243 lines
31 KiB
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243 lines
31 KiB
XML
<document id="97EB22B3794D72D7D2A465F97857D55A" ID-DOI="10.5281/zenodo.12808359" ID-ISSN="1942-1354" ID-Zenodo-Dep="12808359" ID-ZooBank="993F641D-A7D1-4BB9-BBDB-F812852C69B2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721837439321" checkinUser="felipe" docAuthor="Skelley, Paul E." docDate="2021" docId="737B87F6FFEE1E6DFF77F909FC35FED8" docLanguage="en" docName="InsectaMundi.2021.0896.1-5.pdf" docOrigin="Insecta Mundi 2021 (896)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Priscyparus quisqueyensis Skelley 2021, new species" docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="8F42FF8EFFED1E6BFFB7FFB1FFEEFFEB" masterDocTitle="New genus of Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae) in amber from the Dominican Republic" masterLastPageNumber="5" masterPageNumber="1" pageNumber="2" updateTime="1721840385163" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="3E72248074512593136B3DBAC6D5C878" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1FF76C07E68DAFB6C7E6A9D9A3929C0C">
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<mods:title id="46B7A1801AF9FE62DCCD0C64554E5F71">New genus of Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae) in amber from the Dominican Republic</mods:title>
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</mods:titleInfo>
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<mods:name id="B41D5FCF6E3DA9947F5EB0300E24C0FE" type="personal">
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<mods:role id="F4F53544F3B6AFC67FF53EEE89F3CC1B">
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<mods:roleTerm id="2EBB34B82937C24AB51FACB837EBCFAC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="894F0A7A2C12E1CDCB39F798A95FBC40">Skelley, Paul E.</mods:namePart>
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<mods:affiliation id="F08B76670B3DE9B997701F6B7674A0B7">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100 USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="7A3224B1A5D61B442A8516E990694269">text</mods:typeOfResource>
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<mods:relatedItem id="F62EBF368055335C6CA412AD688AAC07" type="host">
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<mods:titleInfo id="5025B96E1E2E18CA012C877D6A12F4CB">
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<mods:title id="72C6C948B8C0BA2E8E4C5D4104926FA1">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="4E6937869FBA45C049BD5856EB96E85D">
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<mods:date id="7DDB32662A6A7EE9D9027F393FA19336">2021</mods:date>
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<mods:detail id="C56B15B8E498982128FE4EB00BC0D910" type="pubDate">
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<mods:number id="519F2B9C8CD1A1B3DDC3E2EEE244394B">2021-11-19</mods:number>
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</mods:detail>
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<mods:detail id="2C13C1928D64503E60EAD85D6A6A102A" type="volume">
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<mods:number id="D67F8D9BD458982A61DD8AFE5FFD2F03">2021</mods:number>
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</mods:detail>
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<mods:detail id="5CD9478F52275D5CD0D287A0DD37D9C6" type="issue">
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<mods:number id="79090AE1F21340D7FA777648C673EB97">896</mods:number>
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</mods:detail>
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<mods:start id="872A78A35D08F500D0A542F119559066">1</mods:start>
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<mods:classification id="91B9677CA9E3F9E1DB16D96EAFF65AB7">journal article</mods:classification>
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<mods:identifier id="4CDB95C2A686C9B0659E87EA7DF1B8B5" type="DOI">10.5281/zenodo.12808359</mods:identifier>
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<mods:identifier id="8F5984DEB61AF24C1A53288A6993380D" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="ABBC23D862765632DCDCD5499157F283" type="Zenodo-Dep">12808359</mods:identifier>
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<mods:identifier id="BDB77E492B8A31AAA4933B7BCD5A6E33" type="ZooBank">993F641D-A7D1-4BB9-BBDB-F812852C69B2</mods:identifier>
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</mods:mods>
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<treatment id="737B87F6FFEE1E6DFF77F909FC35FED8" ID-DOI="http://doi.org/10.5281/zenodo.12808367" ID-Zenodo-Dep="12808367" LSID="urn:lsid:plazi:treatment:737B87F6FFEE1E6DFF77F909FC35FED8" httpUri="http://treatment.plazi.org/id/737B87F6FFEE1E6DFF77F909FC35FED8" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
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<subSubSection id="B3C8656BFFEE1E68FF77F909FCE2F93F" box="[192,780,1719,1748]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="FB6D36E0FFEE1E68FF77F909FCE2F93F" blockId="3.[192,780,1719,1782]" box="[192,780,1719,1748]" pageId="3" pageNumber="2">
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<heading id="A025818CFFEE1E68FF77F909FCE2F93F" box="[192,780,1719,1748]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
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<emphasis id="C9A6EAF2FFEE1E68FF77F909FCE2F93F" bold="true" box="[192,780,1719,1748]" pageId="3" pageNumber="2">
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<taxonomicName id="3CD24D63FFEE1E68FF77F909FD85F93F" ID-CoL="8SSQK" authority="Skelley" authorityName="Skelley" authorityYear="2021" box="[192,619,1719,1748]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="quisqueyensis" status="sp. nov.">
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<emphasis id="C9A6EAF2FFEE1E68FF77F909FDE4F93F" bold="true" box="[192,522,1720,1748]" italics="true" pageId="3" pageNumber="2">Priscyparus quisqueyensis</emphasis>
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Skelley
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</taxonomicName>
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,
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<taxonomicNameLabel id="D2955789FFEE1E68FDC0F909FCE2F93F" box="[631,780,1719,1748]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3C8656BFFEE1E6EFF77F96DFED9FCE2" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="description">
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<paragraph id="FB6D36E0FFEE1E68FF77F96DFEADF91D" blockId="3.[192,780,1719,1782]" box="[192,323,1756,1782]" pageId="3" pageNumber="2">
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<figureCitation id="63E92A65FFEE1E68FF77F96DFEADF91D" box="[192,323,1756,1782]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="3" pageNumber="2">Figures 1–6</figureCitation>
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</paragraph>
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<paragraph id="FB6D36E0FFEE1E6EFF77F8B4FED9FCE2" blockId="3.[192,1440,1797,1927]" lastBlockId="5.[191,1442,214,777]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
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<emphasis id="C9A6EAF2FFEE1E68FF77F8B4FDE5F8CB" bold="true" box="[192,523,1797,1824]" pageId="3" pageNumber="2">
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Description. Male
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<typeStatus id="24698842FFEE1E68FE2AF8B7FDE8F8CB" box="[413,518,1798,1824]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length 4.5 mm, greatest elytral and pronotal width 1.8 mm. Body with encrustations preventing study of some surface details; shape elongate, narrowing posteriorly (
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<figureCitation id="63E92A65FFEE1E68FB0FF899FB10F8A9" box="[1208,1278,1832,1858]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
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).
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<emphasis id="C9A6EAF2FFEE1E68FAA2F899FAB5F8A9" bold="true" box="[1301,1371,1832,1858]" pageId="3" pageNumber="2">Head.</emphasis>
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Head one-third wider than long, clypeus anteriorly with inflexed lower margin, upper margin quadridentate, inner tooth on each side at lateral third, outer tooth on each side adjacent to edge of gena; other characters obscured in encrustation. Vertex with 4 short longitudinal costae, 2 median costae slightly closer. Gena rounded, hidden in encrustation.
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<emphasis id="C9A6EAF2FFE91E6FFEECF975FE34F935" bold="true" box="[347,474,1732,1758]" pageId="4" pageNumber="3">Pronotum.</emphasis>
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Pronotum with 6 irregularly longitudinal carinae. Paramedian costa complete, nearly parallel, tall and blade-like entire length, anterior lobe greatly enlarged but not swollen (
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<figureCitation id="63E92A65FFE91E6FFB24F957FB1CF8EB" box="[1171,1266,1766,1792]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="4" pageNumber="3">Fig. 4–6</figureCitation>
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), in lateral view anterior lobe flattened above and sloping anteriorly, posterior part of costa slightly swollen and lobe-like over posterior pronotal margin, abruptly declivous at margin, surface between costa punctate; discolateral costa interrupted in anterior third by deep irregular pits, anterior lobe similar development but small than paramedian lobe, posterior part of costa low and complete from base to pit; submarginal costa prominent, anterior lobe similar but smaller than discolateral anterior lobe, posterior part of costa prominently swollen and laterally arched, obscuring structures beneath; all costae extending to posterior pronotal margin; lateral margin of pronotum with two lobes, anterior lobe circular and more prominent than tooth-like median lobe, posterior lateral margin and angle obscure.
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<emphasis id="C9A6EAF2FFE81E6EFE91FE8AFE68FEBE" bold="true" box="[294,390,315,341]" pageId="5" pageNumber="4">Elytron.</emphasis>
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Elytron with 3 equally sharply elevated keel-like costa and a weak marginal costa; discomedian costa separated from sutural ridge by transverse rugulae, costa straight and narrow to slightly swollen apex projecting over trichome; discolateral costa weakly arched at base, narrow and straight, not swollen as approaching and touching trichome; posthumeral costa prominent at humerus, sharply elevated on basal half, abruptly declivous and reducing in size, obsolete as nearing trichome; marginal costa fine from base to caudal bulb; caudal bulb prominently projecting posteriorly beyond elytral apex (
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<figureCitation id="63E92A65FFE81E6EFCC1FE53FC57FE16" box="[886,953,482,509]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
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).
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<emphasis id="C9A6EAF2FFE81E6EFC65FE52FBCBFE16" bold="true" box="[978,1061,483,509]" pageId="5" pageNumber="4">Venter.</emphasis>
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Postprosternal apophysis hastate. Metaventrite with distinct median impression and smaller impressions behind mesocoxae (evident by encrustations), visible surface smooth and punctate. Abdomen mostly encrusted, not visible; two sharp tubercles at the base of the last ventrite (
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<figureCitation id="63E92A65FFE81E6EFE62FDF6FDF9FD89" box="[469,535,583,610]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
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).
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<emphasis id="C9A6EAF2FFE81E6EFD9AFDF6FD87FD8A" bold="true" box="[557,617,583,609]" pageId="5" pageNumber="4">Legs.</emphasis>
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Profemur robust; meso- and metafemur narrow, not swollen; metafemur long, reaching elytral apex. Protibia narrow, apically tridentate, each tooth fine, 1 tooth present on each side of apex and 1 medially; mesotibia narrow, weakly enlarging toward apex, inner margin weakly sinuate, inner apical angle with small inwardly projecting tooth (
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<figureCitation id="63E92A65FFE81E6EFD65FD1DFCFDFD2C" box="[722,787,684,711]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
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); metatibia narrow, encrusted obscuring details. Protarsus short; meso- and metatarsus short, half-length of tibia, tarsomeres thickened, basal tarsomeres long as next two combined.
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</paragraph>
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</subSubSection>
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<caption id="AFAD6668FFE91E6FFF08FA5BFE7BF982" ID-DOI="http://doi.org/10.5281/zenodo.12808361" ID-Zenodo-Dep="12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="4" pageNumber="3" startId="4.[191,278,1514,1540]" targetBox="[192,1440,213,1484]" targetPageId="4" targetType="figure">
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<paragraph id="FB6D36E0FFE91E6FFF08FA5BFE7BF982" blockId="4.[191,1441,1514,1641]" pageId="4" pageNumber="3">
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<emphasis id="C9A6EAF2FFE91E6FFF08FA5BFEA1F9EF" bold="true" box="[191,335,1514,1540]" pageId="4" pageNumber="3">Figures 1–6.</emphasis>
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<taxonomicName id="3CD24D63FFE91E6FFEE2FA5AFD9CF9EF" authorityName="Skelley" authorityYear="2021" box="[341,626,1515,1540]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="quisqueyensis" status="sp. nov.">
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<emphasis id="C9A6EAF2FFE91E6FFEE2FA5AFD9CF9EF" box="[341,626,1515,1540]" italics="true" pageId="4" pageNumber="3">Priscyparus quisqueyensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="D2955789FFE91E6FFDC8FA5AFCEBF9EF" box="[639,773,1514,1541]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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, holotype.
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<emphasis id="C9A6EAF2FFE91E6FFCC8FA5BFC79F9EF" bold="true" box="[895,919,1514,1540]" pageId="4" pageNumber="3">1)</emphasis>
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Amber piece with host termite, holotype indicated by arrow.
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<emphasis id="C9A6EAF2FFE91E6FFED9F9BAFE69F9CE" bold="true" box="[366,391,1547,1573]" pageId="4" pageNumber="3">2)</emphasis>
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Dorsal view.
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<emphasis id="C9A6EAF2FFE91E6FFDABF9BAFDDAF9CE" bold="true" box="[540,564,1547,1573]" pageId="4" pageNumber="3">3)</emphasis>
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Right lateral view showing elytral apex.
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<emphasis id="C9A6EAF2FFE91E6FFC42F9BAFBE3F9CE" bold="true" box="[1013,1037,1547,1573]" pageId="4" pageNumber="3">4)</emphasis>
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Right lateral view showing pronotal development.
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<emphasis id="C9A6EAF2FFE91E6FFEE8F99CFE99F9AC" bold="true" box="[351,375,1581,1607]" pageId="4" pageNumber="3">5)</emphasis>
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Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen.
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<emphasis id="C9A6EAF2FFE91E6FFF76F9FEFF37F982" bold="true" box="[193,217,1615,1641]" pageId="4" pageNumber="3">6)</emphasis>
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Left lateral view.
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</paragraph>
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</caption>
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<subSubSection id="B3C8656BFFE81E6EFF77FCA8FEB0FC73" pageId="5" pageNumber="4" type="materials_examined">
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<paragraph id="FB6D36E0FFE81E6EFF77FCA8FEB0FC73" blockId="5.[192,1440,793,920]" pageId="5" pageNumber="4">
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<emphasis id="C9A6EAF2FFE81E6EFF77FCA8FE43FCD8" bold="true" box="[192,429,793,819]" pageId="5" pageNumber="4">Materials examined.</emphasis>
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<materialsCitation id="4BBA3CBDFFE81E6EFE04FCA8FEB0FC73" collectionCode="FSCA" country="Dominican Republic" location="These" pageId="5" pageNumber="4" specimenCount="2" typeStatus="holotype">
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<typeStatus id="24698842FFE81E6EFE04FCA8FDF5FCD8" box="[435,539,793,819]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
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fossil in amber labeled: “[red paper] /
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<typeStatus id="24698842FFE81E6EFC08FCA8FBBFFCD8" box="[959,1105,793,819]" pageId="5" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="3CD24D63FFE81E6EFBD1FCABFB09FCD8" authorityName="Skelley" authorityYear="2021" box="[1126,1255,794,819]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="C9A6EAF2FFE81E6EFBD1FCABFB09FCD8" box="[1126,1255,794,819]" italics="true" pageId="5" pageNumber="4">Priscyparus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="3CD24D63FFE81E6EFB4CFCABFA7EFCD8" authorityName="Skelley" authorityYear="2021" box="[1275,1424,794,819]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="quisqueyensis">
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<emphasis id="C9A6EAF2FFE81E6EFB4CFCABFA7EFCD8" box="[1275,1424,794,819]" italics="true" pageId="5" pageNumber="4">quisqueyensis</emphasis>
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</taxonomicName>
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/ Skelley /”, deposited
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<collectionCode id="9DC3AE25FFE81E6EFE12FC8AFE04FCBE" box="[421,490,827,853]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
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.
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<specimenCount id="EDD4FD69FFE81E6EFE43FC8AFCE7FCBF" box="[500,777,826,853]" count="2" pageId="5" pageNumber="4" type="generic" typeStatus="paratype">Two paratype specimens</specimenCount>
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labeled to be from “Alto de las Piedras,
|
||
<collectingCountry id="83C57670FFE81E6EFB70FC8BFF35FC9D" name="Dominican Republic" pageId="5" pageNumber="4">Dominican Republic</collectingCountry>
|
||
” (
|
||
<collectionCode id="9DC3AE25FFE81E6EFF44FCEDFED4FC9D" box="[243,314,860,886]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
|
||
) were also studied.
|
||
<location id="FE0D603BFFE81E6EFDA2FCEDFDBBFC9D" LSID="urn:lsid:plazi:treatment:737B87F6FFEE1E6DFF77F909FC35FED8:FE0D603BFFE81E6EFDA2FCEDFDBBFC9D" box="[533,597,860,886]" country="Dominican Republic" name="These" pageId="5" pageNumber="4">These</location>
|
||
<typeStatus id="24698842FFE81E6EFDECFCECFD29FC9C" box="[603,711,861,887]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
are mostly enclosed in bubbles (
|
||
<figureCitation id="63E92A65FFE81E6EFB85FCEDFB9DFC9D" box="[1074,1139,860,886]" captionStart="Figure 7" captionStartId="5.[462,538,1900,1926]" captionTargetBox="[477,1169,1337,1871]" captionTargetId="figure-488@5.[462,1171,1337,1871]" captionTargetPageId="5" captionText="Figure 7. Priscyparus quisqueyensis, new species, paratypes." figureDoi="http://doi.org/10.5281/zenodo.12808365" httpUri="https://zenodo.org/record/12808365/files/figure.png" pageId="5" pageNumber="4">Fig. 7</figureCitation>
|
||
), making structural details poorly visible.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3C8656BFFE81E6EFF77FC16FE6AFBEF" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="FB6D36E0FFE81E6EFF77FC16FE6AFBEF" blockId="5.[192,1441,935,1028]" pageId="5" pageNumber="4">
|
||
<emphasis id="C9A6EAF2FFE81E6EFF77FC16FEAAFC2A" bold="true" box="[192,324,935,961]" pageId="5" pageNumber="4">Etymology.</emphasis>
|
||
The species epithet comes from Hispaniola, which was called Quisqueya by the aboriginal Taíno people. The species name is Latinized with -
|
||
<emphasis id="C9A6EAF2FFE81E6EFD04FC7BFD09FC08" box="[691,743,970,995]" italics="true" pageId="5" pageNumber="4">ensis</emphasis>
|
||
, so the full name of this species means the “ancient
|
||
<taxonomicName id="3CD24D63FFE81E6EFA8FFC78FA4FFC08" box="[1336,1441,969,995]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFA8FFC78FA4FFC08" box="[1336,1441,969,995]" italics="true" pageId="5" pageNumber="4">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
from Quisqueya.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3C8656BFFE81E6DFF77FBA5FC35FED8" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="FB6D36E0FFE81E6EFF77FBA5FE58FAF2" blockId="5.[192,1446,1044,1306]" pageId="5" pageNumber="4">
|
||
<emphasis id="C9A6EAF2FFE81E6EFF77FBA5FEA6FBC5" bold="true" box="[192,328,1044,1070]" pageId="5" pageNumber="4">Comments.</emphasis>
|
||
This species is unusual in its combination of characters, as well as having a few unique characters. The pronotal costae are greatly raised, even more than is seen in
|
||
<taxonomicName id="3CD24D63FFE81E6EFC5FFB87FB82FBBB" authorityName="Wasmann" authorityYear="1894" box="[1000,1132,1078,1104]" class="Insecta" family="Scarabaeidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFC5FFB87FB82FBBB" box="[1000,1132,1078,1104]" italics="true" pageId="5" pageNumber="4">Termitodius</emphasis>
|
||
</taxonomicName>
|
||
. Most other characters are similar to
|
||
<taxonomicName id="3CD24D63FFE81E6EFE84FBE6FDC2FB9A" authority="Nakane" authorityName="Nakane" authorityYear="1961" box="[307,556,1111,1137]" class="Insecta" family="Scarabaeidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFE84FBE6FE21FB9A" box="[307,463,1111,1137]" italics="true" pageId="5" pageNumber="4">Termitodiellus</emphasis>
|
||
Nakane
|
||
</taxonomicName>
|
||
from SE Asia,
|
||
<collectingCountry id="83C57670FFE81E6EFD64FBE6FCAFFB9A" box="[723,833,1111,1137]" name="Indonesia" pageId="5" pageNumber="4">Indonesia</collectingCountry>
|
||
and New
|
||
<collectingCountry id="83C57670FFE81E6EFC04FBE6FBE6FB9A" box="[947,1032,1111,1137]" name="Guinea-Bissau" pageId="5" pageNumber="4">Guinea</collectingCountry>
|
||
. The elongate, posteriorly projecting discomedian costa and caudal bulb of the elytra are seen in
|
||
<taxonomicName id="3CD24D63FFE81E6EFCDDFBC8FB0EFB78" authority="(Paulian)" baseAuthorityName="Paulian" baseAuthorityYear="1983" box="[874,1248,1145,1171]" class="Insecta" family="Scarabaeidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="besucheti">
|
||
<emphasis id="C9A6EAF2FFE81E6EFCDDFBC8FB9CFB78" box="[874,1138,1145,1171]" italics="true" pageId="5" pageNumber="4">Termitodiellus besucheti</emphasis>
|
||
(Paulian)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CD24D63FFE81E6EFAACFBC8FD6BFB5F" authority="(Krikken and Huijbregts)" baseAuthorityName="Krikken and Huijbregts" class="Insecta" family="Scarabaeidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hammondi">
|
||
<emphasis id="C9A6EAF2FFE81E6EFAACFBC8FEB1FB5F" italics="true" pageId="5" pageNumber="4">Termitodiellus hammondi</emphasis>
|
||
(Krikken and Huijbregts)
|
||
</taxonomicName>
|
||
from Borneo and Sumatra (
|
||
<bibRefCitation id="9F434B11FFE81E6EFC70FB2BFAFBFB5E" author="Krikken J & Huijbregts J." box="[967,1301,1178,1205]" pageId="5" pageNumber="4" pagination="97 - 111" refId="ref3439" refString="Krikken J, Huijbregts J. 1987. Southeast Asian Termitodius: A taxonomic review, with description of four new species (Coleoptera: Aphodiidae). Zoologische Mededelingen (Leiden) 61 (7): 97 - 111." type="journal article" year="1987">Krikken and Huijbregts 1987</bibRefCitation>
|
||
, fig. 13–14). However,
|
||
<taxonomicName id="3CD24D63FFE81E6EFE87FB0DFE5AFB3D" authorityName="Wasmann" authorityYear="1894" box="[304,436,1212,1238]" class="Insecta" family="Scarabaeidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFE87FB0DFE5AFB3D" box="[304,436,1212,1238]" italics="true" pageId="5" pageNumber="4">Termitodius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CD24D63FFE81E6EFE5BFB0DFD66FB3D" authorityName="Nakane" authorityYear="1961" box="[492,648,1212,1238]" class="Insecta" family="Scarabaeidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFE5BFB0DFD66FB3D" box="[492,648,1212,1238]" italics="true" pageId="5" pageNumber="4">Termitodiellus</emphasis>
|
||
</taxonomicName>
|
||
have different tibial
|
||
<typeStatus id="24698842FFE81E6EFCDAFB0CFC46FB3C" box="[877,936,1213,1239]" pageId="5" pageNumber="4">types</typeStatus>
|
||
than
|
||
<taxonomicName id="3CD24D63FFE81E6EFC5EFB0CFB84FB3D" authorityName="Skelley" authorityYear="2021" box="[1001,1130,1213,1238]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFC5EFB0CFB84FB3D" box="[1001,1130,1213,1238]" italics="true" pageId="5" pageNumber="4">Priscyparus</emphasis>
|
||
</taxonomicName>
|
||
which has a mesotibia with an inwardly directed spine on inner apex. This is the
|
||
<taxonomicName id="3CD24D63FFE81E6EFCA1FB6FFC90FB13" box="[790,894,1246,1272]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFCA1FB6FFC90FB13" box="[790,894,1246,1272]" italics="true" pageId="5" pageNumber="4">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="24698842FFE81E6EFC32FB6FFC56FB13" box="[901,952,1246,1272]" pageId="5" pageNumber="4">type</typeStatus>
|
||
tibia found in
|
||
<taxonomicName id="3CD24D63FFE81E6EFBD6FB6FFB24FB13" box="[1121,1226,1246,1272]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFE81E6EFBD6FB6FFB24FB13" box="[1121,1226,1246,1272]" italics="true" pageId="5" pageNumber="4">Rhyparus</emphasis>
|
||
</taxonomicName>
|
||
and closely related genera (
|
||
<bibRefCitation id="9F434B11FFE81E6EFEAEFB4EFE45FAF2" author="Skelley PE" box="[281,427,1279,1305]" pageId="5" pageNumber="4" pagination="1 - 9" refId="ref3638" refString="Skelley PE. 2007. Generic limits of the Rhyparini with respect to the genus Termitodius Wasmann, 1894 (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi 0009: 1 - 9." type="book chapter" year="2007">Skelley 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AFAD6668FFE81E6EFE79F8DDFB89F86C" ID-DOI="http://doi.org/10.5281/zenodo.12808365" ID-Zenodo-Dep="12808365" box="[462,1127,1900,1927]" httpUri="https://zenodo.org/record/12808365/files/figure.png" pageId="5" pageNumber="4" startId="5.[462,538,1900,1926]" targetBox="[477,1169,1337,1871]" targetPageId="5" targetType="figure">
|
||
<paragraph id="FB6D36E0FFE81E6EFE79F8DDFB89F86C" blockId="5.[462,1127,1900,1927]" box="[462,1127,1900,1927]" pageId="5" pageNumber="4">
|
||
<emphasis id="C9A6EAF2FFE81E6EFE79F8DDFDDBF86D" bold="true" box="[462,565,1900,1926]" pageId="5" pageNumber="4">Figure 7.</emphasis>
|
||
<taxonomicName id="3CD24D63FFE81E6EFD8CF8DCFCB9F86D" authorityName="Skelley" authorityYear="2021" box="[571,855,1901,1926]" class="Insecta" family="Scarabaeidae" genus="Priscyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="quisqueyensis" status="sp. nov.">
|
||
<emphasis id="C9A6EAF2FFE81E6EFD8CF8DCFCB9F86D" box="[571,855,1901,1926]" italics="true" pageId="5" pageNumber="4">Priscyparus quisqueyensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D2955789FFE81E6EFCD3F8DCFC04F86D" box="[868,1002,1900,1927]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
|
||
, paratypes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB6D36E0FFEB1E6DFEB7FF67FC35FED8" blockId="6.[191,1440,214,307]" pageId="6" pageNumber="5">
|
||
The
|
||
<typeStatus id="24698842FFEB1E6DFE86FF67FE7DFF1B" box="[305,403,214,240]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
is in a piece of amber with a host termite (
|
||
<figureCitation id="63E92A65FFEB1E6DFCCFFF67FC59FF1B" box="[888,951,214,240]" captionStart="Figures 1–6" captionStartId="4.[191,278,1514,1540]" captionTargetBox="[192,1440,213,1484]" captionTargetId="figure-128@4.[192,1440,213,1484]" captionTargetPageId="4" captionText="Figures 1–6. Priscyparus quisqueyensis, new species, holotype. 1) Amber piece with host termite, holotype indicated by arrow. 2) Dorsal view. 3) Right lateral view showing elytral apex. 4) Right lateral view showing pronotal development. 5) Right lateral view, arrows point to suspected sexual dimorphisms on mesotibia and abdomen. 6) Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808361" httpUri="https://zenodo.org/record/12808361/files/figure.png" pageId="6" pageNumber="5">Fig. 1</figureCitation>
|
||
). Other rhyparine species are guests in termite colonies (
|
||
<bibRefCitation id="9F434B11FFEB1E6DFED7FF46FD33FEFA" author="Reyes-Castillo P & Martinez A." box="[352,733,247,273]" pageId="6" pageNumber="5" pagination="115 - 133" refId="ref3604" refString="Reyes-Castillo P, Martinez A. 1979. Nuevos Rhyparini Neotropicales, con notas sobre su biologia (Coleoptera, Scarabeidae, Aphodiinae). Folia Entomologica Mexicana 41: 115 - 133." type="journal article" year="1979">Reyes-Castillo and Martínez 1979</bibRefCitation>
|
||
). Amber
|
||
<taxonomicName id="3CD24D63FFEB1E6DFCF7FF49FBAEFEF9" authority="Wasmann" authorityName="Wasmann" authorityYear="1896" box="[832,1088,248,274]" class="Insecta" family="Rhinotermitidae" genus="Coptotermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A6EAF2FFEB1E6DFCF7FF49FC27FEF9" box="[832,969,248,274]" italics="true" pageId="6" pageNumber="5">Coptotermes</emphasis>
|
||
Wasmann
|
||
</taxonomicName>
|
||
(Isoptera:
|
||
<taxonomicName id="3CD24D63FFEB1E6DFB01FF46FA9CFEFA" box="[1206,1394,247,273]" class="Insecta" family="Rhinotermitidae" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="5" phylum="Arthropoda" rank="family">Rhinotermitidae</taxonomicName>
|
||
) are reported from
|
||
<collectingCountry id="83C57670FFEB1E6DFED3FEA8FDE2FED8" box="[356,524,281,307]" name="Dominican Republic" pageId="6" pageNumber="5">the Dominican</collectingCountry>
|
||
Republic by Krisna and Grimaldi (2009).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |