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<document id="F66841BC24C444E0B69583AE69C7C25A" ID-DOI="10.11646/phytotaxa.437.4.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13873796" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727815446411" checkinUser="felipe" docAuthor="Penneckamp, Diego, Chaura, Rodrigo, Rojas, Gloria &amp; Stuessy, Tod" docDate="2020" docId="9B2E87F5FFCCF766FF5FF8BDFB758527" docLanguage="en" docName="phytotaxa.437.4.4.pdf" docOrigin="Phytotaxa 437 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.437.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Plectocephalus formosus Penneck. &amp; Gl. Rojas 2020, comb. nov." docType="treatment" docVersion="2" lastPageNumber="232" masterDocId="6717FF8DFFCFF763FFD7FFE9FFFE8056" masterDocTitle="Taxonomic notes on Plectocephalus (Centaurea s. l., Centaureinae, Asteraceae) from Chile, including new combinations and synonyms" masterLastPageNumber="236" masterPageNumber="227" pageNumber="230" updateTime="1727817782491" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="27101AF9AB7E3DB18D38A23E74CB1A56" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7EDBC12A2EBC29B589BA4934C95184FC">
<mods:title id="46442873B4DD7CD281D65D9C137FE0A6">Taxonomic notes on Plectocephalus (Centaurea s. l., Centaureinae, Asteraceae) from Chile, including new combinations and synonyms</mods:title>
</mods:titleInfo>
<mods:name id="B456D0CFED6AF4B96E8BB638E74B82C5" type="personal">
<mods:role id="B1D27C46B25F6AE04B97C7233FFAF20B">
<mods:roleTerm id="FF265923E8840EF4E34A335DCE940526">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="474595CB144470605FCEAA8C20E03E78">Penneckamp, Diego</mods:namePart>
<mods:affiliation id="199B9F23829E1AFBAF5E387598A32E65">Laboratorio de Biodiversidad y Ecología del Dosel, Facultad de Ciencias Forestales y Recursos Naturales, Universidad Austral de Chile, P. O. Box 567, Valdivia, Chile &amp; Museo Nacional de Historia Natural, Casilla 787, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="FDD2BDB778125004451D9304A1267F03" type="personal">
<mods:role id="D42060FB71B191F4848402D98F460D56">
<mods:roleTerm id="7D342B6426A94EDB318A39BC1E8B1D68">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5520BD2DA2FFF239940CC96E8C1047DE">Chaura, Rodrigo</mods:namePart>
<mods:affiliation id="64C84F0B86400C9D9919F4A9E63538DB">Laboratorio de Biodiversidad y Ecología del Dosel, Facultad de Ciencias Forestales y Recursos Naturales, Universidad Austral de Chile, P. O. Box 567, Valdivia, Chile &amp; Museo Nacional de Historia Natural, Casilla 787, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="4DA469242B650C80211443614667B955" type="personal">
<mods:role id="DBF665FB41403901D5E99A7B5072C52F">
<mods:roleTerm id="DBB8FC44250C3CA558C1969B8134A1B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44D8BDB92310932952D890C78C990152">Rojas, Gloria</mods:namePart>
<mods:affiliation id="139B54689B789B00F98A02E4493A429D">Museo Nacional de Historia Natural, Casilla 787, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="2A284B1CEBB2E027E1A3F9676E13C218" type="personal">
<mods:role id="68A858D1E709734E927EDEB7B512072E">
<mods:roleTerm id="C16D7A589B52C5922596340BCB8B4E62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F0DA09418D38E36638BBAD5D7C693B1">Stuessy, Tod</mods:namePart>
<mods:affiliation id="9A735D03F6BAB52F3BB528FF4137C992">Department of Botany and Biodiversity Research, University of Vienna, Rennweg 14, A- 1030 Vienna, Austria &amp; Herbarium and Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, 1315 Kinnear Avenue, Columbus, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="AF3FC17157E01A5927D4C181D7145A21">text</mods:typeOfResource>
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<mods:titleInfo id="84364C2450D841F852FFA67456569077">
<mods:title id="75E0CB918BCF1C86DAD9B725CDB6F3D3">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="3D68BB261904BA378A92BC89A709A5D9">
<mods:date id="B4021AE1BE69F0C3395851CF4D8B42F8">2020</mods:date>
<mods:detail id="FD92D8E925C2D8E6644906166FFD9393" type="pubDate">
<mods:number id="5724622CB5BE271A23F28FAF8236CA68">2020-03-30</mods:number>
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<mods:detail id="0F39627E3952FDE6B27854FDBC27A6EA" type="volume">
<mods:number id="80861657DD4320E8120365DA7A575C83">437</mods:number>
</mods:detail>
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<mods:number id="76FBB2B3E25450576985ABECF1F1ABD8">4</mods:number>
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<mods:start id="4F3984A9E2DF879651DDB338A6B445E6">227</mods:start>
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<mods:url id="E175B402863881B2F59DCE9F10D593FB">http://dx.doi.org/10.11646/phytotaxa.437.4.4</mods:url>
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<mods:classification id="7D21EE466040CE1939FE04033FB030C8">journal article</mods:classification>
<mods:identifier id="AE37EEBC3AF66333500D1DCA849256CF" type="DOI">10.11646/phytotaxa.437.4.4</mods:identifier>
<mods:identifier id="AC3955AFC24E97C6BBE85062205EDAC6" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="DF0DAD81638FDA81FC504E49F0EF9C42" type="Zenodo-Dep">13873796</mods:identifier>
</mods:mods>
<treatment id="9B2E87F5FFCCF766FF5FF8BDFB758527" ID-DOI="http://doi.org/10.5281/zenodo.13873866" ID-Zenodo-Dep="13873866" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527" httpUri="http://treatment.plazi.org/id/9B2E87F5FFCCF766FF5FF8BDFB758527" lastPageId="5" lastPageNumber="232" pageId="3" pageNumber="230">
<subSubSection id="5B9D6568FFCCF760FF5FF8BDFBD68738" box="[136,1064,1876,1903]" pageId="3" pageNumber="230" type="nomenclature">
<paragraph id="133836E3FFCCF760FF5FF8BDFBD68738" blockId="3.[136,1064,1876,1903]" box="[136,1064,1876,1903]" pageId="3" pageNumber="230">
<heading id="4870818FFFCCF760FF5FF8BDFBD68738" box="[136,1064,1876,1903]" fontSize="11" level="2" pageId="3" pageNumber="230" reason="3">
<taxonomicName id="D4874D60FFCCF760FF5FF8BDFCDC8738" ID-CoL="4K3C4" authority="(Ravenna) Penneck. &amp; Gl. Rojas" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" box="[136,802,1876,1903]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosus" status="comb. nov.">
<emphasis id="21F3EAF1FFCCF760FF5FF8BDFE568738" bold="true" box="[136,424,1876,1903]" italics="true" pageId="3" pageNumber="230">Plectocephalus formosus</emphasis>
(Ravenna) Penneck. &amp; Gl. Rojas
</taxonomicName>
,
<emphasis id="21F3EAF1FFCCF760FCFAF8BDFC568738" bold="true" box="[813,936,1876,1903]" italics="true" pageId="3" pageNumber="230">
<taxonomicNameLabel id="3AC0578AFFCCF760FCFAF8BDFC568738" box="[813,936,1876,1903]" pageId="3" pageNumber="230" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="8BBC2A66FFCCF760FC60F8BDFBE98738" box="[951,1047,1876,1902]" captionStart="FIGURE 3" captionStartId="5.[136,229,1081,1103]" captionTargetBox="[151,1435,245,1057]" captionTargetId="figure-52@5.[151,1435,245,1057]" captionTargetPageId="5" captionText="FIGURE 3. Plectocephalus formosus: A, type specimen; B, living plant in natural habitat, photograph by Gloria Rojas." figureDoi="http://doi.org/10.5281/zenodo.13873802" httpUri="https://zenodo.org/record/13873802/files/figure.png" pageId="3" pageNumber="230">Figure 3</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5B9D6568FFCCF767FF5FF849FC3E8173" lastPageId="4" lastPageNumber="231" pageId="3" pageNumber="230" type="reference_group">
<paragraph id="133836E3FFCCF760FF5FF849FBE28877" blockId="3.[136,1453,1952,2082]" pageId="3" pageNumber="230">
<materialsCitation id="A3EF3CBEFFCCF760FF5FF849FBE28877" collectingDate="1991-10-03" collectingDateMax="2003-10" collectingDateMin="1991-10-03" collectionCode="P, BA, CONC" collectorName="El Arrayan &amp; Ch. von Bohlen" country="Chile" county="Basionym" elevation="135" latitude="-29.65" location="Elqui Province" longLatPrecision="1226" longitude="-71.3" municipality="Type" pageId="3" pageNumber="230" specimenCode="SGO 133272" specimenCount="4" stateProvince="Coquimbo" typeStatus="holotype">
<collectingCounty id="FA594E6FFFCCF760FF5FF849FF1287E0" box="[136,236,1952,1974]" pageId="3" pageNumber="230">Basionym</collectingCounty>
:—
<taxonomicName id="D4874D60FFCCF760FEDFF849FD5787E3" ID-CoL="S6KC" authority="Ravenna (2006: 48 - 49)" authorityName="Ravenna" authorityPageNumber="48 - 49" authorityYear="2006" box="[264,681,1952,1974]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosa">
<emphasis id="21F3EAF1FFCCF760FEDFF849FE3F87E3" box="[264,449,1952,1973]" italics="true" pageId="3" pageNumber="230">Centaurea formosa</emphasis>
<bibRefCitation id="77164B12FFCCF760FE12F849FD5787E3" author="Ravenna, P." box="[453,681,1952,1974]" pageId="3" pageNumber="230" pagination="48 - 49" refId="ref6925" refString="Ravenna, P. (2006) Centaurea formosa and C. huascoensis, two new species of Chilean Asteraceae. Onira (10) 15: 48 - 49." type="journal article" year="2006">Ravenna (2006: 48–49)</bibRefCitation>
</taxonomicName>
.
<collectingMunicipality id="F35CAC99FFCCF760FD66F849FD1A87E3" box="[689,740,1952,1973]" pageId="3" pageNumber="230">Type</collectingMunicipality>
:—
<collectingCountry id="6B907673FFCCF760FCD7F849FCB487E3" box="[768,842,1952,1973]" name="Chile" pageId="3" pageNumber="230">CHILE</collectingCountry>
:
<collectingRegion id="D143F801FFCCF760FC86F849FBFA87E0" box="[849,1028,1952,1974]" country="Chile" name="Coquimbo" pageId="3" pageNumber="230">Coquimbo Region</collectingRegion>
,
<location id="16586038FFCCF760FBDBF849FB6387E0" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FBDBF849FB6387E0" box="[1036,1181,1952,1974]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="Elqui Province" pageId="3" pageNumber="230" stateProvince="Coquimbo">Elqui Province</location>
, bushy field near
<collectorName id="BE725335FFCCF760FA93F849FA5287E0" box="[1348,1452,1952,1974]" pageId="3" pageNumber="230">
<location id="16586038FFCCF760FA93F849FA5287E0" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FA93F849FA5287E0" box="[1348,1452,1952,1974]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="El Arrayan" pageId="3" pageNumber="230" stateProvince="Coquimbo">El Arrayán</location>
</collectorName>
about
<quantity id="D47F9B06FFCCF760FF2FF82DFECA878C" box="[248,308,1988,2010]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="230" unit="km" value="25.0">25 km</quantity>
north from
<location id="16586038FFCCF760FE73F82DFDF8878F" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FE73F82DFDF8878F" box="[420,518,1988,2009]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="La Serena" pageId="3" pageNumber="230" stateProvince="Coquimbo">La Serena</location>
,
<date id="67391023FFCCF760FDD8F82DFD6D878F" box="[527,659,1988,2010]" pageId="3" pageNumber="230" value="2003-10">
<collectingDate id="777DE9CBFFCCF760FDD8F82DFD6D878F" box="[527,659,1988,2010]" pageId="3" pageNumber="230" value="2003-10">October 2003</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCCF760FD4BF82DFCC2878F" box="[668,828,1988,2009]" italics="true" pageId="3" pageNumber="230">
<collectionCode id="7596AE26FFCCF760FD4BF82DFD55878F" box="[668,683,1988,2009]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="230" type="Herbarium">P</collectionCode>
.
<location id="16586038FFCCF760FD64F82DFCF9878F" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FD64F82DFCF9878F" box="[691,775,1988,2009]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="Ravenna" pageId="3" pageNumber="230" stateProvince="Coquimbo">Ravenna</location>
5023
</emphasis>
(
<typeStatus id="CC3C8841FFCCF760FC90F82DFC5C878C" box="[839,930,1988,2010]" pageId="3" pageNumber="230" type="holotype">Holotype</typeStatus>
<collectionCode id="7596AE26FFCCF760FC70F82DFC35878F" box="[935,971,1988,2009]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="230" type="Herbarium">BA</collectionCode>
;
<typeStatus id="CC3C8841FFCCF760FC03F82DFBDD878C" box="[980,1059,1988,2010]" pageId="3" pageNumber="230" type="isotype">isotypes</typeStatus>
Herb. Rav.,
<collectionCode id="7596AE26FFCCF760FB4EF82DFB1E878F" box="[1177,1248,1988,2009]" country="Chile" httpUri="http://grbio.org/cool/mhpx-qnxz" name="Universidad de Concepción" pageId="3" pageNumber="230" type="Herbarium">CONC</collectionCode>
).
<typeStatus id="CC3C8841FFCCF760FB38F82DFABD878F" box="[1263,1347,1988,2009]" pageId="3" pageNumber="230" type="neotype">Neotype</typeStatus>
:—
<collectingCountry id="6B907673FFCCF760FA88F82DFA56878F" box="[1375,1448,1988,2009]" name="Chile" pageId="3" pageNumber="230">CHILE</collectingCountry>
.
<collectingRegion id="D143F801FFCCF760FF6AF801FE8F87A8" box="[189,369,2024,2046]" country="Chile" name="Coquimbo" pageId="3" pageNumber="230">Coquimbo Region</collectingRegion>
:
<location id="16586038FFCCF760FEAEF801FDF487A8" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FEAEF801FDF487A8" box="[377,522,2024,2046]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="Elqui Province" pageId="3" pageNumber="230" stateProvince="Coquimbo">Elqui Province</location>
,
<location id="16586038FFCCF760FDC5F801FD2987AB" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FDC5F801FD2987AB" box="[530,727,2024,2046]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="Panamericana Norte" pageId="3" pageNumber="230" stateProvince="Coquimbo">Panamericana Norte</location>
,
<locationDeviation id="028A2FC1FFCCF760FD08F801FBB287AB" box="[735,1100,2024,2046]" firstLocation="La Serena" pageId="3" pageNumber="230" secondLocation="Caleta Hornos">between La Serena and Caleta Hornos</locationDeviation>
, before
<location id="16586038FFCCF760FB40F801FAE987A8" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCCF760FB40F801FAE987A8" box="[1175,1303,2024,2046]" country="Chile" county="Basionym" latitude="-29.65" longLatPrecision="1226" longitude="-71.3" municipality="Type" name="Juan Soldado" pageId="3" pageNumber="230" stateProvince="Coquimbo">Juan Soldado</location>
bridge, (
<geoCoordinate id="76B35024FFCCF760FABCF801FF328877" degrees="29" direction="south" minutes="39" orientation="latitude" pageId="3" pageNumber="230" precision="925" value="-29.65">29°39’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCCF760FF00F7E5FEC88877" box="[215,310,2060,2081]" degrees="71" direction="west" minutes="18" orientation="longitude" pageId="3" pageNumber="230" precision="925" value="-71.3">71°18’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCCF760FE90F7E5FE548877" box="[327,426,2060,2081]" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.9" metricValueMin="0.8" pageId="3" pageNumber="230" unit="m" value="135.0" valueMax="190.0" valueMin="80.0">
<elevation id="98AAD1D0FFCCF760FE90F7E5FE548877" box="[327,426,2060,2081]" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.9" metricValueMin="0.8" pageId="3" pageNumber="230" unit="m" value="135.0" valueMax="190.0" valueMin="80.0">80–190 m</elevation>
</quantity>
,
<date id="67391023FFCCF760FE64F7E5FDB58877" box="[435,587,2060,2082]" pageId="3" pageNumber="230" value="1991-10-03">
<collectingDate id="777DE9CBFFCCF760FE64F7E5FDB58877" box="[435,587,2060,2082]" pageId="3" pageNumber="230" value="1991-10-03">3 October 1991</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCCF760FD82F7E5FCE18877" box="[597,799,2060,2081]" italics="true" pageId="3" pageNumber="230">
<collectorName id="BE725335FFCCF760FD82F7E5FD178877" box="[597,745,2060,2081]" pageId="3" pageNumber="230">Ch. von Bohlen</collectorName>
1209
</emphasis>
(
<typeStatus id="CC3C8841FFCCF760FCFBF7E5FC7C8874" box="[812,898,2060,2082]" pageId="3" pageNumber="230" type="holotype">holotype</typeStatus>
<specimenCode id="43219E98FFCCF760FC50F7E5FBF08877" box="[903,1038,2060,2081]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="3" pageNumber="230" type="Herbarium">SGO 133272</specimenCode>
!).
</materialsCitation>
</paragraph>
<paragraph id="133836E3FFCBF767FF5FFF77FC3E8173" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="231">
Note:—Although our efforts to locate the
<typeStatus id="CC3C8841FFCBF767FDB4FF76FD6D80EF" box="[611,659,159,185]" pageId="4" pageNumber="231">type</typeStatus>
material in the mentioned institutions have been unfruitful, we accept this as a distinct species based on the detailed description. However the
<typeStatus id="CC3C8841FFCBF767FC04FF2AFBFD808B" box="[979,1027,195,221]" pageId="4" pageNumber="231">type</typeStatus>
material do not exist at the herbaria cited (BA, CONC, pers. comm.) and the P. Ravenna´s personal herbarium was destroyed (Ravenna’s daughter, pers. comm.), so here we designate a new
<typeStatus id="CC3C8841FFCBF767FDF2FEE2FDAB8173" box="[549,597,267,293]" pageId="4" pageNumber="231">type</typeStatus>
specimen for this species name.
</paragraph>
</subSubSection>
<subSubSection id="5B9D6568FFCBF767FF6AFEC7FDCE8317" pageId="4" pageNumber="231" type="description">
<paragraph id="133836E3FFCBF767FF6AFEC7FDCE8317" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="231">
Shrub ca.
<quantity id="D47F9B06FFCBF767FEFAFEC7FE76811F" box="[301,392,302,329]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="231" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</quantity>
tall, branched, lignified at base, perennial. Stems glabrous to glabrescent, striated toward apex. Leaves pinnately-parted to pinnatisect, (2–)2.5–9 ×
<quantity id="D47F9B06FFCBF767FD35FEBBFC9B813B" box="[738,869,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.6" metricValueMin="0.7" pageId="4" pageNumber="231" unit="cm" value="1.65" valueMax="2.6" valueMin="0.7">0.7–2.6 cm</quantity>
, lamina glabrous to glabrescent at adaxial surface and puberulent at abaxial surface with revolute margin. Capitulum borne singly on a glabrescent, striate peduncle, accompanied by some reduced leaves in form of pinnatifid bracts. Involucre globose, hemispheric-campanulate, large, (2–)2.5–3.5(–4) cm diam., woolly-pilose at base. Phyllaries many in 5–7 series. Internal phyllaries of first series sparsely woolly-pilose on adaxial surface at the base and center, becoming glabrous toward margins or rarely glabrescent. External series phyllaries shorter and wider, subcordate, ovate to ovate-lanceolate with acute apex, 5–10 ×
<quantity id="D47F9B06FFCBF767FF5FFDC3FF198213" box="[136,231,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="231" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
, inner phyllaries longer and narrowly lanceolate, linear-lanceolate to subspathulate, 25 ×
<quantity id="D47F9B06FFCBF767FB04FDC3FACC8213" box="[1235,1330,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="4" pageNumber="231" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
, with apex expanded into an erect spreading pectinate-fringed coriaceous appendage, brownish, being c. 1/3 to equaling inner phyllary length or even external phyllary length. Phyllaries with fimbriate appendage with a long mucro at apex. Ray florets sterile, 3.5–5 ×
<quantity id="D47F9B06FFCBF767FE5DFD7FFDF582E7" box="[394,523,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="4" pageNumber="231" unit="cm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 cm</quantity>
, corolla 5-lobed, lobes lanceolate with apex acute, 2.5 ×
<quantity id="D47F9B06FFCBF767FB45FD7FFAED82E7" box="[1170,1299,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="231" unit="cm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 cm</quantity>
, tube
<quantity id="D47F9B06FFCBF767FA80FD7FFF548283" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="4" pageNumber="231" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 cm</quantity>
long. Disc florets tubular, smaller than ray florets, fertile and hermaphroditic, 2–2.3 ×
<quantity id="D47F9B06FFCBF767FBA7FD53FB0E8283" box="[1136,1264,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="231" unit="cm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 cm</quantity>
, corolla 5-lobed, lobes acute to linear-lanceolate separated at least 1/5 to 1/4 of length. Corollas very showy, ligules pink to whitish, disc floret corollas whitish-yellow. Style
<quantity id="D47F9B06FFCBF767FDC9FCEBFD76834B" box="[542,648,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="4" pageNumber="231" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
with bilobed stigma, stigma branches ca.
<quantity id="D47F9B06FFCBF767FB8CFCEBFB60834B" box="[1115,1182,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="231" unit="mm" value="3.0">3 mm</quantity>
. Anthers ca.
<quantity id="D47F9B06FFCBF767FAFBFCEAFA93834B" box="[1324,1389,771,797]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="231" unit="mm" value="7.0">7 mm</quantity>
long. Pappus and mature cypselae not seen.
</paragraph>
</subSubSection>
<caption id="47F8666BFFCBF767FF5FF9E9FED7866C" ID-DOI="http://doi.org/10.5281/zenodo.13873800" ID-Zenodo-Dep="13873800" httpUri="https://zenodo.org/record/13873800/files/figure.png" pageId="4" pageNumber="231" startId="4.[136,229,1536,1558]" targetBox="[145,1442,854,1508]" targetPageId="4" targetType="figure">
<paragraph id="133836E3FFCBF767FF5FF9E9FED7866C" blockId="4.[136,1451,1536,1594]" pageId="4" pageNumber="231">
<emphasis id="21F3EAF1FFCBF767FF5FF9E9FF008643" bold="true" box="[136,254,1536,1558]" pageId="4" pageNumber="231">FIGURE 2.</emphasis>
<taxonomicName id="D4874D60FFCBF767FED2F9E9FD798643" authorityName="Penneck. &amp; Chaura" authorityYear="2020" baseAuthorityName="Penneck. &amp; Chaura" baseAuthorityYear="2020" box="[261,647,1536,1558]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="variety" species="chilensis" variety="brevilobus">
<emphasis id="21F3EAF1FFCBF767FED2F9E9FE0C8643" box="[261,498,1536,1557]" italics="true" pageId="4" pageNumber="231">Plectocephalus chilensis</emphasis>
var.
<emphasis id="21F3EAF1FFCBF767FDF4F9E9FD798643" box="[547,647,1536,1557]" italics="true" pageId="4" pageNumber="231">brevilobus</emphasis>
</taxonomicName>
: A, living plant in natural habitat; B, detail of flowering capitulum. Photograph by Rodrigo Chaura.
</paragraph>
</caption>
<subSubSection id="5B9D6568FFCBF767FF6AF980FC8786F1" pageId="4" pageNumber="231" type="etymology">
<paragraph id="133836E3FFCBF767FF6AF980FC8786F1" blockId="4.[136,1452,1640,2063]" pageId="4" pageNumber="231">
<emphasis id="21F3EAF1FFCBF767FF6AF980FEC086D5" bold="true" box="[189,318,1641,1667]" pageId="4" pageNumber="231">Etymology</emphasis>
:—From the Latin,
<taxonomicName id="D4874D60FFCBF767FDFFF980FD7186D5" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" box="[552,655,1641,1667]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="formosus">
<emphasis id="21F3EAF1FFCBF767FDFFF980FD7186D5" box="[552,655,1641,1667]" italics="true" pageId="4" pageNumber="231">formosus</emphasis>
</taxonomicName>
, meaning beautiful, because this species has large and very showy ornamental flowers, having one of the largest capitula in the genus.
</paragraph>
</subSubSection>
<subSubSection id="5B9D6568FFCBF766FF6AF959FB758527" lastPageId="5" lastPageNumber="232" pageId="4" pageNumber="231" type="distribution">
<paragraph id="133836E3FFCBF767FF6AF959FD078744" blockId="4.[136,1452,1640,2063]" pageId="4" pageNumber="231">
<emphasis id="21F3EAF1FFCBF767FF6AF959FE16869D" bold="true" box="[189,488,1712,1739]" pageId="4" pageNumber="231">Distribution and ecology</emphasis>
:—Restricted to sclerophyllous shrublands in coastal areas of Juan Soldado north of La Serena,
<collectingRegion id="D143F801FFCBF767FEDDF93DFE2486B8" box="[266,474,1748,1774]" country="Chile" name="Coquimbo" pageId="4" pageNumber="231">Coquimbo Region</collectingRegion>
(ca.
<geoCoordinate id="76B35024FFCBF767FDC2F93CFD3886B9" box="[533,710,1749,1775]" degrees="29" direction="south" minutes="39" orientation="latitude" pageId="4" pageNumber="231" precision="1" seconds="31.00" value="-29.658611">29°39’31.00” S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCBF767FD02F93CFC6F86B9" box="[725,913,1749,1775]" degrees="71" direction="west" minutes="18" orientation="longitude" pageId="4" pageNumber="231" precision="1" seconds="42.00" value="-71.31167">71°18’42.00” W</geoCoordinate>
) and in the range of Chañaral de Aceituno (ca.
<geoCoordinate id="76B35024FFCBF767FF5FF911FED48745" box="[136,298,1784,1811]" degrees="29" direction="south" minutes="3" orientation="latitude" pageId="4" pageNumber="231" precision="1" seconds="60.00" value="-29.066666">29°3’60.00” S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCBF767FEE0F910FE1D8745" box="[311,483,1785,1811]" degrees="71" direction="west" minutes="29" orientation="longitude" pageId="4" pageNumber="231" precision="1" seconds="0.00" value="-71.48333">71°29’0.00” W</geoCoordinate>
) in the
<collectingRegion id="D143F801FFCBF767FDE1F910FD088744" box="[566,758,1784,1811]" country="Chile" name="Atacama" pageId="4" pageNumber="231">Atacama Region</collectingRegion>
.
</paragraph>
<paragraph id="133836E3FFCBF767FF6AF8F4FBFE87F4" blockId="4.[136,1452,1640,2063]" pageId="4" pageNumber="231">
<emphasis id="21F3EAF1FFCBF767FF6AF8F4FEFE8761" bold="true" box="[189,256,1821,1847]" pageId="4" pageNumber="231">Notes</emphasis>
:—This species was proposed by
<bibRefCitation id="77164B12FFCBF767FD5DF8F4FCBC8760" author="Ravenna, P." box="[650,834,1820,1847]" pageId="4" pageNumber="231" pagination="48 - 49" refId="ref6925" refString="Ravenna, P. (2006) Centaurea formosa and C. huascoensis, two new species of Chilean Asteraceae. Onira (10) 15: 48 - 49." type="journal article" year="2006">Ravenna (2006)</bibRefCitation>
in a little-known journal, but nonetheless effectively published.The name was also validly published because the author provided a description,
<typeStatus id="CC3C8841FFCBF767FBBBF8A8FB62870D" box="[1132,1180,1857,1883]" pageId="4" pageNumber="231">type</typeStatus>
specimen, and indicated the institution of deposit. Since the species appears to be a good one, and for clarity with regard to the other species of
<taxonomicName id="D4874D60FFCBF767FF5FF861FECD87F4" box="[136,307,1928,1954]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="genus">
<emphasis id="21F3EAF1FFCBF767FF5FF861FECD87F4" box="[136,307,1928,1954]" italics="true" pageId="4" pageNumber="231">Plectocephalus</emphasis>
</taxonomicName>
in
<collectingCountry id="6B907673FFCBF767FE80F861FE6887F4" box="[343,406,1928,1954]" name="Chile" pageId="4" pageNumber="231">Chile</collectingCountry>
, we provide a new comprehensive description and key.
</paragraph>
<paragraph id="133836E3FFCBF767FF6AF845FA6187BC" blockId="4.[136,1452,1640,2063]" pageId="4" pageNumber="231">
<taxonomicName id="D4874D60FFCBF767FF6AF845FE2A8791" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" box="[189,468,1964,1991]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="formosus">
<emphasis id="21F3EAF1FFCBF767FF6AF845FE2A8791" box="[189,468,1964,1991]" italics="true" pageId="4" pageNumber="231">Plectocephalus formosus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="D4874D60FFCBF767FD9BF844FD338790" box="[588,717,1964,1990]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="floccosus">
<emphasis id="21F3EAF1FFCBF767FD9BF844FD338790" box="[588,717,1964,1990]" italics="true" pageId="4" pageNumber="231">P.floccosus</emphasis>
</taxonomicName>
, which differs by lacking woolly-tomentose stems.
<taxonomicName id="D4874D60FFCBF767FAD7F845FF1187BD" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="formosus">
<emphasis id="21F3EAF1FFCBF767FAD7F845FF1187BD" italics="true" pageId="4" pageNumber="231">Plectocephalus formosus</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="D4874D60FFCBF767FE62F838FDCD87BC" box="[437,563,2000,2026]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="21F3EAF1FFCBF767FE62F838FDCD87BC" box="[437,563,2000,2026]" italics="true" pageId="4" pageNumber="231">P. chilensis</emphasis>
</taxonomicName>
, from which the former differs in the abaxial puberulent surface of the leaves.
</paragraph>
<paragraph id="133836E3FFCBF766FF5FF81CFB07808A" blockId="4.[136,1452,1640,2063]" lastBlockId="5.[136,1452,158,221]" lastPageId="5" lastPageNumber="232" pageId="4" pageNumber="231">
One might speculate that based on intermediate morphological features and distribution,
<taxonomicName id="D4874D60FFCBF767FB4EF81CFAE18859" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" box="[1177,1311,2037,2063]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="formosus">
<emphasis id="21F3EAF1FFCBF767FB4EF81CFAE18859" box="[1177,1311,2037,2063]" italics="true" pageId="4" pageNumber="231">P. formosus</emphasis>
</taxonomicName>
might have originated through hybridization between
<taxonomicName id="D4874D60FFCAF766FD8CFF76FD2680EE" box="[603,728,158,184]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="21F3EAF1FFCAF766FD8CFF76FD2680EE" box="[603,728,158,184]" italics="true" pageId="5" pageNumber="232">P. chilensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D4874D60FFCAF766FCDBFF76FC7080EE" box="[780,910,158,184]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="species" species="floccosus">
<emphasis id="21F3EAF1FFCAF766FCDBFF76FC7080EE" box="[780,910,158,184]" italics="true" pageId="5" pageNumber="232">P. floccosus</emphasis>
</taxonomicName>
. This might be an example of a recent speciation radiation that occurred in this region of the distribution of
<taxonomicName id="D4874D60FFCAF766FCC2FF2BFC3E808A" box="[789,960,194,220]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="genus">
<emphasis id="21F3EAF1FFCAF766FCC2FF2BFC3E808A" box="[789,960,194,220]" italics="true" pageId="5" pageNumber="232">Plectocephalus</emphasis>
</taxonomicName>
(A. Susanna, pers. comm.).
</paragraph>
<caption id="47F8666BFFCAF766FF5FFBD0FAFB8406" ID-DOI="http://doi.org/10.5281/zenodo.13873802" ID-Zenodo-Dep="13873802" box="[136,1285,1081,1104]" httpUri="https://zenodo.org/record/13873802/files/figure.png" pageId="5" pageNumber="232" startId="5.[136,229,1081,1103]" targetBox="[151,1435,245,1057]" targetPageId="5" targetType="figure">
<paragraph id="133836E3FFCAF766FF5FFBD0FAFB8406" blockId="5.[136,1285,1081,1104]" box="[136,1285,1081,1104]" pageId="5" pageNumber="232">
<emphasis id="21F3EAF1FFCAF766FF5FFBD0FF008418" bold="true" box="[136,254,1081,1103]" pageId="5" pageNumber="232">FIGURE 3.</emphasis>
<taxonomicName id="D4874D60FFCAF766FED4FBD3FE0A8419" authorityName="Penneck. &amp; Gl. Rojas" authorityYear="2020" baseAuthorityName="Penneck. &amp; Gl. Rojas" baseAuthorityYear="2020" box="[259,500,1082,1103]" class="Magnoliopsida" family="Asteraceae" genus="Plectocephalus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="species" species="formosus">
<emphasis id="21F3EAF1FFCAF766FED4FBD3FE0A8419" box="[259,500,1082,1103]" italics="true" pageId="5" pageNumber="232">Plectocephalus formosus</emphasis>
</taxonomicName>
: A, type specimen; B, living plant in natural habitat, photograph by Gloria Rojas.
</paragraph>
</caption>
<paragraph id="133836E3FFCAF766FF6AFB97FB758527" blockId="5.[136,1452,1150,1393]" pageId="5" pageNumber="232">
<emphasis id="21F3EAF1FFCAF766FF6AFB97FE3C84CE" bold="true" box="[189,450,1150,1176]" pageId="5" pageNumber="232">Specimens examined:</emphasis>
—
<materialsCitation id="A3EF3CBEFFCAF766FE37FB97FB8A84EA" collectingDate="1949-09-08" collectionCode="W" collectorName="de Domeyko &amp; Biese" country="Chile" elevation="400" latitude="-29.116667" location="Chanar de Aceitunas" longLatPrecision="1228" longitude="-71.416664" municipality="Huasco Province" pageId="5" pageNumber="232" specimenCode="SGO 104196" specimenCount="1" stateProvince="Atacama">
<collectingCountry id="6B907673FFCAF766FE37FB97FDCB84CE" box="[480,565,1150,1176]" name="Chile" pageId="5" pageNumber="232">CHILE</collectingCountry>
.
<emphasis id="21F3EAF1FFCAF766FD95FB97FCEB84CE" bold="true" box="[578,789,1150,1176]" pageId="5" pageNumber="232">
<collectingRegion id="D143F801FFCAF766FD95FB97FCEE84CE" box="[578,784,1150,1176]" country="Chile" name="Atacama" pageId="5" pageNumber="232">Atacama Region</collectingRegion>
:
</emphasis>
<collectingMunicipality id="F35CAC99FFCAF766FCF7FB96FC1884CE" box="[800,998,1150,1177]" pageId="5" pageNumber="232">Huasco Province</collectingMunicipality>
:
<location id="16586038FFCAF766FC23FB97FB1884CE" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCAF766FC23FB97FB1884CE" box="[1012,1254,1150,1176]" country="Chile" latitude="-29.116667" longLatPrecision="1228" longitude="-71.416664" municipality="Huasco Province" name="Chanar de Aceitunas" pageId="5" pageNumber="232" stateProvince="Atacama">Chañar de Aceitunas</location>
,
<quantity id="D47F9B06FFCAF766FB23FB97FAC184CE" box="[1268,1343,1150,1176]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="232" unit="km" value="60.0">60 km</quantity>
al
<collectionCode id="7596AE26FFCAF766FABFFB96FA7A84CF" box="[1384,1412,1151,1177]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="232" type="Herbarium">W</collectionCode>
.
<collectorName id="BE725335FFCAF766FA46FB97FF0884EA" pageId="5" pageNumber="232">de Domeyko</collectorName>
(
<geoCoordinate id="76B35024FFCAF766FEDEFB4BFE9E84EA" box="[265,352,1186,1212]" degrees="29" direction="south" minutes="7" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.116667">29°7’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FEB8FB4BFE1E84EA" box="[367,480,1186,1213]" degrees="71" direction="west" minutes="25" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.416664">71°25’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FE20FB4BFD8484EB" box="[503,634,1186,1213]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="232" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="98AAD1D0FFCAF766FE20FB4BFD8484EB" box="[503,634,1186,1213]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="232" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300–500 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FD5FFB4BFCF084EA" box="[648,782,1186,1212]" pageId="5" pageNumber="232" value="1949-09-08">
<collectingDate id="777DE9CBFFCAF766FD5FFB4BFCF084EA" box="[648,782,1186,1212]" pageId="5" pageNumber="232" value="1949-09-08">8 Sep 1949</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FCCCFB4AFC4184EA" box="[795,959,1186,1212]" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FCCCFB4AFCCD84EA" box="[795,819,1187,1212]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="232" type="Herbarium">W</collectionCode>
.
<collectorName id="BE725335FFCAF766FC96FB4AFC8084EA" box="[833,894,1187,1212]" pageId="5" pageNumber="232">Biese</collectorName>
2544
</emphasis>
(
<specimenCode id="43219E98FFCAF766FC07FB4BFB9284EA" box="[976,1132,1186,1212]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 104196</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="A3EF3CBEFFCAF766FB57FB4BFB2A84B7" collectingDate="1965-10-12" collectionCode="C" collectorName="Munoz" country="Chile" elevation="100" latitude="-29.8" location="Juan Soldado" longLatPrecision="1225" longitude="-71.25" municipality="Elqui Province" pageId="5" pageNumber="232" specimenCode="SGO 135306" specimenCount="1" stateProvince="Coquimbo">
<emphasis id="21F3EAF1FFCAF766FB57FB4BFA9B84EA" bold="true" box="[1152,1381,1186,1212]" pageId="5" pageNumber="232">
<collectingRegion id="D143F801FFCAF766FB57FB4BFA9E84EA" box="[1152,1376,1186,1212]" country="Chile" name="Coquimbo" pageId="5" pageNumber="232">Coquimbo Region</collectingRegion>
:
</emphasis>
<collectingMunicipality id="F35CAC99FFCAF766FAB8FB4BFF0E84B6" pageId="5" pageNumber="232">Elqui Province</collectingMunicipality>
:
<location id="16586038FFCAF766FF2BFB2EFE6D84B6" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCAF766FF2BFB2EFE6D84B6" box="[252,403,1222,1249]" country="Chile" latitude="-29.8" longLatPrecision="1225" longitude="-71.25" municipality="Elqui Province" name="Juan Soldado" pageId="5" pageNumber="232" stateProvince="Coquimbo">Juan Soldado</location>
(
<geoCoordinate id="76B35024FFCAF766FE73FB2FFDF684B6" box="[420,520,1222,1248]" degrees="29" direction="south" minutes="48" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.8">29°48’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FDC1FB2FFD7884B6" box="[534,646,1222,1249]" degrees="71" direction="west" minutes="15" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.25">71°15’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FD4CFB2FFD1884B7" box="[667,742,1222,1249]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="232" unit="m" value="100.0">
<elevation id="98AAD1D0FFCAF766FD4CFB2FFD1884B7" box="[667,742,1222,1249]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="232" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FD24FB2FFC7C84B7" box="[755,898,1222,1249]" pageId="5" pageNumber="232" value="1965-10-12">
<collectingDate id="777DE9CBFFCAF766FD24FB2FFC7C84B7" box="[755,898,1222,1249]" pageId="5" pageNumber="232" value="1965-10-12">12 Oct 1965</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FC58FB2FFBDF84B6" box="[911,1057,1222,1248]" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FC58FB2FFC5B84B6" box="[911,933,1222,1248]" country="Denmark" name="University of Copenhagen" pageId="5" pageNumber="232" type="Herbarium">C</collectionCode>
.
<collectorName id="BE725335FFCAF766FC66FB2EFC0384B6" box="[945,1021,1223,1248]" pageId="5" pageNumber="232">Muñoz</collectorName>
48
</emphasis>
(
<specimenCode id="43219E98FFCAF766FBE6FB2FFB3284B7" box="[1073,1228,1222,1249]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 135306</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="A3EF3CBEFFCAF766FB37FB2FFECC857E" collectingDate="1957-09-17" collectionCode="N, C" collectorName="Munoz" country="Chile" elevation="110" latitude="-29.8" location="Quebrada de Juan Soldado" longLatPrecision="1225" longitude="-71.25" pageId="5" pageNumber="232" specimenCode="SGO 118488" specimenCount="1" stateProvince="Coquimbo">
<location id="16586038FFCAF766FB37FB2FFF198552" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCAF766FB37FB2FFF198552" country="Chile" latitude="-29.8" longLatPrecision="1225" longitude="-71.25" name="Quebrada de Juan Soldado" pageId="5" pageNumber="232" stateProvince="Coquimbo">Quebrada de Juan Soldado</location>
, al
<locationDeviation id="028A2FC1FFCAF766FEDAFB02FE428552" box="[269,444,1258,1285]" location="La Serena" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FEDAFB02FEDF8553" box="[269,289,1259,1285]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="232" type="Herbarium">N</collectionCode>
de La Serena
</locationDeviation>
, por la carretera panamericana (
<geoCoordinate id="76B35024FFCAF766FCF5FB03FC7B8552" box="[802,901,1258,1284]" degrees="29" direction="south" minutes="48" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.8">29°48’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FC46FB03FBFE8552" box="[913,1024,1258,1285]" degrees="71" direction="west" minutes="15" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.25">71°15’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FBC4FB03FBA58553" box="[1043,1115,1258,1285]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="232" unit="m" value="110.0">
<elevation id="98AAD1D0FFCAF766FBC4FB03FBA58553" box="[1043,1115,1258,1285]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="232" unit="m" value="110.0">110 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FBB0FB03FB0B8553" box="[1127,1269,1258,1285]" pageId="5" pageNumber="232" value="1957-09-17">
<collectingDate id="777DE9CBFFCAF766FBB0FB03FB0B8553" box="[1127,1269,1258,1285]" pageId="5" pageNumber="232" value="1957-09-17">17 Sep 1957</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FAD7FB03FA558552" box="[1280,1451,1258,1284]" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FAD7FB03FAE88552" box="[1280,1302,1258,1284]" country="Denmark" name="University of Copenhagen" pageId="5" pageNumber="232" type="Herbarium">C</collectionCode>
.
<collectorName id="BE725335FFCAF766FAF7FB02FA928552" box="[1312,1388,1259,1284]" pageId="5" pageNumber="232">Muñoz</collectorName>
4296
</emphasis>
(
<specimenCode id="43219E98FFCAF766FF47FAE7FED4857E" box="[144,298,1294,1320]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 118488</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="A3EF3CBEFFCAF766FEE8FAE6FE8E851A" collectingDate="1971-12-25" collectionCode="N, K, M, J" collectorName="Cheese &amp; Watson" country="Chile" elevation="30" latitude="-29.616667" location="Beckett" longLatPrecision="1226" longitude="-71.28333" municipality="La Serena" pageId="5" pageNumber="232" specimenCode="SGO 110041" specimenCount="1" stateProvince="Coquimbo">
<collectingMunicipality id="F35CAC99FFCAF766FEE8FAE6FE4A857E" box="[319,436,1294,1321]" pageId="5" pageNumber="232">La Serena</collectingMunicipality>
,
<quantity id="D47F9B06FFCAF766FE17FAE7FDF7857E" box="[448,521,1294,1320]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="km" value="30.0">30 km</quantity>
<collectionCode id="7596AE26FFCAF766FDC6FAE6FDDB857F" box="[529,549,1295,1321]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="232" type="Herbarium">N</collectionCode>
(
<geoCoordinate id="76B35024FFCAF766FDE0FAE7FD65857E" box="[567,667,1294,1320]" degrees="29" direction="south" minutes="37" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.616667">29°37’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FD7DFAE7FCE4857E" box="[682,794,1294,1320]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.28333">71°17’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FCF8FAE7FC92857F" box="[815,876,1294,1321]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="m" value="30.0">
<elevation id="98AAD1D0FFCAF766FCF8FAE7FC92857F" box="[815,876,1294,1321]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FCAEFAE7FBF0857E" box="[889,1038,1294,1321]" pageId="5" pageNumber="232" value="1971-12-25">
<collectingDate id="777DE9CBFFCAF766FCAEFAE7FBF0857E" box="[889,1038,1294,1321]" pageId="5" pageNumber="232" value="1971-12-25">25 Dec 1971</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FBCCFAE6FF3E851A" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FBCCFAE6FBCF857E" box="[1051,1073,1295,1320]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="232" type="Herbarium">K</collectionCode>
.
<location id="16586038FFCAF766FBEAFAE7FB6C857E" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCAF766FBEAFAE7FB6C857E" box="[1085,1170,1294,1320]" country="Chile" latitude="-29.616667" longLatPrecision="1226" longitude="-71.28333" municipality="La Serena" name="Beckett" pageId="5" pageNumber="232" stateProvince="Coquimbo">Beckett</location>
,
<collectionCode id="7596AE26FFCAF766FB48FAE6FB44857E" box="[1183,1210,1295,1320]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="232" type="Herbarium">M</collectionCode>
.
<collectorName id="BE725335FFCAF766FB11FAE7FAE9857E" box="[1222,1303,1294,1320]" pageId="5" pageNumber="232">Cheese</collectorName>
&amp;
<collectionCode id="7596AE26FFCAF766FAE9FAE6FAB0857E" box="[1342,1358,1295,1320]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="5" pageNumber="232" type="Herbarium">J</collectionCode>
.
<collectorName id="BE725335FFCAF766FA8DFAE6FA55857E" box="[1370,1451,1295,1320]" pageId="5" pageNumber="232">Watson</collectorName>
4678
</emphasis>
(
<specimenCode id="43219E98FFCAF766FF18FADBFE97851A" box="[207,361,1330,1356]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 110041</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="A3EF3CBEFFCAF766FEABFADAFF178526" collectingDate="1984-10-21" collectionCode="M" collectorName="Munoz" country="Chile" elevation="200" latitude="-29.75" location="Km" longLatPrecision="1225" longitude="-71.316666" pageId="5" pageNumber="232" specimenCode="SGO 108070" specimenCount="1" stateProvince="Coquimbo">
<location id="16586038FFCAF766FEABFADAFE58851B" LSID="urn:lsid:plazi:treatment:9B2E87F5FFCCF766FF5FF8BDFB758527:16586038FFCAF766FEABFADAFE58851B" box="[380,422,1331,1357]" country="Chile" latitude="-29.75" longLatPrecision="1225" longitude="-71.316666" name="Km" pageId="5" pageNumber="232" stateProvince="Coquimbo">Km</location>
490, hacia los roquerios (
<geoCoordinate id="76B35024FFCAF766FD1CFADBFCD1851A" box="[715,815,1330,1357]" degrees="29" direction="south" minutes="45" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.75">29°45’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FCEBFADBFC52851A" box="[828,940,1330,1356]" degrees="71" direction="west" minutes="19" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.316666">71°19’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FC17FADBFBF4851B" box="[960,1034,1330,1357]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="232" unit="m" value="200.0">
<elevation id="98AAD1D0FFCAF766FC17FADBFBF4851B" box="[960,1034,1330,1357]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="232" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FBC1FADBFB5B851A" box="[1046,1189,1330,1356]" pageId="5" pageNumber="232" value="1984-10-21">
<collectingDate id="777DE9CBFFCAF766FBC1FADBFB5B851A" box="[1046,1189,1330,1356]" pageId="5" pageNumber="232" value="1984-10-21">21 Oct 1984</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FB66FADAFA9C851A" box="[1201,1378,1330,1356]" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FB66FADAFB32851A" box="[1201,1228,1331,1356]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="232" type="Herbarium">M</collectionCode>
.
<collectorName id="BE725335FFCAF766FB00FADAFADD851A" box="[1239,1315,1331,1356]" pageId="5" pageNumber="232">Muñoz</collectorName>
1914
</emphasis>
(
<specimenCode id="43219E98FFCAF766FAA5FADBFF1F8526" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 108070</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="A3EF3CBEFFCAF766FF23FABFFB798527" box="[244,1159,1366,1393]" collectingDate="2006-10-15" collectionCode="N, ET" collectorName="El Tofo &amp; Schulz" country="Chile" elevation="300" latitude="-29.45" longLatPrecision="1226" longitude="-71.25" pageId="5" pageNumber="232" specimenCode="SGO 154742" specimenCount="1" stateProvince="Coquimbo">
<collectorName id="BE725335FFCAF766FF23FABFFEB68526" box="[244,328,1366,1392]" pageId="5" pageNumber="232">El Tofo</collectorName>
(
<geoCoordinate id="76B35024FFCAF766FE80FABFFE458526" box="[343,443,1366,1392]" degrees="29" direction="south" minutes="27" orientation="latitude" pageId="5" pageNumber="232" precision="925" value="-29.45">29°27’ S</geoCoordinate>
,
<geoCoordinate id="76B35024FFCAF766FE10FABFFDC88526" box="[455,566,1366,1393]" degrees="71" direction="west" minutes="15" orientation="longitude" pageId="5" pageNumber="232" precision="925" value="-71.25">71°15’ W</geoCoordinate>
),
<quantity id="D47F9B06FFCAF766FD9DFABFFD6D8527" box="[586,659,1366,1393]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="m" value="300.0">
<elevation id="98AAD1D0FFCAF766FD9DFABFFD6D8527" box="[586,659,1366,1393]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="232" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="67391023FFCAF766FD48FABFFCD28526" box="[671,812,1366,1393]" pageId="5" pageNumber="232" value="2006-10-15">
<collectingDate id="777DE9CBFFCAF766FD48FABFFCD28526" box="[671,812,1366,1393]" pageId="5" pageNumber="232" value="2006-10-15">15 Oct 2006</collectingDate>
</date>
,
<emphasis id="21F3EAF1FFCAF766FCE0FABEFC298526" box="[823,983,1366,1392]" italics="true" pageId="5" pageNumber="232">
<collectionCode id="7596AE26FFCAF766FCE0FABEFCB38526" box="[823,845,1367,1392]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="232" type="Herbarium">N</collectionCode>
.
<collectorName id="BE725335FFCAF766FC8FFABFFC5F8526" box="[856,929,1366,1392]" pageId="5" pageNumber="232">Schulz</collectorName>
9
<collectionCode id="7596AE26FFCAF766FC62FABFFC298526" box="[949,983,1366,1392]" country="0" name="East Texas State University" pageId="5" pageNumber="232">ET</collectionCode>
</emphasis>
(
<specimenCode id="43219E98FFCAF766FC32FABFFB818527" box="[997,1151,1366,1393]" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" pageId="5" pageNumber="232" type="Herbarium">SGO 154742</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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