278 lines
36 KiB
XML
278 lines
36 KiB
XML
<document id="E473272371A7D69514DD4382D58015FD" ID-CLB-Dataset="28026" ID-DOI="10.11646/zootaxa.4512.1.1" ID-GBIF-Dataset="26e12e1b-2175-4007-871b-d0b73122e5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607575" ID-ZooBank="E752FB7B-F34C-4D12-B8A2-EA6C791DD6C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586644813" checkinUser="plazi" docAuthor="Kaiser, Christine M., Kaiser, Hinrich & O’Shea, Mark" docDate="2018" docId="C80EBE29FFEBFFDBFF75FED00D2FFC28" docLanguage="en" docName="zootaxa.4512.1.1.pdf" docOrigin="Zootaxa 4512 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stegonotus diehli Lindholm 1905" docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="3437C651FFC7FFF6FFE2FE470F12FFFE" masterDocTitle="The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="45" updateTime="1698713152511" updateUser="ExternalLinkService">
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<mods:title id="546232AC11DDD8264541BE2A56AFB61B">The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)</mods:title>
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<mods:namePart id="9B8541D00EC5C63DF4E64FD8701A2CC2">Kaiser, Christine M.</mods:namePart>
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<mods:namePart id="187B3187AA2FF0C45A1537CD24BDCE96">Kaiser, Hinrich</mods:namePart>
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<mods:namePart id="D512F8205D09B526EF5A5D787F6F5894">O’Shea, Mark</mods:namePart>
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<mods:title id="4E3CD5E4D8F6229DF21453644B183D75">Zootaxa</mods:title>
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<mods:date id="A9711F0AEC345736776CA32F960FD073">2018</mods:date>
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<treatment id="C80EBE29FFEBFFDBFF75FED00D2FFC28" ID-DOI="http://doi.org/10.5281/zenodo.5997409" ID-GBIF-Taxon="154496320" ID-Zenodo-Dep="5997409" LSID="urn:lsid:plazi:treatment:C80EBE29FFEBFFDBFF75FED00D2FFC28" httpUri="http://treatment.plazi.org/id/C80EBE29FFEBFFDBFF75FED00D2FFC28" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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<subSubSection id="08BD5CB4FFEBFFDAFF75FED00EAAFF09" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="40180F3FFFEBFFDAFF75FED00D78FF4C" blockId="44.[151,618,151,178]" box="[151,618,151,178]" pageId="44" pageNumber="45">
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<heading id="1B50B853FFEBFFDAFF75FED00D78FF4C" bold="true" box="[151,618,151,178]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
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<treatmentCitationGroup id="60B72811FFEBFFDAFF75FED00D78FF4C" box="[151,618,151,178]" pageId="44" pageNumber="45">
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<taxonomicName id="87A774BCFFEBFFDAFF75FED00D78FF4C" authority="Lindholm, 1905: 236" authorityName="Lindholm" authorityPageNumber="236" authorityYear="1905" box="[151,618,151,178]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="diehli">
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<emphasis id="72D3D32DFFEBFFDAFF75FED00D78FF4C" bold="true" box="[151,618,151,178]" pageId="44" pageNumber="45">
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<emphasis id="72D3D32DFFEBFFDAFF75FED00E75FF4F" bold="true" box="[151,359,151,177]" italics="true" pageId="44" pageNumber="45">Stegonotus diehli</emphasis>
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<treatmentCitation id="C106292EFFEBFFDAFE8CFEDF0D78FF4C" author="Lindholm, W. A." box="[366,618,152,178]" page="236" pageId="44" pageNumber="45" year="1905">
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<bibRefCitation id="243672CEFFEBFFDAFE8CFEDF0D27FF4C" author="Lindholm, W. A." box="[366,565,152,178]" pageId="44" pageNumber="45" pagination="227 - 240" refId="ref44980" refString="Lindholm, W. A. (1905) Uber einige Eidechsen und Schlangen aus Deutsch-Neuguinea. Jahrbucher des nassauischen Fereins fur Naturkunde, 58, 227 - 240." type="journal article" year="1905">Lindholm, 1905</bibRefCitation>
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:236
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</treatmentCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="40180F3FFFEBFFDAFF75FE990EAAFF09" blockId="44.[151,1437,222,862]" box="[151,440,222,247]" pageId="44" pageNumber="45">
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<emphasis id="72D3D32DFFEBFFDAFF75FE990E7CFF09" bold="true" box="[151,366,222,247]" pageId="44" pageNumber="45">Taxonomic status.</emphasis>
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Valid.
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</paragraph>
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</subSubSection>
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<subSubSection id="08BD5CB4FFEBFFDAFF25FF450D45FD54" pageId="44" pageNumber="45" type="description">
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<paragraph id="40180F3FFFEBFFDAFF25FF450DC2FEE5" blockId="44.[151,1437,222,862]" box="[199,720,258,283]" pageId="44" pageNumber="45">
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<emphasis id="72D3D32DFFEBFFDAFF25FF450E5BFEE5" bold="true" box="[199,329,258,283]" pageId="44" pageNumber="45">Synonyms.</emphasis>
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<taxonomicName id="87A774BCFFEBFFDAFEB2FF440DDEFEE5" authority="Werner, 1924" authorityName="Werner" authorityYear="1924" box="[336,716,258,283]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="dorsalis">
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<emphasis id="72D3D32DFFEBFFDAFEB2FF440D39FEE5" box="[336,555,258,283]" italics="true" pageId="44" pageNumber="45">Stegonotus dorsalis</emphasis>
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<bibRefCitation id="243672CEFFEBFFDAFDD0FF440DDEFEE5" author="Werner, F." box="[562,716,259,283]" pageId="44" pageNumber="45" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werner, 1924</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="40180F3FFFEBFFDAFF25FF610D45FD54" blockId="44.[151,1437,222,862]" pageId="44" pageNumber="45">
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<emphasis id="72D3D32DFFEBFFDAFF25FF610E93FEC1" bold="true" box="[199,385,294,319]" pageId="44" pageNumber="45">Original name.</emphasis>
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<taxonomicName id="87A774BCFFEBFFDAFE6FFF600C5DFEC1" authority="Lindholm, 1905: 236" authorityName="Lindholm" authorityPageNumber="236" authorityYear="1905" box="[397,847,294,319]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="diehli">
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<emphasis id="72D3D32DFFEBFFDAFE6FFF600D46FEC1" box="[397,596,294,319]" italics="true" pageId="44" pageNumber="45">Stegonotus diehli</emphasis>
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<bibRefCitation id="243672CEFFEBFFDAFD82FF610C0FFEC1" author="Lindholm, W. A." box="[608,797,294,319]" pageId="44" pageNumber="45" pagination="227 - 240" refId="ref44980" refString="Lindholm, W. A. (1905) Uber einige Eidechsen und Schlangen aus Deutsch-Neuguinea. Jahrbucher des nassauischen Fereins fur Naturkunde, 58, 227 - 240." type="journal article" year="1905">Lindholm, 1905</bibRefCitation>
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:236
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</taxonomicName>
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. Wilhelm Adolfovich Lindholm (
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<date id="341929FFFFEBFFDAFB3EFF610A4DFEC1" box="[1244,1375,294,319]" pageId="44" pageNumber="45" value="1874" valueMax="1935">1874–1935</date>
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;
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||
<figureCitation id="D89C13BAFFEBFFDAFA92FF610FDAFE9D" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (1840–1913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (1843–1920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (1820–1891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (1846–1926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (1858–1937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (1835–1917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (1871–1943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (1874–1935), author of the species name diehli. (I) Nelly de Rooij (1883–1964), author of the species name florensis. (J) Franz Werner (1867–1939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (1870–1921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (1902–1990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="44" pageNumber="45">Fig. 17H</figureCitation>
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) was a Russian malacologist and herpetologist (
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<bibRefCitation id="243672CEFFEBFFDAFD07FF0D0C78FE9D" author="Adler, K." box="[741,874,330,355]" pageId="44" pageNumber="45" refId="ref41666" refString="Adler, K. (2007) Contributions to the History of Herpetology. Folume 2. Society for the Study of Reptiles and Amphibians, Lawrence, Kansas, USA, 400 pp." type="book" year="2007">Adler 2007</bibRefCitation>
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), who worked as a scientific adviser at the natural history museum in Wiesbaden,
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<collectingCountry id="38B04FAFFFEBFFDAFE18FF280D70FE79" box="[506,610,367,391]" name="Germany" pageId="44" pageNumber="45">Germany</collectingCountry>
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from
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<date id="341929FFFFEBFFDAFD4AFF280C38FE79" box="[680,810,367,391]" pageId="44" pageNumber="45" value="1900" valueMax="1907">1900–1907</date>
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. He then became a member of the curatorial staff at the Zoological Museum of Moscow Imperial University (now Moscow State University) and in 1914 took the position of Curator of Mollusks at the Zoological Museum of the Russian Academy of Sciences in St. Petersburg (later Petrograd, then Leningrad). Lindholm named this species in honor of the German Protestant missionary Wilhelm Diehl (
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<date id="341929FFFFEBFFDAFF0BFFB80E7EFDE9" box="[233,364,511,535]" pageId="44" pageNumber="45" value="1874" valueMax="1940">1874–1940</date>
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), who spent 18 years in German New
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<collectingCountry id="38B04FAFFFEBFFDAFCA2FFB90C86FDE9" box="[832,916,510,535]" name="Guinea-Bissau" pageId="44" pageNumber="45">Guinea</collectingCountry>
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, then known as
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<collectingCountry id="38B04FAFFFEBFFDAFBB9FFB90A5BFDE9" box="[1115,1353,510,535]" name="Papua New Guinea" pageId="44" pageNumber="45">Kaiser-Wilhelmsland</collectingCountry>
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(
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<bibRefCitation id="243672CEFFEBFFDAFABEFFB90FCCFDD1" author="Diehl, W." pageId="44" pageNumber="45" pagination="1906 - 1913" refId="ref43088" refString="Diehl, W. (2015) Pioniermissionar in Kaiser-Wilhelmsland: Wilhelm Diehl berichtet aus Deutsch-Neuguinea. 1906 - 1913. In: Klein, D. (Ed.), Quellen und Forschungen zur Sudsee. Volume 4. Harrassowitz Verlag, Leipzig, Germany, 192 pp." type="book chapter" year="2015">
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Diehl 2015
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<superScript id="B7D2A277FFEBFFDAFF2DFC660FCCFDD1" attach="none" box="[207,222,545,559]" fontSize="6" pageId="44" pageNumber="45">18</superScript>
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</bibRefCitation>
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) and who, in 1905, sent a small herpetological collection to Eduard Lampe (
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<date id="341929FFFFEBFFDAFB9FFC610A12FDC0" box="[1149,1280,550,574]" pageId="44" pageNumber="45" value="1871" valueMax="1919">1871–1919</date>
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), at the time museum curator in Wiesbaden (
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<bibRefCitation id="243672CEFFEBFFDAFDE2FC0E0DA7FD9C" author="Lindholm, W. A." box="[512,693,585,610]" pageId="44" pageNumber="45" pagination="227 - 240" refId="ref44980" refString="Lindholm, W. A. (1905) Uber einige Eidechsen und Schlangen aus Deutsch-Neuguinea. Jahrbucher des nassauischen Fereins fur Naturkunde, 58, 227 - 240." type="journal article" year="1905">Lindholm 1905</bibRefCitation>
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). Lampe asked Lindholm to identify and describe the specimens. The species description was presented in German. The photograph of Lindholm (
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<figureCitation id="D89C13BAFFEBFFDAFBC4FC2A0B9CFD78" box="[1062,1166,621,646]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (1840–1913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (1843–1920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (1820–1891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (1846–1926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (1858–1937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (1835–1917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (1871–1943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (1874–1935), author of the species name diehli. (I) Nelly de Rooij (1883–1964), author of the species name florensis. (J) Franz Werner (1867–1939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (1870–1921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (1902–1990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="44" pageNumber="45">Fig. 17H</figureCitation>
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; public domain) is from the obituary published by
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<bibRefCitation id="243672CEFFEBFFDAFE59FCD50D41FD54" author="Hesse, P." box="[443,595,657,682]" pageId="44" pageNumber="45" pagination="116 - 117" refId="ref44441" refString="Hesse, P. (1936) W. A. Lindhom †. Archiv fur Molluskenkunde, 68 (3), 116 - 117." type="journal article" year="1936">Hesse (1936)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="08BD5CB4FFEBFFDBFF25FCF20BCDFF06" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="materials_examined">
|
||
<paragraph id="40180F3FFFEBFFDAFF25FCF20CF1FD30" blockId="44.[151,1437,222,862]" box="[199,995,693,718]" pageId="44" pageNumber="45">
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||
<materialsCitation id="F0CF0562FFEBFFDAFF25FCF20CF1FD30" ID-GBIF-Occurrence="2028140073" box="[199,995,693,718]" collectionCode="MWNH" pageId="44" pageNumber="45" specimenCode="MWNH 1244" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="72D3D32DFFEBFFDAFF25FCF20E29FD30" bold="true" box="[199,315,693,718]" pageId="44" pageNumber="45">
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||
<typeStatus id="9F1CB19DFFEBFFDAFF25FCF20E24FD30" box="[199,310,693,718]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
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||
.
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</emphasis>
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<specimenCode id="1001A744FFEBFFDAFEA0FCF10EF2FD30" box="[322,480,694,718]" collectionCode="MWNH" country="Germany" httpUri="http://grbio.org/cool/zw2m-hsji" name="Museum Wiesbaden, Department of Natural Science" pageId="44" pageNumber="45">MWNH 1244</specimenCode>
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, a juvenile of unknown sex (
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<figureCitation id="D89C13BAFFEBFFDAFCC3FCF20C67FD30" box="[801,885,693,718]" captionStart="FIGURE 25" captionStartId="44.[151,250,1800,1822]" captionTargetBox="[151,1436,912,1768]" captionTargetId="figure@44.[151,1436,912,1768]" captionTargetPageId="44" captionText="FIGURE 25. Holotype of Stegonotus diehli Lindholm, 1905, MWNH 1244, a juvenile from Bogadjim, Madang Province, Papua New Guinea. Shown are the entire specimen (TTL = 252 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views." httpUri="https://zenodo.org/record/2607629/files/figure.png" pageId="44" pageNumber="45">Fig. 25</figureCitation>
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;
|
||
<tableCitation id="0D253A84FFEBFFDAFC62FCF20CC0FD30" box="[896,978,693,718]" captionStart="TABLE 1" captionStartId="60.[147,229,603,624]" captionText="TABLE 1. Name-bearing types of species in the genus Stegonotus. In this list we present basic data, including common measurements and selected scale counts, for all namebearing types (i.e., holotypes—HT, lectotypes—LT, neotypes—NT, syntypes—ST) to provide an easy overview of species. Additional details and the information for paratypes and paralectotypes can be found in the text. Character abbreviations are explained in the main text. For specimens where sex is indicated in red, we used the SCR value and not direct observation to estimate sex. Missing values are indicated by O. Specimen Type SCR SubC Species Synonym Sex SVL TL TTL V SC Dorsals SCE SL IL ILG Number Status (%) Condition LSUMZ admiraltiensis HT M 658 185 843 208 98 32 paired 17-19-15 4+5 8 10 10 93958 MSNG aruensis HT M 582 233 815 190 103 35 paired 17-17-15 3+4 7 8 4 30186 BMNH australis HT M 503 135 638 205 77 27 paired 17-17-15 4+5 8 9 5 1946.1.14.93 AMS australis darnleyensis HT F 720 O O 198 53+ O paired 17-17-15 4+5 9 10 5 R31894 AMS australis plumbeus HT M 1025 250 1275 220 74 25 paired 17-17-15 3+4+5 9 10 5 R31895 BMNH batjanensis HT M 1239 321 1560 235 86 27 paired 17-17-15 3+4+5 8 10 5 1946.1.11.36 FMNH borneensis HT F 783 221 1004 195 79 29 paired 17-17-15 4+5 10 11 5 164746 MNHN cucullatus HT M 769 254 1023 209 91 30 paired 17-17-15 4+5 8 9 5 3412 MTKD a cucullatus magnus ST F O O O 207 74 26 paired O-19-O 4+5 8 O O 573 ZMB b cucullatus magnus ST M 975 265 1240 211 87 29 paired 19-19-17 3+4 8 10 5 8794 MZB derooijae HT M 455 193 648 181 94 34 paired 17-17-15 3+4 7 8 8 Ophi.3288 MWNH diehli HT juv 196 56 252 179 82 31 paired 15-15-15 3+4 7 8 4 1244 NMW dorsalis HT M 805 245 1050 208 88 30 paired 15-15-15 4+5 8 10 5 14861 ZMA 9R florensis HT M 447 109 556 217 65 23 single 21-21-19 3+4+5 10 5 11080 8L BMNH guentheri LT F 683 165 848 192 64 25 paired 15-15-13 4+5 8 9 4 1946.1.11.38 ……continued on the next page" pageId="44" pageNumber="45">Table 1</tableCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="40180F3FFFEBFFDAFF25FC9E0A3CFCA0" blockId="44.[151,1437,222,862]" pageId="44" pageNumber="45">
|
||
<emphasis id="72D3D32DFFEBFFDAFF25FC9E0E75FD0C" bold="true" box="[199,359,729,754]" pageId="44" pageNumber="45">Type locality.</emphasis>
|
||
“Bogadjim an der Astrolabe Bai in Deutsch-Neuguinea” [Bogadjim,
|
||
<collectingRegion id="8263C1DDFFEBFFDAFB62FC9E0A59FD0C" box="[1152,1355,729,754]" country="Papua New Guinea" name="Madang" pageId="44" pageNumber="45">Madang Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="38B04FAFFFEBFFDAFABAFC9D0E3AFCE8" name="Papua New Guinea" pageId="44" pageNumber="45">Papua New Guinea</collectingCountry>
|
||
, ca.
|
||
<geoCoordinate id="259369F8FFEBFFDAFEBEFCBA0EB9FCE8" box="[348,427,765,790]" direction="south" orientation="latitude" pageId="44" pageNumber="45" precision="555" value="-5.45">5.45°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="259369F8FFEBFFDAFE55FCBA0D34FCE8" box="[439,550,765,790]" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="555" value="145.75">145.75°E</geoCoordinate>
|
||
]. Bogadjim was a native New Guinean settlement, where a Protestant mission was founded in 1887. Adjacent to the village, the German colonial government subsequently built a settlement called Stephansort beginning in 1889, which disappeared again at the end of the German colonial period.
|
||
</paragraph>
|
||
<caption id="14D85FB7FFEBFFDAFF75F94F0D7FF8A2" httpUri="https://zenodo.org/record/2607629/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1800,1822]" targetBox="[151,1436,912,1768]" targetPageId="44">
|
||
<paragraph id="40180F3FFFEBFFDAFF75F94F0D7FF8A2" blockId="44.[151,1436,1800,1884]" pageId="44" pageNumber="45">
|
||
<emphasis id="72D3D32DFFEBFFDAFF75F94F0E30F8E3" bold="true" box="[151,290,1800,1822]" pageId="44" pageNumber="45">FIGURE 25.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="87A774BCFFEBFFDAFE50F94E0C07F8E0" authority="Lindholm, 1905" authorityName="Lindholm" authorityYear="1905" box="[434,789,1800,1822]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEBFFDAFE50F94E0D76F8E0" box="[434,612,1800,1822]" italics="true" pageId="44" pageNumber="45">Stegonotus diehli</emphasis>
|
||
Lindholm, 1905
|
||
</taxonomicName>
|
||
, MWNH 1244, a juvenile from Bogadjim, Madang Province, Papua New Guinea. Shown are the entire specimen (TTL = 252 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="40180F3FFFEBFFDAFF75F9FB0A58F80F" blockId="44.[151,1436,1980,2033]" pageId="44" pageNumber="45">18. This citation is obviously for a posthumous publication, Wilhelm Diehl having died in 1940. However, Diehl kept meticulous diaries, which had been unpublished and were collated and made available for the first time in this volume.</paragraph>
|
||
<paragraph id="40180F3FFFEAFFDBFF25FED00BCDFF06" blockId="45.[151,1436,151,982]" pageId="45" pageNumber="46">
|
||
<emphasis id="72D3D32DFFEAFFDBFF25FED00E55FF4E" bold="true" box="[199,327,151,176]" pageId="45" pageNumber="46">Collection.</emphasis>
|
||
According to the text preceding the formal description of
|
||
<taxonomicName id="87A774BCFFEAFFDBFBE7FEDF0B74FF4E" authorityName="Lindholm" authorityYear="1905" box="[1029,1126,151,176]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEAFFDBFBE7FEDF0B74FF4E" box="[1029,1126,151,176]" italics="true" pageId="45" pageNumber="46">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
, a shipment of specimens collected by Diehl was received in Wiesbaden in
|
||
<date id="341929FFFFEAFFDBFD28FEFB0C2EFF2B" box="[714,828,188,213]" pageId="45" pageNumber="46" value="1905-07">July 1905</date>
|
||
. There is no mention of prior shipments or collection dates, and it is therefore not possible to pinpoint the collection of the
|
||
<typeStatus id="9F1CB19DFFEAFFDBFC7DFE980B13FF06" box="[927,1025,223,248]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
with any precision.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08BD5CB4FFEAFFDBFF25FF430BF0FD4F" pageId="45" pageNumber="46" type="key">
|
||
<paragraph id="40180F3FFFEAFFDBFF25FF430A96FEBD" blockId="45.[151,1436,151,982]" pageId="45" pageNumber="46">
|
||
<emphasis id="72D3D32DFFEAFFDBFF25FF430D78FEE3" bold="true" box="[199,618,260,285]" pageId="45" pageNumber="46">
|
||
Key characteristics of the
|
||
<materialsCitation id="F0CF0562FFEAFFDBFE1EFF430D78FEE3" ID-GBIF-Occurrence="2028140053" box="[508,618,260,285]" collectionCode="SVL" pageId="45" pageNumber="46" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="9F1CB19DFFEAFFDBFE1EFF430D77FEE3" box="[508,613,260,285]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
.
|
||
</materialsCitation>
|
||
</emphasis>
|
||
O (196) mm SVL + O (56) mm TL = O (252) mm TTL. V
|
||
<subScript id="DC230D7AFFEAFFDBFAE6FF550A18FEDE" attach="left" box="[1284,1290,274,288]" fontSize="6" pageId="45" pageNumber="46">?</subScript>
|
||
= 175 (179), SC
|
||
<subScript id="DC230D7AFFEAFFDBFF5BFF7E0FADFEB9" attach="left" box="[185,191,313,327]" fontSize="6" pageId="45" pageNumber="46">?</subScript>
|
||
= 80 (82), SCR
|
||
<subScript id="DC230D7AFFEAFFDBFE90FF7E0E6AFEB9" attach="left" box="[370,376,313,327]" fontSize="6" pageId="45" pageNumber="46">?</subScript>
|
||
= 0.31 (0.31), D = O-15-O (15-15-15), SL
|
||
<subScript id="DC230D7AFFEAFFDBFC82FF7E0C7BFEB9" attach="left" box="[864,873,313,327]" fontSize="6" pageId="45" pageNumber="46">E</subScript>
|
||
= 3+4 (3+4), SL = 7 (7), IL = 8 (8), IL
|
||
<subScript id="DC230D7AFFEAFFDBFAFDFF7E0A38FEB9" attach="left" box="[1311,1322,313,327]" fontSize="6" pageId="45" pageNumber="46">G</subScript>
|
||
= 4 (4).
|
||
</paragraph>
|
||
<paragraph id="40180F3FFFEAFFDBFF25FF160BACFDE0" blockId="45.[151,1436,151,982]" pageId="45" pageNumber="46">
|
||
<emphasis id="72D3D32DFFEAFFDBFF25FF160D4FFE94" bold="true" box="[199,605,337,362]" pageId="45" pageNumber="46">Key characteristics of the species.</emphasis>
|
||
Along with the names
|
||
<taxonomicName id="87A774BCFFEAFFDBFC8BFF160CCEFE94" box="[873,988,337,362]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="cucullatus">
|
||
<emphasis id="72D3D32DFFEAFFDBFC8BFF160CCEFE94" box="[873,988,337,362]" italics="true" pageId="45" pageNumber="46">cucullatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="87A774BCFFEAFFDBFC0EFF160B46FE94" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1004,1108,337,362]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="72D3D32DFFEAFFDBFC0EFF160B46FE94" box="[1004,1108,337,362]" italics="true" pageId="45" pageNumber="46">modestus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="87A774BCFFEAFFDBFB76FF150BF2FE97" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[1172,1248,338,361]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="72D3D32DFFEAFFDBFB76FF150BF2FE97" box="[1172,1248,338,361]" italics="true" pageId="45" pageNumber="46">parvus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="87A774BCFFEAFFDBFB12FF160A22FE94" authorityName="Lindholm" authorityYear="1905" box="[1264,1328,337,362]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEAFFDBFB12FF160A22FE94" box="[1264,1328,337,362]" italics="true" pageId="45" pageNumber="46">diehli</emphasis>
|
||
</taxonomicName>
|
||
is one of four species names that have been applied to a large number of
|
||
<taxonomicName id="87A774BCFFEAFFDBFC8BFF310CF1FE73" authorityName="Guentheri Boulenger" authorityYear="1895" box="[873,995,374,397]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="72D3D32DFFEAFFDBFC8BFF310CF1FE73" box="[873,995,374,397]" italics="true" pageId="45" pageNumber="46">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
specimens in museum collections and, just like the other three, it has been applied across all regions of New
|
||
<collectingCountry id="38B04FAFFFEAFFDBFC45FFDE0CE9FE4C" box="[935,1019,409,434]" name="Guinea-Bissau" pageId="45" pageNumber="46">Guinea</collectingCountry>
|
||
. However, we do not believe that the name has been applied consistently or with the data from the
|
||
<typeStatus id="9F1CB19DFFEAFFDBFCBBFFFA0CA9FE28" box="[857,955,445,470]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
in mind, given the variation in pholidosis and coloration we have observed among specimens bearing the
|
||
<taxonomicName id="87A774BCFFEAFFDBFC8DFFA60CBDFE04" authorityName="Lindholm" authorityYear="1905" box="[879,943,481,506]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEAFFDBFC8DFFA60CBDFE04" box="[879,943,481,506]" italics="true" pageId="45" pageNumber="46">diehli</emphasis>
|
||
</taxonomicName>
|
||
name. We are currently engaged in a more detailed analysis to determine what represents
|
||
<emphasis id="72D3D32DFFEAFFDBFD43FC410C83FDE3" box="[673,913,517,542]" italics="true" pageId="45" pageNumber="46">
|
||
<taxonomicName id="87A774BCFFEAFFDBFD43FC410DEFFDE0" authority="Kaiser & Kaiser & O’Shea, 2018" authorityName="Kaiser & Kaiser & O’Shea" authorityYear="2018" box="[673,765,517,542]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" sensu="stricto" species="diehli">S. diehli</taxonomicName>
|
||
<taxonomicNameLabel id="69E06E56FFEAFFDBFCE6FC410C83FDE3" box="[772,913,518,541]" pageId="45" pageNumber="46" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
</emphasis>
|
||
(C. Kaiser
|
||
<emphasis id="72D3D32DFFEAFFDBFBF0FC410B58FDE0" box="[1042,1098,517,542]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
|
||
, in prep.).
|
||
</paragraph>
|
||
<paragraph id="40180F3FFFEAFFDBFF25FC6D0BF0FD4F" blockId="45.[151,1436,151,982]" pageId="45" pageNumber="46">
|
||
In the meantime, our species-level assessment of
|
||
<taxonomicName id="87A774BCFFEAFFDBFD16FC6D0C42FDBC" authorityName="Kaiser & Kaiser & O’Shea" authorityYear="2018" box="[756,848,553,578]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEAFFDBFD16FC6D0C42FDBC" box="[756,848,553,578]" italics="true" pageId="45" pageNumber="46">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
is limited, but specimens with values of D = 15-15- 15, SL
|
||
<subScript id="DC230D7AFFEAFFDBFF00FC1B0FF9FD94" attach="left" box="[226,235,604,618]" fontSize="6" pageId="45" pageNumber="46">E</subScript>
|
||
= 3+4, SL = 7, IL = 8, and IL
|
||
<subScript id="DC230D7AFFEAFFDBFDA2FC1B0D59FD94" attach="left" box="[576,587,604,618]" fontSize="6" pageId="45" pageNumber="46">G</subScript>
|
||
= 4, with relatively low ventral and subcaudal scale counts (V <180, SC <85; values based on the
|
||
<typeStatus id="9F1CB19DFFEAFFDBFE4CFC330D02FD73" box="[430,528,628,653]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
counts and considering standard deviations in these values in other
|
||
<taxonomicName id="87A774BCFFEAFFDBFAC0FC320A8EFD72" authorityName="Guentheri Boulenger" authorityYear="1895" box="[1314,1436,629,652]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="72D3D32DFFEAFFDBFAC0FC320A8EFD72" box="[1314,1436,629,652]" italics="true" pageId="45" pageNumber="46">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
species). However, values outside of these parameters may, probably rarely, occur in conspecifics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08BD5CB4FFEAFFDBFF25FCFB0D2FFC28" pageId="45" pageNumber="46" type="discussion">
|
||
<paragraph id="40180F3FFFEAFFDBFF25FCFB0D2FFC28" blockId="45.[151,1436,151,982]" pageId="45" pageNumber="46">
|
||
<emphasis id="72D3D32DFFEAFFDBFF25FCFB0E5DFD2B" bold="true" box="[199,335,700,725]" pageId="45" pageNumber="46">Comments.</emphasis>
|
||
<bibRefCitation id="243672CEFFEAFFDBFEBBFCFB0D0EFD2B" author="Lindholm, W. A." box="[345,540,700,725]" pageId="45" pageNumber="46" pagination="227 - 240" refId="ref44980" refString="Lindholm, W. A. (1905) Uber einige Eidechsen und Schlangen aus Deutsch-Neuguinea. Jahrbucher des nassauischen Fereins fur Naturkunde, 58, 227 - 240." type="journal article" year="1905">Lindholm (1905)</bibRefCitation>
|
||
stated in the original description that he considered this specimen to be “ein jüngeres gut erhaltenes Stück” [a juvenile well-preserved exemplar]. We have seen a variety of juvenile
|
||
<taxonomicName id="87A774BCFFEAFFDBFAC0FCA60A8EFD06" authorityName="Guentheri Boulenger" authorityYear="1895" box="[1314,1436,737,760]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="72D3D32DFFEAFFDBFAC0FCA60A8EFD06" box="[1314,1436,737,760]" italics="true" pageId="45" pageNumber="46">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
among the hundreds of specimens we have examined, and the color pattern of the head detailed by Lindholm exists in many of these. We are uncertain about the influence of preservatives on some of these markings, and the color in some juvenile specimens has clearly become bleached over time, but we have ascertained that this pattern is not unique to juveniles of
|
||
<taxonomicName id="87A774BCFFEAFFDBFE71FD360EE2FC77" authorityName="Kaiser & Kaiser & O’Shea" authorityYear="2018" box="[403,496,880,905]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="72D3D32DFFEAFFDBFE71FD360EE2FC77" box="[403,496,880,905]" italics="true" pageId="45" pageNumber="46">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
. It certainly exists in juvenile individuals with D = 17-17-15, SL
|
||
<subScript id="DC230D7AFFEAFFDBFB2FFD390BC4FC72" attach="left" box="[1229,1238,894,908]" fontSize="6" pageId="45" pageNumber="46">E</subScript>
|
||
= 4+5, SL = 8 or 9, IL = 9 or 10, and IL
|
||
<subScript id="DC230D7AFFEAFFDBFE49FDE20EA4FC4D" attach="left" box="[427,438,933,947]" fontSize="6" pageId="45" pageNumber="46">G</subScript>
|
||
= 5. Juvenile head patterning therefore does not present a suitable set of features for species-level diagnosis in
|
||
<taxonomicName id="87A774BCFFEAFFDBFE5EFDF90D24FC2B" authorityName="Guentheri Boulenger" authorityYear="1895" box="[444,566,958,981]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="72D3D32DFFEAFFDBFE5EFDF90D24FC2B" box="[444,566,958,981]" italics="true" pageId="45" pageNumber="46">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |