450 lines
76 KiB
XML
450 lines
76 KiB
XML
<document id="22626CBFE81294CFB1DD11D9D43A2675" ID-CLB-Dataset="86221" ID-DOI="10.11646/zootaxa.5159.3.1" ID-GBIF-Dataset="e19dda1c-86af-404d-9595-c08d8c848abd" ID-ISSN="1175-5326" ID-Zenodo-Dep="6781669" ID-ZooBank="13308111-EF49-4710-9C45-CF69DABE2C5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656566366326" checkinUser="plazi" docAuthor="Kataev, Boris M., Hongbin, Liang & Wrase, David W." docDate="2022" docId="D364737A243E1727FF60B5EF16A718C1" docLanguage="en" docName="zootaxa.5159.3.1.pdf" docOrigin="Zootaxa 5159 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus Kataev, Hongbin & Wrase, 2022, s. str." docType="key" docVersion="5" lastPageNumber="352" masterDocId="2F5D0B02240C1714FFF7B578133B1A36" masterDocTitle="New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies" masterLastPageNumber="353" masterPageNumber="301" pageNumber="351" updateTime="1699353733348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="47A0BAB9A8B7D5D81DDC00B7CF78072C">New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies</mods:title>
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<mods:name id="09CB2ABE5FC787F8BFEAE77BAE7D9AB6" type="personal">
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<mods:roleTerm id="C316F821EE446232B0F0BBF1D2F996AC">Author</mods:roleTerm>
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<mods:namePart id="A58C511BC8F0D8CC8EC590353D197C64">Kataev, Boris M.</mods:namePart>
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<mods:affiliation id="536A60A2CC358307CD12725C911EB6AB">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="F008AA57F7566817572082AD562B03D0">Hongbin, Liang</mods:namePart>
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<mods:nameIdentifier id="15B9FDFA2F0A1DCE758F08E553A07D2B" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
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<mods:affiliation id="C2488E3EB4165D9A81FD09CC016C1C4F">Key laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. lianghb @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 9668 - 1167</mods:affiliation>
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<mods:nameIdentifier id="96440DB888AB6A751937FE7AC21878DC" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:namePart id="214E04129157B40493FFE284C5E3DAF9">Wrase, David W.</mods:namePart>
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<mods:affiliation id="F218084D6483FDF7097F06129AFDFD5B">Oderstr. 2, D- 15306 Gusow-Platkow, Germany.</mods:affiliation>
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<mods:title id="2B5391DE4D65D80F55B8DC1E236EC013">Zootaxa</mods:title>
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<mods:part id="9BCE3F278C4BDE81B3CA4004211B4F85">
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<mods:date id="245A17E5CC8A454B12EF1432299B3F8E">2022</mods:date>
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<mods:number id="ED252994AA4B678C50E8D47F5D8A92B4">2022-06-30</mods:number>
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<treatment id="D364737A243E1727FF60B5EF16A718C1" ID-DOI="http://doi.org/10.5281/zenodo.6787380" ID-GBIF-Taxon="196373180" ID-Zenodo-Dep="6787380" LSID="urn:lsid:plazi:treatment:D364737A243E1727FF60B5EF16A718C1" httpUri="http://treatment.plazi.org/id/D364737A243E1727FF60B5EF16A718C1" lastPageId="51" lastPageNumber="352" pageId="50" pageNumber="351">
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<subSubSection id="13D791E7243E1726FF60B5EF10841A87" box="[151,959,151,178]" pageId="50" pageNumber="351" type="nomenclature">
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<paragraph id="5B72C26C243E1726FF60B5EF10841A87" blockId="50.[151,959,151,178]" box="[151,959,151,178]" pageId="50" pageNumber="351">
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<heading id="003A7500243E1726FF60B5EF10841A87" bold="true" box="[151,959,151,178]" fontSize="11" level="1" pageId="50" pageNumber="351" reason="1">
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<emphasis id="69B91E7E243E1726FF60B5EF10841A87" bold="true" box="[151,959,151,178]" pageId="50" pageNumber="351">
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A preliminary key to species of
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<taxonomicName id="9CCDB9EF243E1726FDE8B5EF11F21A84" ID-CoL="8KW2X" authorityName="Morawitz" authorityYear="1863" box="[543,713,151,178]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="69B91E7E243E1726FDE8B5EF11F21A84" bold="true" box="[543,713,151,178]" italics="true" pageId="50" pageNumber="351">Trichotichnus</emphasis>
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<taxonomicNameLabel id="728AA305243E1726FD26B5EF10291A87" box="[721,786,151,177]" pageId="50" pageNumber="351" sensu="stricto">s. str.</taxonomicNameLabel>
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from
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<collectingRegion id="99090C8E243E1726FCAAB5EF10841A87" box="[861,959,151,177]" country="China" name="Yunnan" pageId="50" pageNumber="351">Yunnan</collectingRegion>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="13D791E7243E1726FF60B5A412B71B0B" pageId="50" pageNumber="351" type="discussion">
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<paragraph id="5B72C26C243E1726FF60B5A412B71B0B" blockId="50.[151,1436,220,317]" pageId="50" pageNumber="351">
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Since the fauna of
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<collectingRegion id="99090C8E243E1726FE9EB5A512841AC1" box="[361,447,221,247]" country="China" name="Yunnan" pageId="50" pageNumber="351">Yunnan</collectingRegion>
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is still poorly known and distinctive characters of external morphology in many species are variable both individually and geographically, the examination of male genitalia is very desirable for accurate species identification.
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</paragraph>
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</subSubSection>
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<subSubSection id="13D791E7243E1727FF60B41016A718C1" lastPageId="51" lastPageNumber="352" pageId="50" pageNumber="351" type="key">
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<keyLead id="E03C6159243E1726FF60B41016A71BE5" pageId="50" pageNumber="351">
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1 Legs relatively short (as in
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<figureCitation id="C3F6DEE9243E1726FE1EB410117F1B4B" box="[489,580,360,382]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1674,1698]" captionTargetBox="[167,1435,450,1643]" captionTargetId="figure-134@5.[151,1436,450,1648]" captionTargetPageId="5" captionText="FIGURES 1–2. Trichotichnus anthracinus Landin, 1955, habitus (1, Yunnan; 2, Myanmar, holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781671" httpUri="https://zenodo.org/record/6781671/files/figure.png" pageId="50" pageNumber="351">Figs. 1, 2</figureCitation>
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,
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<figureCitation id="C3F6DEE9243E1726FDB9B41011BA1B4B" box="[590,641,360,381]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">8–10</figureCitation>
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,
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<figureCitation id="C3F6DEE9243E1726FD7CB41011F21B48" box="[651,713,360,382]" captionStart="FIGURES 35–37" captionStartId="20.[151,264,1335,1359]" captionTargetBox="[158,1428,536,1307]" captionTargetId="figure-178@20.[150,1437,525,1312]" captionTargetPageId="20" captionText="FIGURES 35–37. Trichotichnus spp., habitus. 35, T. cyrtops (Tschitschérine, 1906) (Yunnan); 36, T. doiinthanonensis Ito, 1997 (Dulongjiang); 37, T. fedorenkoi sp. n. (Tam Dao). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781715" httpUri="https://zenodo.org/record/6781715/files/figure.png" pageId="50" pageNumber="351">35–37</figureCitation>
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,
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<figureCitation id="C3F6DEE9243E1726FD24B410102A1B48" box="[723,785,360,382]" captionStart="FIGURES 56–58" captionStartId="27.[151,264,1121,1145]" captionTargetBox="[168,1428,182,1096]" captionTargetId="figure-19@27.[158,1430,181,1098]" captionTargetPageId="27" captionText="FIGURES 56–58. Trichotichnus schmidti sp. n. (Yunnan: 56 & 57, Nankang; 58, S Tengchong), habitus. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781731" httpUri="https://zenodo.org/record/6781731/files/figure.png" pageId="50" pageNumber="351">56–58</figureCitation>
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,
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<figureCitation id="C3F6DEE9243E1726FCECB41010611B48" box="[795,858,360,382]" captionStart="FIGURES 74–76" captionStartId="32.[151,264,1767,1791]" captionTargetBox="[167,1418,922,1737]" captionTargetId="figure-307@32.[158,1429,916,1744]" captionTargetPageId="32" captionText="FIGURES 74–76. Trichotichnus spp., habitus. 74, T. subangulatus Ito, 2021 (NNE Tengchong); 75. T.? subangulatus Ito, 2021 (S Longyuan); 76, T. noctuabundus Habu, 1954 (Vietnam). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781747" httpUri="https://zenodo.org/record/6781747/files/figure.png" pageId="50" pageNumber="351">74–76</figureCitation>
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), pro- and mesotarsomeres
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both sexes with comparatively sparse moderately long ventro-lateral setae (as in
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<figureCitation id="C3F6DEE9243E1726FCF2B4FD10401BAC" box="[773,891,389,411]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Figs. 18, 19</figureCitation>
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). Pronotum not cordiform, in most specimens relatively wider (PWmax/PL 1.45–1.63), with flat or only moderately deepened laterobasal depressions. Legs unicoloured pale or with infuscate femora. Protibia with or without longitudinal sulcus on dorsal side...................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="5B72C26C243E1726FF60B4A316A7181E" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
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<keyLead id="E03C6159243E1726FF60B4A316A7181E" pageId="50" pageNumber="351">
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- Legs relatively long (as in
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<figureCitation id="C3F6DEE9243E1726FE10B4A211671BC6" box="[487,604,474,496]" captionStart="FIGURES 94–95" captionStartId="38.[151,264,1747,1771]" captionTargetBox="[174,1411,469,1715]" captionTargetId="figure-151@38.[158,1429,453,1724]" captionTargetPageId="38" captionText="FIGURES 94–95. Trichotichnus hamulipenis sp. n., habitus. 94, nominotypical subspecies (above Bapo); 95, T. hamulipenis nepalensis ssp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781755" httpUri="https://zenodo.org/record/6781755/files/figure.png" pageId="50" pageNumber="351">Figs. 94, 95</figureCitation>
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,
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<figureCitation id="C3F6DEE9243E1726FD97B4A211831BD9" box="[608,696,474,496]" captionStart="FIGURES 107–108" captionStartId="43.[151,264,1956,1980]" captionTargetBox="[157,1420,650,1924]" captionTargetId="figure-222@43.[153,1433,633,1933]" captionTargetPageId="43" captionText="FIGURES 107–108. Trichotichnus ssp., habitus. 107, T. miwai continentalis ssp. n. (Wuliangshan); 108, T. kavanaughi sp. n. (Kongdang). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="50" pageNumber="351">107, 108</figureCitation>
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), pro- and mesotarsomeres 1–4, particularly in male, with longer and denser ventro-lateral setae (as in
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<figureCitation id="C3F6DEE9243E1726FE11B48F110A183A" box="[486,561,503,525]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 20</figureCitation>
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). Pronotum more or less cordiform, in most specimens relatively narrower (PWmax/PL 1.34–1.48), with deep basolateral depressions. Femora infuscate. Protibia without longitudinal sulcus on dorsal side..... 12
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="E039DAC9243E1726FF60B74816A718AD" pageId="50" pageNumber="351">
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<paragraph id="5B72C26C243E1726FF60B74816A71857" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
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<keyLead id="E03C6159243E1726FF60B74816A71857" pageId="50" pageNumber="351">
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2 Protibia with longitudinal sulcus on dorsal side. Mesotarsomere 1 of male without adhesive scales ventrally. Preapical sinuation of elytra very shallow (as in
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<figureCitation id="C3F6DEE9243E1726FE03B73411001857" box="[500,571,588,610]" captionStart="FIGURES 38–39" captionStartId="21.[151,264,1041,1065]" captionTargetBox="[151,1436,669,1016]" captionTargetId="figure-229@21.[150,1437,669,1018]" captionTargetPageId="21" captionText="FIGURES 38–39. Trichotichnus spp., elytral apex. 38, T. cyrtops (Tschitschérine, 1906); 39, T. schmidti sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781717" httpUri="https://zenodo.org/record/6781717/files/figure.png" pageId="50" pageNumber="351">Fig. 39</figureCitation>
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). Legs unicoloured brownish yellow, femora not darker than tibiae................. 3
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</keyLead>
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</paragraph>
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<paragraph id="5B72C26C243E1726FF60B71116A718AD" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
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<keyLead id="E03C6159243E1726FF60B71116A718AD" pageId="50" pageNumber="351">- Protibia without longitudinal sulcus on dorsal side. Mesotarsomere 1 of male with adhesive scales at apex ventrally. Preapical sinuation of elytra shallow or rather deep. Legs unicoloured pale or with infuscate femora........................... 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="E039DAC9243E1726FF60B7DA16A7191F" pageId="50" pageNumber="351">
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<paragraph id="5B72C26C243E1726FF60B7DA16A718C6" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
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<keyLead id="E03C6159243E1726FF60B7DA16A718C6" pageId="50" pageNumber="351">
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3 Left mandible in frontal view wide and truncate at tip (
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<figureCitation id="C3F6DEE9243E1726FCF6B7DA1070188E" box="[769,843,674,696]" captionStart="FIGURES 53–55" captionStartId="25.[151,264,1845,1869]" captionTargetBox="[156,1410,1314,1813]" captionTargetId="figure-908@25.[155,1433,1314,1820]" captionTargetPageId="25" captionText="FIGURES 53–55. Trichotichnus spp., head, dorso-frontal view. 53, T. schmidti sp. n.; 54, T. noctuabundus Habu, 1954; 55, T. subangulatus Ito, 2021. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781729" httpUri="https://zenodo.org/record/6781729/files/figure.png" pageId="50" pageNumber="351">Fig. 55</figureCitation>
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). Labrum more or less markedly concave anteriorly. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FE79B7C6113918E2" box="[398,514,702,724]" captionStart="FIGURES 77–86" captionStartId="33.[151,264,1903,1927]" captionTargetBox="[175,1410,459,1868]" captionTargetId="figure-134@33.[159,1429,453,1880]" captionTargetPageId="33" captionText="FIGURES 77–86. Trichotichnus subangulatus Ito, 2021 (77–79, 82 & 83, Yunnan; 80 & 81, Vietnam; 84–86, T.? subangulatus, Yunnan, Elidang Vill.); 77, 84, terminal lamella of aedeagus; 78–81, 85 & 86, median lobe of aedeagus; 82, gonocoxite; 83, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (77, 79, 81, 84 & 86), lateral (78, 80, 82 & 85) and ventral (83) views. Scale bars: A = 0.5 mm (77, 82–84), B = 1.0 mm (78–81, 85 & 86)." figureDoi="http://doi.org/10.5281/zenodo.6787363" httpUri="https://zenodo.org/record/6787363/files/figure.png" pageId="50" pageNumber="351">Figs. 77–81</figureCitation>
|
||
) with very apex not bent dorsally or ventrally, internal sac with narrow spiny patches along sides apically; female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FDFAB7A311AF18C6" box="[525,660,731,753]" captionStart="FIGURES 77–86" captionStartId="33.[151,264,1903,1927]" captionTargetBox="[175,1410,459,1868]" captionTargetId="figure-134@33.[159,1429,453,1880]" captionTargetPageId="33" captionText="FIGURES 77–86. Trichotichnus subangulatus Ito, 2021 (77–79, 82 & 83, Yunnan; 80 & 81, Vietnam; 84–86, T.? subangulatus, Yunnan, Elidang Vill.); 77, 84, terminal lamella of aedeagus; 78–81, 85 & 86, median lobe of aedeagus; 82, gonocoxite; 83, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (77, 79, 81, 84 & 86), lateral (78, 80, 82 & 85) and ventral (83) views. Scale bars: A = 0.5 mm (77, 82–84), B = 1.0 mm (78–81, 85 & 86)." figureDoi="http://doi.org/10.5281/zenodo.6787363" httpUri="https://zenodo.org/record/6787363/files/figure.png" pageId="50" pageNumber="351">Figs. 82 & 83</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FCD7B7A3105C18C6" box="[800,871,731,753]" captionStart="FIGURES 74–76" captionStartId="32.[151,264,1767,1791]" captionTargetBox="[167,1418,922,1737]" captionTargetId="figure-307@32.[158,1429,916,1744]" captionTargetPageId="32" captionText="FIGURES 74–76. Trichotichnus spp., habitus. 74, T. subangulatus Ito, 2021 (NNE Tengchong); 75. T.? subangulatus Ito, 2021 (S Longyuan); 76, T. noctuabundus Habu, 1954 (Vietnam). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781747" httpUri="https://zenodo.org/record/6781747/files/figure.png" pageId="50" pageNumber="351">Fig. 74</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FC1AB7A3175B18C6" box="[1005,1120,731,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.6" metricValueMin="7.7" pageId="50" pageNumber="351" unit="mm" value="8.65" valueMax="9.6" valueMin="7.7">7.7–9.6 mm</quantity>
|
||
....
|
||
<taxonomicName id="9CCDB9EF243E1726FB5FB7A416A718C6" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[1192,1436,731,753]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="subangulatus">
|
||
<emphasis id="69B91E7E243E1726FB5FB7A4167B18C6" box="[1192,1344,731,753]" italics="true" pageId="50" pageNumber="351">T. subangulatus</emphasis>
|
||
Ito, 2021
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B78016A7191F" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B78016A7191F" pageId="50" pageNumber="351">
|
||
- Left mandible in frontal view markedly narrowed apically, blunted at tip (
|
||
<figureCitation id="C3F6DEE9243E1726FC51B78F10D5193B" box="[934,1006,759,781]" captionStart="FIGURES 53–55" captionStartId="25.[151,264,1845,1869]" captionTargetBox="[156,1410,1314,1813]" captionTargetId="figure-908@25.[155,1433,1314,1820]" captionTargetPageId="25" captionText="FIGURES 53–55. Trichotichnus spp., head, dorso-frontal view. 53, T. schmidti sp. n.; 54, T. noctuabundus Habu, 1954; 55, T. subangulatus Ito, 2021. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781729" httpUri="https://zenodo.org/record/6781729/files/figure.png" pageId="50" pageNumber="351">Fig. 53</figureCitation>
|
||
). Apical margin of labrum almost straight, at most only slightly concave medially. Male and female genitalia not as above...................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B64916A7198E" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B64916A71949" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B64916A71949" pageId="50" pageNumber="351">
|
||
4 Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FE2AB649116B1970" box="[477,592,816,839]" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 & 50, terminal lamella of aedeagus; 48, 49, 51 & 52, median lobe of aedeagus. Dorsal (47, 49, 50 & 52) and lateral (48 & 51) views. Scale bars: A = 0.5 mm (47 & 50), B = 1.0 mm (48, 49, 51 & 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="50" pageNumber="351">Figs. 47–52</figureCitation>
|
||
) in lateral view slenderer, angularly bent ventrally just at apex and slightly swollen at tip forming a small apical capitulum; internal sac without distinct spiny patches. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FBBAB63517AF1955" box="[1101,1172,845,867]" captionStart="FIGURES 35–37" captionStartId="20.[151,264,1335,1359]" captionTargetBox="[158,1428,536,1307]" captionTargetId="figure-178@20.[150,1437,525,1312]" captionTargetPageId="20" captionText="FIGURES 35–37. Trichotichnus spp., habitus. 35, T. cyrtops (Tschitschérine, 1906) (Yunnan); 36, T. doiinthanonensis Ito, 1997 (Dulongjiang); 37, T. fedorenkoi sp. n. (Tam Dao). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781715" httpUri="https://zenodo.org/record/6781715/files/figure.png" pageId="50" pageNumber="351">Fig. 36</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FAEEB63516B71955" box="[1305,1420,845,867]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="50" pageNumber="351" unit="mm" value="8.35" valueMax="8.5" valueMin="8.2">8.2–8.5 mm</quantity>
|
||
.............................................................................
|
||
<taxonomicName id="9CCDB9EF243E1726FB73B61216A71949" ID-CoL="58LKV" authority="Ito, 1997" authorityName="Ito" authorityYear="1997" box="[1156,1436,874,895]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="doiinthanonensis">
|
||
<emphasis id="69B91E7E243E1726FB73B612167B1949" box="[1156,1344,874,895]" italics="true" pageId="50" pageNumber="351">T. doiinthanonensis</emphasis>
|
||
<bibRefCitation id="3F5CBF9D243E1726FAB1B61216A71949" author="Ito, N." box="[1350,1436,874,895]" pageId="50" pageNumber="351" pagination="45 - 49" refId="ref38089" refString="Ito, N. (1997) Two new species of the genus Trichotichnus from Mt. Doi Inthanon in North Thailand (Coleoptera, Carabidae, Harpalini). The Entomological Review of Japan, 52 (1), 45 - 49." type="journal article" year="1997">Ito, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B6FF16A7198E" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B6FF16A7198E" pageId="50" pageNumber="351">
|
||
- Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FE16B6FE116F19AA" box="[481,596,902,924]" captionStart="FIGURES 59–66" captionStartId="28.[151,264,1477,1501]" captionTargetBox="[155,1428,927,1446]" captionTargetId="figure-343@28.[151,1436,921,1454]" captionTargetPageId="28" captionText="FIGURES 59–66. Trichotichnus schmidti sp. n. (59–61, Yunnan, holotype; 62–66, Nepal, Manaslu). 59 & 62, terminal lamella of aedeagus; 60, 61, 63 & 64, median lobe of aedeagus; 65, gonocoxite; 66, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (59, 61, 62 & 64), lateral (60, 63 & 65) and ventral (66) views. Scale bars: A = 0.5 mm (59, 62, 65 & 66), B = 1.0 mm (60, 61, 63 & 64)." figureDoi="http://doi.org/10.5281/zenodo.6781735" httpUri="https://zenodo.org/record/6781735/files/figure.png" pageId="50" pageNumber="351">Figs. 59–64</figureCitation>
|
||
) more robust, with apex bent dorsally; internal sac with somewhat wide oval (in dorsal view) spiny patches apically. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FD77B6DB11CF198F" box="[640,756,931,953]" captionStart="FIGURES 56–58" captionStartId="27.[151,264,1121,1145]" captionTargetBox="[168,1428,182,1096]" captionTargetId="figure-19@27.[158,1430,181,1098]" captionTargetPageId="27" captionText="FIGURES 56–58. Trichotichnus schmidti sp. n. (Yunnan: 56 & 57, Nankang; 58, S Tengchong), habitus. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781731" httpUri="https://zenodo.org/record/6781731/files/figure.png" pageId="50" pageNumber="351">Figs. 56–58</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FC8DB6DB10D6198E" box="[890,1005,931,952]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="8.4" metricValueMin="7.3" pageId="50" pageNumber="351" unit="mm" value="7.85" valueMax="8.4" valueMin="7.3">7.3–8.4 mm</quantity>
|
||
.....................
|
||
<taxonomicName id="9CCDB9EF243E1726FB00B6DB165B198E" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[1271,1376,931,952]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="schmidti" status="sp. nov.">
|
||
<emphasis id="69B91E7E243E1726FB00B6DB165B198E" box="[1271,1376,931,952]" italics="true" pageId="50" pageNumber="351">T. schmidti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243E1726FA91B6DB16A7198E" bold="true" box="[1382,1436,931,952]" pageId="50" pageNumber="351">
|
||
<taxonomicNameLabel id="728AA305243E1726FA91B6DB16A7198E" box="[1382,1436,931,952]" pageId="50" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B6C716A71E71" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B6C716A71E38" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B6C716A71E38" pageId="50" pageNumber="351">
|
||
5 Preapical sinuation of elytra rather deep (
|
||
<figureCitation id="C3F6DEE9243E1726FD80B6C711FA19E2" box="[631,705,959,981]" captionStart="FIGURES 38–39" captionStartId="21.[151,264,1041,1065]" captionTargetBox="[151,1436,669,1016]" captionTargetId="figure-229@21.[150,1437,669,1018]" captionTargetPageId="21" captionText="FIGURES 38–39. Trichotichnus spp., elytral apex. 38, T. cyrtops (Tschitschérine, 1906); 39, T. schmidti sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781717" httpUri="https://zenodo.org/record/6781717/files/figure.png" pageId="50" pageNumber="351">Fig. 38</figureCitation>
|
||
). Legs brownish yellow, femora not darker than tibiae. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FEC6B6A4128F19C7" box="[305,436,988,1010]" captionStart="FIGURES 40–45" captionStartId="21.[151,264,1897,1921]" captionTargetBox="[158,1430,1129,1868]" captionTargetId="figure-253@21.[151,1436,1125,1874]" captionTargetPageId="21" captionText="FIGURES 40–45. Trichotichnus spp.: 40–43, T. cyrtops (Tschitschérine, 1906) (holotype and female from Sichuan); 44, 45, T. emarginatus Andrewes, 1930 (Nepal). 40, 41, 44 & 45, median lobe of aedeagus; 42, gonocoxite; 43, laterotergite, gonosubcox- ite, and gonocoxite. Lateral (40, 42 & 44), dorsal (41 & 45) and ventral (43) views. Scale bars: A = 0.5 mm (42 & 43), B = 1.0 mm (40, 41, 44 & 45)." figureDoi="http://doi.org/10.5281/zenodo.6781721" httpUri="https://zenodo.org/record/6781721/files/figure.png" pageId="50" pageNumber="351">Figs. 40 & 41</figureCitation>
|
||
) in lateral view almost straight in basal two-thirds, terminal lamella in dorsal view wide and long; female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FE9AB68012CF1E3B" box="[365,500,1016,1038]" captionStart="FIGURES 40–45" captionStartId="21.[151,264,1897,1921]" captionTargetBox="[158,1430,1129,1868]" captionTargetId="figure-253@21.[151,1436,1125,1874]" captionTargetPageId="21" captionText="FIGURES 40–45. Trichotichnus spp.: 40–43, T. cyrtops (Tschitschérine, 1906) (holotype and female from Sichuan); 44, 45, T. emarginatus Andrewes, 1930 (Nepal). 40, 41, 44 & 45, median lobe of aedeagus; 42, gonocoxite; 43, laterotergite, gonosubcox- ite, and gonocoxite. Lateral (40, 42 & 44), dorsal (41 & 45) and ventral (43) views. Scale bars: A = 0.5 mm (42 & 43), B = 1.0 mm (40, 41, 44 & 45)." figureDoi="http://doi.org/10.5281/zenodo.6781721" httpUri="https://zenodo.org/record/6781721/files/figure.png" pageId="50" pageNumber="351">Figs. 42 & 43</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FD76B68011F31E38" box="[641,712,1016,1038]" captionStart="FIGURES 35–37" captionStartId="20.[151,264,1335,1359]" captionTargetBox="[158,1428,536,1307]" captionTargetId="figure-178@20.[150,1437,525,1312]" captionTargetPageId="20" captionText="FIGURES 35–37. Trichotichnus spp., habitus. 35, T. cyrtops (Tschitschérine, 1906) (Yunnan); 36, T. doiinthanonensis Ito, 1997 (Dulongjiang); 37, T. fedorenkoi sp. n. (Tam Dao). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781715" httpUri="https://zenodo.org/record/6781715/files/figure.png" pageId="50" pageNumber="351">Fig. 35</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FCBAB68010FB1E38" box="[845,960,1016,1038]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" metricValueMax="9.2" metricValueMin="7.7" pageId="50" pageNumber="351" unit="mm" value="8.45" valueMax="9.2" valueMin="7.7">7.7–9.2 mm</quantity>
|
||
............
|
||
<taxonomicName id="9CCDB9EF243E1726FB95B68116A71E38" ID-CoL="58LKM" authority="(Tschitscherine, 1906)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1906" box="[1122,1436,1016,1038]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="cyrtops">
|
||
<emphasis id="69B91E7E243E1726FB95B68117FB1E38" box="[1122,1216,1017,1038]" italics="true" pageId="50" pageNumber="351">T. cyrtops</emphasis>
|
||
(
|
||
<bibRefCitation id="3F5CBF9D243E1726FB3BB68016AE1E38" author="Tschitscherine, T." box="[1228,1429,1016,1038]" pageId="50" pageNumber="351" pagination="247 - 292" refId="ref39061" refString="Tschitscherine, T. (1906) Notes detaches sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37, 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B16D16A71E71" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
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<keyLead id="E03C6159243E1726FF60B16D16A71E71" pageId="50" pageNumber="351">- Preapical sinuation of elytra shallow. Legs unicoloured pale or with infuscate femora. Male and female genitalia not as above ................................................................................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B13616A71EE3" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B13616A71EB6" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B13616A71EB6" pageId="50" pageNumber="351">
|
||
6 Protibia in both sexes with three stout, similar sized preapical spines on outer margin (as in
|
||
<figureCitation id="C3F6DEE9243E1726FBBAB136178C1E55" box="[1101,1207,1102,1124]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 18, 19</figureCitation>
|
||
). Legs brownish yellow or brown, at most femora slightly infuscate. Body larger: length
|
||
<quantity id="9C356F89243E1726FCBBB11310F11EB6" box="[844,970,1131,1152]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.14" metricValueMin="0.9199999999999999" pageId="50" pageNumber="351" unit="mm" value="10.3" valueMax="11.4" valueMin="9.2">9.2–11.4 mm</quantity>
|
||
..................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B1FF16A71EE3" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B1FF16A71EE3" pageId="50" pageNumber="351">
|
||
- Protibia on outer margin preapically with one stouter and two very thin spines in male (as in
|
||
<figureCitation id="C3F6DEE9243E1726FBA8B1FF179E1EAA" box="[1119,1189,1159,1181]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 20</figureCitation>
|
||
) and with three comparatively stout spines in female. Legs brownish yellow or reddish brown, with femora distinctly infuscate. Body smaller: length
|
||
<quantity id="9C356F89243E1726FF15B1B8126E1EE0" box="[226,341,1216,1238]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.8" metricValueMin="6.7" pageId="50" pageNumber="351" unit="mm" value="7.75" valueMax="8.8" valueMin="6.7">6.7–8.8 mm</quantity>
|
||
(up to
|
||
<quantity id="9C356F89243E1726FE6DB1B812D41EE0" box="[410,495,1216,1238]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="50" pageNumber="351" unit="mm" value="10.4">10.4 mm</quantity>
|
||
in
|
||
<taxonomicName id="9CCDB9EF243E1726FDF9B1B9114E1EE3" ID-CoL="58LKD" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" box="[526,629,1216,1238]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="bouvieri">
|
||
<emphasis id="69B91E7E243E1726FDF9B1B9114E1EE3" box="[526,629,1216,1238]" italics="true" pageId="50" pageNumber="351">T. bouvieri</emphasis>
|
||
</taxonomicName>
|
||
)................................................................. 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B1A516A71FB6" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B1A516A71F1D" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B1A516A71F1D" pageId="50" pageNumber="351">
|
||
7 Metepisternum (
|
||
<figureCitation id="C3F6DEE9243E1726FE88B1A512FF1EC4" box="[383,452,1245,1267]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 22</figureCitation>
|
||
) shorter, only slightly longer than wide. Body (
|
||
<figureCitation id="C3F6DEE9243E1726FC72B1A510F11EC4" box="[901,970,1245,1267]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Fig. 10</figureCitation>
|
||
) more robust, with appendages shorter; antennae not reaching pronotal basal edge; metatarsus slightly shorter than HWmin. Median lobe of aedeagus with a small transverse apical capitulum (
|
||
<figureCitation id="C3F6DEE9243E1726FE79B06E113F1F1D" box="[398,516,1302,1324]" captionStart="FIGURES 29–33" captionStartId="16.[151,264,1947,1971]" captionTargetBox="[157,1427,1097,1915]" captionTargetId="figure-415@16.[151,1436,1089,1923]" captionTargetPageId="16" captionText="FIGURES 29–33. Trichotichnus subrobustus sp. n. 29, terminal lamella of aedeagus; 30 & 31, median lobe of aedeagus (holo- type); 32, gonocoxite; 33, laterotergite, gonosubcoxite, and gonocoxite (N Dizhengdang). Dorsal (29 & 31), lateral (30 & 32) and ventral (33) views. Scale bars: A = 0.5 mm (29, 32 & 33), B = 1.0 mm (30 & 31)." figureDoi="http://doi.org/10.5281/zenodo.6781711" httpUri="https://zenodo.org/record/6781711/files/figure.png" pageId="50" pageNumber="351">Figs. 29–31</figureCitation>
|
||
), female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FD10B06E106E1F1D" box="[743,853,1302,1324]" captionStart="FIGURES 29–33" captionStartId="16.[151,264,1947,1971]" captionTargetBox="[157,1427,1097,1915]" captionTargetId="figure-415@16.[151,1436,1089,1923]" captionTargetPageId="16" captionText="FIGURES 29–33. Trichotichnus subrobustus sp. n. 29, terminal lamella of aedeagus; 30 & 31, median lobe of aedeagus (holo- type); 32, gonocoxite; 33, laterotergite, gonosubcoxite, and gonocoxite (N Dizhengdang). Dorsal (29 & 31), lateral (30 & 32) and ventral (33) views. Scale bars: A = 0.5 mm (29, 32 & 33), B = 1.0 mm (30 & 31)." figureDoi="http://doi.org/10.5281/zenodo.6781711" httpUri="https://zenodo.org/record/6781711/files/figure.png" pageId="50" pageNumber="351">Figs 32, 33</figureCitation>
|
||
. Body length 9.2–10.0 mm.........
|
||
<taxonomicName id="9CCDB9EF243E1726FB22B06E165B1F1D" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[1237,1376,1302,1323]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="subrobustus" status="sp. nov.">
|
||
<emphasis id="69B91E7E243E1726FB22B06E165B1F1D" box="[1237,1376,1302,1323]" italics="true" pageId="50" pageNumber="351">T. subrobustus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243E1726FA91B06E16A71F1D" bold="true" box="[1382,1436,1302,1323]" pageId="50" pageNumber="351">
|
||
<taxonomicNameLabel id="728AA305243E1726FA91B06E16A71F1D" box="[1382,1436,1302,1323]" pageId="50" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B04B16A71FB6" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B04B16A71FB6" pageId="50" pageNumber="351">
|
||
- Metepisternum markedly longer than wide (as in
|
||
<figureCitation id="C3F6DEE9243E1726FD4EB04A103B1F71" box="[697,768,1330,1352]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 21</figureCitation>
|
||
). Body (
|
||
<figureCitation id="C3F6DEE9243E1726FCA6B04A109E1F71" box="[849,933,1330,1352]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Figs 8, 9</figureCitation>
|
||
) less robust, with appendages longer; antennae slender, surpassing pronotal basal edge; metatarsus almost as long as or slightly shorter than HWmax. Male and female genitalia not as above......................................................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B0F016A71C53" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B0F016A71FC5" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B0F016A71FC5" pageId="50" pageNumber="351">
|
||
8 Pronotum (
|
||
<figureCitation id="C3F6DEE9243E1726FEA7B0F012B01FAB" box="[336,395,1416,1438]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Fig. 8</figureCitation>
|
||
) relatively larger, with shallower laterobasal depressions; microsculpture on disc strongly obliterated, not forming distinct meshes. In male, pro- and mesotarsomeres wider (as in
|
||
<figureCitation id="C3F6DEE9243E1726FC61B0DC10E71F8F" box="[918,988,1444,1466]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 18</figureCitation>
|
||
), last visible abdominal sternite with two pairs of marginal setae. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FD6DB0B910361FE0" box="[666,781,1473,1495]" captionStart="FIGURES 11–16" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[194,1393,1153,1911]" captionTargetId="figure-51@9.[187,1401,1144,1919]" captionTargetPageId="9" captionText="FIGURES 11–16. Trichotichnus fukuharai Habu, 1957 (11, 15 & 16, Yunnan; 12–14, Sichuan). 11 & 12, terminal lamella of aedeagus; 13, 14, median lobe of aedeagus; 15, gonocoxite; 16, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (11, 12 & 14), lateral (13 & 15) and ventral (16) views. Scale bars: A = 0.5 mm (11, 12, 15 & 16), B = 1.0 mm (13 & 14)." figureDoi="http://doi.org/10.5281/zenodo.6781691" httpUri="https://zenodo.org/record/6781691/files/figure.png" pageId="50" pageNumber="351">Figs. 11–14</figureCitation>
|
||
) with apex not bent dorsally and with a large spine in internal sac; female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FE43B0A511121FC5" box="[436,553,1501,1523]" captionStart="FIGURES 11–16" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[194,1393,1153,1911]" captionTargetId="figure-51@9.[187,1401,1144,1919]" captionTargetPageId="9" captionText="FIGURES 11–16. Trichotichnus fukuharai Habu, 1957 (11, 15 & 16, Yunnan; 12–14, Sichuan). 11 & 12, terminal lamella of aedeagus; 13, 14, median lobe of aedeagus; 15, gonocoxite; 16, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (11, 12 & 14), lateral (13 & 15) and ventral (16) views. Scale bars: A = 0.5 mm (11, 12, 15 & 16), B = 1.0 mm (13 & 14)." figureDoi="http://doi.org/10.5281/zenodo.6781691" httpUri="https://zenodo.org/record/6781691/files/figure.png" pageId="50" pageNumber="351">Figs. 15, 16</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FD42B0A511D41FC5" box="[693,751,1501,1523]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Fig. 8</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FC82B0A610C81FC5" box="[885,1011,1502,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.13" metricValueMin="0.97" pageId="50" pageNumber="351" unit="mm" value="10.5" valueMax="11.3" valueMin="9.7">9.7–11.3 mm</quantity>
|
||
...............
|
||
<taxonomicName id="9CCDB9EF243E1726FB46B0A616A71FC5" ID-CoL="58LL4" authority="Habu, 1957" authorityName="Habu" authorityYear="1957" box="[1201,1436,1501,1523]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="fukuharai">
|
||
<emphasis id="69B91E7E243E1726FB46B0A6161D1FC4" box="[1201,1318,1501,1523]" italics="true" pageId="50" pageNumber="351">T. fukuharai</emphasis>
|
||
<bibRefCitation id="3F5CBF9D243E1726FADBB0A516A71FC5" author="Habu, A." box="[1324,1436,1501,1523]" pageId="50" pageNumber="351" pagination="76 - 78" refId="ref37900" refString="Habu, A. (1957) Description of a new Trichotichnus - species. Akitu, 6, 76 - 78." type="journal article" year="1957">Habu, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B08216A71C53" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B08216A71C53" pageId="50" pageNumber="351">
|
||
- Pronotum (
|
||
<figureCitation id="C3F6DEE9243E1726FEB8B08212B21C39" box="[335,393,1530,1552]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Fig. 9</figureCitation>
|
||
) relatively smaller, with deeper laterobasal depressions; microsculpture on disc more distinct, consisting of very fine, more or less isodiametric meshes. In male, pro- and mesotarsomeres narrower (as in
|
||
<figureCitation id="C3F6DEE9243E1726FB85B36E17831C1A" box="[1138,1208,1558,1580]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="50" pageNumber="351">Fig. 19</figureCitation>
|
||
), last visible abdominal sternite with one pair of marginal setae. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FC9CB34B10E41C7F" box="[875,991,1587,1609]" captionStart="FIGURES 23–27" captionStartId="14.[151,264,922,946]" captionTargetBox="[160,1428,186,890]" captionTargetId="figure-32@14.[151,1436,181,899]" captionTargetPageId="14" captionText="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 & 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 & 25), lateral (24 & 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 & 27), B = 1.0 mm (24 & 25)." figureDoi="http://doi.org/10.5281/zenodo.6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="50" pageNumber="351">Figs. 23–25</figureCitation>
|
||
) with apex bent dorsally and without spine in internal sac; female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FDD8B337118D1C53" box="[559,694,1615,1637]" captionStart="FIGURES 23–27" captionStartId="14.[151,264,922,946]" captionTargetBox="[160,1428,186,890]" captionTargetId="figure-32@14.[151,1436,181,899]" captionTargetPageId="14" captionText="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 & 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 & 25), lateral (24 & 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 & 27), B = 1.0 mm (24 & 25)." figureDoi="http://doi.org/10.5281/zenodo.6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="50" pageNumber="351">Figs. 26 & 27</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="C3F6DEE9243E1726FCB5B33710471C53" box="[834,892,1615,1637]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="50" pageNumber="351">Fig. 9</figureCitation>
|
||
. Body length 10.0–
|
||
<quantity id="9C356F89243E1726FBCFB32817B61C53" box="[1080,1165,1616,1637]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="50" pageNumber="351" unit="mm" value="11.4">11.4 mm</quantity>
|
||
.....
|
||
<taxonomicName id="9CCDB9EF243E1726FB3AB328165B1C53" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[1229,1376,1616,1637]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
|
||
<emphasis id="69B91E7E243E1726FB3AB328165B1C53" box="[1229,1376,1616,1637]" italics="true" pageId="50" pageNumber="351">T. angustitarsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243E1726FA91B32816A71C53" bold="true" box="[1382,1436,1616,1637]" pageId="50" pageNumber="351">
|
||
<taxonomicNameLabel id="728AA305243E1726FA91B32816A71C53" box="[1382,1436,1616,1637]" pageId="50" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B31416A71D1A" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B31416A71CE1" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B31416A71CE1" pageId="50" pageNumber="351">
|
||
9 Sides of pronotum (
|
||
<figureCitation id="C3F6DEE9243E1726FE51B31411221CB7" box="[422,537,1644,1666]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1674,1698]" captionTargetBox="[167,1435,450,1643]" captionTargetId="figure-134@5.[151,1436,450,1648]" captionTargetPageId="5" captionText="FIGURES 1–2. Trichotichnus anthracinus Landin, 1955, habitus (1, Yunnan; 2, Myanmar, holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781671" httpUri="https://zenodo.org/record/6781671/files/figure.png" pageId="50" pageNumber="351">Figs. 1 & 2</figureCitation>
|
||
) almost straightly converging in basal half; apical margin very shallowly emarginate; basal angles obtusangular, sharp at apex, generally with a very small obtuse denticle. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243E1726FB06B3F116581CA8" box="[1265,1379,1673,1695]" captionStart="FIGURES 3–6" captionStartId="6.[151,264,1805,1829]" captionTargetBox="[160,1427,1001,1771]" captionTargetId="figure-360@6.[151,1436,993,1782]" captionTargetPageId="6" captionText="FIGURES 3–6. Trichotichnus anthracinus Landin, 1955 (Yunnan). 3, 4, median lobe of aedeagus; 5, gonocoxite; 6, lateroter- gite, gonosubcoxite, and gonocoxite. Lateral (3 & 5), dorsal (4) and ventral (6) views. Scale bars: A = 0.5 mm (5 & 6), B = 1.0 mm (3 & 4)." figureDoi="http://doi.org/10.5281/zenodo.6781673" httpUri="https://zenodo.org/record/6781673/files/figure.png" pageId="50" pageNumber="351">Figs. 3 & 4</figureCitation>
|
||
) with apical portion angularly bent dorsally; female genitalia as in
|
||
<figureCitation id="C3F6DEE9243E1726FCDEB3DD10AC1C8D" box="[809,919,1701,1723]" captionStart="FIGURES 3–6" captionStartId="6.[151,264,1805,1829]" captionTargetBox="[160,1427,1001,1771]" captionTargetId="figure-360@6.[151,1436,993,1782]" captionTargetPageId="6" captionText="FIGURES 3–6. Trichotichnus anthracinus Landin, 1955 (Yunnan). 3, 4, median lobe of aedeagus; 5, gonocoxite; 6, lateroter- gite, gonosubcoxite, and gonocoxite. Lateral (3 & 5), dorsal (4) and ventral (6) views. Scale bars: A = 0.5 mm (5 & 6), B = 1.0 mm (3 & 4)." figureDoi="http://doi.org/10.5281/zenodo.6781673" httpUri="https://zenodo.org/record/6781673/files/figure.png" pageId="50" pageNumber="351">Figs. 5 & 6</figureCitation>
|
||
. Body length
|
||
<quantity id="9C356F89243E1726FBEAB3DD17AB1C8D" box="[1053,1168,1701,1723]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="7.7" metricValueMin="7.2" pageId="50" pageNumber="351" unit="mm" value="7.45" valueMax="7.7" valueMin="7.2">7.2–7.7 mm</quantity>
|
||
...................................................................................................
|
||
<taxonomicName id="9CCDB9EF243E1726FB7BB3BA16A71CE1" ID-CoL="58LK6" authority="Landin, 1955" authorityName="Ldn., B. O. Landin" authorityYear="1955" box="[1164,1436,1729,1752]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="anthracinus">
|
||
<emphasis id="69B91E7E243E1726FB7BB3BA162C1CE1" box="[1164,1303,1730,1751]" italics="true" pageId="50" pageNumber="351">T. anthracinus</emphasis>
|
||
<bibRefCitation id="3F5CBF9D243E1726FAEBB3BA16A71CE1" author="Landin, B. - O." box="[1308,1436,1729,1752]" pageId="50" pageNumber="351" pagination="399 - 472" refId="ref38830" refString="Landin, B. - O. (1955) Entomological results from the Swedish expedition 1934 to Burma and British India. Coleoptera: Carabidae. Collected by Rene Malaise. Arkiv for Zoologi, Series 2, 8, 399 - 472. [1955 - 1956]" type="journal article" year="1955">Landin, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B3A716A71D1A" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B3A716A71D1A" pageId="50" pageNumber="351">
|
||
- Sides of pronotum generally rounded along entire length; apical margin shallowly or moderately deeply emarginate; basal angles of pronotum more obtuse, sharp or blunted at apex, generally without denticle (if sides of pronotum almost straightly converging basally, body length more than
|
||
<quantity id="9C356F89243E1726FD77B26011F61D1A" box="[640,717,1815,1837]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="50" pageNumber="351" unit="mm" value="7.7">7.7 mm</quantity>
|
||
). Male and female genitalia not as above........................ 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243E1726FF60B24C16A71DEF" pageId="50" pageNumber="351">
|
||
<paragraph id="5B72C26C243E1726FF60B24C16A71DA9" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B24C16A71DA9" pageId="50" pageNumber="351">
|
||
10 Body robust, with wide and short elytra (EL/EW about 1.3). Last visible abdominal sternite of male with one pair of marginal setae. Median lobe of aedeagus arcuate in lateral view, with straight or slightly convex median portion of ventral margin; terminal lamella in dorsal view about as long as wide, with very small oblique apical capitulum. Body length
|
||
<quantity id="9C356F89243E1726FB13B215166C1DB4" box="[1252,1367,1901,1923]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="8.4" metricValueMin="6.7" pageId="50" pageNumber="351" unit="mm" value="7.55" valueMax="8.4" valueMin="6.7">6.7–8.4 mm</quantity>
|
||
................................................................................
|
||
<taxonomicName id="9CCDB9EF243E1726FB85B2F216A71DA9" ID-CoL="58LK7" authority="Ito, 2000" authorityName="Ito" authorityYear="2000" box="[1138,1436,1930,1951]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="arcuatomarginatus">
|
||
<emphasis id="69B91E7E243E1726FB85B2F216041DA9" box="[1138,1343,1930,1951]" italics="true" pageId="50" pageNumber="351">T. arcuatomarginatus</emphasis>
|
||
<bibRefCitation id="3F5CBF9D243E1726FAB1B2F216A71DA9" author="Ito, N." box="[1350,1436,1930,1951]" pageId="50" pageNumber="351" pagination="39 - 50" refId="ref38168" refString="Ito, N. (2000) Four new species of the genus Trichotichnus from West and South China (Coleoptera: Carabidae: Harpalini). The Entomological Review of Japan, 55 (1), 39 - 50." type="journal article" year="2000">Ito, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243E1726FF60B2DF16A71DEF" blockId="50.[151,1437,360,2010]" pageId="50" pageNumber="351">
|
||
<keyLead id="E03C6159243E1726FF60B2DF16A71DEF" pageId="50" pageNumber="351">- Body more elongate, with relatively longer elytra (EL/EW about 1.4–1.5). Last visible abdominal sternite of male with one or two pairs of marginal setae............................................................................ 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243F1727FF60B5E016A71B4C" pageId="51" pageNumber="352">
|
||
<paragraph id="5B72C26C243F1727FF60B5E016A71B33" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B5E016A71B33" pageId="51" pageNumber="352">
|
||
11 Pronotum widest at middle; basal fovea separated from lateral depression by convexity. Apex of left mandible in frontal view more or less truncate. Last visible abdominal sternite of male with two pairs of marginal setae. Median lobe of aedeagus arcuate in lateral view, with almost straight median portion of ventral margin; terminal lamella without apical capitulum, in dorsal view longer than wide. Body length
|
||
<quantity id="9C356F89243F1727FDFCB58811B11B33" box="[523,650,240,261]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.4" metricValueMin="8.1" pageId="51" pageNumber="352" unit="mm" value="9.25" valueMax="10.4" valueMin="8.1">8.1–10.4 mm</quantity>
|
||
.....................................
|
||
<taxonomicName id="9CCDB9EF243F1727FBAFB58816A71B33" ID-CoL="58LKD" authority="(Tschitscherine, 1897)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" box="[1112,1436,239,261]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="352" phylum="Arthropoda" rank="species" species="bouvieri">
|
||
<emphasis id="69B91E7E243F1727FBAFB58817FB1B32" box="[1112,1216,239,261]" italics="true" pageId="51" pageNumber="352">T. bouvieri</emphasis>
|
||
(Tschitschérine, 1897)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243F1727FF60B47516A71B4C" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B47516A71B4C" pageId="51" pageNumber="352">
|
||
- Pronotum widest before middle; area between basal fovea and lateral margin flattened. Apex of left mandible in frontal view blunted. Last visible abdominal sternite of male with one pair of marginal setae. Median lobe of aedeagus in lateral view evenly arcuate along dorsal and ventral margins; terminal lamella in dorsal view slightly wider than long, with very small oblique apical capitulum (recognizable in lateral view). Body length
|
||
<quantity id="9C356F89243F1727FCFBB41D10441B4C" box="[780,895,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.8" metricValueMin="7.8" pageId="51" pageNumber="352" unit="mm" value="8.3" valueMax="8.8" valueMin="7.8">7.8–8.8 mm</quantity>
|
||
.......................
|
||
<taxonomicName id="9CCDB9EF243F1727FB52B41D16A71B4C" ID-CoL="8MJFN" authority="Andrewes, 1930" authorityName="Andrewes" authorityYear="1930" box="[1189,1436,357,379]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="352" phylum="Arthropoda" rank="species" species="aquilo">
|
||
<emphasis id="69B91E7E243F1727FB52B41D17C11B4C" box="[1189,1274,357,378]" italics="true" pageId="51" pageNumber="352">T. aquilo</emphasis>
|
||
<bibRefCitation id="3F5CBF9D243F1727FB08B41D16A71B4C" author="Andrewes, H. E." box="[1279,1436,357,379]" pageId="51" pageNumber="352" pagination="1 - 44" refId="ref37802" refString="Andrewes, H. E. (1930) The Carabidae of the third Mount Everest Expedition, 1924. Transactions of the Entomological Society of London, 78, 1 - 44, 1 map. https: // doi. org / 10.1111 / j. 1365 - 2311.1930. tb 01198. x" type="book chapter" year="1930">Andrewes, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243F1727FF60B4FA16A7183B" pageId="51" pageNumber="352">
|
||
<paragraph id="5B72C26C243F1727FF60B4FA16A71BE5" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B4FA16A71BE5" pageId="51" pageNumber="352">
|
||
12 Metepisternum shorter, 1.18–1.25 times as long as wide. Wings wholly or partly reduced. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243F1727FA9BB4FA12101B83" captionStart="FIGURES 98–105" captionStartId="40.[151,264,1637,1661]" captionTargetBox="[158,1430,964,1605]" captionTargetId="figure-323@40.[151,1436,957,1614]" captionTargetPageId="40" captionText="FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105)." figureDoi="http://doi.org/10.5281/zenodo.6781759" httpUri="https://zenodo.org/record/6781759/files/figure.png" pageId="51" pageNumber="352">Figs. 98–100</figureCitation>
|
||
) stout, sharply hooked ventrally at apex, female genitalia as in
|
||
<figureCitation id="C3F6DEE9243F1727FC6CB4E717041B83" box="[923,1087,415,437]" captionStart="FIGURES 98–105" captionStartId="40.[151,264,1637,1661]" captionTargetBox="[158,1430,964,1605]" captionTargetId="figure-323@40.[151,1436,957,1614]" captionTargetPageId="40" captionText="FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105)." figureDoi="http://doi.org/10.5281/zenodo.6781759" httpUri="https://zenodo.org/record/6781759/files/figure.png" pageId="51" pageNumber="352">Figs. 101 & 102</figureCitation>
|
||
. Habitus as in
|
||
<figureCitation id="C3F6DEE9243F1727FB25B4E716211B83" box="[1234,1306,415,437]" captionStart="FIGURES 94–95" captionStartId="38.[151,264,1747,1771]" captionTargetBox="[174,1411,469,1715]" captionTargetId="figure-151@38.[158,1429,453,1724]" captionTargetPageId="38" captionText="FIGURES 94–95. Trichotichnus hamulipenis sp. n., habitus. 94, nominotypical subspecies (above Bapo); 95, T. hamulipenis nepalensis ssp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781755" httpUri="https://zenodo.org/record/6781755/files/figure.png" pageId="51" pageNumber="352">Fig. 94</figureCitation>
|
||
. Body length 9.0–
|
||
<quantity id="9C356F89243F1727FEFAB4C5125A1BE4" box="[269,353,445,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="51" pageNumber="352" unit="mm" value="10.5">10.5 mm</quantity>
|
||
.......................................
|
||
<taxonomicName id="9CCDB9EF243F1727FCC1B4C510F91BE4" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[822,962,445,466]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="352" phylum="Arthropoda" rank="species" species="hamulipenis" status="sp. nov.">
|
||
<emphasis id="69B91E7E243F1727FCC1B4C510F91BE4" box="[822,962,445,466]" italics="true" pageId="51" pageNumber="352">T. hamulipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243F1727FC3FB4C510C41BE4" bold="true" box="[968,1023,445,466]" pageId="51" pageNumber="352">
|
||
<taxonomicNameLabel id="728AA305243F1727FC3FB4C510C41BE4" box="[968,1023,445,466]" pageId="51" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(in
|
||
<collectingRegion id="99090C8E243F1727FBD3B4C5174B1BE4" box="[1060,1136,445,466]" country="China" name="Yunnan" pageId="51" pageNumber="352">Yunnan</collectingRegion>
|
||
, the nominotypical subspecies)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243F1727FF60B4A316A7183B" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B4A316A7183B" pageId="51" pageNumber="352">- Metepisternum longer, 1.40–1.60 times as long as wide. Wings well developed. Median lobe of aedeagus slenderer, not hooked at apex ventrally.................................................................................... 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="E039DAC9243F1727FF60B76D16A718C1" pageId="51" pageNumber="352">
|
||
<paragraph id="5B72C26C243F1727FF60B76D16A718B4" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B76D16A718B4" pageId="51" pageNumber="352">
|
||
13 Body (
|
||
<figureCitation id="C3F6DEE9243F1727FED3B76D1243181C" box="[292,376,533,555]" captionStart="FIGURES 107–108" captionStartId="43.[151,264,1956,1980]" captionTargetBox="[157,1420,650,1924]" captionTargetId="figure-222@43.[153,1433,633,1933]" captionTargetPageId="43" captionText="FIGURES 107–108. Trichotichnus ssp., habitus. 107, T. miwai continentalis ssp. n. (Wuliangshan); 108, T. kavanaughi sp. n. (Kongdang). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="51" pageNumber="352">Fig. 107</figureCitation>
|
||
) with head and pronotum relatively wider, pronotal basal angles in most specimens less protruding laterally. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243F1727FE29B74A115C1871" box="[478,615,562,584]" captionStart="FIGURES 109–120" captionStartId="44.[151,264,1763,1787]" captionTargetBox="[165,1420,194,1727]" captionTargetId="figure-32@44.[151,1436,181,1739]" captionTargetPageId="44" captionText="FIGURES 109–120. Trichotichnus miwai Jedlička, 1949: 109–113, nominotypical subspecies (109 & 110, Nantou; 111 & 113, Hualien); 114–120, T. miwai continentalis ssp. n. (114–116, holotype; 117 & 118, Phia Oac, Vietnam; 119 & 120, Kunming). 109, 111, 114 & 117, terminal lamella of aedeagus; 110, 112, 113, 115, 116 & 118, median lobe of aedeagus; 119, gonocoxite; 120, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (109, 111, 113, 114, 116 & 117), lateral (110, 112, 115, 118 & 119) and ventral (120) views. Scale bars: A = 0.5 mm (109, 111, 114, 117, 119 & 120), B = 1.0 mm (110, 112, 113, 115, 116 & 118)." figureDoi="http://doi.org/10.5281/zenodo.6781767" httpUri="https://zenodo.org/record/6781767/files/figure.png" pageId="51" pageNumber="352">Figs. 114–118</figureCitation>
|
||
) evenly arcuate, with shorter terminal lamella (114, 117); internal sac with a mediumsized elongate spine on the left side behind the middle. Gonocoxite (119, 120) narrower and with wider base. Body length 8.0–
|
||
<quantity id="9C356F89243F1727FEFBB715126E18B4" box="[268,341,621,642]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="51" pageNumber="352" unit="mm" value="9.4">9.4 mm</quantity>
|
||
..................................................................
|
||
<taxonomicName id="9CCDB9EF243F1727FB74B715166C18B4" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[1155,1367,621,642]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="352" phylum="Arthropoda" rank="subSpecies" species="miwai" subSpecies="continentalis">
|
||
<emphasis id="69B91E7E243F1727FB74B715166C18B4" box="[1155,1367,621,642]" italics="true" pageId="51" pageNumber="352">T. miwai continentalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243F1727FAAAB71516A718B4" bold="true" box="[1373,1436,621,642]" pageId="51" pageNumber="352">ssp. n.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="5B72C26C243F1727FF60B7F316A718C1" blockId="51.[151,1437,151,760]" pageId="51" pageNumber="352">
|
||
<keyLead id="E03C6159243F1727FF60B7F316A718C1" pageId="51" pageNumber="352">
|
||
- Body (
|
||
<figureCitation id="C3F6DEE9243F1727FED3B7F2124D18A9" box="[292,374,650,672]" captionStart="FIGURES 107–108" captionStartId="43.[151,264,1956,1980]" captionTargetBox="[157,1420,650,1924]" captionTargetId="figure-222@43.[153,1433,633,1933]" captionTargetPageId="43" captionText="FIGURES 107–108. Trichotichnus ssp., habitus. 107, T. miwai continentalis ssp. n. (Wuliangshan); 108, T. kavanaughi sp. n. (Kongdang). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781763" httpUri="https://zenodo.org/record/6781763/files/figure.png" pageId="51" pageNumber="352">Fig. 108</figureCitation>
|
||
) with head and pronotum relatively narrower, pronotal basal angles in most specimens more protruding laterally. Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9243F1727FDE6B7DF11A5188B" box="[529,670,679,701]" captionStart="FIGURES 122–126" captionStartId="48.[151,264,1717,1741]" captionTargetBox="[159,1429,927,1683]" captionTargetId="figure-339@48.[151,1436,921,1693]" captionTargetPageId="48" captionText="FIGURES 122–126. Trichotichnus kavanaughi sp. n. 122, terminal lamella of aedeagus; 123, 124, median lobe of aedeagus (holotype); 125, gonocoxite; 126, laterotergite, gonosubcoxite, and gonocoxite (Maku). Dorsal (122 & 124), lateral (123 & 125) and ventral (126) views. Scale bars: A = 0.5 mm (122, 125 & 126), B = 1.0 mm (123 & 124)." figureDoi="http://doi.org/10.5281/zenodo.6781771" httpUri="https://zenodo.org/record/6781771/files/figure.png" pageId="51" pageNumber="352">Figs. 122–124</figureCitation>
|
||
) in lateral view more arcuate and slenderer, with much longer terminal lamella (
|
||
<figureCitation id="C3F6DEE9243F1727FF1EB7BD120518EC" box="[233,318,709,731]" captionStart="FIGURES 122–126" captionStartId="48.[151,264,1717,1741]" captionTargetBox="[159,1429,927,1683]" captionTargetId="figure-339@48.[151,1436,921,1693]" captionTargetPageId="48" captionText="FIGURES 122–126. Trichotichnus kavanaughi sp. n. 122, terminal lamella of aedeagus; 123, 124, median lobe of aedeagus (holotype); 125, gonocoxite; 126, laterotergite, gonosubcoxite, and gonocoxite (Maku). Dorsal (122 & 124), lateral (123 & 125) and ventral (126) views. Scale bars: A = 0.5 mm (122, 125 & 126), B = 1.0 mm (123 & 124)." figureDoi="http://doi.org/10.5281/zenodo.6781771" httpUri="https://zenodo.org/record/6781771/files/figure.png" pageId="51" pageNumber="352">Fig. 122</figureCitation>
|
||
); internal sac with a group of small spines instead of a medium-sized spine. Gonocoxite (125, 126) wider and with narrower base. Body length
|
||
<quantity id="9C356F89243F1727FE06B79A115F18CE" box="[497,612,738,760]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.6" metricValueMin="8.5" pageId="51" pageNumber="352" unit="mm" value="9.05" valueMax="9.6" valueMin="8.5">8.5–9.6 mm</quantity>
|
||
...................................................
|
||
<taxonomicName id="9CCDB9EF243F1727FB2EB79B165B18C1" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[1241,1376,738,760]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="352" phylum="Arthropoda" rank="species" species="kavanaughi" status="sp. nov.">
|
||
<emphasis id="69B91E7E243F1727FB2EB79B165B18C1" box="[1241,1376,738,760]" italics="true" pageId="51" pageNumber="352">T. kavanaughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E243F1727FA91B79A16A718C1" bold="true" box="[1382,1436,738,759]" pageId="51" pageNumber="352">
|
||
<taxonomicNameLabel id="728AA305243F1727FA91B79A16A718C1" box="[1382,1436,738,759]" pageId="51" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |