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<mods:title id="7EB36F9D308AB8AA5801B9FECC7AE604">Redescriptions of Indian species of Nigrobaetis Kazlauskas (in Novikova &amp; Kluge) 1987 (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="2F0AA6DA383E663D79F780D624EF713E">Kluge, Nikita</mods:namePart>
<mods:affiliation id="748583BA1D8840252F1E7DB69E490142">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia.</mods:affiliation>
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<mods:namePart id="BA3C4B38BDE013B9AB458FCAC8B17FBA">Sivaruban, T.</mods:namePart>
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<mods:namePart id="82F6412A80064C3681A32DC77D290E34">Isack, Rajasekaran</mods:namePart>
<mods:affiliation id="4834E85B314DB74300E721A5343CF33A">PG and Research department of Zoology, The American College, Madurai- 625002, India.</mods:affiliation>
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<taxonomicName id="87BC4D35FFE7043059C1FC83FC8CFF43" ID-CoL="749XZ" authority="(Muller-Liebenau 1984)" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[151,882,151,178]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="534" phylum="Arthropoda" rank="species" species="minutus" subGenus="Margobaetis">
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<emphasis id="72C8EAA4FFE7043059C1FC83FEDDFF40" bold="true" box="[151,291,151,178]" italics="true" pageId="6" pageNumber="534">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFE704305862FC83FE32FF40" bold="true" box="[308,460,151,178]" italics="true" pageId="6" pageNumber="534">Margobaetis</emphasis>
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<emphasis id="72C8EAA4FFE70430588BFC83FDBEFF43" bold="true" box="[477,576,151,177]" italics="true" pageId="6" pageNumber="534">minutus</emphasis>
(
<bibRefCitation id="242D4B47FFE704305B06FC83FC97FF43" author="Muller-Liebenau, I." box="[592,873,151,177]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
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(
<figureCitation id="D8872A33FFE7043059C9FCAFFEEFFF27" box="[159,273,187,213]" captionStart-0="FIGURES 1–12" captionStart-1="FIGURES 13–18" captionStart-2="FIGURES 19–22" captionStart-3="FIGURES 23–25" captionStart-4="FIGURES 26–32" captionStart-5="FIGURES 33–34" captionStart-6="FIGURES 35–38" captionStart-7="FIGURES 39–43" captionStartId-0="2.[151,264,1942,1966]" captionStartId-1="3.[151,264,1827,1851]" captionStartId-2="4.[151,264,1837,1861]" captionStartId-3="5.[151,264,1831,1855]" captionStartId-4="7.[151,264,1794,1818]" captionStartId-5="8.[151,264,1976,2000]" captionStartId-6="9.[151,264,1914,1938]" captionStartId-7="10.[151,264,1802,1826]" captionTargetBox-0="[151,1436,346,1919]" captionTargetBox-1="[151,1436,181,1804]" captionTargetBox-2="[151,1436,181,1813]" captionTargetBox-3="[151,1436,181,1807]" captionTargetBox-4="[151,1416,181,1770]" captionTargetBox-5="[151,1435,237,1952]" captionTargetBox-6="[166,1436,309,1891]" captionTargetBox-7="[151,1436,309,1779]" captionTargetId-0="figure-128@2.[151,1436,345,1919]" captionTargetId-1="figure-35@3.[151,1436,181,1804]" captionTargetId-2="figure-20@4.[151,1436,181,1813]" captionTargetId-3="figure-35@5.[151,1436,181,1807]" captionTargetId-4="figure-35@7.[151,1435,181,1770]" captionTargetId-5="figure-44@8.[151,1435,237,1952]" captionTargetId-6="figure-91@9.[151,1436,309,1891]" captionTargetId-7="figure-75@10.[151,1436,309,1779]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." captionText-1="FIGURES 13–18. Nigrobaetis (Margobaetis) minutus, larval mouthparts. 13, labrum; 14–15, left and right mandibles; 16, maxilla; 17, paraglossa (focus on dorsal side); 18, labium (focus on ventral side)." captionText-2="FIGURES 19–22. Nigrobaetis (Margobaetis) minutus, exuviae of larval tibia and tarsus. 19–21, fore, middle and hind legs; 22, hind leg of another specimen." captionText-3="FIGURES 23–25. Nigrobaetis (Margobaetis) minutus, exuviae of larval abdomen. 23, sterna VIII–IX of female; 24, sterna VIII–X of male; 25, fragment of cercus." captionText-4="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." captionText-5="FIGURES 33–34. Nigrobaetis (Margobaetis) minutus, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg." captionText-6="FIGURES 35–38. Nigrobaetis (Margobaetis) minutus, genitalia of male imago. 35, ventral view; 36, view with focus on gonovectes and penial bridge; 37, ventral view with focus on sterno-styligeral muscle and median sclerite; 38, lateral view. Abbreviation: m.s, median sterno-styligeral muscle." captionText-7="FIGURES 39–43. Nigrobaetis (Margobaetis) minutus, eggs." figureDoi-0="http://doi.org/10.5281/zenodo.10779997" figureDoi-1="http://doi.org/10.5281/zenodo.10780001" figureDoi-2="http://doi.org/10.5281/zenodo.10780003" figureDoi-3="http://doi.org/10.5281/zenodo.10780007" figureDoi-4="http://doi.org/10.5281/zenodo.10780011" figureDoi-5="http://doi.org/10.5281/zenodo.10780013" figureDoi-6="http://doi.org/10.5281/zenodo.10780015" figureDoi-7="http://doi.org/10.5281/zenodo.10780019" httpUri-0="https://zenodo.org/record/10779997/files/figure.png" httpUri-1="https://zenodo.org/record/10780001/files/figure.png" httpUri-2="https://zenodo.org/record/10780003/files/figure.png" httpUri-3="https://zenodo.org/record/10780007/files/figure.png" httpUri-4="https://zenodo.org/record/10780011/files/figure.png" httpUri-5="https://zenodo.org/record/10780013/files/figure.png" httpUri-6="https://zenodo.org/record/10780015/files/figure.png" httpUri-7="https://zenodo.org/record/10780019/files/figure.png" pageId="6" pageNumber="534">Figs 1–43</figureCitation>
)
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<emphasis id="72C8EAA4FFE7043059C1FCEFFED3FEE0" box="[151,301,251,274]" italics="true" pageId="6" pageNumber="534">Baetis minutus</emphasis>
<bibRefCitation id="242D4B47FFE704305865FCEEFDB5FEE0" author="Muller-Liebenau, I." box="[307,587,250,274]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984: 255</bibRefCitation>
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(larva).
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<emphasis id="72C8EAA4FFE7043059C1FD0EFF28FEC3" box="[151,214,282,305]" italics="true" pageId="6" pageNumber="534">Baetis</emphasis>
(
<emphasis id="72C8EAA4FFE7043059B3FD0DFEA2FEC3" box="[229,348,281,305]" italics="true" pageId="6" pageNumber="534">Nigrobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFE70430583DFD0EFE45FEC3" box="[363,443,282,305]" italics="true" pageId="6" pageNumber="534">minutus</emphasis>
</taxonomicName>
: 
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<bibRefCitation id="242D4B47FFE70430589EFD0DFD04FEC3" author="Novikova, E. A. &amp; Kluge, N. J." box="[456,762,281,305]" pageId="6" pageNumber="534" pagination="623 - 644" refId="ref7366" refString="Novikova, E. A. &amp; Kluge, N. J. (1994) Mayflies of the subgenus Nigrobaetis (Ephemeroptera, Baetidae, Baetis). Entomologicheskoe Obozrenie, 73 (3), 623 - 644. [in Russian]" type="journal article" year="1994">Novikova &amp; Kluge 1994: 627</bibRefCitation>
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.
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<emphasis id="72C8EAA4FFE7043059C1FD2CFEF0FEA2" box="[151,270,312,336]" italics="true" pageId="6" pageNumber="534">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFE70430584BFD2CFE60FEA2" box="[285,414,312,336]" italics="true" pageId="6" pageNumber="534">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFE7043058FAFD2CFE02FEBD" box="[428,508,312,335]" italics="true" pageId="6" pageNumber="534">minutus</emphasis>
</taxonomicName>
: 
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<bibRefCitation id="242D4B47FFE704305B5CFD2CFD4AFEBD" author="Kluge, N. J." box="[522,692,311,336]" pageId="6" pageNumber="534" pagination="151 - 186" refId="ref6828" refString="Kluge, N. J. (2022) Taxonomic significance of microlepides on subimaginal tarsi of Ephemeroptera. Zootaxa, 5159 (2), 151 - 186. https: // doi. org / 10.11646 / zootaxa. 5159.2.1" type="journal article" year="2022">Kluge 2022: 163</bibRefCitation>
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(subimago, misidentified).
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<taxonomicName id="87BC4D35FFE7043059C1FD42FCEAFE9C" authority="Kubendran &amp; Balasubramanian" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[151,788,342,366]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="534" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFE7043059C1FD42FE4BFE9C" box="[151,437,342,366]" italics="true" pageId="6" pageNumber="534">Nigrobaetis paramakalyani</emphasis>
Kubendran &amp; Balasubramanian
</taxonomicName>
<emphasis id="72C8EAA4FFE704305A75FD43FCC9FE9C" box="[803,823,343,366]" italics="true" pageId="6" pageNumber="534">in</emphasis>
<bibRefCitation id="242D4B47FFE704305A13FD42FA63FE9C" author="Kubendran, T. &amp; Balasubramanian, C. &amp; Selvakumar, C. &amp; Gattolliat, J. L. &amp; Sivaramakrishnan, K. G." box="[837,1437,342,366]" pageId="6" pageNumber="534" pagination="188 - 200" refId="ref7026" refString="Kubendran, T., Balasubramanian, C., Selvakumar, C., Gattolliat, J. L. &amp; Sivaramakrishnan, K. G. (2015) Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India). Zootaxa, 3957 (2), 188 - 200. https: // doi. org / 10.11646 / zootaxa. 3957.2.3" type="journal article" year="2015">Kubendran, Balasubramanian, Selvakumar, Gattolliat &amp;</bibRefCitation>
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Sivaramakrishnan 2015: 193 (larva, partim), 
<emphasis id="72C8EAA4FFE704305BCDFD61FD1DFE7F" bold="true" box="[667,739,373,397]" pageId="6" pageNumber="534">syn. n.</emphasis>
; 
<bibRefCitation id="242D4B47FFE704305BA5FD61FB00FE7F" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[755,1278,373,397]" pageId="6" pageNumber="534" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">Sivaruban, Srinivasan, Barathy &amp; Isack 2022: 189</bibRefCitation>
(corrections to larval description). 
<taxonomicName id="87BC4D35FFE7043059C1FDA6FD1DFE38" authority="Kaltenbach &amp; Gattolliat 2023: 196" authorityName="Kaltenbach &amp; Gattolliat" authorityPageNumber="196" authorityYear="2023" box="[151,739,434,458]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="534" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFE7043059C1FDA6FE82FE38" box="[151,380,434,458]" italics="true" pageId="6" pageNumber="534">Nigrobaetis sumbensis</emphasis>
<bibRefCitation id="242D4B47FFE7043058D5FDA6FD1DFE38" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[387,739,434,458]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023: 196</bibRefCitation>
</taxonomicName>
(larva), 
<emphasis id="72C8EAA4FFE704305A6FFDA6FC81FE38" bold="true" box="[825,895,434,458]" pageId="6" pageNumber="534">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="08A6653DFFE7043859C1FDE7FEE1FD3F" lastPageId="14" lastPageNumber="542" pageId="6" pageNumber="534" type="materials_examined">
<paragraph id="400336B6FFE7043059C1FDE7FD07FD33" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
<emphasis id="72C8EAA4FFE7043059C1FDE7FE72FDFF" bold="true" box="[151,396,499,525]" pageId="6" pageNumber="534">Material examined.</emphasis>
<materialsCitation id="F0D43CEBFFE7043058C8FDE7FC4CFDC3" collectingDate="2012-06-28" collectionCode="VI" collectorName="C. Balasubramanian &amp; T. Kubendran &amp; C. Selvakumar" country="India" county="State" location="Gadana river" municipality="Tirunelveli district" pageId="6" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<collectingCountry id="38AB7626FFE7043058C8FDE7FE0CFDFF" box="[414,498,499,525]" name="India" pageId="6" pageNumber="534">INDIA</collectingCountry>
: State 
<collectingRegion id="8278F854FFE704305B04FDE7FD18FDFF" box="[594,742,499,525]" country="India" name="Tamil Nadu" pageId="6" pageNumber="534">Tamil Nadu</collectingRegion>
: 
<collectingMunicipality id="A067ACCCFFE704305BAAFDE7FC25FDFF" box="[764,987,499,525]" pageId="6" pageNumber="534">Tirunelveli district</collectingMunicipality>
, 
<location id="4563606DFFE704305AA7FDE7FB6EFDFF" LSID="urn:lsid:plazi:treatment:C81587A0FFE7043859C1FC83FEE1FD3F:4563606DFFE704305AA7FDE7FB6EFDFF" box="[1009,1168,499,525]" country="India" county="State" municipality="Tirunelveli district" name="Gadana river" pageId="6" pageNumber="534" stateProvince="Tamil Nadu">Gadana river</location>
, 
<date id="34021076FFE704305DF0FDE7FAD7FDFF" box="[1190,1321,499,525]" pageId="6" pageNumber="534" value="2012-06-28">
<collectingDate id="2446E99EFFE704305DF0FDE7FAD7FDFF" box="[1190,1321,499,525]" pageId="6" pageNumber="534" value="2012-06-28">28.VI.2012</collectingDate>
</date>
, coll. 
<collectorName id="ED495360FFE704305CD5FDE7FEA2FDC3" pageId="6" pageNumber="534">C. Balasubramanian</collectorName>
, 
<collectorName id="ED495360FFE704305831FE0CFDFFFDC3" box="[359,513,535,562]" pageId="6" pageNumber="534">T. Kubendran</collectorName>
&amp; 
<collectorName id="ED495360FFE704305B73FE03FD2EFDC3" box="[549,720,535,561]" pageId="6" pageNumber="534">C. Selvakumar</collectorName>
: 
<specimenCount id="56BAFD3FFFE704305B8DFE0CFD0EFDC0" box="[731,752,536,562]" count="1" pageId="6" pageNumber="534" type="male">♂</specimenCount>
larva (
<typeStatus id="9F078814FFE704305A68FE03FC58FDC3" box="[830,934,535,561]" pageId="6" pageNumber="534" type="holotype">holotype</typeStatus>
),
</materialsCitation>
<materialsCitation id="F0D43CEBFFE704305AECFE03FB60FDC3" box="[954,1182,535,561]" collectingDate="2016-02-10" collectionCode="VI" collectorName="Kubendran &amp; Selvakumar &amp; Sheyko" country="India" county="State" location="Gadana river" municipality="Tirunelveli district" pageId="6" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="2" stateProvince="Tamil Nadu" typeStatus="paratype">
<specimenCount id="56BAFD3FFFE704305AECFE03FC22FDC3" box="[954,988,535,561]" count="1" pageId="6" pageNumber="534" type="female">1♀</specimenCount>
larva (
<typeStatus id="9F078814FFE704305D7DFE03FB6CFDC3" box="[1067,1170,535,561]" pageId="6" pageNumber="534" type="paratype">paratype</typeStatus>
);
</materialsCitation>
<materialsCitation id="F0D43CEBFFE704305DF3FE03FCA4FDA7" collectingDate="2016-02-10" collectingDateMax="2022-04-23" collectingDateMin="2016-02-10" collectionCode="ZIN" collectorName="N. Kluge &amp; Sheyko" country="India" county="State" location="river Vaigai" municipality="Madurai district" pageId="6" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<collectingMunicipality id="A067ACCCFFE704305DF3FE03FAA2FDC3" box="[1189,1372,535,561]" pageId="6" pageNumber="534">Madurai district</collectingMunicipality>
, 
<location id="4563606DFFE704305C31FE03FF1FFDA7" LSID="urn:lsid:plazi:treatment:C81587A0FFE7043859C1FC83FEE1FD3F:4563606DFFE704305C31FE03FF1FFDA7" country="India" county="State" municipality="Madurai district" name="river Vaigai" pageId="6" pageNumber="534" stateProvince="Tamil Nadu">river Vaigai</location>
, 
<date id="34021076FFE7043059BBFE2FFE9AFDA7" box="[237,356,571,597]" pageId="6" pageNumber="534" value="2016-02-10">
<collectingDate id="2446E99EFFE7043059BBFE2FFE9AFDA7" box="[237,356,571,597]" pageId="6" pageNumber="534" value="2016-02-10">10.II.2016</collectingDate>
</date>
, coll. 
<taxonomicName id="87BC4D35FFE7043058FEFE28FDF0FDA7" baseAuthorityName="Kluge" baseAuthorityYear="1983" box="[424,526,571,598]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="kluge">
<collectorName id="ED495360FFE7043058FEFE28FDF0FDA7" box="[424,526,571,598]" pageId="6" pageNumber="534">N. Kluge</collectorName>
</taxonomicName>
&amp; L. 
<collectorName id="ED495360FFE704305B04FE2FFD54FDA7" box="[594,682,571,597]" pageId="6" pageNumber="534">Sheyko</collectorName>
: 1 L-S♂ (
<collectionCode id="26ADAE73FFE704305A4FFE2FFCB3FDA7" box="[793,845,571,597]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="534" type="Museum">ZIN</collectionCode>
);
</materialsCitation>
<materialsCitation id="F0D43CEBFFE704305A37FE2FFE3FFD8B" collectingDate="2022-04-23" collectingDateMax="2022-04-23" collectingDateMin="2016-02-10" collectionCode="AMC" collectorName="P. Srinivasan &amp; R. Isack" country="India" county="State" location="river Vaigai" municipality="Madurai district" pageId="6" pageNumber="534" specimenCount="3" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
the same locality, 
<date id="34021076FFE704305D7AFE2FFB54FDA7" box="[1068,1194,571,597]" pageId="6" pageNumber="534" value="2022-04-23">
<collectingDate id="2446E99EFFE704305D7AFE2FFB54FDA7" box="[1068,1194,571,597]" pageId="6" pageNumber="534" value="2022-04-23">23.IV.2022</collectingDate>
</date>
, coll. 
<collectorName id="ED495360FFE704305DB8FE28FA81FDA7" box="[1262,1407,571,598]" pageId="6" pageNumber="534">P. Srinivasan</collectorName>
&amp; 
<collectorName id="ED495360FFE7043059C1FE74FF06FD8B" box="[151,248,607,634]" pageId="6" pageNumber="534">R. Isack</collectorName>
: 
<specimenCount id="56BAFD3FFFE704305852FE4BFEA0FD8B" box="[260,350,607,633]" count="3" pageId="6" pageNumber="534" type="larva">3 larvae</specimenCount>
(
<collectionCode id="26ADAE73FFE704305839FE4BFE4BFD8B" box="[367,437,607,633]" collectionName="AMC" country="USA" httpUri="http://grbio.org/cool/2xyg-zxc2" name="Department of Biologics Research" pageId="6" pageNumber="534">AMC</collectionCode>
).
</materialsCitation>
<materialsCitation id="F0D43CEBFFE70430589EFE4BFD0AFD6F" collectingDate="2016-01-22" collectionCode="ZIN" collectorName="N. Kluge &amp; L. Sheyko" country="India" county="State" location="Erumeli" municipality="Kottayam District" pageId="6" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="2" stateProvince="Kerala" typeStatus="holotype">
State 
<collectingRegion id="8278F854FFE704305B5FFE4BFDA8FD8B" box="[521,598,607,633]" country="India" name="Kerala" pageId="6" pageNumber="534">Kerala</collectingRegion>
, 
<collectingMunicipality id="A067ACCCFFE704305B34FE74FCCEFD8B" box="[610,816,607,634]" pageId="6" pageNumber="534">Kottayam District</collectingMunicipality>
, 
<location id="4563606DFFE704305A6AFE4BFC67FD8B" LSID="urn:lsid:plazi:treatment:C81587A0FFE7043859C1FC83FEE1FD3F:4563606DFFE704305A6AFE4BFC67FD8B" box="[828,921,607,633]" country="India" county="State" municipality="Kottayam District" name="Erumeli" pageId="6" pageNumber="534" stateProvince="Kerala">Erumeli</location>
, 
<date id="34021076FFE704305AF0FE4BFBEAFD8B" box="[934,1044,607,633]" pageId="6" pageNumber="534" value="2016-01-22">
<collectingDate id="2446E99EFFE704305AF0FE4BFBEAFD8B" box="[934,1044,607,633]" pageId="6" pageNumber="534" value="2016-01-22">22.I.2016</collectingDate>
</date>
, coll. 
<taxonomicName id="87BC4D35FFE704305D0FFE74FB3FFD8B" baseAuthorityName="Kluge" baseAuthorityYear="1983" box="[1113,1217,607,634]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="kluge">
<collectorName id="ED495360FFE704305D0FFE74FB3FFD8B" box="[1113,1217,607,634]" pageId="6" pageNumber="534">N. Kluge</collectorName>
</taxonomicName>
&amp; 
<collectorName id="ED495360FFE704305DB1FE74FA9EFD8B" box="[1255,1376,607,634]" pageId="6" pageNumber="534">L. Sheyko</collectorName>
: 1 L-S-I♂, L-S♂, 3 S-I♂, 1 S-I♀, 
<date id="34021076FFE7043058B5FE97FDFDFD6C" box="[483,515,643,670]" pageId="6" pageNumber="534">2 I</date>
♂, 19 S♂, 9 L (
<collectionCode id="26ADAE73FFE704305BE2FE97FD16FD6F" box="[692,744,643,669]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="534" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
<materialsCitation id="F0D43CEBFFE704305BAAFE97FD0AFD33" collectingDate="2009-08-28" collectingDateMax="2009-08-31" collectingDateMin="2009-08-28" collectionCode="ZIN" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" county="State" location="Pinrang" municipality="Tirunelveli district" pageId="6" pageNumber="534" specimenCount="1" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="2" stateProvince="Sulawesi" typeStatus="holotype">
<collectingCountry id="38AB7626FFE704305BAAFE97FC69FD6F" box="[764,919,643,669]" name="Indonesia" pageId="6" pageNumber="534">INDONESIA</collectingCountry>
, 
<collectingRegion id="8278F854FFE704305AF5FE97FBF5FD6F" box="[931,1035,643,669]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="534">Sulawesi</collectingRegion>
, 
<location id="4563606DFFE704305D41FE97FB8FFD6F" LSID="urn:lsid:plazi:treatment:C81587A0FFE7043859C1FC83FEE1FD3F:4563606DFFE704305D41FE97FB8FFD6F" box="[1047,1137,643,669]" country="Indonesia" county="State" municipality="Tirunelveli district" name="Pinrang" pageId="6" pageNumber="534" stateProvince="Sulawesi">Pinrang</location>
, 
<date id="34021076FFE704305D2BFE97FAC2FD6F" box="[1149,1340,643,669]" pageId="6" pageNumber="534" value="2009-08-28" valueMax="2009-08-31" valueMin="2009-08-28">
<collectingDate id="2446E99EFFE704305D2BFE97FAC2FD6F" box="[1149,1340,643,669]" pageId="6" pageNumber="534" value="2009-08-28" valueMax="2009-08-31" valueMin="2009-08-28">28–31.VIII.2009</collectingDate>
</date>
, coll. 
<taxonomicName id="87BC4D35FFE704305CD7FE90FF25FD33" baseAuthorityName="Kluge" baseAuthorityYear="1983" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="548" phylum="Arthropoda" rank="species" species="kluge">
<collectorName id="ED495360FFE704305CD7FE90FF25FD33" pageId="6" pageNumber="534">N. Kluge</collectorName>
</taxonomicName>
&amp; 
<collectorName id="ED495360FFE7043059A9FEBCFE89FD33" box="[255,375,679,706]" pageId="6" pageNumber="534">L. Sheyko</collectorName>
: 1 L/S♂, 1 L-S-I♀, 1 L/S♀ (
<collectionCode id="26ADAE73FFE704305BEEFEB3FD12FD33" box="[696,748,679,705]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="534" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="400336B6FFE704305991FEDFFA96FBDB" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
<emphasis id="72C8EAA4FFE704305991FEDFFDBDFD17" bold="true" box="[199,579,715,741]" pageId="6" pageNumber="534">Additional description of larva.</emphasis>
CUTICULAR COLORATION: With more or less contrasting brown and colorless or light areas (
<figureCitation id="D8872A33FFE7043058F3FEFBFDE7FCFB" box="[421,537,751,777]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Figs 1–12</figureCitation>
). Pronotum and mesonotum with colorless median stripe of variable, composite shape and paired colorless blanks (
<figureCitation id="D8872A33FFE704305B74FF07FD4AFCDF" box="[546,692,787,813]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Figs 6, 8, 12</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305B97FF07FBEAFCDF" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[705,1044,787,813]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 15a). Leg of each pair mostly colorless, with more or less expressed brown transverse band on femur, brown femur-tibia articulation and brown apex of tibia (
<figureCitation id="D8872A33FFE704305864FF4FFE20FC87" box="[306,478,859,885]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Figs 2–4, 9–10</figureCitation>
; 
<bibRefCitation id="242D4B47FFE7043058BCFF4FFD14FC87" author="Muller-Liebenau, I." box="[490,746,859,885]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 2i–j; 
<bibRefCitation id="242D4B47FFE704305A0AFF4FFB52FC87" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[860,1196,859,885]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 19a); sometimes nearly entirely colorless. Abdominal tergum I mostly colorless; terga II– 
<collectionCode id="26ADAE73FFE704305A9CFF94FC18FC68" box="[970,998,896,922]" pageId="6" pageNumber="534">III</collectionCode>
mostly brown, with contrasting paired and unpaired blanks; tergum 
<collectionCode id="26ADAE73FFE7043058B7FFB0FE01FC4C" box="[481,511,932,958]" pageId="6" pageNumber="534">IV</collectionCode>
mostly colorless, with pair of contrasting brown spots near anterior margin; terga 
<collectionCode id="26ADAE73FFE7043059C1FFDCFF55FC10" box="[151,171,968,994]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="534" type="Museum">V</collectionCode>
–VII mostly brown, with contrasting paired and unpaired blanks; terga 
<collectionCode id="26ADAE73FFE704305AB8FFDCFBE0FC10" box="[1006,1054,968,994]" pageId="6" pageNumber="534">VIII</collectionCode>
–X mostly colorless, with brown posterior margin of tergum X (
<figureCitation id="D8872A33FFE7043058BCFFFFFD88FBF7" box="[490,630,1003,1030]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Figs 5, 7, 11</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305BD7FFFFFC7EFBF7" author="Muller-Liebenau, I." box="[641,896,1003,1029]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 17; 
<bibRefCitation id="242D4B47FFE704305AB6FFFFFAD3FBF7" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[992,1325,1003,1029]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 15a). Caudalii mostly colorless, with brown band near middle and apically (
<figureCitation id="D8872A33FFE704305AFCF81BFC10FBD8" box="[938,1006,1039,1066]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Fig. 5</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305AADF81BFB05FBDB" author="Muller-Liebenau, I." box="[1019,1275,1039,1065]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 
<quantity id="87449B53FFE704305C65F81BFAA5FBDB" box="[1331,1371,1039,1065]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="534" unit="m" value="2.0">2m</quantity>
).
</paragraph>
<paragraph id="400336B6FFE70430599EF827FCF2FBBF" blockId="6.[151,1437,499,2038]" box="[200,780,1075,1102]" pageId="6" pageNumber="534">HYPODERMAL COLORATION: Not expressed.</paragraph>
<paragraph id="400336B6FFE70430599EF843FE98FB4B" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
<collectionCode id="26ADAE73FFE70430599EF843FE01FB83" box="[200,511,1111,1137]" pageId="6" pageNumber="534">SHAPE AND SETATION</collectionCode>
: Labrum parallel-sided in proximal half (
<figureCitation id="D8872A33FFE704305A8DF843FBCEFB83" box="[987,1072,1111,1137]" captionStart="FIGURES 13–18" captionStartId="3.[151,264,1827,1851]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-35@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 13–18. Nigrobaetis (Margobaetis) minutus, larval mouthparts. 13, labrum; 14–15, left and right mandibles; 16, maxilla; 17, paraglossa (focus on dorsal side); 18, labium (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.10780001" httpUri="https://zenodo.org/record/10780001/files/figure.png" pageId="6" pageNumber="534">Fig. 13</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305D6BF843FAC1FB83" author="Muller-Liebenau, I." box="[1085,1343,1111,1137]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 2a; 
<bibRefCitation id="242D4B47FFE7043059C1F86FFE10FB67" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[151,494,1147,1173]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: figs 16a–b). Other mouthparts as in photos (
<figureCitation id="D8872A33FFE704305AAEF86FFB83FB67" box="[1016,1149,1147,1173]" captionStart="FIGURES 13–18" captionStartId="3.[151,264,1827,1851]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-35@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 13–18. Nigrobaetis (Margobaetis) minutus, larval mouthparts. 13, labrum; 14–15, left and right mandibles; 16, maxilla; 17, paraglossa (focus on dorsal side); 18, labium (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.10780001" httpUri="https://zenodo.org/record/10780001/files/figure.png" pageId="6" pageNumber="534">Figs 14–18</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305DDDF86FFF2DFB4B" author="Kaltenbach, T. &amp; Gattolliat, J. - L." pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: figs 16–18).
</paragraph>
<paragraph id="400336B6FFE704305991F8D7FC09FB2F" blockId="6.[151,1437,499,2038]" box="[199,1015,1219,1246]" pageId="6" pageNumber="534">
Hind protoptera or their vestiges absent (
<bibRefCitation id="242D4B47FFE704305BC6F8D7FC6CFB2F" author="Muller-Liebenau, I." box="[656,914,1219,1245]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 2h).
</paragraph>
<paragraph id="400336B6FFE704305991F8F3FA7DFABB" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
Each tibia with one stout, pointed seta on outer side near apex (subapical seta); middle and hind tibiae, besides subapical seta, with several (2–10) smaller, pointed setae forming irregular longitudinal row on outer side; outer side of fore tibia with no more than one stout seta other than subapical one (
<figureCitation id="D8872A33FFE704305AD1F93BFBF9FABB" box="[903,1031,1327,1353]" captionStart="FIGURES 19–22" captionStartId="4.[151,264,1837,1861]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-20@4.[151,1436,181,1813]" captionTargetPageId="4" captionText="FIGURES 19–22. Nigrobaetis (Margobaetis) minutus, exuviae of larval tibia and tarsus. 19–21, fore, middle and hind legs; 22, hind leg of another specimen." figureDoi="http://doi.org/10.5281/zenodo.10780003" httpUri="https://zenodo.org/record/10780003/files/figure.png" pageId="6" pageNumber="534">Figs 19–22</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305D44F93BFAF3FABB" author="Muller-Liebenau, I." box="[1042,1293,1327,1353]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: figs 2i–j).
</paragraph>
<paragraph id="400336B6FFE704305991F947FAECF9B7" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
Posterior margins of abdominal terga II–X with triangular denticles, shorter and blunter on anterior segments, longer and pointer on posterior segments (
<bibRefCitation id="242D4B47FFE704305B2EF963FC35FA63" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[632,971,1399,1425]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 2a). Posterior margins of abdominal sterna I– 
<collectionCode id="26ADAE73FFE7043059ADF988FEE7FA44" box="[251,281,1436,1462]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="534">VI</collectionCode>
smooth; posterior margins of sterna VII–IX with sharply pointed, triangular denticles (
<figureCitation id="D8872A33FFE704305C5CF98FFA6FFA47" box="[1290,1425,1435,1461]" captionStart="FIGURES 23–25" captionStartId="5.[151,264,1831,1855]" captionTargetBox="[151,1436,181,1807]" captionTargetId="figure-35@5.[151,1436,181,1807]" captionTargetPageId="5" captionText="FIGURES 23–25. Nigrobaetis (Margobaetis) minutus, exuviae of larval abdomen. 23, sterna VIII–IX of female; 24, sterna VIII–X of male; 25, fragment of cercus." figureDoi="http://doi.org/10.5281/zenodo.10780007" httpUri="https://zenodo.org/record/10780007/files/figure.png" pageId="6" pageNumber="534">Figs 23–24</figureCitation>
). Paraproct with many small denticles (
<figureCitation id="D8872A33FFE704305B0EF9ABFD4FFA2B" box="[600,689,1471,1497]" captionStart="FIGURES 23–25" captionStartId="5.[151,264,1831,1855]" captionTargetBox="[151,1436,181,1807]" captionTargetId="figure-35@5.[151,1436,181,1807]" captionTargetPageId="5" captionText="FIGURES 23–25. Nigrobaetis (Margobaetis) minutus, exuviae of larval abdomen. 23, sterna VIII–IX of female; 24, sterna VIII–X of male; 25, fragment of cercus." figureDoi="http://doi.org/10.5281/zenodo.10780007" httpUri="https://zenodo.org/record/10780007/files/figure.png" pageId="6" pageNumber="534">Fig. 24</figureCitation>
; 
<bibRefCitation id="242D4B47FFE704305B94F9ABFC39FA2B" author="Muller-Liebenau, I." box="[706,967,1471,1497]" pageId="6" pageNumber="534" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 2i; 
<bibRefCitation id="242D4B47FFE704305D65F9ABFAC8FA2B" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[1075,1334,1471,1498]" pageId="6" pageNumber="534" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">
Sivaruban 
<emphasis id="72C8EAA4FFE704305DE7F9D4FB19FA2B" box="[1201,1255,1471,1497]" italics="true" pageId="6" pageNumber="534">et al</emphasis>
. 2022
</bibRefCitation>
: fig. 4d; 
<bibRefCitation id="242D4B47FFE7043059C1F9F7FE13FA0F" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[151,493,1507,1533]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 20b). All 7 pairs of tergalii present, relatively long and narrow (
<figureCitation id="D8872A33FFE704305C55F9F7FA66FA0F" box="[1283,1432,1507,1534]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="6" pageNumber="534">Figs 5, 7, 11</figureCitation>
; 
<bibRefCitation id="242D4B47FFE7043059C1FA13FE77F9D3" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[151,393,1543,1570]" pageId="6" pageNumber="534" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">
Sivaruban 
<emphasis id="72C8EAA4FFE704305859FA1CFEC1F9D3" box="[271,319,1543,1569]" italics="true" pageId="6" pageNumber="534">et al</emphasis>
. 2022
</bibRefCitation>
: figs 4B, E; 
<bibRefCitation id="242D4B47FFE704305B46FA13FCA0F9D3" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[528,862,1543,1569]" pageId="6" pageNumber="534" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: figs 15a, 19d–e). In middle part of cercus, each 4th segment with several enlarged denticles on outer side (
<figureCitation id="D8872A33FFE704305BAAFA3FFCB1F9B4" box="[764,847,1579,1606]" captionStart="FIGURES 23–25" captionStartId="5.[151,264,1831,1855]" captionTargetBox="[151,1436,181,1807]" captionTargetId="figure-35@5.[151,1436,181,1807]" captionTargetPageId="5" captionText="FIGURES 23–25. Nigrobaetis (Margobaetis) minutus, exuviae of larval abdomen. 23, sterna VIII–IX of female; 24, sterna VIII–X of male; 25, fragment of cercus." figureDoi="http://doi.org/10.5281/zenodo.10780007" httpUri="https://zenodo.org/record/10780007/files/figure.png" pageId="6" pageNumber="534">Fig. 25</figureCitation>
); paracercus without enlarged denticles.
</paragraph>
<paragraph id="400336B6FFE704305991FA5BFDD8F99B" blockId="6.[151,1437,499,2038]" box="[199,550,1615,1641]" pageId="6" pageNumber="534">
<emphasis id="72C8EAA4FFE704305991FA5BFDD8F99B" bold="true" box="[199,550,1615,1641]" pageId="6" pageNumber="534">Descriptions of winged stages</emphasis>
</paragraph>
<paragraph id="400336B6FFE704305991FA67FABCF927" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
<emphasis id="72C8EAA4FFE704305991FA67FEBDF97F" bold="true" box="[199,323,1651,1678]" pageId="6" pageNumber="534">
<emphasis id="72C8EAA4FFE704305991FA67FEC2F97C" bold="true" box="[199,316,1651,1678]" italics="true" pageId="6" pageNumber="534">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head colorless, antennal flagellum light brown. Pronotum colorless. Mesonotum light brownish, with brown sutures (
<figureCitation id="D8872A33FFE704305A6FFA83FC71F943" box="[825,911,1687,1713]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="6" pageNumber="534">Fig. 28</figureCitation>
). Thoracic pleura and sterna mostly colorless, with certain sclerites brown (
<figureCitation id="D8872A33FFE70430588BFAAFFDCFF927" box="[477,561,1723,1749]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="6" pageNumber="534">Fig. 27</figureCitation>
). Legs colorless. Abdomen colorless, caudalii and gonostyli colorless.
</paragraph>
<paragraph id="400336B6FFE70430599EFACBFD0AF90B" blockId="6.[151,1437,499,2038]" box="[200,756,1759,1786]" pageId="6" pageNumber="534">HYPODERMAL COLORATION: As in imago.</paragraph>
<paragraph id="400336B6FFE704305991FB10FBD9F807" blockId="6.[151,1437,499,2038]" pageId="6" pageNumber="534">
TEXTURE: On middle and hind legs of both sexes, proximal tarsal segment (corresponding to initial 1st+2nd tarsomeres) in proximal part covered with blunt microlepides, in distal part covered with pointed microlepides; other three segments entirely covered with pointed microlepides (
<figureCitation id="D8872A33FFE704305ADDFB5FFC1CF897" box="[907,994,1867,1893]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,1976,2000]" captionTargetBox="[151,1435,237,1952]" captionTargetId="figure-44@8.[151,1435,237,1952]" captionTargetPageId="8" captionText="FIGURES 33–34. Nigrobaetis (Margobaetis) minutus, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg." figureDoi="http://doi.org/10.5281/zenodo.10780013" httpUri="https://zenodo.org/record/10780013/files/figure.png" pageId="6" pageNumber="534">Fig. 34</figureCitation>
). On fore leg of female, 1st tarsomere covered with pointed microlepides; 2nd tarsomere in proximal part covered with blunt microlepides, in distal part covered with pointed microlepides; 3rd–5th tarsomeres entirely covered with pointed microlepides (similar to 
<figureCitation id="D8872A33FFE704305C26FB87FF49F823" captionStart="FIGURES 33–34" captionStartId="8.[151,264,1976,2000]" captionTargetBox="[151,1435,237,1952]" captionTargetId="figure-44@8.[151,1435,237,1952]" captionTargetPageId="8" captionText="FIGURES 33–34. Nigrobaetis (Margobaetis) minutus, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg." figureDoi="http://doi.org/10.5281/zenodo.10780013" httpUri="https://zenodo.org/record/10780013/files/figure.png" pageId="6" pageNumber="534">Fig. 34</figureCitation>
). On fore leg of male, 1st–4th tarsal segments covered mostly with blunt microlepides, apically with pointed microlepides; 5th tarsomere entirely covered with pointed microlepides (
<figureCitation id="D8872A33FFE704305A91FBCFFBE4F807" box="[967,1050,2011,2037]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,1976,2000]" captionTargetBox="[151,1435,237,1952]" captionTargetId="figure-44@8.[151,1435,237,1952]" captionTargetPageId="8" captionText="FIGURES 33–34. Nigrobaetis (Margobaetis) minutus, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg." figureDoi="http://doi.org/10.5281/zenodo.10780013" httpUri="https://zenodo.org/record/10780013/files/figure.png" pageId="6" pageNumber="534">Fig. 33</figureCitation>
).
</paragraph>
<caption id="14C3663EFFE6043159C1FB16FB53F8CC" ID-DOI="http://doi.org/10.5281/zenodo.10780011" ID-Zenodo-Dep="10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="7" pageNumber="535" startId="7.[151,264,1794,1818]" targetBox="[151,1416,181,1770]" targetPageId="7" targetType="figure">
<paragraph id="400336B6FFE6043159C1FB16FB53F8CC" blockId="7.[151,1436,1794,1854]" pageId="7" pageNumber="535">
<emphasis id="72C8EAA4FFE6043159C1FB16FEAAF8E8" bold="true" box="[151,340,1794,1818]" pageId="7" pageNumber="535">FIGURES 26–32.</emphasis>
<taxonomicName id="87BC4D35FFE60431580CFB16FD41F8E8" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[346,703,1794,1818]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="535" phylum="Arthropoda" rank="species" species="minutus" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFE60431580CFB16FE2FF8E8" box="[346,465,1794,1818]" italics="true" pageId="7" pageNumber="535">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFE6043158B6FB16FD9FF8E8" box="[480,609,1794,1818]" italics="true" pageId="7" pageNumber="535">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFE604315B39FB17FD41F8E8" box="[623,703,1795,1818]" italics="true" pageId="7" pageNumber="535">minutus</emphasis>
</taxonomicName>
. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago.
</paragraph>
</caption>
<paragraph id="400336B6FFE6043E5991FB78FDFAFF27" blockId="7.[151,1437,1899,2035]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="536" pageId="7" pageNumber="535">
<emphasis id="72C8EAA4FFE604315991FB78FEA4F874" bold="true" box="[199,346,1899,1926]" italics="true" pageId="7" pageNumber="535">Imago, male</emphasis>
(
<figureCitation id="D8872A33FFE60431583AFB78FE3DF874" box="[364,451,1900,1926]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="7" pageNumber="535">Fig. 26</figureCitation>
). Head brown. Turbinate eyes dark reddish-brown. Thorax brown with ochre, equally dark dorsally, laterally and ventrally. Wings colorless, veins ochre-brownish, C and Sc+ 
<collectionCode id="26ADAE73FFE604315D3BFB85FB7EF859" box="[1133,1152,1937,1963]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="535">R</collectionCode>
proximad of costal brace brown. Pterostigma with simple, oblique crossveins. Hind wings absent. Legs of all pairs light ochre, either unicolor (
<figureCitation id="D8872A33FFE6043159C9FBCCFF08F800" box="[159,246,2008,2034]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="7" pageNumber="535">Fig. 26</figureCitation>
), or with apex of femur brown (
<figureCitation id="D8872A33FFE604315B27FBCCFD06F800" box="[625,760,2008,2034]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="7" pageNumber="535">Figs 29–30</figureCitation>
). On middle and hind legs, tarsus with 2 apical spines, on 1st+2nd and 3rd tarsomeres. Abdominal terga and sterna I– 
<collectionCode id="26ADAE73FFE9043E5A7DFC83FCB7FF43" box="[811,841,151,177]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="536">VI</collectionCode>
colorless, translucent, with brown spots on spiracles; terga VII–X brown. Cerci ochre.
</paragraph>
<caption id="14C3663EFFE9043E59C1FBACFEB3F806" ID-DOI="http://doi.org/10.5281/zenodo.10780013" ID-Zenodo-Dep="10780013" httpUri="https://zenodo.org/record/10780013/files/figure.png" pageId="8" pageNumber="536" startId="8.[151,264,1976,2000]" targetBox="[151,1435,237,1952]" targetPageId="8" targetType="figure">
<paragraph id="400336B6FFE9043E59C1FBACFEB3F806" blockId="8.[151,1437,1976,2036]" pageId="8" pageNumber="536">
<emphasis id="72C8EAA4FFE9043E59C1FBACFEAAF822" bold="true" box="[151,340,1976,2000]" pageId="8" pageNumber="536">FIGURES 33–34.</emphasis>
<taxonomicName id="87BC4D35FFE9043E580FFBACFD42F822" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[345,700,1976,2000]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="536" phylum="Arthropoda" rank="species" species="minutus" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFE9043E580FFBACFE2EF822" box="[345,464,1976,2000]" italics="true" pageId="8" pageNumber="536">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFE9043E5888FBACFDA1F822" box="[478,607,1976,2000]" italics="true" pageId="8" pageNumber="536">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFE9043E5B3AFBADFD42F822" box="[620,700,1977,2000]" italics="true" pageId="8" pageNumber="536">minutus</emphasis>
</taxonomicName>
, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg.
</paragraph>
</caption>
<paragraph id="400336B6FFE8043F5991FC83FCE8FEEF" blockId="9.[151,1437,151,285]" pageId="9" pageNumber="537">
Male genitalia (
<figureCitation id="D8872A33FFE8043F582EFC83FE02FF43" box="[376,508,151,177]" captionStart-0="FIGURES 33–34" captionStart-1="FIGURES 35–38" captionStartId-0="8.[151,264,1976,2000]" captionStartId-1="9.[151,264,1914,1938]" captionTargetBox-0="[151,1435,237,1952]" captionTargetBox-1="[166,1436,309,1891]" captionTargetId-0="figure-44@8.[151,1435,237,1952]" captionTargetId-1="figure-91@9.[151,1436,309,1891]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 33–34. Nigrobaetis (Margobaetis) minutus, male subimago. 33, 3rd–5th tarsomeres of fore leg, 34, apex of tibia and tarsus of hind leg." captionText-1="FIGURES 35–38. Nigrobaetis (Margobaetis) minutus, genitalia of male imago. 35, ventral view; 36, view with focus on gonovectes and penial bridge; 37, ventral view with focus on sterno-styligeral muscle and median sclerite; 38, lateral view. Abbreviation: m.s, median sterno-styligeral muscle." figureDoi-0="http://doi.org/10.5281/zenodo.10780013" figureDoi-1="http://doi.org/10.5281/zenodo.10780015" httpUri-0="https://zenodo.org/record/10780013/files/figure.png" httpUri-1="https://zenodo.org/record/10780015/files/figure.png" pageId="9" pageNumber="537">Figs 33–38</figureCitation>
): Styliger with well-outlined, brown median sclerite, to which distal end of sternostyligeral muscle is attached (
<figureCitation id="D8872A33FFE8043F588DFCAFFDA4FF27" box="[475,602,186,213]" captionStart="FIGURES 35–38" captionStartId="9.[151,264,1914,1938]" captionTargetBox="[166,1436,309,1891]" captionTargetId="figure-91@9.[151,1436,309,1891]" captionTargetPageId="9" captionText="FIGURES 35–38. Nigrobaetis (Margobaetis) minutus, genitalia of male imago. 35, ventral view; 36, view with focus on gonovectes and penial bridge; 37, ventral view with focus on sterno-styligeral muscle and median sclerite; 38, lateral view. Abbreviation: m.s, median sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.10780015" httpUri="https://zenodo.org/record/10780015/files/figure.png" pageId="9" pageNumber="537">Figs 35, 37</figureCitation>
). 1st segment of gonostylus narrowing from base to apex, angulate apically-medially. 3rd (terminal) segment of gonostylus elongate. Penial bridge with prominent, cone-like, sclerotized median projection. Gonovectes sharply bent, with narrow apices.
</paragraph>
<caption id="14C3663EFFE8043F59C1FB6EFD58F829" ID-DOI="http://doi.org/10.5281/zenodo.10780015" ID-Zenodo-Dep="10780015" httpUri="https://zenodo.org/record/10780015/files/figure.png" pageId="9" pageNumber="537" startId="9.[151,264,1914,1938]" targetBox="[166,1436,309,1891]" targetPageId="9" targetType="figure">
<paragraph id="400336B6FFE8043F59C1FB6EFD58F829" blockId="9.[151,1437,1914,2011]" pageId="9" pageNumber="537">
<emphasis id="72C8EAA4FFE8043F59C1FB6EFEA6F861" bold="true" box="[151,344,1914,1939]" pageId="9" pageNumber="537">FIGURES 35–38.</emphasis>
<taxonomicName id="87BC4D35FFE8043F5834FB6FFD30F860" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[354,718,1915,1939]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="537" phylum="Arthropoda" rank="species" species="minutus" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFE8043F5834FB6FFE27F861" box="[354,473,1915,1939]" italics="true" pageId="9" pageNumber="537">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFE8043F58BDFB6FFD92F861" box="[491,620,1915,1939]" italics="true" pageId="9" pageNumber="537">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFE8043F5B28FB6FFD30F860" box="[638,718,1915,1938]" italics="true" pageId="9" pageNumber="537">minutus</emphasis>
</taxonomicName>
, genitalia of male imago. 35, ventral view; 36, view with focus on gonovectes and penial bridge; 37, ventral view with focus on sterno-styligeral muscle and median sclerite; 38, lateral view. Abbreviation: m.s, median sterno-styligeral muscle.
</paragraph>
</caption>
<paragraph id="400336B6FFEB043C5991FC83FC9EFF27" blockId="10.[151,1436,150,285]" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C5991FC83FE91FF43" bold="true" box="[199,367,150,177]" italics="true" pageId="10" pageNumber="538">Imago, female</emphasis>
. All abdominal terga brown, sterna ochre with pair of brown stripes laterally (
<figureCitation id="D8872A33FFEB043C5DB4FC83FA98FF43" box="[1250,1382,151,177]" captionStart="FIGURES 26–32" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[151,1416,181,1770]" captionTargetId="figure-35@7.[151,1435,181,1770]" captionTargetPageId="7" captionText="FIGURES 26–32. Nigrobaetis (Margobaetis) minutus. 26, male imago; 27, subimaginal exuviae of meso- and metapleura; 28, subimaginal exuviae of mesonotum; 29–30, fore and middle legs of male imago; 31–32, female imago." figureDoi="http://doi.org/10.5281/zenodo.10780011" httpUri="https://zenodo.org/record/10780011/files/figure.png" pageId="10" pageNumber="538">Figs 31–32</figureCitation>
). On fore leg, tarsus with 2 apical spines, on 2nd and 3rd tarsomeres.
</paragraph>
<paragraph id="400336B6FFEB043C5991FCCBFC28FEEF" blockId="10.[151,1436,150,285]" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C5991FCCBFF08FF0B" bold="true" box="[199,246,223,249]" italics="true" pageId="10" pageNumber="538">Egg</emphasis>
(
<figureCitation id="D8872A33FFEB043C5850FCCBFE74FF0B" box="[262,394,223,249]" captionStart="FIGURES 39–43" captionStartId="10.[151,264,1802,1826]" captionTargetBox="[151,1436,309,1779]" captionTargetId="figure-75@10.[151,1436,309,1779]" captionTargetPageId="10" captionText="FIGURES 39–43. Nigrobaetis (Margobaetis) minutus, eggs." figureDoi="http://doi.org/10.5281/zenodo.10780019" httpUri="https://zenodo.org/record/10780019/files/figure.png" pageId="10" pageNumber="538">Figs 39–43</figureCitation>
). Asymmetric, generally irregularly-oval, with one side more or less stretched. Surface with evenly dispersed small round papillae and irregular folds. Micropile wide.
</paragraph>
<caption id="14C3663EFFEB043C59C1FB1EFCFFF8D1" ID-DOI="http://doi.org/10.5281/zenodo.10780019" ID-Zenodo-Dep="10780019" box="[151,769,1802,1827]" httpUri="https://zenodo.org/record/10780019/files/figure.png" pageId="10" pageNumber="538" startId="10.[151,264,1802,1826]" targetBox="[151,1436,309,1779]" targetPageId="10" targetType="figure">
<paragraph id="400336B6FFEB043C59C1FB1EFCFFF8D1" blockId="10.[151,769,1802,1827]" box="[151,769,1802,1827]" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C59C1FB1EFEAAF8D1" bold="true" box="[151,340,1802,1827]" pageId="10" pageNumber="538">FIGURES 39–43.</emphasis>
<taxonomicName id="87BC4D35FFEB043C580CFB1FFD41F8D0" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[346,703,1803,1827]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="minutus" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFEB043C580CFB1FFE2FF8D1" box="[346,465,1803,1827]" italics="true" pageId="10" pageNumber="538">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFEB043C58B6FB1FFD9FF8D1" box="[480,609,1803,1827]" italics="true" pageId="10" pageNumber="538">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFEB043C5B39FB1FFD41F8D0" box="[623,703,1803,1826]" italics="true" pageId="10" pageNumber="538">minutus</emphasis>
</taxonomicName>
, eggs.
</paragraph>
</caption>
<paragraph id="400336B6FFEB043C5991FB45FB3EF899" blockId="10.[151,1437,1873,2044]" box="[199,1216,1873,1900]" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C5991FB45FEB0F899" bold="true" box="[199,334,1873,1899]" pageId="10" pageNumber="538">Dimension.</emphasis>
Fore wing length (and approximate body length): male 3.5 mm, female 
<quantity id="87449B53FFEB043C5D2EFB45FB42F899" box="[1144,1212,1873,1899]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="538" unit="mm" value="4.0">4 mm</quantity>
.
</paragraph>
<paragraph id="400336B6FFEB043C5991FB61FEF2F825" blockId="10.[151,1437,1873,2044]" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C5991FB61FE9FF87D" bold="true" box="[199,353,1909,1935]" pageId="10" pageNumber="538">Distribution.</emphasis>
Oriental Region. Described from West 
<collectingCountry id="38AB7626FFEB043C5A67FB61FC66F87D" box="[817,920,1909,1935]" name="Malaysia" pageId="10" pageNumber="538">Malaysia</collectingCountry>
(as 
<taxonomicName id="87BC4D35FFEB043C5A9BFB62FBB7F87D" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[973,1097,1910,1935]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEB043C5A9BFB62FBB7F87D" box="[973,1097,1910,1935]" italics="true" pageId="10" pageNumber="538">B. minutus</emphasis>
</taxonomicName>
), from Southern 
<collectingCountry id="38AB7626FFEB043C5C42FB61FAB0F87D" box="[1300,1358,1909,1935]" name="India" pageId="10" pageNumber="538">India</collectingCountry>
(as 
<taxonomicName id="87BC4D35FFEB043C5CD5FB62FEBFF841" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEB043C5CD5FB62FEBFF841" italics="true" pageId="10" pageNumber="538">N. paramakalyani</emphasis>
</taxonomicName>
) and from Sumba Island in 
<collectingCountry id="38AB7626FFEB043C5BC6FB8DFD03F841" box="[656,765,1945,1971]" name="Indonesia" pageId="10" pageNumber="538">Indonesia</collectingCountry>
(as 
<taxonomicName id="87BC4D35FFEB043C5A65FB8EFC34F841" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[819,970,1945,1971]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEB043C5A65FB8EFC34F841" box="[819,970,1945,1971]" italics="true" pageId="10" pageNumber="538">N. sumbensis</emphasis>
</taxonomicName>
); reported here from 
<collectingRegion id="8278F854FFEB043C5D9FFB8DFAD0F841" box="[1225,1326,1945,1971]" country="Indonesia" name="Sulawesi Utara" pageId="10" pageNumber="538">Sulawesi</collectingRegion>
island in 
<collectingCountry id="38AB7626FFEB043C59C1FBA9FEF6F825" box="[151,264,1981,2007]" name="Indonesia" pageId="10" pageNumber="538">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="400336B6FFEB043D5991FBF5FA6CFF0B" blockId="10.[151,1437,1873,2044]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="538">
<emphasis id="72C8EAA4FFEB043C5991FBF5FD02F809" bold="true" box="[199,764,2016,2043]" pageId="10" pageNumber="538">
Errors in original description of 
<taxonomicName id="87BC4D35FFEB043C5B78FBF5FD0BF809" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[558,757,2016,2043]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEB043C5B78FBF5FD0BF809" bold="true" box="[558,757,2016,2043]" italics="true" pageId="10" pageNumber="538">N. paramakalyani</emphasis>
</taxonomicName>
.
</emphasis>
Originally(
<bibRefCitation id="242D4B47FFEB043C5A2DFBF5FB98F80E" author="Kubendran, T. &amp; Balasubramanian, C. &amp; Selvakumar, C. &amp; Gattolliat, J. L. &amp; Sivaramakrishnan, K. G." box="[891,1126,2017,2044]" pageId="10" pageNumber="538" pagination="188 - 200" refId="ref7026" refString="Kubendran, T., Balasubramanian, C., Selvakumar, C., Gattolliat, J. L. &amp; Sivaramakrishnan, K. G. (2015) Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India). Zootaxa, 3957 (2), 188 - 200. https: // doi. org / 10.11646 / zootaxa. 3957.2.3" type="journal article" year="2015">
Kubendran 
<emphasis id="72C8EAA4FFEB043C5AA0FBF6FBDCF809" box="[1014,1058,2017,2043]" italics="true" pageId="10" pageNumber="538">et al</emphasis>
.2015
</bibRefCitation>
), 
<taxonomicName id="87BC4D35FFEB043C5D23FBF5FA62F809" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[1141,1436,2017,2043]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="538" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEB043C5D23FBF5FA62F809" box="[1141,1436,2017,2043]" italics="true" pageId="10" pageNumber="538">Nigrobaetis paramakalyani</emphasis>
</taxonomicName>
was described based on 
<specimenCount id="56BAFD3FFFEA043D58FDFC83FDEDFF43" box="[427,531,151,177]" count="14" pageId="11" pageNumber="539" type="larva">14 larvae</specimenCount>
collected in Gadana River in Southern 
<collectingCountry id="38AB7626FFEA043D5A81FC83FBEAFF43" box="[983,1044,151,177]" name="India" pageId="11" pageNumber="539">India</collectingCountry>
. Actually, these larvae represented a mixture of two different species, the second one of which was subsequently described as 
<taxonomicName id="87BC4D35FFEA043D5D9CFCAFFE75FF0B" authority="Sivaruban et al. 2022" authorityName="Sivaruban" authorityYear="2022" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="klugei">
<emphasis id="72C8EAA4FFEA043D5D9CFCAFFA62FF27" box="[1226,1436,187,213]" italics="true" pageId="11" pageNumber="539">Nigrobaetis klugei</emphasis>
<bibRefCitation id="242D4B47FFEA043D59C1FCCBFE75FF0B" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[151,395,223,249]" pageId="11" pageNumber="539" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">
Sivaruban 
<emphasis id="72C8EAA4FFEA043D5846FCCBFEBFFF0B" box="[272,321,223,249]" italics="true" pageId="11" pageNumber="539">et al</emphasis>
. 2022
</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="9F078814FFEA043D58C3FCCBFE03FF0B" box="[405,509,223,249]" pageId="11" pageNumber="539" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="87BC4D35FFEA043D5B74FCCBFD13FF0B" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[546,749,223,249]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEA043D5B74FCCBFD13FF0B" box="[546,749,223,249]" italics="true" pageId="11" pageNumber="539">N. paramakalyani</emphasis>
</taxonomicName>
is the mature male larva (
<bibRefCitation id="242D4B47FFEA043D5D59FCCBFAFAFF0B" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[1039,1284,223,249]" pageId="11" pageNumber="539" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">
Sivaruban 
<emphasis id="72C8EAA4FFEA043D5DDFFCCBFB45FF0B" box="[1161,1211,223,249]" italics="true" pageId="11" pageNumber="539">et al</emphasis>
. 2022
</bibRefCitation>
: figs 4A–E).
</paragraph>
<caption id="14C3663EFFEA043D59C1FB9FFEEBF818" ID-DOI="http://doi.org/10.5281/zenodo.10780023" ID-Zenodo-Dep="10780023" httpUri="https://zenodo.org/record/10780023/files/figure.png" pageId="11" pageNumber="539" startId="11.[151,264,1931,1955]" targetBox="[151,1434,277,1902]" targetPageId="11" targetType="figure">
<paragraph id="400336B6FFEA043D59C1FB9FFEEBF818" blockId="11.[151,1437,1931,2026]" pageId="11" pageNumber="539">
<emphasis id="72C8EAA4FFEA043D59C1FB9FFEAAF851" bold="true" box="[151,340,1931,1955]" pageId="11" pageNumber="539">FIGURES 44–52.</emphasis>
<taxonomicName id="87BC4D35FFEA043D580CFB9FFD53F851" box="[346,685,1931,1955]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="539" phylum="Arthropoda" rank="species" species="klugei" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFEA043D580CFB9FFE2FF851" box="[346,465,1931,1955]" italics="true" pageId="11" pageNumber="539">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFEA043D58B6FB9FFD9FF851" box="[480,609,1931,1955]" italics="true" pageId="11" pageNumber="539">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFEA043D5B39FB9FFD53F851" box="[623,685,1931,1955]" italics="true" pageId="11" pageNumber="539">klugei</emphasis>
</taxonomicName>
, larval exuviae. 44, head; 45, prosternum with fore leg; 46, mesosternum, metasternum and metanotum with hind protoptera, middle and hind legs; 47, 49, 51, abdomen; 48, 50, 52, half of pronotum and mesonotum.
</paragraph>
</caption>
<caption id="14C3663EFFED043A59C1FB27FBBAF8B9" ID-DOI="http://doi.org/10.5281/zenodo.10780027" ID-Zenodo-Dep="10780027" box="[151,1092,1843,1868]" httpUri="https://zenodo.org/record/10780027/files/figure.png" pageId="12" pageNumber="540" startId="12.[151,264,1843,1867]" targetBox="[151,1434,181,1819]" targetPageId="12" targetType="figure">
<paragraph id="400336B6FFED043A59C1FB27FBBAF8B9" blockId="12.[151,1092,1843,1868]" box="[151,1092,1843,1868]" pageId="12" pageNumber="540">
<emphasis id="72C8EAA4FFED043A59C1FB27FEAAF8B9" bold="true" box="[151,340,1843,1867]" pageId="12" pageNumber="540">FIGURES 53–54.</emphasis>
<taxonomicName id="87BC4D35FFED043A580CFB27FD53F8B9" box="[346,685,1843,1868]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="540" phylum="Arthropoda" rank="species" species="klugei" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFED043A580CFB27FE2FF8B9" box="[346,465,1843,1867]" italics="true" pageId="12" pageNumber="540">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFED043A58B6FB27FD9FF8B9" box="[480,609,1843,1867]" italics="true" pageId="12" pageNumber="540">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFED043A5B39FB27FD53F8B9" box="[623,685,1843,1867]" italics="true" pageId="12" pageNumber="540">klugei</emphasis>
</taxonomicName>
, larval exuviae. 53, labrum; 54, labium.
</paragraph>
</caption>
<paragraph id="400336B6FFED043B5991FB6EFA66FEEF" blockId="12.[151,1436,1914,2012]" lastBlockId="13.[151,1437,151,285]" lastPageId="13" lastPageNumber="541" pageId="12" pageNumber="540">
The original description and illustrations of 
<taxonomicName id="87BC4D35FFED043A5BF5FB6EFC2BF866" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[675,981,1914,1940]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="540" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFED043A5BF5FB6EFC2BF866" box="[675,981,1914,1940]" italics="true" pageId="12" pageNumber="540">Nigrobaetis paramakalyani</emphasis>
</taxonomicName>
contain a mixture of characters belonging to both species. The photos of female larva (
<bibRefCitation id="242D4B47FFED043A5BC2FB8AFC69F84B" author="Kubendran, T. &amp; Balasubramanian, C. &amp; Selvakumar, C. &amp; Gattolliat, J. L. &amp; Sivaramakrishnan, K. G." box="[660,919,1950,1977]" pageId="12" pageNumber="540" pagination="188 - 200" refId="ref7026" refString="Kubendran, T., Balasubramanian, C., Selvakumar, C., Gattolliat, J. L. &amp; Sivaramakrishnan, K. G. (2015) Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India). Zootaxa, 3957 (2), 188 - 200. https: // doi. org / 10.11646 / zootaxa. 3957.2.3" type="journal article" year="2015">
Kubendran 
<emphasis id="72C8EAA4FFED043A5A4FFB8BFCB2F84A" box="[793,844,1950,1976]" italics="true" pageId="12" pageNumber="540">et al</emphasis>
. 2015
</bibRefCitation>
: figs 22–23) belong to 
<taxonomicName id="87BC4D35FFED043A5DF6FB8BFA92F84A" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[1184,1388,1950,1976]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="540" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFED043A5DF6FB8BFA92F84A" box="[1184,1388,1950,1976]" italics="true" pageId="12" pageNumber="540">N. paramakalyani</emphasis>
</taxonomicName>
and demonstrate characteristic color pattern of thoracic and abdominal terga. In contrast to this, the drawing of paraproct which bears a few long spines (ibid., fig. 36) belongs to 
<taxonomicName id="87BC4D35FFEC043B5A45FC83FC87FF43" box="[787,889,151,177]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="klugei">
<emphasis id="72C8EAA4FFEC043B5A45FC83FC87FF43" box="[787,889,151,177]" italics="true" pageId="13" pageNumber="541">N. klugei</emphasis>
</taxonomicName>
, while true paraproct of 
<taxonomicName id="87BC4D35FFEC043B5DC7FC83FAA2FF43" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[1169,1372,151,177]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEC043B5DC7FC83FAA2FF43" box="[1169,1372,151,177]" italics="true" pageId="13" pageNumber="541">N. paramakalyani</emphasis>
</taxonomicName>
has a higher number of small spines (
<bibRefCitation id="242D4B47FFEC043B58A6FCAFFCEDFF27" author="Sivaruban, T. &amp; Srinivasan, P. &amp; Barathy, S. &amp; Isack, R." box="[496,787,187,213]" pageId="13" pageNumber="541" pagination="182 - 190" refId="ref7411" refString="Sivaruban, T., Srinivasan, P., Barathy, S. &amp; Isack, R. (2022) A new species of Nigrobaetis Novikova &amp; Kluge, 1987 (Ephemeroptera, Baetidae) from Tamil Nadu, India. Zootaxa, 5091 (1), 182 - 190. https: // doi. org / 10.11646 / zootaxa. 5091.1.8" type="journal article" year="2022">
Sivaruban 
<emphasis id="72C8EAA4FFEC043B5B3EFCAFFD66FF27" box="[616,664,187,213]" italics="true" pageId="13" pageNumber="541">et al</emphasis>
. 2022: 189
</bibRefCitation>
and fig. 4D). The original description of 
<taxonomicName id="87BC4D35FFEC043B5D82FCAFFA63FF27" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[1236,1437,187,213]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEC043B5D82FCAFFA63FF27" box="[1236,1437,187,213]" italics="true" pageId="13" pageNumber="541">N. paramakalyani</emphasis>
</taxonomicName>
states that «Hind wing pads present» and contains the drawing of hind protopteron (
<bibRefCitation id="242D4B47FFEC043B5D6AFCCBFA93FF0B" author="Kubendran, T. &amp; Balasubramanian, C. &amp; Selvakumar, C. &amp; Gattolliat, J. L. &amp; Sivaramakrishnan, K. G." box="[1084,1389,222,249]" pageId="13" pageNumber="541" pagination="188 - 200" refId="ref7026" refString="Kubendran, T., Balasubramanian, C., Selvakumar, C., Gattolliat, J. L. &amp; Sivaramakrishnan, K. G. (2015) Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India). Zootaxa, 3957 (2), 188 - 200. https: // doi. org / 10.11646 / zootaxa. 3957.2.3" type="journal article" year="2015">
Kubendran 
<emphasis id="72C8EAA4FFEC043B5DE9FCCBFB0EFF0B" box="[1215,1264,223,249]" italics="true" pageId="13" pageNumber="541">et al</emphasis>
. 2015: 194
</bibRefCitation>
and fig. 33). However, the 
<typeStatus id="9F078814FFEC043B58C2FD17FE08FEEF" box="[404,502,259,285]" pageId="13" pageNumber="541" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="87BC4D35FFEC043B5B4AFD17FD18FEEF" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[540,742,259,285]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEC043B5B4AFD17FD18FEEF" box="[540,742,259,285]" italics="true" pageId="13" pageNumber="541">N. paramakalyani</emphasis>
</taxonomicName>
has no hind protoptera, and the drawing belongs to 
<taxonomicName id="87BC4D35FFEC043B5C7AFD17FA6FFEEF" box="[1324,1425,259,285]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="klugei">
<emphasis id="72C8EAA4FFEC043B5C7AFD17FA6FFEEF" box="[1324,1425,259,285]" italics="true" pageId="13" pageNumber="541">N. klugei</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="14C3663EFFEC043B59C1FB68FF43F84A" ID-DOI="http://doi.org/10.5281/zenodo.10780029" ID-Zenodo-Dep="10780029" httpUri="https://zenodo.org/record/10780029/files/figure.png" pageId="13" pageNumber="541" startId="13.[151,264,1916,1940]" targetBox="[151,1436,309,1893]" targetPageId="13" targetType="figure">
<paragraph id="400336B6FFEC043B59C1FB68FF43F84A" blockId="13.[151,1436,1916,1976]" pageId="13" pageNumber="541">
<emphasis id="72C8EAA4FFEC043B59C1FB68FEA8F866" bold="true" box="[151,342,1916,1940]" pageId="13" pageNumber="541">FIGURES 55–57.</emphasis>
<taxonomicName id="87BC4D35FFEC043B580BFB68FD4CF866" box="[349,690,1916,1940]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="541" phylum="Arthropoda" rank="species" species="klugei" subGenus="Margobaetis">
<emphasis id="72C8EAA4FFEC043B580BFB68FE2AF866" box="[349,468,1916,1940]" italics="true" pageId="13" pageNumber="541">Nigrobaetis</emphasis>
(
<emphasis id="72C8EAA4FFEC043B58B2FB68FD9BF866" box="[484,613,1916,1940]" italics="true" pageId="13" pageNumber="541">Margobaetis</emphasis>
) 
<emphasis id="72C8EAA4FFEC043B5B22FB68FD4CF866" box="[628,690,1916,1940]" italics="true" pageId="13" pageNumber="541">klugei</emphasis>
</taxonomicName>
, larval exuviae of tibia and tarsus. 55, fore leg; 56, middle leg; 57, hind leg.
</paragraph>
</caption>
<paragraph id="400336B6FFEF04385991FC82FAA4FEEF" blockId="14.[151,1437,150,717]" pageId="14" pageNumber="542">
<emphasis id="72C8EAA4FFEF04385991FC82FDEEFF43" bold="true" box="[199,528,150,177]" pageId="14" pageNumber="542">
Synonymy of 
<taxonomicName id="87BC4D35FFEF04385838FC83FDF7FF43" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[366,521,150,177]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385838FC83FDF7FF43" bold="true" box="[366,521,150,177]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="87BC4D35FFEF04385B4FFC83FB8FFF43" authority="Kaltenbach &amp; Gattolliat 2023" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[537,1137,151,177]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385B4FFC83FCE6FF43" box="[537,792,151,177]" italics="true" pageId="14" pageNumber="542">Nigrobaetis sumbensis</emphasis>
<bibRefCitation id="242D4B47FFEF04385A76FC83FB8FFF43" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[800,1137,151,177]" pageId="14" pageNumber="542" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
</taxonomicName>
was described based on 
<specimenCount id="56BAFD3FFFEF04385CD8FC83FF25FF27" count="3" pageId="14" pageNumber="542" type="larva">3 larvae</specimenCount>
from Sumba Islan in 
<collectingCountry id="38AB7626FFEF04385885FCAFFDBAFF27" box="[467,580,187,213]" name="Indonesia" pageId="14" pageNumber="542">Indonesia</collectingCountry>
. This species was compared with other 
<taxonomicName id="87BC4D35FFEF04385D50FCAFFB1CFF27" box="[1030,1250,187,213]" pageId="14" pageNumber="542">
<emphasis id="72C8EAA4FFEF04385D50FCAFFB77FF27" box="[1030,1161,187,213]" italics="true" pageId="14" pageNumber="542">Nigrobaetis</emphasis>
species
</taxonomicName>
from 
<collectingCountry id="38AB7626FFEF04385C7EFCAFFA66FF27" box="[1320,1432,187,213]" name="Indonesia" pageId="14" pageNumber="542">Indonesia</collectingCountry>
, 
<collectingCountry id="38AB7626FFEF043859C1FCCBFEEBFF0B" box="[151,277,223,249]" name="Philippines" pageId="14" pageNumber="542">Philippines</collectingCountry>
and 
<collectingCountry id="38AB7626FFEF0438580EFCCBFE3FFF0B" box="[344,449,223,249]" name="Malaysia" pageId="14" pageNumber="542">Malaysia</collectingCountry>
, but not with 
<taxonomicName id="87BC4D35FFEF04385B25FCCBFCBAFF0B" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[627,836,223,249]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEF04385B25FCCBFCBAFF0B" box="[627,836,223,249]" italics="true" pageId="14" pageNumber="542">N. paramakalyani</emphasis>
</taxonomicName>
from 
<collectingCountry id="38AB7626FFEF04385AC2FCCBFC2FFF0B" box="[916,977,223,249]" name="India" pageId="14" pageNumber="542">India</collectingCountry>
. Comparison of 
<taxonomicName id="87BC4D35FFEF04385DCBFCCBFAC8FF0B" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[1181,1334,223,249]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385DCBFCCBFAC8FF0B" box="[1181,1334,223,249]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
with 
<taxonomicName id="87BC4D35FFEF04385CD5FCCBFEBFFEEF" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEF04385CD5FCCBFEBFFEEF" italics="true" pageId="14" pageNumber="542">N. paramakalyani</emphasis>
</taxonomicName>
was impossible, because 
<taxonomicName id="87BC4D35FFEF04385B37FD17FCD2FEEF" authorityName="Kubendran &amp; Balasubramanian" authorityYear="2015" box="[609,812,259,285]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="paramakalyani">
<emphasis id="72C8EAA4FFEF04385B37FD17FCD2FEEF" box="[609,812,259,285]" italics="true" pageId="14" pageNumber="542">N. paramakalyani</emphasis>
</taxonomicName>
was wrongly characterized as having hind wings.
</paragraph>
<paragraph id="400336B6FFEF04385991FD33FC1AFE07" blockId="14.[151,1437,150,717]" pageId="14" pageNumber="542">
Comparison of 
<taxonomicName id="87BC4D35FFEF04385821FD33FDF2FEB3" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[375,524,295,321]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385821FD33FDF2FEB3" box="[375,524,295,321]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
with 
<taxonomicName id="87BC4D35FFEF04385B18FD33FD36FEB2" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[590,712,295,320]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEF04385B18FD33FD36FEB2" box="[590,712,295,320]" italics="true" pageId="14" pageNumber="542">N. minutus</emphasis>
</taxonomicName>
was given in the key (
<bibRefCitation id="242D4B47FFEF04385A9DFD33FA6EFEB3" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[971,1424,295,321]" pageId="14" pageNumber="542" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023: 228–229</bibRefCitation>
), according to which 
<taxonomicName id="87BC4D35FFEF04385825FD5FFDF8FE97" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[371,518,331,357]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385825FD5FFDF8FE97" box="[371,518,331,357]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="87BC4D35FFEF04385BCEFD5FFCEFFE96" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[664,785,331,356]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEF04385BCEFD5FFCEFFE96" box="[664,785,331,356]" italics="true" pageId="14" pageNumber="542">N. minutus</emphasis>
</taxonomicName>
and other species by «Fore femur very slender ..., dorsally slightly concave». Actually, the leg described and figures as «foreleg» is actually middle or hind leg, that is testified by the 4-segmented subimaginal tarsus developing inside (
<bibRefCitation id="242D4B47FFEF04385A72FD87FB8CFE5F" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[804,1138,403,429]" pageId="14" pageNumber="542" pagination="175 - 234" refId="ref6534" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2023) New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae). ZooKeys, 1166, 175 - 234. https: // doi. org / 10.3897 / zookeys. 1166.102941" type="journal article" year="2023">Kaltenbach &amp; Gattolliat 2023</bibRefCitation>
: fig. 19a). As well in many other species, in 
<taxonomicName id="87BC4D35FFEF0438580FFDA3FE2AFE22" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[345,468,439,464]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEF0438580FFDA3FE2AFE22" box="[345,468,439,464]" italics="true" pageId="14" pageNumber="542">N. minutus</emphasis>
</taxonomicName>
femur of the foreleg is thicker, with straight outer margin (
<figureCitation id="D8872A33FFEF04385D2AFDA3FB1AFE23" box="[1148,1252,439,465]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="14" pageNumber="542">Figs 2, 9</figureCitation>
), and femora of middle and hind legs are thinner, with concave outer margin (
<figureCitation id="D8872A33FFEF04385A11FDCFFC29FE07" box="[839,983,475,501]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1942,1966]" captionTargetBox="[151,1436,346,1919]" captionTargetId="figure-128@2.[151,1436,345,1919]" captionTargetPageId="2" captionText="FIGURES 1–12. Nigrobaetis (Margobaetis) minutus, larvae. 1–6, specimen from Madurai; 7–8, specimen from Erumeli; 9–12, specimen from Pinrang (Sulawesi). 1, exuviae of head; 2–4, exuviae of fore, middle and hind legs; 5, 7, 11, exuviae of abdomen; 6, 8, 12, exuviae of pronotum and mesonotum; 9, 10, fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.10779997" httpUri="https://zenodo.org/record/10779997/files/figure.png" pageId="14" pageNumber="542">Figs 3–4, 10</figureCitation>
).
</paragraph>
<paragraph id="400336B6FFEF04385991FDEBFAB4FD93" blockId="14.[151,1437,150,717]" pageId="14" pageNumber="542">
Another characters of 
<taxonomicName id="87BC4D35FFEF043858E9FDEBFDACFDEB" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[447,594,511,537]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF043858E9FDEBFDACFDEB" box="[447,594,511,537]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
reported in this key, is the number of denticles on claws; variability of their number has not been reported for 
<taxonomicName id="87BC4D35FFEF04385B43FE37FD71FDCE" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[533,655,547,572]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEF04385B43FE37FD71FDCE" box="[533,655,547,572]" italics="true" pageId="14" pageNumber="542">N. minutus</emphasis>
</taxonomicName>
. The third reported character is a tuft of setae-like processes between prostheca and mola of left mandible; the same tuft is figured for 
<taxonomicName id="87BC4D35FFEF04385A3CFE53FC1AFD92" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1984" box="[874,996,583,608]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="72C8EAA4FFEF04385A3CFE53FC1AFD92" box="[874,996,583,608]" italics="true" pageId="14" pageNumber="542">N. minutus</emphasis>
</taxonomicName>
(
<bibRefCitation id="242D4B47FFEF04385AA4FE53FB0AFD93" author="Muller-Liebenau, I." box="[1010,1268,583,609]" pageId="14" pageNumber="542" pagination="253 - 284" refId="ref7277" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253 - 284." type="journal article" year="1984">Müller-Liebenau 1984</bibRefCitation>
: fig. 2).
</paragraph>
<paragraph id="400336B6FFEF0438599BFE7FFEE1FD3F" blockId="14.[151,1437,150,717]" pageId="14" pageNumber="542">
Comparison of the larval description of 
<taxonomicName id="87BC4D35FFEF04385BD3FE7FFCE9FD77" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2023" box="[645,791,619,645]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="542" phylum="Arthropoda" rank="species" species="sumbensis">
<emphasis id="72C8EAA4FFEF04385BD3FE7FFCE9FD77" box="[645,791,619,645]" italics="true" pageId="14" pageNumber="542">N. sumbensis</emphasis>
</taxonomicName>
with our material from 
<collectingCountry id="38AB7626FFEF04385D4BFE7FFBA9FD77" box="[1053,1111,619,645]" name="India" pageId="14" pageNumber="542">India</collectingCountry>
and Sulawesi does not reveal differences. Comparison of egg structure of specimens from 
<collectingCountry id="38AB7626FFEF04385A1EFE9BFC7CFD5B" box="[840,898,655,681]" name="India" pageId="14" pageNumber="542">India</collectingCountry>
and Sulawesi confirms the wide distribution of this species.
</paragraph>
</subSubSection>
</treatment>
</document>