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<document id="D8C80B5F24A6632FE8E1EE97555BB01F" ID-CLB-Dataset="126748" ID-DOI="10.1111/zoj.12330" ID-GBIF-Dataset="6f1d30f9-d20f-415b-a70c-a65074943adf" ID-ISSN="0024-4082" ID-Zenodo-Dep="5358180" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630223637443" checkinUser="marcus" docAuthor="Hosoishi, Shingo &amp; Ogata, Kazuo" docDate="2016" docId="1E64E12B3E6FFFA37FBDBCE8DC0DFBB4" docLanguage="en" docName="zoj12330.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (3)" docSource="http://dx.doi.org/10.1111/zoj.12330" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Orthocrema (Orthocrema) Santschi" docType="treatment" docVersion="5" lastPageNumber="566" masterDocId="E25D99533E7FFFB07F0BBA1BD82FFFC9" masterDocTitle="Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae)" masterLastPageNumber="606" masterPageNumber="547" pageNumber="563" updateTime="1699216389104" updateUser="ExternalLinkService">
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<mods:title id="1FAF39C1225FFDD744C4519F2B74473A">Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="4C75B0D27007691227845CD9AA9EF514">Hosoishi, Shingo</mods:namePart>
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<mods:namePart id="1B4D70F4744382C0D13D5929B5FC8DC3">Ogata, Kazuo</mods:namePart>
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<treatment id="1E64E12B3E6FFFA37FBDBCE8DC0DFBB4" ID-DOI="http://doi.org/10.5281/zenodo.7015402" ID-GBIF-Taxon="199534311" ID-Zenodo-Dep="7015402" LSID="urn:lsid:plazi:treatment:1E64E12B3E6FFFA37FBDBCE8DC0DFBB4" httpUri="http://treatment.plazi.org/id/1E64E12B3E6FFFA37FBDBCE8DC0DFBB4" lastPageId="19" lastPageNumber="566" pageId="16" pageNumber="563">
<subSubSection id="DED703B63E6FFFA07FBDBCE8DA26F8E1" pageId="16" pageNumber="563" type="nomenclature">
<paragraph id="9672503D3E6FFFA07FBDBCE8DA26F8E1" blockId="16.[182,720,1779,1832]" pageId="16" pageNumber="563">
<heading id="CD3AE7513E6FFFA07FBDBCE8DA26F8E1" centered="true" fontSize="8" level="2" pageId="16" pageNumber="563" reason="8">
SYNOPSIS OF ASIAN SPECIES OF THE SUBGENUS
<taxonomicName id="51CD2BBE3E6FFFA07E75BD08DA26F8E1" authorityName="Santschi" box="[382,521,1811,1832]" class="Insecta" family="Formicidae" genus="Orthocrema" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="563" phylum="Arthropoda" rank="genus">
<emphasis id="A4B98C2F3E6FFFA07E75BD08DA26F8E1" box="[382,521,1811,1832]" italics="true" pageId="16" pageNumber="563">Orthocrema</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E6FFFA27F9BBD22D984FA76" lastPageId="18" lastPageNumber="565" pageId="16" pageNumber="563" type="description">
<paragraph id="9672503D3E6FFFA07F9BBD22DAD8F8A7" blockId="16.[144,759,1849,1902]" pageId="16" pageNumber="563">
The worker caste of the subgenus
<taxonomicName id="51CD2BBE3E6FFFA07D1CBD22DAB2F887" authorityName="Santschi" box="[535,669,1849,1870]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="563" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6FFFA07D1CBD22DAB2F887" box="[535,669,1849,1870]" italics="true" pageId="16" pageNumber="563">Orthocrema</emphasis>
</taxonomicName>
was diagnosed by
<bibRefCitation id="F25C2DCC3E6FFFA07E1CBD43D9E2F8A7" author="Blaimer BB" box="[279,461,1880,1902]" pageId="16" pageNumber="563" pagination="47 - 67" refId="ref38014" refString="Blaimer BB. 2012 c. A subgeneric revision of Crematogaster and discussion of regional species groups (Hymenoptera: Formicidae). Zootaxa 3482: 47 - 67." type="journal article" year="2012">Blaimer (2012c)</bibRefCitation>
on a global scale. For the
</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBFC9DCC7F9CF" blockId="16.[806,1421,1489,1542]" pageId="16" pageNumber="563">purpose of taxonomic convenience, we present the following characteristics for Asian fauna:</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBC3DDBCAF97F" blockId="16.[806,1422,1574,1902]" pageId="16" pageNumber="563">
1. Mandibles with four teeth on the masticatory margin; the teeth located at equal distance each other in most species, but basal tooth located apart from other teeth in subterranean species (
<bibRefCitation id="F25C2DCC3E6FFFA07A26BC99DBF6F97F" author="Hosoishi S &amp; Yamane S &amp; Ogata K" pageId="16" pageNumber="563" pagination="345 - 350" refId="ref39498" refString="Hosoishi S, Yamane S, Ogata K. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13: 345 - 350." type="journal article" year="2010">
Hosoishi
<emphasis id="A4B98C2F3E6FFFA07C5DBCBADBA8F97C" box="[854,903,1696,1718]" italics="true" pageId="16" pageNumber="563">et al</emphasis>
., 2010
</bibRefCitation>
).
</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBCA4DCF8F91C" blockId="16.[806,1422,1574,1902]" box="[806,1239,1727,1749]" pageId="16" pageNumber="563">
2. Palp formula 5, 3 (
<bibRefCitation id="F25C2DCC3E6FFFA07B3EBCA4DCE3F91C" author="Bolton B" box="[1077,1228,1727,1749]" pageId="16" pageNumber="563" pagination="1 - 370" refId="ref38164" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">
<collectingRegion id="54099EDF3E6FFFA07B3EBCA4DCACF91C" box="[1077,1155,1727,1749]" country="United Kingdom" name="Bolton" pageId="16" pageNumber="563">Bolton</collectingRegion>
, 2003
</bibRefCitation>
).
</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBCC5DCCCF8DB" blockId="16.[806,1422,1574,1902]" pageId="16" pageNumber="563">
3. Anterolateral margins of clypeus not protruded anteriorly (
<bibRefCitation id="F25C2DCC3E6FFFA07CB7BCE7DCF7F8DB" author="Hosoishi S &amp; Ogata K" box="[956,1240,1788,1810]" pageId="16" pageNumber="563" pagination="18 - 30" refId="ref39425" refString="Hosoishi S, Ogata K. 2012. Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae). Zootaxa 3349: 18 - 30." type="journal article" year="2012">Hosoishi &amp; Ogata, 2012</bibRefCitation>
).
</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBD00DDA1F886" blockId="16.[806,1422,1574,1902]" pageId="16" pageNumber="563">4. Clypeus usually striated with longitudinal rugulae; the length of rugulae variable in different species.</paragraph>
<paragraph id="9672503D3E6FFFA07C2DBD43DC49F8A7" blockId="16.[806,1422,1574,1902]" box="[806,1126,1880,1902]" pageId="16" pageNumber="563">5. Antenna 11-segmented.</paragraph>
<caption id="C2B200B53E6EFFA17FAFBF63DB94FA42" ID-DOI="http://doi.org/10.5281/zenodo.5358204" ID-Zenodo-Dep="5358204" box="[164,955,1400,1419]" httpUri="https://zenodo.org/record/5358204/files/figure.png" pageId="17" pageNumber="564" startId="17.[164,243,1400,1419]" targetBox="[167,1441,196,1367]" targetPageId="17">
<paragraph id="9672503D3E6EFFA17FAFBF63DB94FA42" blockId="17.[164,955,1400,1419]" box="[164,955,1400,1419]" pageId="17" pageNumber="564">
<emphasis id="A4B98C2F3E6EFFA17FAFBF63D936FA42" bold="true" box="[164,281,1400,1419]" pageId="17" pageNumber="564">Figure 51.</emphasis>
Geographical distribution of the species of the
<taxonomicName id="51CD2BBE3E6EFFA17C16BF63DB41FA42" authorityName="Mayr" authorityYear="1897" box="[797,878,1400,1419]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="A4B98C2F3E6EFFA17C16BF63DB02FA42" box="[797,813,1400,1419]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17C36BF63DB41FA42" box="[829,878,1400,1419]" italics="true" pageId="17" pageNumber="564">biroi</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="9672503D3E6EFFA17FAFBFD1D932F9F2" blockId="17.[164,779,1482,1902]" pageId="17" pageNumber="564">
6. Antennal club 2-segmented. Apical two flagellar segments clearly distinguished in coloration from other flagellar segments especially in the
<taxonomicName id="51CD2BBE3E6EFFA17D94BC13DB24F9D5" authorityName="Forel" authorityYear="1912" box="[671,779,1543,1565]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="baduvi">
<emphasis id="A4B98C2F3E6EFFA17D94BC13DA9FF9D4" box="[671,688,1544,1565]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17DB4BC1CDB24F9D5" box="[703,779,1543,1564]" italics="true" pageId="17" pageNumber="564">baduvi</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="9672503D3E6EFFA17FAFBC5EDA22F951" blockId="17.[164,779,1482,1902]" pageId="17" pageNumber="564">
7. Sensilla basiconica and sensilla trichodea curvata distributed in the apical two flagellar segments (
<bibRefCitation id="F25C2DCC3E6EFFA17FD0BC99DA2DF951" author="Hosoishi S &amp; Ogata K" box="[219,514,1666,1688]" pageId="17" pageNumber="564" pagination="755 - 766" refId="ref39390" refString="Hosoishi S, Ogata K. 2009 c. The relationship between the antennal club and the antennal sensilla in Crematogaster ants (Hymenoptera: Formicidae). Sociobiology 53: 755 - 766." type="journal article" year="2009">Hosoishi &amp; Ogata, 2009c</bibRefCitation>
).
</paragraph>
<paragraph id="9672503D3E6EFFA17FAFBCBBD94DF93D" blockId="17.[164,779,1482,1902]" pageId="17" pageNumber="564">
8. Compound eye generally distinct and large, but strongly reduced in subterranean species (
<bibRefCitation id="F25C2DCC3E6EFFA17DA0BCA4D978F93D" author="Hosoishi S &amp; Yamane S &amp; Ogata K" pageId="17" pageNumber="564" pagination="345 - 350" refId="ref39498" refString="Hosoishi S, Yamane S, Ogata K. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13: 345 - 350." type="journal article" year="2010">
Hosoishi
<emphasis id="A4B98C2F3E6EFFA17FDFBCC5D92BF93B" box="[212,260,1757,1779]" italics="true" pageId="17" pageNumber="564">et al</emphasis>
., 2010
</bibRefCitation>
).
</paragraph>
<paragraph id="9672503D3E6EFFA17FAFBCE7DAC1F8DB" blockId="17.[164,779,1482,1902]" box="[164,750,1788,1810]" pageId="17" pageNumber="564">9. Ocelli rarely present in intermediate workers.</paragraph>
<paragraph id="9672503D3E6EFFA17FAFBD00D9DCF8F8" blockId="17.[164,779,1482,1902]" box="[164,499,1819,1841]" pageId="17" pageNumber="564">10. Occipital carina distinct.</paragraph>
<paragraph id="9672503D3E6EFFA17FAFBD22DB24F8A4" blockId="17.[164,779,1482,1902]" pageId="17" pageNumber="564">
11. Mesonotal ridges developed dorsolaterally, but weakly developed in
<taxonomicName id="51CD2BBE3E6EFFA17EB5BD43DA13F8A4" authorityName="Hosoishi, Yamane &amp; Ogata" authorityYear="2010" box="[446,572,1880,1901]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="masukoi">
<emphasis id="A4B98C2F3E6EFFA17EB5BD43D9E0F8A4" box="[446,463,1880,1901]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17ED5BD43DA13F8A4" box="[478,572,1880,1901]" italics="true" pageId="17" pageNumber="564">masukoi</emphasis>
</taxonomicName>
and
<taxonomicName id="51CD2BBE3E6EFFA17D7DBD43DB2BF8A4" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[630,772,1880,1901]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="moatensis">
<emphasis id="A4B98C2F3E6EFFA17D7DBD43DAA8F8A4" box="[630,647,1880,1901]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17D9DBD43DB2BF8A4" box="[662,772,1880,1901]" italics="true" pageId="17" pageNumber="564">moatensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6EFFA17C31BFD1DB9FFA37" blockId="17.[826,1441,1482,1902]" pageId="17" pageNumber="564">12. Ventrolateral katepisternal ridge distinct (Hosoishi, 2015).</paragraph>
<paragraph id="9672503D3E6EFFA17C31BC1CDD6CF9F2" blockId="17.[826,1441,1482,1902]" pageId="17" pageNumber="564">
13. Propodeal spines usually developed, but undeveloped or small tubercles in
<taxonomicName id="51CD2BBE3E6EFFA17BAFBC3DDD12F9F2" authority="Forel, 1904" authorityName="Forel" authorityYear="1904" box="[1188,1341,1574,1595]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="binghamii">
<emphasis id="A4B98C2F3E6EFFA17BAFBC3DDC99F9F2" box="[1188,1206,1574,1595]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17BCCBC3DDD12F9F2" box="[1223,1341,1574,1595]" italics="true" pageId="17" pageNumber="564">binghamii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6EFFA17C31BC5FDD07F993" blockId="17.[826,1441,1482,1902]" box="[826,1320,1604,1626]" pageId="17" pageNumber="564">14. Petiole usually with subparallel sides.</paragraph>
<paragraph id="9672503D3E6EFFA17C31BC78DD50F9B0" blockId="17.[826,1441,1482,1902]" box="[826,1407,1635,1657]" pageId="17" pageNumber="564">15. Petiole usually with short process posteriorly.</paragraph>
<paragraph id="9672503D3E6EFFA17C31BC9ADD40F97F" blockId="17.[826,1441,1482,1902]" pageId="17" pageNumber="564">16. Postpetiole not bilobed. If weakly bilobed, usually without distinct longitudinal median sulcus.</paragraph>
<paragraph id="9672503D3E6EFFA17C31BCA4DB82F93A" blockId="17.[826,1441,1482,1902]" pageId="17" pageNumber="564">17. Fourth abdominal tergite with erect to suberect setae.</paragraph>
<paragraph id="9672503D3E6EFFA17C31BCE7DC8FF8A4" blockId="17.[826,1441,1482,1902]" pageId="17" pageNumber="564">
18. Body principally yellow, but sometimes partly brown to black in
<taxonomicName id="51CD2BBE3E6EFFA17CE5BD00DC52F8E6" authority="Forel, 1904" authorityName="Forel" authorityYear="1904" box="[1006,1149,1818,1840]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="A4B98C2F3E6EFFA17CE5BD00DC2FF8F9" box="[1006,1024,1819,1840]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17B1BBD01DC52F8E6" box="[1040,1149,1818,1839]" italics="true" pageId="17" pageNumber="564">binghami</emphasis>
</taxonomicName>
group,
<taxonomicName id="51CD2BBE3E6EFFA17BDCBD00DD01F8E6" authorityName="Mayr" authorityYear="1897" box="[1239,1326,1818,1840]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="A4B98C2F3E6EFFA17BDCBD00DCC6F8F9" box="[1239,1257,1819,1840]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17BF2BD01DD01F8E6" box="[1273,1326,1818,1839]" italics="true" pageId="17" pageNumber="564">biroi</emphasis>
</taxonomicName>
group,
<taxonomicName id="51CD2BBE3E6EFFA17A83BD00DBF2F886" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="moatensis">
<emphasis id="A4B98C2F3E6EFFA17A83BD00DDB5F8F9" box="[1416,1434,1819,1840]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17C61BD21DBF2F886" box="[874,989,1850,1871]" italics="true" pageId="17" pageNumber="564">moatensis</emphasis>
</taxonomicName>
group and
<taxonomicName id="51CD2BBE3E6EFFA17B64BD22DD36F887" authorityName="Forel" authorityYear="1911" box="[1135,1305,1849,1871]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E6EFFA17B64BD22DCAEF887" box="[1135,1153,1849,1870]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17B9FBD22DD36F887" box="[1172,1305,1849,1870]" italics="true" pageId="17" pageNumber="564">quadriruga</emphasis>
</taxonomicName>
group, but brown in
<taxonomicName id="51CD2BBE3E6EFFA17CD7BD43DC62F8A4" authorityName="Forel" authorityYear="1912" box="[988,1101,1880,1901]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="564" phylum="Arthropoda" rank="species" species="baduvi">
<emphasis id="A4B98C2F3E6EFFA17CD7BD43DBC1F8A4" box="[988,1006,1880,1901]" italics="true" pageId="17" pageNumber="564">C</emphasis>
.
<emphasis id="A4B98C2F3E6EFFA17CF5BD43DC62F8A4" box="[1022,1101,1880,1901]" italics="true" pageId="17" pageNumber="564">baduvi</emphasis>
</taxonomicName>
group.
</paragraph>
<caption id="C2B200B53E6DFFA27F9BBEE6DBC2FAD8" ID-DOI="http://doi.org/10.5281/zenodo.5358206" ID-Zenodo-Dep="5358206" box="[144,1005,1277,1297]" httpUri="https://zenodo.org/record/5358206/files/figure.png" pageId="18" pageNumber="565" startId="18.[144,223,1277,1296]" targetBox="[147,1422,198,1246]" targetPageId="18">
<paragraph id="9672503D3E6DFFA27F9BBEE6DBC2FAD8" blockId="18.[144,1005,1277,1297]" box="[144,1005,1277,1297]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27F9BBEE6D92AFAD8" bold="true" box="[144,261,1277,1297]" pageId="18" pageNumber="565">Figure 52.</emphasis>
Geographical distribution of the species of the
<taxonomicName id="51CD2BBE3E6DFFA27C02BEE5DB8EFAD8" authorityName="Forel" authorityYear="1911" box="[777,929,1278,1297]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E6DFFA27C02BEE5DB36FAD8" box="[777,793,1278,1297]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C22BEE5DB8EFAD8" box="[809,929,1278,1297]" italics="true" pageId="18" pageNumber="565">quadriruga</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="9672503D3E6DFFA27F9BBF56D950FA48" blockId="18.[144,759,1357,1471]" pageId="18" pageNumber="565">19. Monomorphic in size; intermediate workers occasionally present.</paragraph>
<paragraph id="9672503D3E6DFFA27F9BBF90D984FA76" blockId="18.[144,759,1357,1471]" pageId="18" pageNumber="565">20. Nesting in soil; some species nesting in dead twigs on lower vegetation.</paragraph>
</subSubSection>
<subSubSection id="DED703B63E6DFFA37FBBBFE4D9C1FB9B" lastPageId="19" lastPageNumber="566" pageId="18" pageNumber="565" type="synonymic_list">
<paragraph id="9672503D3E6DFFA27FBBBFE4DAF9F9DD" blockId="18.[176,726,1535,1556]" box="[176,726,1535,1556]" pageId="18" pageNumber="565">
<heading id="CD3AE7513E6DFFA27FBBBFE4DAF9F9DD" box="[176,726,1535,1556]" centered="true" fontSize="8" level="2" pageId="18" pageNumber="565" reason="8">
SYNONYMIC LIST OF ASIAN
<taxonomicName id="51CD2BBE3E6DFFA27EE2BFE4DA5BF9DD" authorityName="Santschi" box="[489,628,1535,1556]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6DFFA27EE2BFE4DA5BF9DD" box="[489,628,1535,1556]" italics="true" pageId="18" pageNumber="565">Orthocrema</emphasis>
</taxonomicName>
SPECIES
</heading>
</paragraph>
<paragraph id="9672503D3E6DFFA27F9BBC3DDDA1FA48" blockId="18.[144,759,1574,1902]" lastBlockId="18.[806,1422,1357,1410]" pageId="18" pageNumber="565">
In this study, five species groups are established within the Asian
<taxonomicName id="51CD2BBE3E6DFFA27E0ABC5FD9ACF990" authorityName="Santschi" box="[257,387,1604,1625]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6DFFA27E0ABC5FD9ACF990" box="[257,387,1604,1625]" italics="true" pageId="18" pageNumber="565">Orthocrema</emphasis>
</taxonomicName>
. The species groups were defined based on putative apomorphies and discrete morphological boundaries. The species groups recognized here will provide a possible basis for new monophyletic taxa. The
<taxonomicName id="51CD2BBE3E6DFFA27FCEBCA4D919F91D" authorityName="Forel" authorityYear="1912" box="[197,310,1727,1748]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="baduvi">
<emphasis id="A4B98C2F3E6DFFA27FCEBCA4D8F8F91D" box="[197,215,1727,1748]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27FECBCA4D919F91D" box="[231,310,1727,1748]" italics="true" pageId="18" pageNumber="565">baduvi</emphasis>
</taxonomicName>
,
<taxonomicName id="51CD2BBE3E6DFFA27E4DBCA4D9F1F91D" box="[326,478,1727,1748]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="binghamii">
<emphasis id="A4B98C2F3E6DFFA27E4DBCA4D977F91D" box="[326,344,1727,1748]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27E63BCA4D9F1F91D" box="[360,478,1727,1748]" italics="true" pageId="18" pageNumber="565">binghamii</emphasis>
</taxonomicName>
and
<taxonomicName id="51CD2BBE3E6DFFA27D10BCA4DA5EF91D" authorityName="Mayr" authorityYear="1897" box="[539,625,1727,1748]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="A4B98C2F3E6DFFA27D10BCA4DA02F91D" box="[539,557,1727,1748]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27D37BCA4DA5EF91D" box="[572,625,1727,1748]" italics="true" pageId="18" pageNumber="565">biroi</emphasis>
</taxonomicName>
groups are relatively easily defined, but the
<taxonomicName id="51CD2BBE3E6DFFA27D00BCC5DA82F93B" authorityName="Forel" authorityYear="1911" box="[523,685,1757,1779]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E6DFFA27D00BCC5DA32F93A" box="[523,541,1758,1779]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27D20BCC6DA82F93B" box="[555,685,1757,1778]" italics="true" pageId="18" pageNumber="565">quadriruga</emphasis>
</taxonomicName>
group is probably not monophyletic as currently defined. The
<taxonomicName id="51CD2BBE3E6DFFA27F9BBD00D932F8F9" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[144,285,1819,1840]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="moatensis">
<emphasis id="A4B98C2F3E6DFFA27F9BBD00D88EF8F9" box="[144,161,1819,1840]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27FA4BD00D932F8F9" box="[175,285,1819,1840]" italics="true" pageId="18" pageNumber="565">moatensis</emphasis>
</taxonomicName>
group was represented by a single species since the position of
<taxonomicName id="51CD2BBE3E6DFFA27E89BD21DA3AF886" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[386,533,1850,1871]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="moatensis">
<emphasis id="A4B98C2F3E6DFFA27E89BD21D9BBF886" box="[386,404,1850,1871]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27EAFBD21DA3AF886" box="[420,533,1850,1871]" italics="true" pageId="18" pageNumber="565">moatensis</emphasis>
</taxonomicName>
in cladograms was always consistent. The taxa should ideally be monophyletic, but also considered as useful units for taxonomic purposes in the case of hyperdiverse genera.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBFB1DC42FA76" blockId="18.[806,1193,1450,1594]" box="[806,1133,1450,1471]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBFB1DC42FA76" box="[806,1133,1450,1471]" italics="true" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBFB1DC0CFA76" authorityName="Forel" authorityYear="1912" box="[806,1059,1450,1471]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="baduvi">Crematogaster baduvi</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBFD2DC16FA17" blockId="18.[806,1193,1450,1594]" box="[806,1081,1480,1502]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBFD2DC1AFA17" authority="Forel, 1912 b" authorityName="Forel" authorityYear="1912" box="[806,1077,1480,1502]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="baduvi">
<emphasis id="A4B98C2F3E6DFFA27C2DBFD2DB17FA17" box="[806,824,1481,1502]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BFD3DBB7FA14" box="[841,920,1480,1501]" italics="true" pageId="18" pageNumber="565">baduvi</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27CAABFD3DC1AFA17" author="Forel A" box="[929,1077,1480,1502]" pageId="18" pageNumber="565" pagination="97 - 112" refId="ref39001" refString="Forel A. 1912 b. Ameisen aus Java beobachtet und gesammelt von Edward Jacobson. 3 Theil. Notes from the Leyden Museum 34: 97 - 112." type="journal article" year="1912">Forel, 1912b</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBFFCDC1FFA35" blockId="18.[806,1193,1450,1594]" box="[806,1072,1511,1533]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBFFCDC1FFA35" bold="true" box="[806,1072,1511,1533]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBFFCDBE9FA34" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[806,966,1511,1533]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="brunensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6DFFA27C2DBFFCDB16FA35" bold="true" box="[806,825,1511,1532]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C41BFFCDBE9FA34" bold="true" box="[842,966,1511,1533]" italics="true" pageId="18" pageNumber="565">brunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6DFFA27CDBBFFCDC1FFA35" box="[976,1072,1511,1532]" pageId="18" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBC1DDC86F9D2" blockId="18.[806,1193,1450,1594]" box="[806,1193,1541,1564]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBC1DDC8AF9D2" authority="Creighton, 1945" authorityName="Creighton" authorityYear="1945" box="[806,1189,1541,1564]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="macracantha">
<emphasis id="A4B98C2F3E6DFFA27C2DBC1DDB17F9D2" box="[806,824,1542,1563]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BC1DDBF0F9D2" box="[841,991,1542,1563]" italics="true" pageId="18" pageNumber="565">macracantha</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27CE3BC1DDC8AF9D2" author="Creighton WS" box="[1000,1189,1541,1564]" pageId="18" pageNumber="565" pagination="109 - 118" refId="ref38438" refString="Creighton WS. 1945. Observations on the subgenus Rhachiocrema with the description of a new species from Borneo. Psyche 52: 109 - 118." type="journal article" year="1945">Creighton, 1945</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBC3FDBD1F9F0" blockId="18.[806,1193,1450,1594]" box="[806,1022,1572,1594]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBC3FDBD1F9F0" bold="true" box="[806,1022,1572,1594]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBC3FDBBBF9F3" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[806,916,1572,1594]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="storki" status="sp. nov.">
<emphasis id="A4B98C2F3E6DFFA27C2DBC3FDB16F9F0" bold="true" box="[806,825,1572,1593]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C41BC3FDBBBF9F3" bold="true" box="[842,916,1572,1594]" italics="true" pageId="18" pageNumber="565">storki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6DFFA27C95BC3FDBD1F9F0" box="[926,1022,1572,1593]" pageId="18" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBC79DCBCF9B1" blockId="18.[806,1171,1634,1748]" box="[806,1171,1634,1656]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBC79DCBCF9B1" box="[806,1171,1634,1656]" italics="true" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBC79DC65F9BE" box="[806,1098,1634,1655]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="binghamii">Crematogaster binghamii</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBC9ADC7DF95F" blockId="18.[806,1171,1634,1748]" box="[806,1106,1665,1687]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBC9ADC61F95F" authority="Forel, 1904" authorityName="Forel" authorityYear="1904" box="[806,1102,1665,1687]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="binghamii">
<emphasis id="A4B98C2F3E6DFFA27C2DBC9ADB17F95F" box="[806,824,1665,1686]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BC9ADB90F95F" box="[841,959,1665,1686]" italics="true" pageId="18" pageNumber="565">binghamii</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27CC3BC9ADC61F95F" author="Forel A" box="[968,1102,1665,1687]" pageId="18" pageNumber="565" pagination="1 - 52" refId="ref38818" refString="Forel A. 1904. Miscellanea myrmecologiques. Revue Suisse de Zoologie 12: 1 - 52." type="journal article" year="1904">Forel, 1904</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBC84DC15F97D" blockId="18.[806,1171,1634,1748]" box="[806,1082,1695,1717]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBC84DC15F97D" bold="true" box="[806,1082,1695,1717]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBC84DBE0F97C" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[806,975,1695,1717]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="brevispina" status="sp. nov.">
<emphasis id="A4B98C2F3E6DFFA27C2DBC84DB16F97D" bold="true" box="[806,825,1695,1716]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BC84DBE0F97C" bold="true" box="[841,975,1695,1717]" italics="true" pageId="18" pageNumber="565">brevispina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6DFFA27CD1BC84DC15F97D" box="[986,1082,1695,1716]" pageId="18" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBCA5DC75F91A" blockId="18.[806,1171,1634,1748]" box="[806,1114,1726,1748]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBCA5DC79F91A" authority="Forel, 1907" authorityName="Forel" authorityYear="1907" box="[806,1110,1726,1748]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="longipilosa">
<emphasis id="A4B98C2F3E6DFFA27C2DBCA5DB17F91A" box="[806,824,1726,1747]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BCA5DBE9F91A" box="[841,966,1726,1747]" italics="true" pageId="18" pageNumber="565">longipilosa</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27CC4BCA5DC79F91A" author="Forel A" box="[975,1110,1726,1748]" pageId="18" pageNumber="565" pagination="1 - 42" refId="ref38872" refString="Forel A. 1907. Formicides du Musee National Hongrois. Annales Historico-Naturales Musei Nationalis Hungarici 5: 1 - 42." type="journal article" year="1907">Forel, 1907</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBCE7DC7CF8DB" blockId="18.[806,1324,1788,1902]" box="[806,1107,1788,1810]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBCE7DC7CF8DB" box="[806,1107,1788,1810]" italics="true" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBCE7DC26F8D8" authorityName="Mayr" authorityYear="1897" box="[806,1033,1788,1809]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="biroi">Crematogaster biroi</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBD00DC3DF8F9" blockId="18.[806,1324,1788,1902]" box="[806,1042,1818,1840]" pageId="18" pageNumber="565">
<taxonomicName id="51CD2BBE3E6DFFA27C2DBD00DC21F8F9" authority="Mayr, 1897" authorityName="Mayr" authorityYear="1897" box="[806,1038,1818,1840]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="A4B98C2F3E6DFFA27C2DBD00DB17F8F9" box="[806,824,1819,1840]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C42BD01DB51F8E6" box="[841,894,1818,1839]" italics="true" pageId="18" pageNumber="565">biroi</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27C83BD00DC21F8F9" author="Mayr G" box="[904,1038,1819,1840]" pageId="18" pageNumber="565" pagination="420 - 436" refId="ref39892" refString="Mayr G. 1897. Formiciden aus Ceylon und Singapur. Termeszetrajzi Fuzetek 20: 420 - 436." type="journal article" year="1897">Mayr, 1897</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBD21DCFFF887" blockId="18.[806,1324,1788,1902]" box="[806,1232,1849,1871]" pageId="18" pageNumber="565">
=
<taxonomicName id="51CD2BBE3E6DFFA27C37BD22DC7CF887" authority="Forel, 1902 a" authorityName="Forel" authorityYear="1902" box="[828,1107,1849,1871]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="species" species="aitkenii">
<emphasis id="A4B98C2F3E6DFFA27C37BD22DB61F887" box="[828,846,1849,1870]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C55BD22DB9AF887" box="[862,949,1849,1870]" italics="true" pageId="18" pageNumber="565">aitkenii</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27CB4BD22DC7CF887" author="Forel A" box="[959,1107,1849,1871]" pageId="18" pageNumber="565" pagination="165 - 249" refId="ref38774" refString="Forel A. 1902 a. Myrmicinae nouveaux de l'Inde et de Ceylan. Revue Suisse de Zoologie 10: 165 - 249." type="journal article" year="1902">Forel, 1902a</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6DFFA27B6ABD22DCFFF887" bold="true" box="[1121,1232,1849,1870]" pageId="18" pageNumber="565">syn. nov.</emphasis>
</paragraph>
<paragraph id="9672503D3E6DFFA27C2DBD43DD03F8A4" blockId="18.[806,1324,1788,1902]" box="[806,1324,1880,1902]" pageId="18" pageNumber="565">
<emphasis id="A4B98C2F3E6DFFA27C2DBD43DB1AF8A4" bold="true" box="[806,821,1880,1901]" pageId="18" pageNumber="565">=</emphasis>
<taxonomicName id="51CD2BBE3E6DFFA27C37BD43DC80F8A4" authority="Forel, 1902 a" authorityName="Forel" authorityYear="1902" box="[828,1199,1880,1902]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="565" phylum="Arthropoda" rank="subSpecies" species="biroi" subSpecies="smythiesii">
<emphasis id="A4B98C2F3E6DFFA27C37BD43DB61F8A4" box="[828,846,1880,1901]" italics="true" pageId="18" pageNumber="565">C</emphasis>
.
<emphasis id="A4B98C2F3E6DFFA27C55BD43DC3FF8A4" box="[862,1040,1880,1901]" italics="true" pageId="18" pageNumber="565">biroi smythiesii</emphasis>
<bibRefCitation id="F25C2DCC3E6DFFA27B11BD43DC80F8A4" author="Forel A" box="[1050,1199,1880,1902]" pageId="18" pageNumber="565" pagination="165 - 249" refId="ref38774" refString="Forel A. 1902 a. Myrmicinae nouveaux de l'Inde et de Ceylan. Revue Suisse de Zoologie 10: 165 - 249." type="journal article" year="1902">Forel, 1902a</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6DFFA27BB7BD43DD03F8A4" bold="true" box="[1212,1324,1880,1901]" pageId="18" pageNumber="565">syn. nov.</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBADEDA67FF13" blockId="19.[164,698,197,493]" box="[164,584,197,219]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBADED89CFF13" bold="true" box="[164,179,197,218]" pageId="19" pageNumber="566">=</emphasis>
<taxonomicName id="51CD2BBE3E6CFFA37FB1BADED9E4FF13" authority="Bharti, 2003" authorityName="Bharti" authorityYear="2003" box="[186,459,197,219]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="urvijae">
<emphasis id="A4B98C2F3E6CFFA37FB1BADED8E3FF13" box="[186,204,197,218]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FD7BADDD903FF12" box="[220,300,198,219]" italics="true" pageId="19" pageNumber="566">urvijae</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E3DBADED9E4FF13" author="Bharti H" box="[310,459,197,219]" pageId="19" pageNumber="566" pagination="85 - 88" refId="ref37846" refString="Bharti H. 2003. A new species of Crematogaster (Hymenoptera; formicidae: myrmicinae) from India. Entomon 28: 85 - 88." type="journal article" year="2003">Bharti, 2003</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37ED3BADEDA67FF13" bold="true" box="[472,584,197,218]" pageId="19" pageNumber="566">syn. nov.</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBAFFD98FFF30" blockId="19.[164,698,197,493]" box="[164,416,228,249]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBAFFD9B3FF30" authority="Emery, 1901" authorityName="Emery" authorityYear="1901" box="[164,412,228,249]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="fritzi">
<emphasis id="A4B98C2F3E6CFFA37FAFBAFFD899FF30" box="[164,182,228,249]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBAFFD8D1FF30" box="[198,254,228,249]" italics="true" pageId="19" pageNumber="566">fritzi</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E03BAFFD9B3FF30" author="Emery C" box="[264,412,228,249]" pageId="19" pageNumber="566" pagination="565 - 580" refId="ref38625" refString="Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbuucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 14: 565 - 580." type="journal article" year="1901">Emery, 1901</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBB19D999FEDE" blockId="19.[164,698,197,493]" box="[164,438,258,280]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBB19D999FEDE" bold="true" box="[164,438,258,280]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB19D963FED1" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,332,258,280]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFBB19D898FEDE" bold="true" box="[164,183,258,279]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCBB19D963FED1" bold="true" box="[199,332,258,280]" italics="true" pageId="19" pageNumber="566">luzonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E5DBB19D999FEDE" box="[342,438,258,279]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBB3AD9B8FE9D" blockId="19.[164,698,197,493]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB3ADA99FEFF" authority="Hosoishi, Yamane &amp; Ogata, 2010" authorityName="Hosoishi, Yamane &amp; Ogata" authorityYear="2010" box="[164,694,289,311]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="masukoi">
<emphasis id="A4B98C2F3E6CFFA37FAFBB3AD899FEFF" box="[164,182,289,310]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBB3AD909FEFF" box="[198,294,289,310]" italics="true" pageId="19" pageNumber="566">masukoi</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E3BBB3ADA99FEFF" author="Hosoishi S &amp; Yamane S &amp; Ogata K" box="[304,694,289,311]" pageId="19" pageNumber="566" pagination="345 - 350" refId="ref39498" refString="Hosoishi S, Yamane S, Ogata K. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13: 345 - 350." type="journal article" year="2010">Hosoishi, Yamane &amp; Ogata, 2010</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37FAFBB24D9B8FE9D" bold="true" box="[164,407,319,341]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB24D902FE9C" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,301,319,341]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="ocellata" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFBB24D898FE9D" bold="true" box="[164,183,319,340]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCBB24D902FE9C" bold="true" box="[199,301,319,341]" italics="true" pageId="19" pageNumber="566">ocellata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E3CBB24D9B8FE9D" box="[311,407,319,340]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBB45D9E9FEBA" blockId="19.[164,698,197,493]" box="[164,454,350,372]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB45D9EEFEBA" authority="Forel, 1900" authorityName="Forel" authorityYear="1900" box="[164,449,350,372]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="osakensis">
<emphasis id="A4B98C2F3E6CFFA37FAFBB45D899FEBA" box="[164,182,350,371]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBB45D91EFEBA" box="[198,305,350,371]" italics="true" pageId="19" pageNumber="566">osakensis</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E30BB45D9EEFEBA" author="Forel A" box="[315,449,350,372]" pageId="19" pageNumber="566" pagination="267 - 287" refId="ref38736" refString="Forel A. 1900. Fourmis du Japon. Nids en toile. Strongylognathus huberti et voisins. Fourmiliere triple. Cyphomyrmex wheeleri. Fourmis importees. Mittheilungen der Schweizerishen Entomologischen Gesellschaft 10: 267 - 287." type="journal article" year="1900">Forel, 1900</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBB66D9C0FE5B" blockId="19.[164,698,197,493]" box="[164,495,380,402]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB66D9C4FE5B" authority="Hosoishi, 2009" authorityName="Hosoishi" authorityYear="2009" box="[164,491,380,402]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="A4B98C2F3E6CFFA37FAFBB66D899FE5B" box="[164,182,381,402]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBB67D91BFE58" box="[198,308,380,401]" italics="true" pageId="19" pageNumber="566">reticulata</emphasis>
Hosoishi, 2009
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBB80D9F0FE79" blockId="19.[164,698,197,493]" box="[164,479,411,433]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBB80D9F4FE79" authority="Forel, 1912 a" authorityName="Forel" authorityYear="1912" box="[164,475,411,433]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="schimmeri">
<emphasis id="A4B98C2F3E6CFFA37FAFBB80D899FE79" box="[164,182,411,432]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBB80D912FE79" box="[198,317,411,432]" italics="true" pageId="19" pageNumber="566">schimmeri</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E4CBB80D9F4FE79" author="Forel A" box="[327,475,411,433]" pageId="19" pageNumber="566" pagination="45 - 81" refId="ref38980" refString="Forel A. 1912 a. H. Sauter's Formosa-Ausbeute: Formicidae. Entomologische Mitteilungen 1: 45 - 81." type="journal article" year="1912">Forel, 1912a</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBBA1D9AAFE06" blockId="19.[164,698,197,493]" box="[164,389,441,464]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBBA1D9AEFE06" authority="Forel, 1905" authorityName="Forel" authorityYear="1905" box="[164,385,441,464]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="udo">
<emphasis id="A4B98C2F3E6CFFA37FAFBBA1D899FE06" box="[164,182,442,463]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDBBA1D8DFFE06" box="[198,240,442,463]" italics="true" pageId="19" pageNumber="566">udo</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37FF1BBA1D9AEFE06" author="Forel A" box="[250,385,441,464]" pageId="19" pageNumber="566" pagination="1 - 26" refId="ref38836" refString="Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin, 1904. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten [Mitteilungen aus dem Naturhistorischen Museum] 22: 1 - 26." type="journal article" year="1905">Forel, 1905</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBBC3D946FE24" blockId="19.[164,698,197,493]" box="[164,361,472,493]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBBC3D946FE24" bold="true" box="[164,361,472,493]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBBC3D8D0FE24" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,255,472,493]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="vieti" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFBBC3D898FE24" bold="true" box="[164,183,472,493]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCBBC3D8D0FE24" bold="true" box="[199,255,472,493]" italics="true" pageId="19" pageNumber="566">vieti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E02BBC3D946FE24" box="[265,361,472,493]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB80DDA22FDE2" blockId="19.[164,525,534,585]" box="[164,525,534,555]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFB80DDA22FDE2" box="[164,525,534,555]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB80DD9EDFDE2" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,450,534,555]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="moatensis">Crematogaster moatensis</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB82FD99FFD80" blockId="19.[164,525,534,585]" box="[164,432,564,585]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFB82FD99FFD80" bold="true" box="[164,432,564,585]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB82FD969FD80" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,326,564,585]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="moatensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFB82FD969FD80" bold="true" box="[164,326,564,585]" italics="true" pageId="19" pageNumber="566">C. moatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E5BB82FD99FFD80" box="[336,432,564,585]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB869DA31FD4E" blockId="19.[164,674,625,983]" box="[164,542,625,647]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFB869DA31FD4E" box="[164,542,625,647]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB869D9FCFD4F" authorityName="Forel" authorityYear="1911" box="[164,467,625,647]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="quadriruga">Crematogaster quadriruga</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB88BDA4CFD6C" blockId="19.[164,674,625,983]" box="[164,611,656,678]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB88BD9CEFD6C" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[164,481,656,678]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="bandarensis">
<emphasis id="A4B98C2F3E6CFFA37FAFB88BD899FD6C" box="[164,182,656,677]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB88BD97EFD6C" box="[198,337,656,677]" italics="true" pageId="19" pageNumber="566">bandarensis</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E50B88BD9CEFD6C" author="Forel A" box="[347,481,656,678]" pageId="19" pageNumber="566" pagination="1 - 148" refId="ref39055" refString="Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien, ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. 2. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel- Reepen in den Jahren 1911 - 1912. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 36: 1 - 148." type="journal article" year="1913">Forel, 1913</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37EE4B88BDA4CFD6C" bold="true" box="[495,611,656,677]" pageId="19" pageNumber="566">stat. nov.</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB8B4D980FD2B" blockId="19.[164,674,625,983]" pageId="19" pageNumber="566">
=
<taxonomicName id="51CD2BBE3E6CFFA37FB2B8B4DA0AFD0D" authority="Santschi, 1928" authorityName="Santschi" authorityYear="1928" box="[185,549,686,708]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="subSpecies" species="biroi" subSpecies="andelis">
<emphasis id="A4B98C2F3E6CFFA37FB2B8B4D8E4FD0D" box="[185,203,687,708]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FD7B8B5D942FD0A" box="[220,365,686,707]" italics="true" pageId="19" pageNumber="566">biroi andelis</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E7CB8B5DA0AFD0D" author="Santschi F" box="[375,549,686,708]" pageId="19" pageNumber="566" pagination="119 - 140" refId="ref40343" refString="Santschi F. 1928. Fourmis de Sumatra, recoltees par Mr. J. B. Corporaal. Tijdschrift Voor Entomologie 71: 119 - 140." type="journal article" year="1928">Santschi, 1928</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37D39B8B4D980FD2B" bold="true" pageId="19" pageNumber="566">
syn. nov.
<taxonomicName id="51CD2BBE3E6CFFA37FAFB8D6D96AFD2A" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,325,717,739]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="celebensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFB8D6D898FD2B" bold="true" box="[164,183,717,738]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCB8D6D96AFD2A" bold="true" box="[199,325,717,739]" italics="true" pageId="19" pageNumber="566">celebensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E44B8D6D980FD2B" box="[335,431,717,738]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB8F7D9CAFCC8" blockId="19.[164,674,625,983]" box="[164,485,747,770]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB8F7D9CEFCC8" authority="Menozzi, 1935" authorityName="Menozzi" authorityYear="1935" box="[164,481,747,770]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="gavapiga">
<emphasis id="A4B98C2F3E6CFFA37FAFB8F7D899FCC8" box="[164,182,748,769]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB8F7D901FCC8" box="[198,302,748,769]" italics="true" pageId="19" pageNumber="566">gavapiga</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E33B8F7D9CEFCC8" author="Menozzi C" box="[312,481,747,770]" pageId="19" pageNumber="566" pagination="103 - 116" refId="ref39911" refString="Menozzi C. 1935. Formiche indo-australiane del genere Crematogaster Lund raccolte da W. Karawaiew. Konowia 14: 103 - 116." type="journal article" year="1935">Menozzi, 1935</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB910D9F1FCE9" blockId="19.[164,674,625,983]" box="[164,478,778,800]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB910D9F5FCE9" authority="Menozzi, 1935" authorityName="Menozzi" authorityYear="1935" box="[164,474,778,800]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="A4B98C2F3E6CFFA37FAFB910D899FCE9" box="[164,182,779,800]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB910D908FCE9" box="[198,295,779,800]" italics="true" pageId="19" pageNumber="566">javanica</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E3AB911D9F5FCE9" author="Menozzi C" box="[305,474,778,800]" pageId="19" pageNumber="566" pagination="103 - 116" refId="ref39911" refString="Menozzi C. 1935. Formiche indo-australiane del genere Crematogaster Lund raccolte da W. Karawaiew. Konowia 14: 103 - 116." type="journal article" year="1935">Menozzi, 1935</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB932D981FCF7" blockId="19.[164,674,625,983]" box="[164,430,809,831]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB932D985FCF7" authority="Forel, 1911 a" authorityName="Forel" authorityYear="1911" box="[164,426,809,831]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="myops">
<emphasis id="A4B98C2F3E6CFFA37FAFB932D899FCF7" box="[164,182,809,830]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB932D921FCF7" box="[198,270,809,830]" italics="true" pageId="19" pageNumber="566">myops</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E1CB932D985FCF7" author="Forel A" box="[279,426,809,831]" pageId="19" pageNumber="566" pagination="23 - 62" refId="ref38894" refString="Forel A. 1911 a. Fourmis de Borneo, Singapore, Ceylan, etc. Revue Suisse de Zoologie 19: 23 - 62." type="journal article" year="1911">Forel, 1911a</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB95CD9C7FC94" blockId="19.[164,674,625,983]" box="[164,488,839,861]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFB95CD9C7FC94" bold="true" box="[164,488,839,861]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB95CD951FC94" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,382,839,861]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFB95CD898FC95" bold="true" box="[164,183,839,860]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCB95CD951FC94" bold="true" box="[199,382,839,861]" italics="true" pageId="19" pageNumber="566">philippinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E83B953D9C7FC94" box="[392,488,840,861]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB97DDA4CFCB2" blockId="19.[164,674,625,983]" box="[164,611,870,892]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB97DD9CDFCB2" authority="Forel, 1911 c" authorityName="Forel" authorityYear="1911" box="[164,482,870,892]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E6CFFA37FAFB97DD899FCB2" box="[164,182,870,891]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB97DD967FCB2" box="[198,328,870,891]" italics="true" pageId="19" pageNumber="566">quadriruga</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E59B97DD9CDFCB2" author="Forel A" box="[338,482,870,892]" pageId="19" pageNumber="566" pagination="445 - 459" refId="ref38945" refString="Forel A. 1911 c. Sur le genre Metapone n. g. nouveau groupe des formicides et sur quelques autres formes nouvelles. Revue Suisse de Zoologie 19: 445 - 459." type="journal article" year="1911">Forel, 1911c</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37EFBB97DDA4CFCB2" bold="true" box="[496,611,870,891]" pageId="19" pageNumber="566">stat. nov.</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB99ED9CBFC53" blockId="19.[164,674,625,983]" box="[164,484,901,922]" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFB99ED9F0FC53" authority="Terayama, 1999" authorityName="Terayama" authorityYear="1999" box="[164,479,901,922]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E6CFFA37FAFB99ED899FC53" box="[164,182,901,922]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCDB99ED936FC53" box="[198,281,901,922]" italics="true" pageId="19" pageNumber="566">suehiro</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E28B99ED9F0FC53" author="Terayama M" box="[291,479,901,922]" pageId="19" pageNumber="566" pagination="726 - 727" refId="ref40436" refString="Terayama M. 1999. Description and record of a new and a little known ants from Japan (Formicidae). In: Yamane SK, Ikudome S, Terayama M, eds. Identification guide to the aculeata of the Nansei Islands, Japan. Sapporo: Hokkaido University Press, 726 - 727." type="book chapter" year="1999">Terayama, 1999</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB9BFD9D5FC1E" blockId="19.[164,674,625,983]" pageId="19" pageNumber="566">
=
<taxonomicName id="51CD2BBE3E6CFFA37FB2B9BFD9DBFC71" authority="Terayama, 2013" authorityName=", Terayama" authorityYear="2013" box="[185,500,931,953]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="miroku">
<emphasis id="A4B98C2F3E6CFFA37FB2B9BFD8E4FC70" box="[185,203,932,953]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FD7B9B8D901FC71" box="[220,302,931,952]" italics="true" pageId="19" pageNumber="566">miroku</emphasis>
<bibRefCitation id="F25C2DCC3E6CFFA37E33B9BFD9DBFC71" author="Terayama M" box="[312,500,931,953]" pageId="19" pageNumber="566" pagination="1 - 24" refId="ref40527" refString="Terayama M. 2013. Additions to knowledge of the ant fauna of Japan (Hymenoptera: Formicidae). Memoirs of the Myrmecological Society of Japan 3: 1 - 24." type="journal article" year="2013">Terayama, 2013</bibRefCitation>
</taxonomicName>
.
<emphasis id="A4B98C2F3E6CFFA37D0AB9B8D9D5FC1E" bold="true" pageId="19" pageNumber="566">
syn. nov.
<taxonomicName id="51CD2BBE3E6CFFA37FAFB9D9D9BFFC1E" authority="Hosoishi &amp; Ogata, 2016" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[164,400,961,983]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="sundalandensis" status="sp. nov.">
<emphasis id="A4B98C2F3E6CFFA37FAFB9D9D898FC1E" bold="true" box="[164,183,962,983]" italics="true" pageId="19" pageNumber="566">C</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37FCCB9DAD9BFFC1E" bold="true" box="[199,400,961,983]" italics="true" pageId="19" pageNumber="566">sundalandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8A31543E6CFFA37E92B9D9D9D5FC1E" box="[409,506,962,983]" pageId="19" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFB9E4D9C1FB9B" blockId="19.[164,779,1023,1106]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFB9E4DAECFBDC" box="[164,707,1023,1045]" italics="true" pageId="19" pageNumber="566">
The species groups of Asian
<taxonomicName id="51CD2BBE3E6CFFA37EECB9E4DA41FBDD" authorityName="Santschi" box="[487,622,1023,1044]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">Orthocrema</taxonomicName>
species
</emphasis>
Apomorphies of the species groups obtained by cladistic analysis are given in italics.
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E6CFFA37FAFBE61DC0DFBB4" pageId="19" pageNumber="566" type="description">
<paragraph id="9672503D3E6CFFA37FAFBE61D9C5FB46" blockId="19.[164,779,1145,1443]" box="[164,490,1145,1167]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBE61D9C5FB46" box="[164,490,1145,1167]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBE61D98FFB47" authorityName="Forel" authorityYear="1912" box="[164,416,1145,1167]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="baduvi">Crematogaster baduvi</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBE83DAA4FAE1" blockId="19.[164,779,1145,1443]" pageId="19" pageNumber="566">
Relatively long scape (SI 98–118). Basal flagellar segment (antennal segment III) longer than broad. Propodeal spines long and
<emphasis id="A4B98C2F3E6CFFA37ECCBECEDA52FB23" box="[455,637,1237,1258]" italics="true" pageId="19" pageNumber="566">directed laterally</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37D80BECEDA60FAC0" italics="true" pageId="19" pageNumber="566">Postpetiolar dorsum highly convex in lateral view</emphasis>
;
<emphasis id="A4B98C2F3E6CFFA37D54BEEFDAABFAE1" italics="true" pageId="19" pageNumber="566">postpetiole distinctly higher than petiole in lateral view</emphasis>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FB7BF2AD8D4FA6A" blockId="19.[164,779,1145,1443]" pageId="19" pageNumber="566">
This species group is easily distinguished by the propodeal spines directed laterally and highly convex postpetiolar dorsum from other Asian
<taxonomicName id="51CD2BBE3E6CFFA37D74BF75DB25FA4A" authorityName="Santschi" box="[639,778,1390,1411]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6CFFA37D74BF75DB25FA4A" box="[639,778,1390,1411]" italics="true" pageId="19" pageNumber="566">Orthocrema</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBFD1DA3EFA29" blockId="19.[164,779,1482,1779]" box="[164,529,1482,1504]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBFD1DA3EFA29" box="[164,529,1482,1504]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBFD1D9E8FA16" authorityName="Forel" authorityYear="1904" box="[164,455,1482,1503]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="binghamii">Crematogaster binghamii</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBFF2DAD9F9B1" blockId="19.[164,779,1482,1779]" pageId="19" pageNumber="566">
Basal flagellar segment (antennal segment III) longer than broad.
<emphasis id="A4B98C2F3E6CFFA37E38BC13DAA1F9F2" italics="true" pageId="19" pageNumber="566">Posterior margins of mesonotum forming short triangle-shaped process in lateral view</emphasis>
. Propodeal spines undeveloped, or developed and directed posteriorly.
<emphasis id="A4B98C2F3E6CFFA37FFBBC78DAC0F9B1" box="[240,751,1635,1657]" italics="true" pageId="19" pageNumber="566">Standing pilosity abundant on body surface</emphasis>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37FB7BC9ADA74F93A" blockId="19.[164,779,1482,1779]" pageId="19" pageNumber="566">
This species group is easily distinguished by the short triangle-shaped process on posterior margins of mesonotum and long and abundant standing pilosity from other Asian
<taxonomicName id="51CD2BBE3E6CFFA37E78BCC6D9D5F93B" authorityName="Santschi" box="[371,506,1757,1778]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6CFFA37E78BCC6D9D5F93B" box="[371,506,1757,1778]" italics="true" pageId="19" pageNumber="566">Orthocrema</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBD00D9FEF8F9" blockId="19.[164,779,1818,1902]" box="[164,465,1818,1840]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37FAFBD00D9FEF8F9" box="[164,465,1818,1840]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37FAFBD00D9A9F8E6" authorityName="Mayr" authorityYear="1897" box="[164,390,1818,1840]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="biroi">Crematogaster biroi</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37FAFBD22DBBEFE9C" blockId="19.[164,779,1818,1902]" lastBlockId="19.[826,1441,197,463]" pageId="19" pageNumber="566">
Relatively short scape (SI 70–89). Basal flagellar segment (antennal segment III) broader than long.
<emphasis id="A4B98C2F3E6CFFA37C31BADEDB95FF30" italics="true" pageId="19" pageNumber="566">Anterior margin of pronotal collar almost straight in dorsal view</emphasis>
.
<emphasis id="A4B98C2F3E6CFFA37CC3BAF8DBA0FEDE" italics="true" pageId="19" pageNumber="566">Dorsal surface of propodeum striated with rugulae</emphasis>
. Mesosoma sculptured. Subpetiolar process developed acutely.
<emphasis id="A4B98C2F3E6CFFA37B0CBB3BDBA5FE9D" italics="true" pageId="19" pageNumber="566">Subpostpetiolar process developed acutely</emphasis>
.
</paragraph>
<paragraph id="9672503D3E6CFFA37C58BB45DBBDFE06" blockId="19.[826,1441,197,463]" pageId="19" pageNumber="566">
This species group is easily distinguished by the subpostpetiolar process developed acutely and sculptured surface of mesosoma from other Asian
<taxonomicName id="51CD2BBE3E6CFFA37A2ABB80DD8EFE79" authorityName="Santschi" box="[1313,1441,411,432]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6CFFA37A2ABB80DD8EFE79" box="[1313,1441,411,432]" italics="true" pageId="19" pageNumber="566">Orthocrema</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="9672503D3E6CFFA37C31BBE6DC8CFDDB" blockId="19.[826,1441,509,806]" box="[826,1187,509,530]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37C31BBE6DC8CFDDB" box="[826,1187,509,530]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37C31BBE6DC76FDDB" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[826,1113,509,530]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="moatensis">Crematogaster moatensis</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37C31B807DBBCFD65" blockId="19.[826,1441,509,806]" pageId="19" pageNumber="566">Two apical flagellar segments not differentiated in coloration. Pronotum distinctly higher than pronotal collar. Metanotal groove not covered by lamellate ridges. Propodeal spiracles oval. Petiole long and slender.</paragraph>
<paragraph id="9672503D3E6CFFA37C58B8AFDC8BFCEF" blockId="19.[826,1441,509,806]" pageId="19" pageNumber="566">
This species group is unique in having the metanotal groove not covered by lamellate ridges, oval-shaped propodeal spiracles and long and slender petiole from other Asian
<taxonomicName id="51CD2BBE3E6CFFA37CC1B90BDC7EFCEC" authorityName="Santschi" box="[970,1105,784,805]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6CFFA37CC1B90BDC7EFCEC" box="[970,1105,784,805]" italics="true" pageId="19" pageNumber="566">Orthocrema</emphasis>
</taxonomicName>
fauna.
</paragraph>
<paragraph id="9672503D3E6CFFA37C31B94FDC9BFCA0" blockId="19.[826,1441,851,1149]" box="[826,1204,851,873]" pageId="19" pageNumber="566">
<emphasis id="A4B98C2F3E6CFFA37C31B94FDC9BFCA0" box="[826,1204,851,873]" italics="true" pageId="19" pageNumber="566">
<taxonomicName id="51CD2BBE3E6CFFA37C31B94FDC45FCA1" authorityName="Forel" authorityYear="1911" box="[826,1130,851,873]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="566" phylum="Arthropoda" rank="species" species="quadriruga">Crematogaster quadriruga</taxonomicName>
group
</emphasis>
</paragraph>
<paragraph id="9672503D3E6CFFA37C31B969DC07FBCB" blockId="19.[826,1441,851,1149]" pageId="19" pageNumber="566">Relatively long scape (SI 81–100). Basal flagellar segment (antennal segment III) broader than long. Subpetiolar process developed. Subpostpetiolar portion wholly convex in lateral view. Mesosoma generally smooth and shining.</paragraph>
<paragraph id="9672503D3E6CFFA37C58BE10DC0DFBB4" blockId="19.[826,1441,851,1149]" pageId="19" pageNumber="566">
This species group is distinguished by the subpostpetiolar portion wholly convex and smooth and shining surface of mesosoma from other Asian
<taxonomicName id="51CD2BBE3E6CFFA37C31BE7CDBEEFBB5" authorityName="Santschi" box="[826,961,1127,1148]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E6CFFA37C31BE7CDBEEFBB5" box="[826,961,1127,1148]" italics="true" pageId="19" pageNumber="566">Orthocrema</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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