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<mods:title id="826550DAB65AB923DA54B406092FC44A">The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)</mods:title>
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<paragraph id="40180F3FFFE8FFD9FF75FED00D44FF4C" blockId="47.[151,598,151,178]" box="[151,598,151,178]" pageId="47" pageNumber="48">
<heading id="1B50B853FFE8FFD9FF75FED00D44FF4C" bold="true" box="[151,598,151,178]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<treatmentCitationGroup id="60B72811FFE8FFD9FF75FED00D44FF4C" box="[151,598,151,178]" pageId="47" pageNumber="48">
<taxonomicName id="87A774BCFFE8FFD9FF75FED00D44FF4C" authority="Werner, 1924: 32" authorityName="Werner" authorityPageNumber="32" authorityYear="1924" box="[151,598,151,178]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FF75FED00D44FF4C" bold="true" box="[151,598,151,178]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF75FED00E92FF4F" bold="true" box="[151,384,151,177]" italics="true" pageId="47" pageNumber="48">Stegonotus dorsalis</emphasis>
<treatmentCitation id="C106292EFFE8FFD9FE6AFEDF0D44FF4C" author="Werner, F." box="[392,598,152,178]" page="32" pageId="47" pageNumber="48" year="1924">
<bibRefCitation id="243672CEFFE8FFD9FE6AFEDF0D23FF4C" author="Werner, F." box="[392,561,152,178]" pageId="47" pageNumber="48" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werner, 1924</bibRefCitation>
:32
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</paragraph>
<paragraph id="40180F3FFFE8FFD9FF75FE9B0EE8FDBA" blockId="47.[151,1437,220,1914]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF75FE9B0E7DFF0B" bold="true" box="[151,367,220,245]" pageId="47" pageNumber="48">Taxonomic status.</emphasis>
<bibRefCitation id="243672CEFFE8FFD9FE95FE9B0D74FF0A" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[375,614,220,245]" pageId="47" pageNumber="48" refId="ref47316" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the World: a Catalogue of Living and Extinct Species. CRC Press, New York, USA, 1237 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach
<emphasis id="72D3D32DFFE8FFD9FE38FE9A0D19FF0B" box="[474,523,220,245]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2014)
</bibRefCitation>
listed
<taxonomicName id="87A774BCFFE8FFD9FD51FE9A0C38FF0B" authorityName="Werner" authorityYear="1924" box="[691,810,220,245]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FD51FE9A0C38FF0B" box="[691,810,220,245]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
in the synonymy of
<taxonomicName id="87A774BCFFE8FFD9FBF4FE9A0B61FF0B" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[1046,1139,220,245]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FBF4FE9A0B61FF0B" box="[1046,1139,220,245]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
without further comment.
<bibRefCitation id="243672CEFFE8FFD9FF75FEB90E29FEE8" author="Werner, F." box="[151,315,254,278]" pageId="47" pageNumber="48" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werner (1924)</bibRefCitation>
himself stated at the beginning of his description that he considered this species “nächstverwandt” [sister species] to
<taxonomicName id="87A774BCFFE8FFD9FE95FF580EC8FEC9" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[375,474,286,311]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FE95FF580EC8FEC9" box="[375,474,286,311]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
but provided a definitive series of differentiating characteristics. Based on the understanding of
<taxonomicName id="87A774BCFFE8FFD9FE87FF070ECDFEA9" authorityName="Guentheri Boulenger" authorityYear="1895" box="[357,479,320,343]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFE8FFD9FE87FF070ECDFEA9" box="[357,479,320,343]" italics="true" pageId="47" pageNumber="48">Stegonotus</emphasis>
</taxonomicName>
we have gained by examining nearly 1500 museum specimens, we consider the characteristics differentiating
<taxonomicName id="87A774BCFFE8FFD9FE13FF250D7CFE84" authorityName="Werner" authorityYear="1924" box="[497,622,353,378]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FE13FF250D7CFE84" box="[497,622,353,378]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
from
<taxonomicName id="87A774BCFFE8FFD9FD5DFF250C30FE84" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[703,802,353,378]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FD5DFF250C30FE84" box="[703,802,353,378]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
, in addition to the uncertain taxonomy of the latter, sufficient to uphold a taxonomic distinction at the species level. None of the well-defined taxa in the genus
<taxonomicName id="87A774BCFFE8FFD9FF75FFE20E03FE42" authorityName="Guentheri Boulenger" authorityYear="1895" box="[151,273,421,444]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFE8FFD9FF75FFE20E03FE42" box="[151,273,421,444]" italics="true" pageId="47" pageNumber="48">Stegonotus</emphasis>
</taxonomicName>
(
<taxonomicName id="87A774BCFFE8FFD9FEC2FFE20EA9FE43" baseAuthorityName="Gunther" baseAuthorityYear="1865" box="[288,443,420,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="batjanensis">
<emphasis id="72D3D32DFFE8FFD9FEC2FFE20EA9FE43" box="[288,443,420,445]" italics="true" pageId="47" pageNumber="48">S. batjanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFE8FFD9FE2BFFE20D5DFE43" authorityName="Boulenger" authorityYear="1895" box="[457,591,420,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="72D3D32DFFE8FFD9FE2BFFE20D5DFE43" box="[457,591,420,445]" italics="true" pageId="47" pageNumber="48">S. guentheri</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFE8FFD9FDBFFFE20DF1FE43" authorityName="Boulenger" authorityYear="1893" box="[605,739,420,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="heterurus">
<emphasis id="72D3D32DFFE8FFD9FDBFFFE20DF1FE43" box="[605,739,420,445]" italics="true" pageId="47" pageNumber="48">S. heterurus</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFE8FFD9FD13FFE20C7BFE43" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" baseAuthorityName="Doria" baseAuthorityYear="1874" box="[753,873,420,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="keyensis">
<emphasis id="72D3D32DFFE8FFD9FD13FFE20C7BFE43" box="[753,873,420,445]" italics="true" pageId="47" pageNumber="48">S. keyensis</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFE8FFD9FC95FFE20CE8FE43" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[887,1018,420,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFE8FFD9FC95FFE20CE8FE43" box="[887,1018,420,445]" italics="true" pageId="47" pageNumber="48">S. modestus</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFE8FFD9FBEAFFE20B92FE43" authorityName=", Boulenger" authorityYear="1893" box="[1032,1152,420,445]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="72D3D32DFFE8FFD9FBEAFFE20B92FE43" box="[1032,1152,420,445]" italics="true" pageId="47" pageNumber="48">S. muelleri</emphasis>
</taxonomicName>
) display ventral scales of sexes combined with a range of 29 scales (
<quantity id="875FA2DAFFE8FFD9FD60FF810DD9FE20" box="[642,715,453,478]" metricMagnitude="0" metricUnit="m" metricValue="4.5466" pageId="47" pageNumber="48" unit="in" value="179.0">179 in</quantity>
the
<typeStatus id="9F1CB19DFFE8FFD9FD1DFF820C73FE20" box="[767,865,453,478]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
of
<taxonomicName id="87A774BCFFE8FFD9FC6BFF810CF5FE20" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[905,999,453,478]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FC6BFF810CF5FE20" box="[905,999,453,478]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
to
<quantity id="875FA2DAFFE8FFD9FBEFFF810B44FE20" box="[1037,1110,453,478]" metricMagnitude="0" metricUnit="m" metricValue="5.2832" pageId="47" pageNumber="48" unit="in" value="208.0">208 in</quantity>
the
<typeStatus id="9F1CB19DFFE8FFD9FB6BFF820BF9FE20" box="[1161,1259,453,478]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
of
<taxonomicName id="87A774BCFFE8FFD9FAF6FF810A99FE20" authorityName="Werner" authorityYear="1924" box="[1300,1419,453,478]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FAF6FF810A99FE20" box="[1300,1419,453,478]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
), which already provides strong evidence for species-level distinction. Furthermore, differences include (features of
<taxonomicName id="87A774BCFFE8FFD9FF75FC4F0FE6FDDE" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[151,244,519,544]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FF75FC4F0FE6FDDE" box="[151,244,519,544]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
provided in parentheses) SL = 8 (7), SL
<subScript id="DC230D7AFFE8FFD9FD26FC510DDFFDDA" attach="left" box="[708,717,534,548]" fontSize="6" pageId="47" pageNumber="48">E</subScript>
= 4+5 (3+4), and IL
<subScript id="DC230D7AFFE8FFD9FC59FC510CD4FDDA" attach="left" box="[955,966,534,548]" fontSize="6" pageId="47" pageNumber="48">G</subScript>
= 5 (4). We therefore remove
<taxonomicName id="87A774BCFFE8FFD9FAC6FC4F0A8EFDDE" authorityName="Werner" authorityYear="1924" box="[1316,1436,519,544]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FAC6FC4F0A8EFDDE" box="[1316,1436,519,544]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
from the synonymy of
<taxonomicName id="87A774BCFFE8FFD9FE75FC6A0EE6FDBB" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[407,500,556,581]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FE75FC6A0EE6FDBB" box="[407,500,556,581]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFE8FFD9FF25FC0A0CE2FCB1" pageId="47" pageNumber="48" type="description">
<paragraph id="40180F3FFFE8FFD9FF25FC0A0E86FD9B" blockId="47.[151,1437,220,1914]" box="[199,404,589,614]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF25FC0A0E5BFD98" bold="true" box="[199,329,589,614]" pageId="47" pageNumber="48">Synonyms.</emphasis>
None.
</paragraph>
<paragraph id="40180F3FFFE8FFD9FF25FC290CE2FCB1" blockId="47.[151,1437,220,1914]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF25FC290E6FFD79" bold="true" box="[199,381,622,647]" pageId="47" pageNumber="48">Original name.</emphasis>
<taxonomicName id="87A774BCFFE8FFD9FE64FC280C3FFD79" authority="Werner, 1924: 32" authorityName="Werner" authorityPageNumber="32" authorityYear="1924" box="[390,813,622,647]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FE64FC280D71FD79" box="[390,611,622,647]" italics="true" pageId="47" pageNumber="48">Stegonotus dorsalis</emphasis>
<bibRefCitation id="243672CEFFE8FFD9FD8EFC370C18FD79" author="Werner, F." box="[620,778,623,647]" pageId="47" pageNumber="48" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werner, 1924</bibRefCitation>
:32
</taxonomicName>
. Franz Werner (
<date id="341929FFFFE8FFD9FC07FC290B7AFD79" box="[997,1128,622,647]" pageId="47" pageNumber="48" value="1867" valueMax="1939">18671939</date>
;
<figureCitation id="D89C13BAFFE8FFD9FB97FC290BC0FD79" box="[1141,1234,622,647]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="47" pageNumber="48">Fig. 17J</figureCitation>
) was an Austrian herpetologist and entomologist, as well as lecturer and professor at the Institute for Zoology at the University of
<collectingRegion id="8263C1DDFFE8FFD9FF75FCF60FF4FD34" box="[151,230,689,714]" country="Austria" name="Wien" pageId="47" pageNumber="48">Vienna</collectingRegion>
from
<date id="341929FFFFE8FFD9FECCFCF50EBDFD34" box="[302,431,690,714]" pageId="47" pageNumber="48" value="1891" valueMax="1933">18911933</date>
. It is noteworthy that Werner was initially unable to use specimens at the NMW for his research, having been barred by Franz Steindachner (
<date id="341929FFFFE8FFD9FD12FC940C61FD15" box="[752,883,723,747]" pageId="47" pageNumber="48" value="1834" valueMax="1919">18341919</date>
), the then-curator. However, after Steindachners death, these restrictions appear to have been loosened, as attested by Werners description of
<taxonomicName id="87A774BCFFE8FFD9FB06FCB20A4CFCF3" authorityName="Werner" authorityYear="1924" box="[1252,1374,756,781]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FB06FCB20A4CFCF3" box="[1252,1374,756,781]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
. The undated portrait of Werner (
<figureCitation id="D89C13BAFFE8FFD9FE33FD520D22FCD0" box="[465,560,789,814]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="47" pageNumber="48">Fig. 17J</figureCitation>
; public domain) was taken by the Viennese portrait photographer Albin Kobé (
<date id="341929FFFFE8FFD9FF7DFD710E0DFCB1" box="[159,287,822,847]" pageId="47" pageNumber="48" value="1884" valueMax="1963">18841963</date>
) and hangs in the office of the herpetology curator at the NMW.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFE8FFD9FF25FD1F0CDEF884" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="40180F3FFFE8FFD9FF25FD1F0C5FFC8F" blockId="47.[151,1437,220,1914]" box="[199,845,856,881]" pageId="47" pageNumber="48">
<materialsCitation id="F0CF0562FFE8FFD9FF25FD1F0C5FFC8F" ID-GBIF-Occurrence="2028140044" box="[199,845,856,881]" collectionCode="NMW" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">
<emphasis id="72D3D32DFFE8FFD9FF25FD1F0E29FC8F" bold="true" box="[199,315,856,881]" pageId="47" pageNumber="48">
<typeStatus id="9F1CB19DFFE8FFD9FF25FD1F0E24FC8F" box="[199,310,856,881]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="26B697FAFFE8FFD9FEA0FD1E0E98FC8E" box="[322,394,857,880]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="47" pageNumber="48">NMW</collectionCode>
14861 (
<figureCitation id="D89C13BAFFE8FFD9FE04FD1F0D2BFC8E" box="[486,569,856,881]" captionStart="FIGURE 27" captionStartId="48.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1050]" captionTargetId="figure@48.[151,1436,193,1050]" captionTargetPageId="48" captionText="FIGURE 27. Holotype of Stegonotus dorsalis Werner, 1924, NMW 14861, an adult male from near Bogia, Madang Province, Papua New Guinea. Shown are the entire specimen (TTL = 1050 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views." httpUri="https://zenodo.org/record/2607633/files/figure.png" pageId="47" pageNumber="48">Fig. 27</figureCitation>
;
<tableCitation id="0D253A84FFE8FFD9FDA7FD1F0D84FC8E" box="[581,662,856,881]" captionStart="TABLE 1" captionStartId="60.[147,229,603,624]" captionText="TABLE 1. Name-bearing types of species in the genus Stegonotus. In this list we present basic data, including common measurements and selected scale counts, for all namebearing types (i.e., holotypes—HT, lectotypes—LT, neotypes—NT, syntypes—ST) to provide an easy overview of species. Additional details and the information for paratypes and paralectotypes can be found in the text. Character abbreviations are explained in the main text. For specimens where sex is indicated in red, we used the SCR value and not direct observation to estimate sex. Missing values are indicated by O. Specimen Type SCR SubC Species Synonym Sex SVL TL TTL V SC Dorsals SCE SL IL ILG Number Status (%) Condition LSUMZ admiraltiensis HT M 658 185 843 208 98 32 paired 17-19-15 4+5 8 10 10 93958 MSNG aruensis HT M 582 233 815 190 103 35 paired 17-17-15 3+4 7 8 4 30186 BMNH australis HT M 503 135 638 205 77 27 paired 17-17-15 4+5 8 9 5 1946.1.14.93 AMS australis darnleyensis HT F 720 O O 198 53+ O paired 17-17-15 4+5 9 10 5 R31894 AMS australis plumbeus HT M 1025 250 1275 220 74 25 paired 17-17-15 3+4+5 9 10 5 R31895 BMNH batjanensis HT M 1239 321 1560 235 86 27 paired 17-17-15 3+4+5 8 10 5 1946.1.11.36 FMNH borneensis HT F 783 221 1004 195 79 29 paired 17-17-15 4+5 10 11 5 164746 MNHN cucullatus HT M 769 254 1023 209 91 30 paired 17-17-15 4+5 8 9 5 3412 MTKD a cucullatus magnus ST F O O O 207 74 26 paired O-19-O 4+5 8 O O 573 ZMB b cucullatus magnus ST M 975 265 1240 211 87 29 paired 19-19-17 3+4 8 10 5 8794 MZB derooijae HT M 455 193 648 181 94 34 paired 17-17-15 3+4 7 8 8 Ophi.3288 MWNH diehli HT juv 196 56 252 179 82 31 paired 15-15-15 3+4 7 8 4 1244 NMW dorsalis HT M 805 245 1050 208 88 30 paired 15-15-15 4+5 8 10 5 14861 ZMA 9R florensis HT M 447 109 556 217 65 23 single 21-21-19 3+4+5 10 5 11080 8L BMNH guentheri LT F 683 165 848 192 64 25 paired 15-15-13 4+5 8 9 4 1946.1.11.38 ……continued on the next page" pageId="47" pageNumber="48">Table 1</tableCitation>
), an adult male.
</materialsCitation>
</paragraph>
<paragraph id="40180F3FFFE8FFD9FF25FD3E0C49FA59" blockId="47.[151,1437,220,1914]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF25FD3E0E77FC6C" bold="true" box="[199,357,889,914]" pageId="47" pageNumber="48">Type locality.</emphasis>
Neither jar label nor specimen catalog entry provide any information regarding the
<typeStatus id="9F1CB19DFFE8FFD9FAEEFD3D0A2EFC6C" box="[1292,1340,890,914]" pageId="47" pageNumber="48">type</typeStatus>
locality. The historic jar label states “Fundort?” [locality?], indicating that no information about the
<typeStatus id="9F1CB19DFFE8FFD9FB41FDDC0BC1FC4D" box="[1187,1235,923,947]" pageId="47" pageNumber="48">type</typeStatus>
locality appeared to be available at the time the specimen was accessioned into the collection. However, based on the specimens included in
<bibRefCitation id="243672CEFFE8FFD9FEF9FD990EC0FC08" author="Werner, F." box="[283,466,990,1014]" pageId="47" pageNumber="48" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werners (1924)</bibRefCitation>
account of snakes in the Vienna museum it becomes clear that the only possible origin for specimens of
<taxonomicName id="87A774BCFFE8FFD9FE86FDB80ECCFBE8" authorityName="Guentheri Boulenger" authorityYear="1895" box="[356,478,1023,1046]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFE8FFD9FE86FDB80ECCFBE8" box="[356,478,1023,1046]" italics="true" pageId="47" pageNumber="48">Stegonotus</emphasis>
</taxonomicName>
housed there is New
<collectingCountry id="38B04FAFFFE8FFD9FD01FDB90C2AFBE9" box="[739,824,1022,1047]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
. Furthermore, the only zoological collection in New
<collectingCountry id="38B04FAFFFE8FFD9FF75FA670FFAFBC7" box="[151,232,1056,1081]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
that would have found its way exclusively into the Vienna museum was made by the anthropologist and explorer Rudolf Pöch (
<date id="341929FFFFE8FFD9FE7FFA050D32FBA4" box="[413,544,1090,1114]" pageId="47" pageNumber="48" value="1870" valueMax="1921">18701921</date>
;
<figureCitation id="D89C13BAFFE8FFD9FDCFFA060D84FBA4" box="[557,662,1089,1114]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="47" pageNumber="48">Fig. 17K</figureCitation>
), later a professor of anthropology at Vienna University, who spent several months in New
<collectingCountry id="38B04FAFFFE8FFD9FE46FA250EE7FB85" box="[420,501,1122,1147]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
in 1904 (
<date id="341929FFFFE8FFD9FD86FA240C76FB85" box="[612,868,1122,1147]" pageId="47" pageNumber="48">28 July25 November</date>
;
<bibRefCitation id="243672CEFFE8FFD9FC93FA250CFEFB85" author="Poch, R." box="[881,1004,1122,1147]" pageId="47" pageNumber="48" pagination="609 - 616" refId="ref45924" refString="Poch, R. (1907) Travels in German, British, and Dutch New Guinea. The Geographical Journal, 30 (6), 609 - 616. https: // doi. org / 10.2307 / 1776812" type="journal article" year="1907">Pöch 1907</bibRefCitation>
,
<bibRefCitation id="243672CEFFE8FFD9FC1AFA240B21FB85" author="Poch, R." box="[1016,1075,1123,1147]" pageId="47" pageNumber="48" pagination="169 - 173" refId="ref45965" refString="Poch, R. (1908) Reisen an der Nordkuste von Kaiser Wilhelmsland. Globus, 93 (9), 139 - 143, continued in 93 (10), 149 - 155, continued in 93 (11), 169 - 173." type="journal article" year="1908">1908</bibRefCitation>
), and whose name is associated with the (falsely identified) specimen of “
<taxonomicName id="87A774BCFFE8FFD9FD85FAC20C5DFB63" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[615,847,1156,1181]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFE8FFD9FD85FAC20C5DFB63" box="[615,847,1156,1181]" italics="true" pageId="47" pageNumber="48">Stegonotus modestus</emphasis>
</taxonomicName>
” that
<bibRefCitation id="243672CEFFE8FFD9FC70FAC20B26FB62" author="Werner, F." box="[914,1076,1156,1180]" pageId="47" pageNumber="48" pagination="29 - 56" refId="ref47499" refString="Werner, F. (1924) Neue oder wenig bekannte Schlangen aus dem Naturhistorischen Staatsmuseum in Wien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 133, 29 - 56." type="journal article" year="1924">Werner (1924)</bibRefCitation>
included in his account. Further research we conducted in the archives of the Vienna collection led to the discovery of historic specimen inventory slips indicating New
<collectingCountry id="38B04FAFFFE8FFD9FE73FA810EF0FB21" box="[401,482,1222,1247]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
as the collection locality, as well as letters from Pöch to the then-curator, Franz Steindachner, in which the former recounts some of his travels. In one of these letters, Pöch wrote that specimens that had been sent without any specific locality information (i.e., those merely listing New
<collectingCountry id="38B04FAFFFE8FFD9FB3AFB4E0A38FADC" box="[1240,1322,1289,1314]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
) were all collected at “Monumbo, Küstenstrich von Potsdamhafen, Deutsch-Neu-Guinea” [Monumbo, in the coastal region around Potsdamhafen, German New
<collectingCountry id="38B04FAFFFE8FFD9FDA5FB0B0D89FA9B" box="[583,667,1356,1381]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
, now
<collectingCountry id="38B04FAFFFE8FFD9FD04FB0A0CDFFA9B" box="[742,973,1356,1381]" name="Papua New Guinea" pageId="47" pageNumber="48">Papua New Guinea</collectingCountry>
]. A notation in the margin of the letter reveals that one package of specimens from this area was indeed inventoried. Alas, modern maps and gazetteers of
<collectingCountry id="38B04FAFFFE8FFD9FF75FBD70E7DFA59" box="[151,367,1422,1447]" name="Papua New Guinea" pageId="47" pageNumber="48">Papua New Guinea</collectingCountry>
show neither Monumbo nor Potsdamhafen.
</paragraph>
<paragraph id="40180F3FFFE8FFD9FF25FBF70A18F96F" blockId="47.[151,1437,220,1914]" pageId="47" pageNumber="48">
German maps of the early 1900s and, specifically, one created by Pöch himself in accordance with the
<emphasis id="72D3D32DFFE8FFD9FAC9FBF70E3EFA14" italics="true" pageId="47" pageNumber="48">Deutscher Kolonialatlas</emphasis>
[German colonial atlas] (see
<bibRefCitation id="243672CEFFE8FFD9FD94FB960DFDFA14" author="Poch, R." box="[630,751,1489,1514]" pageId="47" pageNumber="48" pagination="169 - 173" refId="ref45965" refString="Poch, R. (1908) Reisen an der Nordkuste von Kaiser Wilhelmsland. Globus, 93 (9), 139 - 143, continued in 93 (10), 149 - 155, continued in 93 (11), 169 - 173." type="journal article" year="1908">Pöch 1908</bibRefCitation>
), reveal that the location of Potsdamhafen was in a small bay across from Manam Island,
<collectingRegion id="8263C1DDFFE8FFD9FE2FFBB50D81F9F5" box="[461,659,1522,1547]" country="Papua New Guinea" name="Madang" pageId="47" pageNumber="48">Madang Province</collectingRegion>
,
<collectingCountry id="38B04FAFFFE8FFD9FD7DFBB40C67F9F5" box="[671,885,1522,1547]" name="Papua New Guinea" pageId="47" pageNumber="48">Papua New Guinea</collectingCountry>
(ca.
<geoCoordinate id="259369F8FFE8FFD9FC4FFBB40B04F9F5" box="[941,1046,1523,1547]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="-4.228">4.2280°S</geoCoordinate>
,
<geoCoordinate id="259369F8FFE8FFD9FBC3FBB40BBBF9F5" box="[1057,1193,1523,1547]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="144.9333">144.9333°E</geoCoordinate>
).
<materialsCitation id="F0CF0562FFE8FFD9FB5EFBB50A14F96F" ID-GBIF-Occurrence="2028140041" country="Austria" location="Vienna" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Wien">
Pöchs map and text also show that he used the term “Monumbo” for both a New Guinean tribe and the mountains bordering Potsdamhafen on the landward side. Pöch made several excursions into and through these mountains and included some biological aspects of his surveys in his travelogue (
<bibRefCitation id="243672CEFFE8FFD9FCF1F8110C9CF991" author="Poch, R." box="[787,910,1622,1647]" pageId="47" pageNumber="48" pagination="169 - 173" refId="ref45965" refString="Poch, R. (1908) Reisen an der Nordkuste von Kaiser Wilhelmsland. Globus, 93 (9), 139 - 143, continued in 93 (10), 149 - 155, continued in 93 (11), 169 - 173." type="journal article" year="1908">Pöch 1908</bibRefCitation>
). The image of Pöch (
<figureCitation id="D89C13BAFFE8FFD9FB6AF8110BFDF991" box="[1160,1263,1622,1647]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="47" pageNumber="48">Fig. 17K</figureCitation>
; licensed under CC-BY-SA-4.0) is from the photographic collection of the archives at the University of
<collectingRegion id="8263C1DDFFE8FFD9FBB5F83F0BBAF96F" box="[1111,1192,1656,1681]" country="Austria" name="Wien" pageId="47" pageNumber="48">Vienna</collectingRegion>
,
<collectingCountry id="38B04FAFFFE8FFD9FB50F83F0A14F96F" box="[1202,1286,1656,1681]" name="Austria" pageId="47" pageNumber="48">Austria</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="40180F3FFFE8FFD9FF25F8DE0CDEF884" blockId="47.[151,1437,220,1914]" pageId="47" pageNumber="48">
<emphasis id="72D3D32DFFE8FFD9FF25F8DE0E55F94C" bold="true" box="[199,327,1689,1714]" pageId="47" pageNumber="48">Collection.</emphasis>
The label affixed to the jar lists only the comment “Coll.?” [collector?], indicating that no information was available. The most likely scenario for the collection of the
<typeStatus id="9F1CB19DFFE8FFD9FC12F8FD0B40F92D" box="[1008,1106,1722,1747]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
of
<taxonomicName id="87A774BCFFE8FFD9FB94F8FC0A43F92D" authorityName="Werner" authorityYear="1924" box="[1142,1361,1722,1747]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FB94F8FC0A43F92D" box="[1142,1361,1722,1747]" italics="true" pageId="47" pageNumber="48">Stegonotus dorsalis</emphasis>
</taxonomicName>
is that Pöch encountered the snake on one of his forays into the interior during the months of AugustNovember 1904. With no opportunity to specify a locality on these expeditions, and with a focus on anthropological research, Pöch could have grouped his collected zoological specimens and provided the general locality of Monumbo for any specimens from the area inland of Potsdamhafen. The type locality of
<taxonomicName id="87A774BCFFE8FFD9FC49F9060B33F8A7" authorityName="Werner" authorityYear="1924" box="[939,1057,1856,1881]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFE8FFD9FC49F9060B33F8A7" box="[939,1057,1856,1881]" italics="true" pageId="47" pageNumber="48">S. dorsalis</emphasis>
</taxonomicName>
(see below) is ca.
<quantity id="875FA2DAFFE8FFD9FB11F9070A5AF8A7" box="[1267,1352,1856,1881]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="47" pageNumber="48" unit="km" value="160.0">160 km</quantity>
NW of the type locality of
<taxonomicName id="87A774BCFFE8FFD9FE92F9250EDEF884" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[368,460,1889,1914]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFE8FFD9FE92F9250EDEF884" box="[368,460,1889,1914]" italics="true" pageId="47" pageNumber="48">S. diehli</emphasis>
</taxonomicName>
, across the mountains of the Adelbert Range.
</paragraph>
</subSubSection>
<caption id="14D85FB7FFF7FFC6FF75FA680D7FFB77" httpUri="https://zenodo.org/record/2607633/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1071,1093]" targetBox="[151,1436,193,1050]" targetPageId="48">
<paragraph id="40180F3FFFF7FFC6FF75FA680D7FFB77" blockId="48.[151,1436,1071,1161]" pageId="48" pageNumber="49">
<emphasis id="72D3D32DFFF7FFC6FF75FA680E32FBBA" bold="true" box="[151,288,1071,1093]" pageId="48" pageNumber="49">FIGURE 27.</emphasis>
Holotype of
<taxonomicName id="87A774BCFFF7FFC6FE45FA770C12FBBB" authority="Werner, 1924" authorityName="Werner" authorityYear="1924" box="[423,768,1071,1093]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFF7FFC6FE45FA770D7CFBBB" box="[423,622,1071,1093]" italics="true" pageId="48" pageNumber="49">Stegonotus dorsalis</emphasis>
Werner, 1924
</taxonomicName>
, NMW 14861, an adult male from near Bogia, Madang Province, Papua New Guinea. Shown are the entire specimen (TTL = 1050 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views.
</paragraph>
</caption>
<subSubSection id="08BD5CB4FFF7FFC6FF25FA9D0B13FA3D" pageId="48" pageNumber="49" type="key">
<paragraph id="40180F3FFFF7FFC6FF25FA9D0E32FAC5" blockId="48.[151,1436,1242,1511]" pageId="48" pageNumber="49">
<emphasis id="72D3D32DFFF7FFC6FF25FA9D0D7CFB0D" bold="true" box="[199,622,1242,1267]" pageId="48" pageNumber="49">
Key characteristics of the
<materialsCitation id="F0CF0562FFF7FFC6FDE2FA9D0D7CFB0D" ID-GBIF-Occurrence="2028140070" box="[512,622,1242,1267]" collectionCode="SVL" pageId="48" pageNumber="49" specimenCount="1" typeStatus="holotype">
<typeStatus id="9F1CB19DFFF7FFC6FDE2FA9D0D7BFB0D" box="[512,617,1242,1267]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</emphasis>
805 (805) mm SVL + 245 (245) mm TL = 1050 (1050) mm TTL.
<materialsCitation id="F0CF0562FFF7FFC6FA87FA9C0EE1FAE9" ID-GBIF-Occurrence="2028140040" collectionCode="SCR" pageId="48" pageNumber="49" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
V
<specimenCount id="56A1C4B6FFF7FFC6FA9BFAAF0A97FB08" box="[1401,1413,1256,1270]" pageId="48" pageNumber="49" type="male">
<subScript id="DC230D7AFFF7FFC6FA9BFAAF0A97FB08" attach="left" box="[1401,1413,1256,1270]" fontSize="6" pageId="48" pageNumber="49"></subScript>
</specimenCount>
= 208 (208), SC
<subScript id="DC230D7AFFF7FFC6FED7FB4B0E53FAE4" attach="left" box="[309,321,1292,1306]" fontSize="6" pageId="48" pageNumber="49">
<specimenCount id="56A1C4B6FFF7FFC6FED7FB4B0E53FAE4" box="[309,321,1292,1306]" pageId="48" pageNumber="49" type="male"></specimenCount>
</subScript>
= 88 (88),
<collectionCode id="26B697FAFFF7FFC6FE5CFAB80EE1FAE9" box="[446,499,1279,1303]" country="USA" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography, University of California" pageId="48" pageNumber="49" type="Herbarium">SCR</collectionCode>
</materialsCitation>
<materialsCitation id="F0CF0562FFF7FFC6FE11FB4B0A38FAE8" ID-GBIF-Occurrence="2028140067" box="[499,1322,1278,1306]" collectionCode="IL" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<subScript id="DC230D7AFFF7FFC6FE11FB4B0EEDFAE4" attach="left" box="[499,511,1292,1306]" fontSize="6" pageId="48" pageNumber="49">
<specimenCount id="56A1C4B6FFF7FFC6FE11FB4B0EEDFAE4" box="[499,511,1292,1306]" pageId="48" pageNumber="49" type="male"></specimenCount>
</subScript>
= 0.30 (0.30), D = O-15-O (15-15-15), SL
<subScript id="DC230D7AFFF7FFC6FC04FB4B0CFDFAE4" attach="left" box="[998,1007,1292,1306]" fontSize="6" pageId="48" pageNumber="49">E</subScript>
= 4+5 (4+5), SL = 8 (8),
<collectionCode id="26B697FAFFF7FFC6FAEDFAB80A38FAE8" box="[1295,1322,1279,1302]" pageId="48" pageNumber="49">IL</collectionCode>
</materialsCitation>
= O (10), IL
<subScript id="DC230D7AFFF7FFC6FF53FB760FAEFAC1" attach="left" box="[177,188,1329,1343]" fontSize="6" pageId="48" pageNumber="49">G</subScript>
= O (5).
</paragraph>
<paragraph id="40180F3FFFF7FFC6FF25FB010B13FA3D" blockId="48.[151,1436,1242,1511]" pageId="48" pageNumber="49">
<emphasis id="72D3D32DFFF7FFC6FF25FB010D49FAA1" bold="true" box="[199,603,1350,1375]" pageId="48" pageNumber="49">Key characteristics of the species.</emphasis>
With only a single specimen available, any species-level characterization must remain very general. It appears that
<taxonomicName id="87A774BCFFF7FFC6FD81FB2F0DCAFA7E" authorityName="Werner" authorityYear="1924" box="[611,728,1383,1408]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="72D3D32DFFF7FFC6FD81FB2F0DCAFA7E" box="[611,728,1383,1408]" italics="true" pageId="48" pageNumber="49">S. dorsalis</emphasis>
</taxonomicName>
has a relatively high ventral count (V = 208) compared to other species with 15 middorsal scale rows (
<taxonomicName id="87A774BCFFF7FFC6FD87FBCD0DD7FA5C" authorityName="Kaiser &amp; Kaiser &amp; OShea" authorityYear="2018" box="[613,709,1417,1442]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="diehli">
<emphasis id="72D3D32DFFF7FFC6FD87FBCD0DD7FA5C" box="[613,709,1417,1442]" italics="true" pageId="48" pageNumber="49">S. diehli</emphasis>
</taxonomicName>
,
<taxonomicName id="87A774BCFFF7FFC6FD3BFBCD0C77FA5C" authorityName="Boulenger" authorityYear="1895" box="[729,869,1417,1442]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="72D3D32DFFF7FFC6FD3BFBCD0C77FA5C" box="[729,869,1417,1442]" italics="true" pageId="48" pageNumber="49">S. guentheri</emphasis>
</taxonomicName>
), whose counts reach only the mid-190s. Other characteristics include SL
<subScript id="DC230D7AFFF7FFC6FE5AFBFE0ED3FA39" attach="left" box="[440,449,1465,1479]" fontSize="6" pageId="48" pageNumber="49">E</subScript>
= 4+5, IL
<subScript id="DC230D7AFFF7FFC6FDD1FBFE0D2CFA39" attach="left" box="[563,574,1465,1479]" fontSize="6" pageId="48" pageNumber="49">G</subScript>
= 5, and a relatively high IL count (10).
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFF7FFC6FF25FB890A57FA19" box="[199,1349,1486,1511]" pageId="48" pageNumber="49" type="discussion">
<paragraph id="40180F3FFFF7FFC6FF25FB890A57FA19" blockId="48.[151,1436,1242,1511]" box="[199,1349,1486,1511]" pageId="48" pageNumber="49">
<emphasis id="72D3D32DFFF7FFC6FF25FB890E56FA19" bold="true" box="[199,324,1486,1511]" pageId="48" pageNumber="49">Comment.</emphasis>
This species was not mentioned in the works of
<bibRefCitation id="243672CEFFF7FFC6FC84FB890B3DFA19" author="McDowell, S. B." box="[870,1071,1486,1511]" pageId="48" pageNumber="49" pagination="6 - 26" refId="ref45102" refString="McDowell, S. B. (1972) The species of Stegonotus (Serpentes, Colubridae) in Papua New Guinea. Zoologische Mededelingen, 47 (2), 6 - 26." type="journal article" year="1972">McDowell (1972)</bibRefCitation>
and Ruane
<emphasis id="72D3D32DFFF7FFC6FB56FB880BF7FA19" box="[1204,1253,1486,1511]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2017).
</paragraph>
</subSubSection>
</treatment>
</document>