treatments-xml/data/AA/E4/16/AAE41682797B55719EB05CFA4076CFA2.xml

<document id="34D3B043A68D62CAAC4AE910ABD54059" ID-DOI="10.3897/contrib.entomol.74.e126799" ID-ZooBank="339AB68C-DCB1-435F-B88E-2C42D0A7B05E" ID-publisher-id="126799" URI-arpha="B2861E35-EA80-53AF-A4AD-1BA2816CBD9C" URI-zoobank="http://zoobank.org/339AB68C-DCB1-435F-B88E-2C42D0A7B05E" article-type="research-article" checkinTime="1722264657988" checkinUser="pensoft" docAuthor="da Costa-Silva, Vinícius, Strümpher, Werner P., Barclay, Maxwell V. L. &amp; Vaz-de-Mello, Fernando Z." docDate="2024" docId="AAE41682797B55719EB05CFA4076CFA2" docLanguage="en" docName="Contributions to Entomology 74(1): 81-101" docOrigin="Contributions to Entomology 74 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/126799/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Omorgus (Haroldomorgus) batesi" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="101" masterDocId="B2861E35EA8053AFA4AD1BA2816CBD9C" masterDocTitle="An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes" masterLastPageNumber="101" masterPageNumber="81" pageNumber="81" updateTime="1722265173734" updateUser="ExternalLinkService">
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<mods:title id="02665A194A241E7AE298A4413CDBE0AD">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:namePart id="BD01ABA884B1A69DC181FC966FD37045">da Costa-Silva, Vinícius</mods:namePart>
<mods:nameIdentifier id="C9896533757A1192FE3389CBD7858F58" type="ORCID">0000-0002-4556-3793</mods:nameIdentifier>
<mods:affiliation id="B6A6FF1F37E9CD58792414124E984C94">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:namePart id="13D63A53883F5F05FE6B0A8D140CF06F">Strümpher, Werner P.</mods:namePart>
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<mods:affiliation id="0B75A7C72D72D1DB424E49055E95E158">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
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<mods:namePart id="030B52F96FE1B5892CC0CB60B4F454D6">Barclay, Maxwell V. L.</mods:namePart>
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<mods:affiliation id="361F1AD070F860E21E49F7DB323249D3">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
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<mods:namePart id="7F6D71D98F5BE4BFEF2D0ACA3953417C">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="F395CDCD6FEBECAE265BFE553406A90F">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:title id="2A938B6077AB5ADFE0FC0A9CF7A60F2B">Contributions to Entomology</mods:title>
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<mods:date id="A084E4D8D22526EDCE8BE6F2EF0D8DBB">2024</mods:date>
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<mods:number id="D5A3255A48134C231704809EE9837B9E">2024-07-29</mods:number>
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<mods:number id="13745A20A9F632733014155226ED8DE4">74</mods:number>
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<subSubSection id="E05724007124FDC08A328DBE4A79B0AD" type="nomenclature">
<paragraph id="28F2B34AC059E70A768251F6F9C0C984">
<taxonomicName id="3F49B37964F30FFDDF2C97376FC19A74" ID-CoL="49DQD" authority="(Harold, 1872)" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="18A0905C01B2B97105D042950FB926E4" italics="true">Omorgus</emphasis>
(
<emphasis id="D289EF1589062A0BAFDED198C81CE9D0" italics="true">Haroldomorgus</emphasis>
) 
<emphasis id="ADE265465134BF2FD509D4D3514763B2" italics="true">batesi</emphasis>
(
<bibRefCitation id="2179EACC62224DF209BDA66A6DD52BD0" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F098D270F818191E31EEB89FCA860700" type="reference_group">
<paragraph id="2734DCE8896DBF476A3789525C29088D">
<treatmentCitationGroup id="915A9A3F65FDC9CFDEF97AD56673F485">
<treatmentCitation id="4A9C7525FC2A560FECB1ED2BD00A361E" author="Harold E" page="126" year="1872">
<taxonomicName id="213F023B508B350A9B40488159C1E4E4" ID-CoL="7CWTQ" authority="Harold, 1872: 126" authorityName="Harold" authorityPageNumber="126" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="4AF456D29BA5DC8AC5AFA2308027A199" italics="true">Trox batesi</emphasis>
<comment id="2E56592B4811A3B2D7AB9B67207843D4">
<bibRefCitation id="F817089CEA34F4F71DF0BB82AC7E18DB" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872: 126</bibRefCitation>
(original description); 
<treatmentCitation id="239DDA19717C9EC90F54CFE751592B1C" author="Arrow GJ" page="54" year="1912">
<bibRefCitation id="45ADC035C5E4F5871F9663711D7CCB68" author="Arrow GJ" firstAuthor="Arrow" journalOrPublisher="Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk" page="54" refId="B3" refString="Arrow GJ (1912) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk, 66 pp." title="Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43." year="1912">Arrow 1912: 54</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="CEE4D830B1C1DDAB569A99AEB3E4FFC8" author="Blackwelder RE" page="218" year="1944">
<bibRefCitation id="0766F4392EF3517A57937CA337A4A17A" DOI="10.5479/si.03629236.185.2" author="Blackwelder RE" firstAuthor="Blackwelder" journalOrPublisher="Bulletin of the United States National Museum" page="218" pagination="218-219" refId="B10" refString="Blackwelder RE (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae. Bulletin of the United States National Museum 185: 218–219. https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae." volume="185" year="1944">Blackwelder 1944: 218</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="C5106E47BB89F04CC7F2BDAC1E83786D" author="Vaurie P" page="154" year="1962">
<bibRefCitation id="CCF4719E9D0B04AC59B2D6EEF60869C2" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="154" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 154</bibRefCitation>
</treatmentCitation>
(redescription).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="1FEF7DA64A61FB96DB335678AE25DAA5">
<treatmentCitationGroup id="CDFE8CCDD95705D629AD9AB21AA80484">
<treatmentCitation id="6798AC2A9E9A92A89B68B4C55D45502D">
<taxonomicName id="B79E35AC5192185C0E3D3C24BBC5BC0D" ID-CoL="7CWTQ" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Omorgus">
<emphasis id="66FF06731FBEA2EBD818E3D985AC730B" italics="true">Trox</emphasis>
(
<emphasis id="354C0FB483F3977EA2EA92F618E9EBDF" italics="true">Omorgus</emphasis>
) 
<emphasis id="0A4ED761E89B0B3BDF523A4A3C2F7D6C" italics="true">batesi</emphasis>
</taxonomicName>
<comment id="0B3EC5E1435751E3261F1B655B29FABD">
: 
<treatmentCitation id="2C0BDC587ADC88549CD84C990960894F" author="Scholtz CH" page="8" year="1982">
<bibRefCitation id="9D9BF70B015E18B53B35D00A2EE2296E" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="8" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 8</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E552B1F79CB0BF14010A8984E9D59C50">
<treatmentCitationGroup id="489BA9630C7115C7BEA3DAF5719E8637">
<treatmentCitation id="00651084B88D851637C277FCC1BAEBBB">
<taxonomicName id="A11BC77431E4CDC71906759F27053F7F" ID-CoL="49DQD" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="0582ACB32A01E8D39BD257ACC3617CDA" italics="true">Omorgus</emphasis>
(
<emphasis id="E6248EB8D433B501A64E226037F68690" italics="true">Haroldomorgus</emphasis>
) 
<emphasis id="04BB2877B3F1F142CAFD8056E3F9C8F3" italics="true">batesi</emphasis>
</taxonomicName>
<comment id="39E5D5B0EB51354EF40906981B749332">
: 
<treatmentCitation id="71EB29838CC2C4765CB2C54222C4F117" author="Scholtz CH" page="362" year="">
<bibRefCitation id="05D83253B29A3F4904A2799EDE2F1437" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz 1986 a</bibRefCitation>
: 362
</treatmentCitation>
(phylogenetics); 
<treatmentCitation id="9B7AD59A87EA40B88C7054D375CB6578" author="Scholtz CH" page="1418" year="1990">
<bibRefCitation id="437066C765781BB89A018AD672A376B4" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1418" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1418</bibRefCitation>
</treatmentCitation>
(redescription); 
<treatmentCitation id="C6DC83BBDAB422F88D143BB3D70F129A" author="Zidek J" page="7" year="2013">
<bibRefCitation id="3874947F874FA806A22B77A1293E7289" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="7" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 7</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="27D203EE47F7BEF2EBE49402EFC37D0C" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" page="549" year="2014">
<bibRefCitation id="4820DB48A01CE31F5A152A50AE25F147" DOI="10.1111/syen.12074" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="3" journalOrPublisher="Systematic Entomology" page="549" pagination="548-562" refId="B104" refString="Strümpher WP, Sole CL, Villet MH, Scholtz CH (2014) Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. Systematic Entomology 39 (3): 548–562. https://doi.org/10.1111/syen.12074" title="Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data." volume="39" year="2014">Strümpher et al. 2014: 549</bibRefCitation>
</treatmentCitation>
(systematics); 
<treatmentCitation id="1F3F915DA6484672EF592E924FDFFC78" author="Strümpher WP &amp; Villet MH &amp; Sole CL &amp; Scholtz CH" page="62" year="2016">
<bibRefCitation id="D0F7218971895599127F4F45C6DF0A59" DOI="10.1163/1876312X-46052133" author="Strümpher WP &amp; Villet MH &amp; Sole CL &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="1" journalOrPublisher="Insect Systematics &amp; Evolution" page="62" pagination="53-82" refId="B105" refString="Strümpher WP, Villet MH, Sole CL, Scholtz CH (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics &amp; Evolution 47 (1): 53–82. https://doi.org/10.1163/1876312X-46052133" title="Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea)." volume="47" year="2016">Strümpher et al. 2016: 62</bibRefCitation>
</treatmentCitation>
(systematics); 
<treatmentCitation id="3E09C5246716990D974F80A36C0ADCB0" author="Zidek J" page="98" year="2017">
<bibRefCitation id="34DCA27B00B23F688E262119DD6813C7" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="98" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 98</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="E6D16DDCFE33C5301932A7BD18EEDA07" author="Hielkema AJ &amp; Hielkema MA" page="10" year="2019">
<bibRefCitation id="3029D5D0547DDCE5814341F9E3E6C7FB" author="Hielkema AJ &amp; Hielkema MA" firstAuthor="Hielkema" journalOrPublisher="Insecta Mundi" page="10" pagination="1-306" refId="B53" refString="Hielkema AJ, Hielkema MA (2019) An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732: 1–306." title="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas." volume="732" year="2019">Hielkema and Hielkema 2019: 10</bibRefCitation>
</treatmentCitation>
(checklist for the Guianas); 
<treatmentCitation id="11CBBCF171BD5B876F32D002CB857293" ID-DOI="http://doi.org/10.5281/zenodo.5030752" ID-Zenodo-Dep="5030752" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFAAFFE3FE95FE32FD431855" page="1997" year="2021">
<bibRefCitation id="E999BF3E46A51F460CBFF75630FE2EEF" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1997" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 1997</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F2BFDCF0C7F933D6BB7FEA4CA30ADDA7">
<treatmentCitationGroup id="A56E8793CB213AFB437E41016FB53726">
<treatmentCitation id="3E555C8DDF9A3EAFE005F9279AECA269">
<taxonomicName id="F8DE3757E6E23C040DAEEEF86F591B70" ID-CoL="49DQD" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="B9B9B003D7E3AE8F0505A735E057B2CD" italics="true">Omorgus batesi</emphasis>
</taxonomicName>
<comment id="1A90622963B1F14EB8952E7715BFE2A9">
: 
<treatmentCitation id="D5493AA2EF33D14F4D6A4AD34574CA2D" author="Deloya C" page="122" year="2005">
<bibRefCitation id="A56520A56F70DA122DF40CA0A042076B" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESHAE" type="Type specimen examined">
<paragraph id="54B66FCCFAFE2500EB8B568E4B97628F">
<heading id="6E59C1732FAA405479C6F3FC4AC02A70" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="1F52F2EA4FF344F7547EA0CE47BC448B">
<materialsCitation id="B0699F9D505DE7A0E4C67E48A369981A" collectionCode="MNHN" location="See" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="97E75DBBE64257BA4D2C087C0063A43B" bold="true" italics="true">
<typeStatus id="B7AE2040C56DB441D98DAD959A2130C3" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="E2B0157FC712C4EA1960F9E51E4E4CC8" count="1" type="male">♂</specimenCount>
<named-content id="A64F786CC72E9580701AE5E670E70EE4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<emphasis id="98E72B59F2BF38459DC2FA93791EBFC3" bold="true">
<collectionCode id="465F18F8DAA9B43E1830EF5B1A96440D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</emphasis>
</named-content>
). 
<location id="FCACA370276EF1D7D07440550412AB42" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:FCACA370276EF1D7D07440550412AB42" name="See" stateProvince="Amazonas">See</location>
<bibRefCitation id="D0036923992B3EDB9854EED693A3408A" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1997" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 1997)</bibRefCitation>
for high-quality images of the type specimen. 
<emphasis id="759151E0DEA0E6D0B2AE25532050756C" bold="true">Type locality</emphasis>
: “ Amazonestrom ” [
<collectingRegion id="19C4BB1046DC8B3BE33002FE690C6C39" country="Peru" name="Amazonas">Amazonas</collectingRegion>
River].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIAE" type="distribution">
<paragraph id="2FACC83005367B2E2957795391E05EA6">
<heading id="51878925EF05374370DEE0A335A7B196" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="3496C75F20C14C92C47006454221F112">
<collectingCountry id="46A1F7963692B5116DBB2304AE66FDFF" name="Argentina">Argentina</collectingCountry>
, 
<collectingCountry id="E07B7FC7676D921699F2600EAD05793D" name="Bolivia">Bolivia</collectingCountry>
, 
<collectingCountry id="0866953D3D982EDE9F8FC4A7132E8348" name="Brazil">Brazil</collectingCountry>
and 
<collectingCountry id="67722E7B65FF79275F7545766DCFFCBC" name="Paraguay">Paraguay</collectingCountry>
(
<bibRefCitation id="AB40FA841FF9D3CE5B0B05671C8D515E" author="Zidek" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017</bibRefCitation>
; 
<bibRefCitation id="F92A5997C1C4994F2091DC64ACC268D9" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWIAE" type="type material">
<paragraph id="2930A1EC1954BF7380153AE6F0FC88FD">
<heading id="60284A0E5C2F7424EA560C28F485250C" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="A9ACC2DE8B54796CA65C55D3847AC080">
<emphasis id="B24FDD138C753F035E174AE089122149" bold="true">
(
<specimenCount id="483D60EBC816F1CE3666C997A9D1F38E" count="4" type="generic">four specimens</specimenCount>
additionally to 
<bibRefCitation id="49BFD1558FE4E943188AFEC646715D76" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz (1990)</bibRefCitation>
and 
<bibRefCitation id="9D77B9BEEE9C0537F841026521CAF9D9" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021)</bibRefCitation>
).
</emphasis>
<materialsCitation id="1C04C9A866F4D95EF706A062EA2257B0" country="Brazil" location="Amazonas" specimenCount="1" stateProvince="Amazonas">
<collectingCountry id="81D96254E8C59CE9512F61C8B34004B9" name="Brazil">BRAZIL</collectingCountry>
– 
<emphasis id="604E570B3E1D66EC050A101E5E42502D" bold="true">
<collectingRegion id="691B60F8459693157383967D0F30AC0D">Amazonas</collectingRegion>
</emphasis>
</materialsCitation>
• 
<materialsCitation id="C596DD577D8A061E0E358F1B5152210B" collectingDate="1969-11-08" collectionCode="A, CNC" collectorName="Jameson ML &amp; Campbell" country="Brazil" location="Eva" specimenCount="1" stateProvince="Parana">
1; 
<collectingRegion id="F61D637CC6346328D480D2686D4BD2B5" country="Brazil" name="Parana">Parana</collectingRegion>
da 
<location id="5EBBDF5FC2C89D28ACC225224EFE520B" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:5EBBDF5FC2C89D28ACC225224EFE520B" country="Brazil" name="Eva" stateProvince="Parana">Eva</location>
; 
<date id="1F4AC7AE4F0C1A5AADC40E0D430D96C7" value="1969-11-08">
<collectingDate id="4961A59397E60CF71440315A066CAEFA" value="1969-11-08">8 Nov. 1969</collectingDate>
</date>
; 
<collectorName id="F21EF3035214F6938EBC642146FE4A09" name="Jameson ML">J. M.</collectorName>
&amp; B. 
<collectionCode id="C7A360541499C4308F575536981BD3B8">A</collectionCode>
. 
<collectorName id="DE831129E126E66B8578732E0AAACEAF">Campbell</collectorName>
leg.; 
<named-content id="A0FBB658F861BDB889F1265E4758524F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="368F226CA005239B79DDC9229A05BDAE" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
. – 
<emphasis id="6FF498D99D04E89C46867106D47030F5" bold="true">
<collectingRegion id="A7627F64786029B29F793B78A70CDE61" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
</emphasis>
</materialsCitation>
• 
<materialsCitation id="5F6A384B081D57B69FAF82523A56111F" collectingDate="2022-10-01" collectionCode="CEMT" collectorName="K. R. Ferreira" country="Brazil" location="Juina" specimenCount="1" stateProvince="Amazonas">
1; 
<location id="4F819FB0167B2C8450793960ED8477F0" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:4F819FB0167B2C8450793960ED8477F0" country="Brazil" name="Juina" stateProvince="Amazonas">Juina</location>
; 
<date id="EA13D9AE731F9D3BD460A02BB68C9543" value="2022-10-01">
<collectingDate id="493493A2698F5C767EF9145806DD8ADB" value="2022-10-01">1 Oct. 2022</collectingDate>
</date>
; 
<collectorName id="2EC9E1B572BDE27BB1ED1FE92107C38A">K. R. Ferreira</collectorName>
leg; 
<abbrev id="ABBRID0EPJAE" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="4DB5D43D5F2552A68C934D9952648640">CEMT</collectionCode>
</abbrev>
</materialsCitation>
• 
<materialsCitation id="E9DFF68FFEF76BCE6BCAB59BC6AE24F7" collectingDate="2024-01-11" collectingDateMax="2024-01-17" collectingDateMin="2024-01-11" collectionCode="CEMT" collectorName="D. F. Rodrigues &amp; T. I. Vitor-da-Silva" country="Brazil" latitude="-9.761389" location="Faz. da Ilha" longLatPrecision="21" longitude="-52.26139" municipality="Santa Cruz do Xingu" specimenCount="1" stateProvince="Rio de Janeiro">
1; 
<collectingMunicipality id="9EA9DA32C7EED12DA116ECE21F406FE4">Santa Cruz do Xingu</collectingMunicipality>
, 
<location id="8BCF9027FAB8EF3E2B00AAD941214C59" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:8BCF9027FAB8EF3E2B00AAD941214C59" country="Brazil" latitude="-9.761389" longLatPrecision="21" longitude="-52.26139" municipality="Santa Cruz do Xingu" name="Faz. da Ilha" stateProvince="Rio de Janeiro">Faz. da Ilha</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWJAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-52.261389,-9.761389]}">
<geoCoordinate id="A87E4344BDBD7ACD0E134A8B63DDB42C" degrees="9" direction="south" minutes="45" orientation="latitude" precision="15" seconds="41" value="-9.761389">9 ° 45 ' 41 &quot; S</geoCoordinate>
, 
<geoCoordinate id="81083514922CE537C8ECF11BA81D3F5E" degrees="52" direction="west" minutes="15" orientation="longitude" precision="15" seconds="41" value="-52.26139">52 ° 15 ' 41 &quot; W</geoCoordinate>
</named-content>
; 
<collectingMethod id="95B49C19D90A5763E56F3ED60CDA6D91">UV light trap</collectingMethod>
; 
<date id="A7C32D1D6CC95E1834676A267FB6266C" value="2024-01-11" valueMax="2024-01-17" valueMin="2024-01-11">
<collectingDate id="00972538014D0C9CA055371648D6446A" value="2024-01-11" valueMax="2024-01-17" valueMin="2024-01-11">11–17 Jan. 2024</collectingDate>
</date>
; 
<collectorName id="442F052A9A0FD2B5C70FFC5112DBAEA9">D. F. Rodrigues</collectorName>
&amp; 
<collectorName id="09FB234272DF1CFB048E1690EFAF116D">T. I. Vitor-da-Silva</collectorName>
leg.; 
<abbrev id="ABBRID0E2JAE" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="B0008BB4687853B58A080868F1241F4A">CEMT</collectionCode>
</abbrev>
. – 
<emphasis id="836133B69DA0E4E041A1D82BBDC76C72" bold="true">
<collectingRegion id="76A87E0D415349E688BFC023FC53C646" country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
</emphasis>
</materialsCitation>
• 
<materialsCitation id="01D6DC5AB2E502C72F68CACC7343C1B6" collectingDate="1999-10-15" collectingDateMax="1999-11-10" collectingDateMin="1999-10-15" collectionCode="NHMUK" collectorName="W. Grosser" country="Brazil" location="Mage" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro">
<specimenCount id="1B8876D3B5554028EBF86FC9FC99AEED" count="1" type="female">1 ♀</specimenCount>
; 
<location id="9F97267D65E2569387024A491E0E2B8A" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:9F97267D65E2569387024A491E0E2B8A" country="Brazil" name="Mage" stateProvince="Rio de Janeiro">Magé</location>
, 
<quantity id="64542373FFE67E3B1F4F4A9E50F54B55" metricMagnitude="4" metricUnit="m" metricValue="7.0" unit="km" value="70.0">70 km</quantity>
N [north] of 
<collectingRegion id="D672974F6221E9DAF74F23D0F57ADF04" country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
; 
<date id="FC3ACC23E81456089A4A0BCF6463DA50" value="1999-10-15" valueMax="1999-11-10" valueMin="1999-10-15">
<collectingDate id="5B0BE7A3D73FEDD51F16B3BFE217542F" value="1999-10-15" valueMax="1999-11-10" valueMin="1999-10-15">15 Oct. – 10 Nov. 1999</collectingDate>
</date>
; 
<collectorName id="C2F1E24E107F6A5274508E8535E8C64F">W. Grosser</collectorName>
leg.; 
<named-content id="28E757B24B40347A777EC92E5344945E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="9061BB9BA0BE0D7303CF40186BE78BDF" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFKAE" type="notes">
<paragraph id="9C12FF3D4E641624810228FD12090448">
<heading id="5A727C8499411A245A21365875D79F37" reason="title">Note.</heading>
</paragraph>
<paragraph id="FA819C789A590BE073DE666F79E51A6B">
The monotypic subgenus 
<taxonomicName id="1191D229115F5DE0D32C73596305B57F" authority="Scholtz, 1986 a" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="030685FA542DF0CBB1862C6F7DC8B1F1" italics="true">Haroldomorgus</emphasis>
Scholtz, 1986 a
</taxonomicName>
was established to accommodate 
<taxonomicName id="98F2235B1AC6BB14C93D0F993887CFB4" authority="Harold, 1872" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="70E51C81F4AD5631F3D9F64D25299CAB" italics="true">Trox batesi</emphasis>
<bibRefCitation id="5C5200841EA701E055F503BF8B3E7DDE" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
</taxonomicName>
, following a morphological phylogenetic analysis aimed at elucidating the suprageneric relationships within 
<taxonomicName id="8988907B2F0F6D953645983D047FBE11" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
(refer to Scholtz, 1986 a for details). One of the most distinctive features of 
<taxonomicName id="86EC15DBF40EAAF49E49CF9D69B7EDA6" authorityName="Scholtz" authorityYear="1986" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="7D70C348A148E13282BDCDFF06666EEE" italics="true">Omorgus</emphasis>
(
<emphasis id="FBDB75CD4942CE3178DBD2FD8D4D80B7" italics="true">Haroldomorgus</emphasis>
)
</taxonomicName>
species is a complete absence of tubercles or ridges on the pronotum and elytra. Additionally, this species exhibits an oval-shaped scutellum that is not constricted at the base, which distinguishes it from the typical modern concept of 
<taxonomicName id="4BA3C87E50059230618065039D1EFB8E" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C6EA6C9FC187D5219D89A6629E13C3CB" italics="true">Omorgus</emphasis>
species.
</taxonomicName>
Following a morphological phylogenetic analysis conducted by some of the authors (
<collectionCode id="5E0C3343E9AE24BE74AB8488A083531F">VCS</collectionCode>
, WPS, and FZVM – 
<bibRefCitation id="DD2A7732C2543293748529819F10B38A" author="Costa-Silva V &amp; Strümpher WP &amp; Thyssen PJ &amp; Vaz-de-Mello FZ" refId="B28" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (in press) Phylogenetic systematics of the keratin-feeding genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Zoological Journal of the Linnean Society.">Costa-Silva et al. in press</bibRefCitation>
), the genus 
<taxonomicName id="7C26A7022E01C3DA8EC872C74C6DFE68" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0A17E395206F26D152CEA9A7DA8D8231" italics="true">Omorgus</emphasis>
</taxonomicName>
was recovered to be paraphyletic, restoring 
<taxonomicName id="EA2E9B6DF4C8965868285E5F7004844B" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="6C091A55D3620B9DBB34209FA0B3C5AA" italics="true">Haroldomorgus</emphasis>
</taxonomicName>
as a sister group to 
<taxonomicName id="64126C94FFD2526C398778F9E8859AB0" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="55D5EA04CAB9911D7C0FE5DAB8BC5A37" italics="true">Omorgus</emphasis>
</taxonomicName>
+ 
<taxonomicName id="2E8F67F4D087DEB75EC590E404056D07" authorityName="Burmeister" authorityYear="1876" class="Insecta" family="Trogidae" genus="Polynoncus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="8D52D92AB81C5C5AB439E1FF7CE71F48" italics="true">Polynoncus</emphasis>
</taxonomicName>
. In the first and only molecular phylogeny of 
<taxonomicName id="41FE7EDBD778203F8D6D605DA9E2EB0A" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
performed to date, by 
<bibRefCitation id="81E2070E34F893CCC367BFC8D0C71B0E" DOI="10.1111/syen.12074" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="3" journalOrPublisher="Systematic Entomology" pagination="548-562" refId="B104" refString="Strümpher WP, Sole CL, Villet MH, Scholtz CH (2014) Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. Systematic Entomology 39 (3): 548–562. https://doi.org/10.1111/syen.12074" title="Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data." volume="39" year="2014">Strümpher et al. (2014)</bibRefCitation>
, no specimens of 
<taxonomicName id="A84DF574A01CD9351EED5EB7C8E95C48" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="8AAF9D706FAD7BBE758A6FC92D638B6A" italics="true">Omorgus</emphasis>
(
<emphasis id="5D679551323FABE55EFFFC0A28EE723E" italics="true">Haroldomorgus</emphasis>
) 
<emphasis id="AA23D8D4B43B9864AABC1C2B127CC5CD" italics="true">batesi</emphasis>
</taxonomicName>
were used in the analysis. However, we have now obtained fresh specimens to revisit the molecular analysis and finally test the 
<bibRefCitation id="670856A99D9105E9BE9D8CABF78F7581" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz’s (1986 a</bibRefCitation>
) hypothesis and the position of 
<taxonomicName id="4A4535816AC1F4140677D82FC8544AB2" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="C0F7A454FF5BFD4A2C47A18FDED47DB8" italics="true">Haroldomorgus</emphasis>
</taxonomicName>
within 
<taxonomicName id="8D1B4137ED52B76F8DA9FDA49502A5E2" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>