118]" gridcol="9" gridrow="4" pageId="5" pageNumber="394">0.0 1 8</td>
<td id="7720C87CFFD9D7B27D78BF4DCD80FBFC" box="[1892,1998,1100,1118]" gridcol="10" gridrow="4" pageId="5" pageNumber="394">0.0 1 5</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF63CD80FBD1" box="[151,1998,1122,1139]" gridrow="5" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF63CB6EFBD1" box="[151,288,1122,1139]" gridcol="0" gridrow="5" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF63CAACFBD0" box="[151,226,1122,1138]" italics="true" pageId="5" pageNumber="394">N. g ra yii</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF63CB86FBD1" box="[350,456,1122,1139]" gridcol="1" gridrow="5" pageId="5" pageNumber="394">0.0 7 6</td>
<td id="7720C87CFFD9D7B27BF2BF63C83DFBD1" box="[494,627,1122,1139]" gridcol="2" gridrow="5" pageId="5" pageNumber="394">0.0 4 0</td>
<td id="7720C87CFFD9D7B27885BF63C951FBD1" box="[665,799,1122,1139]" gridcol="3" gridrow="5" pageId="5" pageNumber="394">0.0 3 0</td>
<td id="7720C87CFFD9D7B27958BF63C9E3FBD1" box="[836,941,1122,1139]" gridcol="4" gridrow="5" pageId="5" pageNumber="394">0.0 1 7</td>
<td id="7720C87CFFD9D7B279ECBF63CE3BFBD1" box="[1008,1141,1122,1139]" gridcol="5" gridrow="5" pageId="5" pageNumber="394">0.0 0 8 \0.0 0 2</td>
<td id="7720C87CFFD9D7B27E87BF63CF6AFBD1" box="[1179,1316,1122,1139]" gridcol="6" gridrow="5" pageId="5" pageNumber="394">0.0 1 9</td>
<td id="7720C87CFFD9D7B27F5ABF63CF82FBD1" box="[1350,1484,1122,1139]" gridcol="7" gridrow="5" pageId="5" pageNumber="394">0.0 1 6</td>
<td id="7720C87CFFD9D7B27FEEBF63CC15FBD1" box="[1522,1627,1122,1139]" gridcol="8" gridrow="5" pageId="5" pageNumber="394">0.0 2 5</td>
<td id="7720C87CFFD9D7B27C81BF63CD48FBD1" box="[1693,1798,1122,1139]" gridcol="9" gridrow="5" pageId="5" pageNumber="394">0.0 2 1</td>
<td id="7720C87CFFD9D7B27D78BF63CD80FBD1" box="[1892,1998,1122,1139]" gridcol="10" gridrow="5" pageId="5" pageNumber="394">0.0 1 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF76CD80FB2B" box="[151,1998,1143,1161]" gridrow="6" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF76CB6EFB2B" box="[151,288,1143,1161]" gridcol="0" gridrow="6" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF76CB6EFB25" box="[151,288,1143,1159]" italics="true" pageId="5" pageNumber="394">
N. tu b e rcu la tu 
<geoCoordinate id="EFF5C585FFD928487B0BBF76CB6EFB25" box="[279,288,1143,1159]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF76CB86FB2B" box="[350,456,1143,1161]" gridcol="1" gridrow="6" pageId="5" pageNumber="394">0.0 7 9</td>
<td id="7720C87CFFD9D7B27BF2BF76C83DFB2B" box="[494,627,1143,1161]" gridcol="2" gridrow="6" pageId="5" pageNumber="394">0.0 5 5</td>
<td id="7720C87CFFD9D7B27885BF76C951FB2B" box="[665,799,1143,1161]" gridcol="3" gridrow="6" pageId="5" pageNumber="394">0.0 6 3</td>
<td id="7720C87CFFD9D7B27958BF76C9E3FB2B" box="[836,941,1143,1161]" gridcol="4" gridrow="6" pageId="5" pageNumber="394">0.0 6 1</td>
<td id="7720C87CFFD9D7B279ECBF76CE3BFB2B" box="[1008,1141,1143,1161]" gridcol="5" gridrow="6" pageId="5" pageNumber="394">0.0 6 0</td>
<td id="7720C87CFFD9D7B27E87BF76CF6AFB2B" box="[1179,1316,1143,1161]" gridcol="6" gridrow="6" pageId="5" pageNumber="394">0.0 0 7 \0.0 0 2</td>
<td id="7720C87CFFD9D7B27F5ABF76CF82FB2B" box="[1350,1484,1143,1161]" gridcol="7" gridrow="6" pageId="5" pageNumber="394">0.0 1 6</td>
<td id="7720C87CFFD9D7B27FEEBF76CC15FB2B" box="[1522,1627,1143,1161]" gridcol="8" gridrow="6" pageId="5" pageNumber="394">0.0 2 4</td>
<td id="7720C87CFFD9D7B27C81BF76CD48FB2B" box="[1693,1798,1143,1161]" gridcol="9" gridrow="6" pageId="5" pageNumber="394">0.0 2 0</td>
<td id="7720C87CFFD9D7B27D78BF76CD80FB2B" box="[1892,1998,1143,1161]" gridcol="10" gridrow="6" pageId="5" pageNumber="394">0.0 1 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBF8CCD80FB3C" box="[151,1998,1165,1182]" gridrow="7" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBF8CCB6EFB3C" box="[151,288,1165,1182]" gridcol="0" gridrow="7" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBF8CCAB2FB3F" box="[151,252,1165,1181]" italics="true" pageId="5" pageNumber="394">
<taxonomicName id="4DC1D8C1FFD928487A8BBF8CCAAEFB3F" box="[151,224,1165,1181]" pageId="5" pageNumber="394" rank="subSpecies" species="ste" subSpecies="lla">N. ste lla</taxonomicName>
tu 
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</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BF8CCB86FB3C" box="[350,456,1165,1182]" gridcol="1" gridrow="7" pageId="5" pageNumber="394">0.0 8 1</td>
<td id="7720C87CFFD9D7B27BF2BF8CC83DFB3C" box="[494,627,1165,1182]" gridcol="2" gridrow="7" pageId="5" pageNumber="394">0.0 6 1</td>
<td id="7720C87CFFD9D7B27885BF8CC951FB3C" box="[665,799,1165,1182]" gridcol="3" gridrow="7" pageId="5" pageNumber="394">0.0 6 9</td>
<td id="7720C87CFFD9D7B27958BF8CC9E3FB3C" box="[836,941,1165,1182]" gridcol="4" gridrow="7" pageId="5" pageNumber="394">0.0 6 7</td>
<td id="7720C87CFFD9D7B279ECBF8CCE3BFB3C" box="[1008,1141,1165,1182]" gridcol="5" gridrow="7" pageId="5" pageNumber="394">0.0 6 6</td>
<td id="7720C87CFFD9D7B27E87BF8CCF6AFB3C" box="[1179,1316,1165,1182]" gridcol="6" gridrow="7" pageId="5" pageNumber="394">0.0 4 0</td>
<td id="7720C87CFFD9D7B27F5ABF8CCF82FB3C" box="[1350,1484,1165,1182]" gridcol="7" gridrow="7" pageId="5" pageNumber="394">0.0 2 6 \0.0 0 5</td>
<td id="7720C87CFFD9D7B27FEEBF8CCC15FB3C" box="[1522,1627,1165,1182]" gridcol="8" gridrow="7" pageId="5" pageNumber="394">0.0 1 4</td>
<td id="7720C87CFFD9D7B27C81BF8CCD48FB3C" box="[1693,1798,1165,1182]" gridcol="9" gridrow="7" pageId="5" pageNumber="394">0.0 1 0</td>
<td id="7720C87CFFD9D7B27D78BF8CCD80FB3C" box="[1892,1998,1165,1182]" gridcol="10" gridrow="7" pageId="5" pageNumber="394">0.0 0 7</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBFA3CD80FB16" box="[151,1998,1186,1204]" gridrow="8" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBFA3CB6EFB16" box="[151,288,1186,1204]" gridcol="0" gridrow="8" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBFA3CA93FB10" box="[151,221,1186,1202]" italics="true" pageId="5" pageNumber="394">N. ru d is</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BFA3CB86FB16" box="[350,456,1186,1204]" gridcol="1" gridrow="8" pageId="5" pageNumber="394">0.0 7 5</td>
<td id="7720C87CFFD9D7B27BF2BFA3C83DFB16" box="[494,627,1186,1204]" gridcol="2" gridrow="8" pageId="5" pageNumber="394">0.0 5 7</td>
<td id="7720C87CFFD9D7B27885BFA3C951FB16" box="[665,799,1186,1204]" gridcol="3" gridrow="8" pageId="5" pageNumber="394">0.0 6 4</td>
<td id="7720C87CFFD9D7B27958BFA3C9E3FB16" box="[836,941,1186,1204]" gridcol="4" gridrow="8" pageId="5" pageNumber="394">0.0 6 3</td>
<td id="7720C87CFFD9D7B279ECBFA3CE3BFB16" box="[1008,1141,1186,1204]" gridcol="5" gridrow="8" pageId="5" pageNumber="394">0.0 6 2</td>
<td id="7720C87CFFD9D7B27E87BFA3CF6AFB16" box="[1179,1316,1186,1204]" gridcol="6" gridrow="8" pageId="5" pageNumber="394">0.0 3 6</td>
<td id="7720C87CFFD9D7B27F5ABFA3CF82FB16" box="[1350,1484,1186,1204]" gridcol="7" gridrow="8" pageId="5" pageNumber="394">0.0 3 2</td>
<td id="7720C87CFFD9D7B27FEEBFA3CC15FB16" box="[1522,1627,1186,1204]" gridcol="8" gridrow="8" pageId="5" pageNumber="394">— \—</td>
<td id="7720C87CFFD9D7B27C81BFA3CD48FB16" box="[1693,1798,1186,1204]" gridcol="9" gridrow="8" pageId="5" pageNumber="394">0.0 0 4</td>
<td id="7720C87CFFD9D7B27D78BFA3CD80FB16" box="[1892,1998,1186,1204]" gridcol="10" gridrow="8" pageId="5" pageNumber="394">0.0 0 9</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBFB9CD80FB6B" box="[151,1998,1208,1225]" gridrow="9" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBFB9CB86FB6B" box="[151,456,1208,1225]" colspan="2" colspanRight="1" gridcol="0" gridrow="9" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBFB9CB79FB6A" box="[151,311,1208,1224]" italics="true" pageId="5" pageNumber="394">
N. p o e cilo ch lo ru 
<geoCoordinate id="EFF5C585FFD928487B32BFB9CB79FB6A" box="[302,311,1208,1224]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
0.0 7 4
</th>
<td id="7720C87CFFD9D7B27BF2BFB9C83DFB6B" box="[494,627,1208,1225]" gridcol="2" gridrow="9" pageId="5" pageNumber="394">0.0 5 3</td>
<td id="7720C87CFFD9D7B27885BFB9C951FB6B" box="[665,799,1208,1225]" gridcol="3" gridrow="9" pageId="5" pageNumber="394">0.0 6 1</td>
<td id="7720C87CFFD9D7B27958BFB9C9E3FB6B" box="[836,941,1208,1225]" gridcol="4" gridrow="9" pageId="5" pageNumber="394">0.0 5 9</td>
<td id="7720C87CFFD9D7B279ECBFB9CE3BFB6B" box="[1008,1141,1208,1225]" gridcol="5" gridrow="9" pageId="5" pageNumber="394">0.0 5 7</td>
<td id="7720C87CFFD9D7B27E87BFB9CF6AFB6B" box="[1179,1316,1208,1225]" gridcol="6" gridrow="9" pageId="5" pageNumber="394">0.0 3 2</td>
<td id="7720C87CFFD9D7B27F5ABFB9CF82FB6B" box="[1350,1484,1208,1225]" gridcol="7" gridrow="9" pageId="5" pageNumber="394">0.0 2 8</td>
<td id="7720C87CFFD9D7B27FEEBFB9CC15FB6B" box="[1522,1627,1208,1225]" gridcol="8" gridrow="9" pageId="5" pageNumber="394">0.0 1 0</td>
<td id="7720C87CFFD9D7B27C81BFB9CD48FB6B" box="[1693,1798,1208,1225]" gridcol="9" gridrow="9" pageId="5" pageNumber="394">— \—</td>
<td id="7720C87CFFD9D7B27D78BFB9CD80FB6B" box="[1892,1998,1208,1225]" gridcol="10" gridrow="9" pageId="5" pageNumber="394">0.0 0 6</td>
</tr>
<tr id="34F1A100FFD9D7B27A8BBFCCCD80FB7E" box="[151,1998,1229,1244]" gridrow="10" pageId="5" pageNumber="394">
<th id="7720C87CFFD9D7B27A8BBFCCCB6EFB7E" box="[151,288,1229,1244]" gridcol="0" gridrow="10" pageId="5" pageNumber="394">
<emphasis id="B8B57F50FFD928487A8BBFCCCB53FB79" box="[151,285,1229,1243]" italics="true" pageId="5" pageNumber="394">
N. m a n u ka n u 
<geoCoordinate id="EFF5C585FFD928487B08BFCCCB53FB79" box="[276,285,1229,1243]" degrees="0.0" direction="south" orientation="latitude" pageId="5" pageNumber="394" precision="5555" value="-0.0">s</geoCoordinate>
</emphasis>
</th>
<td id="7720C87CFFD9D7B27B42BFCCCB86FB7E" box="[350,456,1229,1244]" gridcol="1" gridrow="10" pageId="5" pageNumber="394">0.0 7 8</td>
<td id="7720C87CFFD9D7B27BF2BFCCC83DFB7E" box="[494,627,1229,1244]" gridcol="2" gridrow="10" pageId="5" pageNumber="394">0.0 5 4</td>
<td id="7720C87CFFD9D7B27885BFCCC951FB7E" box="[665,799,1229,1244]" gridcol="3" gridrow="10" pageId="5" pageNumber="394">0.0 6 3</td>
<td id="7720C87CFFD9D7B27958BFCCC9E3FB7E" box="[836,941,1229,1244]" gridcol="4" gridrow="10" pageId="5" pageNumber="394">0.0 5 8</td>
<td id="7720C87CFFD9D7B279ECBFCCCE3BFB7E" box="[1008,1141,1229,1244]" gridcol="5" gridrow="10" pageId="5" pageNumber="394">0.0 5 9</td>
<td id="7720C87CFFD9D7B27E87BFCCCF6AFB7E" box="[1179,1316,1229,1244]" gridcol="6" gridrow="10" pageId="5" pageNumber="394">0.0 3 4</td>
<td id="7720C87CFFD9D7B27F5ABFCCCF82FB7E" box="[1350,1484,1229,1244]" gridcol="7" gridrow="10" pageId="5" pageNumber="394">0.0 3 6</td>
<td id="7720C87CFFD9D7B27FEEBFCCCC15FB7E" box="[1522,1627,1229,1244]" gridcol="8" gridrow="10" pageId="5" pageNumber="394">0.0 2 5</td>
<td id="7720C87CFFD9D7B27C81BFCCCD48FB7E" box="[1693,1798,1229,1244]" gridcol="9" gridrow="10" pageId="5" pageNumber="394">0.0 2 0</td>
<td id="7720C87CFFD9D7B27D78BFCCCD80FB7E" box="[1892,1998,1229,1244]" gridcol="10" gridrow="10" pageId="5" pageNumber="394">0.0 0 5 \0.0 0 0</td>
</tr>
</table>
</paragraph>
<caption id="DEBEF3CAFFDA284B7A8BBC96C9C7F876" ID-DOI="http://doi.org/10.5281/zenodo.4456588" ID-Zenodo-Dep="4456588" httpUri="https://zenodo.org/record/4456588/files/figure.png" pageId="6" pageNumber="395" startId="6.[151,250,1943,1967]" targetBox="[158,1429,181,1905]" targetPageId="6">
<paragraph id="8A7EA342FFDA284B7A8BBC96C9C7F876" blockId="6.[151,1437,1943,2004]" pageId="6" pageNumber="395">
<emphasis id="B8B57F50FFDA284B7A8BBC96CB5AF80D" bold="true" box="[151,276,1943,1967]" pageId="6" pageNumber="395">FIGURE 1.</emphasis>
Adult female holotype of 
<taxonomicName id="4DC1D8C1FFDA284B783ABC99C8B9F812" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[550,759,1944,1968]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDA284B783ABC99C8B9F812" box="[550,759,1944,1968]" italics="true" pageId="6" pageNumber="395">Naultinus flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDA284B78E2BC99C901F812" bold="true" box="[766,847,1944,1968]" pageId="6" pageNumber="395">
<taxonomicNameLabel id="A386C22BFFDA284B78E2BC99C901F812" box="[766,847,1944,1968]" pageId="6" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
RE003322 (original field number G.789): A.) dorsal, B.) ventral, and C.) lateral views. Photographs by J-C. Stahl. Scale bar = 1cm.
</paragraph>
</caption>
<caption id="DEBEF3CAFFDB284A7A8BBC96CECDF85A" ID-DOI="http://doi.org/10.5281/zenodo.4456590" ID-Zenodo-Dep="4456590" httpUri="https://zenodo.org/record/4456590/files/figure.png" pageId="7" pageNumber="396" startId="7.[151,250,1943,1967]" targetBox="[246,1313,181,1920]" targetPageId="7">
<paragraph id="8A7EA342FFDB284A7A8BBC96CECDF85A" blockId="7.[151,1437,1943,2040]" pageId="7" pageNumber="396">
<emphasis id="B8B57F50FFDB284A7A8BBC96CB5DF80D" bold="true" box="[151,275,1943,1967]" pageId="7" pageNumber="396">FIGURE 2.</emphasis>
Comparison of head scalation between relevant 
<taxonomicName id="4DC1D8C1FFDB284A7918BC99C929F812" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[772,871,1944,1968]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFDB284A7918BC99C929F812" box="[772,871,1944,1968]" italics="true" pageId="7" pageNumber="396">Naultinus</emphasis>
</taxonomicName>
spp.: A.) 
<taxonomicName id="4DC1D8C1FFDB284A79D7BC99CF6FF812" authority="(MONZ RE. 2013)" baseAuthorityName="MONZ RE." baseAuthorityYear="2013" box="[971,1313,1944,1968]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="manukanus">
<emphasis id="B8B57F50FFDB284A79D7BC99CE12F812" box="[971,1116,1944,1968]" italics="true" pageId="7" pageNumber="396">N. manukanus</emphasis>
(MONZ RE.2013)
</taxonomicName>
; B.) 
<taxonomicName id="4DC1D8C1FFDB284A7F4FBC99CA86F876" authorityName="Hutton" authorityYear="1872" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B8B57F50FFDB284A7F4FBC99CA86F876" italics="true" pageId="7" pageNumber="396">N. stellatus</emphasis>
</taxonomicName>
(RE.005617; original field number RAH466); C.) 
<taxonomicName id="4DC1D8C1FFDB284A78D3BCBDC91AF876" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[719,852,1980,2004]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFDB284A78D3BCBDC91AF876" box="[719,852,1980,2004]" italics="true" pageId="7" pageNumber="396">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFDB284A7947BCBDC9E2F876" bold="true" box="[859,940,1980,2004]" pageId="7" pageNumber="396">
<taxonomicNameLabel id="A386C22BFFDB284A7947BCBDC9E2F876" box="[859,940,1980,2004]" pageId="7" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); D.) 
<taxonomicName id="4DC1D8C1FFDB284A7E4EBCBDCEF3F876" authorityName="Gray" authorityYear="1842" box="[1106,1213,1980,2004]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B8B57F50FFDB284A7E4EBCBDCEF3F876" box="[1106,1213,1980,2004]" italics="true" pageId="7" pageNumber="396">N. elegans</emphasis>
</taxonomicName>
(RE.002658; original field number G.102); and E.) 
<taxonomicName id="4DC1D8C1FFDB284A7BD8BCE1C850F855" authorityName="Bell" authorityYear="1843" box="[452,542,2016,2039]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFDB284A7BD8BCE1C850F855" box="[452,542,2016,2039]" italics="true" pageId="7" pageNumber="396">N. grayii</emphasis>
</taxonomicName>
(RE.002771; original field number G.231). Photos by SVN.
</paragraph>
</caption>
<caption id="DEBEF3CAFFD428457A8BBE4FC84AFA28" ID-DOI="http://doi.org/10.5281/zenodo.4456592" ID-Zenodo-Dep="4456592" httpUri="https://zenodo.org/record/4456592/files/figure.png" pageId="8" pageNumber="397" startId="8.[151,250,1358,1382]" targetBox="[157,1431,181,1319]" targetPageId="8">
<paragraph id="8A7EA342FFD428457A8BBE4FC84AFA28" blockId="8.[151,1436,1358,1418]" pageId="8" pageNumber="397">
<emphasis id="B8B57F50FFD428457A8BBE4FCB5DFAC4" bold="true" box="[151,275,1358,1382]" pageId="8" pageNumber="397">FIGURE 3.</emphasis>
In-life photographs of two 
<taxonomicName id="4DC1D8C1FFD428457832BE4EC8FDFAC4" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[558,691,1358,1382]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="397" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFD428457832BE4EC8FDFAC4" box="[558,691,1358,1382]" italics="true" pageId="8" pageNumber="397">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFD4284578A6BE4FC945FAC4" bold="true" box="[698,779,1358,1382]" pageId="8" pageNumber="397">
<taxonomicNameLabel id="A386C22BFFD4284578A6BE4FC945FAC4" box="[698,779,1358,1382]" pageId="8" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
individuals from Te Paki (both male; not vouchered). Photos by SVN (A, B) and D. van Winkel (C).
</paragraph>
</caption>
<paragraph id="8A7EA342FFD428457ADBBEB9CB13F9C0" blockId="8.[151,1437,1464,2031]" pageId="8" pageNumber="397">
<emphasis id="B8B57F50FFD428457ADBBEB9CBF2FA70" bold="true" box="[199,444,1464,1490]" pageId="8" pageNumber="397">Conservation status.</emphasis>
This species is listed by Hitchmough 
<emphasis id="B8B57F50FFD428457971BEB8C9E8FA70" box="[877,934,1464,1490]" italics="true" pageId="8" pageNumber="397">et al.</emphasis>
(2016) under the 
<collectingCountry id="F2D6E3D2FFD428457E6EBEB8CF47FA70" box="[1138,1289,1464,1491]" name="New Zealand" pageId="8" pageNumber="397">New Zealand</collectingCountry>
Threat Classification System as “At Risk – Declining” on the basis of criterion C (2/1) (total area of occupancy&gt; 
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<superScript id="7DB40E0AFFD428457F91BEDDCFDBFA48" attach="left" box="[1421,1429,1500,1514]" fontSize="6" pageId="8" pageNumber="397">2</superScript>
, predicted decline 10–70%), and with the Qualifiers Data Poor and One Location. The One Location qualifier means that the species occupies a single area of continuous habitat, so a new threat could very quickly spread to affect the entire population.
</paragraph>
<paragraph id="8A7EA342FFD428457ADBBD6DCB55F8FC" blockId="8.[151,1437,1464,2031]" pageId="8" pageNumber="397">
Under the IUCN Redlist system, we suggest the species is Endangered under the criteria B1a,b(v), B2a,b(v): Geographic range in the form of (B1) extent of occurrence estimated to be (1) less than 
<quantity id="4D390EA7FFD428457E96BD91CEB6F908" box="[1162,1272,1680,1707]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="397" unit="km" value="5000.0">5,000 km</quantity>
2, and (a) estimates indicating severely fragmented or known to exist at no more than five locations and (b) continuing decline, observed, inferred or projected, in (v) number of mature individuals. Geographic range in the form of (B2) area of occupancy estimated to be (1) less than 
<quantity id="4D390EA7FFD42845784DBDFDC8EBF8B4" box="[593,677,1788,1815]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="397" unit="km" value="500.0">500 km</quantity>
<superScript id="7DB40E0AFFD4284578B9BDFDC8E3F8A8" attach="left" box="[677,685,1788,1802]" fontSize="6" pageId="8" pageNumber="397">2</superScript>
, and estimates indicating (a) severely fragmented or known to exist at no more than five locations and (b) continuing decline, observed, inferred or projected, in (v) number of mature individuals.
</paragraph>
<paragraph id="8A7EA342FFD428447ADBBC69CAA5FEE3" blockId="8.[151,1437,1464,2031]" lastBlockId="9.[151,1437,150,1257]" lastPageId="9" lastPageNumber="398" pageId="8" pageNumber="397">
The presumed original habitat of 
<taxonomicName id="4DC1D8C1FFD428457822BC68C89EF820" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[574,720,1896,1922]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="397" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFD428457822BC68C89EF820" box="[574,720,1896,1922]" italics="true" pageId="8" pageNumber="397">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFD4284578CBBC69C97FF820" bold="true" box="[727,817,1896,1922]" pageId="8" pageNumber="397">
<taxonomicNameLabel id="A386C22BFFD4284578CBBC69C97FF820" box="[727,817,1896,1922]" pageId="8" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, tall subtropical rainforest, has been almost entirely removed since human arrival in the 13 
<superScript id="7DB40E0AFFD428457833BC8DC872F838" attach="left" box="[559,572,1932,1946]" fontSize="6" pageId="8" pageNumber="397">th</superScript>
Century AD, and now only exists in a handful of tiny remnant patches (3.2% of the Ecological District; 
<bibRefCitation id="EE50DEB3FFD428457BDEBCB0C83AF868" author="Lux, J. &amp; Holland, W. &amp; Rate, S. &amp; Beadel, S." box="[450,628,1968,1995]" pageId="8" pageNumber="397" refId="ref8422" refString="Lux, J., Holland, W., Rate, S. &amp; Beadel, S. (2009) Natural areas of Te Paki Ecological District: reconnaissance survey report for the Protected Natural Areas Programme. Department of Conservation, Northland Conservancy, Whangarei, viii + 319 pp." type="book" year="2009">
Lux 
<emphasis id="B8B57F50FFD428457BEBBCB0C87EF868" box="[503,560,1968,1994]" italics="true" pageId="8" pageNumber="397">et al.</emphasis>
2009
</bibRefCitation>
). Fires set by Māori removed most of the original forest on the Aupōuri Peninsula soon after their arrival in 
<collectingCountry id="F2D6E3D2FFD42845783FBCD4C8F6F84C" box="[547,696,2004,2031]" name="New Zealand" pageId="8" pageNumber="397">New Zealand</collectingCountry>
in about the 12 
<superScript id="7DB40E0AFFD42845797EBCD5C921F840" attach="left" box="[866,879,2004,2018]" fontSize="6" pageId="8" pageNumber="397">th</superScript>
Century CE; initially it was replaced by extensive areas of bracken fern (
<taxonomicName id="4DC1D8C1FFD528447B88BB96C9B9FF13" authority="(G. Forster)) (Elliot et al. 1995)" baseAuthorityName="G. Forster" box="[404,1015,150,177]" class="Polypodiopsida" family="Dennstaedtiaceae" genus="Pteridium" kingdom="Animalia" order="Polypodiales" pageId="9" pageNumber="398" phylum="Tracheophyta" rank="species" species="esculentum">
<emphasis id="B8B57F50FFD528447B88BB96C8C7FF13" box="[404,649,151,177]" italics="true" pageId="9" pageNumber="398">Pteridium esculentum</emphasis>
(G. Forster)) (
<bibRefCitation id="EE50DEB3FFD528447930BB96C9A0FF13" author="Elliot, M. B. &amp; Striewski, B. &amp; Flenley, J. R. &amp; Sutton, D. G." box="[812,1006,150,177]" pageId="9" pageNumber="398" pagination="899 - 916" refId="ref7961" refString="Elliot, M. B., Striewski, B., Flenley, J. R. &amp; Sutton, D. G. (1995) Palynological and sedimentological evidence for a radiocarbon chronology of environmental change and Polynesian deforestation from Lake Taumatawhana, Northland, New Zealand. Radiocarbon, 37, 899 - 916. https: // doi. org / 10.1017 / S 0033822200014983" type="journal article" year="1995">
Elliot 
<emphasis id="B8B57F50FFD52844796FBB96C9E2FF13" box="[883,940,151,177]" italics="true" pageId="9" pageNumber="398">et al.</emphasis>
1995
</bibRefCitation>
)
</taxonomicName>
. The Te Paki area is now largely covered in fire-maintained secondary shrublands dominated by 
<taxonomicName id="4DC1D8C1FFD52844795FBBBACE7EFF77" authority="Reichenbach" authorityName="Reichenbach" box="[835,1072,187,213]" class="Magnoliopsida" family="Myrtaceae" genus="Kunzea" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="398" phylum="Tracheophyta" rank="genus">
<emphasis id="B8B57F50FFD52844795FBBBAC9D9FF76" box="[835,919,187,212]" italics="true" pageId="9" pageNumber="398">Kunzea</emphasis>
Reichenbach
</taxonomicName>
and 
<taxonomicName id="4DC1D8C1FFD528447E76BBBACBD1FF5B" authority="J. R. Forster &amp; G. Forster. However" authorityName="J. R. Forster &amp; G. Forster. However" class="Magnoliopsida" family="Myrtaceae" genus="Leptospermum" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="398" phylum="Tracheophyta" rank="genus">
<emphasis id="B8B57F50FFD528447E76BBBACF5EFF76" box="[1130,1296,187,212]" italics="true" pageId="9" pageNumber="398">Leptospermum</emphasis>
J.R. Forster &amp; G. Forster. However
</taxonomicName>
, this shrubland is readily occupied by 
<taxonomicName id="4DC1D8C1FFD528447946BBDEC989FF5B" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[858,967,223,249]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFD528447946BBDEC989FF5B" box="[858,967,223,249]" italics="true" pageId="9" pageNumber="398">Naultinus</emphasis>
</taxonomicName>
, so the total area of habitat has probably changed relatively little. Only parts of the available habitat are known to be occupied, probably as a result of fire history.
</paragraph>
<paragraph id="8A7EA342FFD528447ADBBA4ACE6DFD53" blockId="9.[151,1437,150,1257]" pageId="9" pageNumber="398">
With fire now better controlled, threats are almost entirely from invasive species. Almost the full suite of small predatory introduced mammals which have decimated native lizard populations across mainland 
<collectingCountry id="F2D6E3D2FFD528447EF5BA6ECFCFFE2B" box="[1257,1409,367,393]" name="New Zealand" pageId="9" pageNumber="398">New Zealand</collectingCountry>
is present and common: cats (
<taxonomicName id="4DC1D8C1FFD528447BC8BA92C88BFE0C" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[468,709,403,430]" class="Mammalia" family="Felidae" genus="Felis" kingdom="Animalia" order="Carnivora" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="catus">
<emphasis id="B8B57F50FFD528447BC8BA92C81FFE0E" box="[468,593,403,429]" italics="true" pageId="9" pageNumber="398">Felis catus</emphasis>
Linnaeus
</taxonomicName>
), hedgehogs (
<taxonomicName id="4DC1D8C1FFD528447979BA92CE86FE0C" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[869,1224,403,430]" class="Mammalia" family="Erinaceidae" genus="Erinaceus" kingdom="Animalia" order="Erinaceomorpha" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="europaeus">
<emphasis id="B8B57F50FFD528447979BA92CE1AFE0E" box="[869,1108,403,428]" italics="true" pageId="9" pageNumber="398">Erinaceus europaeus</emphasis>
Linnaeus
</taxonomicName>
), brush-tailed possums (
<taxonomicName id="4DC1D8C1FFD528447AFDBAB6C87DFE73" authority="(Kerr))" authorityName="" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[225,563,439,465]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="vulpecula">
<emphasis id="B8B57F50FFD528447AFDBAB6CB94FE73" box="[225,474,439,465]" italics="true" pageId="9" pageNumber="398">Trichosurus vulpecula</emphasis>
(Kerr))
</taxonomicName>
, stoats (
<taxonomicName id="4DC1D8C1FFD528447892BAB6C9F2FE70" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[654,956,439,466]" class="Mammalia" family="Mustelidae" genus="Mustela" kingdom="Animalia" order="Carnivora" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="erminea">
<emphasis id="B8B57F50FFD528447892BAB6C905FE72" box="[654,843,439,465]" italics="true" pageId="9" pageNumber="398">Mustela erminea</emphasis>
Linnaeus
</taxonomicName>
), weasels (
<taxonomicName id="4DC1D8C1FFD528447E2ABAB6CF1DFE70" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1766" box="[1078,1363,439,466]" class="Mammalia" family="Mustelidae" genus="Mustela" kingdom="Animalia" order="Carnivora" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="nivalis">
<emphasis id="B8B57F50FFD528447E2ABAB6CEACFE73" box="[1078,1250,439,465]" italics="true" pageId="9" pageNumber="398">Mustela nivalis</emphasis>
Linnaeus
</taxonomicName>
), mice (
<taxonomicName id="4DC1D8C1FFD528447ABDBADACBFFFE54" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[161,433,475,502]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="musculus">
<emphasis id="B8B57F50FFD528447ABDBADACB71FE57" box="[161,319,475,501]" italics="true" pageId="9" pageNumber="398">Mus musculus</emphasis>
Linnaeus
</taxonomicName>
), and rats (both 
<taxonomicName id="4DC1D8C1FFD528447878BADAC93AFE54" authority="rattus (Linnaeus)" baseAuthorityName="Linnaeus" box="[612,884,475,502]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="rattus">
<emphasis id="B8B57F50FFD528447878BADAC8BDFE56" box="[612,755,475,500]" italics="true" pageId="9" pageNumber="398">Rattus rattus</emphasis>
(Linnaeus)
</taxonomicName>
and 
<taxonomicName id="4DC1D8C1FFD5284479B6BADACEA3FE54" authority="(Berkenhout))" authorityName="" baseAuthorityName="Berkenhout" baseAuthorityYear="1769" box="[938,1261,475,502]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="norvegicus">
<emphasis id="B8B57F50FFD5284479B6BADACE0DFE56" box="[938,1091,475,500]" italics="true" pageId="9" pageNumber="398">R. norvegicus</emphasis>
(Berkenhout))
</taxonomicName>
. Feral pigs (
<emphasis id="B8B57F50FFD528447F69BADACA92FDBB" italics="true" pageId="9" pageNumber="398">Sus scrofa</emphasis>
Linnaeus), which have also recently been demonstrated to eat large numbers of native lizards (J. Reardon pers. comm.), are widespread and abundant. Te Paki was one of the last areas of the country reached by possums, which have become abundant only in the last 20 years. In 1985 they were still recorded as absent but were spreading rapidly northward up the Aupōuri Peninsula, where they had reached about 
<quantity id="4D390EA7FFD528447E30B96ACE3BFD27" box="[1068,1141,619,645]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="398" unit="km" value="10.0">10 km</quantity>
south of Waitiki Landing (
<bibRefCitation id="EE50DEB3FFD528447A83B98ECB0FFD0B" author="Mitchell, N." box="[159,321,654,681]" pageId="9" pageNumber="398" pagination="11 - 12" refId="ref8653" refString="Mitchell, N. (1985) The northward movement of possums. New Zealand Ecological Society Newsletter, 43, 11 - 12." type="journal article" year="1985">Mitchell 1985</bibRefCitation>
). The relatively abundant (compared to neighbouring mainland areas) 
<taxonomicName id="4DC1D8C1FFD528447E57B98ECF5AFD0B" authorityName="Gray" authorityYear="1842" box="[1099,1300,655,681]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B8B57F50FFD528447E57B98ECF5AFD0B" box="[1099,1300,655,681]" italics="true" pageId="9" pageNumber="398">Naultinus elegans</emphasis>
</taxonomicName>
populations on Waiheke Island, where possums are absent but most other invasive small mammals are abundant, suggests that arboreal possums may be particularly significant predators of arboreal 
<taxonomicName id="4DC1D8C1FFD5284479B3B9D6CE52FD53" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[943,1052,727,753]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFD5284479B3B9D6CE52FD53" box="[943,1052,727,753]" italics="true" pageId="9" pageNumber="398">Naultinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8A7EA342FFD528447ADBB9FAC9C4FB97" blockId="9.[151,1437,150,1257]" pageId="9" pageNumber="398">
In addition to the threats from mammalian predators, social insects are an emerging threat. Introduced yellowjacket (
<taxonomicName id="4DC1D8C1FFD528447B09B81ECB93FC9B" authority="Thomson" authorityName="Thomson" box="[277,477,799,825]" class="Insecta" family="Vespidae" genus="Vespula" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B8B57F50FFD528447B09B81ECB22FC9B" box="[277,364,799,825]" italics="true" pageId="9" pageNumber="398">Vespula</emphasis>
Thomson
</taxonomicName>
spp.) and paper wasps (
<taxonomicName id="4DC1D8C1FFD5284478F1B81EC9E2FC9B" authority="Latreille" authorityName="Latreille" authorityYear="1802" box="[749,940,799,825]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="B8B57F50FFD5284478F1B81EC908FC9B" box="[749,838,799,825]" italics="true" pageId="9" pageNumber="398">Polistes</emphasis>
Latreille
</taxonomicName>
spp.) are now very abundant in Te Paki and prey on the same insect species as 
<taxonomicName id="4DC1D8C1FFD528447802B842C906FCFF" authority="(Dymock 2000)" baseAuthorityName="Dymock" baseAuthorityYear="2000" box="[542,840,835,861]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFD528447802B842C8C5FCFF" box="[542,651,835,861]" italics="true" pageId="9" pageNumber="398">Naultinus</emphasis>
(
<bibRefCitation id="EE50DEB3FFD528447886B842C90EFCFF" author="Dymock, J. J." box="[666,832,835,861]" pageId="9" pageNumber="398" refId="ref7898" refString="Dymock, J. J. (2000) Risk assessment for establishment of polistine (Polistes spp.) and vespine (Vespula spp.) wasps on the Three Kings Islands in the far north of New Zealand. In: Science for Conservation. Vol. 156. Department of Conservation, Wellington, pp. 1 - 18." type="book" year="2000">Dymock 2000</bibRefCitation>
)
</taxonomicName>
. 
<collectingCountry id="F2D6E3D2FFD52844794EB842C98FFCFF" box="[850,961,835,861]" name="Argentina" pageId="9" pageNumber="398">Argentine</collectingCountry>
ants (
<taxonomicName id="4DC1D8C1FFD528447E1AB842CF08FCFF" authority="(Mayr))" baseAuthorityName="Mayr" box="[1030,1350,835,861]" class="Insecta" family="Formicidae" genus="Linepithema" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="398" phylum="Arthropoda" rank="species" species="humile">
<emphasis id="B8B57F50FFD528447E1AB842CEA8FCFF" box="[1030,1254,835,861]" italics="true" pageId="9" pageNumber="398">Linepithema humile</emphasis>
(Mayr))
</taxonomicName>
(both a competitor and a predator) are beginning to invade the area (https://www.doc.govt.nz/news/media-releases/2011/ doc-in-far-north-winning-battle-against-ruthless-ecological-invader/). The invasive Australian plague skink (
<taxonomicName id="4DC1D8C1FFD528447F4FB88ACB8DFC68" authority="(De Vis))" authorityName="" baseAuthorityName="De Vis" baseAuthorityYear="1888" class="Reptilia" family="Scincidae" genus="Lampropholis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="delicata">
<emphasis id="B8B57F50FFD528447F4FB88ACB1AFC6B" italics="true" pageId="9" pageNumber="398">Lampropholis delicata</emphasis>
(De Vis))
</taxonomicName>
is spreading rapidly up the Aupōuri Peninsula (Kamera Raharaha, (dec.) pers. comm. to RAH) and is likely to invade Te Paki within the next few years. This species reaches enormous population densities, particularly near its invasion front (
<bibRefCitation id="EE50DEB3FFD528447835B8F6C97FFBB3" author="van Winkel, D. &amp; Baling, M. &amp; Hitchmough, R." box="[553,817,1015,1041]" pageId="9" pageNumber="398" refId="ref9042" refString="van Winkel, D., Baling, M. &amp; Hitchmough, R. (2018) Reptiles and Amphibians of New Zealand, a Field Guide. Auckland University Press, Auckland, 366 pp." type="book" year="2018">
van Winkel 
<emphasis id="B8B57F50FFD5284478A8B8F6C8A8FBB3" box="[692,742,1015,1041]" italics="true" pageId="9" pageNumber="398">et al</emphasis>
. 2018
</bibRefCitation>
), and is likely to be a serious competitor for 
<taxonomicName id="4DC1D8C1FFD528447F33B8F6CFD2FBB3" baseAuthorityName="Dymock" baseAuthorityYear="2000" box="[1327,1436,1015,1041]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="genus">
<emphasis id="B8B57F50FFD528447F33B8F6CFD2FBB3" box="[1327,1436,1015,1041]" italics="true" pageId="9" pageNumber="398">Naultinus</emphasis>
</taxonomicName>
and other native lizards in the area, despite being mainly terrestrial.
</paragraph>
</subSubSection>
<subSubSection id="C2DBF0C9FFD528447ADBBF3FCB15FB4B" pageId="9" pageNumber="398" type="discussion">
<paragraph id="8A7EA342FFD528447ADBBF3FCB15FB4B" blockId="9.[151,1437,150,1257]" pageId="9" pageNumber="398">
<emphasis id="B8B57F50FFD528447ADBBF3FCB01FBFA" bold="true" box="[199,335,1086,1112]" pageId="9" pageNumber="398">Comments.</emphasis>
When 
<bibRefCitation id="EE50DEB3FFD528447BB9BF3EC8A5FBFB" author="Robb, J. &amp; Hitchmough, R. A." box="[421,747,1087,1113]" pageId="9" pageNumber="398" pagination="189 - 200" refId="ref8947" refString="Robb, J. &amp; Hitchmough, R. A. (1980) Review of the genus Naultinus Gray (Reptilia: Gekkonidae). Records of the Auckland Institute and Museum, 16, 189 - 200." type="journal article" year="1980">Robb &amp; Hitchmough (1980)</bibRefCitation>
resurrected 
<taxonomicName id="4DC1D8C1FFD528447966BF3EC9AEFBFA" authorityName="Bell" authorityYear="1843" box="[890,992,1087,1112]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="grayii">
<emphasis id="B8B57F50FFD528447966BF3EC9AEFBFA" box="[890,992,1087,1112]" italics="true" pageId="9" pageNumber="398">N. grayii</emphasis>
</taxonomicName>
from synonymy with 
<taxonomicName id="4DC1D8C1FFD528447EFFBF3ECF12FBFB" authorityName="Gray" authorityYear="1842" box="[1251,1372,1087,1113]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B8B57F50FFD528447EFFBF3ECF12FBFB" box="[1251,1372,1087,1113]" italics="true" pageId="9" pageNumber="398">N. elegans</emphasis>
</taxonomicName>
, they included MONZ RE003322, the 
<typeStatus id="557A1DE0FFD52844781ABF62C826FBDF" box="[518,616,1123,1149]" pageId="9" pageNumber="398" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4DC1D8C1FFD528447890BF62C950FBDF" authority="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer, 2021" authorityName="Hitchmough &amp; Nielsen &amp; Lysaght &amp; Bauer" authorityYear="2021" box="[652,798,1123,1149]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="flavirictus" status="sp. nov.">
<emphasis id="B8B57F50FFD528447890BF62C950FBDF" box="[652,798,1123,1149]" italics="true" pageId="9" pageNumber="398">N. flavirictus</emphasis>
</taxonomicName>
<emphasis id="B8B57F50FFD528447939BF62C930FBDF" bold="true" box="[805,894,1123,1149]" pageId="9" pageNumber="398">
<taxonomicNameLabel id="A386C22BFFD528447939BF62C930FBDF" box="[805,894,1123,1149]" pageId="9" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, among the material examined. Its mouth colour was not visible, but the smaller size, smaller snout scales and blunter snout were all noted; however,the small size of the specimen was interpreted as an indication that it was immature or subadult, and this was thought to explain these differences.
</paragraph>
</subSubSection>
</treatment>
</document>