290 lines
31 KiB
XML
290 lines
31 KiB
XML
<document id="C086D4005B8E643841779F0305197B89" ID-DOI="10.11646/phytotaxa.307.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13690119" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725504643557" checkinUser="felipe" docAuthor="Moonlight, P. W. & Jara-Muñoz, A." docDate="2017" docId="074FC101DF7EFF81FF6653DC4CF5492C" docLanguage="en" docName="phytotaxa.307.1.1.pdf" docOrigin="Phytotaxa 307 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.307.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia tepuiensis Moonlight & Jara-Muñoz 2017, spec. nov." docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="FB76B979DF6FFF90FFEE534348014F53" masterDocTitle="A revision and recircumscription of Begonia Section Pilderia including one new species" masterLastPageNumber="22" masterPageNumber="1" pageNumber="18" updateTime="1725505101269" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="1D08C4400C0C450BE51118220C4AB5B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9A52866D9FFBA15307130C68A7BBC363">
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<mods:title id="CD10C4E1D16CE55877D3A07F9508234A">A revision and recircumscription of Begonia Section Pilderia including one new species</mods:title>
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</mods:titleInfo>
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<mods:name id="A68B395FABBF38B57E6761FDD0E4021E" type="personal">
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<mods:role id="A5FC16DB85B43A05C60E24E67DB79691">
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<mods:roleTerm id="125185C130B4351C6F7D5C34FE100859">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="11C3E71303AF50B2B19D4D51BC6603E1">Moonlight, P. W.</mods:namePart>
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<mods:affiliation id="E31A03FF2014239E87B5C3D515DD035A">Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, G 12 8 QQ, UK</mods:affiliation>
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</mods:name>
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<mods:name id="7299A8B4CFB43D1F050297E721FCE53A" type="personal">
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<mods:roleTerm id="A7CE838C04DD122833B39C7412D4BE2B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E19DBBC0D99B16DC4082EAA276BDE2DD">Jara-Muñoz, A.</mods:namePart>
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<mods:typeOfResource id="B8C457C34F0C6242FEC16680D455AA71">text</mods:typeOfResource>
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<mods:title id="E6B53F5BC56FADC815CC4DE92B4E496C">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="28C45A19E35D5A6A55291E25B0F6AF65">
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<mods:date id="FFB35BD10127BB77E0184745B1F3DFEF">2017</mods:date>
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<mods:number id="A840494109852E91D48C6F1C286292FF">2017-05-19</mods:number>
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<mods:detail id="C382C3FB5C03784BA7867219893FE2FE" type="volume">
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<mods:number id="4613A16D56EE906C5AE414DC40D625E8">307</mods:number>
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<mods:number id="0E56696F427F88AD38DB570C28C709C4">1</mods:number>
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<mods:start id="1AE8D2F77C632C15BBC650F57E7F0D84">1</mods:start>
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<mods:url id="9497AE0FBBB67FB471BE0E6B3C54292B">http://dx.doi.org/10.11646/phytotaxa.307.1.1</mods:url>
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<mods:classification id="E038D42CA044893F119578ADBEA4BC6B">journal article</mods:classification>
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<mods:identifier id="87E42CE4066C6432AC23CB241EAA9EC5" type="DOI">10.11646/phytotaxa.307.1.1</mods:identifier>
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<treatment id="074FC101DF7EFF81FF6653DC4CF5492C" ID-DOI="http://doi.org/10.5281/zenodo.13690143" ID-Zenodo-Dep="13690143" LSID="urn:lsid:plazi:treatment:074FC101DF7EFF81FF6653DC4CF5492C" httpUri="http://treatment.plazi.org/id/074FC101DF7EFF81FF6653DC4CF5492C" lastPageNumber="18" pageId="17" pageNumber="18">
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<subSubSection id="C7FC239CDF7EFF81FF6653DC4B804FEB" box="[136,897,158,184]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8F597017DF7EFF81FF6653DC4B804FEB" blockId="17.[136,1452,158,471]" box="[136,897,158,184]" pageId="17" pageNumber="18">
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<heading id="D411C77BDF7EFF81FF6653DC4B804FEB" box="[136,897,158,184]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
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<emphasis id="BD92AC05DF7EFF81FF6653DC489C4FEB" bold="true" box="[136,157,159,184]" pageId="17" pageNumber="18">6.</emphasis>
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<taxonomicName id="48E60B94DF7EFF81FF4A53DD4A4C4FEB" ID-CoL="7SHS9" authority="Moonlight & Jara" authorityName="Moonlight & Jara-Muñoz" authorityYear="2017" box="[164,589,158,184]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tepuiensis" status="sp. nov.">
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<emphasis id="BD92AC05DF7EFF81FF4A53DD497E4FEB" bold="true" box="[164,383,158,184]" italics="true" pageId="17" pageNumber="18">Begonia tepuiensis</emphasis>
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Moonlight & Jara
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</taxonomicName>
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<emphasis id="BD92AC05DF7EFF81FDBA53DD4AC24FEB" bold="true" box="[596,707,158,184]" italics="true" pageId="17" pageNumber="18">
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<taxonomicNameLabel id="A6A1117EDF7EFF81FDBA53DD4AC24FEB" box="[596,707,158,184]" pageId="17" pageNumber="18" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<emphasis id="BD92AC05DF7EFF81FD3A53DC4B1F4FEB" bold="true" box="[724,798,159,184]" pageId="17" pageNumber="18">
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<figureCitation id="17DD6C92DF7EFF81FD3A53DC4B194FEB" box="[724,792,159,184]" captionStart="FIGURE 6" captionStartId="18.[136,229,1953,1974]" captionTargetBox="[252,1327,214,1916]" captionTargetId="figure-17@18.[236,1351,190,1929]" captionTargetPageId="18" captionText="FIGURE 6. Begonia tepuiensis Moonlight & Jara. A. Habit, flowering top; B. Habit, shoot with rooting nodes; C. Female flower (side view); D. Stigma (front view); E. Stigma (back view); F.Bract; G. Detail of leaf hairs, upper lamina; H. Detail of leaf hairs, lower lamina. Drawn from type collection R.S. Cowen & J.J. Wurdack 31443 by Claire Banks." figureDoi="http://doi.org/10.5281/zenodo.13690133" httpUri="https://zenodo.org/record/13690133/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
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.
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</emphasis>
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,
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<figureCitation id="17DD6C92DF7EFF81FCC253DC4B764FEB" box="[812,887,159,184]" captionStart="FIGURE 7" captionStartId="19.[136,229,1953,1974]" captionTargetBox="[180,1403,195,1928]" captionTargetId="figure-14@19.[180,1407,190,1928]" captionTargetPageId="19" captionText="FIGURE 7. Distributions of species in Begonia sect. Pilderia (Klotzsch) A.DC. A. B. buddleiifolia A.DC.; B. B. glandulifera Griseb.; C. B. humillima L.B.Sm. & Wassh.; D. B. jenmanii Tutin; E. B. mariannensis Wassh. & T.McClellan; F.B. tepuiensis Moonlight & Jara." figureDoi="http://doi.org/10.5281/zenodo.13690135" httpUri="https://zenodo.org/record/13690135/files/figure.png" pageId="17" pageNumber="18">
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<emphasis id="BD92AC05DF7EFF81FCC253DC4B764FEB" bold="true" box="[812,887,159,184]" pageId="17" pageNumber="18">Fig. 7f</emphasis>
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</figureCitation>
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)
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C7FC239CDF7EFF81FF6653854C034ADA" pageId="17" pageNumber="18" type="description">
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<paragraph id="8F597017DF7EFF81FF665385491D4EDC" blockId="17.[136,1452,158,471]" pageId="17" pageNumber="18">
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<taxonomicName id="48E60B94DF7EFF81FF665385493E4F88" authorityName="Moonlight & Jara-Muñoz" authorityYear="2017" box="[136,319,198,219]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tepuiensis">
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<emphasis id="BD92AC05DF7EFF81FF665385493E4F88" box="[136,319,198,219]" italics="true" pageId="17" pageNumber="18">Begonia tepuiensis</emphasis>
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||
</taxonomicName>
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||
is most similar to
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<taxonomicName id="48E60B94DF7EFF81FE1D53854A8B4F88" box="[499,650,198,219]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
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<emphasis id="BD92AC05DF7EFF81FE1D53854A8B4F88" box="[499,650,198,219]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
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</taxonomicName>
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but is distinguished from that species by its unbranched habit (
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<taxonomicName id="48E60B94DF7EFF81FB0153854D844F88" box="[1263,1413,198,219]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
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<emphasis id="BD92AC05DF7EFF81FB0153854D844F88" box="[1263,1413,198,219]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
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</taxonomicName>
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has frequently-branching stems); larger leaves to
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||
<quantity id="481EDDF2DF7EFF81FD9253A94AB84FAC" box="[636,697,234,255]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="18" unit="cm" value="18.0">18 cm</quantity>
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||
(those of
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||
<taxonomicName id="48E60B94DF7EFF81FCCE53A94BB94FAC" box="[800,952,234,255]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
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||
<emphasis id="BD92AC05DF7EFF81FCCE53A94BB94FAC" box="[800,952,234,255]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
|
||
</taxonomicName>
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||
rarely exceed
|
||
<quantity id="481EDDF2DF7EFF81FBA553A94C8D4FAC" box="[1099,1164,234,255]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="cm" value="12.0">12 cm</quantity>
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||
); its internodes, petioles, and inflorescences that are densely lanate throughout (those of
|
||
<taxonomicName id="48E60B94DF7EFF81FD15524D4B904E70" box="[763,913,270,291]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
|
||
<emphasis id="BD92AC05DF7EFF81FD15524D4B904E70" box="[763,913,270,291]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
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||
</taxonomicName>
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||
are sparsely to densely lanate); its lateral inflorescence with only cymose lateral branches branching one time (lower lateral branches in
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||
<taxonomicName id="48E60B94DF7EFF81FC3252714C754E14" box="[988,1140,306,327]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
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<emphasis id="BD92AC05DF7EFF81FC3252714C754E14" box="[988,1140,306,327]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
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||
</taxonomicName>
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||
are tyrsoid with lateral cymose branches branching two to four times); and its longer, lanceolate bracteoles from
|
||
<quantity id="481EDDF2DF7EFF81FC2252154C2F4E38" box="[972,1070,342,363]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="17" pageNumber="18" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8 to 9 mm</quantity>
|
||
(those of
|
||
<taxonomicName id="48E60B94DF7EFF81FB6152154D244E38" box="[1167,1317,342,363]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="buddleiifolia">
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||
<emphasis id="BD92AC05DF7EFF81FB6152154D244E38" box="[1167,1317,342,363]" italics="true" pageId="17" pageNumber="18">B. buddleiifolia</emphasis>
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||
</taxonomicName>
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||
are ovate and to
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||
<quantity id="481EDDF2DF7EFF81FF38523949104EDC" box="[214,273,378,399]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
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||
).
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</paragraph>
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||
<paragraph id="8F597017DF7EFF81FF6652DD4B914E84" blockId="17.[136,1452,158,471]" pageId="17" pageNumber="18">
|
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Type:
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<collectingCountry id="F7F13087DF7EFF81FF2D52DD49544EE0" box="[195,341,414,435]" name="Venezuela" pageId="17" pageNumber="18">VENEZUELA</collectingCountry>
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||
.
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<collectingRegion id="4D22BEF5DF7EFF81FEB252DD49C44EE0" box="[348,453,414,435]" country="Venezuela" name="Amazonas" pageId="17" pageNumber="18">Amazonas</collectingRegion>
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: Serranía Parú, Cerro Parú: top camp to caño camp, alt. ca.
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<quantity id="481EDDF2DF7EFF81FC1952DD4C404EE0" box="[1015,1089,414,435]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="m" value="1200.0">1200 m</quantity>
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||
,
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<date id="FB5856D7DF7EFF81FBA452DD4CF44EE0" box="[1098,1269,414,435]" pageId="17" pageNumber="18" value="1951-02-13">13 February 1951</date>
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||
,
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<emphasis id="BD92AC05DF7EFF81FB1052DD49554E84" italics="true" pageId="17" pageNumber="18">R.S. Cowen & J.J. Wurdack 31443</emphasis>
|
||
(
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||
<typeStatus id="505DCEB5DF7EFF81FE8E528149B74E84" box="[352,438,450,471]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
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||
MO [MO-036500],
|
||
<typeStatus id="505DCEB5DF7EFF81FD9552814AC14E84" box="[635,704,450,471]" pageId="17" pageNumber="18" type="isotype">isotype</typeStatus>
|
||
NY [NY02497219]).
|
||
</paragraph>
|
||
<paragraph id="8F597017DF7EFF81FF6651434C034ADA" blockId="17.[136,1452,512,1663]" pageId="17" pageNumber="18">
|
||
<emphasis id="BD92AC05DF7EFF81FF66514348D14D4A" box="[136,208,512,537]" italics="true" pageId="17" pageNumber="18">Plants</emphasis>
|
||
caulescent, rhizomatous herbs,
|
||
<quantity id="481EDDF2DF7EFF81FDC651434A954D4A" box="[552,660,512,537]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="17" pageNumber="18" unit="cm" value="27.5" valueMax="40.0" valueMin="15.0">15–40 cm</quantity>
|
||
high;
|
||
<emphasis id="BD92AC05DF7EFF81FD3B51434B094D4A" box="[725,776,512,537]" italics="true" pageId="17" pageNumber="18">stem</emphasis>
|
||
repent, rooting at the nodes, flexuous, unbranched;
|
||
<emphasis id="BD92AC05DF7EFF81FADB51434DAA4D4A" box="[1333,1451,512,537]" italics="true" pageId="17" pageNumber="18">internodes</emphasis>
|
||
to
|
||
<quantity id="481EDDF2DF7EFF81FF4B516048DC4D6F" box="[165,221,547,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="cm" value="1.0">1 cm</quantity>
|
||
long and rhizomatous at the base of the stem, woody, to
|
||
<quantity id="481EDDF2DF7EFF81FC8C51604BBD4D6F" box="[866,956,547,572]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="cm" value="0.75">0.75 cm</quantity>
|
||
thick, to
|
||
<quantity id="481EDDF2DF7EFF81FBC651604C5E4D6F" box="[1064,1119,547,572]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="cm" value="3.0">3 cm</quantity>
|
||
long and trailing towards the apex, to
|
||
<quantity id="481EDDF2DF7EFF81FF0B510549314D0C" box="[229,304,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="cm" value="0.5">0.5 cm</quantity>
|
||
thick, succulent, very densely lanate, hairs to
|
||
<quantity id="481EDDF2DF7EFF81FCC551054B9F4D0C" box="[811,926,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="17" pageNumber="18" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
|
||
, red;
|
||
<emphasis id="BD92AC05DF7EFF81FC3751054C2E4D0C" box="[985,1071,582,607]" italics="true" pageId="17" pageNumber="18">stipules</emphasis>
|
||
persistent, ovate, 8–12 ×
|
||
<quantity id="481EDDF2DF7EFF81FAA751054DA64D0C" box="[1353,1447,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="17" pageNumber="18" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
, apex acute, margins entire, densely ciliate, cilia recurved.
|
||
<emphasis id="BD92AC05DF7EFF81FCE5512A4B584DD1" box="[779,857,617,642]" italics="true" pageId="17" pageNumber="18">Leaves</emphasis>
|
||
alternate, clustered towards the apex of the stem, less than 5, basifixed;
|
||
<emphasis id="BD92AC05DF7EFF81FEBB51CE49AD4DF5" box="[341,428,653,678]" italics="true" pageId="17" pageNumber="18">petioles</emphasis>
|
||
<quantity id="481EDDF2DF7EFF81FE5B51CE4A1E4DF5" box="[437,543,653,678]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" pageNumber="18" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
|
||
long, very densely lanate, hairs
|
||
<quantity id="481EDDF2DF7EFF81FC7D51CE4C094DF5" box="[915,1032,653,678]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="17" pageNumber="18" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
|
||
, red;
|
||
<emphasis id="BD92AC05DF7EFF81FBA751CE4C964DF5" box="[1097,1175,653,678]" italics="true" pageId="17" pageNumber="18">lamina</emphasis>
|
||
asymmetric at the base, straight, oblique, lanceolate, 8–18 ×
|
||
<quantity id="481EDDF2DF7EFF81FDCF51F34A8A4D9A" box="[545,651,688,713]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.5" metricValueMin="3.0" pageId="17" pageNumber="18" unit="cm" value="4.25" valueMax="5.5" valueMin="3.0">3–5.5 cm</quantity>
|
||
, apex acuminate, base cuneate on the narrow side of the blade, rounded on the wide side of the blade, acuspidate, margins double-dentate, teeth
|
||
<quantity id="481EDDF2DF7EFF81FC4B51904C174DBF" box="[933,1046,723,748]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="17" pageNumber="18" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 mm</quantity>
|
||
long, densely long-ciliate, the upper surface bullate, with 5 to 15 dense, glandular-pilose hairs between each tertiary vein connected to one to four three-fid to eight-fid glands visible within the leaf lamina between each tertiary vein, the glands drying black, the veins sparsely lanate to densely lanate towards the petiolar insertion, the lower surface densely glandular-pilose between the tertiary veins, the veins very densely lanate; venation pinnate, with ca. 15 lateral veins on the widest side of the lamina, and ca. 10 lateral veins on the narrow side.
|
||
<emphasis id="BD92AC05DF7EFF81FDF650C04AC34CCF" box="[536,706,899,924]" italics="true" pageId="17" pageNumber="18">Inflorescences:</emphasis>
|
||
axillary, terminal, erect, to
|
||
<quantity id="481EDDF2DF7EFF81FC1950C04C3E4CCF" box="[1015,1087,899,924]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="18" unit="cm" value="18.0">18 cm</quantity>
|
||
, very densely lanate throughout, thyrsoid with 6–10 lateral branches, lateral branches cymose, branching 1 time, protandrous;
|
||
<emphasis id="BD92AC05DF7EFF81FB4650E54D0C4CEC" box="[1192,1293,934,959]" italics="true" pageId="17" pageNumber="18">peduncle</emphasis>
|
||
to
|
||
<quantity id="481EDDF2DF7EFF81FADC50E54D6B4CEC" box="[1330,1386,934,959]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="cm" value="8.0">8 cm</quantity>
|
||
long, internodes to
|
||
<quantity id="481EDDF2DF7EFF81FEF0508A49554CB1" box="[286,340,969,994]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="cm" value="3.0">3 cm</quantity>
|
||
long, decreasing to ca.
|
||
<quantity id="481EDDF2DF7EFF81FDB6508A4A994CB1" box="[600,664,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
|
||
long at the apex, peduncles of cymes
|
||
<quantity id="481EDDF2DF7EFF81FBD4508A4C984CB1" box="[1082,1177,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
|
||
;
|
||
<emphasis id="BD92AC05DF7EFF81FB4A508A4CEB4CB1" box="[1188,1258,969,994]" italics="true" pageId="17" pageNumber="18">bracts</emphasis>
|
||
persistent, ovate, 4–5 ×
|
||
<quantity id="481EDDF2DF7EFF81FF3E50AE49314B55" box="[208,304,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
|
||
, membranous, glabrous, apex obtuse, margin lacerate, ciliate;
|
||
<emphasis id="BD92AC05DF7EFF81FC0250AE4CFA4B55" box="[1004,1275,1005,1030]" italics="true" pageId="17" pageNumber="18">pedicels of male flowers</emphasis>
|
||
ca.
|
||
<quantity id="481EDDF2DF7EFF81FAC450AE4D6D4B55" box="[1322,1388,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="mm" value="4.0">4 mm</quantity>
|
||
long, fibrous;
|
||
<emphasis id="BD92AC05DF7EFF81FF0957534A0A4B7A" box="[231,523,1040,1065]" italics="true" pageId="17" pageNumber="18">pedicels of female flowers</emphasis>
|
||
<quantity id="481EDDF2DF7EFF81FDFD57534A7F4B7A" box="[531,638,1040,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="17" pageNumber="18" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
|
||
long, fibrous.
|
||
<emphasis id="BD92AC05DF7EFF81FCCD57534BB74B7A" box="[803,950,1040,1065]" italics="true" pageId="17" pageNumber="18">Male flowers</emphasis>
|
||
:
|
||
<emphasis id="BD92AC05DF7EFF81FC2857534C084B7A" box="[966,1033,1040,1065]" italics="true" pageId="17" pageNumber="18">tepals</emphasis>
|
||
known only from bud, membranous, glabrous, the margins entire, aciliate, the outer 2 ovate, 4 ×
|
||
<quantity id="481EDDF2DF7EFF81FCF257704B754B1F" box="[796,884,1075,1100]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, apex acute, white, the inner unknown and perhaps non-existent;
|
||
<emphasis id="BD92AC05DF7EFF81FECE5715497B4B3C" box="[288,378,1110,1135]" italics="true" pageId="17" pageNumber="18">stamens</emphasis>
|
||
unknown.
|
||
<emphasis id="BD92AC05DF7EFF81FE1A57154AA14B3C" box="[500,672,1110,1135]" italics="true" pageId="17" pageNumber="18">Female flowers</emphasis>
|
||
:
|
||
<emphasis id="BD92AC05DF7EFF81FD4057154B224B3C" box="[686,803,1110,1135]" italics="true" pageId="17" pageNumber="18">bracteoles</emphasis>
|
||
2 persistent, lanceolate, 8–9 × 2–2.5, membranous, villous outside, glabrous inside, margins serrate, densely ciliate;
|
||
<emphasis id="BD92AC05DF7EFF81FCE2573A4B4E4BC1" box="[780,847,1145,1170]" italics="true" pageId="17" pageNumber="18">tepals</emphasis>
|
||
5, persistent in fruit, same colour as males, subequal, lanceolate to ovate, 6–9 × 2–4, margins entire, ciliate;
|
||
<emphasis id="BD92AC05DF7EFF81FD0D57DE4B5E4BE5" box="[739,863,1181,1206]" italics="true" pageId="17" pageNumber="18">ovary body</emphasis>
|
||
globose, 4.5–7 ×
|
||
<quantity id="481EDDF2DF7EFF81FBCB57DE4C824BE5" box="[1061,1155,1181,1206]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="18" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
, sparsely pilose, 3-locular, unequally 3-winged, largest wing triangular, not ascending, widest 1/3 of the length towards the apex, 5–12 ×
|
||
<quantity id="481EDDF2DF7EFF81FA6C578348B74BAF" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="17" pageNumber="18" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
, apex acute, base cordate to rounded, the upper margin entire, ciliate, the lower margin lacerate, ciliate, smallest 2 marginiform,
|
||
<quantity id="481EDDF2DF7EFF81FED5564549984A4C" box="[315,409,1286,1311]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="17" pageNumber="18" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
|
||
wide, rounded;
|
||
<emphasis id="BD92AC05DF7EFF81FDBC56454ABE4A4C" box="[594,703,1286,1311]" italics="true" pageId="17" pageNumber="18">placentae</emphasis>
|
||
axile, simple, ovuliferous all over;
|
||
<emphasis id="BD92AC05DF7EFF81FBBB56454C924A4C" box="[1109,1171,1286,1311]" italics="true" pageId="17" pageNumber="18">styles</emphasis>
|
||
3,
|
||
<quantity id="481EDDF2DF7EFF81FB5756454D164A4C" box="[1209,1303,1286,1311]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="17" pageNumber="18" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
long, bifid
|
||
<quantity id="481EDDF2DF7EFF81FA73564548B54A11" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="2.0">2 mm</quantity>
|
||
from base, the branches erect, spirally twisted two to three times, persistent in fruit.
|
||
<emphasis id="BD92AC05DF7EFF81FBB0566A4D124A11" box="[1118,1299,1321,1346]" italics="true" pageId="17" pageNumber="18">Fruiting pedicel</emphasis>
|
||
as the female flowers.
|
||
<emphasis id="BD92AC05DF7EFF81FF04560E49254A35" box="[234,292,1357,1382]" italics="true" pageId="17" pageNumber="18">Fruit</emphasis>
|
||
ovate, enlarging to 8 ×
|
||
<quantity id="481EDDF2DF7EFF81FDC2560E4A6F4A35" box="[556,622,1357,1382]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="mm" value="7.0">7 mm</quantity>
|
||
, largest wing the same shape as in the ovary, enlarging to 12 ×
|
||
<quantity id="481EDDF2DF7EFF81FAC3560E4D7F4A35" box="[1325,1406,1357,1382]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="12.0">12 mm</quantity>
|
||
, the smallest enlarging to a triangular wing 9 ×
|
||
<quantity id="481EDDF2DF7EFF81FD8456334AAF4ADA" box="[618,686,1392,1417]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
<emphasis id="BD92AC05DF7EFF81FD5656334AF94ADA" box="[696,760,1392,1417]" italics="true" pageId="17" pageNumber="18">Seeds</emphasis>
|
||
globose, 0.2 ×
|
||
<quantity id="481EDDF2DF7EFF81FC4B56334BFF4ADA" box="[933,1022,1392,1417]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7FC239CDF7EFF81FF5356D04A2E4A9C" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="8F597017DF7EFF81FF5356D04A2E4A9C" blockId="17.[136,1452,512,1663]" pageId="17" pageNumber="18">
|
||
<emphasis id="BD92AC05DF7EFF81FF5356D04A094AFF" bold="true" box="[189,520,1427,1452]" pageId="17" pageNumber="18">Distribution and habitat:—</emphasis>
|
||
Veneuela. Known only from talus slopes at ca.
|
||
<quantity id="481EDDF2DF7EFF81FBF156D04C744AFF" box="[1055,1141,1427,1452]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="m" value="1200.0">1200 m</quantity>
|
||
on the Cerro Parú Tepui in the
|
||
<collectingRegion id="4D22BEF5DF7EFF81FF5E56F5498E4A9C" box="[176,399,1462,1487]" country="Venezuela" name="Amazonas" pageId="17" pageNumber="18">Amazonas province</collectingRegion>
|
||
of
|
||
<collectingCountry id="F7F13087DF7EFF81FE5A56F54A2A4A9C" box="[436,555,1462,1487]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7FC239CDF7EFF81FF53569A4CF5492C" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="8F597017DF7EFF81FF53569A4B49496A" blockId="17.[136,1452,512,1663]" pageId="17" pageNumber="18">
|
||
<emphasis id="BD92AC05DF7EFF81FF53569A49654AA1" bold="true" box="[189,356,1497,1522]" pageId="17" pageNumber="18">Etymology:—</emphasis>
|
||
The genus
|
||
<taxonomicName id="48E60B94DF7EFF81FE0D569A4A414AA1" baseAuthorityName="Linnaeus 1753:" baseAuthorityYear="1056" box="[483,576,1497,1522]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="BD92AC05DF7EFF81FE0D569A4A414AA1" box="[483,576,1497,1522]" italics="true" pageId="17" pageNumber="18">Begonia</emphasis>
|
||
</taxonomicName>
|
||
is relatively poorly known from the tepuis of northern Amazonia. We name this species
|
||
<taxonomicName id="48E60B94DF7EFF81FEE056BE499C4945" authorityName="Moonlight & Jara-Muñoz" authorityYear="2017" box="[270,413,1533,1558]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tepuiensis">
|
||
<emphasis id="BD92AC05DF7EFF81FEE056BE499C4945" box="[270,413,1533,1558]" italics="true" pageId="17" pageNumber="18">B. tepuiensis</emphasis>
|
||
</taxonomicName>
|
||
as it is only the third
|
||
<taxonomicName id="48E60B94DF7EFF81FD6356BE4B434945" box="[653,834,1533,1558]" pageId="17" pageNumber="18">
|
||
<emphasis id="BD92AC05DF7EFF81FD6356BE4AEB4945" box="[653,746,1533,1558]" italics="true" pageId="17" pageNumber="18">Begonia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described exclusively from tepuis after
|
||
<taxonomicName id="48E60B94DF7EFF81FAEF56BE4984496A" authority="L. B. Sm. & B. G. Schub" authorityName="L. B. Sm. & B. G. Schub" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="steyermarkii">
|
||
<emphasis id="BD92AC05DF7EFF81FAEF56BE4DAA4945" box="[1281,1451,1533,1558]" italics="true" pageId="17" pageNumber="18">B. steyermarkii</emphasis>
|
||
L.B.Sm. & B.G.Schub
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="48E60B94DF7EFF81FE5555634B49496A" authority="L. B. Sm. & B. G. Schub." authorityName="L. B. Sm. & B. G. Schub." box="[443,840,1568,1593]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="nubicola">
|
||
<emphasis id="BD92AC05DF7EFF81FE5555634A3D496A" box="[443,572,1568,1593]" italics="true" pageId="17" pageNumber="18">B. nubicola</emphasis>
|
||
L.B.Sm. & B.G.Schub.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8F597017DF7EFF81FF5355004CF5492C" blockId="17.[136,1452,512,1663]" pageId="17" pageNumber="18">
|
||
<emphasis id="BD92AC05DF7EFF81FF5355004A14490F" bold="true" box="[189,533,1603,1628]" pageId="17" pageNumber="18">IUCN Redlist Assessment:—</emphasis>
|
||
No information is available about population size or trends in
|
||
<taxonomicName id="48E60B94DF7EFF81FB3E55004DA5490F" authorityName="Moonlight & Jara-Muñoz" authorityYear="2017" box="[1232,1444,1603,1628]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tepuiensis">
|
||
<emphasis id="BD92AC05DF7EFF81FB3E55004DA5490F" box="[1232,1444,1603,1628]" italics="true" pageId="17" pageNumber="18">Begonia tepuiensis</emphasis>
|
||
</taxonomicName>
|
||
, which is known from only a single site. Accordingly, we assess
|
||
<taxonomicName id="48E60B94DF7EFF81FCBC55254BE3492C" authorityName="Moonlight & Jara-Muñoz" authorityYear="2017" box="[850,994,1638,1663]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tepuiensis">
|
||
<emphasis id="BD92AC05DF7EFF81FCBC55254BE3492C" box="[850,994,1638,1663]" italics="true" pageId="17" pageNumber="18">B. tepuiensis</emphasis>
|
||
</taxonomicName>
|
||
as Vulnerable (VU D2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |