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<document id="421D8938DD4F6B8C900414D04EFC80B5" ID-DOI="10.5852/ejt.2017.368" ID-GBIF-Dataset="7d707e31-1104-4ed3-8493-ef9302b98796" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838412" ID-ZooBank="E4AEAA16-A546-48B8-99BA-2E0152C8C6F0" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1590087673171" checkinUser="carolina" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2017" docId="03E9996C14484E77B5116EE6806BDBF0" docLanguage="en" docName="ejt-2017-368.pdf.imf" docOrigin="European Journal of Taxonomy 368 (368)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phtheochroa alpinana Zlatkov &amp; Huemer 2017, sp. nov." docType="treatment" docUuid="704B04D3-3AAA-4554-A4D4-0F7C9439173D" docUuidSource="ZooBank" docVersion="11" lastPageNumber="11" masterDocId="FFD0E114144F4E7DB7706C11845EDC35" masterDocTitle="Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="8" updateTime="1737036911417" updateUser="carolina" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A587E9A9504E8BDE712D5FA7213C1802">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="EB0D19779BD1430536EEB6A79862EB39" type="personal">
<mods:role id="2FC4263F43EBCE686DF437AFF9AF7F8E">
<mods:roleTerm id="A3FCEDCE6960034944FA812F9423A709">Author</mods:roleTerm>
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<mods:namePart id="352B198F2AFEE38AC0327CD1CA9E255F">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="F93AECCEAB908E0A684A643E62AFCF7F">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria.</mods:affiliation>
<mods:nameIdentifier id="8B8360992DB15689CDD29ACCEC8A5977" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="34A8A2EF808052EFD70A414D6BA38BE1">Huemer, Peter</mods:namePart>
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<mods:affiliation id="CB4BAA30F4EFBAFA4372B1E5F4411FD6">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria.</mods:affiliation>
<mods:nameIdentifier id="D8901EB36F313D21D446AC56E0FCC029" type="email">p.huemer@tiroler-landesmuseen.at</mods:nameIdentifier>
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<mods:title id="BF227175C6BB9B1C545FCC993A20B630">European Journal of Taxonomy</mods:title>
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<mods:date id="572F1DEA3BA96578D43891FB2829A23A">2017</mods:date>
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<mods:number id="FE3CCC99AABCB7A5C25B5767E6924C2F">2017-11-21</mods:number>
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<mods:number id="8804899B71FEC546C06B828DC48D36ED">368</mods:number>
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<mods:number id="6E762CDDD7ED75A1AA7AD79B7A16D1F7">368</mods:number>
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<treatment id="03E9996C14484E77B5116EE6806BDBF0" ID-DOI="http://doi.org/10.5281/zenodo.3852276" ID-GBIF-Taxon="164246400" ID-Zenodo-Dep="3852276" LSID="urn:lsid:zoobank.org:act:704B04D3-3AAA-4554-A4D4-0F7C9439173D" httpUri="http://treatment.plazi.org/id/03E9996C14484E77B5116EE6806BDBF0" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" scope_class="Insecta" scope_family="Tortricidae" scope_order="Lepidoptera" scope_subFamily="Tortricinae" scope_superFamily="Tortricoidea" scope_tribe="Cochylini">
<subSubSection id="C35A7BF114484E7AB5116EE6878DDF27" box="[609,979,759,786]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BFF287A14484E7AB5116EE6878DDF27" blockId="7.[364,1223,759,856]" box="[609,979,759,786]" pageId="7" pageNumber="8">
<heading id="D0B79F1614484E7AB5116EE6878DDF27" box="[609,979,759,786]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C4053F914484E7AB5116EE6872FDF27" ID-CoL="B3ZCD" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[609,881,759,786]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
<emphasis id="B934F46814484E7AB5116EE6872FDF27" bold="true" box="[609,881,759,786]" italics="true" pageId="7" pageNumber="8">Phtheochroa alpinana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314484E7AB4086EE9878DDF27" box="[888,979,760,786]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114484E7AB61C6F0B8014DF6D" pageId="7" pageNumber="8" type="description">
<paragraph id="8BFF287A14484E7AB61C6F0B8099DF00" blockId="7.[364,1223,759,856]" box="[364,1223,794,821]" pageId="7" pageNumber="8">
<uri id="FFD1247814484E7AB61C6F0B8099DF00" box="[364,1223,794,821]" pageId="7" pageNumber="8">
urn:lsid:zoobank.org:act:
<uuid id="FFE612AF14484E7AB5FB6F0B8099DF00" box="[651,1223,794,821]" pageId="7" pageNumber="8">704B04D3-3AAA-4554-A4D4-0F7C9439173D</uuid>
</uri>
</paragraph>
<paragraph id="8BFF287A14484E7AB6996F2F8014DF6D" blockId="7.[364,1223,759,856]" box="[489,1098,830,856]" pageId="7" pageNumber="8">
<figureCitation id="137B34FF14484E7AB6996F2F8672DF6D" box="[489,556,830,856]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="7" pageNumber="8">Figs 1</figureCitation>
E–F, 2C, 3C, 4D–E, 6A–B, 8B, 9, 10;
<tableCitation id="C6C21DC114484E7AB4836F2F8014DF6D" box="[1011,1098,830,856]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="7" pageNumber="8" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114484E7AB7CD6F6E8668D8D3" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BFF287A14484E7AB7CD6F6E8568DFAC" blockId="7.[189,310,895,921]" box="[189,310,895,921]" pageId="7" pageNumber="8">
<heading id="D0B79F1614484E7AB7CD6F6E8568DFAC" bold="true" box="[189,310,895,921]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B934F46814484E7AB7CD6F6E8568DFAC" bold="true" box="[189,310,895,921]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14484E7AB7CD6FA08668D8D3" blockId="7.[189,1399,945,1254]" pageId="7" pageNumber="8">
The forewing upperside is paler than in
<taxonomicName id="4C4053F914484E7AB5FD6FA3876ADFFE" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[653,820,945,971]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814484E7AB5FD6FA3876ADFFE" box="[653,820,945,971]" italics="true" pageId="7" pageNumber="8">P. schawerdae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F914484E7AB41C6FA3804DDFFE" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[876,1043,946,971]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814484E7AB41C6FA3804DDFFE" box="[876,1043,946,971]" italics="true" pageId="7" pageNumber="8">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314484E7AB3696FA3802DDFF9" box="[1049,1139,946,972]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
and somewhat deeper yellow than in
<taxonomicName id="4C4053F914484E7AB6186FC785AEDFDA" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[360,496,981,1007]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814484E7AB6186FC785AEDFDA" box="[360,496,981,1007]" italics="true" pageId="7" pageNumber="8">P. frigidana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F914484E7AB5566FC7868FDFDA" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[550,721,981,1007]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814484E7AB5566FC7868FDFDA" box="[550,721,981,1007]" italics="true" pageId="7" pageNumber="8">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314484E7AB5A76FC4876EDFDA" box="[727,816,981,1007]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
The hindwings are pale grey as in
<taxonomicName id="4C4053F914484E7AB3CE6FC78118DFDA" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[1214,1350,981,1007]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814484E7AB3CE6FC78118DFDA" box="[1214,1350,981,1007]" italics="true" pageId="7" pageNumber="8">P. frigidana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F914484E7AB7CD6FE88536D827" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[189,360,1016,1042]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814484E7AB7CD6FE88536D827" box="[189,360,1016,1042]" italics="true" pageId="7" pageNumber="8">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314484E7AB61F6FE98597D827" box="[367,457,1016,1042]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
The vesica easily distinguishes this species from all other members of the group: with large dorsal and ventral diverticula;
<taxonomicName id="4C4053F914484E7AB5ED680D8778D800" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[669,806,1051,1077]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814484E7AB5ED680D8778D800" box="[669,806,1051,1077]" italics="true" pageId="7" pageNumber="8">P. frigidana</emphasis>
</taxonomicName>
has only one diverticulum (dorsal); the diverticula on the other three species are oriented more or less laterally. The shape of the valva resembles those of
<taxonomicName id="4C4053F914484E7AB7CD68738519D84E" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[189,327,1121,1147]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814484E7AB7CD68738519D84E" box="[189,327,1121,1147]" italics="true" pageId="7" pageNumber="8">P. frigidana</emphasis>
</taxonomicName>
. The phallic process is slightly curved to the right and resembles that of
<taxonomicName id="4C4053F914484E7AB3D168738113D84E" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1185,1357,1121,1147]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814484E7AB3D168738113D84E" box="[1185,1357,1121,1147]" italics="true" pageId="7" pageNumber="8">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314484E7AB225687384B1D8AA" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
The female genitalia are generally similar to these of
<taxonomicName id="4C4053F914484E7AB41768978050D8AA" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[871,1038,1157,1183]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814484E7AB41768978050D8AA" box="[871,1038,1157,1183]" italics="true" pageId="7" pageNumber="8">P. schawerdae</emphasis>
</taxonomicName>
, but the sclerotized area of the corpus bursae is larger, with three long folds on the right ventral side; the folds in
<taxonomicName id="4C4053F914484E7AB3EF68B88117D8F7" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[1183,1353,1192,1218]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814484E7AB3EF68B88117D8F7" box="[1183,1353,1192,1218]" italics="true" pageId="7" pageNumber="8">P. schawerdae</emphasis>
</taxonomicName>
are short and more numerous (5–6).
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114484E7AB7CD6905801AD955" box="[189,1092,1300,1377]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BFF287A14484E7AB7CD6905851AD91B" blockId="7.[189,324,1300,1326]" box="[189,324,1300,1326]" pageId="7" pageNumber="8">
<heading id="D0B79F1614484E7AB7CD6905851AD91B" bold="true" box="[189,324,1300,1326]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B934F46814484E7AB7CD6905851AD91B" bold="true" box="[189,324,1300,1326]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14484E7AB7CD6957801AD955" blockId="7.[189,1092,1350,1377]" box="[189,1092,1350,1377]" pageId="7" pageNumber="8">The specific name is a feminine adjective derived from the name of the Alps.</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114484E75B7CD699E8703DE9E" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BFF287A14484E7AB7CD699E85F4D99C" blockId="7.[189,426,1423,1449]" box="[189,426,1423,1449]" pageId="7" pageNumber="8">
<heading id="D0B79F1614484E7AB7CD699E85F4D99C" bold="true" box="[189,426,1423,1449]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B934F46814484E7AB7CD699E85F4D99C" bold="true" box="[189,426,1423,1449]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B28222714484E7AB79A69D38578DA4D" ID-GBIF-Occurrence="2621725302" collectingDate="1991-07-20" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" country="France" elevation="2200" location="2200 m NE" municipality="Alpes Maritimes Tete Chaudon" pageId="7" pageNumber="8" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<paragraph id="8BFF287A14484E7AB79A69D38502D9E9" blockId="7.[234,348,1474,1500]" box="[234,348,1474,1500]" pageId="7" pageNumber="8">
<emphasis id="B934F46814484E7AB79A69D38502D9E9" bold="true" box="[234,348,1474,1500]" pageId="7" pageNumber="8">
<typeStatus id="54FB96D814484E7AB79A69D38502D9E9" box="[234,348,1474,1500]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A14484E7AB7CD69E58578DA4D" blockId="7.[189,1399,1524,1656]" pageId="7" pageNumber="8">
<collectingCountry id="F35768EA14484E7AB7CD69E5856BDA3B" box="[189,309,1524,1550]" name="France" pageId="7" pageNumber="8">FRANCE</collectingCountry>
:
<specimenCount id="9D46E3F314484E7AB63469E48502DA3A" box="[324,348,1525,1551]" count="1" pageId="7" pageNumber="8" type="male">♂</specimenCount>
, pinned, spread, well-preserved, three labels: // FRANKREICH
<collectingMunicipality id="6B9BB20014484E7AB30169E58574DA04" country="France" name="Alpes Maritimes Tete Chaudon" pageId="7" pageNumber="8">Alpes Maritimes Tete Chaudon</collectingMunicipality>
,
<location id="8E9F7EA114484E7AB6466A06859DDA07" LSID="urn:lsid:plazi:treatment:03E9996C14484E77B5116EE6806BDBF0:8E9F7EA114484E7AB6466A06859DDA07" box="[310,451,1559,1586]" country="France" municipality="Alpes Maritimes Tete Chaudon" name="2200 m NE" pageId="7" pageNumber="8">
<quantity id="4CB8859F14484E7AB6466A0685CEDA07" box="[310,400,1559,1586]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="m" value="2200.0">2200 m</quantity>
NE
</location>
.
<location id="8E9F7EA114484E7AB6BE6A0686DDDA04" LSID="urn:lsid:plazi:treatment:03E9996C14484E77B5116EE6806BDBF0:8E9F7EA114484E7AB6BE6A0686DDDA04" box="[462,643,1559,1585]" country="France" municipality="Alpes Maritimes Tete Chaudon" name="Col de la Boira" pageId="7" pageNumber="8">Col de la Boira</location>
<date id="FFFE0EBA14484E7AB5FB6A06875EDA04" box="[651,768,1559,1585]" pageId="7" pageNumber="8" value="1991-07-20">
<collectingDate id="EFBAF75214484E7AB5FB6A06875EDA04" box="[651,768,1559,1585]" pageId="7" pageNumber="8" value="1991-07-20">20.7.1991</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14484E7AB44B6A0987C7DA07" box="[827,921,1560,1586]" pageId="7" pageNumber="8">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14484E7AB4B06A098076DA07" box="[960,1064,1560,1586]" pageId="7" pageNumber="8">Tarmann</collectorName>
// Gen. prep.
<specimenCount id="9D46E3F314484E7AB3BC6A0980BCDA07" box="[1228,1250,1560,1586]" count="1" pageId="7" pageNumber="8" type="male">♂</specimenCount>
<collectingDate id="EFBAF75214484E7AB39A6A068129DA04" box="[1258,1399,1559,1585]" pageId="7" pageNumber="8">
1/
<date id="FFFE0EBA14484E7AB38D6A068129DA04" box="[1277,1399,1559,1585]" pageId="7" pageNumber="8" value="1991-07-20">20.7.1991</date>
</collectingDate>
[genitalia slide number] // HOLOTYPE
<taxonomicName id="4C4053F914484E7AB5EE6A2B8084DA60" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[670,1242,1594,1621]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814484E7AB5EE6A2B87FBDA61" box="[670,933,1594,1620]" italics="true" pageId="7" pageNumber="8">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14484E7AB7B56A4F8544DA4D" box="[197,282,1630,1656]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="7" pageNumber="8">TLMF</collectionCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B28222714484E7AB79A6AB680ECDB0C" collectingDate="1991-07-20" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" country="France" location="2200 m NE. Col de la Boira" municipality="Alpes Maritimes Tete Chaudon" pageId="7" pageNumber="8" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<paragraph id="8BFF287A14484E7AB79A6AB68539DAF4" blockId="7.[234,359,1703,1729]" box="[234,359,1703,1729]" pageId="7" pageNumber="8">
<emphasis id="B934F46814484E7AB79A6AB68539DAF4" bold="true" box="[234,359,1703,1729]" pageId="7" pageNumber="8">
<typeStatus id="54FB96D814484E7AB79A6AB68539DAF4" box="[234,359,1703,1729]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A14484E75B7CD6AC88165DDBA" blockId="7.[189,1399,1753,2026]" lastBlockId="8.[189,1399,267,400]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<collectingCountry id="F35768EA14484E7AB7CD6AC8856FDAC6" box="[189,305,1753,1779]" name="France" pageId="7" pageNumber="8">FRANCE</collectingCountry>
:
<specimenCount id="9D46E3F314484E7AB63C6AC88522DAC6" box="[332,380,1753,1779]" count="1" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, three labels: // FRANKREICH
<collectingMunicipality id="6B9BB20014484E7AB4736AC88022DAC6" box="[771,1148,1753,1779]" pageId="7" pageNumber="8">Alpes Maritimes Tete Chaudon</collectingMunicipality>
,
<location id="8E9F7EA114484E7AB3FA6AC88540DB23" LSID="urn:lsid:plazi:treatment:03E9996C14484E77B5116EE6806BDBF0:8E9F7EA114484E7AB3FA6AC88540DB23" country="France" municipality="Alpes Maritimes Tete Chaudon" name="2200 m NE. Col de la Boira" pageId="7" pageNumber="8">
<quantity id="4CB8859F14484E7AB3FA6AC880BBDAC6" box="[1162,1253,1753,1779]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="m" value="2200.0">2200 m</quantity>
NE. Col de la Boira
</location>
<date id="FFFE0EBA14484E7AB6596AED85C1DB23" box="[297,415,1788,1814]" pageId="7" pageNumber="8" value="1991-07-20">
<collectingDate id="EFBAF75214484E7AB6596AED85C1DB23" box="[297,415,1788,1814]" pageId="7" pageNumber="8" value="1991-07-20">20.7.1991</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14484E7AB6906AEC8660DB22" box="[480,574,1789,1815]" pageId="7" pageNumber="8">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14484E7AB51B6AEC868DDB22" box="[619,723,1789,1815]" pageId="7" pageNumber="8">Tarmann</collectorName>
// Gen. prep.
<specimenCount id="9D46E3F314484E7AB4F36AEC87C7DB22" box="[899,921,1789,1815]" count="1" pageId="7" pageNumber="8" type="male">♂</specimenCount>
2/
<date id="FFFE0EBA14484E7AB4CB6AED806FDB23" box="[955,1073,1788,1814]" pageId="7" pageNumber="8" value="1991-07-20">20.7.1991</date>
[genitalia slide number] // PARATYPE
<taxonomicName id="4C4053F914484E7AB6296B0E87C3DB0C" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[345,925,1823,1850]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814484E7AB6296B0E863CDB0C" box="[345,610,1823,1849]" italics="true" pageId="7" pageNumber="8">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14484E7AB3216B0E80F8DB0C" box="[1105,1190,1823,1849]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="7" pageNumber="8">TLMF</collectionCode>
);
<materialsCitation id="3B28222714484E7AB3CD6B0E80F8DBB5" collectingDateMax="1991-07-20" collectingDateMin="1991-07-03" collectionCode="TLMF" country="France" location="2200 m NE. Col de la Boira" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D46E3F314484E7AB3CD6B0E80B2DB0F" box="[1213,1260,1823,1850]" count="1" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, with three labels: // locality data label as preceding // Gen. prep.
<specimenCount id="9D46E3F314484E7AB4066B5287D2DB68" box="[886,908,1859,1885]" count="1" pageId="7" pageNumber="8" type="female">♀</specimenCount>
3/
<date id="FFFE0EBA14484E7AB4DE6B52807BDB68" box="[942,1061,1859,1885]" pageId="7" pageNumber="8" value="1991-07-20">20.7.1991</date>
[genitalia slide number] // PARATYPE
<taxonomicName id="4C4053F914484E7AB6286B7787C8DBB5" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[344,918,1894,1920]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814484E7AB6286B778601DBB5" box="[344,607,1894,1920]" italics="true" pageId="7" pageNumber="8">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14484E7AB3346B7780C8DBB5" box="[1092,1174,1894,1920]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="7" pageNumber="8">TLMF</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B28222714484E7AB3DF6B778595DBDC" collectingDate="1990-07-23" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" elevation="2150" location="Dep. Alpes-Maritimes Marguareis W-Hang Navela" municipality="Dep. Alpes-Maritimes Marguareis W-Hang Navela" pageId="7" pageNumber="8" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D46E3F314484E7AB3DF6B7780ABDBB5" box="[1199,1269,1894,1920]" count="2" pageId="7" pageNumber="8" type="male">2 ♂♂</specimenCount>
, each with two labels: // FRANKREICH
<collectingMunicipality id="6B9BB20014484E7AB55A6B9880D1DB96" box="[554,1167,1929,1955]" pageId="7" pageNumber="8">Dep. Alpes-Maritimes Marguareis W-Hang Navela</collectingMunicipality>
,
<quantity id="4CB8859F14484E7AB3EE6B988160DB96" box="[1182,1342,1929,1955]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="7" pageNumber="8" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">
<elevation id="006DCF4914484E7AB3EE6B988160DB96" box="[1182,1342,1929,1955]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="7" pageNumber="8" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200 m</elevation>
</quantity>
21.-
<date id="FFFE0EBA14484E7AB7CD6BBD856CDBF3" box="[189,306,1964,1990]" pageId="7" pageNumber="8" value="1990-07-23">
<collectingDate id="EFBAF75214484E7AB7CD6BBD856CDBF3" box="[189,306,1964,1990]" pageId="7" pageNumber="8" value="1990-07-23">23.7.1990</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14484E7AB6056BBC858DDBF2" box="[373,467,1965,1991]" pageId="7" pageNumber="8">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14484E7AB5706BBC8636DBF2" box="[512,616,1965,1991]" pageId="7" pageNumber="8">Tarmann</collectorName>
// PARATYPE
<taxonomicName id="4C4053F914484E7AB45D6BBD8129DBF3" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[813,1399,1964,1991]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814484E7AB45D6BBD8069DBF3" box="[813,1079,1964,1990]" italics="true" pageId="7" pageNumber="8">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14484E7AB61A6BDE85E1DBDC" box="[362,447,1999,2025]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="7" pageNumber="8">TLMF</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B28222714484E75B6A76BDE873ADD59" collectingDateMax="2008-07-05" collectingDateMin="2008-07-01" collectionCode="ZMUC" collectorName="O. Karsholt" elevation="2000" lastPageId="8" lastPageNumber="9" location="Alpes Maritimes" municipality="Alpes Maritimes" pageId="7" pageNumber="8" specimenCode="ZMUC 00400703" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D46E3F314484E7AB6A76BDE8659DBDF" box="[471,519,1999,2026]" count="1" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, with five labels: // France,
<collectingMunicipality id="6B9BB20014484E7AB4156BDE806FDBDC" box="[869,1073,1999,2025]" pageId="7" pageNumber="8">Alpes Maritimes</collectingMunicipality>
,
<quantity id="4CB8859F14484E7AB3316BDE80C0DBDF" box="[1089,1182,1999,2026]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="m" value="2000.0">
<elevation id="006DCF4914484E7AB3316BDE80C0DBDF" box="[1089,1182,1999,2026]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="m" value="2000.0">2000 m</elevation>
</quantity>
<locationDeviation id="9A4D315814484E75B3D96BDE850FDD10" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<quantity id="4CB8859F14484E7AB3D96BDE80B6DBDC" box="[1193,1256,1999,2025]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="8" unit="km" value="6.0">6 km</quantity>
NW Tende Mont Chajol
</locationDeviation>
<date id="FFFE0EBA14474E75B6286D1A8591DD13" box="[344,463,267,294]" pageId="8" pageNumber="9" value="2008-07-05">5.vii.2008</date>
<collectorName id="26B54DAC14474E75B6A66D1A8601DD10" box="[470,607,267,293]" pageId="8" pageNumber="9">O. Karsholt</collectorName>
//
<specimenCode id="DBE6800114474E75B50D6D1A8713DD10" box="[637,845,267,293]" pageId="8" pageNumber="9">ZMUC 00400703</specimenCode>
// Coll. ZMUC, Copenhagen Denmark [yellow label] // DK Copenhagen Zool. Museum ♀ Genitalia slide No. 1/
<date id="FFFE0EBA14474E75B4CA6D3F807BDD7C" box="[954,1061,302,329]" pageId="8" pageNumber="9" value="2008-07-05">5.7.2008</date>
// PARATYPE
<taxonomicName id="4C4053F914474E75B3906D3E860BDD59" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814474E75B3906D3E857BDD59" italics="true" pageId="8" pageNumber="9">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14474E75B58D6D438706DD59" box="[765,856,338,364]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="9">ZMUC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B28222714474E75B41C6D438165DDBA" collectingDateMax="1991-07-05" collectingDateMin="1991-07-02" collectionCode="ZMUC" collectorName="O. Karsholt" elevation="2000" location="Alpes Maritimes" municipality="Alpes Maritimes" pageId="8" pageNumber="9" specimenCode="ZMUC 00400704" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D46E3F314474E75B41C6D4387C6DD59" box="[876,920,338,364]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
, five labels, all same as preceding but: //
<specimenCode id="DBE6800114474E75B7CD6D6485D0DDBA" box="[189,398,373,399]" pageId="8" pageNumber="9">ZMUC 00400704</specimenCode>
// DK Copenhagen Zool. Museum
<specimenCount id="9D46E3F314474E75B45D6D64871DDDBA" box="[813,835,373,399]" count="1" pageId="8" pageNumber="9" type="female">♀</specimenCount>
Genitalia slide No. 2/
<date id="FFFE0EBA14474E75B3356D6480F2DDA5" box="[1093,1196,373,400]" pageId="8" pageNumber="9" value="2008-07-05">5.7.2008</date>
// (
<collectionCode id="ED51B0BF14474E75B3A36D648171DDBA" box="[1235,1327,373,399]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="9">ZMUC</collectionCode>
).
</materialsCitation>
</paragraph>
</materialsCitation>
<paragraph id="8BFF287A14474E75B7CD6DAF8703DE9E" blockId="8.[189,1399,446,684]" pageId="8" pageNumber="9">
<materialsCitation id="3B28222714474E75B7CD6DAF8654DE2B" collectingDate="1991-07-19" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" country="Italy" elevation="2325" location="Prov. Imperia Punta Marguareis S-Hang" municipality="Prov. Imperia Punta Marguareis S-Hang" pageId="8" pageNumber="9" specimenCount="6" specimenCount-female="1" specimenCount-male="5" typeStatus="paratype">
<collectingCountry id="F35768EA14474E75B7CD6DAF854CDDED" box="[189,274,446,472]" name="Italy" pageId="8" pageNumber="9">ITALY</collectingCountry>
:
<specimenCount id="9D46E3F314474E75B66F6DAF853BDDED" box="[287,357,446,473]" count="5" pageId="8" pageNumber="9" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D46E3F314474E75B6056DAF85FDDDED" box="[373,419,446,472]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
, each with two labels: // ITALIA
<collectingMunicipality id="6B9BB20014474E75B4306DAF817ADDED" box="[832,1316,446,472]" pageId="8" pageNumber="9">Prov. Imperia Punta Marguareis S-Hang</collectingMunicipality>
,
<quantity id="4CB8859F14474E75B2426DAF8546DDC9" metricMagnitude="3" metricUnit="m" metricValue="2.325" metricValueMax="2.4" metricValueMin="2.25" pageId="8" pageNumber="9" unit="m" value="2325.0" valueMax="2400.0" valueMin="2250.0">
<elevation id="006DCF4914474E75B2426DAF8546DDC9" metricMagnitude="3" metricUnit="m" metricValue="2.325" metricValueMax="2.4" metricValueMin="2.25" pageId="8" pageNumber="9" unit="m" value="2325.0" valueMax="2400.0" valueMin="2250.0">2250- 2400 m</elevation>
</quantity>
<date id="FFFE0EBA14474E75B6506DF085CBDDCE" box="[288,405,481,507]" pageId="8" pageNumber="9" value="1991-07-19">
<collectingDate id="EFBAF75214474E75B6506DF085CBDDCE" box="[288,405,481,507]" pageId="8" pageNumber="9" value="1991-07-19">19.7.1991</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14474E75B6A36DF3866FDDC9" box="[467,561,482,508]" pageId="8" pageNumber="9">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14474E75B5296DF3869FDDC9" box="[601,705,482,508]" pageId="8" pageNumber="9">Tarmann</collectorName>
// PARATYPE
<taxonomicName id="4C4053F914474E75B40D6DF084A6DE2B" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814474E75B40D6DF080DBDDCE" box="[893,1157,481,507]" italics="true" pageId="8" pageNumber="9">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14474E75B6D96E1585A0DE2B" box="[425,510,516,542]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="8" pageNumber="9">TLMF</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B28222714474E75B5656E1587A8DE50" collectingDate="1990-07-23" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" country="Italy" elevation="2525" location="Prov. Imperia Punta Marguareis" municipality="Prov. Imperia Punta Marguareis" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D46E3F314474E75B5656E15861BDE2A" box="[533,581,516,543]" count="1" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
, three labels: //
<collectingCountry id="F35768EA14474E75B4796E148739DE2A" box="[777,871,517,543]" name="Italy" pageId="8" pageNumber="9">ITALIA</collectingCountry>
<collectingMunicipality id="6B9BB20014474E75B4006E1480B0DE2B" box="[880,1262,516,543]" pageId="8" pageNumber="9">Prov. Imperia Punta Marguareis</collectingMunicipality>
<quantity id="4CB8859F14474E75B3876E15848ADE77" metricMagnitude="3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="8" pageNumber="9" unit="m" value="2525.0" valueMax="2600.0" valueMin="2450.0">
<elevation id="006DCF4914474E75B3876E15848ADE77" metricMagnitude="3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="8" pageNumber="9" unit="m" value="2525.0" valueMax="2600.0" valueMin="2450.0">2450-2600 m</elevation>
</quantity>
<date id="FFFE0EBA14474E75B7AB6E39850FDE77" box="[219,337,552,578]" pageId="8" pageNumber="9" value="1990-07-23">
<collectingDate id="EFBAF75214474E75B7AB6E39850FDE77" box="[219,337,552,578]" pageId="8" pageNumber="9" value="1990-07-23">23.7.1990</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14474E75B6FB6E3985B7DE77" box="[395,489,552,578]" pageId="8" pageNumber="9">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14474E75B5606E398626DE77" box="[528,632,552,578]" pageId="8" pageNumber="9">Tarmann</collectorName>
// TOR 105 ♂ P.
<collectorName id="26B54DAC14474E75B4366E3987FADE77" box="[838,932,552,578]" pageId="8" pageNumber="9">Huemer</collectorName>
[genitalia slide number] // PARATYPE
<taxonomicName id="4C4053F914474E75B7CD6E5A86AEDE50" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[189,752,587,613]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814474E75B7CD6E5A859CDE50" box="[189,450,587,613]" italics="true" pageId="8" pageNumber="9">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14474E75B4E56E5A87B4DE50" box="[917,1002,587,613]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="8" pageNumber="9">TLMF</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B28222714474E75B48E6E5A8703DE9E" collectingDate="1990-07-23" collectionCode="TLMF" collectorName="Huemer &amp; Tarmann" country="Italy" elevation="2525" location="Prov. Imperia Punta Marguareis" municipality="Prov. Imperia Punta Marguareis" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D46E3F314474E75B48E6E5A8077DE50" box="[1022,1065,587,613]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
, two labels: //
<collectingCountry id="F35768EA14474E75B3A56E5A816DDE50" box="[1237,1331,587,613]" name="Italy" pageId="8" pageNumber="9">ITALIA</collectingCountry>
<collectingMunicipality id="6B9BB20014474E75B24A6E5A85B3DEBD" pageId="8" pageNumber="9">Prov. Imperia Punta Marguareis</collectingMunicipality>
<quantity id="4CB8859F14474E75B6836E7F86D1DEBD" box="[499,655,622,649]" metricMagnitude="3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="8" pageNumber="9" unit="m" value="2525.0" valueMax="2600.0" valueMin="2450.0">
<elevation id="006DCF4914474E75B6836E7F86D1DEBD" box="[499,655,622,649]" metricMagnitude="3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="8" pageNumber="9" unit="m" value="2525.0" valueMax="2600.0" valueMin="2450.0">2450-2600 m</elevation>
</quantity>
<date id="FFFE0EBA14474E75B5E56E7F8755DEBD" box="[661,779,622,648]" pageId="8" pageNumber="9" value="1990-07-23">
<collectingDate id="EFBAF75214474E75B5E56E7F8755DEBD" box="[661,779,622,648]" pageId="8" pageNumber="9" value="1990-07-23">23.7.1990</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14474E75B4326E7F87FEDEBD" box="[834,928,622,648]" pageId="8" pageNumber="9">Huemer</collectorName>
&amp;
<collectorName id="26B54DAC14474E75B4B36E7F8075DEBD" box="[963,1067,622,648]" pageId="8" pageNumber="9">Tarmann</collectorName>
// PARATYPE
<taxonomicName id="4C4053F914474E75B3906E7F860ADE9E" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alpinana">
<emphasis id="B934F46814474E75B3906E7F857BDE9E" italics="true" pageId="8" pageNumber="9">Phtheochroa alpinana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14474E75B58C6E80870FDE9E" box="[764,849,657,683]" collectionName="Austria, Innsbruck, Tiroler Landesmuseum Ferdinandeum" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="8" pageNumber="9">TLMF</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114474E77B7CD6ECB80BDD9ED" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="8BFF287A14474E75B7CD6ECB8510DEC1" blockId="8.[189,334,730,756]" box="[189,334,730,756]" pageId="8" pageNumber="9">
<heading id="D0B79F1614474E75B7CD6ECB8510DEC1" bold="true" box="[189,334,730,756]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B934F46814474E75B7CD6ECB8510DEC1" bold="true" box="[189,334,730,756]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14474E75B79A6F1C85C5DF12" blockId="8.[234,411,780,807]" box="[234,411,780,807]" pageId="8" pageNumber="9">
<emphasis id="B934F46814474E75B79A6F1C8574DF12" bold="true" box="[234,298,781,807]" pageId="8" pageNumber="9">Male</emphasis>
(
<figureCitation id="137B34FF14474E75B64A6F1D85CFDF12" box="[314,401,780,807]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="8" pageNumber="9">Fig. 1E</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14474E75B7CD6F2E8644DFAA" blockId="8.[189,1398,831,928]" pageId="8" pageNumber="9">HEAD. Frons and vertex covered with creamy scales. Labial palps pointed anterad, long, dorsally creamy, laterally and ventrally brown. Antennae with scapus and pedicellus dorsally brown, ventrally white; flagellum with creamy scales.</paragraph>
<paragraph id="8BFF287A14474E77B7CD6FDE85BBDD59" blockId="8.[189,1399,974,1071]" lastBlockId="10.[189,1399,267,364]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
THORAX. Dorsally creamy with brown scales in middle, ventrally creamy with brown scales. Tegula yellowish, with brown scales laterally. Forewing length
<quantity id="4CB8859F14474E75B4166FE0805BD839" box="[870,1029,1009,1036]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" metricValueMax="10.4" metricValueMin="8.4" pageId="8" pageNumber="9" unit="mm" value="9.4" valueMax="10.4" valueMin="8.4">8.4–10.4 mm</quantity>
(x=9.0, n =9). Forewing long and narrow, with pointed apex, upperside ground colour yellow with paler distal half and inconspicuous rust dorsal blotch, underside dark grey with scattered white scales in apical area, cilia pale yellow. Hindwings with upperside grey, underside grey with white costa and white longitudinal lines, cilia white with pale grey basal part.
</paragraph>
<caption id="DF3F78F214474E75B7CD6B31873EDBDF" ID-DOI="http://doi.org/10.5281/zenodo.3838420" ID-Zenodo-Dep="3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1824,1850]" targetBox="[189,1398,1117,1787]" targetPageId="8">
<paragraph id="8BFF287A14474E75B7CD6B31873EDBDF" blockId="8.[189,1399,1824,2027]" pageId="8" pageNumber="9">
<emphasis id="B934F46814474E75B7CD6B318553DB0F" bold="true" box="[189,269,1824,1850]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
Ventral phallic process of
<taxonomicName id="4C4053F914474E75B5216B3086B9DB0E" authorityName="Stephens" authorityYear="1829" box="[593,743,1825,1851]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B934F46814474E75B5216B3086B9DB0E" box="[593,743,1825,1851]" italics="true" pageId="8" pageNumber="9">Phtheochroa</emphasis>
</taxonomicName>
spp., ventral view.
<emphasis id="B934F46814474E75B4A56B318050DB0F" bold="true" box="[981,1038,1824,1850]" pageId="8" pageNumber="9">A–C</emphasis>
.
<taxonomicName id="4C4053F914474E75B36F6B308128DB0E" authority="(Rebel, 1908)" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[1055,1398,1824,1851]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
<emphasis id="B934F46814474E75B36F6B308097DB0E" box="[1055,1225,1825,1851]" italics="true" pageId="8" pageNumber="9">P. schawerdae</emphasis>
(Rebel, 1908)
</taxonomicName>
<taxonomicNameLabel id="A207491314474E75B7CD6B558563DB6B" box="[189,317,1860,1886]" pageId="8" pageNumber="9" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="B934F46814474E75B6326B558527DB6B" bold="true" box="[322,377,1860,1886]" pageId="8" pageNumber="9">A–B</emphasis>
. Bulgaria, Rila Mts.
<emphasis id="B934F46814474E75B51B6B5586DEDB6B" bold="true" box="[619,640,1860,1886]" pageId="8" pageNumber="9">C</emphasis>
. Republic of Macedonia, Korab Mts. –
<emphasis id="B934F46814474E75B3396B5580DEDB6B" bold="true" box="[1097,1152,1860,1886]" pageId="8" pageNumber="9">D–E</emphasis>
.
<taxonomicName id="4C4053F914474E75B3FD6B54814EDB6B" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1165,1296,1860,1886]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
<emphasis id="B934F46814474E75B3FD6B54814EDB6B" box="[1165,1296,1860,1886]" italics="true" pageId="8" pageNumber="9">P. alpinana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314474E75B2666B55812CDB6B" box="[1302,1394,1860,1886]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, France, Alpes Maritimes.
<emphasis id="B934F46814474E75B6826B768659DBB4" bold="true" box="[498,519,1895,1921]" pageId="8" pageNumber="9">D</emphasis>
. Holotype.
<emphasis id="B934F46814474E75B5E46B7686F6DBB4" bold="true" box="[660,680,1895,1921]" pageId="8" pageNumber="9">E</emphasis>
. Paratype. –
<emphasis id="B934F46814474E75B4376B768720DBB4" bold="true" box="[839,894,1895,1921]" pageId="8" pageNumber="9">F–G</emphasis>
.
<taxonomicName id="4C4053F914474E75B4FE6B798066DBB4" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[910,1080,1896,1921]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814474E75B4FE6B798066DBB4" box="[910,1080,1896,1921]" italics="true" pageId="8" pageNumber="9">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314474E75B3306B7680C1DBB4" box="[1088,1183,1895,1921]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, Italy, Gran Sasso National Park.
<emphasis id="B934F46814474E75B61C6B9B8520DB91" bold="true" box="[364,382,1930,1956]" pageId="8" pageNumber="9">F</emphasis>
. Paratype.
<emphasis id="B934F46814474E75B5716B9B8646DB91" bold="true" box="[513,536,1930,1956]" pageId="8" pageNumber="9">G</emphasis>
. Holotype. –
<emphasis id="B934F46814474E75B5C66B9B8693DB91" bold="true" box="[694,717,1930,1956]" pageId="8" pageNumber="9">H</emphasis>
.
<taxonomicName id="4C4053F914474E75B5AB6B9A807EDB90" authority="(Guenee, 1845)" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[731,1056,1930,1957]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="frigidana" status="stat. nov.">
<emphasis id="B934F46814474E75B5AB6B9A873ADB91" box="[731,868,1930,1956]" italics="true" pageId="8" pageNumber="9">P. frigidana</emphasis>
(Guenée, 1845)
</taxonomicName>
<taxonomicNameLabel id="A207491314474E75B3576B9A80D2DB90" box="[1063,1164,1931,1957]" pageId="8" pageNumber="9">stat. rev.</taxonomicNameLabel>
, Andorra, Pyrenees, neotype of
<taxonomicName id="4C4053F914474E75B6326BBC8668DBF2" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[322,566,1965,1991]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814474E75B6326BBC8668DBF2" box="[322,566,1965,1991]" italics="true" pageId="8" pageNumber="9">Eupoecilia frigidana</emphasis>
</taxonomicName>
. –
<emphasis id="B934F46814474E75B52C6BBF86DBDBFD" bold="true" box="[604,645,1966,1992]" pageId="8" pageNumber="9">I–J</emphasis>
.
<taxonomicName id="4C4053F914474E75B5E46BBF871EDBF2" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[660,832,1965,1991]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814474E75B5E46BBF871EDBF2" box="[660,832,1965,1991]" italics="true" pageId="8" pageNumber="9">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314474E75B4396BBF87F8DBFD" box="[841,934,1966,1992]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, Spain, Picos de Europa National Park.
<emphasis id="B934F46814474E75B7CD6BC08496DBDE" bold="true" box="[189,200,2001,2027]" pageId="8" pageNumber="9">I</emphasis>
. Holotype.
<emphasis id="B934F46814474E75B6236BC0853CDBDE" bold="true" box="[339,354,2001,2027]" pageId="8" pageNumber="9">J</emphasis>
. Paratype. Scale bar = 100 µm, all to scale.
</paragraph>
</caption>
<caption id="DF3F78F214464E74B7CD6B51869CDBF1" ID-DOI="http://doi.org/10.5281/zenodo.3838422" ID-Zenodo-Dep="3838422" httpUri="https://zenodo.org/record/3838422/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1856,1882]" targetBox="[199,1390,280,1807]" targetPageId="9">
<paragraph id="8BFF287A14464E74B7CD6B51869CDBF1" blockId="9.[189,1399,1856,1988]" pageId="9" pageNumber="10">
<emphasis id="B934F46814464E74B7CD6B518552DB6F" bold="true" box="[189,268,1856,1882]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
Phalli with vesica everted,
<taxonomicName id="4C4053F914464E74B5286B518070DB6F" authority="(Rebel, 1908)" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[600,1070,1856,1882]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
<emphasis id="B934F46814464E74B5286B5187DCDB6F" box="[600,898,1856,1882]" italics="true" pageId="9" pageNumber="10">Phtheochroa schawerdae</emphasis>
(Rebel, 1908)
</taxonomicName>
<taxonomicNameLabel id="A207491314464E74B3476B5180E4DB6E" box="[1079,1210,1856,1883]" pageId="9" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="B934F46814464E74B3B26B5180A7DB6F" bold="true" box="[1218,1273,1856,1882]" pageId="9" pageNumber="10">A–B</emphasis>
. Bulgaria, Rila Mts. –
<emphasis id="B934F46814464E74B64D6B728528DB48" bold="true" box="[317,374,1891,1917]" pageId="9" pageNumber="10">C–D</emphasis>
. Republic of Macedonia, Korab Mts.A, C: left.B, D: dorsal.Abbreviations:gs = gonopore sclerotization; ld = left diverticulum; rd = right diverticulum; svd = small ventral diverticulum; vpp = ventral phallic process. Scale bar = 250 µm.
</paragraph>
</caption>
<paragraph id="8BFF287A14454E77B7CD6D8A8524DD80" blockId="10.[189,378,411,437]" box="[189,378,411,437]" pageId="10" pageNumber="11">ABDOMEN. Grey.</paragraph>
<paragraph id="8BFF287A14454E77B7CD6DF5866DDF6B" blockId="10.[189,1399,484,862]" pageId="10" pageNumber="11">
GENITALIA (
<figureCitation id="137B34FF14454E77B6326DF585FFDDCB" box="[322,417,484,510]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
). Curvature of valva prominent. Rest of valva, uncus and socii resembles those in
<taxonomicName id="4C4053F914454E77B7CD6E198539DE14" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[189,359,519,545]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814454E77B7CD6E198539DE14" box="[189,359,519,545]" italics="true" pageId="10" pageNumber="11">P. schawerdae</emphasis>
</taxonomicName>
. Transtilla (
<figureCitation id="137B34FF14454E77B68A6E168608DE14" box="[506,598,519,545]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="10" pageNumber="11">Fig. 3C</figureCitation>
) wide, rectangular apically, with shallow median incision. Phallus (
<figureCitation id="137B34FF14454E77B7B66E3B8555DE70" box="[198,267,554,581]" captionStart="Fig" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[262,1281,269,1845]" captionTargetId="figure@12.[253,1334,265,1851]" captionTargetPageId="12" captionText="Fig. 6. Phalli with vesica everted of Phtheochroa spp. A–B. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – C–D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype.A, C: left. B, D: dorsal. Abbreviations: dd = dorsal diverticulum; vd = ventral diverticulum; other abbreviations as in Fig. 5. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838424" httpUri="https://zenodo.org/record/3838424/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
A–B) curved ventrally, with long, slender medioventral process (
<figureCitation id="137B34FF14454E77B37D6E3B800ADE71" box="[1037,1108,554,580]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
D–E), with apex slightly curved to the right. Vesica with subspherical proximal part and large part protruded from the right, bearing two large diverticula ending with cornuti. A furrow divides the two parts. Gonopore sunk in dorsal portion of furrow and surrounded by large sclerotized plate with uneven surface (distal part of ductus ejaculatorius). Right part of vesica protrudes in wider, conical, almost straight dorsal and narrower, cylindrical, curved ventral diverticulum. Cornuti aciculate and capitate, very robust, large, ventral one larger and slightly curved near apex. Proximal part of vesica bears a large conical ventral protrusion (diverticulum) with small processes. Acanthae present on posterior side of proximal part of vesica and at end of diverticula.
</paragraph>
<paragraph id="8BFF287A14454E77B79A6F9C85EBDF92" blockId="10.[234,437,909,935]" box="[234,437,909,935]" pageId="10" pageNumber="11">
<emphasis id="B934F46814454E77B79A6F9C851BDF92" bold="true" box="[234,325,909,935]" pageId="10" pageNumber="11">Female</emphasis>
(
<figureCitation id="137B34FF14454E77B6266F9C85F5DF92" box="[342,427,909,935]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="10" pageNumber="11">Fig. 1F</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14454E77B7CD6FAE8048D90C" blockId="10.[189,1399,959,1338]" pageId="10" pageNumber="11">
Considerably smaller than male. Head and thorax resemble those of male, but with forewing length
<quantity id="4CB8859F14454E77B7CD6FF38515DFC8" box="[189,331,994,1021]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="9.2" metricValueMin="7.2" pageId="10" pageNumber="11" unit="mm" value="8.2" valueMax="9.2" valueMin="7.2">7.2–9.2 mm</quantity>
(x= 7.9, n= 5), upperside with rust brown dorsal blotch and subterminal fascia. Hindwing upperside grey, underside white with scattered grey scales.Abdomen grey. Female genitalia (
<figureCitation id="137B34FF14454E77B3AE6814816BD815" box="[1246,1333,1029,1056]" captionStart="Fig" captionStartId="16.[189,232,1930,1956]" captionTargetBox="[188,1400,281,1861]" captionTargetId="figure@16.[180,1408,269,1871]" captionTargetPageId="16" captionText="Fig. 8. Female genitalia of Phtheochroa spp. A. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – B. P. alpinana sp. nov., France, Alpes Maritimes, paratype. Arrow: ventral diverticulum of ductus bursae. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838428" httpUri="https://zenodo.org/record/3838428/files/figure.png" pageId="10" pageNumber="11">Fig. 8B</figureCitation>
) with tergum 8 medially membranous, sterigma with two lateral protuberances, antrum nearly rectangular, with round anterior angles. Apophyses anteriores longer than apophyses posteriores. Ductus bursae ventrally membranous, with conical diverticulum of thick cuticle slightly curved to the left. Corpus bursae with large sclerite that starts on right side of ductus bursae with folds, then expands anteroventrally forming large ventral sclerotization and laterally to the left, then passes on dorsal side where it forms several folds on ductus bursae. Anterior medial part of corpus bursae with folded membranous area extending to the right. Ductus of accessory bursa emerges medioventrally, folded cuticle and small spines present to the left of its emerging area. Ductus seminalis inserted on ventral side.
</paragraph>
<paragraph id="8BFF287A14454E77B7CD6979861FD9B7" blockId="10.[189,577,1384,1410]" box="[189,577,1384,1410]" pageId="10" pageNumber="11">
<emphasis id="B934F46814454E77B7CD69798525D9B7" bold="true" box="[189,379,1384,1410]" pageId="10" pageNumber="11">
<heading id="D0B79F1614454E77B7CD69798563D9B7" box="[189,317,1384,1410]" pageId="10" pageNumber="11">Molecular</heading>
data
</emphasis>
(
<tableCitation id="C6C21DC114454E77B6FB697985BAD9B7" box="[395,484,1384,1410]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="10" pageNumber="11" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
,
<figureCitation id="137B34FF14454E77B68269798669D9B7" box="[498,567,1384,1410]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[190,1397,1281,1884]" captionTargetId="figure@18.[172,1412,1251,1902]" captionTargetPageId="18" captionText="Fig. 9. Maximum likelihood tree (built with MEGA6) of cytochrome c oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates." figureDoi="http://doi.org/10.5281/zenodo.3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14454E77B7CD698A80BDD9ED" blockId="10.[189,1399,1434,1496]" pageId="10" pageNumber="11">
BIN URI: BOLD:AAL5381. The intraspecific divergence of the barcode region is unknown (n =1). The minimum distance to the nearest neighbour,
<taxonomicName id="4C4053F914454E77B5B669AF8730D9ED" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[710,878,1470,1496]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814454E77B5B669AF8730D9ED" box="[710,878,1470,1496]" italics="true" pageId="10" pageNumber="11">P. schawerdae</emphasis>
</taxonomicName>
from the Dinaric Mts is 4.01%.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114454E77B7CD6A168579DB13" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="8BFF287A14454E77B7CD6A16857EDA14" blockId="10.[189,288,1543,1569]" box="[189,288,1543,1569]" pageId="10" pageNumber="11">
<heading id="D0B79F1614454E77B7CD6A16857EDA14" bold="true" box="[189,288,1543,1569]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B934F46814454E77B7CD6A16857EDA14" bold="true" box="[189,288,1543,1569]" pageId="10" pageNumber="11">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14454E77B7CD6A288579DB13" blockId="10.[189,1399,1593,1830]" pageId="10" pageNumber="11">
Preimaginal stages and larval host plant are unknown. The
<typeStatus id="54FB96D814454E77B4146A2887C9DA66" box="[868,919,1593,1619]" pageId="10" pageNumber="11">type</typeStatus>
material was collected in July at altitudes of about
<quantity id="4CB8859F14454E77B6546A4D8522DA42" box="[292,380,1628,1655]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="11" unit="m" value="2200.0">2200 m</quantity>
a.s.l. According to
<bibRefCitation id="EFD1558B14454E77B52D6A4D87E5DA43" author="Bassi G. &amp; Scaramozzino P. L." box="[605,955,1628,1654]" pageId="10" pageNumber="11" pagination="155 - 163" refId="ref11161" refString="Bassi G. &amp; Scaramozzino P. L. 1989. Contributo allo studio della microlepidotterofauna Piemontese. 1. Note sul alcuni interessante perperti di Tortricidae (Lepidoptera). Rivista Piemontese di Storia naturale 10: 155 - 163." type="journal article" year="1989">Bassi &amp; Scaramozzino (1989)</bibRefCitation>
the moths fly in two generations from April to July. However, material collected by these authors in the Italian Alps dates from mid- to late May and was collected at relatively low altitudes from
<quantity id="4CB8859F14454E77B4316AB287A4DA88" box="[833,1018,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.35" metricValueMin="1.2" pageId="10" pageNumber="11" unit="m" value="1275.0" valueMax="1350.0" valueMin="1200.0">
<date id="FFFE0EBA14454E77B4316AB28782DA88" box="[833,988,1698,1725]" pageId="10" pageNumber="11" value="1200" valueMax="1350">1200 to 1350</date>
m
</quantity>
a.s.l. From available data it thus seems more likely that the species is univoltine with a flight period from May to July, depending on climatic conditions and elevation. Habitat: the
<typeStatus id="54FB96D814454E77B5AE6AF8874FDB36" box="[734,785,1769,1795]" pageId="10" pageNumber="11">type</typeStatus>
material was collected in alpine grassland above the tree-line.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114454E77B7CD6B44806BDBF0" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BFF287A14454E77B7CD6B44859ADB5A" blockId="10.[189,452,1877,1903]" box="[189,452,1877,1903]" pageId="10" pageNumber="11">
<heading id="D0B79F1614454E77B7CD6B448508DB5A" box="[189,342,1877,1903]" pageId="10" pageNumber="11">
<emphasis id="B934F46814454E77B7CD6B448508DB5A" bold="true" box="[189,342,1877,1903]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
(
<figureCitation id="137B34FF14454E77B6166B4485E5DB5A" box="[358,443,1877,1903]" captionStart="Fig" captionStartId="20.[189,232,1990,2016]" captionTargetBox="[189,1398,1296,1948]" captionTargetId="figure@20.[183,1404,1290,1954]" captionTargetPageId="20" captionText="Fig. 10. Distribution of the Phtheochroa frigidana s. lat. species complex based on examined material." figureDoi="http://doi.org/10.5281/zenodo.3838432" httpUri="https://zenodo.org/record/3838432/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14454E77B7CD6B96806BDBF0" blockId="10.[189,1398,1927,1989]" pageId="10" pageNumber="11">
So far known only from the south-western Alps (
<collectingCountry id="F35768EA14454E77B4786B968702DB94" box="[776,860,1927,1953]" name="France" pageId="10" pageNumber="11">France</collectingCountry>
,
<collectingCountry id="F35768EA14454E77B41A6B9687FBDB97" box="[874,933,1927,1954]" name="Italy" pageId="10" pageNumber="11">Italy</collectingCountry>
). A record from the Slovenian Alps (as
<taxonomicName id="4C4053F914454E77B7CD6BBA8519DBF0" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[189,327,1963,1989]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814454E77B7CD6BBA8519DBF0" box="[189,327,1963,1989]" italics="true" pageId="10" pageNumber="11">P. frigidana</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFD1558B14454E77B6106BBA8676DBF0" author="Lesar T. &amp; Habeler H. &amp; Arenberger E." box="[352,552,1963,1989]" pageId="10" pageNumber="11" pagination="39 - 60" refId="ref11882" refString="Lesar T., Habeler H. &amp; Arenberger E. 2009. Prispevek k poznavanju metuljev (Lepidoptera) Slovenije II: nove vrste metuljckov (Microlepidoptera). Natura Sloveniae 11 (2): 39 - 60." type="journal article" year="2009">
Lesar
<emphasis id="B934F46814454E77B6DA6BBA8580DBF0" box="[426,478,1963,1989]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 2009
</bibRefCitation>
) is doubtful and may refer to
<taxonomicName id="4C4053F914454E77B4F66BBA8070DBF0" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[902,1070,1963,1989]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814454E77B4F66BBA8070DBF0" box="[902,1070,1963,1989]" italics="true" pageId="10" pageNumber="11">P. schawerdae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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